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<document id="05E8F889B7BE885E098B2C60CB3D97F1" ID-CLB-Dataset="28026" ID-DOI="10.11646/zootaxa.4512.1.1" ID-GBIF-Dataset="26e12e1b-2175-4007-871b-d0b73122e5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607575" ID-ZooBank="E752FB7B-F34C-4D12-B8A2-EA6C791DD6C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586644813" checkinUser="plazi" docAuthor="Kaiser, Christine M., Kaiser, Hinrich &amp; OShea, Mark" docDate="2018" docId="C80EBE29FFEAFFD8FF75FA6E0CCDF81A" docLanguage="en" docName="zootaxa.4512.1.1.pdf" docOrigin="Zootaxa 4512 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Stegonotus florensis" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="3437C651FFC7FFF6FFE2FE470F12FFFE" masterDocTitle="The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="46" updateTime="1698713152511" updateUser="ExternalLinkService">
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<mods:title id="E84BA789BFFB3D0E21E55D57B5909ED0">The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874)</mods:title>
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<mods:namePart id="2B570E3318D3B83CCF6063D4367E5848">Kaiser, Christine M.</mods:namePart>
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<mods:namePart id="86982E1AC8F41ADE2DDEFF9E1883B749">Kaiser, Hinrich</mods:namePart>
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<treatment id="C80EBE29FFEAFFD8FF75FA6E0CCDF81A" ID-DOI="http://doi.org/10.5281/zenodo.5997411" ID-GBIF-Taxon="154496323" ID-Zenodo-Dep="5997411" LSID="urn:lsid:plazi:treatment:C80EBE29FFEAFFD8FF75FA6E0CCDF81A" httpUri="http://treatment.plazi.org/id/C80EBE29FFEAFFD8FF75FA6E0CCDF81A" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<paragraph id="40180F3FFFEAFFDBFF75FA6E0D99FBBA" blockId="45.[151,651,1065,1092]" box="[151,651,1065,1092]" pageId="45" pageNumber="46">
<heading id="1B50B853FFEAFFDBFF75FA6E0D99FBBA" bold="true" box="[151,651,1065,1092]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<taxonomicName id="87A774BCFFEAFFDBFF75FA6E0D99FBBA" authority="(de Rooij, 1917: 110)" authorityPageNumber="110" baseAuthorityName="de Rooij" baseAuthorityPageNumber="110" baseAuthorityYear="1917" box="[151,651,1065,1092]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFEAFFDBFF75FA6E0D99FBBA" bold="true" box="[151,651,1065,1092]" pageId="45" pageNumber="46">
<emphasis id="72D3D32DFFEAFFDBFF75FA6E0E98FBBD" bold="true" box="[151,394,1065,1091]" italics="true" pageId="45" pageNumber="46">Stegonotus florensis</emphasis>
(de
<treatmentCitation id="C106292EFFEAFFDBFE22FA6D0D91FBBA" author="de Rooij, N." box="[448,643,1066,1092]" page="110" pageId="45" pageNumber="46" year="1917">
<bibRefCitation id="243672CEFFEAFFDBFE22FA6D0D42FBBA" author="de Rooij, N." box="[448,592,1066,1092]" pageId="45" pageNumber="46" refId="ref43040" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. Volume II. Ophidia. E. J. Brill, Leiden, The Netherlands, 334 pp. https: // doi. org / 10.5962 / bhl. title. 24239" type="book" year="1917">Rooij, 1917</bibRefCitation>
:110
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)
</emphasis>
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</heading>
</paragraph>
<paragraph id="40180F3FFFEAFFDBFF75FA370EAAFB77" blockId="45.[151,1437,1136,1953]" box="[151,440,1136,1161]" pageId="45" pageNumber="46">
<emphasis id="72D3D32DFFEAFFDBFF75FA370E7CFB77" bold="true" box="[151,366,1136,1161]" pageId="45" pageNumber="46">Taxonomic status.</emphasis>
Valid.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFEAFFDBFF25FAD30E91F9C7" pageId="45" pageNumber="46" type="description">
<paragraph id="40180F3FFFEAFFDBFF25FAD30DAFFB53" blockId="45.[151,1437,1136,1953]" box="[199,701,1172,1197]" pageId="45" pageNumber="46">
<emphasis id="72D3D32DFFEAFFDBFF25FAD30E5BFB53" bold="true" box="[199,329,1172,1197]" pageId="45" pageNumber="46">Synonyms.</emphasis>
<taxonomicName id="87A774BCFFEAFFDBFEB2FAD20DA4FB53" authority="Forcart, 1953" authorityName="Forcart" authorityYear="1953" box="[336,694,1172,1197]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="sutteri">
<emphasis id="72D3D32DFFEAFFDBFEB2FAD20D0BFB52" box="[336,537,1173,1196]" italics="true" pageId="45" pageNumber="46">Stegonotus sutteri</emphasis>
