536 lines
67 KiB
XML
536 lines
67 KiB
XML
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<mods:title id="ABACEED8FD2EBFB0C4E56F7EC6BFCFA9">Notes on the genus Liebstadia Oudemans, 1906 (Acarina, Oribatida) in Central Europe</mods:title>
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<mods:date id="32C5B19AE9F6D13BA3B4633421F77B92">1996</mods:date>
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<treatment id="B624D40484E7952C7C2C602BF37A897A" ID-DOI="http://doi.org/10.5281/zenodo.6280793" ID-GBIF-Taxon="100112126" ID-Zenodo-Dep="6280793" LSID="urn:lsid:plazi:treatment:B624D40484E7952C7C2C602BF37A897A" httpUri="http://treatment.plazi.org/id/B624D40484E7952C7C2C602BF37A897A" lastPageId="25" lastPageNumber="98" pageId="21" pageNumber="94">
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<subSubSection id="48DC9A40108E084322B2866D907D31D8" pageId="21" pageNumber="94" type="nomenclature">
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<paragraph id="A8CC0E73B4FC87DDD1F8EB29CD6F7069" pageId="21" pageNumber="94">
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[Genus
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<taxonomicName id="61242964B9B636233B20626EFFC79755" ID-CoL="62X3X" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia OUDEMANS</taxonomicName>
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]
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</paragraph>
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</subSubSection>
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<subSubSection id="B7A1CB41AFEFA87EC95C77B49EB0D7C2" lastPageId="23" lastPageNumber="96" pageId="21" pageNumber="94" type="discussion">
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<paragraph id="C14FF0E5A76784A7218508991F34CDE4" pageId="21" pageNumber="94">Discussion</paragraph>
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<paragraph id="6BEDA5605BCFED4E1A091DB33640B815" pageId="21" pageNumber="94">
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The special lamellar-sublamellar complex with the lamella touching the Insertion point of interlamellar seta is regarded as a specific and apomorphic character of the genus
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<taxonomicName id="F29C9A5BBDAD9FE2C295E2CEE2ABDF8D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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. Other characters with diagnostic value are less specific and obviously often developed convergently in different genera within the
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<taxonomicName id="16D1E865BFC85478F48C5F854840E3C2" pageId="21" pageNumber="94" rank="superFamily" superFamily="Oripodoidea">Oripodoidea</taxonomicName>
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: i.e. having prolamella and sublamella, 10 notogastral setae, 3-4 areae porosae. 4 genital setae, legs monodactyl - some of which are considered typical for scheloribatid mites.
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</paragraph>
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<paragraph id="B687B823921030E3253CD144A4D0D96A" pageId="21" pageNumber="94">
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The monodactyly of the
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<taxonomicName id="98B7D55FB3165FE272D6C7B0E8446E20" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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species is (along with the habitual similarity in dorsal view) the reason that some of them have been placed within the genus
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<taxonomicName id="89176D0380FE1D792531646BB8B581AD" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
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BERLESE, 1908. This is the only generic character given by Berlese (1908) to separate
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<taxonomicName id="1C1B76F232FAC9711B4A5F44EB89E5FA" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
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from
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<taxonomicName id="B4378CC96445D0F1DE89AFCABC8E369F" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Scheloribates</taxonomicName>
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. Recent research has shown that some
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<taxonomicName id="A1672E8199A8804B54CFFD3C4C24F918" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
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auct. species in the literature belong to the revised genus
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<taxonomicName id="7400B5F26A7E0DDA4A4435E72D6076A4" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
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(Weigmann et al. 1993), and some other species with the specific lamellar complex ("
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<taxonomicName id="DBBC7A47363E6F194BE7EEDF0D3900BD" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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type") belong to
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<taxonomicName id="AC11E4458F85C8C88BE76D24BBB7913D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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as defined in this paper, in accordance with Weigmann (1969) and Wunderle et al. (1990). Additional species of "
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<taxonomicName id="DECA7482C5B08B49C8BB79242027F726" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
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",
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<taxonomicName id="ADE0FC7A35E88C1270055DA5E1530109" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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and other related genera, described worldwide, will likely belong to the genus
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<taxonomicName id="1DA08B8D4CC0AD4F7631B313CABD5614" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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as described here. However, usually no lateral drawings present in the literature, and for this reason it is very difficult to determinate the generic Status. Further more, at least in some recently described species of the genus
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<taxonomicName id="900AD0DB6156E49C9BDAD83037DFEEC7" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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the position in the genus cannot be proven from the original description (see discussion of the species
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<taxonomicName id="3A7DF308D2EDB1EB19FA39280E6E5E46" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="963" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
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below).
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</paragraph>
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<paragraph id="96A51EE41218578EEDD4C60D10A05D58" pageId="21" pageNumber="94">
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As we do not intend to produce a worldwide revision of the
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<taxonomicName id="27A54DF77E6364633DA939B3907E4800" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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species we have focused only on the Central European species on which we have enough information. Some other species are discussed, using the original descriptions and redescriptions from the literature, or (as in the case of Willmann's species
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<taxonomicName id="2C9305EA417F61218F2D11EDCD297EC3" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="novus">Protoribates novus</taxonomicName>
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and
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<taxonomicName id="192E11E390F2B1CEC8C25A3A8338DC8C" class="Arachnida" family="Protoribatidae" genus="Protoribates" genus.bestMatchDistance="769" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="austriacus">P. austriacus</taxonomicName>
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) on the basis of the original material, which is unfortunately not sufficient for redescription. Generally, the most important fact is that the always immovable pteromorphs of
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<taxonomicName id="EDD776B6FF53FC583678A9C77909B235" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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species show a high diversity in shape, forming a coutinuous row from small toothlike blades (
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<taxonomicName id="C7D82302C5FCA5500BCDF5E0101FF8B9" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="1308" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
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.
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<taxonomicName id="42E11C2E304C33C91B3519BE2481AB02" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="861" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="similis">L. similis</taxonomicName>
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), over small rounded blades, not bent downwards (
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<taxonomicName id="84D9294F9168AD5EBC6634BFC31F7861" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="2366" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="longior">L. longior</taxonomicName>
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) to large pteromorphs bent downwards (
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<taxonomicName id="0CE43CEF6D7E3B591A623FFE803740A8" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="2149" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="pannonica">L. pannonica</taxonomicName>
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,
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<taxonomicName id="9EB61EB6BF94AA0062507C5176C10AE9" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="2228" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="willmanni">L. willmanni</taxonomicName>
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<taxonomicNameLabel id="E32715F984CF93048C0E99A639E267D5" pageId="21" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
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), without recognizable gaps (see also Perez-Inigo 1993). Therefore, the shape of pteromorphs cannot be used as a generic character. This result raises the immediate question of which diagnostic character of the pteromorph development should be used in the other scheloribatid genera, e.g.
