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<mods:title id="8ADC37281487CF8E660C9821F5886BE2">A review of the families and genera of the hyperiidean amphipod superfamily Phronimoidea Bowman &amp; Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea)</mods:title>
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<treatment id="03931615EC60FFE0FEDFFDA3FAB8FE50" ID-DOI="http://doi.org/10.5281/zenodo.5029994" ID-GBIF-Taxon="184952228" ID-Zenodo-Dep="5029994" LSID="urn:lsid:plazi:treatment:03931615EC60FFE0FEDFFDA3FAB8FE50" httpUri="http://treatment.plazi.org/id/03931615EC60FFE0FEDFFDA3FAB8FE50" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C320F488EC60FFE1FEDFFDA3FD5BFDDB" box="[264,713,603,629]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B85A703EC60FFE1FEDFFDA3FD5BFDDB" blockId="16.[264,713,603,629]" box="[264,713,603,629]" pageId="16" pageNumber="17">
<heading id="D0CD106FEC60FFE1FEDFFDA3FD5BFDDB" bold="true" box="[264,713,603,629]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B94E7B11EC60FFE1FEDFFDA3FD5BFDDB" bold="true" box="[264,713,603,629]" pageId="16" pageNumber="17">
Family
<taxonomicName id="4C3ADC80EC60FFE1FEB0FDA3FD5BFDDB" ID-CoL="7NKG4" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[359,713,603,629]" class="Malacostraca" family="Phrosinidae" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">
PHROSINIDAE
<bibRefCitation id="EFABDAF2EC60FFE1FDEAFDA3FD5BFDDB" author="Dana, J. D." box="[573,713,603,629]" pageId="16" pageNumber="17" pagination="297 - 316" refId="ref25481" refString="Dana, J. D. (1852) On the classification of the Crustacea Choristopoda or Tetradecapoda. American Journal of Sciences and Arts, series 2, 14 (41), 297 - 316." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="C320F488EC60FFE1FEDFFD5FFE46F907" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B85A703EC60FFE1FEDFFD5FFEEDFD6E" blockId="16.[264,1325,679,1305]" box="[264,383,679,704]" pageId="16" pageNumber="17">
<emphasis id="B94E7B11EC60FFE1FEDFFD5FFEEDFD6E" box="[264,383,679,704]" italics="true" pageId="16" pageNumber="17">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B85A703EC60FFE1FEEFFD37FCB7FB5F" blockId="16.[264,1325,679,1305]" pageId="16" pageNumber="17">
Body length
<quantity id="4CC20AE6EC60FFE1FE05FD37FDC1FD47" box="[466,595,719,745]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="16" pageNumber="17" unit="mm" value="20.0" valueMax="30.0" valueMin="10.0">1030 mm</quantity>
, rather compact with relatively thick cuticle, relatively transparent. Head large, globular, height more than length. Eyes large, occupying most of head surface. Pereonites all separate, or pereonites 1 &amp; 2 fused. Coxae separate from pereonites. Antennae 1 reduced to two articles in females; multi­articulate, in males with enlarged callynophore, with aesthetasc brush composed of two asymmetrical fields located ventromedially and ventrolaterally. Antennae 2 rudimentary, or absent in females; multiarticulate in males. Mandibles with palp in males, without palp in females; molar welldeveloped. Maxillae 1 with palp and well­developed outer lobe, inner lobe absent. Maxillae 2 bilobed, well­developed. Maxilliped with slender outer lobes; inner lobe about halflength outer lobes. Gnathopods 1 &amp; 2 simple. Pereopods 36 prehensile, or subchelate. Pereopod 5 the longest with large, denticulate subchela. Pereopods 57 with broad basis. Pereopod 7 reduced in size, sometimes with reduced number of articles. Uropods composed of single, foliaceous article. Telson small, not longer than half­length U3. Gills on pereonites 26. Oostegites on pereonites 25.
</paragraph>
<paragraph id="8B85A703EC60FFE1FEEFFB07FC1CFAB6" blockId="16.[264,1325,679,1305]" box="[312,910,1279,1305]" pageId="16" pageNumber="17">
Three genera:
<emphasis id="B94E7B11EC60FFE1FE36FB07FD64FAB6" box="[481,758,1279,1304]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C3ADC80EC60FFE1FE36FB07FDDCFAB6" authorityName="Risso" authorityYear="1822" box="[481,590,1279,1304]" class="Malacostraca" family="Phrosinidae" genus="Phrosina" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Phrosina</taxonomicName>
,
<taxonomicName id="4C3ADC80EC60FFE1FD8DFB07FD64FAB6" authorityName="Milne-Edwards" authorityYear="1830" box="[602,758,1279,1304]" class="Malacostraca" family="Phrosinidae" genus="Anchylomera" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Anchylomera</taxonomicName>
</emphasis>
and
<emphasis id="B94E7B11EC60FFE1FCF8FB07FC1CFAB6" box="[815,910,1279,1304]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C3ADC80EC60FFE1FCF8FB07FC18FAB6" authorityName="Guerin-Meneville" authorityYear="1836" box="[815,906,1279,1304]" class="Malacostraca" family="Phrosinidae" genus="Primno" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Primno</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B85A703EC60FFE1FEDFFAB7FEFCFAC6" blockId="16.[264,1322,1359,1705]" box="[264,366,1359,1384]" pageId="16" pageNumber="17">
<emphasis id="B94E7B11EC60FFE1FEDFFAB7FEFCFAC6" box="[264,366,1359,1384]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
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<paragraph id="8B85A703EC60FFE1FEEFFA8FFE5EF9A7" blockId="16.[264,1322,1359,1705]" pageId="16" pageNumber="17">
Species of this family are very distinctive, and are often present in plankton collections, sometimes in very large numbers.
<taxonomicName id="4C3ADC80EC60FFE1FD2FFA67FCF0FA16" authorityName="Risso" authorityYear="1822" box="[760,866,1439,1464]" class="Malacostraca" family="Phrosinidae" genus="Phrosina" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC60FFE1FD2FFA67FCF0FA16" box="[760,866,1439,1464]" italics="true" pageId="16" pageNumber="17">Phrosina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3ADC80EC60FFE1FC48FA67FBA9FA16" authorityName="Milne-Edwards" authorityYear="1830" box="[927,1083,1439,1464]" class="Malacostraca" family="Phrosinidae" genus="Anchylomera" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC60FFE1FC48FA67FBA9FA16" box="[927,1083,1439,1464]" italics="true" pageId="16" pageNumber="17">Anchylomera</emphasis>
</taxonomicName>
are monotypic, and
<taxonomicName id="4C3ADC80EC60FFE1FEDFFA3FFEF2FA4E" authorityName="Guerin-Meneville" authorityYear="1836" box="[264,352,1479,1504]" class="Malacostraca" family="Phrosinidae" genus="Primno" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC60FFE1FEDFFA3FFEF2FA4E" box="[264,352,1479,1504]" italics="true" pageId="16" pageNumber="17">Primno</emphasis>
</taxonomicName>
has been revised by
<bibRefCitation id="EFABDAF2EC60FFE1FD8DFA3FFC8FFA4F" author="Bowman, T. E." box="[602,797,1479,1505]" pageId="16" pageNumber="17" pagination="1 - 23" refId="ref24383" refString="Bowman, T. E. (1978). Revision of the pelagic amphipod genus Primno (Hyperiidea: Phrosinidae). Smithsonian Contributions to Zoology No. 275, 1 - 23." type="book chapter" year="1978">Bowman (1978)</bibRefCitation>
. Thus, only minimal additional information is provided here.
</paragraph>
<paragraph id="8B85A703EC60FFE1FEEFF9EFFE46F907" blockId="16.[264,1322,1359,1705]" pageId="16" pageNumber="17">
The structure of the uropods, each consisting of a single leaf­like article, is a unique feature amongst the
<taxonomicName id="4C3ADC80EC60FFE1FDDDF9C7FD02F9F7" box="[522,656,1599,1625]" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subOrder" subOrder="Hyperiidea">Hyperiidea</taxonomicName>
. They may serve as effective locomotory organs as phrosinids are known to be active swimmers, and sometimes occur in large swarms (
<bibRefCitation id="EFABDAF2EC60FFE1FB35F99FFE55F907" author="Lobel, P. S. &amp; Randall, J. E." pageId="16" pageNumber="17" pagination="253 - 262" refId="ref28008" refString="Lobel, P. S. &amp; Randall, J. E. (1986) Swarming behaviour of the hyperiid amphipod Anchylomera blossevillei. Journal of Plankton Research, 8 (2), 253 - 262." type="journal article" year="1986">Lobel &amp; Randall 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="8B85A703EC60FFE1FEDFF8FEFCFAF88E" blockId="16.[264,872,1798,1824]" box="[264,872,1798,1824]" pageId="16" pageNumber="17">
<heading id="D0CD106FEC60FFE1FEDFF8FEFCFAF88E" bold="true" box="[264,872,1798,1824]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B94E7B11EC60FFE1FEDFF8FEFCFAF88E" bold="true" box="[264,872,1798,1824]" pageId="16" pageNumber="17">
Key to the genera of the Family
<taxonomicName id="4C3ADC80EC60FFE1FD4EF8FEFCFAF88E" ID-CoL="7NKG4" authorityName="Dana" authorityYear="1852" box="[665,872,1798,1824]" class="Malacostraca" family="Phrosinidae" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">PHROSINIDAE</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C320F488EC60FFE0FEDFF8A9FAB8FE50" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="key">
<keyStep id="30CEBFA6EC60FFE0FEDFF8A9FABFFE28" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<paragraph id="8B85A703EC60FFE0FEDFF8A9FAB8FE98" blockId="16.[264,1321,1873,1899]" lastBlockId="17.[264,1325,284,510]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<keyLead id="30CB0436EC60FFE0FEDFF8A9FAB8FE98" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
1. Pereopods 3 &amp; 4 distinctly subchelate, with large teeth on distal margin of carpus. Pereopod 7 reduced to basis and one tiny additional article
<emphasis id="B94E7B11EC61FFE0FC3CFEE4FBBFFE9B" box="[1003,1069,284,309]" italics="true" pageId="17" pageNumber="18">.........</emphasis>
<taxonomicName id="4C3ADC80EC61FFE0FBFAFEE4FAB8FE98" ID-CoL="7PRM2" authority="Risso, 1822" authorityName="Risso" authorityYear="1822" box="[1069,1322,284,310]" class="Malacostraca" family="Phrosinidae" genus="Phrosina" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC61FFE0FBFAFEE4FB0BFE9B" box="[1069,1177,284,309]" italics="true" pageId="17" pageNumber="18">Phrosina</emphasis>
<bibRefCitation id="EFABDAF2EC61FFE0FB77FEE4FAB8FE98" author="Risso, A." box="[1184,1322,284,310]" pageId="17" pageNumber="18" pagination="241 - 248" refId="ref28904" refString="Risso, A. (1822) Memoire sur quelques nouveaux Crustaces observes dans la mer de Nice. Journal de Physique, de Chimie et d'Histoire Naturelle, 95, 241 - 248." type="journal article" year="1822">Risso, 1822</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8B85A703EC61FFE0FEDFFEBCFABFFE28" blockId="17.[264,1325,284,510]" pageId="17" pageNumber="18">
<keyLead id="30CB0436EC61FFE0FEDFFEBCFABFFE28" pageId="17" pageNumber="18">­ Pereopods 3 &amp; 4 simple, or if subchelate, with very small teeth, or bristles on distal margin of carpus. Pereopod 7 reduced, with at least 2 articles in addition to basis...... 2</keyLead>
</paragraph>
</keyStep>
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<keyLead id="30CB0436EC61FFE0FEDFFE6CFAB8FE00" box="[264,1322,404,430]" pageId="17" pageNumber="18">
2. Pereopods 3 &amp; 4 simple. Pereonites 1 &amp; 2 separate
<emphasis id="B94E7B11EC61FFE0FC5BFE6CFC3BFE03" box="[908,937,404,429]" italics="true" pageId="17" pageNumber="18">....</emphasis>
<taxonomicName id="4C3ADC80EC61FFE0FC67FE6CFAB8FE00" ID-CoL="7PV3J" authority="Guerin-Meneville, 1836" authorityName="Guerin-Meneville" authorityYear="1836" box="[944,1322,404,430]" class="Malacostraca" family="Phrosinidae" genus="Primno" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC61FFE0FC67FE6CFB9AFE03" box="[944,1032,404,429]" italics="true" pageId="17" pageNumber="18">Primno</emphasis>
Guérin­Méneville, 1836
</taxonomicName>
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</paragraph>
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<keyLead id="30CB0436EC61FFE0FEDFFE44FAB8FE50" pageId="17" pageNumber="18">
­ Pereopods 3 &amp; 4 distinctly subchelate. Pereonites 1 &amp; 2 fused ......................................
<emphasis id="B94E7B11EC61FFE0FEECFE1CFC17FE53" box="[315,901,484,509]" italics="true" pageId="17" pageNumber="18">................................................................................</emphasis>
<taxonomicName id="4C3ADC80EC61FFE0FC5FFE1CFAB8FE50" ID-CoL="7NP2M" authority="Milne-Edwards, 1830" authorityName="Milne-Edwards" authorityYear="1830" box="[904,1322,484,510]" class="Malacostraca" family="Phrosinidae" genus="Anchylomera" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC61FFE0FC5FFE1CFBB6FE53" box="[904,1060,484,509]" italics="true" pageId="17" pageNumber="18">Anchylomera</emphasis>
<bibRefCitation id="EFABDAF2EC61FFE0FBFCFE1CFAB8FE50" author="Milne-Edwards, H." box="[1067,1322,484,510]" pageId="17" pageNumber="18" pagination="353 - 399" refId="ref28202" refString="Milne-Edwards, H. (1830) Extrait de recherches pour servir a l'histoire naturelle des Crustaces Amphipodes. Annales des Sciences Naturelles, 20, 353 - 399, pls 10, 11." type="book chapter" year="1830">Milne­Edwards, 1830</bibRefCitation>
</taxonomicName>
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