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<document id="4A55D0ED4494D334D9AA98F4D7602896" ID-DOI="10.11646/zootaxa.5322.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203657" ID-ZooBank="DD295EE6-AB3F-4377-900E-CC76F1C6BC2D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690885128246" checkinUser="plazi" docAuthor="Klompen, Hans &amp; Gerdeman, Beverly S." docDate="2023" docId="C55587C9FFEDB81A86C9CC7BF12AFE26" docLanguage="en" docName="zootaxa.5322.1.1.pdf" docOrigin="Zootaxa 5322 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5322.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Allozercon Vitzthum 1926" docType="treatment" docVersion="1" lastPageNumber="37" masterDocId="396CFFB1FFCEB83E865EC861F07CFFD3" masterDocTitle="Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="36" updateTime="1726239911995" updateUser="tatiana">
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<mods:title id="290EE4B986223DB4A38EA5241DB91224">Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)</mods:title>
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<mods:namePart id="1C564285CA5C2B88AA6894DE6ACE47BC">Klompen, Hans</mods:namePart>
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<mods:namePart id="849F2D9ADA8C9FA9F6A74E1DA6D0E012">Gerdeman, Beverly S.</mods:namePart>
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<mods:title id="A56353246E01EDA263BD9245D9302E7F">Zootaxa</mods:title>
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<mods:date id="07F9DCCF3F6CA6EA0E1F0E2B8559C56D">2023</mods:date>
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<mods:identifier id="776AFAD927CFA286EA63EA807EDCC2E7" type="DOI">10.11646/zootaxa.5322.1.1</mods:identifier>
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<heading id="160B81B3FFEDB81D86C9CC7BF1A5FBE7" bold="true" box="[151,473,1050,1077]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CC7BF1A5FBE7" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[151,473,1050,1077]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC7BF1A5FBE7" bold="true" box="[151,473,1050,1077]" pageId="35" pageNumber="36">
<emphasis id="7F88EACDFFEDB81D86C9CC7BF165FBE6" bold="true" box="[151,281,1050,1077]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D877ECC7BF1A5FBE7" author="Vitzthum, H. &amp; Graf" box="[288,473,1050,1076]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</emphasis>
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<treatmentCitation id="CC5D10CEFFEDB81D86C9CC3EF1A9FBA4" author="Vitzthum, H. &amp; Graf" box="[151,469,1119,1143]" page="104" pageId="35" pageNumber="36" year="1926">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CC3EF1A9FBA4" authority="Vitzthum 1926: 104" authorityName="Vitzthum" authorityPageNumber="104" authorityYear="1926" box="[151,469,1119,1143]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC3EF178FBA4" box="[151,260,1119,1143]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D8754CC3EF1A9FBA4" author="Vitzthum, H. &amp; Graf" box="[266,469,1119,1143]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926: 104</bibRefCitation>
</taxonomicName>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D8785CC01F56CFBA4" box="[475,1296,1119,1144]" pageId="35" pageNumber="36" type="type_taxon">
<paragraph id="4D4336DFFFEDB81D8785CC01F56CFBA4" blockId="35.[151,1437,1119,1297]" box="[475,1296,1119,1144]" pageId="35" pageNumber="36">
[
<typeStatus id="9247887DFFEDB81D87BFCC01F273FBAB" box="[481,527,1120,1144]" pageId="35" pageNumber="36">type</typeStatus>
species.
<taxonomicName id="8AFC4D5CFFEDB81D8435CC3EF470FBA4" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[619,1036,1119,1143]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="fecundissimus">
<emphasis id="7F88EACDFFEDB81D8435CC3EF311FBA4" box="[619,877,1119,1143]" italics="true" pageId="35" pageNumber="36">Allozercon fecundissimus</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D852DCC3EF470FBA4" author="Vitzthum, H. &amp; Graf" box="[883,1036,1119,1143]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</taxonomicName>
, by original designation].
