The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region Author Biranvand, Amir Young Researchers and Elite Club, Khorramabad Branch, Islamic Azad University, Khorramabad, Iran Author Nedvěd, Oldřich Faculty of Science, University of South Bohemia, CZ- 37005 České Budějovice, Czech Republic & Biology Centre, Czech Academy of Sciences, Institute of Entomology, CZ- 37005 České Budějovice, Czech Republic Author Tomaszewska, Wioletta Museum and Institute of Zoology, Polish Academy of Sciences, PL- 00 - 679 Warsaw, Poland Author Al Ansi, Amin N. King Saud University Museum of Arthropods, Department of Plant Protection, King Saud University, SA- 11451 Riyadh, Saudi Arabia Author Fekrat, Lida Department of Plant Protection, Ferdowsi University of Mashhad, Mashhad, Iran Author Haghghadam, Zahra Mojib Agricultural and Natural Resources Research Center of Gilan Province, Rasht, Iran Author Khormizi, Mehdi Zare Yazd Provincial Office, Department of Environment, Iran Author Noorinahad, Sara Department of Plant Protection, Lorestan University, Iran Author Şenal, Derya Author Shakarami, Jahanshir Department of Plant Protection, Lorestan University, Iran Author Haelewaters, Danny Faculty of Science, University of South Bohemia, CZ- 37005 České Budějovice, Czech Republic text Acta Entomologica Musei Nationalis Pragae 2019 2019-04-17 59 1 163 170 journal article 8093 10.2478/aemnp-2019-0014 02d54075-6c21-4d05-b8f8-40828a70b492 1804-6487 4552749 605E5337-395B-4C88-AC75-E761330518FE Harmonia Mulsant, 1846 = Leis Mulsant, 1850 ; Ballia Mulsant, 1853 ; Callineda Crotch, 1871 ; Stictoleis Crotch, 1874 ; Ptychanatis Crotch, 1874 ; Rhopaloneda Timberlake, 1943 ( sensu ŚLIPIŃSKI 2007 ). Diagnosis. Length 4–8 mm . Elytra moderately to strongly convex, glabrous; elytral color pattern variable. Clypeus anteriorly straight between lateral projections. Antenna composed of 11 antennomeres, slightly shorter than the width of head capsule, with club moderately compact, composed of three antennomeres ( Fig. 9 ). Terminal maxillary palpomere securiform ( Fig. 11 ). Pronotal disc evenly convex, transverse with narrowly upturned and/or slightly thickenedexternal borders. Prothoracichypomeronwithout fovea near anterior angles; prosternal process without or withdistinctlateral carinae extendinganteriorly ( Figs 6–7 ). Anteriormarginof mesoventriteweaklyfoveate mediallyto straight. Elytral lateral marginsnarrowly raised; epipleuron not foveate. Apices of middle and hindtibiae without spurs ( Fig. 8 ). Abdominalpostcoxallinesnot recurved, incomplete laterally ( Fig. 12 ). Penis stout, consists of basal sclerite and additionalapical piece. Ovipositor withcoxitesclub-handle type ; infundibulum present; spermatheca C-shaped.