<bibRefCitation id="243672CEFFEAFFDBFDFDFAD20DA4FB53" author="Forcart, L." box="[543,694,1172,1197]" pageId="45" pageNumber="46" pagination="356 - 388" refId="ref43614" refString="Forcart, L. (1953) Die Amphibien und Reptilien von Sumba, ihre zoogeographischen Beziehungen und Revision der Unterarten von Typhlops polygrammicus. Ferhandlungen der naturforschenden Gesellschaft zu Basel, 64, 356 - 388." type="journal article" year="1953">Forcart, 1953</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="40180F3FFFEAFFDBFF25FAFF0E91F9C7" blockId="45.[151,1437,1136,1953]" pageId="45" pageNumber="46">
<emphasis id="72D3D32DFFEAFFDBFF25FAFF0E6EFB2F" bold="true" box="[199,380,1208,1233]" pageId="45" pageNumber="46">Original name.</emphasis>
<taxonomicName id="87A774BCFFEAFFDBFE61FAFF0C3EFB2F" authority="de Rooij, 1917: 110" authorityName="de Rooij" authorityPageNumber="110" authorityYear="1917" box="[387,812,1208,1233]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFEAFFDBFE61FAFF0D5BFB2F" box="[387,585,1208,1233]" italics="true" pageId="45" pageNumber="46">Lycodon florensis</emphasis>
de
<bibRefCitation id="243672CEFFEAFFDBFD91FAFF0DEEFB2F" author="de Rooij, N." box="[627,764,1208,1233]" pageId="45" pageNumber="46" refId="ref43040" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. Volume II. Ophidia. E. J. Brill, Leiden, The Netherlands, 334 pp. https: // doi. org / 10.5962 / bhl. title. 24239" type="book" year="1917">Rooij, 1917</bibRefCitation>
:110
</taxonomicName>
. Petronella Johanna “Nelly” de Rooij (
<date id="341929FFFFEAFFDBFB03FAFF0A76FB2F" box="[1249,1380,1208,1233]" pageId="45" pageNumber="46" value="1883" valueMax="1964">18831964</date>
;
<figureCitation id="D89C13BAFFEAFFDBFA92FAFF0FAFFB0B" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (18401913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (18431920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (18201891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (18461926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (18581937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (18351917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (18711943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (18741935), author of the species name diehli. (I) Nelly de Rooij (18831964), author of the species name florensis. (J) Franz Werner (18671939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (18701921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (19021990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="45" pageNumber="46">
Fig.
<date id="341929FFFFEAFFDBFF75FA9A0FAFFB0B" box="[151,189,1245,1269]" pageId="45" pageNumber="46" value="2017-01">17I</date>
</figureCitation>
) was a Dutch zoologist and, from
<date id="341929FFFFEAFFDBFDADFA9A0DDDFB0B" box="[591,719,1245,1269]" pageId="45" pageNumber="46" value="1907" valueMax="1922">19071922</date>
, the curator of the collection in the Zoological Museum at the University of Amsterdam. During this time, de Rooij compiled two important volumes, presenting the first comprehensive descriptions of the reptiles in the Indo-Australian Archipelago (de
<bibRefCitation id="243672CEFFEAFFDBFB98FB630A13FAC3" author="de Rooij, N." box="[1146,1281,1316,1341]" pageId="45" pageNumber="46" refId="ref42988" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Folume I. Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, The Netherlands, 384 pp. https: // doi. org / 10.5962 / bhl. title. 24239" type="book" year="1915">Rooij 1915</bibRefCitation>
,
<bibRefCitation id="243672CEFFEAFFDBFAF1FB620A5DFAC3" author="de Rooij, N." box="[1299,1359,1317,1341]" pageId="45" pageNumber="46" refId="ref43040" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. Volume II. Ophidia. E. J. Brill, Leiden, The Netherlands, 334 pp. https: // doi. org / 10.5962 / bhl. title. 24239" type="book" year="1917">1917</bibRefCitation>
). It is noteworthy that during her career de Rooij experienced heavy discrimination by policies then in place that discriminated against women (
<bibRefCitation id="243672CEFFEAFFDBFE18FB2B0DF1FA7B" author="Beolens, B. &amp; Watkins, M. &amp; Grayson, M." box="[506,739,1388,1413]" pageId="45" pageNumber="46" refId="ref42294" refString="Beolens, B., Watkins, M. &amp; Grayson, M. (2011) The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, Maryland, USA, 312 pp." type="book" year="2011">
Beolens
<emphasis id="72D3D32DFFEAFFDBFD83FB290D84FA7B" box="[609,662,1388,1413]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
. 2011