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D86C9CC1FF190FB00" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="4D4336DFFFEDB81D86C9CC1FF18BFB66" blockId="35.[151,1437,1119,1297]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="6DEC11F1FFEDB81D86C9CC1FF18BFB66" pageId="35" pageNumber="36">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CC1FF178FB45" authorityName="Vitzthum" authorityYear="1926" box="[151,260,1150,1174]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC1FF178FB45" box="[151,260,1150,1174]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
</taxonomicName>
.—
<treatmentCitation id="CC5D10CEFFEDB81D877FCC1FF27BFB45" author="Womersley, H." box="[289,519,1150,1174]" page="129" pageId="35" pageNumber="36" year="1958">
<bibRefCitation id="296D4B2EFFEDB81D877FCC1FF27BFB45" author="Womersley, H." box="[289,519,1150,1174]" pageId="35" pageNumber="36" pagination="115 - 130" refId="ref39986" refString="Womersley, H. (1958) Some new or little known Mesostigmata (Acarina) from Australia, New Zealand and Malaya. Transactions of the Royal Society of South Australia, 81, 115 - 130." type="journal article" year="1958">Womersley 1958: 129</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CC5D10CEFFEDB81D844DCC1FF30BFB45" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[531,887,1150,1174]" page="147" pageId="35" pageNumber="36" year="1990">
<bibRefCitation id="296D4B2EFFEDB81D844DCC1FF30BFB45" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[531,887,1150,1174]" pageId="35" pageNumber="36" pagination="143 - 148" refId="ref37819" refString="Flechtmann, C. H. W. &amp; Johnston, D. E. (1990) Zeterohercon, a new genus of Heterozerconidae (Acari: Mesostigmata) and the description of Zeterohercon amphisbaenae n. sp. from Brasil. International Journal of Acarology, 16, 143 - 148. https: // doi. org / 10.1080 / 01647959008683526" type="journal article" year="1990">Flechtmann &amp; Johnston 1990: 147</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CC5D10CEFFEDB81D85DCCC1FF4C9FB45" author="Gerdeman, B. S. &amp; Garcia, R." box="[898,1205,1150,1174]" page="93" pageId="35" pageNumber="36" year="2010">
<bibRefCitation id="296D4B2EFFEDB81D85DCCC1FF4C9FB45" author="Gerdeman, B. S. &amp; Garcia, R." box="[898,1205,1150,1174]" pageId="35" pageNumber="36" pagination="93 - 96" refId="ref37991" refString="Gerdeman, B. S. &amp; Garcia, R. (2010) Heterozerconidae: A comparison between a temperate and a tropical species. In: Sabelis, M. W. &amp; Bruin, J. (Eds.), Trends in Acarology, Proceedings of the 12 th International Congress. Springer Science, Dordrecht, pp. 93 - 96. https: // doi. org / 10.1007 / 978 - 90 - 481 - 9837 - 5 _ 14" type="book chapter" year="2010">Gerdeman &amp; Garcia 2010: 93</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CC5D10CEFFEDB81D829FCC1FF08CFB67" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." page="640" pageId="35" pageNumber="36" year="2015">
<bibRefCitation id="296D4B2EFFEDB81D829FCC1FF08CFB67" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." pageId="35" pageNumber="36" pagination="639 - 655" refId="ref37329" refString="Di Palma, A., de Moraes, G. J., Gerdeman, B. S., Huber, S., Kitajima, E. W. &amp; Alberti, G. (2015) Ultrastructural and functional adaptations of the female reproductive system in the family Heterozerconidae (Acari, Anactinotrichida, Gamasida, Heterozerconina) and implications for the systematic position of the group. Arthropod Structure &amp; Development, 44, 639 - 655. https: // doi. org / 10.1016 / j. asd. 2015.09.002" type="journal article" year="2015">
