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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1014.60426" ID-GBIF-Dataset="97cd82f3-4a3a-43ca-a68f-92a5a2074d07" ID-PMC="PMC7875960" ID-Pensoft-Pub="1313-2970-1014-1" ID-Pensoft-UUID="62961664CAED5F15B9C8A9990D7D388D" ID-PubMed="33603534" ModsDocID="1313-2970-1014-1" checkinTime="1612410909014" checkinUser="pensoft" docAuthor="Achterberg, Cornelis van" docDate="2021" docId="45A7FD2BBC9F59B5A10788318F55B241" docLanguage="en" docName="ZooKeys 1014: 1-118" docOrigin="ZooKeys 1014" docPubDate="2021-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1014.60426" docTitle="Phanerotoma brunneivena Achterberg 2021, sp. nov." docType="treatment" docVersion="4" id="62961664CAED5F15B9C8A9990D7D388D" lastPageNumber="1" masterDocId="62961664CAED5F15B9C8A9990D7D388D" masterDocTitle="The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen" masterLastPageNumber="118" masterPageNumber="1" pageNumber="1" updateTime="1668149759257" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Achterberg, Cornelis van</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, Postbus 9517, 2300 RA Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">kees@vanachterberg.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1014</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>118</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1014.60426</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1014.60426</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1014-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">62961664CAED5F15B9C8A9990D7D388D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177209574" LSID="urn:lsid:zoobank.org:act:3233C1CD-45CE-4ABB-8A4D-5841BFF2CAFC" httpUri="http://treatment.plazi.org/id/45A7FD2BBC9F59B5A10788318F55B241" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/3233C1CD-45CE-4ABB-8A4D-5841BFF2CAFC" authority="Achterberg, 2021" authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma brunneivena" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="brunneivena" status="sp. nov.">Phanerotoma brunneivena</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 55–57" captionStartId="F9" captionText="Figures 55 - 57. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype (but 57 of paratype and 56 of paratype ♂) 55 habitus lateral 56, 57 apical half of metasoma lateral." figureDoi="10.3897/zookeys..60426.figures55-57" httpUri="https://binary.pensoft.net/fig/504632" pageId="0" pageNumber="1">Figs 55-57</figureCitation>
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">, 58-68</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
, ♀ (RMNH), &quot;
<emphasis bold="true" pageId="0" pageNumber="1">United Arab Emirates</emphasis>
, Fujairah (1484), light trap, 19.iv.-2.v.2005,
<geoCoordinate degrees="25" direction="north" minutes="08" orientation="latitude" precision="925" value="25.133333">25°08'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="21" orientation="longitude" precision="925" value="56.35">56°21'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 28♀, 1♂: Idem, 2.v.-5.vi.2005; 14♀: Idem, 19.iv.-2.v.2005; 4♀: Idem, 5-24.iii.2005; 3♀, 1♂: Idem, 24.iii.-6.iv.2005; 7♀, 2♂: Idem, 13-19.iv.2005; 4♀: Idem, 2-13.v.2005; 1♀: Idem, 16-24.ii.2005; 3♀: Idem, 13-29.xi.2005; 2♀: Idem, 29.xi.2005-2.i.2006; 1♀: Idem, 2-13.v.2005; 1♀: Idem, 24.ii.-5.iii.2005; 1♀: &quot;United Arab Emirates, Sharjah Desert Park (1556), light tr[ap], 30.iv.-7.v.2005,
<geoCoordinate degrees="25" direction="north" minutes="17" orientation="latitude" precision="925" value="25.283333">25°17'N</geoCoordinate>
,
<geoCoordinate degrees="55" direction="east" minutes="42" orientation="longitude" precision="925" value="55.7">55°42'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♀: Idem, 21-28.v.2007; 1♀: Idem, 20-21.iv.2006, M. Fibiger; 1♀: &quot;Wadi Safad (5015), white &amp; yellow pan tr., 2-26.i.2006,
<geoCoordinate degrees="25" direction="north" minutes="13" orientation="latitude" precision="925" value="25.216667">25°13'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="19" orientation="longitude" precision="925" value="56.316666">56°19'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♀: &quot;Wadi Maidaq (3808), at light, 21.xii.2005-2.ii.2006,
<geoCoordinate degrees="25" direction="north" minutes="18" orientation="latitude" precision="925" value="25.3">25°18'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="07" orientation="longitude" precision="925" value="56.116665">56°07'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 2♀: Idem, 27.iv.-4.v.2006; 1♀: &quot;United Arab Emirates, Sharjah x Khor Kalba (3851), light trap,
<geoCoordinate degrees="24" direction="north" minutes="59" orientation="latitude" precision="925" value="24.983334">24°59'N</geoCoordinate>
,
<geoCoordinate degrees="56" direction="east" minutes="09" orientation="longitude" precision="925" value="56.15">56°09'E</geoCoordinate>
, 17-18.iv.2006, M. Fibiger,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♀: &quot;United Arab Emirates, NARC near Sweihan (1299), light trap, 9-20.iv.2005,
<geoCoordinate degrees="24" direction="north" minutes="24" orientation="latitude" precision="925" value="24.4">24°24'N</geoCoordinate>
,
<geoCoordinate degrees="55" direction="east" minutes="26" orientation="longitude" precision="925" value="55.433334">55°26'E</geoCoordinate>
, A. v. Harten,
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♀, &quot;
<emphasis bold="true" pageId="0" pageNumber="1">Yemen</emphasis>
(7501), Al Kadan, light trap, i.2003, A. v. Harten &amp; T. Abdul-Haq,
<normalizedToken originalValue="RMNH’03”">RMNH'03&quot;</normalizedToken>
; 1♀: Idem, x.2001; 1♀: Idem, xi.2001; 1♀: Idem, v.2002; 1♀, &quot;Yemen, Al Kowd, light tr[ap], v.-vi.2000, no. 4719, A. v. Harten &amp; S. Al Haruri,
<normalizedToken originalValue="RMNH’01”">RMNH'01&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comparative diagnosis">
<paragraph pageId="0" pageNumber="1">Comparative diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Very similar to
<taxonomicName family="Braconidae" kingdom="Animalia" lsidName="P. flavivena" order="Hymenoptera" pageId="0" pageNumber="1" rank="species" species="flavivena">
<emphasis italics="true" pageId="0" pageNumber="1">P. flavivena</emphasis>
</taxonomicName>
because of the needle-shaped ovipositor sheath of which the apical half is yellow, the yellow pterostigma, the long third tergite (1.8-2.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as second tergite), the distinctly convex temple and short inner tooth of the mandible. It differs by the rugulose or striate and matt to slightly shiny temple (mostly smooth and shiny in
<taxonomicName family="Braconidae" kingdom="Animalia" lsidName="P. flavivena" order="Hymenoptera" pageId="0" pageNumber="1" rank="species" species="flavivena">
<emphasis italics="true" pageId="0" pageNumber="1">P. flavivena</emphasis>
</taxonomicName>
), the dark brown vein r of fore wing ca. 0.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as vein 3-SR (yellow and 0.4-0.7
<normalizedToken originalValue="×">x</normalizedToken>
), the vein SR1 of fore wing distinctly curved (straight or nearly so) and the longer apical triangular appendage of the hypopygium.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female, holotype, length of body (excluding ovipositor) 5.7 mm; antenna 3.8 mm (but apical segment missing); fore wing 3.9 mm; visible part of ovipositor sheath 0.85 mm (only apex setose).</paragraph>
<caption doi="10.3897/zookeys..60426.figures55-57" httpUri="https://binary.pensoft.net/fig/504632" pageId="0" pageNumber="1" start="Figures 55–57" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 55-57.</emphasis>
<taxonomicName authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma brunneivena" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="brunneivena">
<emphasis italics="true" pageId="0" pageNumber="1">Phanerotoma brunneivena</emphasis>
</taxonomicName>
van Achterberg, sp. nov., ♀, holotype (but
<emphasis bold="true" pageId="0" pageNumber="1">57</emphasis>
of paratype and
<emphasis bold="true" pageId="0" pageNumber="1">56</emphasis>
of paratype ♂)
<emphasis bold="true" pageId="0" pageNumber="1">55</emphasis>
habitus lateral
<emphasis bold="true" pageId="0" pageNumber="1">56, 57</emphasis>
apical half of metasoma lateral.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Width 1.5
<normalizedToken originalValue="×">x</normalizedToken>
median length in anterior view and part of head above eye in lateral view 0.3
<normalizedToken originalValue="×">x</normalizedToken>
height of eye (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">65</figureCitation>
); antenna with 23 segments and 1.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than fore wing, segments gradually shortened, narrowed apically and five apical segments moniliform (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">67</figureCitation>
), third, fourth and penultimate segments 3.2, 3.0 and 1.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide in lateral view, respectively; area of stemmaticum coriaceous; OOL: diameter of posterior ocellus: POL = 20: 4: 10; length of eye 1.7
<normalizedToken originalValue="×">x</normalizedToken>
temple in dorsal view (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">63</figureCitation>
); frons coarsely rugose and with median carina; vertex coarsely and densely reticulate-rugose, and rather dull; temple densely and finely rugose, dull; face coarsely rugose and without distinct median ridge or median carina; clypeus mostly smooth, rather shiny and with three minute teeth medio-ventrally, partly hidden by conspicuous fringe (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">64</figureCitation>
); eye medium-sized, strongly convex and in lateral view 1.3
<normalizedToken originalValue="×">x</normalizedToken>
(measured medially) temple (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">65</figureCitation>
), in anterior view 0.8
<normalizedToken originalValue="×">x</normalizedToken>
minimum width of face; upper condyle of mandible near lower level of eyes (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">64</figureCitation>
); malar space rugose, shiny and 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as basal width of mandible; lower tooth of mandible 0.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as apical tooth (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">68</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1" start="Figures 58–68" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 58-68.</emphasis>
<taxonomicName authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma brunneivena" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="brunneivena">
<emphasis italics="true" pageId="0" pageNumber="1">Phanerotoma brunneivena</emphasis>
</taxonomicName>
van Achterberg, sp. nov., ♀, holotype
<emphasis bold="true" pageId="0" pageNumber="1">58</emphasis>
wings
<emphasis bold="true" pageId="0" pageNumber="1">59</emphasis>
mesosoma dorsal
<emphasis bold="true" pageId="0" pageNumber="1">60</emphasis>
first-third metasomal tergites dorsal
<emphasis bold="true" pageId="0" pageNumber="1">61</emphasis>
metasoma lateral
<emphasis bold="true" pageId="0" pageNumber="1">62</emphasis>
apex of third metasomal tergite dorsal
<emphasis bold="true" pageId="0" pageNumber="1">63</emphasis>
head, dorsal
<emphasis bold="true" pageId="0" pageNumber="1">64</emphasis>
head anterior
<emphasis bold="true" pageId="0" pageNumber="1">65</emphasis>
head lateral
<emphasis bold="true" pageId="0" pageNumber="1">66</emphasis>
antenna lateral
<emphasis bold="true" pageId="0" pageNumber="1">67</emphasis>
apical third of antenna lateral
<emphasis bold="true" pageId="0" pageNumber="1">68</emphasis>
mandible ventral.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
(Figs
<figureCitation captionStart="Figures 55–57" captionStartId="F9" captionText="Figures 55 - 57. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype (but 57 of paratype and 56 of paratype ♂) 55 habitus lateral 56, 57 apical half of metasoma lateral." figureDoi="10.3897/zookeys..60426.figures55-57" httpUri="https://binary.pensoft.net/fig/504632" pageId="0" pageNumber="1">55</figureCitation>
,
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">59</figureCitation>
). Length 1.5
<normalizedToken originalValue="×">x</normalizedToken>
its width in lateral view; side of pronotum mainly rugose, but dorsally nearly smooth; mesoscutum densely rugose and rather shiny, densely setose; notauli hardly indicated; scutellar sulcus wide and with seven carinae (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">59</figureCitation>
); scutellum triangular, largely punctate and rather shiny; metanotum with short median carina anteriorly and small tooth protruding posteriorly; propodeum coarsely reticulate-rugose, without distinct median and transverse carinae, latero-posteriorly tuberculate.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Wings</emphasis>
.
</emphasis>
Fore wing 2.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than its maximum width; length of 1-R1 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as pterostigma; r issued far beyond middle of pterostigma and 0.2
<normalizedToken originalValue="×">x</normalizedToken>
3-SR; 2-SR curved and distally subparallel with posterior margin of pterostigma (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">58</figureCitation>
); SR1 curved; 2-SR+M absent because of interstitial m-cu; parastigma large; 1-CU1 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as vein 2-CU1; r:3-SR:SR1 = 2:10:25; 2-SR:3-SR:r-m = 25:10:7; r-m reclivous; 2-M weakly curved (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">58</figureCitation>
). Hind wing: M+CU:1-M:1r-m = 21:20:10; r feebly indicated.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
.
</emphasis>
Hind femur 3.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide and widened subbasally; middle tibia with small ivory blister; inner spur of middle tibia 0.5
<normalizedToken originalValue="×">x</normalizedToken>
its basitarsus; hind coxa mostly smooth and shiny.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Metasoma</emphasis>
(Figs
<figureCitation captionStart="Figures 55–57" captionStartId="F9" captionText="Figures 55 - 57. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype (but 57 of paratype and 56 of paratype ♂) 55 habitus lateral 56, 57 apical half of metasoma lateral." figureDoi="10.3897/zookeys..60426.figures55-57" httpUri="https://binary.pensoft.net/fig/504632" pageId="0" pageNumber="1">57</figureCitation>
,
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">60-62</figureCitation>
). Elongate elliptical in dorsal view, twice as long as wide and 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as mesosoma; first and second tergites coarsely longitudinally rugose and rugae with interconnections; third tergite 2.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than second tergite and laterally nearly straight, in lateral view rather flat, densely reticulate-rugulose and medio-posteriorly distinctly concave (Figs
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">60</figureCitation>
,
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">62</figureCitation>
), lateral lamella narrow, protruding latero-apically and medio-apically absent (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">62</figureCitation>
); ovipositor sheath narrow, needle-shaped (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">61</figureCitation>
), its visible part 0.22
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore wing and 0.38
<normalizedToken originalValue="×">x</normalizedToken>
metasomal carapace and only its apex with small cluster of setae; hypopygium apically with medium-sized and slender bent up triangle, without apical spine and with medium-sized sparse setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Pale brownish yellow; palpi, mandible (except dark brown teeth), clypeus, pronotum, legs (but hind tibia rather darkened apically) first and second tergites largely and metasoma ventrally pale yellow or ivory; apical antennal segments and stemmaticum brown; pterostigma brownish yellow but anteriorly pale yellowish (Fig.
<figureCitation captionStart="Figures 58–68" captionStartId="F10" captionText="Figures 58 - 68. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype 58 wings 59 mesosoma dorsal 60 first-third metasomal tergites dorsal 61 metasoma lateral 62 apex of third metasomal tergite dorsal 63 head, dorsal 64 head anterior 65 head lateral 66 antenna lateral 67 apical third of antenna lateral 68 mandible ventral." figureDoi="10.3897/zookeys..60426.figures58-68" httpUri="https://binary.pensoft.net/fig/504633" pageId="0" pageNumber="1">58</figureCitation>
); wing membrane subhyaline; parastigma (but brownish posteriorly), veins 1-M, 2-CU1 and m-cu of fore wing yellow and veins r, 1-CU1, cu-a, 2-SR, 3-SR and 2-M dark brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="male">
<paragraph pageId="0" pageNumber="1">Male.</paragraph>
<paragraph pageId="0" pageNumber="1">
Similar to female but third tergite narrowed posteriorly, rounded and convex apically (Fig.
<figureCitation captionStart="Figures 55–57" captionStartId="F9" captionText="Figures 55 - 57. Phanerotoma brunneivena van Achterberg, sp. nov., ♀, holotype (but 57 of paratype and 56 of paratype ♂) 55 habitus lateral 56, 57 apical half of metasoma lateral." figureDoi="10.3897/zookeys..60426.figures55-57" httpUri="https://binary.pensoft.net/fig/504632" pageId="0" pageNumber="1">56</figureCitation>
; one male has a subapical bump) and pterostigma (except anteriorly) brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variations.</paragraph>
<paragraph pageId="0" pageNumber="1">
Length of fore wing of ♀ 3.1-4.3 mm, of ♂ 3.0-3.5 mm; antenna of ♀ with 23 segments; third tergite 1.8-2.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than second tergite, of ♀ medio-apically concave or obtuse and with no or a narrow lamella, flat or with a small subapical bump; ovipositor sheath up to 0.36
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore wing if exserted; rarely scutellum laterally and propodeum apically dark brown, wing membrane below pterostigma and near vein 1-M somewhat infuscate; third tergite rarely dark brown basally or with dark triangle; temple and mesoscutum medially pale brownish yellow or ivory; sometimes pterostigma darkened but anteriorly pale yellowish.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">United Arabian Emirates, Yemen.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Named after the brownish vein r of the fore wing (
<emphasis italics="true" pageId="0" pageNumber="1">brunneus</emphasis>
is Latin for brown).
</paragraph>
</subSubSection>
</treatment>
</document>
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