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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.526.6037" ID-GBIF-Dataset="46cfb24b-675a-4bae-88a3-b55240ef8f8d" ID-PMC="PMC4607843" ID-Pensoft-Pub="1313-2970-526-19" ID-PubMed="26487823" ID-ZBK="648DED714CAD42E0A19DF2E7DB27EE05" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-526-19" ModsDocOrigin="ZooKeys 526" ModsDocTitle="Diagnostic survey of Malagasy Nesomyrmex species-groups and revision of hafahafa group species via morphology based cluster delimitation protocol" checkinTime="1451243915363" checkinUser="pensoft" docAuthor="Csosz, Sandor &amp; Fisher, Brian L." docDate="2015" docId="F5495C1F75CDABB78AEF4B7FB50DABE4" docLanguage="en" docName="ZooKeys 526: 19-59" docOrigin="ZooKeys 526" docSource="http://dx.doi.org/10.3897/zookeys.526.6037" docTitle="Nesomyrmex hafahafa Csosz &amp; Fisher, sp. n." docType="treatment" docUuid="C2249F7A-0FFE-4C76-A2E8-905A4B1EA754" docUuidSource="ZooBank" docVersion="4" lastPageNumber="35" masterDocId="FF89FFBBD738BB54FFC1FF9EFFE7FF9C" masterDocTitle="Diagnostic survey of Malagasy Nesomyrmex species-groups and revision of hafahafa group species via morphology based cluster delimitation protocol" masterLastPageNumber="59" masterPageNumber="19" pageNumber="32" updateTime="1668160833139" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Diagnostic survey of Malagasy Nesomyrmex species-groups and revision of hafahafa group species via morphology based cluster delimitation protocol</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Csosz, Sandor</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>526</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>59</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.526.6037</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.526.6037</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-526-19</mods:identifier>
<mods:identifier type="ZBK">648DED714CAD42E0A19DF2E7DB27EE05</mods:identifier>
<mods:identifier type="ZooBank">648DED714CAD42E0A19DF2E7DB27EE05</mods:identifier>
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<treatment ID-GBIF-Taxon="152062666" LSID="urn:lsid:zoobank.org:act:C2249F7A-0FFE-4C76-A2E8-905A4B1EA754" httpUri="http://treatment.plazi.org/id/F5495C1F75CDABB78AEF4B7FB50DABE4" lastPageId="16" lastPageNumber="35" pageId="13" pageNumber="32">
<subSubSection pageId="13" pageNumber="32" type="multiple">
<paragraph pageId="13" pageNumber="32">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="32" type="nomenclature">
<paragraph pageId="13" pageNumber="32">
<taxonomicName LSID="http://zoobank.org/C2249F7A-0FFE-4C76-A2E8-905A4B1EA754" authority="Csosz &amp; Fisher" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex hafahafa" order="Hymenoptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="hafahafa">
Nesomyrmex hafahafa
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
&amp; Fisher
</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="32">sp. n.</taxonomicNameLabel>
Figs 22-24, Table 4
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="33" type="etymology">
<paragraph pageId="14" pageNumber="33">
<pageBreakToken pageId="14" pageNumber="33" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="14" pageNumber="33">
This Malagasy word
<normalizedToken originalValue="“hafahafa”">&quot;hafahafa&quot;</normalizedToken>
means weird, and refers to the unusual morphology of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="33" type="type material">
<paragraph pageId="14" pageNumber="33">Type material investigated.</paragraph>
<paragraph pageId="14" pageNumber="33">
Holotype worker.CASENT0460666, collection code: BLF06010;
<normalizedToken originalValue="MADGR">MADG'R</normalizedToken>
: Prov. Toliara,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
de Tsinjoriaky, 6.2 km 84° E Tsifota,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-22.8">22°48'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="43.416668">43°25'E</geoCoordinate>
[-
<geoCoordinate direction="north" orientation="latitude" precision="1" value="22.80222">22.80222N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="43.42067">43.42067E</geoCoordinate>
], 70 m, 6-10.iii.2002 Fisher et al. (CAS)
</paragraph>
<paragraph pageId="14" pageNumber="33">
Paratypes. Ten workers, a single gyne and two males with the same label data as the holotype under CASENT codes: CASENT0746771, BLF06010, (2w, CAS); CASENT0460667, BLF06010, (3w, CAS); CASENT0460668, BLF06010, (3w, CAS); CASENT0460669, BLF06010, (1q, CAS); CASENT0451364,
<normalizedToken originalValue="“6019”">&quot;6019&quot;</normalizedToken>
, (2w, CAS); CASENT0451364,
<normalizedToken originalValue="“6019”">&quot;6019&quot;</normalizedToken>
, (2m, CAS);
</paragraph>
<paragraph pageId="14" pageNumber="33">The list of 44 non-type individuals belonging to 25 nest samples of other material investigated is given in Table 1.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="33" type="diagnosis">
<paragraph pageId="14" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="14" pageNumber="33">In key.</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="35" pageId="14" pageNumber="33" type="description of workers">
<paragraph pageId="14" pageNumber="33">Description of workers.</paragraph>
<paragraph lastPageId="16" lastPageNumber="35" pageId="14" pageNumber="33">
Body color: yellow; brown. Body color pattern: body concolorous, only clava darker. Absolute cephalic size: 1062 [974, 1142]
<normalizedToken originalValue="µm">µm</normalizedToken>
(n = 48). Cephalic length vs. maximum width of head capsule (CL/CWb): 1.224 [1.193-1.254]. Postocular distance vs. cephalic length (PoOc/CL): 0.388 [0.361, 0.406]. Postocular sides of cranium contour frontal view orientation: converging posteriorly. Postocular sides of cranium contour frontal view shape: broadly convex. Vertex contour line in frontal view shape: straight; slightly concave. Vertex sculpture: main sculpture
<pageBreakToken pageId="15" pageNumber="34" start="start">rugose</pageBreakToken>
, ground sculpture areolate. Gena contour line in frontal view shape: feebly convex. Genae contour from anterior view orientation: converging. Gena sculpture: rugo-reticulate with areolate ground sculpture. Concentric carinae laterally surrounding antennal foramen count: present. Eye length vs. absolute cephalic size (EL/CS): 0.230 [0.212, 0.248]. Frontal carina distance vs. absolute cephalic size (FRS/CS): 0.316 [0.289, 0.333]. Longitudinal carinae on median region of frons count: present. Longitudinal carinae on medial region of frons shape: forked. Smooth median region on frons count: absent. Antennomere count: 12. Scape length vs. absolute cephalic size (SL/CS): 0.895 [0.861, 0.927]. Facial area of the scape absolute setal angle: setae absent, pubescence only. Median clypeal notch count: present. Median clypeal notch depth vs. absolute cephalic size (Cdep/CS): 0.022 [0.015, 0.029]. Ground sculpture of submedian area of clypeus: smooth. Median carina of clypeus count: present. Lateral carinae of clypeus count: present. Median anatomical line of propodeal spine an
<pageBreakToken pageId="16" pageNumber="35" start="start">gle</pageBreakToken>
value to Weber length in lateral view: 55-60°. Spine length vs. absolute cephalic size (SPST/CS): 0.398 [0.355, 0.427]. Minimum spine distance vs. absolute cephalic size (SPBA/CS): 0.287 [0.257, 0.311]. Apical spine distance vs. absolute cephalic size (SPTI/CS): 0.543 [0.463, 0.607]. Propodeal spine shape: strongly bent. Apical distance of pronotal spines vs. absolute cephalic size (PSTI/CS): 0.724 [0.631, 0.776]. Metanotal depression count: absent. Dorsal region of mesosoma sculpture: areolate ground sculpture, superimposed by dispersed rugae. Lateral region of pronotum sculpture: areolate ground sculpture, superimposed by dispersed rugae. Mesopleuron sculpture: areolate ground sculpture superimposed by dispersed rugulae; areolate ground sculpture, superimposed by dispersed rugae. Metapleuron sculpture: areolate ground sculpture, superimposed by dispersed rugae. Petiole width vs. absolute cephalic size (PEW/CS): 0.307 [0.275, 0.357]. Anterodorsal spines on petiolar node angle of deviation from each other: 80°. Apical distance of anterodorsal spines on petiolar node vs. absolute cephalic size (NSTI/CS): 0.514 [0.473, 0.563]. Frontal profile of petiolar node contour line in lateral view shape: convex. Dorso-caudal petiolar profile contour line in lateral view shape: convex. Dorsal region of petiole sculpture: ground sculpture areolate, main sculpture dispersed rugose. Postpetiole width vs. absolute cephalic size (PPW/CS): 0.538 [0.494, 0.576]. Dorsal region of postpetiole sculpture: ground sculpture areolate, main sculpture dispersed rugose.
</paragraph>
<caption pageId="16" pageNumber="35">
<paragraph pageId="16" pageNumber="35">
Figures 22-24.
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex hafahafa" order="Hymenoptera" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="hafahafa">Nesomyrmex hafahafa</taxonomicName>
sp. n. holotype worker (CASENT0460666). Lateral view of the body (22) head of the holotype worker in full-face view (23), dorsal view of the body (24). Scale 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="16" pageNumber="35" type="distribution">
<paragraph pageId="16" pageNumber="35">Distribution.</paragraph>
<paragraph pageId="16" pageNumber="35">This species is widely distributed along the western forests of Madagascar (Fig. 13) between the 23rd and 20th southern latitudes.</paragraph>
</subSubSection>
</treatment>
</document>