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<document id="28E7F88523ABFBCF10BC6D71F80E66B7" ID-DOI="10.5252/geodiversitas2024v46a15" ID-ISSN="1638-9395" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFC4E425FC73FAC2FD1EFE63" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Ponderia Houart 1986" docType="treatment" docVersion="1" lastPageNumber="527" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="527" updateTime="1729758832705" updateUser="GgImagineBatch" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="B3E2A4F052003F8C80490F2BB4932F44">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="EDC88F9908ABD7E831347D1506C56730">Ledon, Daniel</mods:namePart>
<mods:affiliation id="7A00540B533FF8B5275B744B7D96BF92">Muséum national dhistoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:title id="AB76D55C6272BA5AE0B85D9FDE5FAA9E">Geodiversitas</mods:title>
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<mods:date id="FA0FCB13E731BBDB0AF823CD8123FAAF">2024</mods:date>
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<paragraph id="8B9836CCFFC4E424FC73FAC2FA90FA9E" blockId="34.[971,1297,1299,1326]" box="[971,1297,1299,1326]" pageId="34" pageNumber="527">
<heading id="D0D081A0FFC4E424FC73FAC2FA90FA9E" box="[971,1297,1299,1326]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="527" reason="2">
Genus
<taxonomicName id="4C274D4FFFC4E424FBA0FAC5FA90FA9E" ID-CoL="7PD4Q" authority="Houart, 1986" authorityName="Houart" authorityYear="1986" box="[1048,1297,1299,1326]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FBA0FAC5FBF5FA9E" box="[1048,1140,1299,1325]" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
Houart, 1986
</taxonomicName>
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</paragraph>
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<typeStatus id="549C886EFFC4E424FC95FA9BFCDCFAD1" box="[813,861,1357,1380]" pageId="34" pageNumber="527">TYPE</typeStatus>
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. —
<taxonomicName id="4C274D4FFFC4E424FC62FA9AFA90FAD6" authority="Hutton, 1873" authorityName="Hutton" authorityYear="1873" box="[986,1297,1356,1381]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="zealandica">
<emphasis id="B953EADEFFC4E424FC62FA9AFB01FAD7" box="[986,1152,1356,1380]" italics="true" pageId="34" pageNumber="527">Typhis zealandica</emphasis>
Hutton, 1873
</taxonomicName>
by original designation. Pleistocene to Recent:
<collectingCountry id="F330765CFFC4E424FBD0FAB1FB6FFACC" box="[1128,1262,1383,1407]" name="New Zealand" pageId="34" pageNumber="527">New Zealand</collectingCountry>
.
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<smallCapsWord id="8D7EA010FFC4E424FC95FA65FC2DFA78" baselines="1478,1478" box="[813,940,1459,1485]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="34" pageNumber="527">COMMENTS</smallCapsWord>
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Subfamilial placement of the genus
<taxonomicName id="4C274D4FFFC4E424FB0EFA05FA2EFA5E" authority="Houart, 1986" authorityName="Houart" authorityYear="1986" box="[1206,1455,1491,1517]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB0EFA05FA93FA5E" box="[1206,1298,1491,1517]" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
Houart, 1986
</taxonomicName>
has varied in the literature. DAttilio &amp; Hertz (1988) placed it in the subfamily
<taxonomicName id="4C274D4FFFC4E424FC48F9C4FA2DF99E" authority="D'Attilio &amp; Hertz, 1988" authorityName="d'Attilio &amp; Hertz" authorityYear="1988" box="[1008,1452,1554,1581]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="subFamily" subFamily="Tripterotyphinae">Tripterotyphinae DAttilio &amp; Hertz, 1988</taxonomicName>
, whereas
<bibRefCitation id="EFB64B3DFFC4E424FC3DF9E4FB93F9FF" author="VOKES E. H." box="[901,1042,1586,1612]" pageId="34" pageNumber="527" pagination="4 - 6" refId="ref49051" refString="VOKES E. H. 1996. - One last look at the Muricidae. American Conchologist 24 (2): 4 - 6." type="journal article" year="1996">Vokes (1996)</bibRefCitation>
preferred placement it in the subfamily
<taxonomicName id="4C274D4FFFC4E424FC94F984FC1BF9DF" authorityName="Rafinesque" authorityYear="1815" box="[812,922,1618,1644]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="subFamily" subFamily="Muricinae">Muricinae</taxonomicName>
.More recently,
<bibRefCitation id="EFB64B3DFFC4E424FB8EF984FB75F9DF" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[1078,1268,1618,1644]" pageId="34" pageNumber="527" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFC4E424FB8EF984FBF2F9DF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1078,1139,1618,1644]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFC4E424FBC2F985FB28F9DF" box="[1146,1193,1618,1644]" italics="true" pageId="34" pageNumber="527">et al.</emphasis>
(2011)
</bibRefCitation>
included
<taxonomicName id="4C274D4FFFC4E424FAECF984FA2CF9DF" authorityName="Houart" authorityYear="1986" box="[1364,1453,1618,1644]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FAECF984FA2CF9DF" box="[1364,1453,1618,1644]" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
</taxonomicName>
in a basal muricid group (without subfamilial assignment) and excluded it from the typical
<taxonomicName id="4C274D4FFFC4E424FBFAF944FB31F91F" authorityName="Rafinesque" authorityYear="1815" box="[1090,1200,1682,1708]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="subFamily" subFamily="Muricinae">Muricinae</taxonomicName>
.
<bibRefCitation id="EFB64B3DFFC4E424FB0FF945FACFF91F" author="HOUART R." box="[1207,1358,1682,1709]" pageId="34" pageNumber="527" pagination="37 - 66" refId="ref45005" refString="HOUART R. 2018. - Historique et classification des especes actuelles de Muricidae (Neogastropoda, Muricoidea). Novapex 19 (2): 37 - 66." type="journal article" year="2018">Houart (2018)</bibRefCitation>
included five genera in the
<taxonomicName id="4C274D4FFFC4E424FC53F964FB21F97F" authorityName="d'Attilio &amp; Hertz" authorityYear="1988" box="[1003,1184,1714,1740]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="subFamily" subFamily="Tripterotyphinae">Tripterotyphinae</taxonomicName>
:
<taxonomicName id="4C274D4FFFC4E424FB17F964FCE7F95F" authority="DuShane, 1969" authorityName="DuShane" authorityYear="1969" class="Gastropoda" family="Muricidae" genus="Cinclidotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB17F964FABEF97F" box="[1199,1343,1714,1740]" italics="true" pageId="34" pageNumber="527">Cinclidotyphis</emphasis>
DuShane, 1969
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC4E424FCD7F904FBF1F95F" authority="Ponder, 1972" authorityName="Ponder" authorityYear="1972" box="[879,1136,1746,1772]" class="Gastropoda" family="Muricidae" genus="Prototyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FCD7F904FC5FF95F" box="[879,990,1746,1772]" italics="true" pageId="34" pageNumber="527">Prototyphis</emphasis>
Ponder, 1972
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC4E424FBC0F904FA2AF95F" authority="Jousseaume, 1880" authorityName="Jousseaume" authorityYear="1880" box="[1144,1451,1746,1772]" class="Gastropoda" family="Muricidae" genus="Pterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FBC0F904FB67F95F" box="[1144,1254,1746,1772]" italics="true" pageId="34" pageNumber="527">Pterotyphis</emphasis>
<bibRefCitation id="EFB64B3DFFC4E424FB54F904FA2AF95F" author="JOUSSEAUME F. P." box="[1260,1451,1746,1772]" pageId="34" pageNumber="527" pagination="335 - 338" refId="ref45378" refString="JOUSSEAUME F. P. 1880. - Division methodique de la famille des Purpurides. Le Naturaliste 2 (42): 335 - 338. http: // biodiversitylibrary. org / page / 33928744" type="journal article" year="1880">Jousseaume, 1880</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC4E424FC93F924FB31F8BF" authority="Pilsbry &amp; Lowe, 1932" authorityName="Pilsbry &amp; Lowe" authorityYear="1932" box="[811,1200,1778,1804]" class="Gastropoda" family="Muricidae" genus="Tripterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FC93F924FC37F8BF" box="[811,950,1778,1804]" italics="true" pageId="34" pageNumber="527">Tripterotyphis</emphasis>
