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<document id="FC735DDAD375E705309DA73B7413D064" ID-DOI="10.11646/zootaxa.1094.1.2" ID-ISSN="1175­5334" ID-Zenodo-Dep="5051332" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625109556518" checkinUser="felipe" docAuthor="Shelley, Rowland M. &amp; Mercurio, Randy" docDate="2005" docId="924A8786BD08FFA6FEAA60935E04C90D" docLanguage="en" docName="zt01094p040.pdf" docOrigin="Zootaxa 1094 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1094.1.2" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Ectonocryptoides Shelley &amp; Mercurio 2005, new genus" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="6E73FFFEBD00FFAFFFA2630F5F73CB7B" masterDocTitle="Ectonocryptoides quadrimeropus, a new centipede genus and species from Jalisco, Mexico; proposal of Ectonocryptopinae, analysis of subfamilial relationships, and a key to subfamilies and genera of the Scolopocryptopidae (Scolopendromorpha)" masterLastPageNumber="40" masterPageNumber="25" pageNumber="33" updateTime="1698981460483" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4754F9EE1E61C949F8D01EEBC4CD9A6A">Ectonocryptoides quadrimeropus, a new centipede genus and species from Jalisco, Mexico; proposal of Ectonocryptopinae, analysis of subfamilial relationships, and a key to subfamilies and genera of the Scolopocryptopidae (Scolopendromorpha)</mods:title>
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<mods:namePart id="8E437B3763FC198F834484B52D940F7B">Shelley, Rowland M.</mods:namePart>
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<mods:namePart id="5A90D04C237CBEECE9C336CAC720961C">Mercurio, Randy</mods:namePart>
<mods:affiliation id="A7FF7094A2087D27C683C28E7F3470E4">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 St., New York, NY 10024 USA. email mercurio @ amnh. org</mods:affiliation>
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<paragraph id="1A5C3690BD08FFA7FEAA60935D1BC8CD" blockId="8.[264,616,924,950]" box="[264,616,924,950]" pageId="8" pageNumber="33">
<heading id="411481FCBD08FFA7FEAA60935D1BC8CD" bold="true" box="[264,616,924,950]" fontSize="11" level="1" pageId="8" pageNumber="33" reason="1">
<emphasis id="2897EA82BD08FFA7FEAA60935D1BC8CD" bold="true" box="[264,616,924,950]" pageId="8" pageNumber="33">
<taxonomicName id="DDE34D13BD08FFA7FEAA60935EABC8CD" ID-CoL="49YB" authority="Shelley &amp; Mercurio, 2005" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[264,472,924,950]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="2897EA82BD08FFA7FEAA60935EABC8CD" bold="true" box="[264,472,924,950]" italics="true" pageId="8" pageNumber="33">Ectonocryptoides</emphasis>
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,
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<paragraph id="1A5C3690BD08FFA7FEAA60E85CA0CF7A" blockId="8.[264,979,999,1025]" box="[264,979,999,1025]" pageId="8" pageNumber="33">
<emphasis id="2897EA82BD08FFA7FEAA60E85EE9CF7B" box="[264,410,999,1024]" italics="true" pageId="8" pageNumber="33">
<typeStatus id="C5588832BD08FFA7FEAA60E85E4CCF7B" box="[264,319,999,1024]" pageId="8" pageNumber="33">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="DDE34D13BD08FFA7FE0860E85C43CF7B" authority="Shelley &amp; Mercurio, 2005" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[426,816,999,1024]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="quadrimeropus" status="sp. nov.">
<emphasis id="2897EA82BD08FFA7FE0860E85C43CF7B" box="[426,816,999,1024]" italics="true" pageId="8" pageNumber="33">Ectonocryptoides quadrimeropus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="33A457F9BD08FFA7FC9C60E85CBDCF7A" box="[830,974,999,1025]" pageId="8" pageNumber="33" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
<paragraph id="1A5C3690BD08FFA7FEAA67355DCCCF8D" blockId="8.[264,1325,1078,1756]" pageId="8" pageNumber="33">
<emphasis id="2897EA82BD08FFA7FEAA67355E0CCF28" box="[264,383,1082,1107]" italics="true" pageId="8" pageNumber="33">Diagnosis</emphasis>
.
<taxonomicName id="DDE34D13BD08FFA7FE3267355D1ECF2F" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[400,621,1082,1108]" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="subFamily" subFamily="Ectonocryptopinae">Ectonocryptopinae</taxonomicName>
with 4 podomeres on ultimate legs, 2
<superScript id="ED969BD8BD08FFA7FB9567395B34CF3F" attach="left" box="[1079,1095,1078,1092]" fontSize="6" pageId="8" pageNumber="33">nd</superScript>
tarsi absent, tibiae and 1
<superScript id="ED969BD8BD08FFA7FEE8676D5E26CF0B" attach="left" box="[330,341,1122,1136]" fontSize="6" pageId="8" pageNumber="33">st</superScript>
tarsi inflated and bulbous, imparting subclavate appearance to overall appendage, latter with dorsal excavations forming terminal lobes and with short, fine hairs arising in shallow apical depressions; prefemora and femora heavily armored with small spines in addition to ventral spinous processes.
</paragraph>
<paragraph id="1A5C3690BD08FFA7FE9A660A5EA4CE65" blockId="8.[264,1325,1078,1756]" box="[312,471,1284,1310]" pageId="8" pageNumber="33">
<emphasis id="2897EA82BD08FFA7FE9A660A5EE3CE65" box="[312,400,1285,1310]" italics="true" pageId="8" pageNumber="33">Species</emphasis>
. One.
</paragraph>
<paragraph id="1A5C3690BD08FFA7FE9A66225C2CCE3D" blockId="8.[264,1325,1078,1756]" box="[312,863,1324,1350]" pageId="8" pageNumber="33">
<emphasis id="2897EA82BD08FFA7FE9A66225EB4CE3D" box="[312,455,1325,1350]" italics="true" pageId="8" pageNumber="33">Distribution</emphasis>
. Known only from coastal
<collectingRegion id="D827F872BD08FFA7FCA566235C28CE3D" box="[775,859,1324,1350]" country="Mexico" name="Jalisco" pageId="8" pageNumber="33">Jalisco</collectingRegion>
.
</paragraph>
<paragraph id="1A5C3690BD08FFA7FE9A665A5B68CEED" blockId="8.[264,1325,1078,1756]" pageId="8" pageNumber="33">
<emphasis id="2897EA82BD08FFA7FE9A665A5EC0CE15" box="[312,435,1365,1390]" italics="true" pageId="8" pageNumber="33">Etymology</emphasis>
. The generic name reflects the close anatomical similarity to
<taxonomicName id="DDE34D13BD08FFA7FB6C665A5E1ECEED" authorityName="Crabill" authorityYear="1977" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FB6C665A5E1ECEED" italics="true" pageId="8" pageNumber="33">Ectonocryptops</emphasis>
</taxonomicName>
, based on Crabill's written description (1977) of the latter.
</paragraph>
<paragraph id="1A5C3690BD08FFA7FE9A66AA5A5BCDA7" blockId="8.[264,1325,1078,1756]" pageId="8" pageNumber="33">
<emphasis id="2897EA82BD08FFA7FE9A66AA5EEDCEC5" box="[312,414,1445,1470]" italics="true" pageId="8" pageNumber="33">Remarks</emphasis>
. The discovery of a new scolopendromorph genus is a rare event. Only five years into the new millenium, two and three new genus­group names have already been erected in the Lithobiomorpha and
<taxonomicName id="DDE34D13BD08FFA7FD0766FB5C1CCD75" box="[677,879,1524,1550]" order="Geophilomorpha" pageId="8" pageNumber="32" rank="order">Geophilomorpha</taxonomicName>
, respectively;
<taxonomicName id="DDE34D13BD08FFA7FBB466FA5B90CD75" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[1046,1251,1525,1550]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FBB466FA5B90CD75" box="[1046,1251,1525,1550]" italics="true" pageId="8" pageNumber="33">Ectonocryptoides</emphasis>
</taxonomicName>
is not only the first in the 21
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century for the
<taxonomicName id="DDE34D13BD08FFA7FCA765105C85CD42" authorityName=", Crabill" authorityYear="1977" box="[773,1014,1567,1593]" class="Chilopoda" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
, it is also the first since
<bibRefCitation id="7E724B61BD08FFA7FEAA65485EA7CD1A" author="Schileyko, A. A." box="[264,468,1607,1633]" pageId="8" pageNumber="33" pagination="5 - 19" refId="ref7438" refString="Schileyko, A. A. (1992) Scolopenders of Viet-Nam and some aspects of the system of Scolopendromorpha (Chilopoda Epimorpha). Part 1. Arthropoda Selecta, 1 (1), 5 - 19." type="journal article" year="1992">Schileyko (1992)</bibRefCitation>
established
<taxonomicName id="DDE34D13BD08FFA7FDCB65475C60CD1A" authorityName="Schileyko" authorityYear="1992" box="[617,787,1608,1633]" class="Chilopoda" family="Cryptopidae" genus="Tonkinodentus" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FDCB65475C60CD1A" box="[617,787,1608,1633]" italics="true" pageId="8" pageNumber="33">Tonkinodentus</emphasis>
</taxonomicName>
for a Vietnamese species. Only nine generic names have been proposed for scolopendromorphs since 1950
<superScript id="ED969BD8BD08FFA7FC4265605C9BCD06" attach="none" box="[992,1000,1647,1661]" fontSize="6" pageId="8" pageNumber="33">1</superScript>
, whereas around 36 and 78 have been proposed in the Lithobiomorpha and
<taxonomicName id="DDE34D13BD08FFA7FC9865955B77CDCF" box="[826,1028,1690,1716]" order="Geophilomorpha" pageId="8" pageNumber="32" rank="order">Geophilomorpha</taxonomicName>
, respectively, during this same 55 year period, though some are junior subjective synonyms of older names (Jeekel
</paragraph>
<paragraph id="1A5C3690BD08FFA7FEAA64125BC0CC16" blockId="8.[264,1323,1821,1901]" pageId="8" pageNumber="33">
1. They are
<taxonomicName id="DDE34D13BD08FFA7FE2764125DE1CC49" authority="Crabill, 1953" authorityName="Crabill" authorityYear="1953" box="[389,658,1821,1842]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FE2764125D7BCC49" box="[389,520,1821,1842]" italics="true" pageId="8" pageNumber="33">Dinocryptops</emphasis>
Crabill, 1953
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FD3F64125CEECC49" authority="Crabill, 1960" authorityName="Crabill" authorityYear="1960" box="[669,925,1821,1842]" class="Chilopoda" family="Scolopocryptopidae" genus="Thalkethops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FD3F64125C61CC49" box="[669,786,1821,1842]" italics="true" pageId="8" pageNumber="33">Thalkethops</emphasis>
<bibRefCitation id="7E724B61BD08FFA7FCB864125CEECC49" author="Crabill, R. E., Jr." box="[794,925,1821,1842]" pageId="8" pageNumber="33" pagination="1 - 15" refId="ref6854" refString="Crabill, R. E., Jr. (1960). A new American genus of cryptopid centipede, with an annotated key to the scolopendromorph genera from American north of Mexico. Proceedings of the United States National Museum, 111, 1 - 15." type="journal article" year="1960">Crabill, 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FC0A64125A67CC49" authority="Demange, 1963" authorityName="Demange" authorityYear="1963" box="[936,1300,1821,1842]" class="Chilopoda" family="Cryptopidae" genus="Paratrigonocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FC0A64125B02CC49" box="[936,1137,1821,1842]" italics="true" pageId="8" pageNumber="33">Paratrigonocryptops</emphasis>
<bibRefCitation id="7E724B61BD08FFA7FBDB64125A67CC49" author="Demange, J. - M." box="[1145,1300,1821,1842]" pageId="8" pageNumber="33" pagination="41 - 118" refId="ref6959" refString="Demange, J. - M. (1963) La Reserve Naturelle Integrale de Mont Nimba. III. Chilopoda. Memoires de l'Institut Francais d'Afrique Noire, 66, 41 - 118." type="journal article" year="1963">Demange, 1963</bibRefCitation>
</taxonomicName>
, a subgenus of
<taxonomicName id="DDE34D13BD08FFA7FE2964345E90CC2B" authorityName="Leach" authorityYear="1815" box="[395,483,1851,1872]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FE2964345E90CC2B" box="[395,483,1851,1872]" italics="true" pageId="8" pageNumber="33">Cryptops</emphasis>
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FE5664345C61CC34" authority="Dobroruka, 1968" authorityName="Dobroruka" authorityYear="1968" box="[500,786,1850,1872]" class="Chilopoda" family="Scolopendridae" genus="Congobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FE5664345D2FCC2B" box="[500,604,1851,1872]" italics="true" pageId="8" pageNumber="33">Congobius</emphasis>
Dobroruka, 1968
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FC8264345B40CC34" authority="Wurmli, 1972" authorityName="Wurmli" authorityYear="1972" box="[800,1075,1850,1872]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FC8264345CEACC2B" box="[800,921,1851,1872]" italics="true" pageId="8" pageNumber="33">Asanadopsis</emphasis>
<bibRefCitation id="7E724B61BD08FFA7FC0664355B40CC34" author="Wurmli, M." box="[932,1075,1850,1871]" pageId="8" pageNumber="33" pagination="88 - 104" refId="ref8007" refString="Wurmli, M. (1972) Chilopoda von Sumba und Flores I. Scolopendromorpha. Verhandlungen der Naturforschenden Gesellschaft in Basel, 82 (1), 88 - 104." type="journal article" year="1972">Würmli, 1972</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FBE364345E48CC16" authority="Crabill, 1977" authorityName="Crabill" authorityYear="1977" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FBE364345BA4CC2B" box="[1089,1239,1851,1872]" italics="true" pageId="8" pageNumber="33">Ectonocryptops</emphasis>
<bibRefCitation id="7E724B61BD08FFA7FB4064355E48CC16" author="Crabill, R. E., Jr." pageId="8" pageNumber="33" pagination="346 - 349" refId="ref6903" refString="Crabill, R. E., Jr. (1977) A new cryptopid genus, with a key to the genera known to occur in North America including Mexico (Chilopoda: Scolopendromorpha: Cryptopidae). Proceedings of the Entomological Society of Washington, 79 (3), 346 - 349." type="journal article" year="1977">Crabill, 1977</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FEE764575D5ECC16" authority="Koch, 1985" authorityName="Koch" authorityYear="1985" box="[325,557,1880,1901]" class="Chilopoda" family="Scolopendridae" genus="Notiasemus" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FEE764575EC6CC16" box="[325,437,1880,1901]" italics="true" pageId="8" pageNumber="33">Notiasemus</emphasis>
Koch, 1985
</taxonomicName>
;
<taxonomicName id="DDE34D13BD08FFA7FD9564575C14CC16" authority="Schileyko, 1992" authorityName="Schileyko" authorityYear="1992" box="[567,871,1880,1901]" class="Chilopoda" family="Cryptopidae" genus="Tonkinodentus" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="2897EA82BD08FFA7FD9564575DB1CC16" box="[567,706,1880,1901]" italics="true" pageId="8" pageNumber="33">Tonkinodentus</emphasis>
<bibRefCitation id="7E724B61BD08FFA7FD6A64575C14CC16" author="Schileyko, A. A." box="[712,871,1880,1901]" pageId="8" pageNumber="33" pagination="5 - 19" refId="ref7438" refString="Schileyko, A. A. (1992) Scolopenders of Viet-Nam and some aspects of the system of Scolopendromorpha (Chilopoda Epimorpha). Part 1. Arthropoda Selecta, 1 (1), 5 - 19." type="journal article" year="1992">Schileyko, 1992</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="DDE34D13BD08FFA7FC3B64575B32CC16" authority="Shelley &amp; Mercurio, 2005" authorityName="Shelley &amp; Mercurio" authorityYear="2005" box="[921,1089,1880,1901]" class="Chilopoda" family="Scolopocryptopidae" genus="Ectonocryptoides" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="2897EA82BD08FFA7FC3B64575B32CC16" box="[921,1089,1880,1901]" italics="true" pageId="8" pageNumber="33">Ectonocryptoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A457F9BD08FFA7FBE464575BDCCC16" box="[1094,1199,1880,1901]" pageId="8" pageNumber="33" rank="genus">new genus</taxonomicNameLabel>
.
</paragraph>
<paragraph id="1A5C3690BD09FFA6FEAA62135E04C90D" blockId="9.[264,1324,284,630]" pageId="9" pageNumber="34">
2005 plus searches in Zoological Record, the Nomenclator Zoologicus website, and references in RMS' personal library); conversely, only three new genera have been proposed for the Scutigeromorpha, the least diverse global order. As the
<taxonomicName id="DDE34D13BD09FFA6FEAA629B5E86CAD5" authorityName=", Crabill" authorityYear="1977" box="[264,501,404,430]" class="Chilopoda" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="34" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
contains the largest and most visible chilopods, with lengths ranging to over
<quantity id="DD1B9B75BD09FFA6FECE62B35EA1CAAD" box="[364,466,444,470]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="34" unit="mm" value="300.0">300 mm</quantity>
(
<quantity id="DD1B9B75BD09FFA6FE4A62B35D58CAAD" box="[488,555,444,470]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="9" pageNumber="34" unit="in" value="12.0">12 in</quantity>
) (Shelley and Kiser 2000), perhaps man has now discovered, named, and diagnosed most of the obvious genera, even with substantial unexplored areas in the tropics. However, minute, litter­dwelling scolopendromorphs are a different and virtually untouched realm, and where, it would seem, additional genera are most likely to be found.
</paragraph>
</subSubSection>
</treatment>
</document>