187 lines
20 KiB
XML
187 lines
20 KiB
XML
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<mods:title id="C4CB90E023BBBC1E66EAC4A0E59BFA20">The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region</mods:title>
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<mods:namePart id="FBF7823B7B99C0ACC29F7F9B28C4B4A9">Biranvand, Amir</mods:namePart>
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<mods:affiliation id="9193D43E3514FD95AD212845585EF888">Young Researchers and Elite Club, Khorramabad Branch, Islamic Azad University, Khorramabad, Iran</mods:affiliation>
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<mods:namePart id="182F9C3F7E9280E934B97D87E3E9EEA4">Nedvěd, Oldřich</mods:namePart>
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<mods:affiliation id="86D9DD1343E61F3D30844339B58A55FC">Faculty of Science, University of South Bohemia, CZ- 37005 České Budějovice, Czech Republic & Biology Centre, Czech Academy of Sciences, Institute of Entomology, CZ- 37005 České Budějovice, Czech Republic</mods:affiliation>
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<mods:namePart id="88126FDE6728AE8C89623934E96CF26A">Tomaszewska, Wioletta</mods:namePart>
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<mods:affiliation id="913D2E43F0FC2ED571325B41A02B4385">Museum and Institute of Zoology, Polish Academy of Sciences, PL- 00 - 679 Warsaw, Poland</mods:affiliation>
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<mods:namePart id="EA57931B8F556FF4CFE085C931DACFCE">Al Ansi, Amin N.</mods:namePart>
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<mods:affiliation id="96167FDCCDB9BFC6BA3C31D0BD380A49">King Saud University Museum of Arthropods, Department of Plant Protection, King Saud University, SA- 11451 Riyadh, Saudi Arabia</mods:affiliation>
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<mods:namePart id="64FEB59755B17B1D1C6B4546832069D5">Fekrat, Lida</mods:namePart>
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<mods:affiliation id="FB5F46421F12539F1A8995C3B33F1242">Department of Plant Protection, Ferdowsi University of Mashhad, Mashhad, Iran</mods:affiliation>
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<mods:namePart id="AB4E3AC1E4506AFDEF6A7236A8035A60">Haghghadam, Zahra Mojib</mods:namePart>
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<mods:affiliation id="1E02E678CB7C3FA058450BE6C232501A">Agricultural and Natural Resources Research Center of Gilan Province, Rasht, Iran</mods:affiliation>
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<mods:affiliation id="2EF1B2BEF514CB1F1D61C54BF7BE6AC1">Yazd Provincial Office, Department of Environment, Iran</mods:affiliation>
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<mods:affiliation id="46E771410AA3F2FC2721E49CA1E75D6B">Department of Plant Protection, Lorestan University, Iran</mods:affiliation>
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<mods:namePart id="A3626CF2C5661FB72A8C9F4033791C59">Şenal, Derya</mods:namePart>
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<mods:namePart id="AE8001A0AE909FEF1F2C40A0B90A26F1">Shakarami, Jahanshir</mods:namePart>
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<mods:affiliation id="CD2D13A34FFFD394A51C1FAC8DA81CC9">Department of Plant Protection, Lorestan University, Iran</mods:affiliation>
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<mods:namePart id="4E42B3A825DC80921F6BEB39AD7B4DD8">Haelewaters, Danny</mods:namePart>
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<mods:affiliation id="F37999FCB73D305A743EE36D2C7C3F87">Faculty of Science, University of South Bohemia, CZ- 37005 České Budějovice, Czech Republic</mods:affiliation>
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<mods:title id="5E4A1BF5B54BACA6BC5C849E88CE05F5">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="46743B137C14DA3D0D537F4D416A930A">2019</mods:date>
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<mods:number id="C9BFAC6E6D6AB1345AFC4B1E98C0020A">2019-04-17</mods:number>
