289 lines
37 KiB
XML
289 lines
37 KiB
XML
<document id="789B3030118C95304681195F78D1B8EE" ID-CLB-Dataset="51580" ID-DOI="10.11646/zootaxa.4092.3.2" ID-GBIF-Dataset="dd843b99-1701-4485-b235-1e344c0799db" ID-ISSN="1175-5326" ID-Zenodo-Dep="254902" ID-ZooBank="547625D1-70CC-48A1-9C36-3A0DD41B83A5" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458217128088" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Rahayu, Dwi Listyo" docDate="2016" docId="F56987A9FFBBFFE9FB8FF89EFF717AAC" docLanguage="en" docName="zootaxa.4092.3.2.pdf.imf" docOrigin="Zootaxa 4092 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gandoa Kammerer 2006" docType="treatment" docVersion="9" lastPageNumber="364" masterDocId="0950FFD1FFA3FFF0FB18FF91FFAE7D15" masterDocTitle="On the genera Selwynia Borradaile, 1903, and Gandoa Kammerer, 2006, with descriptions of two new species from Papua New Guinea and French Polynesia (Crustacea: Decapoda: Brachyura: Aphanodactylidae)" masterLastPageNumber="370" masterPageNumber="339" pageNumber="363" updateTime="1698187634807" updateUser="ExternalLinkService">
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<mods:titleInfo id="2672B912E8A1B6F3A2ED7C6D6B7AE7DC">
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<mods:title id="04D9273F0CCCD1F392EB569CEC9A9CEA">On the genera Selwynia Borradaile, 1903, and Gandoa Kammerer, 2006, with descriptions of two new species from Papua New Guinea and French Polynesia (Crustacea: Decapoda: Brachyura: Aphanodactylidae)</mods:title>
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<mods:namePart id="031B5F64217F7E757E47F33CA97D18AE">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="CD312FE628B1FA37DABD7716B358EC9E">Rahayu, Dwi Listyo</mods:namePart>
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<mods:typeOfResource id="069283D9247071616CC1FBBA72CF16D3">text</mods:typeOfResource>
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<mods:title id="A62B70EA4EAB9D30F0504EC4110D7E28">Zootaxa</mods:title>
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<mods:part id="B3F81C4D4F7A1A5B4108182B7F8B95C2">
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<mods:date id="D790B1FFF60D234DA91DD69E728F1F90">2016</mods:date>
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<mods:number id="BAB98F3B2EB5945E1553F42F5B5740E0">4092</mods:number>
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<mods:identifier id="2172DA07609AC8305C720116590D4B94" type="DOI">10.11646/zootaxa.4092.3.2</mods:identifier>
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<treatment id="F56987A9FFBBFFE9FB8FF89EFF717AAC" ID-DOI="http://doi.org/10.5281/zenodo.6081368" ID-GBIF-Taxon="117914237" ID-Zenodo-Dep="6081368" LSID="urn:lsid:plazi:treatment:F56987A9FFBBFFE9FB8FF89EFF717AAC" httpUri="http://treatment.plazi.org/id/F56987A9FFBBFFE9FB8FF89EFF717AAC" lastPageId="25" lastPageNumber="364" pageId="24" pageNumber="363">
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<subSubSection id="35DA6534FFBBFFE8FB8FF89EFDB97A9E" pageId="24" pageNumber="363" type="nomenclature">
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<paragraph id="7D7F36BFFFBBFFE8FB8FF89EFE617A3F" blockId="24.[151,463,1807,1834]" box="[151,463,1807,1834]" pageId="24" pageNumber="363">
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<heading id="263781D3FFBBFFE8FB8FF89EFE617A3F" bold="true" box="[151,463,1807,1834]" fontSize="11" level="1" pageId="24" pageNumber="363" reason="1">
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<taxonomicName id="BAC04D3CFFBBFFE8FB8FF89EFE617A3F" ID-CoL="7PGJH" authority="Kammerer, 2006" authorityName="Kammerer" authorityYear="2006" box="[151,463,1807,1834]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="363" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBBFFE8FB8FF89EFE617A3F" bold="true" box="[151,463,1807,1834]" pageId="24" pageNumber="363">
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<emphasis id="4FB4EAADFFBBFFE8FB8FF89EFF597A3C" bold="true" box="[151,247,1807,1833]" italics="true" pageId="24" pageNumber="363">Gandoa</emphasis>
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Kammerer, 2006
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="7D7F36BFFFBBFFE8FB8FF8C7FDB97A9E" blockId="24.[151,1436,1878,1992]" pageId="24" pageNumber="363">
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<taxonomicName id="BAC04D3CFFBBFFE8FB8FF8C7FE137A79" authority="Lenz, 1905: 364" authorityName="Lenz" authorityPageNumber="364" authorityYear="1905" box="[151,445,1878,1900]" class="Reptilia" family="Scincidae" genus="Voeltzkowia" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="363" phylum="Chordata" rank="genus">
