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<document ID-CLB-Dataset="6710" ID-DOI="http://dx.doi.org/10.3897/dez.63.10241" ID-GBIF-Dataset="0a6965a7-b124-4871-95d4-9e35c49073de" ID-Pensoft-Pub="1860-1324-2-211" ID-Pensoft-UUID="51243BBBE9F158F58BBC9BF3EA964017" ID-Zenodo-Dep="167458" ID-ZooBank="45552C4EC6AE4F9499980C2D492333B3" ModsDocID="1860-1324-63-2-211" checkinTime="1621372567443" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2016" docId="B6338877213E40F656322D1F28D54757" docLanguage="en" docName="DeutEntomolZeit 63(2): 211-270" docOrigin="Deutsche Entomologische Zeitschrift 63 (2)" docPubDate="2016-11-18" docSource="http://dx.doi.org/10.3897/dez.63.10241" docTitle="Cyphocoleus monteithi Liebherr 2016, sp. n." docType="treatment" docUuid="B88459B2-D6F8-486A-B85A-37EA2EA0135A" docUuidSource="ZooBank" docVersion="5" id="51243BBBE9F158F58BBC9BF3EA964017" lastPageNumber="236" masterDocId="51243BBBE9F158F58BBC9BF3EA964017" masterDocTitle="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia" masterLastPageNumber="270" masterPageNumber="211" pageNumber="236" updateTime="1691866435949" updateUser="ExternalLinkService">
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<mods:title id="E822E834DE18B005CC593CAFBE65BB12">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
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<mods:namePart id="2FB2CF0518352103BFB6BD0906A14BBF">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="DCEB81ACDF484CAECDB321037721743C">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:date id="2A7A608463A06BD6B6AB6DEDDCE51BA8">2016</mods:date>
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<mods:classification id="892A93497F0A42927ED2BEA466135C85">journal article</mods:classification>
<mods:identifier id="E851FA2EEA50C305AC8B9987E6951233" type="DOI">http://dx.doi.org/10.3897/dez.63.10241</mods:identifier>
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<treatment id="B6338877213E40F656322D1F28D54757" ID-GBIF-Taxon="182223713" LSID="urn:lsid:zoobank.org:act:B88459B2-D6F8-486A-B85A-37EA2EA0135A" httpUri="http://treatment.plazi.org/id/B6338877213E40F656322D1F28D54757" lastPageNumber="236" pageId="25" pageNumber="236">
<subSubSection id="D1108F3080864600823FE8E3658F1EF3" pageId="25" pageNumber="236" type="nomenclature">
<paragraph id="607255A6F5251FE62BFB43757361A860" pageId="25" pageNumber="236">
8.
<taxonomicName id="A60C3B4CAB30F71363CE8B0CE4587534" ID-CoL="6C8YQ" LSID="http://zoobank.org/B88459B2-D6F8-486A-B85A-37EA2EA0135A" authority="Liebherr, 2016" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus monteithi" order="Coleoptera" pageId="25" pageNumber="236" phylum="Arthropoda" rank="species" species="monteithi" status="sp. n.">Cyphocoleus monteithi</taxonomicName>
<taxonomicNameLabel id="12B9B3AB7A58B750E3F3A959BC784DFA" pageId="25" pageNumber="236">sp. n.</taxonomicNameLabel>
<figureCitation id="304C6205CBEAE638F8FB7C31E38A2FBD" captionStart="Figures 6065" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="25" pageNumber="236">Figures 64</figureCitation>
<figureCitation id="49E95BC87CAF65034A1F1260536C280F" captionStart="Figures 6675" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="25" pageNumber="236">, 72</figureCitation>
<figureCitation id="7E0BBE49A09E15F230C7BFF89EFE0789" captionStart="Figures 7681" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="25" pageNumber="236">, 79</figureCitation>
<figureCitation id="1AD13A161985F0E1F4C8A9E262D88176" captionStart="Figures 8285" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="25" pageNumber="236">, 85</figureCitation>
<figureCitation id="A05E1B1D160C17586084D71CEA8FA373" captionStart="Figures 8698" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="25" pageNumber="236">, 88</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="70BF31EDA615B6EF6257B5036206DE3C" pageId="25" pageNumber="236" type="diagnosis">
<paragraph id="78F6C4C3646986FED028981050F42017" pageId="25" pageNumber="236">Diagnosis.</paragraph>
<paragraph id="232CFE7E7408AF916A2AC5B100F55695" pageId="25" pageNumber="236">
This species (Fig.
<figureCitation id="2E3D48BAD590D11A0A22BF2B58D5EAB2" captionStart="Figures 8285" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="25" pageNumber="236">85</figureCitation>
) shares the very elongate prothorax and narrow head with
<taxonomicName id="BC03254EBA37EBA4230BA30CE6A9C89F" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus prolixus" order="Diptera" pageId="25" pageNumber="236" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="9AFE9DA56778B56284C8D8B0A9EA6F73" italics="true" pageId="25" pageNumber="236">Cyphocoleus prolixus</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="6557BD97829BD4F3414539DF7EB72E3B" captionStart="Figures 4750" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="25" pageNumber="236">48</figureCitation>
), however the pronotal disc is transversely wrinkled and the pronotal front angles are acutely extended as in
<taxonomicName id="F49124F83F5D1F375363679912BE5813" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus miricollis" order="Diptera" pageId="25" pageNumber="236" phylum="Arthropoda" rank="species" species="miricollis">
<emphasis id="76D46860490B18D29A5E599D80EAA385" italics="true" pageId="25" pageNumber="236">Cyphocoleus miricollis</emphasis>
</taxonomicName>
below. These characters support this
<normalizedToken id="C4B685F9492258DBA5D22678DD1CF1A6" originalValue="species">species'</normalizedToken>
phylogenetic intercalation subordinate to the former, and as adelphotaxon to the latter plus all cladistically associated species (Fig.
<figureCitation id="195BF8FBDAB8C66582D21F4B86DE8D79" captionStart="Figures 4445" captionStartId="F8" captionText="Figures 44 - 45. Parsimony cladograms comprising exemplar taxa of Odacanthini used in phylogenetic analysis, rooted at the outgroup Celaenephes linearis (tribe Lebiini): 44, 852 - step length strict consensus of 29 equally parsimonious trees of 835 - step length (CI = 22, RI = 74) [A = subtribe Actenonycina; H = subtribe Homethina subtrib. n.; P = subtribe Pentagonicina; O = subtribe Odacanthina]; 45, majority-rule consensus of 29 equally parsimonious trees with all resolved nodes present in 100 % of the multiple equally parsimonious trees; i. e. the majority-rule consensus is the result of collapsing 8 nodes. Abbreviations for geographical distributions include: Af, Africa; As, Asia; Eu, Europe; NC, New Caledonia; Nea, Nearctic; Neo, Neotropical; NZ, New Zealand; Oz, Australia." figureDoi="10.3897/dez.63.10241.figures44-45" httpUri="https://binary.pensoft.net/fig/111545" pageId="25" pageNumber="236">45</figureCitation>
). The elytra are distinctly ovate and very convex, domed, the lateral intervals and margins very depressed relative to the disc. This is also the first of the species to be treated in this revision that bears an environmental patina: in this species a varnish-like coating that covers the body surface of the beetles, this coating deepest in depressed areas such as the transverse pronotal wrinkles and elytral striae. Body surfaces on the mouthparts, antennae, and at the base of the head between the constricted neck and anterior prothoracic margin are dramatically clear of this varnish. Standardized body length 8.5-10.5 mm.
</paragraph>
</subSubSection>
<subSubSection id="85296F6E2743BB35509F94A627C43029" pageId="25" pageNumber="236" type="description">
<paragraph id="18C650E4149229361C2FBFA34E11446B" pageId="25" pageNumber="236">Description</paragraph>
<paragraph id="C393512415B2DA62187CF15717BE9092" pageId="25" pageNumber="236">
(n = 5). Head capsule ellipsoid, genal surfaces behind eyes subparallel, then converging to distinctly constricted neck; frons with broad medial crest between eyes, fine arcuate wrinkles curving outward and posterad from crest, frontal grooves parallel, broadly depressed from position between hind margins of eyes to clypeus, broadest just posterad frontoclypeal suture; eyes small, convex, 20 ommatidia across horizontal diameter, ocular ratio 1.39-1.50; supraorbital setae absent; mandibles moderately elongate, length 2.4
<normalizedToken id="48CED78CA0FAB634C1545736FA817858" originalValue="×">x</normalizedToken>
distance from antennal articulatory socket to anterolateral margin of labrum; antennae moderately elongate, scape length 2.6
<normalizedToken id="00D66D42ADAD0FE546179E38A1F65BCE" originalValue="×">x</normalizedToken>
maximal breadth; gena setose, subgenal seta present just laterad gula about 2/3 distance from hind margin of eye to constricted neck. Pronotum elongate, tubular, MPW/PL = 0.59-0.69, base constricted relative to apex, APW/BPW = 1.28-1.40; median longitudinal impression finely and shallowly incised, causing interruptions in orientation of transverse wrinkles in some instances; anterior transverse impression broad, shallow, extended laterally to behind front angles; proepisternum bulging, extended beyond marginal bead of notum, with distinct, deeply engraved vertical wrinkles; prosternal process distinctly depressed medially anterad procoxae, flat to narrowly depressed medially on ventral surface, narrowly convex on posterior surface. Elytra inflated, based inflexed at scutellum and depressed along suture, intervals 3-5 convexly elevated; striae smooth with very widely spaced micropunctures along length (view cleaned specimen); parascutellar seta present, articulatory socket papillate, elevated above surrounding elytral surface; dorsal elytral setae absent; lateral elytral setae arranged as 6 + 7, their articulatory sockets slightly upraised above surrounding cuticle, often appearing clear of cuticular varnish; subapical and apical elytral setae present, articulatory sockets papillate and clear of varnish; subapical sinuation moderately concave, meeting lateral margin in broad curve; elytral apices rounded, moderately separated by invaginated margin at fused suture. Mesepisternum impunctate, metepisternal dorsal length 1.1
<normalizedToken id="C352E80EE7127A005E96DA67E7205C02" originalValue="×">x</normalizedToken>
diagonal width. Abdomen with apical ventrite broadly, slightly concave medially; males with one seta each side apical margin of apical ventrite, females with two setae each side. Legs gracile, moderately elongate, mt1 length/tibial length = 0.27; metacoxae bisetose; tarsomeres with dorsal surface glabrous; mt4 with length to apex of outer lobe 1.8
<normalizedToken id="0FB74D6F3F9C49CCDDCB785CE68E0D8F" originalValue="×">x</normalizedToken>
median tarsomere length, 3-4 ventrolateral setae each side. Microsculpture of head transverse, dense; pronotal and elytral disc with fine transverse lines; pro-, meso-, and metasternum plus abdominal ventrites with sparse pelage of extremely short microsetae, such extremely short setae also visible on head on frons, pronotum, and elytra (though cleaning the surface may remove setae). Coloration of head piceous, mandibles rufobrunneous, palps and antennae rufoflavous; pronotum, elytra and body ventrites and coxae piceous; femora piceous with rufobrunneous base and apex, tibiae rufopiceous dorsally, rufobrunneous on lateral and ventral surfaces, tibiae rufobrunneous.
</paragraph>
<paragraph id="F2D23DB80120384360EEA2231D657E30" pageId="25" pageNumber="236">
<emphasis id="3A8B3C3AE6ACE217CCD6C3B7017EF849" italics="true" pageId="25" pageNumber="236">Male genitalia</emphasis>
(n = 6). Male aedeagal median lobe robust, broadly parallel sided from base to apex of ostial opening, dorsoventral breadth at midlength 0.25
<normalizedToken id="4841CA6CF45EF389ED147A7FAECC5D9A" originalValue="×">x</normalizedToken>
distance from tip to base of closed basal bulb (Fig.
<figureCitation id="A8E5A2755799433BE69E4CC610ED7D88" captionStart="Figures 8698" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="25" pageNumber="236">88</figureCitation>
); lobe shaft slightly melanized, the internal sac visible through lobe wall in uneverted position; lobe apex parallel-sided, length distad ostial opening subequal to dorsoventral breadth; tip of lobe evenly, narrowly rounded.
</paragraph>
<paragraph id="785FA8CD97B5068A50237D81D5CA3295" pageId="25" pageNumber="236">
<emphasis id="31A8FECE84797C76BE1E5758F77F4E20" italics="true" pageId="25" pageNumber="236">Female reproductive tract</emphasis>
(n = 1). Bursa copulatrix vase-shaped, basally stalked, distance from base of gonocoxites to spermathecal basal sclerite 1.8
<normalizedToken id="32E480C0B50AD4B05B048F5382258707" originalValue="×">x</normalizedToken>
maximum breadth (dissection compressed under cover slip) (Fig.
<figureCitation id="F200B8E5DD5C2B085CA8E59E459F2D44" captionStart="Figures 6065" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="25" pageNumber="236">64</figureCitation>
); bursal walls thin, translucent basally, thicker and more heavily stained with Chlorazol Black stain near midlength, lumen without spicules, but bursal surface wrinkled in distal half; basal gonocoxite with apical fringe of seven to eight setae (Fig.
<figureCitation id="77522589FEF5C80DDF2C3FF55519BC49" captionStart="Figures 6675" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="25" pageNumber="236">72</figureCitation>
); apical gonocoxite acuminate with narrowly rounded apex and three lateral ensiform setae.
</paragraph>
</subSubSection>
<subSubSection id="C77AD75C109BCF1CB654A2C2A9AAA504" pageId="25" pageNumber="236" type="types">
<paragraph id="C477991F1A8DBE44E6CA94F6FFA8E13B" pageId="25" pageNumber="236">Types.</paragraph>
<paragraph id="C3600F6A78E1C580C3DCD5F1EC9AE4DB" pageId="25" pageNumber="236">
Holotype male (QMB deposited in MNHN): NEW CALEDONIA
<emphasis id="48658389A5C45E4A6153B956B72990F3" bold="true" pageId="25" pageNumber="236">11152</emphasis>
/ 22°02
<normalizedToken id="A11888FED7DA5B9A73FA03542331DD6D" originalValue="Sx166°28">
'Sx
<geoCoordinate id="4F3E2597D5E76F2F1F29301F8E3284E2" degrees="166" direction="east" minutes="28" orientation="longitude" precision="925" value="166.46666">166°28'E</geoCoordinate>
</normalizedToken>
. 950m. / Dzumac Rd., junction / 26Jan2004. G.Monteith / pyrethrum, trees &amp; logs //
<emphasis id="9BF47E41E6FD2E4B1965C8875A3F36A4" bold="true" pageId="25" pageNumber="236">QUEENSLAND / MUSEUM LOAN</emphasis>
/ DATE: April 2004 / No. LE 04.16 (green label) // male habitus photo 1 / J.K.
<bibRefCitation id="CF411D85BB1D8A9C34AFFCFD6E425ABD" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="49" pageNumber="260" pagination="1 - 407" publicationUrl="10.3897/zookeys.544.6074" refId="B33" refString="Liebherr, JK, 2015. The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1 - 407, DOI: 10.3897/zookeys.544.6074" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation." url="10.3897/zookeys.544.6074" volume="544" year="2015">Liebherr 2015</bibRefCitation>
// HOLOTYPE /
<taxonomicName id="725EBDFC0B5C1AEE4EF888A295F6F890" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="25" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="6FEC9F7AC87BC5D524835EC0981FECBA" italics="true" pageId="25" pageNumber="236">Cyphocoleus</emphasis>
</taxonomicName>
/
<taxonomicName id="949A398362AB4F6E84B06DEC955438FE" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="monteithi" order="Diptera" pageId="25" pageNumber="236" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="0E2E98F629BFC37AA02B2EA5E771827A" italics="true" pageId="25" pageNumber="236">monteithi</emphasis>
</taxonomicName>
/ J.K. Liebherr 2016 (black-bordered red label).
</paragraph>
<paragraph id="18FE72115738F9C581DA501E36E54E59" pageId="25" pageNumber="236">
Paratypes (22 specimens). NEW CALEDONIA: Mt. Ouin, 1100 m el., pyrethrum trees &amp; logs,
<geoCoordinate id="DC1FC0DA1E543D4594904034E4599CB3" degrees="22" direction="south" minutes="01" orientation="latitude" precision="925" value="-22.016666">22°01'S</geoCoordinate>
,
<geoCoordinate id="1EF1FC4A2CFD8DB72830B9DC41259FF3" degrees="166" direction="east" minutes="28" orientation="longitude" precision="925" value="166.46666">166°28'E</geoCoordinate>
, 09-xi-2002, Burwell &amp; Monteith (QMB, 1; lot no. 11150); Dzumac Road junction, 950 m el.,
<geoCoordinate id="64CCF3BD50ED670A05423ABC8C077E37" degrees="22" direction="south" minutes="02" orientation="latitude" precision="925" value="-22.033333">22°02'S</geoCoordinate>
,
<geoCoordinate id="6EDC2A1BB0D0AC416ABB46CC838A178F" degrees="166" direction="east" minutes="28" orientation="longitude" precision="925" value="166.46666">166°28'E</geoCoordinate>