Pilsbry &amp; Lowe, 1932
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC4E424FB52F924FAC2F8BF" authorityName="Houart" authorityYear="1986" box="[1258,1347,1778,1804]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB52F924FAC2F8BF" box="[1258,1347,1778,1804]" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
</taxonomicName>
. All these genera share trivarical morphology, but the construction of the anal tube in
<emphasis id="B953EADEFFC4E424FC16F8E7FBC1F8F8" box="[942,1088,1841,1867]" italics="true" pageId="34" pageNumber="527">Cynclidotyphis</emphasis>
and
<taxonomicName id="4C274D4FFFC4E424FBCCF8E7FB63F8F8" authorityName="Jousseaume" authorityYear="1880" box="[1140,1250,1841,1867]" class="Gastropoda" family="Muricidae" genus="Pterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FBCCF8E7FB63F8F8" box="[1140,1250,1841,1867]" italics="true" pageId="34" pageNumber="527">Pterotyphis</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4C274D4FFFC4E424FC94F887FC1AF8D8" authorityName="Ponder" authorityYear="1972" box="[812,923,1873,1899]" class="Gastropoda" family="Muricidae" genus="Prototyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FC94F887FC1AF8D8" box="[812,923,1873,1899]" italics="true" pageId="34" pageNumber="527">Prototyphis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC4E424FC11F887FB82F8D8" authorityName="Houart" authorityYear="1986" box="[937,1027,1873,1899]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FC11F887FB82F8D8" box="[937,1027,1873,1899]" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC4E424FB81F887FB42F8D8" authorityName="Pilsbry &amp; Lowe" authorityYear="1932" box="[1081,1219,1873,1899]" class="Gastropoda" family="Muricidae" genus="Tripterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB81F887FB42F8D8" box="[1081,1219,1873,1899]" italics="true" pageId="34" pageNumber="527">Tripterotyphis</emphasis>
</taxonomicName>
. In
<emphasis id="B953EADEFFC4E424FB57F887FA00F8D8" box="[1263,1409,1873,1899]" italics="true" pageId="34" pageNumber="527">Cynclidotyphis</emphasis>
and
<emphasis id="B953EADEFFC4E424FC95F8A7FC20F838" box="[813,929,1905,1931]" italics="true" pageId="34" pageNumber="527">
<taxonomicName id="4C274D4FFFC4E424FC95F8A7FC1CF838" authorityName="Jousseaume" authorityYear="1880" box="[813,925,1905,1931]" class="Gastropoda" family="Muricidae" genus="Pterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">Pterotyphis</taxonomicName>
,
</emphasis>
the anal tube is built just below the suture probably not with P1, whereas in
<taxonomicName id="4C274D4FFFC4E424FB98F847FB0FF818" authorityName="Ponder" authorityYear="1972" box="[1056,1166,1937,1963]" class="Gastropoda" family="Muricidae" genus="Prototyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB98F847FB0FF818" box="[1056,1166,1937,1963]" italics="true" pageId="34" pageNumber="527">Prototyphis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFC4E424FB24F847FB77F818" authorityName="Houart" authorityYear="1986" box="[1180,1270,1937,1963]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB24F847FB77F818" box="[1180,1270,1937,1963]" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC4E424FA9CF847FA2CF818" authorityName="Pilsbry &amp; Lowe" authorityYear="1932" box="[1316,1453,1937,1963]" class="Gastropoda" family="Muricidae" genus="Tripterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FA9CF847FA2CF818" box="[1316,1453,1937,1963]" italics="true" pageId="34" pageNumber="527">Tripterotyphis</emphasis>
</taxonomicName>
it is clearly built with P1.
<taxonomicName id="4C274D4FFFC4E424FB85F867FB46F878" authorityName="Pilsbry &amp; Lowe" authorityYear="1932" box="[1085,1223,1969,1995]" class="Gastropoda" family="Muricidae" genus="Tripterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FB85F867FB46F878" box="[1085,1223,1969,1995]" italics="true" pageId="34" pageNumber="527">Tripterotyphis</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C274D4FFFC4E424FAC7F864FCE0F858" authorityName="Houart" authorityYear="1986" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FAC7F864FCE0F858" italics="true" pageId="34" pageNumber="527">Ponderia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C274D4FFFC4E424FC2AF807FB80F858" authorityName="Ponder" authorityYear="1972" box="[914,1025,2001,2027]" class="Gastropoda" family="Muricidae" genus="Prototyphis" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FC2AF807FB80F858" box="[914,1025,2001,2027]" italics="true" pageId="34" pageNumber="527">Prototyphis</emphasis>
</taxonomicName>
in having more strongly developed spiral sculpture. Thus, this group of five genera is likely to be polyphyletic. In their molecular phylogenetic work,
<bibRefCitation id="EFB64B3DFFC5E425FD38FF21FF4CFE82" author="RUSSINI V. &amp; FASSIO G. &amp; NOCELLA E. &amp; HOUART R. &amp; BARCO A. &amp; PUILLANDRE N. &amp; MODICA M. V. &amp; M. OLIVERIO" pageId="35" pageNumber="528" pagination="856 - 868" refId="ref48453" refString="RUSSINI V., FASSIO G., NOCELLA E., HOUART R., BARCO A., PUILLANDRE N., LOZOUET P, MODICA M. V. &amp; M. OLIVERIO 2023. - Whelks, rock-snails, and allied: a new phylogenetic framework for the family Muricidae (Mollusca: Gastropoda). The European Zoological Journal 90 (2): 856 - 868. https: // doi. org / 10.1080 / 24750263.2023.2283517" type="journal article" year="2023">
Russini
<emphasis id="B953EADEFFC5E425FD6CFF2EFC86FEA2" box="[724,775,247,273]" italics="true" pageId="35" pageNumber="528">et al.</emphasis>
(2023)
</bibRefCitation>
confirm this hypothesis showing that: 1)
<taxonomicName id="4C274D4FFFC5E425FDC3FEC1FC84FE82" authorityName="Pilsbry &amp; Lowe" authorityYear="1932" box="[635,773,279,305]" class="Gastropoda" family="Muricidae" genus="Tripterotyphis" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="528" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC5E425FDC3FEC1FC84FE82" box="[635,773,279,305]" italics="true" pageId="35" pageNumber="528">Tripterotyphis</emphasis>
</taxonomicName>
belongs to a clade containing
<taxonomicName id="4C274D4FFFC5E425FE0FFEEEFDA2FEE2" authorityName="Bucquoy &amp; Dautzenberg" authorityYear="1882" box="[439,547,312,337]" class="Gastropoda" family="Muricidae" genus="Muricopsis" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="528" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC5E425FE0FFEEEFDA2FEE2" box="[439,547,312,337]" italics="true" pageId="35" pageNumber="528">Muricopsis</emphasis>
</taxonomicName>
; and 2)
<taxonomicName id="4C274D4FFFC5E425FDC1FEE1FD52FEE2" authorityName="Houart" authorityYear="1986" box="[633,723,311,337]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="528" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC5E425FDC1FEE1FD52FEE2" box="[633,723,311,337]" italics="true" pageId="35" pageNumber="528">Ponderia</emphasis>
</taxonomicName>
may represent a plesiomorphic lineage at the base of the pagodulinehaustrine diversification. Thus, those authors (p. 866) placed
<taxonomicName id="4C274D4FFFC5E425FF3DFE40FF5EFE03" authorityName="Houart" authorityYear="1986" box="[133,223,406,432]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="528" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC5E425FF3DFE40FF5EFE03" box="[133,223,406,432]" italics="true" pageId="35" pageNumber="528">Ponderia</emphasis>
</taxonomicName>
in
<emphasis id="B953EADEFFC5E425FEB9FE41FE03FE03" box="[257,386,406,432]" italics="true" pageId="35" pageNumber="528">incertae sedis</emphasis>
, and considered it likely to represent an independent lineage worthy of subfamilial rank.
</paragraph>
</subSubSection>
</treatment>
</document>