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<emphasis id="4FB4EAADFFBBFFE8FB8FF8C7FEBE7A79" box="[151,272,1878,1900]" italics="true" pageId="24" pageNumber="363">Voeltzkowia</emphasis>
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Lenz, 1905: 364
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</taxonomicName>
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[
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<typeStatus id="A27B881DFFBBFFE8FAD2F8C6FE567A79" box="[458,504,1879,1900]" pageId="24" pageNumber="363">type</typeStatus>
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species:
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<taxonomicName id="BAC04D3CFFBBFFE8F94EF8C7FC747A79" authority="Lenz, 1905" authorityName="Lenz" authorityYear="1905" box="[598,986,1878,1900]" class="Reptilia" family="Scincidae" genus="Voeltzkowia" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="363" phylum="Chordata" rank="species" species="zanzibarensis">
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<emphasis id="4FB4EAADFFBBFFE8F94EF8C7FCF17A79" box="[598,863,1878,1900]" italics="true" pageId="24" pageNumber="363">Voeltzkowia zanzibarensis</emphasis>
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Lenz, 1905
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</taxonomicName>
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, by monotypy]; Schmitt
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<emphasis id="4FB4EAADFFBBFFE8FFC0F8C9FAA27A79" box="[1240,1292,1878,1901]" italics="true" pageId="24" pageNumber="363">et al.</emphasis>
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1973: 135 (in list); Ng
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<emphasis id="4FB4EAADFFBBFFE8FA39F8E7FEFA7A9E" box="[289,340,1909,1931]" italics="true" pageId="24" pageNumber="363">et al.</emphasis>
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2008: 247 (in list).
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</paragraph>
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</subSubSection>
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<subSubSection id="35DA6534FFBBFFE8FB8FF805FAB77ADD" pageId="24" pageNumber="363" type="description">
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<paragraph id="7D7F36BFFFBBFFE8FB8FF805FAB77ADD" blockId="24.[151,1436,1878,1992]" pageId="24" pageNumber="363">
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<taxonomicName id="BAC04D3CFFBBFFE8FB8FF805FE797ABF" authority="Kammerer, 2006: 270" authorityName="Kammerer" authorityPageNumber="270" authorityYear="2006" box="[151,471,1940,1962]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="363" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBBFFE8FB8FF805FF477ABF" box="[151,233,1940,1962]" italics="true" pageId="24" pageNumber="363">Gandoa</emphasis>
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Kammerer, 2006: 270
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</taxonomicName>
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[replacement name of
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<taxonomicName id="BAC04D3CFFBBFFE8F9D7F805FC677ABF" authority="Lenz, 1905" authorityName="Lenz" authorityYear="1905" box="[719,969,1940,1962]" class="Reptilia" family="Scincidae" genus="Voeltzkowia" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="363" phylum="Chordata" rank="genus">
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<emphasis id="4FB4EAADFFBBFFE8F9D7F805FCE67ABF" box="[719,840,1940,1962]" italics="true" pageId="24" pageNumber="363">Voeltzkowia</emphasis>
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Lenz, 1905
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</taxonomicName>
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, preoccupied by
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<taxonomicName id="BAC04D3CFFBBFFE8FF98F805FE837ADD" authority="Boettger, 1893" authorityName="Boettger" authorityYear="1893" class="Reptilia" family="Scincidae" genus="Voeltzkowia" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="363" phylum="Chordata" rank="genus">
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<emphasis id="4FB4EAADFFBBFFE8FF98F805FB577ABF" box="[1152,1273,1940,1962]" italics="true" pageId="24" pageNumber="363">Voeltzkowia</emphasis>
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Boettger, 1893 (Reptilia)
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</taxonomicName>
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]; Ng & Naruse 2009: 288; Ahyong & Ng 2009: 38; Ng & Clark 2014: 3; Castro 2015: 553 (in list).
