217 lines
32 KiB
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217 lines
32 KiB
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<document id="706983C9B671FEA0A60C4A23C395DDC6" ID-DOI="10.5281/zenodo.186299" ID-GBIF-Dataset="3c61a031-b0b6-4dda-ac37-a20e488ff85f" ID-ISSN="1175-5326" ID-Zenodo-Dep="186299" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461049713889" checkinUser="plazi" docAuthor="Penttinen, Jouni & Jaschhof, Mathias" docDate="2009" docId="03E0C852FFB46C6E23A90ED50DAD1A63" docLanguage="en" docName="zt02032p054.pdf" docOrigin="Zootaxa 2032" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sylvenomyia Mamaev & Zaitzev 1998" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="FFD9B02AFFB56C6C233E0F3A0E2E1E31" masterDocTitle="On the systematics of Sylvenomyia Mamaev & Zaitzev (Diptera, Cecidomyiidae, “ Porricondylinae ”), with the description of a new species from Finland" masterLastPageNumber="54" masterPageNumber="48" pageNumber="49" updateTime="1698578325124" updateUser="plazi">
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<mods:title id="064388D4A9F2E8F8C48A2DB66CA3F37C">On the systematics of Sylvenomyia Mamaev & Zaitzev (Diptera, Cecidomyiidae, “ Porricondylinae ”), with the description of a new species from Finland</mods:title>
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<mods:namePart id="0C9B44DFD30E2F5228EEFBC83B87F2CB">Penttinen, Jouni</mods:namePart>
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<mods:namePart id="AB2B522B7DF7D2532FA8088F662A45C3">Jaschhof, Mathias</mods:namePart>
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<mods:typeOfResource id="751444D662ABD72ED2DE44552BB8FE1A">text</mods:typeOfResource>
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<mods:title id="BAB03CB701698324DD1120DA1B2AA315">Zootaxa</mods:title>
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<mods:part id="A871D95307989B753ED7BAC62A838D03">
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<mods:date id="231E3741EFBD25EAD3211313915441A5">2009</mods:date>
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<mods:detail id="C7387AC7E9208063A1E2F3AE602FF84A" type="volume">
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<mods:number id="8CD8272140488BA1DF573E112CAB7AA7">2032</mods:number>
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<mods:identifier id="8B835202063590CDE90BD8B303402902" type="DOI">10.5281/zenodo.186299</mods:identifier>
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<mods:identifier id="BFEC4697542DF64FD1A5D65F59C9A856" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D30591FFE996BC3A0EB7C76A41EC1A96" type="Zenodo-Dep">186299</mods:identifier>
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<treatment id="03E0C852FFB46C6E23A90ED50DAD1A63" ID-DOI="http://doi.org/10.5281/zenodo.6214793" ID-GBIF-Taxon="119611536" ID-Zenodo-Dep="6214793" LSID="urn:lsid:plazi:treatment:03E0C852FFB46C6E23A90ED50DAD1A63" httpUri="http://treatment.plazi.org/id/03E0C852FFB46C6E23A90ED50DAD1A63" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49">
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<subSubSection id="C3532ACFFFB46C6D23A90ED50FF01C61" pageId="1" pageNumber="49" type="nomenclature">
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<paragraph id="8BF67944FFB46C6D23A90ED50CEE1C38" blockId="1.[151,704,495,521]" box="[151,704,495,521]" pageId="1" pageNumber="49">
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<heading id="D0BECE28FFB46C6D23A90ED50CEE1C38" bold="true" box="[151,704,495,521]" fontSize="11" level="1" pageId="1" pageNumber="49" reason="1">
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<emphasis id="B93DA556FFB46C6D23A90ED50CEE1C38" bold="true" box="[151,704,495,521]" pageId="1" pageNumber="49">
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Genus
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<taxonomicName id="4C4902C7FFB46C6D23D00ED50CEE1C38" ID-CoL="7QQL" ID-ENA="2338634" authority="Mamaev & Zaitzev 1998" authorityName="Mamaev & Zaitzev" authorityYear="1998" box="[238,704,495,521]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus">
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<emphasis id="B93DA556FFB46C6D23D00ED50FA91C38" bold="true" box="[238,391,495,521]" italics="true" pageId="1" pageNumber="49">Sylvenomyia</emphasis>
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<bibRefCitation id="EFD804B5FFB46C6D22B00ED50CEE1C38" author="Mamaev" box="[398,704,495,521]" pageId="1" pageNumber="54" refString="Mamaev, B. M. & Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev & Zaitzev 1998</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF67944FFB46C6D23A90D030FF01C61" blockId="1.[151,1246,569,625]" box="[151,478,569,592]" pageId="1" pageNumber="49">
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<bibRefCitation id="EFD804B5FFB46C6D23A90D030F881C61" author="Mamaev" box="[151,422,569,592]" pageId="1" pageNumber="54" refString="Mamaev, B. M. & Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev & Zaitzev 1998</bibRefCitation>
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: 211.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3532ACFFFB46C6D23A90D630AF01C41" box="[151,1246,601,625]" pageId="1" pageNumber="49" type="materials_examined">
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<paragraph id="8BF67944FFB46C6D23A90D630AF01C41" blockId="1.[151,1246,569,625]" box="[151,1246,601,625]" pageId="1" pageNumber="49">
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<emphasis id="B93DA556FFB46C6D23A90D630F031C40" bold="true" box="[151,301,601,625]" pageId="1" pageNumber="49">
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<typeStatus id="54F2C7E6FFB46C6D23A90D630EE11C40" box="[151,207,601,625]" pageId="1" pageNumber="49">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C4902C7FFB46C6D220D0D630D0C1C41" authority="Mamaev & Zaitzev 1998" authorityName="Mamaev & Zaitzev" authorityYear="1998" box="[307,802,601,624]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="sueciae">
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<emphasis id="B93DA556FFB46C6D220D0D630C3F1C41" box="[307,529,601,624]" italics="true" pageId="1" pageNumber="49">Sylvenomyia sueciae</emphasis>
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<bibRefCitation id="EFD804B5FFB46C6D21290D630D0C1C41" author="Mamaev" box="[535,802,601,624]" pageId="1" pageNumber="54" refString="Mamaev, B. M. & Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev & Zaitzev 1998</bibRefCitation>
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</taxonomicName>
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[=
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<taxonomicName id="4C4902C7FFB46C6D20760D630AFD1C41" authority="Spungis 1985" authorityName="Spungis" authorityYear="1985" box="[840,1235,601,624]" class="Insecta" family="Cecidomyiidae" genus="Chastomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="spinigera">
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<emphasis id="B93DA556FFB46C6D20760D630A181C41" box="[840,1078,601,624]" italics="true" pageId="1" pageNumber="49">Chastomera spinigera</emphasis>
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<bibRefCitation id="EFD804B5FFB46C6D27020D630AFD1C41" author="Spungis" box="[1084,1235,601,624]" pageId="1" pageNumber="54" refString="Spungis, V. (1985) Gall midges of the subtribe Diallactina (Diptera, Cecidomyiidae) in Latvia. Latvijas Entomologs, 28, 38 - 53 (in Russian, with English summary)." type="journal article" year="1985">Spungis 1985</bibRefCitation>
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C3532ACFFFB46C6D23A90DA50AD31D65" pageId="1" pageNumber="49" type="diagnosis">
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<paragraph id="8BF67944FFB46C6D23A90DA50AD31D65" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
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<emphasis id="B93DA556FFB46C6D23A90DA50F341C88" bold="true" box="[151,282,671,697]" pageId="1" pageNumber="49">Diagnosis.</emphasis>
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The strong anterior portion of C (antC) extends clearly beyond R5 and is followed by a costal break; M and CuA1 are absent (
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<figureCitation id="137265C1FFB46C6D211F0DFC0C491CD1" box="[545,615,710,736]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Fig. 1</figureCitation>
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A). The antennae in males have 11 flagellomeres; the flagellomere nodes bear crenulate whorls of sensory hairs and hair-shaped translucent sensilla (Fig. 2A). The gonostylus bears a strong apical claw (
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<figureCitation id="137265C1FFB46C6D22E50C2E0C0A1D1F" box="[475,548,788,814]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Figs 1</figureCitation>
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B, 2B, C). The simple, small tegmen is largely membranous and of broadconical shape (
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<figureCitation id="137265C1FFB46C6D227B0C000FA51D65" box="[325,395,826,852]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Figs 1</figureCitation>
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D, 2C). Females and preimaginal stages of
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<taxonomicName id="4C4902C7FFB46C6D20B40C000A301D62" box="[906,1054,826,851]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus">
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<emphasis id="B93DA556FFB46C6D20B40C000A301D62" box="[906,1054,826,851]" italics="true" pageId="1" pageNumber="49">Sylvenomyia</emphasis>
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</taxonomicName>
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spp. are unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3532ACFFFB46C6E23F80C5A0DAD1A63" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49" type="description">
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<paragraph id="8BF67944FFB46C6D23F80C5A0A861ABB" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
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<emphasis id="B93DA556FFB46C6D23F80C5A0F9F1D4B" bold="true" box="[198,433,864,890]" pageId="1" pageNumber="49">Description. Head.</emphasis>
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Occiput with short setae. Postocular bristles absent. One pair of large top setae (see
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<bibRefCitation id="EFD804B5FFB46C6D23A90CB20F6B1D93" author="Panelius" box="[151,325,904,930]" pageId="1" pageNumber="54" refString="Panelius, S. (1965) A revision of the European gall midges of the subfamily Porricondylinae (Diptera: Itonididae). Acta Zoologica Fennica, 113, 1 - 157." type="journal article" year="1965">Panelius 1965</bibRefCitation>
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). Postfrons convex, bilobate, asetose. Prefrons large, asetose. Eye bridge up to 4 ommatidia long. Remnants of larval stemmata not discernible. Antennal scape and pedicel subequal in size; scape setose ventrally; pedicel asetose; 11 flagellomeres, the 2 apical flagellomere fused; flagellomere nodes longer than necks, barrel-shaped, fully covered with microtrichia, 1 whorl of basal setae, 1 complete and 1–2 incomplete crenulate whorls of long sensory hairs, several hair-shaped translucent sensilla distally (Fig. 2A). Clypeus smaller than prefrons, projecting in lateral profile, with large setae. Maxillary palpus long, 4-segmented, setae long and curved or short and straight; first segment with hair-shaped translucent sensilla.
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</paragraph>
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<paragraph id="8BF67944FFB46C6D23FB0BAC0ACA1802" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
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<emphasis id="B93DA556FFB46C6D23FB0BAC0F021A81" bold="true" box="[197,300,1174,1200]" pageId="1" pageNumber="49">Thorax.</emphasis>
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Pronotum asetose. Scutum with sparse dorsocentral and lateral setae. Pleural setae absent. A portion in between laterotergite and mediotergite covered with strikingly large microtrichia.
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<emphasis id="B93DA556FFB46C6D27E50B870B311AE6" bold="true" box="[1243,1311,1213,1239]" pageId="1" pageNumber="49">Wing</emphasis>
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(
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<figureCitation id="137265C1FFB46C6D26110B870B5C1AE6" box="[1327,1394,1213,1239]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Fig. 1</figureCitation>
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A). Comparatively short and broad, about as long as body. Membrane covered with microtrichia and setae. AntC extending clearly beyond R5, followed by break; Sc long, evanescent apically; h absent; Rs with vertical rather than horizontal inclination; r-m + m-cu horizontal, in juxtaposition with R5; M and CuA1 absent; CuA2 slightly curved, weak or evanescent apically; CuP short, running very close to CuA2; 1 branch of A present, albeit short; dorsal setae present on Sc basally, stem vein, R, R1, R5, and CuA2; ventral setae absent. Pattern of sensory buds: R1, 2 distal; R5, 1 basal, 2 distal; further 1 mesal sensory bud on Sc. Halter comparatively short, stem and node subequal in length.
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<emphasis id="B93DA556FFB46C6D21B60AF10CE51BD4" bold="true" box="[648,715,1483,1509]" pageId="1" pageNumber="49">Legs.</emphasis>
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Foreleg slightly longer than body, femur shorter than tibia, tibia shorter than tarsus; first segment of foretarsus 1/3 the length of second segment, with short, blunt apical projection. Pretarsal claws slightly curved, with minute teeth at midlength. Empodia vestigial.
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</paragraph>
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<paragraph id="8BF67944FFB46C6D23FB09050AF61987" blockId="1.[151,1438,671,2012]" pageId="1" pageNumber="49">
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<emphasis id="B93DA556FFB46C6D23FB09050F671868" bold="true" box="[197,329,1599,1625]" pageId="1" pageNumber="49">Abdomen.</emphasis>
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All sclerites covered sparsely with large setae. Pattern of tergal plaques not fully resolved, presumably 0/2/2/1/1/1/1/0.
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<emphasis id="B93DA556FFB46C6D22DA095C0C4318B1" bold="true" box="[484,621,1638,1664]" pageId="1" pageNumber="49">Terminalia</emphasis>
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(
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<figureCitation id="137265C1FFB46C6D2142095C0CED18B1" box="[636,707,1638,1664]" captionStart="FIGURE 1" captionStartId="3.[151,256,1220,1244]" captionTargetBox="[297,1266,192,1169]" captionTargetId="figure@3.[264,1323,157,1215]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Sylvenomyia spp. A: Wing of S. fennica sp. n., dorsal view (holotype, 0.5 mm). B: Male terminalia of S. spinigera, ventral view (specimen from Lieksa, 0.05 mm). C: Ejaculatory apodeme and tegmen of S. s pinigera, ventral view (specimen from Kondopoga, 0.05 mm). acgld = accessory gland duct, antC = anterior portion of costa, antGA = anterior portion of gonocoxal apodeme, cbr = costal break, cerc = cercus, dtrbr = dorsal transverse bridge, memcp = membranous cap, parap = parameral apodeme, postC = posterior portion of costa, postGA = posterior portion of gonocoxal apodeme, vgxbr = ventral gonocoxal bridge." httpUri="https://zenodo.org/record/186300/files/figure.png" pageId="1" pageNumber="49">Figs 1</figureCitation>
|
|||
|
B, C, 2B–D). St9 not traceble. Tg9 subtrapezoid, with straight apical margin. Gonocoxites with deep ventral emargination extending beyond midlength; ventral bridge membranous; posterior portion of gonocoxal apodeme (postGA) long, extending to ventrobasal gonocoxal margin; anterior portion of gonocoxal apodeme (antGA) moderately long; dorsal transverse bridge unsclerotized. Gonostylus slightly tapered towards apex, with strong, multipointed apical claw. Ejaculatory apodeme as long as gonocoxites, strongly sclerotized, with membranous, tulip-shaped cap apically. Ducts of accessory glands distinctive. Tegmen small, largely membranous, broad-conical, rounded apically, ventrolateral margins serrate or with scaly surface, apex weak, parameral apodemes directed ventroanteriorly. St10 weak, bilobate, pubescent, asetose. Cerci not or not much extending beyond tg9, setose.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8BF67944FFB46C6E23F808F80D271CFE" blockId="1.[151,1438,671,2012]" lastBlockId="2.[151,1437,152,1106]" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49">
|
|||
|
<emphasis id="B93DA556FFB46C6D23F808F80FE619ED" bold="true" box="[198,456,1986,2012]" pageId="1" pageNumber="49">Systematic position.</emphasis>
|
|||
|
<bibRefCitation id="EFD804B5FFB46C6D22E908F80D1319ED" author="Mamaev" box="[471,829,1986,2012]" pageId="1" pageNumber="54" refString="Mamaev, B. M. & Zaitzev, A. I. (1998) Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica, 9, 211 - 213." type="journal article" year="1998">Mamaev and Zaitzev (1998)</bibRefCitation>
|
|||
|
classified
|
|||
|
<taxonomicName id="4C4902C7FFB46C6D20F408F90A7019ED" box="[970,1118,1987,2012]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB46C6D20F408F90A7019ED" box="[970,1118,1987,2012]" italics="true" pageId="1" pageNumber="49">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
with the
|
|||
|
<taxonomicName id="4C4902C7FFB46C6D27DB08F80B5519ED" box="[1253,1403,1986,2012]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
|
|||
|
, a decision presumably influenced by the wing venation, the presence in males of less than 14 flagellomeres, and the simple, hair-shaped antennal sensilla. On first sight, this classification seems plausible and supported by further features, such as the absence of pleurothoracal setae and the
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E20FA0FDF0A6D1ECF" box="[964,1091,229,254]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E20FA0FDF0A6D1ECF" box="[964,1091,229,254]" italics="true" pageId="2" pageNumber="50">Winnertzia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
-like male terminalia, i.e. the gonostyli equipped with an apical claw and the parameres fused to form a simple, membranous tegmen. However, several other features of
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E217B0E080CF01F7A" box="[581,734,306,331]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E217B0E080CF01F7A" box="[581,734,306,331]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
are not known from
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E20DF0E080A4B1F7A" box="[993,1125,306,331]" class="Insecta" family="Cecidomyiidae" genus="Winnertzia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E20DF0E080A4B1F7A" box="[993,1125,306,331]" italics="true" pageId="2" pageNumber="50">Winnertzia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
or other
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E27E80E080B5A1F7D" box="[1238,1396,306,332]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
|
|||
|
: an asetose pedicel, crenulate whorls of sensory hairs, hair-shaped translucent sensilla on the maxillary palpus, and an asetose pronotum. Also, the first tarsal segments in
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E20560EBA0A2E1FA8" box="[872,1024,384,409]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E20560EBA0A2E1FA8" box="[872,1024,384,409]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
are longer than those in the other porricondylines known to us, with the exception of some Heteropezini. A unique feature of
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E27D70E9D0BAE1FF1" box="[1257,1408,423,448]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E27D70E9D0BAE1FF1" box="[1257,1408,423,448]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
is antC extends beyond R5, whereas the apices of these two veins are confluent in the other porricondylines with a costal break. In Heteropezini, which lack the costal break, the costa is gradually narrowed near the apex of the wing. Altogether, there is little support for
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E21F60D200D711C02" box="[712,863,538,563]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E21F60D200D711C02" box="[712,863,538,563]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
being correctly placed among the
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E27C10D200BB71C05" box="[1279,1433,538,564]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
|
|||
|
. An affiliation of
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E22560D7B0C2C1C6B" box="[360,514,577,602]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E22560D7B0C2C1C6B" box="[360,514,577,602]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
to the Heteropezini would also be inappropriate, because the heteropezine imago is variously degenerated (
|
|||
|
<bibRefCitation id="EFD804B5FFB76C6E212D0D520C8F1CB3" author="Wyatt" box="[531,673,616,642]" pageId="2" pageNumber="54" refString="Wyatt, I. J. (1967) Pupal paedogenesis in the Cecidomyiidae (Diptera) 3 - A reclassification of the Heteropezini. Transactions of the Royal Entomological Society of London, 119 (3), 71 - 98." type="journal article" year="1967">Wyatt 1967</bibRefCitation>
|
|||
|
), unlike the imago in
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E209F0D520A1B1CB0" box="[929,1077,616,641]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E209F0D520A1B1CB0" box="[929,1077,616,641]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
. The Diallactini, with
|
|||
|
<specimenCount id="9D4FB2CDFFB76C6E26000D520BB21CB3" box="[1342,1436,616,642]" pageId="2" pageNumber="50" type="male">14 male</specimenCount>
|
|||
|
flagellomeres, cannot accommodate
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E21570DB50D261C99" box="[617,776,655,680]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E21570DB50D261C99" box="[617,776,655,680]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
either. We do not see a better solution than leaving
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E23A90D8F0F051CFF" box="[151,299,693,718]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E23A90D8F0F051CFF" box="[151,299,693,718]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
unplaced to any tribe for the time being.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8BF67944FFB76C6E23F80DE60DAD1A63" blockId="2.[151,1437,152,1106]" pageId="2" pageNumber="50">
|
|||
|
As regards the
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E22470DE60C041CC7" box="[377,554,732,758]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
|
|||
|
other than “
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E21820DE60DAE1CC7" box="[700,896,732,758]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
|
|||
|
”, sylvenomyias bear an uncanny resemblance to
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E238A0C380F7F1D2D" box="[180,337,770,796]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Lestremiinae">Lestremiinae</taxonomicName>
|
|||
|
. If the first tarsal segments were longer than the second segments – the feature that unfailingly separates
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E22340C130F8D1D72" box="[266,419,809,835]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Lestremiinae">Lestremiinae</taxonomicName>
|
|||
|
from “
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E22C80C130C991D72" box="[502,695,809,835]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
|
|||
|
” and
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E21C20C130D831D72" box="[764,941,809,835]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Cecidomyiinae">Cecidomyiinae</taxonomicName>
|
|||
|
– then sylvenomyias could be considered lestremiines. A venation pattern similar to that in
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E21C20C6A0DBB1D58" box="[764,917,848,873]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E21C20C6A0DBB1D58" box="[764,917,848,873]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
is known from some unusual lestremiines classified with the tribe
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E22860C4C0C641DA1" box="[440,586,886,912]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Strobliellini">Strobliellini</taxonomicName>
|
|||
|
. The terminalia of
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E20100C4D0DEA1DA1" box="[814,964,887,912]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E20100C4D0DEA1DA1" box="[814,964,887,912]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
males correspond largely with those in some species of the
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E22BB0CA70FD41D86" box="[389,506,925,951]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Catochini">Catochini</taxonomicName>
|
|||
|
, another lestremiine tribe. Last but not least, crenulate whorls of sensory hairs, the presence of which is exceptional among porricondylines, is a feature typical of lestremiines. We do not stress these similarities to suggest a close interrelationship between
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E208C0CD00A681A32" box="[946,1094,1002,1027]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E208C0CD00A681A32" box="[946,1094,1002,1027]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and certain
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E27E80CD00B411A35" box="[1238,1391,1002,1028]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Lestremiinae">Lestremiinae</taxonomicName>
|
|||
|
but to support our assumption that
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E213C0B2B0CB81A1B" box="[514,662,1041,1066]" class="Insecta" family="Cecidomyiidae" genus="Sylvenomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B93DA556FFB76C6E213C0B2B0CB81A1B" box="[514,662,1041,1066]" italics="true" pageId="2" pageNumber="50">Sylvenomyia</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
belongs to a basal porricondyline lineage distinct from other such lineages, such as the Diallactini,
|
|||
|
<taxonomicName id="4C4902C7FFB76C6E21290B020C841A63" box="[535,682,1080,1106]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Winnertziini">Winnertziini</taxonomicName>
|
|||
|
and Heteropezini.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|