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<taxonomicName id="970D9630F6494414B58BB8CD415B70C5" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Scheloribates</taxonomicName>
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vs.
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<taxonomicName id="61572E481A65B29949EAA8A09A03D462" class="Arachnida" family="Hemileiidae" genus="Hemileius" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Hemileius</taxonomicName>
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, as the use of this character for family separation seems to be unacceptable (e.g. Perez-Inigo 1993). This question needs further detailed investigation and will be discussed in a later paper.
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</paragraph>
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<paragraph id="A96E3BE596AA0D8D6D60FF8DE343A1F5" pageId="21" pageNumber="94">
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<taxonomicName id="06D9237BEB1DBE8066DCF28BE292C459" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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belongs to the family
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<taxonomicName id="23280E548026EAB900740A6D35FF5D8E" family="Scheloribatidae" pageId="21" pageNumber="94" rank="family">Scheloribatidae</taxonomicName>
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GRANDJEAN, 1933 in the conservative view of Grandjean (1958). Balogh and Balogh (1992) put
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<taxonomicName id="271C6F752FF7C57C1790189AEEB273F7" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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into the family
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<taxonomicName id="0C1ED06C346F447ED5DF0C2E1C7CA236" family="Protoribatidae" pageId="21" pageNumber="94" rank="family">Protoribatidae</taxonomicName>
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BALOGH and BALOGH. 1984, which we believe to be a mixed composition of members from different families. The nominal genus
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<taxonomicName id="6AEBA6FDA28E076A4C0C88769363BAAD" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
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belongs to
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<taxonomicName id="A09734B5DC6483A167395DD9743A8FA9" family="Haplozetidae" pageId="21" pageNumber="94" rank="family">Haplozetidae</taxonomicName>
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, while
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<taxonomicName id="62C0B163A287C5077A3E462041345124" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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definitely does not (see Weigmann et al. 1993 for further discussion).
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</paragraph>
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<paragraph id="1C134170AE0B5506D3AD85208783313A" pageId="21" pageNumber="94">
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Except for the continual change of pteromorph shapes, two trends are easily recognizable: the changes in dorsoventral thickness and the form and shape of the sensillus (see figs 1c, 5b, 12a, 17b). Considering the known facts on the ecology of the different species, we recognized two typical "life forms" of
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<taxonomicName id="AA137921FC0218AD19934ED266F4403A" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
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:
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</paragraph>
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<paragraph id="551A6DB8337C36F2640B13C08098B158" pageId="21" pageNumber="94">
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(a) the species with large body size, dorsoventrally thick, with a longer sensillus stalk and moreless lanceolate sensillus head (e.g.
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<taxonomicName id="1BAF957BCF0D581B4F8514E41566F90A" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="560" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="similis">L. similis</taxonomicName>
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).
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</paragraph>
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<paragraph id="AA0EF7A3F2D8CF4B7FDCDFE35DE97609" pageId="21" pageNumber="94">
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(b) the species with very flat, oblonged body of medium size, with the sensillus stalk short and sphaerical or sub-sphaerical sensillus head (
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<taxonomicName id="97036D2518187F314D2E94AC63F095EC" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="1212" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
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.
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<taxonomicName id="BEFFF369F7398ADE93731581E741A674" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="2043" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="longior">L. longior</taxonomicName>
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).
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</paragraph>
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<paragraph id="D67551E0596523B8DDDD825E1650C473" pageId="21" pageNumber="94">
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In addition to the two mentioned forms intermediates could also be found (e.g.
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<taxonomicName id="8C7C937FBAA5059D66FDA0C623FD995D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="1814" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="pannonica">L. pannonica</taxonomicName>
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|
). Probably, the life-form (a) belongs to the species living in the litter, soil surface or surface layer of the soil, and the life-form (b) seems to be an adaption for life in flat moss/lichen growths or the crevices and slots of tree bark. The intermediate forms were found mostly in grasslands or in shrub formations.
|
|
</paragraph>
|
|
<paragraph id="D6B21520E579721EA6A410F3034ED876" lastPageId="22" lastPageNumber="95" pageId="21" pageNumber="94">
|
|
Some other "
|
|
<taxonomicName id="3F6529C0F0FEBBE56693716ED239ACAC" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Protoribates</taxonomicName>
|
|
" auct. species known from central Europe, are considered to be in fact
|
|
<taxonomicName id="4F0195AD4A90B33D96EAD206BC69880D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
. Both individuals of
|
|
<taxonomicName id="69BD618AB3F584BD5B52172D5F0805E9" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="nova">Liebstadia nova</taxonomicName>
|
|
(WILLMANN 1953)
|
|
<taxonomicNameLabel id="BA95C6435F0DDB190A002EB9F691D093" pageId="21" pageNumber="94">comb. nov.</taxonomicNameLabel>
|
|
(=
|
|
<taxonomicName id="047C32E18FFF55181565370BDFA1BBC5" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="novus">Protoribates novus</taxonomicName>
|
|
WILLMANN, 1953) studied from Willmanns original slide (Staatssammlungen Munich, labeled: "K6-91,
|
|
<taxonomicName id="72DFAD396EF1A0A619B3F99633D184AC" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="novus">Protoribates novus</taxonomicName>
|
|
<taxonomicNameLabel id="C2600E92F241CC18871C59CC8B92E896" pageId="21" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
|
|
, det. C. Willmann, G 294'') axe strongly damaged, and no longer usable for redescription. Nevertheless. from the slide it is clear that the species has no 11
|
|
<pageBreakToken id="5AE0DD1C0A449E87A1DA99712E080343" pageId="22" pageNumber="95" start="start">notogastral</pageBreakToken>
|
|
setae. The eleventh seta in Willmann's description is most probably the lyrifissure, which is also visible on the damaged specimens in the slide. From the characters visible, the species belongs to
|
|
<taxonomicName id="33C3C35777870853D0FB934E37F425DC" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
as defined here, and probably is a synonym of
|
|
<taxonomicName id="9B7DE657F491FCCF74EC4D4B8296978D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="1595" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="pannonica">L. pannonica</taxonomicName>
|
|
. A definitive classification requires the study of topotypical material. Mahunka (1987) reported on the species from
|
|
<normalizedToken id="3B669BF29C33460CBECF82EC539DF0C2" originalValue="Kiskunság" pageId="22" pageNumber="95">Kiskunsag</normalizedToken>
|
|
National Park in Hungary, but without notes on specific characters. The species was reported as being sylvan (found in plant association Junipero-Populetum albae).