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</paragraph>
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</subSubSection>
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<subSubSection id="35DA6534FFBAFFE9FB8FFF06FB797FE5" pageId="25" pageNumber="364" type="diagnosis">
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<paragraph id="7D7F36BFFFBAFFE9FB8FFF06FB797FE5" blockId="25.[151,1437,151,1977]" pageId="25" pageNumber="364">
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<emphasis id="4FB4EAADFFBAFFE9FB8FFF06FEA47DA5" bold="true" box="[151,266,151,176]" pageId="25" pageNumber="364">Diagnosis</emphasis>
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. Carapace wide, ovate, females ca. 1.4 times wider than long; males unknown; lateral margins subcristate; surface with numerous prominent, relatively deep pits. Front narrow, deflexed, broadly triangular in frontal view; straight to weakly sinuous in dorsal view. Orbit transverse; margins entire, unarmed; infraorbital margin mesially terminating in acute angle; supraorbital margin demarcated from antennular fossa by gently curving low ridge; lateral part of orbit distinctly extended laterally to meet the anterolateral margin; inner part of supraorbital margin forms distinct groove which extends to protogastric region. Eyes short, mobile. Antennules folding slightly obliquely. Epistome short, medially sunken. Third maxilliped ischium subquadrate; merus rounded. Thoracic sternites 1, 2 completely fused, broadly angular anteriorly; sternites 2/3 demarcated by shallow groove; sternites 3/4 fused, laterally unarmed; sternites 4–8 demarcated by narrow grooves, those demarcating sternites 4–7 incomplete medially; without longitudinal groove. Cheliped merus, carpus unarmed; outer surface of chela, notably at bases of fingers with dense pubescence. P2–P5 similar in shape, P3 longest, P5 shortest; merus about twice as long as high, as long as or slightly longer than respective carpi, propodi combined; with plumose setae on flexor, extensor margins; merus of P2–P4 usually unarmed or with 1 submedian tooth along flexor margin. P2–P5 without short spines on distoflexor angle of propodus opposing dactylus; dactylus very short, claw-like. Male abdomen simple, triangular; telson, all somites free; somite 1 widest; thoracic sternite 8 exposed when abdomen closed. Female abdomen with telson, all somites freely articulating, widest at somite 4. Vulva ovate, maximum width not exceeding half width of sternite 6. (Modified from Ahyong & Ng 2009: 38).
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</paragraph>
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</subSubSection>
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<subSubSection id="35DA6534FFBAFFE9FBDFFD6DFF717AAC" pageId="25" pageNumber="364" type="discussion">
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<paragraph id="7D7F36BFFFBAFFE9FBDFFD6DFD3979B5" blockId="25.[151,1437,151,1977]" pageId="25" pageNumber="364">
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<emphasis id="4FB4EAADFFBAFFE9FBDFFD6DFE9A7E00" bold="true" box="[199,308,764,789]" pageId="25" pageNumber="364">Remarks</emphasis>
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.