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<treatment id="03F81C7CA446C5233B9BFDC1FAF1FAAD" ID-DOI="http://doi.org/10.5281/zenodo.4504998" ID-GBIF-Taxon="177430888" ID-Zenodo-Dep="4504998" LSID="urn:lsid:plazi:treatment:03F81C7CA446C5233B9BFDC1FAF1FAAD" httpUri="http://treatment.plazi.org/id/03F81C7CA446C5233B9BFDC1FAF1FAAD" lastPageNumber="164" pageId="1" pageNumber="164">
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<subSubSection id="C34BFEE1A446C5233B9BFDC1FB5EFDC2" box="[991,1273,589,613]" pageId="1" pageNumber="164" type="nomenclature">
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<paragraph id="8BEEAD6AA446C5233B9BFDC1FB5EFDC2" blockId="1.[991,1273,589,613]" box="[991,1273,589,613]" pageId="1" pageNumber="164">
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<heading id="D0A61A06A446C5233B9BFDC1FB5EFDC2" box="[991,1273,589,613]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="164" reason="3">
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<taxonomicName id="4C51D6E9A446C5233B9BFDC1FB5EFDC2" ID-CoL="4TR2" ID-ENA="115356" authority="Mulsant, 1846" authorityName="Mulsant" authorityYear="1846" box="[991,1273,589,613]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B9257178A446C5233B9BFDC1FB5EFDC2" bold="true" box="[991,1273,589,613]" pageId="1" pageNumber="164">
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<emphasis id="B9257178A446C5233B9BFDC1FBF6FDC3" bold="true" box="[991,1105,589,613]" italics="true" pageId="1" pageNumber="164">Harmonia</emphasis>
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Mulsant, 1846
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C34BFEE1A446C5233B78FDFFFB1DFD1D" pageId="1" pageNumber="164" type="reference_group">
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<paragraph id="8BEEAD6AA446C5233B78FDFFFB1DFD1D" blockId="1.[828,1437,627,699]" pageId="1" pageNumber="164">
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=
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<taxonomicName id="4C51D6E9A446C5233B0BFDFFFC5EFD20" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[847,1017,627,646]" class="Insecta" family="Coccinellidae" genus="Leis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B9257178A446C5233B0BFDFFFCD4FD20" box="[847,883,627,646]" italics="true" pageId="1" pageNumber="164">Leis</emphasis>
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Mulsant, 1850
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</taxonomicName>
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;
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<taxonomicName id="4C51D6E9A446C5233C40FDFFFB18FD20" authority="Mulsant, 1853" authorityName="Mulsant" authorityYear="1853" box="[1028,1215,627,646]" class="Insecta" family="Coccinellidae" genus="Ballia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B9257178A446C5233C40FDFFFB9FFD20" box="[1028,1080,627,646]" italics="true" pageId="1" pageNumber="164">Ballia</emphasis>
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Mulsant, 1853
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</taxonomicName>
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;
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<taxonomicName id="4C51D6E9A446C5233C8EFDFFFA3EFD20" authority="Crotch, 1871" authorityName="Crotch" authorityYear="1871" box="[1226,1433,627,646]" class="Insecta" family="Coccinellidae" genus="Callineda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B9257178A446C5233C8EFDFFFAB9FD20" box="[1226,1310,627,646]" italics="true" pageId="1" pageNumber="164">Callineda</emphasis>
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Crotch, 1871
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</taxonomicName>
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;
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<taxonomicName id="4C51D6E9A446C5233B1EFD02FB8DFD07" authority="Crotch, 1874" authorityName="Crotch" authorityYear="1874" box="[858,1066,654,673]" class="Insecta" family="Coccinellidae" genus="Stictoleis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B9257178A446C5233B1EFD02FC0DFD07" box="[858,938,654,673]" italics="true" pageId="1" pageNumber="164">Stictoleis</emphasis>
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Crotch, 1874
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</taxonomicName>
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||
;
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||
<taxonomicName id="4C51D6E9A446C5233C72FD02FABBFD07" authority="Crotch, 1874" authorityName="Crotch" authorityYear="1874" box="[1078,1308,654,673]" class="Insecta" family="Coccinellidae" genus="Ptychanatis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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<emphasis id="B9257178A446C5233C72FD02FB3AFD07" box="[1078,1181,654,673]" italics="true" pageId="1" pageNumber="164">Ptychanatis</emphasis>
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Crotch, 1874
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</taxonomicName>
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||
;
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<taxonomicName id="4C51D6E9A446C5233D6CFD02FC56FD1D" authority="Timberlake, 1943" authorityName="Timberlake" authorityYear="1943" class="Insecta" family="Coccinellidae" genus="Rhopaloneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9257178A446C5233D6CFD02FA3BFD07" box="[1320,1436,654,673]" italics="true" pageId="1" pageNumber="164">Rhopaloneda</emphasis>
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||
Timberlake, 1943
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</taxonomicName>
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||
(
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<emphasis id="B9257178A446C5233BB9FD24FB8BFD1D" box="[1021,1068,680,699]" italics="true" pageId="1" pageNumber="164">sensu</emphasis>
|
||
<bibRefCitation id="EFC0D09BA446C5233C76FD24FB08FD1D" author="SLIPINSKI A." bookContentInfo="286 pp." box="[1074,1199,680,699]" journalOrPublisher="Australian Biological Resources Study, Canberra" pageId="1" pageNumber="164" refId="ref8230" refString="SLIPINSKI A. 2007: Australian ladybird beetles (Coleoptera: Coccinellidae): their biology and classification. Australian Biological Resources Study, Canberra, 286 pp." title="Australian ladybird beetles (Coleoptera: Coccinellidae): their biology and classification" type="book" year="2007">
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<subScript id="17D5AF2FA446C5233C76FD24FBDBFD1C" attach="right" box="[1074,1148,680,699]" fontSize="6" pageId="1" pageNumber="164">ŚLIPIŃSKI</subScript>
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||
2007
|
||
</bibRefCitation>
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).
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C34BFEE1A446C5233B78FD58FAF1FAAD" pageId="1" pageNumber="164" type="diagnosis">
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<paragraph id="8BEEAD6AA446C5233B78FD58FAF1FAAD" blockId="1.[828,1437,724,1291]" pageId="1" pageNumber="164">
|
||
<emphasis id="B9257178A446C5233B78FD58FC0AFD4D" bold="true" box="[828,941,724,747]" pageId="1" pageNumber="164">Diagnosis.</emphasis>
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Length
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<quantity id="4CA9008FA446C5233C41FD58FBF8FD4D" box="[1029,1119,724,747]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="1" pageNumber="164" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
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. Elytra moderately to strongly convex, glabrous; elytral color pattern variable. Clypeus anteriorly straight between lateral projections. Antenna composed of 11 antennomeres, slightly shorter than the width of head capsule, with club moderately compact, composed of three antennomeres (
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<figureCitation id="136AB1EFA446C5233CE5FCF8FB46FC2D" box="[1185,1249,884,907]" captionStart="Figs 6–13" captionStartId="3.[151,188,1252,1271]" captionTargetBox="[314,1274,195,1221]" captionTargetId="figure-501@3.[292,1296,189,1241]" captionTargetPageId="3" captionText="Figs 6–13. Morphological characters of Harmonia species: 6 – H. axyridis (Pallas, 1773), prosternum. 7–13 – Harmonia quadripunctata (Pontoppidan, 1763): 7 – prosternum; 8 – hind leg; 9 – antenna; 10 – meso- and metaventrites; 11 – maxillary palp; 12 – abdominal ventrite 1 with postcoxal lines; 13 – mandible." figureDoi="http://doi.org/10.5281/zenodo.4552757" httpUri="https://zenodo.org/record/4552757/files/figure.png" pageId="1" pageNumber="164">Fig. 9</figureCitation>
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). Terminal maxillary palpomere securiform (
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<figureCitation id="136AB1EFA446C5233C27FC18FB17FC0D" box="[1123,1200,916,939]" captionStart="Figs 6–13" captionStartId="3.[151,188,1252,1271]" captionTargetBox="[314,1274,195,1221]" captionTargetId="figure-501@3.[292,1296,189,1241]" captionTargetPageId="3" captionText="Figs 6–13. Morphological characters of Harmonia species: 6 – H. axyridis (Pallas, 1773), prosternum. 7–13 – Harmonia quadripunctata (Pontoppidan, 1763): 7 – prosternum; 8 – hind leg; 9 – antenna; 10 – meso- and metaventrites; 11 – maxillary palp; 12 – abdominal ventrite 1 with postcoxal lines; 13 – mandible." figureDoi="http://doi.org/10.5281/zenodo.4552757" httpUri="https://zenodo.org/record/4552757/files/figure.png" pageId="1" pageNumber="164">Fig. 11</figureCitation>
|
||
). Pronotal disc evenly convex, transverse with narrowly upturned and/or slightly thickenedexternal borders. Prothoracichypomeronwithout fovea near anterior angles; prosternal process without or withdistinctlateral carinae extendinganteriorly (
|
||
<figureCitation id="136AB1EFA446C5233D77FB98FA36FB8D" box="[1331,1425,1044,1067]" captionStart="Figs 6–13" captionStartId="3.[151,188,1252,1271]" captionTargetBox="[314,1274,195,1221]" captionTargetId="figure-501@3.[292,1296,189,1241]" captionTargetPageId="3" captionText="Figs 6–13. Morphological characters of Harmonia species: 6 – H. axyridis (Pallas, 1773), prosternum. 7–13 – Harmonia quadripunctata (Pontoppidan, 1763): 7 – prosternum; 8 – hind leg; 9 – antenna; 10 – meso- and metaventrites; 11 – maxillary palp; 12 – abdominal ventrite 1 with postcoxal lines; 13 – mandible." figureDoi="http://doi.org/10.5281/zenodo.4552757" httpUri="https://zenodo.org/record/4552757/files/figure.png" pageId="1" pageNumber="164">Figs 6–7</figureCitation>
|
||
). Anteriormarginof mesoventriteweaklyfoveate mediallyto straight. Elytral lateral marginsnarrowly raised; epipleuron not foveate. Apices of middle and hindtibiae without spurs (
|
||
<figureCitation id="136AB1EFA446C5233B07FB18FCD8FB0D" box="[835,895,1172,1195]" captionStart="Figs 6–13" captionStartId="3.[151,188,1252,1271]" captionTargetBox="[314,1274,195,1221]" captionTargetId="figure-501@3.[292,1296,189,1241]" captionTargetPageId="3" captionText="Figs 6–13. Morphological characters of Harmonia species: 6 – H. axyridis (Pallas, 1773), prosternum. 7–13 – Harmonia quadripunctata (Pontoppidan, 1763): 7 – prosternum; 8 – hind leg; 9 – antenna; 10 – meso- and metaventrites; 11 – maxillary palp; 12 – abdominal ventrite 1 with postcoxal lines; 13 – mandible." figureDoi="http://doi.org/10.5281/zenodo.4552757" httpUri="https://zenodo.org/record/4552757/files/figure.png" pageId="1" pageNumber="164">Fig. 8</figureCitation>
|
||
). Abdominalpostcoxallinesnot recurved, incomplete laterally (
|
||
<figureCitation id="136AB1EFA446C5233BDAFB38FC4DFB6D" box="[926,1002,1204,1227]" captionStart="Figs 6–13" captionStartId="3.[151,188,1252,1271]" captionTargetBox="[314,1274,195,1221]" captionTargetId="figure-501@3.[292,1296,189,1241]" captionTargetPageId="3" captionText="Figs 6–13. Morphological characters of Harmonia species: 6 – H. axyridis (Pallas, 1773), prosternum. 7–13 – Harmonia quadripunctata (Pontoppidan, 1763): 7 – prosternum; 8 – hind leg; 9 – antenna; 10 – meso- and metaventrites; 11 – maxillary palp; 12 – abdominal ventrite 1 with postcoxal lines; 13 – mandible." figureDoi="http://doi.org/10.5281/zenodo.4552757" httpUri="https://zenodo.org/record/4552757/files/figure.png" pageId="1" pageNumber="164">Fig. 12</figureCitation>
|
||
). Penis stout, consists of basal sclerite and additionalapical piece. Ovipositor withcoxitesclub-handle
|
||
<typeStatus id="54EA13C8A446C5233B78FB78FCCBFAAD" box="[828,876,1268,1291]" pageId="1" pageNumber="164">type</typeStatus>
|
||
; infundibulum present; spermatheca C-shaped.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |