treatments-xml/data/03/E0/C8/03E0C852FFB46C6E23A90ED50DAD1A63.xml
2024-06-21 12:22:17 +02:00

217 lines
32 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="706983C9B671FEA0A60C4A23C395DDC6" ID-DOI="10.5281/zenodo.186299" ID-GBIF-Dataset="3c61a031-b0b6-4dda-ac37-a20e488ff85f" ID-ISSN="1175-5326" ID-Zenodo-Dep="186299" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461049713889" checkinUser="plazi" docAuthor="Penttinen, Jouni &amp; Jaschhof, Mathias" docDate="2009" docId="03E0C852FFB46C6E23A90ED50DAD1A63" docLanguage="en" docName="zt02032p054.pdf" docOrigin="Zootaxa 2032" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sylvenomyia Mamaev &amp; Zaitzev 1998" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="FFD9B02AFFB56C6C233E0F3A0E2E1E31" masterDocTitle="On the systematics of Sylvenomyia Mamaev &amp; Zaitzev (Diptera, Cecidomyiidae, “ Porricondylinae ”), with the description of a new species from Finland" masterLastPageNumber="54" masterPageNumber="48" pageNumber="49" updateTime="1698578325124" updateUser="plazi">
<mods:mods id="101A40D2D0EE05F97CEE81E5C72BB262" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BBF3CB7578A04751446931C7F4146F58">
<mods:title id="064388D4A9F2E8F8C48A2DB66CA3F37C">On the systematics of Sylvenomyia Mamaev &amp; Zaitzev (Diptera, Cecidomyiidae, “ Porricondylinae ”), with the description of a new species from Finland</mods:title>
</mods:titleInfo>
<mods:name id="1405FE952368F0E0A23A95460B544212" type="personal">
<mods:role id="B4624FCBE12C9626C1D95109A1FB5D68">
<mods:roleTerm id="C643852764CE11DEF34D569366F0D8BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C9B44DFD30E2F5228EEFBC83B87F2CB">Penttinen, Jouni</mods:namePart>
</mods:name>
<mods:name id="3D6308E5AE8AF02799B4701EC87322A4" type="personal">
<mods:role id="EA6354581BA9F51FF692C8DAA5EAE06A">
<mods:roleTerm id="DE34E3452724005649F2EB99E6A81C47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB2B522B7DF7D2532FA8088F662A45C3">Jaschhof, Mathias</mods:namePart>
</mods:name>
<mods:typeOfResource id="751444D662ABD72ED2DE44552BB8FE1A">text</mods:typeOfResource>
<mods:relatedItem id="71EAA671713995A986C9A7370D1D9066" type="host">
<mods:titleInfo id="38E73707EFFB9611DE32664C7D2312AB">
<mods:title id="BAB03CB701698324DD1120DA1B2AA315">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A871D95307989B753ED7BAC62A838D03">
<mods:date id="231E3741EFBD25EAD3211313915441A5">2009</mods:date>
<mods:detail id="C7387AC7E9208063A1E2F3AE602FF84A" type="volume">
<mods:number id="8CD8272140488BA1DF573E112CAB7AA7">2032</mods:number>
</mods:detail>
<mods:extent id="7EC870FBB44EABF3B741867CBAA619B0" unit="page">
<mods:start id="FEEECA43A19E52519E3805BB88FD07CF">48</mods:start>
<mods:end id="2972CD62CA8F05D4272207D2D7D95D95">54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EB2F904C5B9E31CA520EC3F7A7EEDE26">journal article</mods:classification>
<mods:identifier id="8B835202063590CDE90BD8B303402902" type="DOI">10.5281/zenodo.186299</mods:identifier>
<mods:identifier id="BC7C6769B245E98DC6B97D17995266FA" type="GBIF-Dataset">3c61a031-b0b6-4dda-ac37-a20e488ff85f</mods:identifier>
<mods:identifier id="BFEC4697542DF64FD1A5D65F59C9A856" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D30591FFE996BC3A0EB7C76A41EC1A96" type="Zenodo-Dep">186299</mods:identifier>
</mods:mods>
<treatment id="03E0C852FFB46C6E23A90ED50DAD1A63" ID-DOI="http://doi.org/10.5281/zenodo.6214793" ID-GBIF-Taxon="119611536" ID-Zenodo-Dep="6214793" LSID="urn:lsid:plazi:treatment:03E0C852FFB46C6E23A90ED50DAD1A63" httpUri="http://treatment.plazi.org/id/03E0C852FFB46C6E23A90ED50DAD1A63" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49">
<subSubSection id="C3532ACFFFB46C6D23A90ED50FF01C61" pageId="1" pageNumber="49" type="nomenclature">
<paragraph id="8BF67944FFB46C6D23A90ED50CEE1C38" blockId="1.[151,704,495,521]" box="[151,704,495,521]" pageId="1" pageNumber="49">
<heading id="D0BECE28FFB46C6D23A90ED50CEE1C38" bold="true" box="[151,704,495,521]" fontSize="11" level="1" pageId="1" pageNumber="49" reason="1">
<emphasis id="B93DA556FFB46C6D23A90ED50CEE1C38" bold="true" box="[151,704,495,521]" pageId="1" pageNumber="49">
Genus
<taxonomicName id="4C4902C7FFB46C6D23D00ED50CEE1C38" ID-CoL="7QQL" ID-ENA="2338634" authority="Mamaev &amp; Zaitzev 1998" authorityName="Mamaev &amp; Zaitzev" authorityYear="1998" box="[238,704,495,521]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB46C6D23D00ED50FA91C38" bold="true" box="[238,391,495,521]" italics="true" pageId="1" pageNumber="49">Sylvenomyia</emphasis>
<bibRefCitation id="EFD804B5FFB46C6D22B00ED50CEE1C38" author="Mamaev" box="[398,704,495,521]" pageId="1" pageNumber="54" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev &amp; Zaitzev 1998</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF67944FFB46C6D23A90D030FF01C61" blockId="1.[151,1246,569,625]" box="[151,478,569,592]" pageId="1" pageNumber="49">
<bibRefCitation id="EFD804B5FFB46C6D23A90D030F881C61" author="Mamaev" box="[151,422,569,592]" pageId="1" pageNumber="54" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev &amp; Zaitzev 1998</bibRefCitation>
: 211.
</paragraph>
</subSubSection>
<subSubSection id="C3532ACFFFB46C6D23A90D630AF01C41" box="[151,1246,601,625]" pageId="1" pageNumber="49" type="materials_examined">
<paragraph id="8BF67944FFB46C6D23A90D630AF01C41" blockId="1.[151,1246,569,625]" box="[151,1246,601,625]" pageId="1" pageNumber="49">
<emphasis id="B93DA556FFB46C6D23A90D630F031C40" bold="true" box="[151,301,601,625]" pageId="1" pageNumber="49">
<typeStatus id="54F2C7E6FFB46C6D23A90D630EE11C40" box="[151,207,601,625]" pageId="1" pageNumber="49">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C4902C7FFB46C6D220D0D630D0C1C41" authority="Mamaev &amp; Zaitzev 1998" authorityName="Mamaev &amp; Zaitzev" authorityYear="1998" box="[307,802,601,624]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="sueciae">
<emphasis id="B93DA556FFB46C6D220D0D630C3F1C41" box="[307,529,601,624]" italics="true" pageId="1" pageNumber="49">Sylvenomyia sueciae</emphasis>
<bibRefCitation id="EFD804B5FFB46C6D21290D630D0C1C41" author="Mamaev" box="[535,802,601,624]" pageId="1" pageNumber="54" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev &amp; Zaitzev 1998</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4C4902C7FFB46C6D20760D630AFD1C41" authority="Spungis 1985" authorityName="Spungis" authorityYear="1985" box="[840,1235,601,624]" class="Insecta" family="Cecidomyiidae" genus="Chastomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="spinigera">
<emphasis id="B93DA556FFB46C6D20760D630A181C41" box="[840,1078,601,624]" italics="true" pageId="1" pageNumber="49">Chastomera spinigera</emphasis>
<bibRefCitation id="EFD804B5FFB46C6D27020D630AFD1C41" author="Spungis" box="[1084,1235,601,624]" pageId="1" pageNumber="54" refString="Spungis, V. (1985) Gall midges of the subtribe Diallactina (Diptera, Cecidomyiidae) in Latvia. Latvijas Entomologs, 28, 38 - 53 (in Russian, with English summary)." type="journal article" year="1985">Spungis 1985</bibRefCitation>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C3532ACFFFB46C6D23A90DA50AD31D65" pageId="1" pageNumber="49" type="diagnosis">
<paragraph id="8BF67944FFB46C6D23A90DA50AD31D65" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
<emphasis id="B93DA556FFB46C6D23A90DA50F341C88" bold="true" box="[151,282,671,697]" pageId="1" pageNumber="49">Diagnosis.</emphasis>
The strong anterior portion of C (antC) extends clearly beyond R5 and is followed by a costal break; M and CuA1 are absent (
<figureCitation id="137265C1FFB46C6D211F0DFC0C491CD1" box="[545,615,710,736]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Fig. 1</figureCitation>
A). The antennae in males have 11 flagellomeres; the flagellomere nodes bear crenulate whorls of sensory hairs and hair-shaped translucent sensilla (Fig. 2A). The gonostylus bears a strong apical claw (
<figureCitation id="137265C1FFB46C6D22E50C2E0C0A1D1F" box="[475,548,788,814]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Figs 1</figureCitation>
B, 2B, C). The simple, small tegmen is largely membranous and of broadconical shape (
<figureCitation id="137265C1FFB46C6D227B0C000FA51D65" box="[325,395,826,852]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Figs 1</figureCitation>
D, 2C). Females and preimaginal stages of
<taxonomicName id="4C4902C7FFB46C6D20B40C000A301D62" box="[906,1054,826,851]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB46C6D20B40C000A301D62" box="[906,1054,826,851]" italics="true" pageId="1" pageNumber="49">Sylvenomyia</emphasis>
</taxonomicName>
spp. are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3532ACFFFB46C6E23F80C5A0DAD1A63" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49" type="description">
<paragraph id="8BF67944FFB46C6D23F80C5A0A861ABB" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
<emphasis id="B93DA556FFB46C6D23F80C5A0F9F1D4B" bold="true" box="[198,433,864,890]" pageId="1" pageNumber="49">Description. Head.</emphasis>
Occiput with short setae. Postocular bristles absent. One pair of large top setae (see
<bibRefCitation id="EFD804B5FFB46C6D23A90CB20F6B1D93" author="Panelius" box="[151,325,904,930]" pageId="1" pageNumber="54" refString="Panelius, S. (1965) A revision of the European gall midges of the subfamily Porricondylinae (Diptera: Itonididae). Acta Zoologica Fennica, 113, 1 - 157." type="journal article" year="1965">Panelius 1965</bibRefCitation>
). Postfrons convex, bilobate, asetose. Prefrons large, asetose. Eye bridge up to 4 ommatidia long. Remnants of larval stemmata not discernible. Antennal scape and pedicel subequal in size; scape setose ventrally; pedicel asetose; 11 flagellomeres, the 2 apical flagellomere fused; flagellomere nodes longer than necks, barrel-shaped, fully covered with microtrichia, 1 whorl of basal setae, 1 complete and 12 incomplete crenulate whorls of long sensory hairs, several hair-shaped translucent sensilla distally (Fig. 2A). Clypeus smaller than prefrons, projecting in lateral profile, with large setae. Maxillary palpus long, 4-segmented, setae long and curved or short and straight; first segment with hair-shaped translucent sensilla.