Di Palma
<emphasis id="7F88EACDFFEDB81D8376CC1EF529FB45" box="[1320,1365,1150,1174]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2015: 640
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CC5D10CEFFEDB81D86A5CCFDF18EFB66" ID-DOI="http://doi.org/10.5281/zenodo.5965975" ID-Zenodo-Dep="5965975" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[251,498,1180,1205]" httpUri="http://treatment.plazi.org/id/03A2D654FF9DFFAFF4FCFB07FD09CABC" page="9" pageId="35" pageNumber="36" year="2018">
<bibRefCitation id="296D4B2EFFEDB81D86A5CCFDF18EFB66" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[251,498,1180,1205]" pageId="35" pageNumber="36" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEDB81D8734CCFCF1EBFB67" box="[362,407,1180,1204]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2018: 9
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4D4336DFFFEDB81D86C9CCDAF190FB00" blockId="35.[151,1437,1119,1297]" box="[151,492,1211,1235]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="6DEC11F1FFEDB81D86C9CCDAF190FB00" box="[151,492,1211,1235]" pageId="35" pageNumber="36">
<treatmentCitation id="CC5D10CEFFEDB81D86C9CCDAF190FB00" author="Fain, A." box="[151,492,1211,1235]" page="146" pageId="35" pageNumber="36" year="1989">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CCDAF190FB00" authority="Fain 1989: 146" authorityName="Fain" authorityPageNumber="146" authorityYear="1989" box="[151,492,1211,1235]" class="Arachnida" family="Heterozerconidae" genus="Asioheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CCDAF13AFB00" box="[151,326,1211,1235]" italics="true" pageId="35" pageNumber="36">Asioheterozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D8711CCDAF190FB00" author="Fain, A." box="[335,492,1211,1235]" pageId="35" pageNumber="36" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 146</bibRefCitation>
</taxonomicName>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D87ABCCDAF3B9FB21" pageId="35" pageNumber="36" type="multiple">
<paragraph id="4D4336DFFFEDB81D87ABCCDAF3B9FB21" blockId="35.[151,1437,1119,1297]" pageId="35" pageNumber="36">
(
<typeStatus id="9247887DFFEDB81D87A2CCDAF255FB00" box="[508,553,1211,1235]" pageId="35" pageNumber="36">type</typeStatus>
species
<taxonomicName id="8AFC4D5CFFEDB81D84DDCCDDF39CFB00" authority="Berlese 1910" authorityName="Berlese" authorityYear="1910" box="[643,992,1211,1235]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="audax">
<emphasis id="7F88EACDFFEDB81D84DDCCDDF32DFB00" box="[643,849,1211,1235]" italics="true" pageId="35" pageNumber="36">Heterozercon audax</emphasis>
Berlese 1910
</taxonomicName>
by original designation);
<treatmentCitation id="CC5D10CEFFEDB81D82B0CCDDF5E0FB00" author="Evans, G. O." box="[1262,1436,1211,1235]" page="426" pageId="35" pageNumber="36" year="1992">
<bibRefCitation id="296D4B2EFFEDB81D82B0CCDDF5E0FB00" author="Evans, G. O." box="[1262,1436,1211,1235]" pageId="35" pageNumber="36" refId="ref37637" refString="Evans, G. O. (1992) Principles of acarology. CAB International, Wallingford, 563 pp. https: // doi. org / 10.1079 / 9780851988221.0000" type="book" year="1992">Evans 1992: 426</bibRefCitation>
</treatmentCitation>
[junior synonym of
<taxonomicName id="8AFC4D5CFFEDB81D87CFCCBBF2E7FB21" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[401,667,1242,1266]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D87CFCCBBF182FB21" box="[401,510,1242,1266]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D845ACCBBF2E7FB21" author="Vitzthum, H. &amp; Graf" box="[516,667,1242,1266]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</taxonomicName>