|
|
</paragraph>
|
|
<paragraph id="F0CB41D0B55123C8419009321F11020C" pageId="22" pageNumber="95">
|
|
The classification of
|
|
<taxonomicName id="544936594D33C68135BF2CE0E44D77A2" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="austriaca">Liebstadia austriaca</taxonomicName>
|
|
(WILLMANN, 1953)
|
|
<taxonomicNameLabel id="08683490495EF7B0A92F6C24EEF3AA90" pageId="22" pageNumber="95">comb. n.</taxonomicNameLabel>
|
|
(=
|
|
<taxonomicName id="287C93CED9ED8023135B2CA504E85082" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="novus">Protoribates novus</taxonomicName>
|
|
WILLMANN, 1953) is very similar. The original material of Willmann (3 slides from Staatssammlungen Munich labeled: "K6-68 (E4/0-3)
|
|
<taxonomicName id="70CA2C60756A1D7E07884EF34A52FB3A" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="austriacus">Protoribates austriacus</taxonomicName>
|
|
<taxonomicNameLabel id="F8EFE209C73617DEFDB3C05B64A0989E" pageId="22" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
|
|
", "K6-69
|
|
<taxonomicName id="9990156B3969C8A16B3F6C23A58D647D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
, A 77/0-3, Admont, det. E. Leitner" with "
|
|
<taxonomicName id="56457D530341A356A4662D25472EFCE2" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="austriacus">Protoribates austriacus</taxonomicName>
|
|
" added by pencil, and "K6-70
|
|
<taxonomicName id="64D428E5E50B91C73F71A0D002018A20" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="austriacus">Protoribates austriacus</taxonomicName>
|
|
<taxonomicNameLabel id="AAA6ABB1FA95B430F2B3FC8D85FF2B71" pageId="22" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
|
|
") cannot be used for redescription, but from the crushed individuals in the slides it is clear, that it is a species of
|
|
<taxonomicName id="C981B4A84D27D6982646468C9FB81942" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
, according to the present description. Moreover, considering the body shape and form of the sensillus, the species is probably also a synonym of the presently redescribed
|
|
<taxonomicName id="149090FB0DF92C4CB28F92E912682E23" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="1383" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="pannonica">L. pannonica</taxonomicName>
|
|
. As in previous species, definitive classification is possible only after the study of topotypical material.
|
|
</paragraph>
|
|
<paragraph id="06A05BD7A233227F84822644701DC758" pageId="22" pageNumber="95">
|
|
The species
|
|
<taxonomicName id="12ADDC08E96C5B0E20B29EB678D82E0D" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="serratomarginatus">Protoribates serratomarginatus</taxonomicName>
|
|
, described by Mahunka (1983) from the
|
|
<normalizedToken id="E7AC508167B220683E188423C9B1FA33" originalValue="Hortobágy" pageId="22" pageNumber="95">Hortobagy</normalizedToken>
|
|
National Park in Hungary is definitely
|
|
<taxonomicName id="809EACC83C12E39BF1ACE101E5091FFC" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
(see figs 30 and 32 of original description). The species is nearly identical (from comparison of available description and drawings) with the nominal species
|
|
<taxonomicName id="99E406430CA7958223BAC470DCB59900" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="similis">Liebstadia similis</taxonomicName>
|
|
. Typical characters of the species, as given by author, are: serrated to denticulated edge of pteromorphs. comparatively long notogastral and prodorsal setae and body size. We did not study the Hungarian material, but the description and pictures closely resemble our material from Germany (2 specimens - 1 female and 1 male from Leipzig, 14. 7. 1991, leg. W. Dunger), which belongs to
|
|
<taxonomicName id="64D5336C5129EC1D47FACEC62733D9FE" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="similis">Liebstadia similis</taxonomicName>
|
|
sensu lato. We also note the remark of Wunderle et al. (1990), that the border of the pteromorphs of
|
|
<taxonomicName id="929EDDE9A507E503916F297368E4864B" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="78" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="similis">L. similis</taxonomicName>
|
|
is slightly undulated or denticulated. The only other notable difference is in body size: Mahunka gives a length of 431 - 498
|
|
<normalizedToken id="0F60888609FC16051B0F8B513A4011DD" originalValue="µm" pageId="22" pageNumber="95">ym</normalizedToken>
|
|
and a width of 245-302
|
|
<normalizedToken id="06974A13442E9CE9BE212A55342042FE" originalValue="µm" pageId="22" pageNumber="95">ym</normalizedToken>
|
|
for
|
|
<taxonomicName id="09C03E29B8D0BF4F8F6FBBA2CAF9C4A5" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="serratomarginatus">Protoribates serratomarginatus</taxonomicName>
|
|
, while the measurements for
|
|
<taxonomicName id="7247ECD1784A8BBB3C30E304C2283A6A" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="166" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="similis">L. similis</taxonomicName>
|
|
are 500- 565 x 330-360
|
|
<normalizedToken id="57B72A0E5CDD0CBEBFBEB2CD403B8728" originalValue="µm" pageId="22" pageNumber="95">ym</normalizedToken>
|
|
(Wunderle et al., 1990). Measurements of specimens studied from Leipzig were more or less intermediate: length 500
|
|
<normalizedToken id="B3302E4B66D5DF4B2B071D15CD45373F" originalValue="µm" pageId="22" pageNumber="95">ym</normalizedToken>
|
|
(male) and 525
|
|
<normalizedToken id="891EE8B2BF383D2676CE775FC42A3F7A" originalValue="µm" pageId="22" pageNumber="95">ym</normalizedToken>
|
|
(female). respectively, and width 320
|
|
<normalizedToken id="3236CC600A00CC7C99A771A300008197" originalValue="µm" pageId="22" pageNumber="95">ym</normalizedToken>
|
|
, so the body size is not usable as a differential character. Remaining characters, i.e. slender body and comparatively longer notogastral and prodorsal setae, are insufficient to establish a species. Differences between this form and
|
|
<taxonomicName id="1CB4997FF0CEC5A65AB821BDCE62C778" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="72" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="similis">L. similis</taxonomicName>
|
|
are in the same range as differences between the nominate form and form with long notogastral and prodorsal setae, described here ("
|
|
<taxonomicName id="9EE8E3490F2C3FFF9A4204376FD425F3" class="Arachnida" family="Liebstadiidae" forma="longisetosa" genus="Liebstadia" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="forma" species="similis">longisetosa</taxonomicName>
|
|
"). Also, the identity of a Hungarian form and "
|
|
<taxonomicName id="B3F253E7A94DB6F008EA1386AF27F849" class="Arachnida" family="Liebstadiidae" forma="longisetosa" genus="Liebstadia" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="forma" species="similis">longisetosa</taxonomicName>
|
|
" is not fully excluded. All the questions mentioned could be solved by the further study of the Hungarian material. At present stage of our knowledge the name
|
|
<taxonomicName id="BBDAEF3AD655181686C756A158DDE540" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="similis" subSpecies="serratomarginata">Liebstadia similis serratomarginata</taxonomicName>
|
|
<taxonomicNameLabel id="97DAD587071DEB8AA33787F99B9A9EDF" pageId="22" pageNumber="95">comb. nov.</taxonomicNameLabel>
|
|
,
|
|
<taxonomicNameLabel id="BBF4470512886758F675B5E29A8D4DD4" pageId="22" pageNumber="95">stat. nov.</taxonomicNameLabel>
|
|
is proposed for the Hungarian form.