</paragraph>
<paragraph id="8BF67944FFB46C6D23FB0BAC0ACA1802" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
<emphasis id="B93DA556FFB46C6D23FB0BAC0F021A81" bold="true" box="[197,300,1174,1200]" pageId="1" pageNumber="49">Thorax.</emphasis>
Pronotum asetose. Scutum with sparse dorsocentral and lateral setae. Pleural setae absent. A portion in between laterotergite and mediotergite covered with strikingly large microtrichia.
<emphasis id="B93DA556FFB46C6D27E50B870B311AE6" bold="true" box="[1243,1311,1213,1239]" pageId="1" pageNumber="49">Wing</emphasis>
(
<figureCitation id="137265C1FFB46C6D26110B870B5C1AE6" box="[1327,1394,1213,1239]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Fig. 1</figureCitation>
A). Comparatively short and broad, about as long as body. Membrane covered with microtrichia and setae. AntC extending clearly beyond R5, followed by break; Sc long, evanescent apically; h absent; Rs with vertical rather than horizontal inclination; r-m + m-cu horizontal, in juxtaposition with R5; M and CuA1 absent; CuA2 slightly curved, weak or evanescent apically; CuP short, running very close to CuA2; 1 branch of A present, albeit short; dorsal setae present on Sc basally, stem vein, R, R1, R5, and CuA2; ventral setae absent. Pattern of sensory buds: R1, 2 distal; R5, 1 basal, 2 distal; further 1 mesal sensory bud on Sc. Halter comparatively short, stem and node subequal in length.
<emphasis id="B93DA556FFB46C6D21B60AF10CE51BD4" bold="true" box="[648,715,1483,1509]" pageId="1" pageNumber="49">Legs.</emphasis>
Foreleg slightly longer than body, femur shorter than tibia, tibia shorter than tarsus; first segment of foretarsus 1/3 the length of second segment, with short, blunt apical projection. Pretarsal claws slightly curved, with minute teeth at midlength. Empodia vestigial.
</paragraph>
<paragraph id="8BF67944FFB46C6D23FB09050AF61987" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
<emphasis id="B93DA556FFB46C6D23FB09050F671868" bold="true" box="[197,329,1599,1625]" pageId="1" pageNumber="49">Abdomen.</emphasis>
All sclerites covered sparsely with large setae. Pattern of tergal plaques not fully resolved, presumably 0/2/2/1/1/1/1/0.
<emphasis id="B93DA556FFB46C6D22DA095C0C4318B1" bold="true" box="[484,621,1638,1664]" pageId="1" pageNumber="49">Terminalia</emphasis>
(
<figureCitation id="137265C1FFB46C6D2142095C0CED18B1" box="[636,707,1638,1664]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Figs 1</figureCitation>
B, C, 2BD). St9 not traceble. Tg9 subtrapezoid, with straight apical margin. Gonocoxites with deep ventral emargination extending beyond midlength; ventral bridge membranous; posterior portion of gonocoxal apodeme (postGA) long, extending to ventrobasal gonocoxal margin; anterior portion of gonocoxal apodeme (antGA) moderately long; dorsal transverse bridge unsclerotized. Gonostylus slightly tapered towards apex, with strong, multipointed apical claw. Ejaculatory apodeme as long as gonocoxites, strongly sclerotized, with membranous, tulip-shaped cap apically. Ducts of accessory glands distinctive. Tegmen small, largely membranous, broad-conical, rounded apically, ventrolateral margins serrate or with scaly surface, apex weak, parameral apodemes directed ventroanteriorly. St10 weak, bilobate, pubescent, asetose. Cerci not or not much extending beyond tg9, setose.
</paragraph>
<paragraph id="8BF67944FFB46C6E23F808F80D271CFE" blockId="1.[151,1438,671,2012]" lastBlockId="2.[151,1437,152,1106]" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49">
<emphasis id="B93DA556FFB46C6D23F808F80FE619ED" bold="true" box="[198,456,1986,2012]" pageId="1" pageNumber="49">Systematic position.</emphasis>
<bibRefCitation id="EFD804B5FFB46C6D22E908F80D1319ED" author="Mamaev" box="[471,829,1986,2012]" pageId="1" pageNumber="54" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev and Zaitzev (1998)</bibRefCitation>
classified
<taxonomicName id="4C4902C7FFB46C6D20F408F90A7019ED" box="[970,1118,1987,2012]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB46C6D20F408F90A7019ED" box="[970,1118,1987,2012]" italics="true" pageId="1" pageNumber="49">Sylvenomyia</emphasis>
</taxonomicName>
with the
<taxonomicName id="4C4902C7FFB46C6D27DB08F80B5519ED" box="[1253,1403,1986,2012]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
, a decision presumably influenced by the wing venation, the presence in males of less than 14 flagellomeres, and the simple, hair-shaped antennal sensilla. On first sight, this classification seems plausible and supported by further features, such as the absence of pleurothoracal setae and the
<taxonomicName id="4C4902C7FFB76C6E20FA0FDF0A6D1ECF" box="[964,1091,229,254]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E20FA0FDF0A6D1ECF" box="[964,1091,229,254]" italics="true" pageId="2" pageNumber="50">Winnertzia</emphasis>
</taxonomicName>
-like male terminalia, i.e. the gonostyli equipped with an apical claw and the parameres fused to form a simple, membranous tegmen. However, several other features of
<taxonomicName id="4C4902C7FFB76C6E217B0E080CF01F7A" box="[581,734,306,331]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E217B0E080CF01F7A" box="[581,734,306,331]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
are not known from
<taxonomicName id="4C4902C7FFB76C6E20DF0E080A4B1F7A" box="[993,1125,306,331]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E20DF0E080A4B1F7A" box="[993,1125,306,331]" italics="true" pageId="2" pageNumber="50">Winnertzia</emphasis>
</taxonomicName>
or other
<taxonomicName id="4C4902C7FFB76C6E27E80E080B5A1F7D" box="[1238,1396,306,332]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
: an asetose pedicel, crenulate whorls of sensory hairs, hair-shaped translucent sensilla on the maxillary palpus, and an asetose pronotum. Also, the first tarsal segments in
<taxonomicName id="4C4902C7FFB76C6E20560EBA0A2E1FA8" box="[872,1024,384,409]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E20560EBA0A2E1FA8" box="[872,1024,384,409]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
are longer than those in the other porricondylines known to us, with the exception of some Heteropezini. A unique feature of
<taxonomicName id="4C4902C7FFB76C6E27D70E9D0BAE1FF1" box="[1257,1408,423,448]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E27D70E9D0BAE1FF1" box="[1257,1408,423,448]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
is antC extends beyond R5, whereas the apices of these two veins are confluent in the other porricondylines with a costal break. In Heteropezini, which lack the costal break, the costa is gradually narrowed near the apex of the wing. Altogether, there is little support for
<taxonomicName id="4C4902C7FFB76C6E21F60D200D711C02" box="[712,863,538,563]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E21F60D200D711C02" box="[712,863,538,563]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
being correctly placed among the
<taxonomicName id="4C4902C7FFB76C6E27C10D200BB71C05" box="[1279,1433,538,564]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
. An affiliation of
<taxonomicName id="4C4902C7FFB76C6E22560D7B0C2C1C6B" box="[360,514,577,602]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E22560D7B0C2C1C6B" box="[360,514,577,602]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
to the Heteropezini would also be inappropriate, because the heteropezine imago is variously degenerated (
<bibRefCitation id="EFD804B5FFB76C6E212D0D520C8F1CB3" author="Wyatt" box="[531,673,616,642]" pageId="2" pageNumber="54" refString="Wyatt, I. J. (1967) Pupal paedogenesis in the Cecidomyiidae (Diptera) 3 - A reclassification of the Heteropezini. Transactions of the Royal Entomological Society of London, 119 (3), 71 - 98." type="journal article" year="1967">Wyatt 1967</bibRefCitation>
), unlike the imago in
<taxonomicName id="4C4902C7FFB76C6E209F0D520A1B1CB0" box="[929,1077,616,641]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E209F0D520A1B1CB0" box="[929,1077,616,641]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
. The Diallactini, with
<specimenCount id="9D4FB2CDFFB76C6E26000D520BB21CB3" box="[1342,1436,616,642]" pageId="2" pageNumber="50" type="male">14 male</specimenCount>
flagellomeres, cannot accommodate
<taxonomicName id="4C4902C7FFB76C6E21570DB50D261C99" box="[617,776,655,680]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E21570DB50D261C99" box="[617,776,655,680]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
either. We do not see a better solution than leaving
<taxonomicName id="4C4902C7FFB76C6E23A90D8F0F051CFF" box="[151,299,693,718]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E23A90D8F0F051CFF" box="[151,299,693,718]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
unplaced to any tribe for the time being.
</paragraph>
<paragraph id="8BF67944FFB76C6E23F80DE60DAD1A63" blockId="2.[151,1437,152,1106]" pageId="2" pageNumber="50">
As regards the
<taxonomicName id="4C4902C7FFB76C6E22470DE60C041CC7" box="[377,554,732,758]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
other than “
<taxonomicName id="4C4902C7FFB76C6E21820DE60DAE1CC7" box="[700,896,732,758]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
”, sylvenomyias bear an uncanny resemblance to
<taxonomicName id="4C4902C7FFB76C6E238A0C380F7F1D2D" box="[180,337,770,796]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Lestremiinae">Lestremiinae</taxonomicName>
. If the first tarsal segments were longer than the second segments the feature that unfailingly separates
<taxonomicName id="4C4902C7FFB76C6E22340C130F8D1D72" box="[266,419,809,835]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Lestremiinae">Lestremiinae</taxonomicName>
from “
<taxonomicName id="4C4902C7FFB76C6E22C80C130C991D72" box="[502,695,809,835]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
” and
<taxonomicName id="4C4902C7FFB76C6E21C20C130D831D72" box="[764,941,809,835]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Cecidomyiinae">Cecidomyiinae</taxonomicName>
then sylvenomyias could be considered lestremiines. A venation pattern similar to that in
<taxonomicName id="4C4902C7FFB76C6E21C20C6A0DBB1D58" box="[764,917,848,873]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E21C20C6A0DBB1D58" box="[764,917,848,873]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
is known from some unusual lestremiines classified with the tribe
<taxonomicName id="4C4902C7FFB76C6E22860C4C0C641DA1" box="[440,586,886,912]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Strobliellini">Strobliellini</taxonomicName>
. The terminalia of
<taxonomicName id="4C4902C7FFB76C6E20100C4D0DEA1DA1" box="[814,964,887,912]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E20100C4D0DEA1DA1" box="[814,964,887,912]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
males correspond largely with those in some species of the
<taxonomicName id="4C4902C7FFB76C6E22BB0CA70FD41D86" box="[389,506,925,951]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Catochini">Catochini</taxonomicName>
, another lestremiine tribe. Last but not least, crenulate whorls of sensory hairs, the presence of which is exceptional among porricondylines, is a feature typical of lestremiines. We do not stress these similarities to suggest a close interrelationship between
<taxonomicName id="4C4902C7FFB76C6E208C0CD00A681A32" box="[946,1094,1002,1027]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E208C0CD00A681A32" box="[946,1094,1002,1027]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
and certain
<taxonomicName id="4C4902C7FFB76C6E27E80CD00B411A35" box="[1238,1391,1002,1028]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Lestremiinae">Lestremiinae</taxonomicName>
but to support our assumption that
<taxonomicName id="4C4902C7FFB76C6E213C0B2B0CB81A1B" box="[514,662,1041,1066]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B93DA556FFB76C6E213C0B2B0CB81A1B" box="[514,662,1041,1066]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
</taxonomicName>
belongs to a basal porricondyline lineage distinct from other such lineages, such as the Diallactini,
<taxonomicName id="4C4902C7FFB76C6E21290B020C841A63" box="[535,682,1080,1106]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
and Heteropezini.
</paragraph>
</subSubSection>
</treatment>
</document>