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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.252.3588" ID-GBIF-Dataset="a4366e50-a271-44c0-bdef-b52c8cad0b85" ID-PMC="PMC3560839" ID-Pensoft-Pub="1313-2970-252-1" ID-PubMed="23378811" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-252-1" ModsDocOrigin="ZooKeys 252" ModsDocTitle="Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)" checkinTime="1451247847790" checkinUser="pensoft" docAuthor="Bond, Jason E." docDate="2012" docId="70D6B31BA35C6B80240A7900C8AB9055" docLanguage="en" docName="ZooKeys 252: 1-209" docOrigin="ZooKeys 252" docSource="http://dx.doi.org/10.3897/zookeys.252.3588" docTitle="Aptostichus sarlacc Bond, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="122" masterDocId="FFAD871BFFD4FF8CFFED1A5CFFE7FFA0" masterDocTitle="Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)" masterLastPageNumber="209" masterPageNumber="1" pageNumber="119" updateTime="1668155039594" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bond, Jason E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>252</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>209</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.252.3588</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.252.3588</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-252-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152039290" LSID="urn:lsid:zoobank.org:act:2FCA4B5E-04D4-4D23-A829-617B0DCCFDA5" httpUri="http://treatment.plazi.org/id/70D6B31BA35C6B80240A7900C8AB9055" lastPageId="121" lastPageNumber="122" pageId="118" pageNumber="119">
<subSubSection pageId="118" pageNumber="119" type="nomenclature">
<paragraph pageId="118" pageNumber="119">
<taxonomicName LSID="The Sarlacc Trapdoor Spider" class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sarlacc" order="Araneae" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="sarlacc">Aptostichus sarlacc</taxonomicName>
<taxonomicNameLabel pageId="118" pageNumber="119">sp. n.</taxonomicNameLabel>
Figures 236-242Map 29
</paragraph>
</subSubSection>
<subSubSection pageId="118" pageNumber="119" type="types">
<paragraph pageId="118" pageNumber="119">Types.</paragraph>
<paragraph pageId="118" pageNumber="119">Male holotype (AP417) from California, San Bernardino County, 14.5km N, 16km E of Ridgecrest sand dunes, 35.7553, -117.5006 4, 960m, coll. D. Giuliani 15.ii.1981-12.iv.1981; male paratype (AP416) from California, Kern County, 11.3km, N 9.6km W of Inyokern, 35.7362, -117.9849 4, 2193m, coll. D. Giuliani 4.iv.1986; male paratype from California, Inyo County, Owens Valley, 9.6km S-SW of Independence, 36.7414, -118.2623 5, 1961m, coll. D. Giuliani 1.xi.1986-12.vi.1987. Deposited in CAS.</paragraph>
</subSubSection>
<subSubSection pageId="118" pageNumber="119" type="etymology">
<paragraph pageId="118" pageNumber="119">Etymology.</paragraph>
<paragraph pageId="118" pageNumber="119">
The specific epithet is a noun in apposition taken from the fictional creature in George
<normalizedToken originalValue="Lucas">Lucas'</normalizedToken>
science fiction saga, Star Wars: Return of the Jedi.
</paragraph>
</subSubSection>
<subSubSection lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119" type="diagnosis">
<paragraph pageId="118" pageNumber="119">Diagnosis.</paragraph>
<paragraph lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119">
Males can be distinguished from other known closely related species of
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus" order="Araneae" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">Aptostichus</taxonomicName>
(e.g.,
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus cahuilla" order="Araneae" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="cahuilla">Aptostichus cahuilla</taxonomicName>
,
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus aguacaliente" order="Araneae" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="aguacaliente">Aptostichus aguacaliente</taxonomicName>
) by having a long curved metatarsus IV
<pageBreakToken pageId="119" pageNumber="120" start="start">relative</pageBreakToken>
to femur IV length and by having an abdomen devoid of any dorsal markings (Fig. 236). This species can be distinguished from geographical proximate members of the
<taxonomicName class="Insecta" family="Staphylinidae" genus="Simus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simus" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="genus">Simus</taxonomicName>
species group, with similarly long fourth tarsi and by having a longer,
<pageBreakToken pageId="120" pageNumber="121" start="start">more</pageBreakToken>
slender palpal tibia (Figs 241, 242). The distribution of
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sarlacc" order="Araneae" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="sarlacc">Aptostichus sarlacc</taxonomicName>
is distributed considerably further to the north of the aforementioned
<taxonomicName class="Insecta" family="Staphylinidae" genus="Hesperus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesperus" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">Hesperus</taxonomicName>
species group taxa.