|
|
</paragraph>
|
|
<paragraph id="0ECDE94F304F7F0D4DEFFD52BAE9D978" pageId="22" pageNumber="95">
|
|
Mahunka (1987) described the species
|
|
<taxonomicName id="B0C48FBC8B564072E46DC00C755A7D77" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
ahumerata from the juniper litter from
|
|
<normalizedToken id="0359079B548D0E681EEBF72FB4835E22" originalValue="Kiskunság" pageId="22" pageNumber="95">Kiskunsag</normalizedToken>
|
|
Nat. Park. Hungary. The differential diagnosis in the description is lacking. The placement of the species in
|
|
<taxonomicName id="89CEA9527F438AE88746832209780C5F" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
is probable, but in the original drawing of the lateral prodorsum (fig. 53) the interlamellar seta does not touch the lamella. As seen from the dorsal view (fig. 51), the setae in cannot be placed in so large distance from the lamella, as shown in the fig. 53, so very probably the proximal (right) seta in is omitted and instead the distal (left) seta is depicted. If the Position of seta in is on the lamella, the species could be placed in
|
|
<taxonomicName id="7C10AF5E90114126FDA399EF3F482A3F" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
. In this case the difference from other similar
|
|
<taxonomicName id="24E3943E273960DA599DB4FA1680A137" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
species must be defined. The author states in the description that the species has only three pairs of small areae porosae (similarly as
|
|
<taxonomicName id="6455F83C2F931BABF43D9F1B31BB1B00" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="1011" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="longior">L. longior</taxonomicName>
|
|
.
|
|
<taxonomicName id="EF6ED01CDAFA75C1D3A884227C2DAE1B" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="172" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
|
|
or
|
|
<taxonomicName id="262FDDB1D53C9A7771A122BD9337EEF0" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="879" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="willmanni">L. willmanni</taxonomicName>
|
|
<taxonomicNameLabel id="98C9B3515693063555C581BFFC04D69F" pageId="22" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
|
|
), on the other hand, the depicted specimen (fig. 51) has four pairs of areae (along with the small size, and somewhat different shape of the sensillus head it could be a specific character). To determine, if the species is or is not a synonym of some of the known species, further evaluation must be done.
|
|
</paragraph>
|
|
<paragraph id="951FC333FE9A166BFC8CC9CE2E1C306C" lastPageId="24" lastPageNumber="97" pageId="22" pageNumber="95">
|
|
Several addidonal species of
|
|
<taxonomicName id="558C3BABA0CED0B0C4122471BC0D7ED1" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
were reported from the Mediterranean area and the Balkan Penninsula. According to Perez-Inigo (1993), 5 species of
|
|
<taxonomicName id="A9ECB1506B8DF207D7A0C18D23068930" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
are present in the Spanish fauna. The species
|
|
<taxonomicName id="33E4D80312A1974A52FA64340A850336" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="57" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
|
|
, sensu the mentioned papers differs from the species, as redescribed by Wunderle et al. (1990) by its 4 pairs of areae porosae and possibly (according to fig. 77B) by longer interamellar and lamellar setae. On the other hand, it is nearly identical with
|
|
<taxonomicName id="8F7ACBD5E6410FB0167417EAFCFCFED0" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="humerata">Liebstadia humerata</taxonomicName>
|
|
sensu Seniczak (1990, fig. 9). Because no more details of adult morphology are given in hoth papers (especially the lateral view: bothridial area and pteromorphs, ventral side,
|
|
<pageBreakToken id="00E1AADFFFA5AA8304C7EF41E28919D2" pageId="24" pageNumber="97" start="start">leg</pageBreakToken>
|
|
chaetotaxy etc.), the real position of these species is uncertain. The body shape, position of seta lp, sensillus shape and the classification of bothridia is very similar to the new species
|
|
<taxonomicName id="809980EDEC1590EEA3ABECE2C6A2983E" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="635" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="willmanni">L. willmanni</taxonomicName>
|
|
(especially the form "A" from Slovakia), described here. On the other hand, the presence of 4 pairs of areae porosae resembles that of the species
|
|
<taxonomicName id="3EC7C6187F82FCFD583877F3B1D0DC28" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="517" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="pannonica">L. pannonica</taxonomicName>
|
|
. It is not excluded, that presence of 3 pairs of areae in
|
|
<taxonomicName id="F6D9F142B79A5EB1B804FCE4A06CEC4B" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="384" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
|
|
or
|
|
<taxonomicName id="AC65DF75D8694045A80F9BA5DF5AFA38" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="579" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="willmanni">L. willmanni</taxonomicName>
|
|
could be variable, and that populations with 4 pairs could exist (see also the opinion of Wunderle et al. 1990). More detailed study may also show, that
|
|
<taxonomicName id="AA5A11232B79FA90426A043357D1192B" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="504" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
|
|
sensu Seniczak and
|
|
<taxonomicName id="DD63FE520A7C8A90BBA5E29793219687" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="522" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
|
|
sensu Perez-Inigo represent a new, undescribed species of
|
|
<taxonomicName id="E29F02A21E6E1910C1077B447F7BA4F4" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
. The next two species from Spain (
|
|
<taxonomicName id="6393A241FC49057F087D58D13EC94B1F" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="gallardoi">L. gallardoi</taxonomicName>
|
|
and
|
|
<taxonomicName id="986128B9883384E00B592C27887AC721" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="microptera">L. microptera</taxonomicName>
|
|
) are most probably Mediterranean forms, not present in Central Europe. The pteromorphs of last two species are developed similarly to
|
|
<taxonomicName id="0948E2ACC83625E324D04001675AD78D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="610" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="97" phylum="Arthropoda" rank="species" species="longior">L. longior</taxonomicName>
|
|
, however, the differences in number of areae porosae, shape of sensillus and prodorsal setae are present.