, 09-xi-2002, Burwell, Monteith &amp; Wright (QMB, 1; lot no. 11145), pyrethrum trees &amp; logs, 09-xi-2002 Burwell &amp; Monteith (QMB, 3; lot no. 11143), pyrethrum mossy trees, 04-xii-2003, Monteith (QMB, 3; lot no. 11458), pyrethrum trees &amp; logs, 26-i-2004, Monteith (QMB, 8; lot no. 11522), flight intercept trap, 05-xii-2003-26-i-2004, Monteith (QMB, 1; lot no. 11465); Mt. Dzumac road, 700 m el., pyrethrum trunks &amp; logs,
<geoCoordinate id="828C3140335AED1E24E9D5F395B1239B" degrees="22" direction="south" minutes="03" orientation="latitude" precision="925" value="-22.05">22°03'S</geoCoordinate>
,
<geoCoordinate id="B750B726189807ACC0EA2DB0434B3C8A" degrees="166" direction="east" minutes="28" orientation="longitude" precision="925" value="166.46666">166°28'E</geoCoordinate>
, 01-xii-2000, Monteith (QMB, 1; lot no. 9913); Mt. Mou summit, 1200 m el.,
<geoCoordinate id="6C8B36C2E0EEA5985986640F7538E82C" degrees="22" direction="south" minutes="04" orientation="latitude" precision="925" value="-22.066668">22°04'S</geoCoordinate>
,
<geoCoordinate id="EE38128134BE99427D4BA44364ABC6A6" degrees="166" direction="east" minutes="21" orientation="longitude" precision="925" value="166.35">166°21'E</geoCoordinate>
, 24-v-1984, Monteith &amp; Cook (ANIC, 1; QMB, 1), moss forest, 1200 m el., hand collecting, 27-28-xii-2004, Monteith (QMB, 1; lot no. 12019);
<normalizedToken id="BC4B9F102D7558E02B84910EB5A21AE1" originalValue="Rivière">Riviere</normalizedToken>
Bleue, Haute Pourina, 800 m el., pyrethrum trees &amp; logs,
<geoCoordinate id="B34D0D511E08FFA015E73096F9E1C371" degrees="22" direction="south" minutes="06" orientation="latitude" precision="925" value="-22.1">22°06'S</geoCoordinate>
,
<geoCoordinate id="4D13EE930B77D53C692928A2CED742AE" degrees="166" direction="east" minutes="38" orientation="longitude" precision="925" value="166.63333">166°38'E</geoCoordinate>
, 22-xi-2002, Monteith (QMB, 1; lot no. 11210).
</paragraph>
</subSubSection>
<subSubSection id="8A656A60D8BE00C2CCEA2FBA05CBAFB3" pageId="25" pageNumber="236" type="etymology">
<paragraph id="7258F2259F5A9D4ECD6547B675B546BE" pageId="25" pageNumber="236">Etymology.</paragraph>
<paragraph id="0178F97807CCED7A9FA4709A77885F18" pageId="25" pageNumber="236">
I take great pleasure in naming this extremely distinctive species (Fig.
<figureCitation id="74D2438993E3A113CD788D9E3E0FF476" captionStart="Figures 8285" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="25" pageNumber="236">85</figureCitation>
) for Dr. Geoff Monteith, Senior Curator of Insects Emeritus, Queensland Museum. His collecting activities involving numerous field expeditions to New Caledonia formed the taxonomic basis for this study.
</paragraph>
</subSubSection>
<subSubSection id="F4694909311478BEDEA50E9C7655DCC7" pageId="25" pageNumber="236" type="distribution">
<paragraph id="8264A9C003E07A14FBDBCB2435B12EA5" pageId="25" pageNumber="236">Distribution and habitat.</paragraph>
<paragraph id="5AD5B9D0FEEEA5D641244BDBEE672E39" pageId="25" pageNumber="236">
This species is known only from the southern portion of Grande Terre: localities range from Mt. Ouin on the north to Parc Provincial
<normalizedToken id="312219D65795C73D7C88A16796D2FEA6" originalValue="Rivière">Riviere</normalizedToken>
Bleue on the south (Fig.
<figureCitation id="9FFD3CBEF02AFA5928EA6AE03733D264" captionStart="Figures 7681" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="25" pageNumber="236">79</figureCitation>
). Collection localities range from 700-1200 m el., with beetles collected via pyrethrin fog of trees and logs, hand collecting, and in a flight intercept trap into which one beetle was able to climb.
</paragraph>
</subSubSection>
</treatment>
</document>