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<taxonomicName id="BAC04D3CFFBAFFE9FA5CFD6DFE317E01" box="[324,415,764,788]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9FA5CFD6DFE317E01" box="[324,415,764,788]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
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</taxonomicName>
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is superficially similar to
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<taxonomicName id="BAC04D3CFFBAFFE9F9CBFD6DFC967E01" box="[723,824,764,788]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9F9CBFD6DFC967E01" box="[723,824,764,788]" italics="true" pageId="25" pageNumber="364">Selwynia</emphasis>
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</taxonomicName>
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in carapace and ambulatory leg shape and structure; however, it can be immediately distinguished in having the lateral edges of the orbit extended laterally to meet the anterolateral margin (
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<figureCitation id="E5FB2A3AFFBAFFE9FA96FCD5FE437E49" box="[398,493,836,861]" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Figs. 17</figureCitation>
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D, 18B, 19B, 20D) and the inner part of the supraorbital margin forms a distinct groove which extends to the protogastric region of the carapace (
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<figureCitation id="E5FB2A3AFFBAFFE9F86FFCF6FC7A7E95" box="[887,980,871,896]" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Figs. 17</figureCitation>
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B, 18B, 19A, B, 20D). The merus of the third maxilliped of
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<taxonomicName id="BAC04D3CFFBAFFE9FA61FC1DFE7D7EB1" box="[377,467,908,932]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9FA61FC1DFE7D7EB1" box="[377,467,908,932]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
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</taxonomicName>
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(
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<figureCitation id="E5FB2A3AFFBAFFE9FAFEFC1DFDEA7EB1" box="[486,580,908,933]" captionStart="FIGURE 18" captionStartId="27.[151,250,1168,1190]" captionTargetBox="[353,1236,203,1138]" captionTargetId="figure@27.[340,1247,193,1147]" captionTargetPageId="27" captionText="FIGURE 18. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, right third maxilliped; B, right orbit and eye; C, right chela; D, right P 3; E, left P 4; F, left P 5. Scale: A, B = 0.5 mm; C – F = 1.0 mm." httpUri="https://zenodo.org/record/254920/files/figure.png" pageId="25" pageNumber="364">Figs. 18</figureCitation>
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A, 20C) is distinctly more rounded than any
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<taxonomicName id="BAC04D3CFFBAFFE9FF49FC1DFB187EB1" box="[1105,1206,908,932]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9FF49FC1DFB187EB1" box="[1105,1206,908,932]" italics="true" pageId="25" pageNumber="364">Selwynia</emphasis>
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</taxonomicName>
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species. The dorsal surface of the carapace and surfaces of the chelipeds of
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<taxonomicName id="BAC04D3CFFBAFFE9F822FC21FC3A7EDD" box="[826,916,944,968]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9F822FC21FC3A7EDD" box="[826,916,944,968]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
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</taxonomicName>
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are distinctly punctate (
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<figureCitation id="E5FB2A3AFFBAFFE9FFADFC3EFAB87EDD" box="[1205,1302,943,968]" captionStart="FIGURE 19" captionStartId="28.[151,250,1601,1623]" captionTargetBox="[151,1436,213,1554]" captionTargetId="figure@28.[151,1436,213,1555]" captionTargetPageId="28" captionText="FIGURE 19. Gandoa zanzibarensis (Lenz, 1905), female holotype (7.2 × 5.0 mm) (ZSM 1277 / 1), Zanzibar. A, overall habitus; B, frontal view of cephalothorax; C, ventral view of cephalothorax showing sternum and vulvae; D, outer view of left cheliped." httpUri="https://zenodo.org/record/254921/files/figure.png" pageId="25" pageNumber="364">Figs. 19</figureCitation>
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A, B, 20A) distinguishing it from