by
<treatmentCitation id="CC5D10CEFFEDB81D849FCCBBF3C6FB21" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[705,954,1242,1266]" page="9" pageId="35" pageNumber="36" year="2018">
<bibRefCitation id="296D4B2EFFEDB81D849FCCBBF3C6FB21" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[705,954,1242,1266]" pageId="35" pageNumber="36" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEDB81D856FCCBAF321FB21" box="[817,861,1242,1266]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2018: 9
</bibRefCitation>
</treatmentCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D86C9CC99F382FAC3" box="[151,1022,1272,1297]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="4D4336DFFFEDB81D86C9CC99F382FAC3" blockId="35.[151,1437,1119,1297]" box="[151,1022,1272,1297]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="6DEC11F1FFEDB81D86C9CC99F382FAC3" box="[151,1022,1272,1297]" pageId="35" pageNumber="36">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CC99F138FAC3" authority="Fain 1989" box="[151,324,1272,1296]" class="Arachnida" family="Heterozerconidae" genus="Alloheterozercon" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC99F138FAC3" box="[151,324,1272,1296]" italics="true" pageId="35" pageNumber="36">Alloheterozercon</emphasis>
</taxonomicName>
.—
<bibRefCitation id="296D4B2EFFEDB81D873FCC99F2C4FAC3" author="Gerdeman, B. S. &amp; Klompen, H." box="[353,696,1272,1297]" pageId="35" pageNumber="36" pagination="351 - 370" refId="ref38078" refString="Gerdeman, B. S. &amp; Klompen, H. (2003) A new North American heterozerconid, Narceoheterozercon ohioensis, n. gen., n. sp., with first description of immatures of Heterozerconidae (Acari: Mesostigmata). International Journal of Acarology, 29, 351 - 370. https: // doi. org / 10.1080 / 01647950308684352" type="journal article" year="2003">Gerdeman &amp; Klompen 2003: 353</bibRefCitation>
[incorrect subsequent spelling].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D86C9CD5EF1F4F9DE" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="4D4336DFFFEDB81D86C9CD5EF1F4F9DE" blockId="35.[151,1437,1343,2017]" pageId="35" pageNumber="36">
Diagnosis. Axial outgrowth of palp trochanter in male distinct and sclerotized, solid (9); setae
<emphasis id="7F88EACDFFEDB81D8297CD5EF483FA8A" box="[1225,1279,1343,1369]" italics="true" pageId="35" pageNumber="36">hyp2</emphasis>
at least twice as long as setae
<emphasis id="7F88EACDFFEDB81D870ACD02F1F6FAAE" box="[340,394,1379,1405]" italics="true" pageId="35" pageNumber="36">hyp3</emphasis>
(22); insertion sternal setae
<emphasis id="7F88EACDFFEDB81D848CCD02F28EFAAE" box="[722,754,1379,1405]" italics="true" pageId="35" pageNumber="36">st2</emphasis>
and
<emphasis id="7F88EACDFFEDB81D8573CD02F332FAAE" box="[813,846,1379,1405]" italics="true" pageId="35" pageNumber="36">
st
<quantity id="8A049B3AFFEDB81D851ECD02F332FAAE" box="[832,846,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="35" pageNumber="36" unit="in" value="3.0">3</quantity>
</emphasis>
in female and
<emphasis id="7F88EACDFFEDB81D85A3CD02F462FAAE" box="[1021,1054,1379,1405]" italics="true" pageId="35" pageNumber="36">
st
<quantity id="8A049B3AFFEDB81D824ECD02F462FAAE" box="[1040,1054,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="35" pageNumber="36" unit="in" value="3.0">3</quantity>
</emphasis>
in male on isolated platelets (38, 40, 52); insertion sternal setae
