Sky islands of the Cameroon Volcanic Line support the westernmost clade of five new Typoderus weevils (Coleoptera: Curculionidae: Molytinae)
Author
Grebennikov, Vasily V.
Canadian Food Inspection Agency, 960 Carling Ave., Ottawa, ON, K 1 A 0 Y 9, Canada
vasily.grebennikov@canada.ca
text
Arthropod Systematics & Phylogeny
2021
2021-04-20
79
57
74
http://dx.doi.org/10.3897/asp.79.e66021
journal article
http://dx.doi.org/10.3897/asp.79.e66021
1864-8312-79-57
A42504B7FFAF4856884A7B74F3E03AA3
B1CDE34C960C59639EA47231EE68A71F
=
Subanchonidium Hoffmann, 1968
syn. nov.
Anchonidium Subanchonidium
Hoffmann 1968
: 23 (as subgenus of
Anchonidium
Bedel, 1884; species included:
Anchonidium baloghi
). Type species:
Anchonidium baloghi
Hoffmann, 1968 by monotypy.
Available evidence.
The genus
Anchonidium
was recently reduced to include only five West Palaearctic species (of them two from Portugal recently described, Germann 2020), while all but two
Anchonidium
from the Afrotropical Region were transferred to a re-defined
Aparopionella
Hustache, 1939 (
Grebennikov 2018
). Two
Anchonidium
species described by
Hoffmann (1968)
from the Republic of the Congo as type species of two monotypic subgenera of
Anchonidium
were noted as belonging to neither
Anchonidium
nor
Aparopionella
(
Grebennikov 2018
). A decision on the taxonomic status of these four names was long delayed by the unavailability of the type series. These historical specimens were borrowed in 2007 from Hungarian Natural History Museum (= HNHM, Budapest, Hungary) by Nicolas Maughan (Aix-Marseille University, Marseille, France; loan #4240/7343) and not returned as of September 2019, despite numerous demands (personal communication,
Otto
Merkl, curator of HNHM
Coleoptera
collection). Requests for high-resolution images of the type specimens resulted in a low-resolution dorsal view of the holotype of
A. distinctum
(Fig.
1
). The image revealed a weevil consistent with
Typoderus
in its appearance (parallel-sided body, effaced elytral shoulders, two longitudinal ridges on each side of pronotum, of them the internal one zig-zag shaped). The last feature is synapomorphic for the clade of
Typoderus
plus
Lupangus
. The beetle in Fig.
1
differs from
Lupangus
by having an anteriorly directed rostrum (more ventrally directed in
Lupangus
), by lacking transverse dorsal groove behind eyes (present in
Lupangus
) and by the more rounded eyes (vertical in
Lupangus
, fig. 4 in
Grebennikov 2017
). These considerations, together with sympatry and great similarity of this species with
A. baloghi
(
"extremement
voisin de
Anchonidium baloghi
...",
Hoffmann 1968
: 24) are the only data available to conclude that both species taxonomically belong to the genus
Typoderus
and are, therefore herein transferred to the later as
Typoderus distinctus
(Hoffmann, 1968)
comb. nov.
and
Typoderus baloghi
(Hoffmann, 1968)
comb. nov.