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<taxonomicName id="BAC04D3CFFBAFFE9FA85FC45FDAC7EF9" box="[413,514,980,1004]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9FA85FC45FDAC7EF9" box="[413,514,980,1004]" italics="true" pageId="25" pageNumber="364">Selwynia</emphasis>
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</taxonomicName>
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. Although the dorsal surface of the carapace of
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<taxonomicName id="BAC04D3CFFBAFFE9FF5AFC45FB097EF9" box="[1090,1191,980,1004]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="4FB4EAADFFBAFFE9FF5AFC45FB097EF9" box="[1090,1191,980,1004]" italics="true" pageId="25" pageNumber="364">Selwynia</emphasis>
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</taxonomicName>
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is also punctate, the depressions are relatively smaller and shallower. In addition, the outer surface of the chela of the female
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<typeStatus id="A27B881DFFBAFFE9FE2DFC66FA327905" box="[1333,1436,1015,1040]" pageId="25" pageNumber="364" type="lectotype">lectotype</typeStatus>
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of
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<taxonomicName id="BAC04D3CFFBAFFE9FBACFB8DFE9F7921" box="[180,305,1052,1076]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
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<emphasis id="4FB4EAADFFBAFFE9FBACFB8DFE9F7921" box="[180,305,1052,1076]" italics="true" pageId="25" pageNumber="364">G. brevipes</emphasis>
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</taxonomicName>
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and
|
||
<typeStatus id="A27B881DFFBAFFE9FA7DFB8DFE697920" box="[357,455,1052,1077]" pageId="25" pageNumber="364" type="holotype">holotype</typeStatus>
|
||
female of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F926FB8DFC987921" box="[574,822,1052,1076]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="zanzibarensis">
|
||
<emphasis id="4FB4EAADFFBAFFE9F926FB8DFC987921" box="[574,822,1052,1076]" italics="true" pageId="25" pageNumber="364">Gandoa zanzibarensis</emphasis>
|
||
</taxonomicName>
|
||
have dense pubescence at the base of the fingers (
|
||
<figureCitation id="E5FB2A3AFFBAFFE9FE7BFB8DFF1D794D" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Figs. 17</figureCitation>
|
||
D, E, 18C, 19D), which easily distinguishes it from
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F81FFBD1FCC2794D" box="[775,876,1088,1112]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FB4EAADFFBAFFE9F81FFBD1FCC2794D" box="[775,876,1088,1112]" italics="true" pageId="25" pageNumber="364">Selwynia</emphasis>
|
||
</taxonomicName>
|
||
. The distoflexor angle of the propodus of P2–P5 opposing the dactylus is also unarmed in
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F998FBF5FD747969" box="[640,730,1124,1148]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FB4EAADFFBAFFE9F998FBF5FD747969" box="[640,730,1124,1148]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E5FB2A3AFFBAFFE9F9F5FBF5FCEF7969" box="[749,833,1124,1149]" captionStart="FIGURE 18" captionStartId="27.[151,250,1168,1190]" captionTargetBox="[353,1236,203,1138]" captionTargetId="figure@27.[340,1247,193,1147]" captionTargetPageId="27" captionText="FIGURE 18. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, right third maxilliped; B, right orbit and eye; C, right chela; D, right P 3; E, left P 4; F, left P 5. Scale: A, B = 0.5 mm; C – F = 1.0 mm." httpUri="https://zenodo.org/record/254920/files/figure.png" pageId="25" pageNumber="364">Fig. 18</figureCitation>
|
||
E, F, 20E–G) but almost always has short spines in
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FB8FFB19FF5279B5" box="[151,252,1160,1184]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FB4EAADFFBAFFE9FB8FFB19FF5279B5" box="[151,252,1160,1184]" italics="true" pageId="25" pageNumber="364">Selwynia</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<figureCitation id="E5FB2A3AFFBAFFE9FA5BFB16FE3E79B5" box="[323,400,1159,1184]" captionStart="FIGURE 4" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[170,1419,204,1881]" captionTargetId="figure@8.[151,1436,193,1896]" captionTargetPageId="8" captionText="FIGURE 4. Selwynia sibogae (Tesch, 1918). A – E, male (7.7 × 5.3 mm) (ZRC 2010.0298), Lombok, Indonesia; F – K, ovigerous female lectotype (10.9 × 6.1 mm). A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, left P 3; H, left P 4; I, left P 5; J, right P 3; K, right P 4. Scales: A – E = 1.0 mm; F = 2.0 mm." httpUri="https://zenodo.org/record/254906/files/figure.png" pageId="25" pageNumber="364">Figs. 4</figureCitation>
|
||
J, 7C–H, 12C, 16C, D).
|
||
</paragraph>
|
||
<paragraph id="7D7F36BFFFBAFFE9FBDFFB3DFCEA7B61" blockId="25.[151,1437,151,1977]" pageId="25" pageNumber="364">
|
||
Examining photographs of the preserved
|
||
<typeStatus id="A27B881DFFBAFFE9F9BAFB3CFD7C79D0" box="[674,722,1197,1221]" pageId="25" pageNumber="364">type</typeStatus>
|
||
specimen, Ng & Naruse (2009: 288–289) transferred
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FE5DFB3CFDB279FD" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9FE5DFB3CFF5B79FD" italics="true" pageId="25" pageNumber="364">Pinnixa brevipes</emphasis>
|
||
H. Milne Edwards, 1853
|
||
</taxonomicName>
|
||
, to
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F950FB41FD0C79FD" box="[584,674,1232,1256]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FB4EAADFFBAFFE9F950FB41FD0C79FD" box="[584,674,1232,1256]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
|
||
</taxonomicName>
|
||
because its dorsal carapace surface was described as punctate (H. Milne Edwards, 1853: 220), and the carapace appeared to be more rectangular with the orbits more elongate. They also noted that the only aphanodactylid known from the Indian Ocean thus far was
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FF48FA89FB047825" box="[1104,1194,1304,1328]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FB4EAADFFBAFFE9FF48FA89FB047825" box="[1104,1194,1304,1328]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
|
||
</taxonomicName>
|
||
. Ng & Clark (2014), who found that
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FA5FFAADFD157841" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" box="[327,699,1340,1365]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="4FB4EAADFFBAFFE9FA5FFAADFE5B7841" box="[327,501,1340,1364]" italics="true" pageId="25" pageNumber="364">Selwynia laevis</emphasis>
|
||
Borradaile, 1903
|
||
</taxonomicName>
|
||
, from the
|
||
<collectingCountry id="05D7762FFFBAFFE9F835FAADFC3B7840" box="[813,917,1340,1365]" name="Maldives" pageId="25" pageNumber="364">Maldives</collectingCountry>
|
||
was congeneric with
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FF91FAADFE8E786D" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="sibogae">
|
||
<emphasis id="4FB4EAADFFBAFFE9FF91FAADFA327841" box="[1161,1436,1340,1364]" italics="true" pageId="25" pageNumber="364">Aphanodactylus sibogae</emphasis>
|
||
Tesch, 1918
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="05D7762FFFBAFFE9FA71FACEFE77786D" box="[361,473,1375,1400]" name="Indonesia" pageId="25" pageNumber="364">Indonesia</collectingCountry>
|
||
, followed this classification but noted that there was now a second genus known from the Indian Ocean and the generic status of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F962FA15FD597889" box="[634,759,1412,1436]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9F962FA15FD597889" box="[634,759,1412,1436]" italics="true" pageId="25" pageNumber="364">G. brevipes</emphasis>
|
||
</taxonomicName>
|
||
needs to be re-evaluated. The
|
||
<typeStatus id="A27B881DFFBAFFE9FF4AFA15FB177888" box="[1106,1209,1412,1437]" pageId="25" pageNumber="364" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FFC7FA14FA327889" box="[1247,1436,1412,1436]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9FFC7FA14FA327889" box="[1247,1436,1412,1436]" italics="true" pageId="25" pageNumber="364">Pinnixa brevipes</emphasis>
|
||
</taxonomicName>
|
||
is in poor condition (
|
||