</bibRefCitation>
), first against her academic advance, which required her to complete her doctorate (earned in 1907) in
<collectingRegion id="8263C1DDFFEAFFDBFD9DFBD70DDFFA57" box="[639,717,1424,1449]" country="Switzerland" name="Zuerich" pageId="45" pageNumber="46">Zürich</collectingRegion>
,
<collectingCountry id="38B04FAFFFEAFFDBFD3AFBD70C73FA57" box="[728,865,1424,1449]" name="Switzerland" pageId="45" pageNumber="46">Switzerland</collectingCountry>
, and later in her position as curator in Amsterdam, from which she elected to resign in 1922 at the age of only 39 due to discriminatory administrative reforms (
<bibRefCitation id="243672CEFFEAFFDBFAB9FBF30FC6FA0F" author="Adler, K." pageId="45" pageNumber="46" refId="ref41666" refString="Adler, K. (2007) Contributions to the History of Herpetology. Folume 2. Society for the Study of Reptiles and Amphibians, Lawrence, Kansas, USA, 400 pp." type="book" year="2007">Adler 2007</bibRefCitation>
). Unfortunately, this effectively ended de Rooijs research career. De Rooijs portrait (
<figureCitation id="D89C13BAFFEAFFDBFB52FB9F0A1FFA0F" box="[1200,1293,1496,1521]" captionStart="FIGURE 17" captionStartId="32.[151,250,1497,1519]" captionTargetBox="[151,1436,193,1473]" captionTargetId="figure@32.[151,1436,193,1473]" captionTargetPageId="32" captionText="FIGURE 17. Selected 19 th and 20 th Century authors and collectors of Stegonotus species. (A) Giacomo Doria (18401913), author of the species names aruensis and keyensis. (B) Odoardo Beccari (18431920), collector of the Stegonotus aruensis and S. keyensis type specimens. (C) William Macleay (18201891), author of the genus name Herbertophis and the species names darnleyensis and plumbeus. (D) John Archibald Boyd (18461926), collector of the Herbertophis plumbeus holotype. (E) George Albert Boulenger (18581937), author of the species names guentheri, heterurus, and reticulatus. Image licensed for use, © National Portrait Galley, London, United Kingdom. (F) The Reverend George Brown (18351917), collector of the Stegonotus heterurus syntypes from Duke of York Island. (G) Albert Stewart Meek (18711943), collector of the Stegonotus guentheri and S. reticulatus syntypes. (H) Wilhelm Adolfovich Lindholm (18741935), author of the species name diehli. (I) Nelly de Rooij (18831964), author of the species name florensis. (J) Franz Werner (18671939), author of the species names dorsalis and poechi. (K) Rudolf Pöch (18701921), collector of the Stegonotus dorsalis and S. poechi holotypes, and in whose honor the latter was named. (L) Lothar Forcart (19021990), author of the species name sutteri." httpUri="https://zenodo.org/record/2607609/files/figure.png" pageId="45" pageNumber="46">
Fig.
<date id="341929FFFFEAFFDBFB04FB9E0A1FFA0F" box="[1254,1293,1497,1521]" pageId="45" pageNumber="46" value="2017-01">17I</date>
</figureCitation>
) is from the archives of the RMNH. The species name reflects the
<typeStatus id="9F1CB19DFFEAFFDBFCF6FBBA0C56F9EB" box="[788,836,1533,1557]" pageId="45" pageNumber="46">type</typeStatus>
locality,
<collectingCountry id="38B04FAFFFEAFFDBFC50FBBB0CEAF9EB" box="[946,1016,1532,1557]" name="Indonesia" pageId="45" pageNumber="46">Flores</collectingCountry>
Island. The species description was presented in English.
</paragraph>
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<paragraph id="40180F3FFFEAFFDBFF25F8030A52F97F" blockId="45.[151,1437,1136,1953]" pageId="45" pageNumber="46">
<materialsCitation id="F0CF0562FFEAFFDBFF25F8030A52F97F" ID-GBIF-Occurrence="2028140086" collectionCode="ZMA" country="Netherlands" county="The" location="The" municipality="Leiden" pageId="45" pageNumber="46" specimenCount="1" typeStatus="holotype">
<emphasis id="72D3D32DFFEAFFDBFF25F8030E29F9A3" bold="true" box="[199,315,1604,1629]" pageId="45" pageNumber="46">
<typeStatus id="9F1CB19DFFEAFFDBFF25F8030E24F9A3" box="[199,310,1604,1629]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="26B697FAFFEAFFDBFEA4F8020E96F9A3" box="[326,388,1605,1629]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="45" pageNumber="46">ZMA</collectionCode>
11080, an adult female (