|
|
</paragraph>
|
|
<caption id="E860736A0F77CB208D3F20E25C749B17" pageId="23" pageNumber="96">
|
|
<paragraph id="9CA30775AAF7EC7AEB2A5A5F2FE9F7EB" pageId="23" pageNumber="96">
|
|
<pageBreakToken id="573464E078CF30A0730E6AD8F8DBBBA1" pageId="23" pageNumber="96" start="start">Fig</pageBreakToken>
|
|
. 20: Differences of length/width ratios in studied specimens of
|
|
<taxonomicName id="CBE91A9A12FDAFB6CEEF4E90254FE4D9" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="366" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="pannonica">L. pannonica</taxonomicName>
|
|
,
|
|
<taxonomicName id="661F4F9676A6702246A5432A8B1DC777" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="445" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="willmanni">L. willmanni</taxonomicName>
|
|
and
|
|
<taxonomicName id="92D0F2D950EAEE7CE1F8A488CADAFC87" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="565" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="longior">L. longior</taxonomicName>
|
|
(linear regression).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="F8E44D725A30C19F7D7904148DD34EBF" pageId="23" pageNumber="96">
|
|
<paragraph id="B8D6374D6902ABE60B1FDB6867FE14B0" pageId="23" pageNumber="96">
|
|
Fig. 21: Comparison of length/width ratios of 3 studied species and
|
|
<taxonomicName id="0C916FEE255EAA8477AD7B5966F9A98D" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="494" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="humerata">L. humerata</taxonomicName>
|
|
(data after Wunderle et al. 1990).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="E092DC13859DD408EDD318689708F5A3" pageId="23" pageNumber="96">
|
|
The species
|
|
<taxonomicName id="3F56A9357DCD16FA701F159A071A3768" class="Arachnida" family="Protoribatidae" genus="Protoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="gratiosus">Protoribates gratiosus</taxonomicName>
|
|
VASILIU et CALUGAR (1973) is most likely also a species of the genus
|
|
<taxonomicName id="A80397ED8A9A29AE02A8EEAF14D07E3E" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
(but the lateral view is absent in the description, only the formulation "... et les poils interlamellaires fixes au milieu de la lamelle ..." is present in the description). If the species is in fact
|
|
<taxonomicName id="A958EE4600B83C24DD8062BF098887C9" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
, the correct name should be
|
|
<taxonomicName id="59E5D2E249D03D2160ECC03D516BF0F6" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="gratiosa">Liebstadia gratiosa</taxonomicName>
|
|
(VASILIU et CALUGAR 1973)
|
|
<taxonomicNameLabel id="867F121777EFC35FC4607359D587EB9E" pageId="23" pageNumber="96">comb. nov.</taxonomicNameLabel>
|
|
In this case the species stands near the new species
|
|
<taxonomicName id="B0028DDFD87FEBE7B93837CB41DDE9D4" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.bestMatchDistance="315" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="willmanni">L. willmanni</taxonomicName>
|
|
. The differences are in the length of prodorsal setae, the shape of bothridium, the shape of sensillus, and general body length and form: the presence of only 9 notogastral setae (seta lp missing) seems to be improbable.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</pageBreakToken>
|
|
<subSubSection id="314BD9B86FC1BE8B7A428F004085EFCF" lastPageId="25" lastPageNumber="98" pageId="23" pageNumber="96" type="key">
|
|
<paragraph id="2683C35E85C6E98801005BA9F2C97D7D" pageId="23" pageNumber="96">
|
|
In conclusion, the following key could be used for the determination of Central-European species of
|
|
<taxonomicName id="EEA3C972ADF5E54759B2C3B6314A40BA" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="genus">Liebstadia</taxonomicName>
|
|
(see also the table 1 with characters of studied species):
|
|
</paragraph>
|
|
<paragraph id="6044C101F20FD14A4F27B9A75C26012C" pageId="23" pageNumber="96">
|
|
1 Body more or less elongated and flat, ratio length:dorsoventral thickness over 3.3:1. Three pairs of areae porosae developed usually, sensillus with short stalk and globular, globular-fusiform or fusiform head. Bothridial lamellae (bl, bl') missing or indistinct in lateral view. Length 310-440
|
|
<normalizedToken id="0AAC2AED09B89B35E74FCE4EC42876BC" originalValue="µm" pageId="23" pageNumber="96">ym</normalizedToken>
|
|
..............................................................3
|
|
</paragraph>
|
|
<paragraph id="1C26FA3BC78740FC69902BEFF5F1FBBB" pageId="23" pageNumber="96">
|
|
- Body more robust, not flat, ratio length:dorsoventral thickness under 3.1:1. Four pairs of areae porosae present, sensillus with longer stalk and fusiform or lanceolate head. Two bothridial lamellae (bl, bl') well developed, bothridium usually with posterolateral lobe or scale. Length 356 - 600
|
|
<normalizedToken id="C697CC4B24923F3BE7E4DBC770314BAB" originalValue="µm" pageId="23" pageNumber="96">ym</normalizedToken>
|
|
...........................................................2
|
|
</paragraph>
|
|
<paragraph id="3B5ED3EB938B7CAD83B3B6EF8DDCBFA3" pageId="23" pageNumber="96">
|
|
2(1) Pteromorphs well developed, bent downwards, in lateral view reaching the acetabular area. Body size 350 - 430
|
|
<normalizedToken id="9772F854845443C8E778A3B1AD65B961" originalValue="µm" pageId="23" pageNumber="96">ym</normalizedToken>
|
|
, sensillus usually fusiform, rarely lanceolate ....................................................
|
|
<taxonomicName id="58CAE906411F1B7AAE290034DE022DFB" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="pannonica">Liebstadia pannonica</taxonomicName>
|
|
(WILLMANN, 1951)
|
|
</paragraph>
|
|
<paragraph id="35FF278F8E0D9012B15F5B929BE78136" pageId="23" pageNumber="96">
|
|
- Pteromorphs developed as horizontal humeral blades, not bent downwards. Body size over 430
|
|
<normalizedToken id="71424830B45A3D457745DE516499227E" originalValue="µm" pageId="23" pageNumber="96">ym</normalizedToken>
|
|
, sensillus lanceolate........................................
|
|
<taxonomicName id="FF82EA69E1FDAA5A394B03422C2D9C87" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="similis">Liebstadia similis</taxonomicName>
|
|
(MICHAEL, 1888)
|
|
</paragraph>
|
|
<paragraph id="1CE92A6706DEC97E3C9F81BFFCA3088B" pageId="23" pageNumber="96">
|
|
3 (1) Pteromorphs well developed, bent downwards, in lateral view reaching or almost reaching the acetabular region. Sensillus fusiform or globular-fusiform in lateral view. Ventral surface microstriated, adanal area with distinct adanal ridge. Body size 352 - 439
|
|
<normalizedToken id="27928C5698832B3075628A5348331635" originalValue="µm" pageId="23" pageNumber="96">ym</normalizedToken>
|
|
..................................................
|
|
<taxonomicName id="D983A29DE54636E3ADACB6BFB7A41928" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="willmanni">Liebstadia willmanni</taxonomicName>
|
|
<taxonomicNameLabel id="3AF0EFF2E011F9B1A94B4E6D7DACCD2B" pageId="23" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="E5C0C65F8D995C41FF4DC03B784CC202" pageId="23" pageNumber="96">
|
|
- Pteromorphs not bent downwards, slightly developed or absent. Sensillus with more-less globular head. Ventral surface without a clear microstriation. Body size 310 - 395
|
|
<normalizedToken id="735424237C21D0CAA24572B5A9369FDC" originalValue="µm" pageId="23" pageNumber="96">ym</normalizedToken>
|
|
...................................................4
|
|
</paragraph>
|
|
<paragraph id="2D95CF78EEBECA3850604BAACE32B572" pageId="23" pageNumber="96">
|
|
4(3) Pteromorphs developed as a small triangular tubercle in humeral area or nearly absent. Tarsus III with 12 setae, genu I with 2+1 setae. Length-width ratio 1.85 - 2.40 .......................................................................
|
|
<taxonomicName id="5C42323EA5228D7D6434902D228B80C4" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="humerata">Liebstadia humerata</taxonomicName>
|
|
SELLNICK, 1928
|
|
</paragraph>
|
|
<paragraph id="C78AB68C68EA2DFCBF57FDF3927E237C" pageId="23" pageNumber="96">
|
|
- Pteromorphs developed as distinct, rounded horizontal blades. Tarsus III with 15 setae, genu 1 with 3 + 1 setae. Length-width ratio 2.25 - 2.45, body extremely flat (length:dorsoventral thickness ratio over 4.5) ..............................................
|
|
<taxonomicName id="48DCA523C96DC620C7B83F17E9B0F3F1" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="96" phylum="Arthropoda" rank="species" species="longior">Liebstadia longior</taxonomicName>
|
|
(BERLESE, 1908)
|
|
</paragraph>
|
|
<paragraph id="830CC6C3AF153D15B0C31949C5A29BD4" pageId="25" pageNumber="98">
|
|
<table id="1AAF4967AF362AA0893BF954552E757C" border="1" frame="box" pageId="25" pageNumber="98" rules="all">
|
|
<tr id="07C2AA64496CAA25C6EABC152E0718DC" pageId="25" pageNumber="98">
|
|
<td id="D98FA66D777E2DC34905D5CAC13D4913" isEmpty="true" pageId="25" pageNumber="98">
|
|
<pageBreakToken id="05DC707A38C2A2A60A5B38D237C2B3AA" pageId="25" pageNumber="98" start="start">EmTaCe</pageBreakToken>
|
|
</td>
|
|
<td id="9C5DB0910206776AFC4D9CC3BFF83274" pageId="25" pageNumber="98">
|
|
<taxonomicName id="F7660B773B0155B0C37C06F29542810A" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="98" phylum="Arthropoda" rank="species" species="similis">similis</taxonomicName>
|
|
</td>
|
|
<td id="42E3344332419FD14F4BB5514A9396F5" pageId="25" pageNumber="98">
|
|
<taxonomicName id="F299C8D330A003B7851BCCBF2108177A" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="98" phylum="Arthropoda" rank="species" species="pannonica">pannonica</taxonomicName>
|
|
</td>
|
|
<td id="80A17A2B9543B3D6784D6BAF73868803" pageId="25" pageNumber="98">
|
|
<taxonomicName id="0D3141B12310174FE808023D80553E35" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="98" phylum="Arthropoda" rank="species" species="willmanni">willmanni</taxonomicName>
|
|
</td>
|
|
<td id="B8A0A44FE8B60235FBC410A219C636A4" pageId="25" pageNumber="98">
|
|
<taxonomicName id="D0B8447E6C30C6F1779CAD0C06DB452E" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="98" phylum="Arthropoda" rank="species" species="longior">longior</taxonomicName>
|
|
</td>
|
|
<td id="2A43442E02CEB1D0E15A71DE1CF14FF8" pageId="25" pageNumber="98">
|
|
<taxonomicName id="3FE037967EC1BFB64F54B54B5B728614" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="98" phylum="Arthropoda" rank="species" species="humerata">humerata</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="EF1CA9E79B33A0EBDA012B649286C2BF" pageId="25" pageNumber="98">
|
|
<td id="ECCE74E4CFD6591DE95E53785DCA8E55" pageId="25" pageNumber="98">body form</td>
|
|
<td id="49B96C5C74FE0D73D70951E87C106DD6" pageId="25" pageNumber="98">robust, wide, arched</td>
|
|
<td id="A0701E866144D1D19E2D1753CD1A7220" pageId="25" pageNumber="98">robust, moderately, arched</td>
|
|
<td id="C57DAA4D69A1863C922CF6307E5F37BE" pageId="25" pageNumber="98">slender, flat</td>
|
|
<td id="12CFEC6C6D181562550B07C22FD4C866" pageId="25" pageNumber="98">elongated, very flat</td>
|
|
<td id="A76B8482D4DFA54686A7017919B50F18" pageId="25" pageNumber="98">slender, flat</td>
|
|
</tr>
|
|
<tr id="345F72FCB2098B0FE5AA5BAF7C57CFDD" pageId="25" pageNumber="98">
|
|
<td id="2FFC50E9E1BC12FEDB46EAD4E2FA542E" pageId="25" pageNumber="98">length (L)</td>
|
|
<td id="885262B3E0E56D84AAB64BEE02A23CFF" pageId="25" pageNumber="98">500-600</td>
|
|
<td id="2DF006B83870527FBD7BC58D3DEF2B6E" pageId="25" pageNumber="98">350-430</td>
|
|
<td id="0FEA40CAAB082B9CC5687E1B17023112" pageId="25" pageNumber="98">352-439</td>
|
|
<td id="622063918D57D6D7AD6A288C185C38F2" pageId="25" pageNumber="98">325-395</td>
|
|
<td id="46CC18D3F1CD52259218887FFDC9DA23" pageId="25" pageNumber="98">310-380</td>
|
|
</tr>
|
|
<tr id="345049A090E6E1433B026EB5264844FF" pageId="25" pageNumber="98">
|
|
<td id="B04FA75F96FC7EA1CDCE0EF397FD3C0D" pageId="25" pageNumber="98">width (W)</td>
|
|
<td id="3900D6C3C098A87AB40AD04B1F254985" pageId="25" pageNumber="98">330-360</td>
|
|
<td id="75D17D9F6E44DB08ECD4F8BE510CA27A" pageId="25" pageNumber="98">185-250</td>
|
|
<td id="0842B4A698E748E54334D5527C0768A7" pageId="25" pageNumber="98">159-214</td>
|
|
<td id="960B9E05CFA4A28822493F7ACD7A3BE3" pageId="25" pageNumber="98">133-162</td>
|
|
<td id="E58521E9A839FDE90030A0F236D34C37" pageId="25" pageNumber="98">140-200</td>
|
|
</tr>
|
|
<tr id="255A11E54AE42F4B98B4D6F4DA985D95" pageId="25" pageNumber="98">
|
|
<td id="B86F343F02D1242570BF95D80A716E3A" pageId="25" pageNumber="98">thickness (T)</td>
|
|
<td id="1F1CF81CFC0311365B922E111F34F0AF" pageId="25" pageNumber="98">ca 240</td>
|
|
<td id="D6A7D39597C784B5FAC0D397B3FFCA6D" pageId="25" pageNumber="98">125-160</td>
|
|
<td id="47FC8CAD00ABADAC71EBC8AC0D498829" pageId="25" pageNumber="98">95-105</td>
|
|
<td id="1EAFEE2DE6F46973C7490D0DB50478C2" pageId="25" pageNumber="98">75-85</td>
|
|
<td id="A142A9638EA1E850A9E6727D94B59FD2" pageId="25" pageNumber="98">ca 90</td>
|
|
</tr>
|
|
<tr id="E3AE65C597E7FA3B96B437C1169FB0E5" pageId="25" pageNumber="98">
|
|
<td id="ED2D3F782B3A54C695558AF516E68453" pageId="25" pageNumber="98">L/W ratio</td>
|
|
<td id="2A4A35B971D5632B05B29449ADBCB6BE" pageId="25" pageNumber="98">1,45-1,60</td>
|
|
<td id="C43B4A3183C97179374DEA251B906433" pageId="25" pageNumber="98">1,7-1.9</td>
|
|
<td id="B6745222348E43E8C35F8FAA60442FF2" pageId="25" pageNumber="98">2,0-2,3</td>
|
|
<td id="BB84BA8467176B55E2B4DE9E466009D6" pageId="25" pageNumber="98">2,25-2,45</td>
|
|
<td id="6A3ABFFAD26CE7DB3851FDB9EB516265" pageId="25" pageNumber="98">1,85-2,40</td>
|
|
</tr>
|
|
<tr id="1C06DBB75FCCEA92C9D7C4530DA8BE80" pageId="25" pageNumber="98">
|
|
<td id="75A0663D8B48C7771EAD2C7EAA5DDBBC" pageId="25" pageNumber="98">L/T ratio</td>
|
|
<td id="AAF9B1066FEC6871BEF5B47AB217D659" pageId="25" pageNumber="98">about 2,2</td>
|
|
<td id="D768180D6868D27B8B375097DF7801E1" pageId="25" pageNumber="98">about 2,7</td>
|
|
<td id="0322CC416D3E8DFCA5DACE71F769656C" pageId="25" pageNumber="98">about 3,6</td>
|
|
<td id="AEED57D399740332E763CA9AFC750C83" pageId="25" pageNumber="98">about 4,9</td>
|
|
<td id="FF231D707EC81BCF166B999E1E189DAC" pageId="25" pageNumber="98">about 4,0</td>
|
|
</tr>
|
|
<tr id="DEEE535D3269BAB55C6AC5746D21E797" pageId="25" pageNumber="98">
|
|
<td id="B4CE3B753B38DAD3B038004F9E0E55F2" pageId="25" pageNumber="98">length of in</td>
|
|
<td id="81649909CA87662F2F5BADCF62727EDD" pageId="25" pageNumber="98">80-90</td>
|
|
<td id="8179A5ADE85E16059F1263A022593A04" pageId="25" pageNumber="98">45-56</td>
|
|
<td id="AC5405DD7C46ED1D8E2B02711EB9F25E" pageId="25" pageNumber="98">30-50</td>
|
|
<td id="A74B7E935B971D34D099485B45DC2392" pageId="25" pageNumber="98">30-40</td>
|
|
<td id="079F6CB1C64768934DCBFDA042D633F4" pageId="25" pageNumber="98">ca 40-50</td>
|
|
</tr>
|
|
<tr id="FB933F62FB4450C75ACF1CD990FF1C3C" pageId="25" pageNumber="98">
|
|
<td id="B3D5764CCD48A37A193497829C698DC6" pageId="25" pageNumber="98">length of ng setae</td>
|
|
<td id="2618E4B6613B367A93AC97E838658892" pageId="25" pageNumber="98">30-35</td>
|
|
<td id="02073A877F12E3D4917E6181F50291CD" pageId="25" pageNumber="98">18-32</td>
|
|
<td id="F16A142E1E36172559AE30A61C9AFC16" pageId="25" pageNumber="98">21-28</td>
|
|
<td id="7C9A1270166E7CFEDEF3BAAC6256F254" pageId="25" pageNumber="98">10-20</td>
|
|
<td id="2C36BB121E49A57B0D6B2F6A8AC7363F" pageId="25" pageNumber="98">20-25</td>
|
|
</tr>
|
|
<tr id="C118236E8016AAF3331ADD1A9771C37E" pageId="25" pageNumber="98">
|
|
<td id="9B95A5ED2E532DCFBC25D7A33E66C87A" pageId="25" pageNumber="98">h3 - A1/lp - A1 ratio</td>
|
|
<td id="AAFF235E20AA368ABBD24449B6E3A5C4" pageId="25" pageNumber="98">0,2-0,3</td>
|
|
<td id="7E649136226DCED0FB572AB923EEE52C" pageId="25" pageNumber="98">1,1-2,6</td>
|
|
<td id="4F9BD5B77B9A62BF875CEC8425F95977" pageId="25" pageNumber="98">2,5-7,5</td>
|
|
<td id="218E32F11D5F3B53F54FC3F15C8CD7B6" pageId="25" pageNumber="98">about 3</td>
|
|
<td id="4ACEC1B03B23A2EED973EB42E0EC3B46" pageId="25" pageNumber="98">about 2</td>
|
|
</tr>
|
|
<tr id="0564BD4D7913850E26B94D4E02D0A573" pageId="25" pageNumber="98">
|
|
<td id="DCEBA15C19369A197159E08AD01129BC" pageId="25" pageNumber="98">pairs of areae porosae</td>
|
|
<td id="13ED5A49D003705D4D330D72306157BE" pageId="25" pageNumber="98">4</td>
|
|
<td id="598209CF2974E32840EFC1549E0C53C4" pageId="25" pageNumber="98">4</td>
|
|
<td id="68372058A7ADD10706E20E83C4B06EF3" pageId="25" pageNumber="98">3 (rarery 4)</td>
|
|
<td id="6734CC410418222C14BF7AEB8F067029" pageId="25" pageNumber="98">3</td>
|
|
<td id="26ABFFFF3C096FF8A2FF5E0A6CA7FE6E" pageId="25" pageNumber="98">3</td>
|
|
</tr>
|
|
<tr id="9086902C51EDDEA3AD17EF4ED8D9DE1A" pageId="25" pageNumber="98">
|
|
<td id="E65D2A77EBF3FA0DA36CE5D66B5EF192" pageId="25" pageNumber="98">pteromorphs</td>
|
|
<td id="A5B026733940A6B4C813CA96FC472EAE" pageId="25" pageNumber="98">small, horizontal</td>
|
|
<td id="2B617153C6A377CC6F485DE0A3CBF9C6" pageId="25" pageNumber="98">well developed, bent downwards</td>
|
|
<td id="6F7515822BD0FB799CB5B1529F26975F" pageId="25" pageNumber="98">well developed, bent downwards</td>
|
|
<td id="72191F20BA1ECE30B67AD7286375A57C" pageId="25" pageNumber="98">horizontal blades</td>
|
|
<td id="32ECDE6C95C4863A46F347EE679BCD94" pageId="25" pageNumber="98">very small to absent</td>
|
|
</tr>
|
|
<tr id="4B8AC905E2A5C861064B5F9C6E27202D" pageId="25" pageNumber="98">
|
|
<td id="865B7B16ED9716B4A4CC93125842CB1C" pageId="25" pageNumber="98">length of sensilus, form of sens. head</td>
|
|
<td id="0AAC99677B2D2A7E0EF53EF9CA9E1B63" pageId="25" pageNumber="98">long, lanceolate</td>
|
|
<td id="F703954FA50EEA56245631072D68FBB8" pageId="25" pageNumber="98">medium to long, clavate</td>
|
|
<td id="82BE938BEEC94D5A1A314CAAD04060B1" pageId="25" pageNumber="98">medium, fusiform to subglobular</td>
|
|
<td id="FFF4F6F97B2CBC8902D6EC153B375393" pageId="25" pageNumber="98">Short, subglobular to globular</td>
|
|
<td id="569750378B23C34332D0DEAEF5CC8CBF" pageId="25" pageNumber="98">short, subglobular to globular</td>
|
|
</tr>
|
|
<tr id="D1E0B328AF1D1FED7CCB25E84CEECECE" pageId="25" pageNumber="98">
|
|
<td id="58F9B35653EB85BC7D47D178E1097346" pageId="25" pageNumber="98">setae on targus I</td>
|
|
<td id="6BEC88459430A09E6F069BF2050DD9FB" pageId="25" pageNumber="98">19+2</td>
|
|
<td id="7DDE1A9877492C49414528CF377FAC11" pageId="25" pageNumber="98">18+2</td>
|
|
<td id="8581FB2F06C32F74E8B804CB3E8C23A1" pageId="25" pageNumber="98">18+2</td>
|
|
<td id="DCD38514E34EAC292444C9B4B021C904" pageId="25" pageNumber="98">18+2</td>
|
|
<td id="C920AB5E423856462E860694EF0856C0" pageId="25" pageNumber="98">18+2</td>
|
|
</tr>
|
|
<tr id="444333EA7C69211A1518CD43E605B51C" pageId="25" pageNumber="98">
|
|
<td id="E032600597E6B52CAE9B065B09FA3E51" pageId="25" pageNumber="98">setae on targus II</td>
|
|
<td id="DDC943E974D61F1F6887EC926C12BC6D" pageId="25" pageNumber="98">15</td>
|
|
<td id="CFDC4E33870C48C265BA5078E4E7D1C8" pageId="25" pageNumber="98">15</td>
|
|
<td id="6387C02945130D3DA6D48596A1B6BD2B" pageId="25" pageNumber="98">15</td>
|
|
<td id="26CF17F6B5C1F967354E19F49C90AA96" pageId="25" pageNumber="98">15</td>
|
|
<td id="6675ADB6C54E093E8A57B904C89301A7" pageId="25" pageNumber="98">12</td>
|
|
</tr>
|
|
<tr id="7C1C047C4AE9F383AA7DDAEBFB2762AC" pageId="25" pageNumber="98">
|
|
<td id="25B0902BE7DBD476AF0C2134DB374352" pageId="25" pageNumber="98">special characters</td>
|
|
<td id="BF06F138292448C4B63E81E85F7E5BFC" isEmpty="true" pageId="25" pageNumber="98">EmTaCe</td>
|
|
<td id="76D1E75694A449827325A202A4C856BF" pageId="25" pageNumber="98">posterolateral scale on bothridium</td>
|
|
<td id="D7CF7BFE8AC25A65D663004F253F70FC" pageId="25" pageNumber="98">microstriated ventrally</td>
|
|
<td id="3F3E25DE259ADB0A889DD1AE99BA00A6" isEmpty="true" pageId="25" pageNumber="98">EmTaCe</td>
|
|
<td id="1993E62C7EB28C6E2FB4AFB80E150E27" pageId="25" pageNumber="98">genu I with 2+1 setae only</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FE7AA0DE042A9B85B7C87AC4F18073E7" pageId="25" pageNumber="98" type="multiple">
|
|
<caption id="051230C6C67AC1F618F810572BE4347F" pageId="25" pageNumber="98">
|
|
<paragraph id="E977F88DE7AD4DFA52E9B6F6813DF887" pageId="25" pageNumber="98">
|
|
Table 1: Characters of Central-European species of the genus
|
|
<taxonomicName id="DC85F19C9523226297106059CBFDE510" class="Arachnida" family="Liebstadiidae" genus="Liebstadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="98" phylum="Arthropoda" rank="genus">Leibstadia</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |