189 lines
25 KiB
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189 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.362.6300" ID-PMC="PMC3867105" ID-Pensoft-Pub="1313-2970-362-29" ID-Pensoft-UUID="EA63FFD35254FFFC603C7A5E8D08FFA0" ID-PubMed="24363599" ID-Zenodo-Dep="578109" ID-ZooBank="569812E982B54BCF84E0D4C43AEE8273" ModsDocID="1313-2970-362-29" checkinTime="1451246566598" checkinUser="pensoft" docAuthor="Erwin, Terry L. & Amundson, Lauren M." docDate="2013" docId="62A40F885EC957CE961FD67BC748BB78" docLanguage="en" docName="ZooKeys 362: 29-54" docOrigin="ZooKeys 362" docPubDate="2013-12-13" docSource="http://dx.doi.org/10.3897/zookeys.362.6300" docTitle="Pseudomorpha (Pseudomorpha) patagonia Erwin & Amundson 2013, sp. n." docType="treatment" docUuid="B32D721B-B86C-4A56-8D4A-CCAAAF8A1318" docUuidSource="ZooBank" docVersion="3" id="EA63FFD35254FFFC603C7A5E8D08FFA0" lastPageNumber="38" masterDocId="EA63FFD35254FFFC603C7A5E8D08FFA0" masterDocTitle="Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group" masterLastPageNumber="54" masterPageNumber="29" pageNumber="36" updateTime="1668157171424" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Erwin, Terry L.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Amundson, Lauren M.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group Summer Intern, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-12-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>362</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.362.6300</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.362.6300</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-362-29</mods:identifier>
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<mods:identifier type="ZooBank">569812E982B54BCF84E0D4C43AEE8273</mods:identifier>
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<mods:identifier type="Pensoft-UUID">EA63FFD35254FFFC603C7A5E8D08FFA0</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578109</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182235302" LSID="urn:lsid:zoobank.org:act:B32D721B-B86C-4A56-8D4A-CCAAAF8A1318" httpUri="http://treatment.plazi.org/id/62A40F885EC957CE961FD67BC748BB78" lastPageId="9" lastPageNumber="38" pageId="7" pageNumber="36">
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<subSubSection pageId="7" pageNumber="36" type="nomenclature">
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<paragraph pageId="7" pageNumber="36">
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<taxonomicName LSID="http://zoobank.org/B32D721B-B86C-4A56-8D4A-CCAAAF8A1318" authority="Erwin & Amundson" authorityName="Erwin & Amundson" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pseudomorpha (Pseudomorpha) patagonia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="patagonia" status="sp. n." subGenus="Pseudomorpha">Pseudomorpha (Pseudomorpha) patagonia Erwin & Amundson</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="36">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="7" pageNumber="36">Figures 2</figureCitation>
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="7" pageNumber="36">, 8</figureCitation>
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="7" pageNumber="36">, 14</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F4" captionText="Figure 18. Distribution map for species of the santarita group of Pseudomorpha." httpUri="https://binary.pensoft.net/fig/27292" pageId="7" pageNumber="36">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="36" type="holotype">
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<paragraph pageId="7" pageNumber="36">Holotype.</paragraph>
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<paragraph pageId="7" pageNumber="36">
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<emphasis bold="true" pageId="7" pageNumber="36">USA</emphasis>
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. Arizona,
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<emphasis italics="true" pageId="7" pageNumber="36">Santa Cruz County</emphasis>
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, Patagonia Mountains, Harshaw Creek,
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<geoCoordinate degrees="31.439" direction="north" orientation="latitude" precision="55" value="31.439">31.439°N</geoCoordinate>
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,
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<geoCoordinate degrees="110.696" direction="west" orientation="longitude" precision="55" value="-110.696">110.696°W</geoCoordinate>
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, 1577m, 1 August 1979 (S. McCleve) (UATC: ADP110694, male).
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</paragraph>
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<paragraph pageId="7" pageNumber="36">A paratype is listed below; see other specimens examined.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="36" type="derivation of scientific epithet">
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<paragraph pageId="7" pageNumber="36">Derivation of scientific epithet.</paragraph>
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<paragraph pageId="7" pageNumber="36">
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The epithet
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<normalizedToken originalValue="“patagonia”">"patagonia"</normalizedToken>
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is a singular feminine noun used in apposition and refers to the Patagonian Mountain range in Arizona where the type specimen was collected. The area was part of the Apache homeland before being settled by those interested in mining the wealth of minerals nearby.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="37" type="proposed english vernacular name">
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<paragraph pageId="8" pageNumber="37">
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<pageBreakToken pageId="8" pageNumber="37" start="start">Proposed</pageBreakToken>
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English vernacular name.
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</paragraph>
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<paragraph pageId="8" pageNumber="37">Patagonia False-form beetle.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="37" type="diagnosis">
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<paragraph pageId="8" pageNumber="37">Diagnosis.</paragraph>
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<paragraph pageId="8" pageNumber="37">Color tone of dorsum alutaceous with head, pronotum and elytral suture paler; body rectangulate, lateral margins of elytra parallel, slightly tapering to an apically truncated and laterally slightly rounded apex; dorsum mostly glabrous with irregularly and wide-spaced short erect setae; pronotum with lateral margins broadly explanate, wider than elytra across humeri, disc markedly convex and medially planar; elytral interneurs minimally impressed yet easily visible under low magnification, 10 umbilicate setae present near lateral margin, dorsal edge of epipleuron lined with long laterally erect setae.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="37" type="description">
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<paragraph pageId="8" pageNumber="37">Description.</paragraph>
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<paragraph pageId="8" pageNumber="37">
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(
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="8" pageNumber="37">Figs 2</figureCitation>
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,
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="8" pageNumber="37">8</figureCitation>
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,
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="8" pageNumber="37">14</figureCitation>
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).
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Tables with measurements and ratios for Pseudomorpha patagonia sp. n.; all measurements are in millimeters." httpUri="http://table.plazi.org/id/0943E647FA7ECEEBD983A41073106F6F" pageId="8" pageNumber="37" tableUuid="0943E647FA7ECEEBD983A41073106F6F">Table 2</tableCitation>
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.
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<emphasis italics="true" pageId="8" pageNumber="37">Size</emphasis>
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: Medium for genus, ABL = 4.9 to 5.2 mm, SBL = 4.9 to 5.1 mm, TW = 4.0 to 4.2 mm. Preocular lobe-eye ratio: 0.3 to 0.39. Pronotum ratio (L/W): 0.29. Elytron ratio (L/W): 1.7.
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<emphasis italics="true" pageId="8" pageNumber="37">Color</emphasis>
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: Dorsum alutaceous with head, pronotum and elytral suture paler.
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<emphasis italics="true" pageId="8" pageNumber="37">Luster</emphasis>
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: Dorsum dull, slightly matte.
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<emphasis italics="true" pageId="8" pageNumber="37">Microsculpture</emphasis>
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: Small isodiametric sculpticells throughout dorsal surface.
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</paragraph>
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<paragraph pageId="8" pageNumber="37">
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<emphasis bold="true" pageId="8" pageNumber="37">Head:</emphasis>
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Genal lobe obsolete, rim posteriad and below eye bearing at least five robust setae directed perpendicular to head; preocular lobe distinct and slightly arching (
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="8" pageNumber="37">Fig. 8</figureCitation>
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); eye exceeding preocular lobe/gena boundary, shallowly arcuate; clypeus fused to frons with pigmented furrow entire and visible, bisetose, setae laterad on margin; labrum with four setae projecting anteriorly (
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="8" pageNumber="37">Fig. 7</figureCitation>
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); antennal flagellum markedly setose, antennomeres 1-3 bisetose.
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</paragraph>
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<paragraph pageId="8" pageNumber="37">
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<emphasis bold="true" pageId="8" pageNumber="37">Prothorax:</emphasis>
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Pronotum (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="8" pageNumber="37">Fig. 2</figureCitation>
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) mostly glabrous with irregularly and wide-spaced short erect setae, apex slightly rounded medially and narrower than ocular boundary, disk markedly convex and medially planar, width coequal to or slightly wider than elytra across humeri, base and apex fringed with more or less evenly spaced setae, pigmented median line ending about
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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before basal margin, lateral margins of pronotum with wide explanate sides, anterior angle 77.82°; prosternal apex fringed with short, evenly spaced setae.
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<emphasis bold="true" pageId="8" pageNumber="37">Pterothorax:</emphasis>
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Scutellum visible, small, triangulate with slightly rounded lateral margins; elytra smooth, interneurs very shallow, clearly visible under low magnification, markedly zig-zagged, intervals slightly convex on disc, lateral margin slightly sinuate at basal third or not, 10 umbilicate erect setae on the ventrally directed curvature of the elytral lateral portion (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="8" pageNumber="37">Fig. 2</figureCitation>
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).
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<emphasis bold="true" pageId="8" pageNumber="37">Abdomen:</emphasis>
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All sterna sparsely setiferous, sternum III densely so; male sternum IV with broad median dense row of posteriorly decumbent setae, sternum VII with two pair of two setae each along posterior edge; female with 2 pairs of 4 setae on sternum, and numerous longer setae on sterna IV, V, and VI.
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<emphasis bold="true" pageId="8" pageNumber="37">Legs:</emphasis>
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Legs flattened, setiferous, tibia bearing fringed ring of setae on distal end, femur with distinct lateral sulcus, femora and tibiae sparsely setose.
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<emphasis bold="true" pageId="8" pageNumber="37">Male Genitalia:</emphasis>
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(
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="8" pageNumber="37">Fig. 14</figureCitation>
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) Basal orifice hooded by phallobase, orifice recessed and small, phalloshaft arching, shaft narrows toward apex and slightly constricted at apical third; parameres co-equal in length, with the left paramere only slightly longer than right, both asetose; apical orifice small, about 1/5 the length of shaft.
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<emphasis bold="true" pageId="8" pageNumber="37">Female Genitalia:</emphasis>
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Not investigated.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="37" type="dispersal potential">
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<paragraph pageId="8" pageNumber="37">Dispersal potential.</paragraph>
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<paragraph pageId="8" pageNumber="37">These beetles are macropterous and have been recorded at lights, hence capable of flight; they are swift and agile runners. Accordingly, it is expected that this species be more broadly distributed across a wider geographical range than current records indicate.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="38" type="way of life">
|
||
<paragraph pageId="9" pageNumber="38">
|
||
<pageBreakToken pageId="9" pageNumber="38" start="start">Way</pageBreakToken>
|
||
of life.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="38">
|
||
Adults are likely found in ant nests and the surrounding vicinity; females are ovoviviparous (
|
||
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Journal of Natural History" pageId="16" pageNumber="45" pagination="1079 - 1086" publicationUrl="10.1080/00222938500770681" refId="B18" refString="Liebherr, JK, Kavanaugh, DH, 1985. Ovoviviparity in carabid beetles of the genus Pseudomorpha (Insecta: Coleoptera). Journal of Natural History 19: 1079 - 1086, 10.1080/00222938500770681" title="Ovoviviparity in carabid beetles of the genus Pseudomorpha (Insecta: Coleoptera)." url="10.1080/00222938500770681" volume="19" year="1985">Liebherr and Kavanaugh 1985</bibRefCitation>
|
||
); larvae are ant nest inquilines (
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="The Coleopterists Bulletin" pageId="16" pageNumber="45" pagination="53 - 68" refId="B8" refString="Erwin, TL, 1981. A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants. The Coleopterists Bulletin 35 (1): 53 - 68" title="A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants." volume="35" year="1981">Erwin 1981</bibRefCitation>
|
||
). Members of
|
||
<taxonomicName authorityName="Erwin & Amundson" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="patagonia">
|
||
<emphasis italics="true" pageId="9" pageNumber="38">Pseudomorpha patagonia</emphasis>
|
||
</taxonomicName>
|
||
occur at upland altitudes in between the Sonoran and Chihuahuan Deserts on oak dominated slopes. See: http://hikearizona.com/photo.php?ZIP=259631. Adults are active in July-August, very hot months in this area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="38" type="other specimens examined">
|
||
<paragraph pageId="9" pageNumber="38">Other specimens examined.</paragraph>
|
||
<paragraph pageId="9" pageNumber="38">
|
||
<emphasis bold="true" pageId="9" pageNumber="38">USA</emphasis>
|
||
, Arizona,
|
||
<emphasis italics="true" pageId="9" pageNumber="38">Cochise County</emphasis>
|
||
, Huachinera Mountains, Copper Canyon,
|
||
<geoCoordinate degrees="31.363" direction="north" orientation="latitude" precision="55" value="31.363">31.363°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="110.300" direction="west" orientation="longitude" precision="55" value="-110.3">110.300°W</geoCoordinate>
|
||
, 1882m, 16 July 1979 (S. McCleve) (UATC: ADP110736, male paratype).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="38" type="geographic distribution">
|
||
<paragraph pageId="9" pageNumber="38">Geographic distribution.</paragraph>
|
||
<paragraph pageId="9" pageNumber="38">
|
||
(
|
||
<figureCitation captionStart="Figure 18" captionStartId="F4" captionText="Figure 18. Distribution map for species of the santarita group of Pseudomorpha." httpUri="https://binary.pensoft.net/fig/27292" pageId="9" pageNumber="38">Fig. 18</figureCitation>
|
||
). This species is currently known from Arizona.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |