treatments-xml/data/FA/49/F1/FA49F17761B956148B45774DE0868FD8.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e66021" ID-GBIF-Dataset="1f858329-d759-403a-829c-08f2ba4e1403" ID-GBIF-Taxon="181520721" ID-Pensoft-Pub="1864-8312-79-57" ID-Pensoft-UUID="B1CDE34C960C59639EA47231EE68A71F" ID-ZooBank="A42504B7FFAF4856884A7B74F3E03AA3" ModsDocID="1864-8312-79-57" checkinTime="1618938684678" checkinUser="pensoft" docAuthor="Grebennikov, Vasily V." docDate="2021" docId="FA49F17761B956148B45774DE0868FD8" docLanguage="en" docName="ArthropodSystPhyl 79: 57-74" docOrigin="Arthropod Systematics &amp; Phylogeny 79" docPubDate="2021-04-20" docSource="http://dx.doi.org/10.3897/asp.79.e66021" docTitle="Subanchonidium Hoffmann 1968, syn. nov." docType="treatment" docVersion="4" id="B1CDE34C960C59639EA47231EE68A71F" lastPageNumber="57" masterDocId="B1CDE34C960C59639EA47231EE68A71F" masterDocTitle="Sky islands of the Cameroon Volcanic Line support the westernmost clade of five new Typoderus weevils (Coleoptera: Curculionidae: Molytinae)" masterLastPageNumber="74" masterPageNumber="57" pageId="0" pageNumber="57" updateTime="1632015666906" updateUser="ExternalLinkService">
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<mods:title>Sky islands of the Cameroon Volcanic Line support the westernmost clade of five new Typoderus weevils (Coleoptera: Curculionidae: Molytinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grebennikov, Vasily V.</mods:namePart>
<mods:affiliation>Canadian Food Inspection Agency, 960 Carling Ave., Ottawa, ON, K 1 A 0 Y 9, Canada</mods:affiliation>
<mods:nameIdentifier type="email">vasily.grebennikov@canada.ca</mods:nameIdentifier>
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<mods:title>Arthropod Systematics &amp; Phylogeny</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-20</mods:number>
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<mods:detail type="volume">
<mods:number>79</mods:number>
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<mods:start>57</mods:start>
<mods:end>74</mods:end>
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<mods:url>http://dx.doi.org/10.3897/asp.79.e66021</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e66021</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-79-57</mods:identifier>
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<treatment ID-GBIF-Taxon="181520721" LSID="urn:lsid:plazi:treatment:FA49F17761B956148B45774DE0868FD8" httpUri="http://treatment.plazi.org/id/FA49F17761B956148B45774DE0868FD8" lastPageNumber="57" pageId="0" pageNumber="57">
<subSubSection pageId="0" pageNumber="57" type="nomenclature">
<paragraph pageId="0" pageNumber="57">
= 
<taxonomicName LSID="FA49F177-61B9-5614-8B45-774DE0868FD8" authority="Hoffmann, 1968" authorityName="Hoffmann" authorityYear="1968" class="Insecta" family="Curculionidae" genus="Subanchonidium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Subanchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus" status="syn. nov.">Subanchonidium Hoffmann, 1968</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="57">syn. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="reference_group">
<paragraph pageId="0" pageNumber="57">
<taxonomicName class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium (Subanchonidium)" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subGenus" status="syn. nov." subGenus="Subanchonidium">Anchonidium Subanchonidium</taxonomicName>
<bibRefCitation author="Hoffmann, A" journalOrPublisher="Opuscula Zoologica, Budapest" pageId="0" pageNumber="57" pagination="11 - 29" refId="B29" refString="Hoffmann, A, 1968. The scientific results of the Hungarian soil zoological expedition to the Brazzaville-Congo. 32. Especes de la famille Curculionidae (Coleoptera). Opuscula Zoologica, Budapest 8: 11 - 29" title="The scientific results of the Hungarian soil zoological expedition to the Brazzaville-Congo. 32. Especes de la famille Curculionidae (Coleoptera)." volume="8" year="1968">Hoffmann 1968</bibRefCitation>
: 23 (as subgenus of 
<taxonomicName authorityName="Bedel" authorityYear="1884" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus" status="syn. nov.">Anchonidium</taxonomicName>
Bedel, 1884; species included: 
<taxonomicName authorityName="Hoffmann" authorityYear="1968" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium baloghi" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="baloghi" status="syn. nov.">Anchonidium baloghi</taxonomicName>
). Type species: 
<taxonomicName authorityName="Hoffmann" authorityYear="1968" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium baloghi" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="baloghi" status="syn. nov.">Anchonidium baloghi</taxonomicName>
Hoffmann, 1968 by monotypy.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="available evidence">
<paragraph pageId="0" pageNumber="57">Available evidence.</paragraph>
<paragraph pageId="0" pageNumber="57">
The genus 
<taxonomicName authorityName="Bedel" authorityYear="1884" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Anchonidium</emphasis>
</taxonomicName>
was recently reduced to include only five West Palaearctic species (of them two from Portugal recently described, Germann 2020), while all but two 
<taxonomicName authorityName="Bedel" authorityYear="1884" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Anchonidium</emphasis>
</taxonomicName>
from the Afrotropical Region were transferred to a re-defined 
<taxonomicName authorityName="Hustache" authorityYear="1939" class="Insecta" family="Curculionidae" genus="Aparopionella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aparopionella" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Aparopionella</emphasis>
</taxonomicName>
Hustache, 1939 (
<bibRefCitation author="Grebennikov, VV" journalOrPublisher="Entomologische Blaetter und Coleoptera" pageId="0" pageNumber="57" pagination="219 - 231" refId="B19" refString="Grebennikov, VV, 2018. Re-defined Aparopionella from Afrotropical forest floors comprises 20 poorly known nominal species (Coleoptera, Curculionidae, Molytinae). Entomologische Blaetter und Coleoptera 114: 219 - 231" title="Re-defined Aparopionella from Afrotropical forest floors comprises 20 poorly known nominal species (Coleoptera, Curculionidae, Molytinae)." volume="114" year="2018">Grebennikov 2018</bibRefCitation>
). Two 
<taxonomicName authorityName="Bedel" authorityYear="1884" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Anchonidium</emphasis>
</taxonomicName>
species described by 
<bibRefCitation author="Hoffmann, A" journalOrPublisher="Opuscula Zoologica, Budapest" pageId="0" pageNumber="57" pagination="11 - 29" refId="B29" refString="Hoffmann, A, 1968. The scientific results of the Hungarian soil zoological expedition to the Brazzaville-Congo. 32. Especes de la famille Curculionidae (Coleoptera). Opuscula Zoologica, Budapest 8: 11 - 29" title="The scientific results of the Hungarian soil zoological expedition to the Brazzaville-Congo. 32. Especes de la famille Curculionidae (Coleoptera)." volume="8" year="1968">Hoffmann (1968)</bibRefCitation>
from the Republic of the Congo as type species of two monotypic subgenera of 
<taxonomicName authorityName="Bedel" authorityYear="1884" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Anchonidium</emphasis>
</taxonomicName>
were noted as belonging to neither 
<taxonomicName authorityName="Bedel" authorityYear="1884" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Anchonidium</emphasis>
</taxonomicName>
nor 
<taxonomicName authorityName="Hustache" authorityYear="1939" class="Insecta" family="Curculionidae" genus="Aparopionella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aparopionella" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Aparopionella</emphasis>
</taxonomicName>
(
<bibRefCitation author="Grebennikov, VV" journalOrPublisher="Entomologische Blaetter und Coleoptera" pageId="0" pageNumber="57" pagination="219 - 231" refId="B19" refString="Grebennikov, VV, 2018. Re-defined Aparopionella from Afrotropical forest floors comprises 20 poorly known nominal species (Coleoptera, Curculionidae, Molytinae). Entomologische Blaetter und Coleoptera 114: 219 - 231" title="Re-defined Aparopionella from Afrotropical forest floors comprises 20 poorly known nominal species (Coleoptera, Curculionidae, Molytinae)." volume="114" year="2018">Grebennikov 2018</bibRefCitation>
). A decision on the taxonomic status of these four names was long delayed by the unavailability of the type series. These historical specimens were borrowed in 2007 from Hungarian Natural History Museum (= HNHM, Budapest, Hungary) by Nicolas Maughan (Aix-Marseille University, Marseille, France; loan #4240/7343) and not returned as of September 2019, despite numerous demands (personal communication, 
<normalizedToken originalValue="Ottó">Otto</normalizedToken>
Merkl, curator of HNHM 
<taxonomicName authorityName="Hustache" authorityYear="1939" class="Insecta" genus="Aparopionella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
collection). Requests for high-resolution images of the type specimens resulted in a low-resolution dorsal view of the holotype of 
<taxonomicName genus="A." lsidName="A. distinctum" pageId="0" pageNumber="57" rank="species" species="distinctum">
<emphasis italics="true" pageId="0" pageNumber="57">A. distinctum</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the genus Typoderus. Map designed using the online SimpleMappr tool (Shorthouse 2010)." figureDoi="10.3897/asp.79.e66021.figure1" httpUri="https://binary.pensoft.net/fig/533979" pageId="0" pageNumber="57">1</figureCitation>
). The image revealed a weevil consistent with 
<taxonomicName authorityName="Marshall" authorityYear="1953" class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Typoderus</emphasis>
</taxonomicName>
in its appearance (parallel-sided body, effaced elytral shoulders, two longitudinal ridges on each side of pronotum, of them the internal one zig-zag shaped). The last feature is synapomorphic for the clade of 
<taxonomicName authorityName="Marshall" authorityYear="1953" class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Typoderus</emphasis>
</taxonomicName>
plus 
<taxonomicName authority="" authorityName="" class="Insecta" family="Curculionidae" genus="Lupangus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lupangus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Lupangus</emphasis>
</taxonomicName>
. The beetle in Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the genus Typoderus. Map designed using the online SimpleMappr tool (Shorthouse 2010)." figureDoi="10.3897/asp.79.e66021.figure1" httpUri="https://binary.pensoft.net/fig/533979" pageId="0" pageNumber="57">1</figureCitation>
differs from 
<taxonomicName authority="" authorityName="" class="Insecta" family="Curculionidae" genus="Lupangus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lupangus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Lupangus</emphasis>
</taxonomicName>
by having an anteriorly directed rostrum (more ventrally directed in 
<taxonomicName authority="" authorityName="" class="Insecta" family="Curculionidae" genus="Lupangus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lupangus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Lupangus</emphasis>
</taxonomicName>
), by lacking transverse dorsal groove behind eyes (present in 
<taxonomicName authority="" authorityName="" class="Insecta" family="Curculionidae" genus="Lupangus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lupangus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Lupangus</emphasis>
</taxonomicName>
) and by the more rounded eyes (vertical in 
<taxonomicName authority="" authorityName="" class="Insecta" family="Curculionidae" genus="Lupangus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lupangus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Lupangus</emphasis>
</taxonomicName>
, fig. 4 in 
<bibRefCitation DOI="https://doi.org/10.4081/fe.2017.229" author="Grebennikov, VV" journalOrPublisher="Fragmenta entomologica" pageId="0" pageNumber="57" pagination="37 - 55" refId="B18" refString="Grebennikov, VV, 2017. Phylogeography and sister group of Lupangus, a new genus for three new flightless allopatric forest litter weevils endemic to the Eastern Arc Mountains, Tanzania (Coleoptera: Curculionidae, Molytinae). Fragmenta entomologica 49: 37 - 55, DOI: https://doi.org/10.4081/fe.2017.229" title="Phylogeography and sister group of Lupangus, a new genus for three new flightless allopatric forest litter weevils endemic to the Eastern Arc Mountains, Tanzania (Coleoptera: Curculionidae, Molytinae)." url="https://doi.org/10.4081/fe.2017.229" volume="49" year="2017">Grebennikov 2017</bibRefCitation>
). These considerations, together with sympatry and great similarity of this species with 
<taxonomicName genus="A." lsidName="A. baloghi" pageId="0" pageNumber="57" rank="species" species="baloghi">
<emphasis italics="true" pageId="0" pageNumber="57">A. baloghi</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="“extrêmement">&quot;extremement</normalizedToken>
voisin de 
<taxonomicName authorityName="Hoffmann" authorityYear="1968" class="Insecta" family="Curculionidae" genus="Anchonidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchonidium baloghi" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis italics="true" pageId="0" pageNumber="57">Anchonidium baloghi</emphasis>
</taxonomicName>
...&quot;, 
<bibRefCitation author="Hoffmann, A" journalOrPublisher="Opuscula Zoologica, Budapest" pageId="0" pageNumber="57" pagination="11 - 29" refId="B29" refString="Hoffmann, A, 1968. The scientific results of the Hungarian soil zoological expedition to the Brazzaville-Congo. 32. Especes de la famille Curculionidae (Coleoptera). Opuscula Zoologica, Budapest 8: 11 - 29" title="The scientific results of the Hungarian soil zoological expedition to the Brazzaville-Congo. 32. Especes de la famille Curculionidae (Coleoptera)." volume="8" year="1968">Hoffmann 1968</bibRefCitation>
: 24) are the only data available to conclude that both species taxonomically belong to the genus 
<taxonomicName authorityName="Marshall" authorityYear="1953" class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Typoderus</emphasis>
</taxonomicName>
and are, therefore herein transferred to the later as 
<taxonomicName class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus distinctus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis italics="true" pageId="0" pageNumber="57">Typoderus distinctus</emphasis>
</taxonomicName>
(Hoffmann, 1968) 
<emphasis bold="true" pageId="0" pageNumber="57">comb. nov.</emphasis>
and 
<taxonomicName class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus baloghi" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="baloghi">
<emphasis italics="true" pageId="0" pageNumber="57">Typoderus baloghi</emphasis>
</taxonomicName>
(Hoffmann, 1968) 
<emphasis bold="true" pageId="0" pageNumber="57">comb. nov.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>