</paragraph>
</subSubSection>
<subSubSection lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121" type="description">
<paragraph pageId="120" pageNumber="121">Description of male holotype.</paragraph>
<paragraph pageId="120" pageNumber="121">Specimen preparation and condition. Specimen collected dead from pitfall trap, preserved 70% EtOH. Coloration likely faded. Pedipalp, leg I left side removed, stored in vial with specimen. General coloration. Carapace, chelicerae, legs strong brown 7.5YR 4/6. Abdomen uniform very pale brown, lacking distinct dorsal markings (e.g., paratype coloration pattern, Fig. 236). Cephalothorax. Carapace 3.41 long, 2.95 wide, glabrous with only sparse thin black setae, stout black bristles along fringe; surface smooth, pars cephalica elevated. Fringe, posterior margin with black bristles. Foveal groove deep, moderately procurved. Eyes on low mound. AER slightly procurved, PER slightly recurved. PME, AME subequal diameter. Sternum moderately setose, STRl 1.90, STRw 1.62. Posterior sternal sigilla small, positioned towards lateral margin, anterior sigilla pairs small, oval, marginal. Chelicerae with distinct anterior tooth row comprising 5 teeth, posterior margin with patch of very small denticles. Palpal endites with patch of small cuspules on proximal, inner margin, labium with 3 small cuspules, LBw 0.60, LBl 0.29. Rastellum consists of 6 stout spines, 2 offset prolaterally. Abdomen. Setose, heavy black setae intermingled with fine black setae. Legs. Leg I: 3.60, 2.40, 2.14, 1.67, 1.12; leg IV: 3.44, 2.16. Light tarsal scopulae on tarsi legs I, II. Tarsus I with single, slightly staggered row of 13 trichobothria. Leg I spination pattern illustrated in Figures 239, 240; TSp 4, TSr 4, TSrd 3. Pedipalp. Articles slender, lacking distinct spines (Figs 241, 242). PTw 0.48, PTl 1.40, Bl 0.70. Embolus slender, curved at midpoint, with slight curve distally, lacking serrations (Figs 241, 242).</paragraph>
<paragraph pageId="121" pageNumber="122">
<pageBreakToken pageId="121" pageNumber="122" start="start">Variation</pageBreakToken>
(2). Cl 3.41-4.85, Cw 2.95-4.28, STRl 1.9-2.45, STRw 1.62-2.19, LBw 0.60-0.85, LBl 0.29-0.46, leg I: 3.60-5.00, 2.40-3.19, 2.14-2.97, 1.67-2.25, 1.12-1.53; leg IV: 3.44-4.75, 2.16-2.88; PTl 1.40-1.98, PTw 0.48-0.63, Bl 0.70-1.01, TSp 4-4, TSr 3-4, TSrd 3-3.
</paragraph>
<caption pageId="121" pageNumber="122">
<paragraph pageId="121" pageNumber="122">
Figures 236-242.
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sarlacc" order="Araneae" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="sarlacc">Aptostichus sarlacc</taxonomicName>
sp. n.; scale bars = 1.0mm. 236-238 male paratype (AP416) from Kern Co. 236 habitus [806006] 237 retrolateral aspect, right leg I [806000] 238 prolateral aspect, right leg I [806004] 239-242 male holotype (AP417) 239 retrolateral aspect, leg I [805992] 240 prolateral aspect, leg I [805996] 241 retrolateral aspect, pedipalp [805998] 242 line drawings (in descending order) of holotype leg I metatarsus and tibia spination pattern (retro and prolateral views), pedipalp (retrolateral view), lower inset, line drawing of paratype TSrd spination pattern.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="121" pageNumber="122" type="description">
<paragraph pageId="121" pageNumber="122">Description of female.</paragraph>
<paragraph pageId="121" pageNumber="122">Known only from male specimens.</paragraph>
</subSubSection>
<subSubSection pageId="121" pageNumber="122" type="material examined">
<paragraph pageId="121" pageNumber="122">Material examined.</paragraph>
<paragraph pageId="121" pageNumber="122">Known only from the type material.</paragraph>
</subSubSection>
<subSubSection pageId="121" pageNumber="122" type="distribution">
<paragraph pageId="121" pageNumber="122">Distribution and natural history.</paragraph>
<paragraph pageId="121" pageNumber="122">Little is known about this species; it is very rare in collections and is only known from two specimens collected in the Mojave Desert in Inyo, Kern and San Bernardino Counties (Map 29).</paragraph>
<caption pageId="121" pageNumber="122">
<paragraph pageId="121" pageNumber="122">
Map 29. Distribution of
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sarlacc" order="Araneae" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="sarlacc">Aptostichus sarlacc</taxonomicName>
sp. n. (filled squares) and
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus derhamgiulianii" order="Araneae" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="derhamgiulianii">Aptostichus derhamgiulianii</taxonomicName>
sp. n. (filled circles).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="121" pageNumber="122" type="conservation status">
<paragraph pageId="121" pageNumber="122">Conservation status.</paragraph>
<paragraph pageId="121" pageNumber="122">This species is rare in collections and is known from only two localities; its status is considered imperiled.</paragraph>
</subSubSection>
<subSubSection pageId="121" pageNumber="122" type="species concept applied">
<paragraph pageId="121" pageNumber="122">Species concept applied.</paragraph>
<paragraph pageId="121" pageNumber="122">Morphological.</paragraph>
</subSubSection>
</treatment>
</document>