<emphasis id="7F88EACDFFEDB81D87A8CDE6F26BFA72" box="[502,535,1415,1441]" italics="true" pageId="35" pageNumber="36">
st
<quantity id="8A049B3AFFEDB81D8455CDE6F26BFA72" box="[523,535,1415,1441]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1</quantity>
</emphasis>
in male on isolated platelets (48); opisthogastral setae
<emphasis id="7F88EACDFFEDB81D82D4CDE6F4CDFA72" box="[1162,1201,1415,1441]" italics="true" pageId="35" pageNumber="36">Jv5</emphasis>
inserted posterior to ventrianal fusion line (71); all marginal opisthosomal setae setiform (77, reversal); femora II setae
<emphasis id="7F88EACDFFEDB81D82B3CDCAF57FFA16" box="[1261,1283,1451,1477]" italics="true" pageId="35" pageNumber="36">al</emphasis>
,
<emphasis id="7F88EACDFFEDB81D834FCDCAF55BFA16" box="[1297,1319,1451,1477]" italics="true" pageId="35" pageNumber="36">pl</emphasis>
, and
<emphasis id="7F88EACDFFEDB81D833BCDCAF503FA17" box="[1381,1407,1451,1476]" italics="true" pageId="35" pageNumber="36">av</emphasis>
in male spines (105–107, shared with
<taxonomicName id="8AFC4D5CFFEDB81D8479CDAEF2B5FA3B" authorityName="Fain" authorityYear="1989" box="[551,713,1487,1512]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D8479CDAEF2B5FA3B" box="[551,713,1487,1512]" italics="true" pageId="35" pageNumber="36">Maracazercon</emphasis>
</taxonomicName>
and
<taxonomicName id="8AFC4D5CFFEDB81D855FCDAEF3EBFA3B" authorityName="Berlese" authorityYear="1888" box="[769,919,1487,1512]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D855FCDAEF3EBFA3B" box="[769,919,1487,1512]" italics="true" pageId="35" pageNumber="36">Heterozercon</emphasis>
</taxonomicName>
); genual and tibial setae
<emphasis id="7F88EACDFFEDB81D82EDCDAEF4B1FA3B" box="[1203,1229,1487,1512]" italics="true" pageId="35" pageNumber="36">pv</emphasis>
on legs II in male spine-like (109, 110).
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81A8699CE76F12AFE26" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="discussion">
<paragraph id="4D4336DFFFEDB81D8699CE76F450F86E" blockId="35.[151,1437,1343,2017]" pageId="35" pageNumber="36">
Remarks. A re-evaluation of the 11 characters listed by
<bibRefCitation id="296D4B2EFFEDB81D856DCE76F3CAF9E2" author="Fain, A." box="[819,950,1559,1585]" pageId="35" pageNumber="36" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain (1989)</bibRefCitation>
for the genus
<taxonomicName id="8AFC4D5CFFEDB81D8208CE76F564F9E2" authorityName="Fain" authorityYear="1989" box="[1110,1304,1559,1585]" class="Arachnida" family="Heterozerconidae" genus="Asioheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D8208CE76F564F9E2" box="[1110,1304,1559,1585]" italics="true" pageId="35" pageNumber="36">Asioheterozercon</emphasis>
</taxonomicName>
, shows that two are included in the updated diagnosis of
<taxonomicName id="8AFC4D5CFFEDB81D84D2CE5AF378F986" authorityName="Vitzthum" authorityYear="1926" box="[652,772,1595,1621]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D84D2CE5AF378F986" box="[652,772,1595,1621]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
</taxonomicName>
listed above: palpal trochanter with its apico-internal angle strongly produced and bearing 2 setae (our character 9) and anal shield with five setae (71). Three of the remaining characters are variable among the S.E. Asian populations: tarsus and tibia of legs I much narrower than the other segments of that leg (81.2); anterior region of the dorsum with 20–40 long and stiff setae (29.1); antero-lateral margin of dorsal shield in male with numerous small spines (30.1). One is incorrect: palp trochanter in female bears only one seta. Seta
<emphasis id="7F88EACDFFEDB81D8764CE8EF128F8DA" box="[314,340,1775,1801]" italics="true" pageId="35" pageNumber="36">v2</emphasis>
is in fact present and is inserted on the membranous axial outgrowth of the trochanter. However, it is highly reduced (
<figureCitation id="D5C72A5AFFEDB81D87DACF72F1A6F8FE" box="[388,474,1811,1837]" captionStart="FIGURES 39–42" captionStartId="45.[151,264,1909,1933]" captionTargetBox="[226,1361,184,1882]" captionTargetId="figure-21@45.[223,1364,181,1885]" captionTargetPageId="45" captionText="FIGURES 39–42. Heterozerconidae, gnathosoma, adults: 39, Allozercon leytensis n. sp., female (OSAL 053243); 40, Allozercon leytensis n. sp., male (UPLB MNH ACA-01335; red arrow: sclerotized axial extension of palp trochanter); 41, Al. (Philippinozercon) makilingensis, female (OSAL 053271); 42, Al. (Philippinozercon) makilingensis male (OSAL 102161). Inset: gnathotectum and labrum + salivary stylets." figureDoi="http://doi.org/10.5281/zenodo.8203685" httpUri="https://zenodo.org/record/8203685/files/figure.png" pageId="35" pageNumber="36">Fig. 39</figureCitation>
). The remaining characters are imprecise or refer to primitive character states at this taxonomic level. To further clarify the status of
<taxonomicName id="8AFC4D5CFFEDB81D84E6CF56F34CF882" authorityName="Vitzthum" authorityYear="1926" box="[696,816,1847,1873]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D84E6CF56F34CF882" box="[696,816,1847,1873]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
</taxonomicName>
a complete description is provided for one of the nine geographically separate populations for which we have specimens available. These nine populations originate in the
<collectingCountry id="35EB764FFFEDB81D869ECF1EF142F84A" box="[192,318,1919,1945]" name="Philippines" pageId="35" pageNumber="36">Philippines</collectingCountry>
(
<collectingRegion id="8F38F83DFFEDB81D8710CF1EF1EBF84A" box="[334,407,1919,1945]" country="Philippines" name="Bohol" pageId="35" pageNumber="36">Bohol</collectingRegion>
,
<collectingRegion id="8F38F83DFFEDB81D87FFCF1EF198F84A" box="[417,484,1919,1945]" country="Philippines" name="Southern Leyte" pageId="35" pageNumber="36">Leyte</collectingRegion>
, Mindoro,
<collectingRegion id="8F38F83DFFEDB81D8401CF1EF2D4F84A" box="[607,680,1919,1945]" country="Philippines" name="Samar" pageId="35" pageNumber="36">Samar</collectingRegion>
, and
<collectingRegion id="8F38F83DFFEDB81D84BDCF1EF341F84A" box="[739,829,1919,1945]" country="Philippines" name="Siquijor" pageId="35" pageNumber="36">Siquijor</collectingRegion>
islands),
<collectingCountry id="35EB764FFFEDB81D85F7CF1EF471F84A" box="[937,1037,1919,1945]" name="Thailand" pageId="35" pageNumber="36">Thailand</collectingCountry>
(
<collectingRegion id="8F38F83DFFEDB81D8245CF1EF400F84A" box="[1051,1148,1919,1945]" country="Thailand" name="Sara Buri" pageId="35" pageNumber="36">Saraburi</collectingRegion>
and
<collectingRegion id="8F38F83DFFEDB81D82EDCF1EF561F84A" box="[1203,1309,1919,1945]" country="Thailand" name="Krung Thep Mahanakhon" pageId="35" pageNumber="36">Bangkok</collectingRegion>
),
<collectingCountry id="35EB764FFFEDB81D8371CF1EF514F84A" box="[1327,1384,1919,1945]" name="Laos" pageId="35" pageNumber="36">Laos</collectingCountry>
, and
<collectingCountry id="35EB764FFFEDB81D86C9CFC2F0A8F86E" box="[151,212,1955,1981]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
. The remaining taxa will be described in an upcoming revision of the genus.
</paragraph>
<paragraph id="4D4336DFFFEDB81A8699CFA6F20CFE92" blockId="35.[151,1437,1343,2017]" lastBlockId="36.[151,1437,151,501]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<bibRefCitation id="296D4B2EFFEDB81D8699CFA6F1B9F832" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." box="[199,453,1990,2017]" pageId="35" pageNumber="36" pagination="639 - 655" refId="ref37329" refString="Di Palma, A., de Moraes, G. J., Gerdeman, B. S., Huber, S., Kitajima, E. W. &amp; Alberti, G. (2015) Ultrastructural and functional adaptations of the female reproductive system in the family Heterozerconidae (Acari, Anactinotrichida, Gamasida, Heterozerconina) and implications for the systematic position of the group. Arthropod Structure &amp; Development, 44, 639 - 655. https: // doi. org / 10.1016 / j. asd. 2015.09.002" type="journal article" year="2015">
Di Palma
<emphasis id="7F88EACDFFEDB81D8766CFA6F116F832" box="[312,362,1991,2017]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. (2015)
</bibRefCitation>
, in a study of the anatomy of the female genital system, found no indication of secondary genital openings (solenostomes) in females of an unidentified
<taxonomicName id="8AFC4D5CFFEAB81A852AC8F6F43FFF62" box="[884,1091,151,177]" pageId="36" pageNumber="37">
<emphasis id="7F88EACDFFEAB81A852AC8F6F390FF62" box="[884,1004,151,177]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
species
</taxonomicName>
from
<collectingRegion id="8F38F83DFFEAB81A82D8C8F6F4BDFF62" box="[1158,1217,151,177]" country="Philippines" name="Cebu" pageId="36" pageNumber="37">Cebu</collectingRegion>
island,
<collectingCountry id="35EB764FFFEAB81A8349C8F6F5E5FF62" box="[1303,1433,151,177]" name="Philippines" pageId="36" pageNumber="37">Philippines</collectingCountry>
. Examination of all available slide mounted
<taxonomicName id="8AFC4D5CFFEAB81A8429C8DAF293FF06" authorityName="Vitzthum" authorityYear="1926" box="[631,751,187,213]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8429C8DAF293FF06" box="[631,751,187,213]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
</taxonomicName>
females also failed to find any indication of secondary genital openings either near the opisthogastral suckers (as in
<taxonomicName id="8AFC4D5CFFEAB81A84B7C8BEF303FF2B" authorityName="Berlese" authorityYear="1888" box="[745,895,223,248]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A84B7C8BEF303FF2B" box="[745,895,223,248]" italics="true" pageId="36" pageNumber="37">Heterozercon</emphasis>
</taxonomicName>
or
<taxonomicName id="8AFC4D5CFFEAB81A85FDC8BEF4F8FF2A" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" box="[931,1156,223,249]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A85FDC8BEF4F8FF2A" box="[931,1156,223,249]" italics="true" pageId="36" pageNumber="37">Narceoheterozercon</emphasis>
</taxonomicName>
) or posterior to coxae IV (as in
<taxonomicName id="8AFC4D5CFFEAB81A8685C962F1E0FECE" authorityName="Fain" authorityYear="1989" box="[219,412,259,285]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8685C962F1E0FECE" box="[219,412,259,285]" italics="true" pageId="36" pageNumber="37">Afroheterozercon</emphasis>
</taxonomicName>
). In the absence of evidence to the contrary, we assume that
<taxonomicName id="8AFC4D5CFFEAB81A821DC962F56FFECE" box="[1091,1299,259,285]" pageId="36" pageNumber="37">
<emphasis id="7F88EACDFFEAB81A821DC962F4C7FECE" box="[1091,1211,259,285]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
species
</taxonomicName>
do not have a secondary genital system in the females.