<figureCitation id="D89C13BAFFEAFFDBFD4DF8030C14F9A3" box="[687,774,1604,1629]" captionStart="FIGURE 26" captionStartId="46.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1050]" captionTargetId="figure@46.[151,1436,193,1050]" captionTargetPageId="46" captionText="FIGURE 26. Holotype of Stegonotus florensis (de Rooij, 1917), ZMA 11080, an adult female from Flores Island, East Nusa Tenggara Province, Indonesia. Shown are the entire specimen (TTL = 556 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views." httpUri="https://zenodo.org/record/2607631/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
;
<tableCitation id="0D253A84FFEAFFDBFCF4F8030C7EF9A3" box="[790,876,1604,1629]" captionStart="TABLE 1" captionStartId="60.[147,229,603,624]" captionText="TABLE 1. Name-bearing types of species in the genus Stegonotus. In this list we present basic data, including common measurements and selected scale counts, for all namebearing types (i.e., holotypes—HT, lectotypes—LT, neotypes—NT, syntypes—ST) to provide an easy overview of species. Additional details and the information for paratypes and paralectotypes can be found in the text. Character abbreviations are explained in the main text. For specimens where sex is indicated in red, we used the SCR value and not direct observation to estimate sex. Missing values are indicated by O. Specimen Type SCR SubC Species Synonym Sex SVL TL TTL V SC Dorsals SCE SL IL ILG Number Status (%) Condition LSUMZ admiraltiensis HT M 658 185 843 208 98 32 paired 17-19-15 4+5 8 10 10 93958 MSNG aruensis HT M 582 233 815 190 103 35 paired 17-17-15 3+4 7 8 4 30186 BMNH australis HT M 503 135 638 205 77 27 paired 17-17-15 4+5 8 9 5 1946.1.14.93 AMS australis darnleyensis HT F 720 O O 198 53+ O paired 17-17-15 4+5 9 10 5 R31894 AMS australis plumbeus HT M 1025 250 1275 220 74 25 paired 17-17-15 3+4+5 9 10 5 R31895 BMNH batjanensis HT M 1239 321 1560 235 86 27 paired 17-17-15 3+4+5 8 10 5 1946.1.11.36 FMNH borneensis HT F 783 221 1004 195 79 29 paired 17-17-15 4+5 10 11 5 164746 MNHN cucullatus HT M 769 254 1023 209 91 30 paired 17-17-15 4+5 8 9 5 3412 MTKD a cucullatus magnus ST F O O O 207 74 26 paired O-19-O 4+5 8 O O 573 ZMB b cucullatus magnus ST M 975 265 1240 211 87 29 paired 19-19-17 3+4 8 10 5 8794 MZB derooijae HT M 455 193 648 181 94 34 paired 17-17-15 3+4 7 8 8 Ophi.3288 MWNH diehli HT juv 196 56 252 179 82 31 paired 15-15-15 3+4 7 8 4 1244 NMW dorsalis HT M 805 245 1050 208 88 30 paired 15-15-15 4+5 8 10 5 14861 ZMA 9R florensis HT M 447 109 556 217 65 23 single 21-21-19 3+4+5 10 5 11080 8L BMNH guentheri LT F 683 165 848 192 64 25 paired 15-15-13 4+5 8 9 4 1946.1.11.38 ……continued on the next page" pageId="45" pageNumber="46">Table 1</tableCitation>
).
<location id="457859E4FFEAFFDBFC6AF8030CA6F9A3" LSID="urn:lsid:plazi:treatment:C80EBE29FFEAFFD8FF75FA6E0CCDF81A:457859E4FFEAFFDBFC6AF8030CA6F9A3" box="[904,948,1604,1629]" country="Netherlands" county="The" municipality="Leiden" name="The" pageId="45" pageNumber="46">The</location>
specimen is now housed in the
<location id="457859E4FFEAFFDBFAD7F8030E66F97F" LSID="urn:lsid:plazi:treatment:C80EBE29FFEAFFD8FF75FA6E0CCDF81A:457859E4FFEAFFDBFAD7F8030E66F97F" country="Netherlands" county="The" municipality="Leiden" name="Naturalis Biodiversity Center" pageId="45" pageNumber="46">Naturalis Biodiversity Center</location>
,
<collectingMunicipality id="A07C9545FFEAFFDBFE62F82F0EC3F97F" box="[384,465,1640,1665]" pageId="45" pageNumber="46">Leiden</collectingMunicipality>
,
<collectingCountry id="38B04FAFFFEAFFDBFE3EF82F0D8BF97F" box="[476,665,1640,1665]" name="Netherlands" pageId="45" pageNumber="46">The Netherlands</collectingCountry>
.
<collectingCounty id="A97977B3FFEAFFDBFD41F82F0DDDF97F" box="[675,719,1640,1665]" pageId="45" pageNumber="46">The</collectingCounty>
specimen was figured by de
<bibRefCitation id="243672CEFFEAFFDBFBF4F82F0A2EF97F" author="de Rooij, N." box="[1046,1340,1640,1665]" pageId="45" pageNumber="46" refId="ref43040" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. Volume II. Ophidia. E. J. Brill, Leiden, The Netherlands, 334 pp. https: // doi. org / 10.5962 / bhl. title. 24239" type="book" year="1917">Rooij (1917: Figs. 46, 47)</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="40180F3FFFEAFFDBFF25F8CB0EC0F8CB" blockId="45.[151,1437,1136,1953]" pageId="45" pageNumber="46">
<emphasis id="72D3D32DFFEAFFDBFF25F8CB0E79F95B" bold="true" box="[199,363,1676,1701]" pageId="45" pageNumber="46">Type locality.</emphasis>
<collectingCountry id="38B04FAFFFEAFFDBFE97F8CB0EAFF95B" box="[373,445,1676,1701]" name="Indonesia" pageId="45" pageNumber="46">Flores</collectingCountry>
(near Kotting)” [Koting, ca.
<quantity id="875FA2DAFFEAFFDBFCFDF8CA0C4CF95B" box="[799,862,1676,1701]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="45" pageNumber="46" unit="km" value="3.0">3 km</quantity>
S Maumere,
<collectingCountry id="38B04FAFFFEAFFDBFBE4F8CB0B5EF95B" box="[1030,1100,1676,1701]" name="Indonesia" pageId="45" pageNumber="46">Flores</collectingCountry>
Island,
<collectingRegion id="8263C1DDFFEAFFDBFB53F8CA0FEFF937" country="Indonesia" name="Nusa Tenggara Timur" pageId="45" pageNumber="46">East Nusa Tenggara Province</collectingRegion>
,
<collectingCountry id="38B04FAFFFEAFFDBFEEFF8F70E6FF937" box="[269,381,1712,1737]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
, ca.
<geoCoordinate id="259369F8FFEAFFDBFE58F8F70D1AF937" box="[442,520,1712,1737]" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="555" value="-8.68">8.68°S</geoCoordinate>
,
<geoCoordinate id="259369F8FFEAFFDBFDFBF8F60D9BF937" box="[537,649,1713,1737]" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="555" value="122.19">122.19°E</geoCoordinate>
]. The catalog entry for the specimen in the ZMA collection reads “
<collectingCountry id="38B04FAFFFEAFFDBFF43F8930FFEF913" box="[161,236,1748,1773]" name="Indonesia" pageId="45" pageNumber="46">Flores</collectingCountry>
, Omgeving van Kotting, Maumeri-Rivier” [
<collectingCountry id="38B04FAFFFEAFFDBFD3EF8930C35F913" box="[732,807,1748,1773]" name="Indonesia" pageId="45" pageNumber="46">Flores</collectingCountry>
, vicinity of Koting, Maumere River], and in the Koting area there are numerous small tributaries that drain into one larger riverbed that runs through the city of Maumere on the north coast of
<collectingCountry id="38B04FAFFFEAFFDBFE66F95B0EDCF8CB" box="[388,462,1820,1845]" name="Indonesia" pageId="45" pageNumber="46">Flores</collectingCountry>
.
</paragraph>
<paragraph id="40180F3FFFEAFFDBFF25F9070A6BF85F" blockId="45.[151,1437,1136,1953]" pageId="45" pageNumber="46">
<emphasis id="72D3D32DFFEAFFDBFF25F9070E55F8A7" bold="true" box="[199,327,1856,1881]" pageId="45" pageNumber="46">Collection.</emphasis>
According to the catalog entry in the ZMA collection, the specimen was collected by G.A.J. van de Sande, a member of the Koninklijk Instituut voor de Taal-, Land- en Volkenkunde van Nederlandsch-Indië [Royal Institute for Linguistics, Geography, and Ethnology of the Dutch Indies], who died in 1909 (
<bibRefCitation id="243672CEFFEAFFDBFB42F9CE0A7FF85F" author="Anonymous" box="[1184,1389,1929,1953]" pageId="45" pageNumber="46" refId="ref41701" refString="Anonymous (1911). Bestuursvergadering van 19 Februari 1910. Bijdragen tot de Taal-, Land- en Folkenkunde van Nederlandsch-Indie, 65 (1 / 2), vi-viii." type="book" year="1911">Anonymous 1911</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="14D85FB7FFE9FFD8FF75FA680DF7FB77" httpUri="https://zenodo.org/record/2607631/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,1071,1093]" targetBox="[151,1436,193,1050]" targetPageId="46">
<paragraph id="40180F3FFFE9FFD8FF75FA680DF7FB77" blockId="46.[151,1436,1071,1161]" pageId="46" pageNumber="47">
<emphasis id="72D3D32DFFE9FFD8FF75FA680E33FBBA" bold="true" box="[151,289,1071,1093]" pageId="46" pageNumber="47">FIGURE 26.</emphasis>
Holotype of
<taxonomicName id="87A774BCFFE9FFD8FE49FA770C38FBBB" authority="(de Rooij, 1917)" authorityName="de Rooij" authorityYear="1917" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[427,810,1071,1093]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FE49FA770D65FBBB" box="[427,631,1071,1093]" italics="true" pageId="46" pageNumber="47">Stegonotus florensis</emphasis>
(de Rooij, 1917)
</taxonomicName>
, ZMA 11080, an adult female from Flores Island, East Nusa Tenggara Province, Indonesia. Shown are the entire specimen (TTL = 556 mm) in (A) dorsal and (B) ventral views, and its head in (C) dorsal, (D) right lateral, and (E) ventral views.
</paragraph>
</caption>
<subSubSection id="08BD5CB4FFE9FFD8FF25FAFE0D9CF9B1" pageId="46" pageNumber="47" type="key">
<paragraph id="40180F3FFFE9FFD8FF25FAFE0EE8FAC1" blockId="46.[151,1437,1209,2021]" pageId="46" pageNumber="47">
<emphasis id="72D3D32DFFE9FFD8FF25FAFE0D78FB2C" bold="true" box="[199,618,1209,1234]" pageId="46" pageNumber="47">
Key characteristics of the
<materialsCitation id="F0CF0562FFE9FFD8FE1FFAFE0D78FB2C" ID-GBIF-Occurrence="2028140077" box="[509,618,1209,1234]" collectionCode="SVL, TTL, SCR, IL" pageId="46" pageNumber="47" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<typeStatus id="9F1CB19DFFE9FFD8FE1FFAFE0D74FB2C" box="[509,614,1209,1234]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</emphasis>
456 (447) mm SVL + 112 (109) mm TL = 568 (556) mm TTL. V
<subScript id="DC230D7AFFE9FFD8FAABFA8F0A47FB28" attach="left" box="[1353,1365,1224,1238]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FAABFA8F0A47FB28" box="[1353,1365,1224,1238]" pageId="46" pageNumber="47" type="female"></specimenCount>
</subScript>
= 218 (217), SC
<subScript id="DC230D7AFFE9FFD8FEE1FAAB0E1DFB04" attach="left" box="[259,271,1260,1274]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FEE1FAAB0E1DFB04" box="[259,271,1260,1274]" pageId="46" pageNumber="47" type="female"></specimenCount>
</subScript>
= 66 (65), SCR
<subScript id="DC230D7AFFE9FFD8FE22FAAB0EDEFB04" attach="left" box="[448,460,1260,1274]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FE22FAAB0EDEFB04" box="[448,460,1260,1274]" pageId="46" pageNumber="47" type="female"></specimenCount>
</subScript>
= 0.23 (0.23), D = O-21-O (21-21-19), SL
<subScript id="DC230D7AFFE9FFD8FC4DFAAB0CAAFB04" attach="left" box="[943,952,1260,1274]" fontSize="6" pageId="46" pageNumber="47">E</subScript>
= 3+4+5 (3+4+5), SL = 9R 8L (9R 8L), IL = O (10), IL
<subScript id="DC230D7AFFE9FFD8FEC7FB570E22FAE0" attach="left" box="[293,304,1296,1310]" fontSize="6" pageId="46" pageNumber="47">G</subScript>
= O (5). Other than
<taxonomicName id="87A774BCFFE9FFD8FDFCFB440DB4FAE5" authorityName="Boulenger" authorityYear="1893" box="[542,678,1282,1307]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="heterurus">
<emphasis id="72D3D32DFFE9FFD8FDFCFB440DB4FAE5" box="[542,678,1282,1307]" italics="true" pageId="46" pageNumber="47">S. heterurus</emphasis>
</taxonomicName>
and
<taxonomicName id="87A774BCFFE9FFD8FD3FFB440C50FAE4" authorityName="Forcart" authorityYear="1953" box="[733,834,1283,1306]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="sutteri">
<emphasis id="72D3D32DFFE9FFD8FD3FFB440C50FAE4" box="[733,834,1283,1306]" italics="true" pageId="46" pageNumber="47">S. sutteri</emphasis>
</taxonomicName>
(see below), this species is the only other
<taxonomicName id="87A774BCFFE9FFD8FAC0FB440A8EFAE4" authorityName="Guentheri Boulenger" authorityYear="1895" box="[1314,1436,1283,1306]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFE9FFD8FAC0FB440A8EFAE4" box="[1314,1436,1283,1306]" italics="true" pageId="46" pageNumber="47">Stegonotus</emphasis>
</taxonomicName>
with unpaired subcaudal scales.
</paragraph>
<paragraph id="40180F3FFFE9FFD8FF25FB000D9CF9B1" blockId="46.[151,1437,1209,2021]" pageId="46" pageNumber="47">
<emphasis id="72D3D32DFFE9FFD8FF25FB000D49FA9E" bold="true" box="[199,603,1351,1376]" pageId="46" pageNumber="47">Key characteristics of the species.</emphasis>
We were only able to find and examine two specimens of this form from
<collectingCountry id="38B04FAFFFE9FFD8FF75FB2F0FCFFA7F" box="[151,221,1384,1409]" name="Indonesia" pageId="46" pageNumber="47">Flores</collectingCountry>
Island (the terra typica) in collections accessible to us, the female
<typeStatus id="9F1CB19DFFE9FFD8FC0DFB2F0B43FA7F" box="[1007,1105,1384,1409]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
and a male specimen in the Western Australian Museum, Perth,
<collectingCountry id="38B04FAFFFE9FFD8FDA1FBCD0DB8FA5D" box="[579,682,1418,1443]" name="Australia" pageId="46" pageNumber="47">Australia</collectingCountry>
(WAM 104606) collected in 1991. The dearth of specimens in collections makes any assessment of the characteristics of
<taxonomicName id="87A774BCFFE9FFD8FCBEFBEB0CCCFA3A" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[860,990,1451,1476]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FCBEFBEB0CCCFA3A" box="[860,990,1451,1476]" italics="true" pageId="46" pageNumber="47">S. florensis</emphasis>
</taxonomicName>
at the species level tenuous at best. Characteristics include V
<subScript id="DC230D7AFFE9FFD8FE50FB9C0EACFA17" attach="left" box="[434,446,1499,1513]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FE50FB9C0EACFA17" box="[434,446,1499,1513]" pageId="46" pageNumber="47" type="female"></specimenCount>
</subScript>
= 217, V
<subScript id="DC230D7AFFE9FFD8FDCAFB9C0D26FA17" attach="left" box="[552,564,1499,1513]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FDCAFB9C0D26FA17" box="[552,564,1499,1513]" pageId="46" pageNumber="47" type="male"></specimenCount>
</subScript>
= 230; SC
<subScript id="DC230D7AFFE9FFD8FD4EFB9C0DAAFA17" attach="left" box="[684,696,1499,1513]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FD4EFB9C0DAAFA17" box="[684,696,1499,1513]" pageId="46" pageNumber="47" type="female"></specimenCount>
</subScript>
= 65, SC
<subScript id="DC230D7AFFE9FFD8FCC1FB9C0C3DFA17" attach="left" box="[803,815,1499,1513]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FCC1FB9C0C3DFA17" box="[803,815,1499,1513]" pageId="46" pageNumber="47" type="male"></specimenCount>
</subScript>
= 82; SCR
<subScript id="DC230D7AFFE9FFD8FC4EFB9C0CAAFA17" attach="left" box="[940,952,1499,1513]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FC4EFB9C0CAAFA17" box="[940,952,1499,1513]" pageId="46" pageNumber="47" type="female"></specimenCount>
</subScript>
= 0.23, SCR
<subScript id="DC230D7AFFE9FFD8FBA8FB9C0B44FA17" attach="left" box="[1098,1110,1499,1513]" fontSize="6" pageId="46" pageNumber="47">
<specimenCount id="56A1C4B6FFE9FFD8FBA8FB9C0B44FA17" box="[1098,1110,1499,1513]" pageId="46" pageNumber="47" type="male"></specimenCount>
</subScript>
= 0.26; D = 21-21-19; SL
<subScript id="DC230D7AFFE9FFD8FA9EFB9C0A97FA17" attach="left" box="[1404,1413,1499,1513]" fontSize="6" pageId="46" pageNumber="47">E</subScript>
= 3+4+5; SL = 8 or 9; IL = 9 or 10; IL
<subScript id="DC230D7AFFE9FFD8FDBBFBB80D76F9F3" attach="left" box="[601,612,1535,1549]" fontSize="6" pageId="46" pageNumber="47">G</subScript>
= 5. It appears that sexual dimorphism in the numbers of ventral and subcaudal scales might exist, with males possessing higher numbers of both and a relatively longer tail. However, given the low sample size this is speculative.
</paragraph>
</subSubSection>
<subSubSection id="08BD5CB4FFE9FFD8FF25F8100CCDF81A" pageId="46" pageNumber="47" type="discussion">
<paragraph id="40180F3FFFE9FFD8FF25F8100CCDF81A" blockId="46.[151,1437,1209,2021]" pageId="46" pageNumber="47">
<emphasis id="72D3D32DFFE9FFD8FF25F8100E5DF98E" bold="true" box="[199,335,1623,1648]" pageId="46" pageNumber="47">Comments.</emphasis>
Based on x-ray images,
<bibRefCitation id="243672CEFFE9FFD8FD92F81F0C07F98E" author="Forcart, L." box="[624,789,1623,1648]" pageId="46" pageNumber="47" pagination="7 - 8" refId="ref43652" refString="Forcart, L. (1954) Die Taxonomie von Lycodon florensis Rooij und Stegonotus sutteri Forcart (Ophidia, Colubridae). Ferhandlungen der naturforschenden Gesellschaft zu Basel, 65, 7 - 8." type="journal article" year="1954">Forcart (1954)</bibRefCitation>
noted that
<taxonomicName id="87A774BCFFE9FFD8FC78F81E0B08F98E" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[922,1050,1623,1648]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FC78F81E0B08F98E" box="[922,1050,1623,1648]" italics="true" pageId="46" pageNumber="47">L. florensis</emphasis>
</taxonomicName>
did not possess the characteristic separation dividing the maxillary teeth into two sets, with some enlarged teeth anterior to the gap. Instead, he considered the maxillary dentition of
<taxonomicName id="87A774BCFFE9FFD8FDA2F8DC0DACF94D" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[576,702,1690,1715]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FDA2F8DC0DACF94D" box="[576,702,1690,1715]" italics="true" pageId="46" pageNumber="47">L. florensis</emphasis>
</taxonomicName>
to reflect the characteristics used by
<bibRefCitation id="243672CEFFE9FFD8FB86F8DD0A46F94D" author="Dumeril, A. - M. - C. &amp; Bibron, G. &amp; Dumeril, A. - H. - A." box="[1124,1364,1690,1715]" pageId="46" pageNumber="47" refId="ref43291" refString="Dumeril, A. - M. - C., Bibron, G. &amp; Dumeril, A. - H. - A. (1854) Erpetologie generale ou histoire naturelle complete des reptiles. Tome septieme, Partie 1. Librairie Encyclopedique de Roret, Paris, France, 780 pp. https: // doi. org / 10.5962 / bhl. title. 112229" type="book" year="1854">
Duméril
<emphasis id="72D3D32DFFE9FFD8FB2BF8DC0BE9F94D" box="[1225,1275,1690,1715]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. (1854)
</bibRefCitation>
to the genus
<taxonomicName id="87A774BCFFE9FFD8FF3DF8FB0E4BF92D" authorityName="Guentheri Boulenger" authorityYear="1895" box="[223,345,1724,1747]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="72D3D32DFFE9FFD8FF3DF8FB0E4BF92D" box="[223,345,1724,1747]" italics="true" pageId="46" pageNumber="47">Stegonotus</emphasis>
</taxonomicName>
. It should be noted that this specimens presents with an aberration in the set of supralabial scales on both sides of the head. Unrecognized by de
<bibRefCitation id="243672CEFFE9FFD8FD48F89A0C28F908" author="de Rooij, N." box="[682,826,1757,1782]" pageId="46" pageNumber="47" refId="ref43040" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. Volume II. Ophidia. E. J. Brill, Leiden, The Netherlands, 334 pp. https: // doi. org / 10.5962 / bhl. title. 24239" type="book" year="1917">Rooij (1917)</bibRefCitation>
and drawn incorrectly in her line drawing of the right lateral side of the head, SL5 appears to be divided into two parts. While this does not change the overall supralabial count, it influences whether SL5 contacts the eye or not. In the second specimen of
<taxonomicName id="87A774BCFFE9FFD8FBBBF9670BC4F8C6" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[1113,1238,1823,1848]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FBBBF9670BC4F8C6" box="[1113,1238,1823,1848]" italics="true" pageId="46" pageNumber="47">S. florensis</emphasis>
</taxonomicName>
we examined, as well as in all specimens of the likely related
<taxonomicName id="87A774BCFFE9FFD8FD46F9050C19F8A7" authorityName="Forcart" authorityYear="1953" box="[676,779,1858,1881]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="sutteri">
<emphasis id="72D3D32DFFE9FFD8FD46F9050C19F8A7" box="[676,779,1858,1881]" italics="true" pageId="46" pageNumber="47">S. sutteri</emphasis>
</taxonomicName>
(see below), SL5 is a single, large supralabial. De
<bibRefCitation id="243672CEFFE9FFD8FA80F9050FF2F885" author="de Lang, R." pageId="46" pageNumber="47" refId="ref42893" refString="de Lang, R. (2011) The Snakes of the Lesser Sunda Islands (Nusa Tenggara), Indonesia: a Field Guide to the Terrestrial and Semi-Aquatic Snakes with Identification Key. Edition Chimaira, Frankfurt am Main, Germany, 349 pp." type="book" year="2011">Lang (2011)</bibRefCitation>
stated that specimens of
<taxonomicName id="87A774BCFFE9FFD8FE1DF9240D69F885" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[511,635,1890,1915]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FE1DF9240D69F885" box="[511,635,1890,1915]" italics="true" pageId="46" pageNumber="47">S. florensis</emphasis>
</taxonomicName>
he studied varied in SL
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, with some specimens showing values of 3+4 and 4+
<quantity id="875FA2DAFFE9FFD8FF0AF9C10E07F861" box="[232,277,1926,1951]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="46" pageNumber="47" unit="in" value="5.0">5 in</quantity>
addition to the diagnostic 3+4+5. We cannot confirm or deny this, but none of the specimens of
<taxonomicName id="87A774BCFFE9FFD8FA65F9C00FE4F83E" baseAuthorityName="de Rooij" baseAuthorityYear="1917" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FA65F9C00FE4F83E" italics="true" pageId="46" pageNumber="47">S. florensis</emphasis>
</taxonomicName>
(as well as of
<taxonomicName id="87A774BCFFE9FFD8FE7FF9EF0D10F83E" authorityName="Forcart" authorityYear="1953" box="[413,514,1960,1984]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="sutteri">
<emphasis id="72D3D32DFFE9FFD8FE7FF9EF0D10F83E" box="[413,514,1960,1984]" italics="true" pageId="46" pageNumber="47">S. sutteri</emphasis>
</taxonomicName>
, see below) we examined had conditions of SL
<subScript id="DC230D7AFFE9FFD8FBFEF9F10B37F83A" attach="left" box="[1052,1061,1974,1988]" fontSize="6" pageId="46" pageNumber="47">E</subScript>
other than 3+4+5. All other data listed by de
<bibRefCitation id="243672CEFFE9FFD8FEFDF98A0EBBF81A" author="de Lang, R." box="[287,425,1996,2021]" pageId="46" pageNumber="47" refId="ref42893" refString="de Lang, R. (2011) The Snakes of the Lesser Sunda Islands (Nusa Tenggara), Indonesia: a Field Guide to the Terrestrial and Semi-Aquatic Snakes with Identification Key. Edition Chimaira, Frankfurt am Main, Germany, 349 pp." type="book" year="2011">Lang (2011)</bibRefCitation>
conform to our concept of the species
<taxonomicName id="87A774BCFFE9FFD8FCBEF98A0CC5F81B" baseAuthorityName="de Rooij" baseAuthorityYear="1917" box="[860,983,1996,2021]" class="Reptilia" family="Colubridae" genus="Stegonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="florensis">
<emphasis id="72D3D32DFFE9FFD8FCBEF98A0CC5F81B" box="[860,983,1996,2021]" italics="true" pageId="46" pageNumber="47">S. florensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>