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699C92AF3C0FE5A" blockId="36.[151,1437,151,501]" pageId="36" pageNumber="37">
Immatures of an
<taxonomicName id="8AFC4D5CFFEAB81A87DFC92AF22CFEB6" box="[385,592,331,357]" pageId="36" pageNumber="37">
<emphasis id="7F88EACDFFEAB81A87DFC92AF185FEB6" box="[385,505,331,357]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
species
</taxonomicName>
from
<collectingRegion id="8F38F83DFFEAB81A84CFC92AF285FEB6" box="[657,761,331,357]" country="Thailand" name="Krung Thep Mahanakhon" pageId="36" pageNumber="37">Bangkok</collectingRegion>
,
<collectingCountry id="35EB764FFFEAB81A855CC92AF31AFEB6" box="[770,870,331,357]" name="Thailand" pageId="36" pageNumber="37">Thailand</collectingCountry>
were partially described in
<bibRefCitation id="296D4B2EFFEAB81A82CBC92AF5E4FEB6" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[1173,1432,331,357]" pageId="36" pageNumber="37" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEAB81A8351C92AF542FEB6" box="[1295,1342,331,357]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (2018)
</bibRefCitation>
, but a complete description of
<taxonomicName id="8AFC4D5CFFEAB81A87BBC90EF221FE5A" authorityName="Vitzthum" authorityYear="1926" box="[485,605,367,393]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A87BBC90EF221FE5A" box="[485,605,367,393]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
</taxonomicName>
immatures is not yet available.
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699C9F2F12AFE26" blockId="36.[151,1437,151,501]" pageId="36" pageNumber="37">
<taxonomicName id="8AFC4D5CFFEAB81A8699C9F2F201FE7E" authority="Blaszak 1984" authorityName="Blaszak" authorityYear="1984" box="[199,637,403,429]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8699C9F2F143FE7E" box="[199,319,403,429]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEAB81A8717C9F2F19FFE7E" author="Blaszak, C." box="[329,483,403,429]" pageId="36" pageNumber="37" pagination="587 - 601" refId="ref37228" refString="Blaszak, C. (1984) Three new genera of zerconid mites (Acari, Mesostigmata, Zerconidae) from the United States of America with a key to the American genera. Polskie Pismo Entomologiczne, 53, 587 - 601." type="journal article" year="1984">Błaszak 1984</bibRefCitation>
(Zerconidae)
</taxonomicName>
is a junior homonym of
<taxonomicName id="8AFC4D5CFFEAB81A85FCC9F2F4B1FE7E" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[930,1229,403,429]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A85FCC9F2F466FE7E" box="[930,1050,403,429]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEAB81A827DC9F2F4B1FE7E" author="Vitzthum, H. &amp; Graf" box="[1059,1229,403,429]" pageId="36" pageNumber="37" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="296D4B2EFFEAB81A8280C9F2F0AEFE02" author="Kemal, M. &amp; Kocak, A. O." pageId="36" pageNumber="37" pagination="147 - 148" refId="ref38442" refString="Kemal, M. &amp; Kocak, A. O. (2009) A replacement name for Blaszakia Kocak &amp; Kemal, 2008 in the family Zerconidae (Acarina, Mesostigmata). Miscellaneous Papers of the Centre for Entomological Studies Ankara, 147 - 148, 10." type="book chapter" year="2009">Kemal &amp; Koçak 2009</bibRefCitation>
;
<bibRefCitation id="296D4B2EFFEAB81A8681C9D6F1A2FE02" author="Kocak, A. O. &amp; Kemal, M." box="[223,478,439,465]" pageId="36" pageNumber="37" pagination="1 - 6" refId="ref38707" refString="Kocak, A. O. &amp; Kemal, M. (2008) Nomenclatural notes on the genus group names of the order Acarina. Miscellaneous Papers of the Centre for Entomological Studies Ankara, 145, 1 - 6." type="journal article" year="2008">Koçak &amp; Kemal 2008</bibRefCitation>
). The zerconid genus name was replaced by
<taxonomicName id="8AFC4D5CFFEAB81A8584C9D6F12EFE26" authority="Kemal &amp; Kocak (Gerdeman et al. 2018)" class="Arachnida" family="Zerconidae" genus="Blaszakzercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8584C9D6F407FE02" box="[986,1147,439,465]" italics="true" pageId="36" pageNumber="37">Blaszakzercon</emphasis>
Kemal &amp; Koçak (
<bibRefCitation id="296D4B2EFFEAB81A8313C9D6F136FE26" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." pageId="36" pageNumber="37" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEAB81A8690C9BAF17CFE26" box="[206,256,475,501]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. 2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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