<figureCitation id="E5FB2A3AFFBAFFE9FA66FA36FE6078D5" box="[382,462,1447,1472]" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Fig. 17</figureCitation>
|
||
A, D), and although the carapace is soft, the key characters are still visible. We can now confirm the observations of Ng & Naruse (2009) that it is a member of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F8ABFA5DFBA378F1" box="[947,1037,1484,1508]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FB4EAADFFBAFFE9F8ABFA5DFBA378F1" box="[947,1037,1484,1508]" italics="true" pageId="25" pageNumber="364">Gandoa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FF03FA5CFB7978F1" box="[1051,1239,1484,1508]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9FF03FA5CFB7978F1" box="[1051,1239,1484,1508]" italics="true" pageId="25" pageNumber="364">Pinnixa brevipes</emphasis>
|
||
</taxonomicName>
|
||
clearly possesses the post supraorbital-protogastric groove present on the anterior part of the carapace and the orbit clearly has the lateral extension, diagnostic features for the genus (cf.
|
||
<figureCitation id="E5FB2A3AFFBAFFE9F803F985FCD47B39" box="[795,890,1556,1581]" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Figs. 17</figureCitation>
|
||
B, D, 18B). In addition, the merus of the third maxilliped is also rounded in
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FAF9F9A8FDF97B45" box="[481,599,1592,1616]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="subSpecies" species="brevi" subSpecies="pes">
|
||
<emphasis id="4FB4EAADFFBAFFE9FAF9F9A8FDF97B45" box="[481,599,1592,1616]" italics="true" pageId="25" pageNumber="364">P. brevi pes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E5FB2A3AFFBAFFE9F971F9A6FD177B45" box="[617,697,1591,1616]" captionStart="FIGURE 18" captionStartId="27.[151,250,1168,1190]" captionTargetBox="[353,1236,203,1138]" captionTargetId="figure@27.[340,1247,193,1147]" captionTargetPageId="27" captionText="FIGURE 18. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, right third maxilliped; B, right orbit and eye; C, right chela; D, right P 3; E, left P 4; F, left P 5. Scale: A, B = 0.5 mm; C – F = 1.0 mm." httpUri="https://zenodo.org/record/254920/files/figure.png" pageId="25" pageNumber="364">Fig. 18</figureCitation>
|
||
A), like
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F80DF9A9FBA37B45" box="[789,1037,1592,1616]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="zanzibarensis">
|
||
<emphasis id="4FB4EAADFFBAFFE9F80DF9A9FBA37B45" box="[789,1037,1592,1616]" italics="true" pageId="25" pageNumber="364">Gandoa zanzibarensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E5FB2A3AFFBAFFE9FF04F9A6FBC17B45" box="[1052,1135,1591,1616]" captionStart="FIGURE 20" captionStartId="29.[151,250,1800,1822]" captionTargetBox="[165,1409,208,1766]" captionTargetId="figure@29.[150,1436,193,1779]" captionTargetPageId="29" captionText="FIGURE 20. Gandoa zanzibarensis (Lenz, 1905), female holotype (7.2 × 5.0 mm) (ZSM 1277 / 1), Zanzibar. A, overall habitus; B, frontal view of cephalothorax; C, left third maxilliped; D, right orbit and eye; E, right P 2; F, right P 3; G, left P 5; H, abdomen. Scales = 1.0 mm. A, B, G after Lenz, 1905: pl. 47, fig. 9." httpUri="https://zenodo.org/record/254922/files/figure.png" pageId="25" pageNumber="364">Fig. 20</figureCitation>
|
||
C); and both have the outer surface of the chela distinctly pubescent (
|
||
<figureCitation id="E5FB2A3AFFBAFFE9F97FF9CDFD6C7B61" box="[615,706,1628,1653]" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Figs. 17</figureCitation>
|
||
D, E, 19D).
|
||
</paragraph>
|
||
<paragraph id="7D7F36BFFFBAFFE9FBDFF910FF717AAC" blockId="25.[151,1437,151,1977]" pageId="25" pageNumber="364">
|
||
Ng & Naruse (2009: 289) provisionally recognized
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F828F911FC1C7B8D" box="[816,946,1664,1688]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9F828F911FC1C7B8D" box="[816,946,1664,1688]" italics="true" pageId="25" pageNumber="364">G. brevipes</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species, commenting that it appeared to differ from
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FAAEF935FDDF7BA9" box="[438,625,1700,1724]" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="zanzibarensis">
|
||
<emphasis id="4FB4EAADFFBAFFE9FAAEF935FDDF7BA9" box="[438,625,1700,1724]" italics="true" pageId="25" pageNumber="364">G. zanzibarensis</emphasis>
|
||
</taxonomicName>
|
||
in having the front less developed and the carapace proportionately broader. The examination of the
|
||
<typeStatus id="A27B881DFFBAFFE9F91FF956FDC37BF5" box="[519,621,1735,1760]" pageId="25" pageNumber="364" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F98CF959FCBF7BF5" box="[660,785,1736,1760]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9F98CF959FCBF7BF5" box="[660,785,1736,1760]" italics="true" pageId="25" pageNumber="364">G. brevipes</emphasis>
|
||
</taxonomicName>
|
||
shows that these differences are merely a consequence of the soft carapace, with the frontal part partially compressed and the posterior margin of the carapace damaged (
|
||
<figureCitation id="E5FB2A3AFFBAFFE9FE77F97DFF1D7A3D" captionStart="FIGURE 17" captionStartId="26.[151,250,1864,1886]" captionTargetBox="[151,1436,263,1842]" captionTargetId="figure@26.[151,1436,263,1842]" captionTargetPageId="26" captionText="FIGURE 17. Selwynia brevipes (H. Milne Edwards, 1853), ovigerous female lectotype (7.6 × 5.3 mm, carapace soft) (MNHN B- 10616), Mayotte. A, overall habitus; B, frontal regions of carapace; C, ventral view of cephalothorax; D, frontal view of cephalothorax; E, outer view of left chela." httpUri="https://zenodo.org/record/254919/files/figure.png" pageId="25" pageNumber="364">Fig. 17</figureCitation>
|
||
A, C, D). The revised measurements show both have the same carapace width to length proportions (ca. 1.4). There are no other obvious differences and we are confident
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F85BF8A4FAB07A58" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" box="[835,1310,1844,1869]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9F85BF8A4FC517A59" box="[835,1023,1844,1868]" italics="true" pageId="25" pageNumber="364">Pinnixa brevipes</emphasis>
|
||
H. Milne Edwards, 1853
|
||
</taxonomicName>
|
||
, is a senior subjective synonym of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FAB8F8C9FCFF7A65" authority="Lenz, 1905" authorityName="Lenz" authorityYear="1905" box="[416,849,1879,1904]" class="Reptilia" family="Scincidae" genus="Voeltzkowia" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="364" phylum="Chordata" rank="species" species="zanzibarensis">
|
||
<emphasis id="4FB4EAADFFBAFFE9FAB8F8C9FD6B7A65" box="[416,709,1880,1904]" italics="true" pageId="25" pageNumber="364">Voeltzkowia zanzibarensis</emphasis>
|
||
Lenz, 1905
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="A27B881DFFBAFFE9F889F8C9FC6F7A65" box="[913,961,1880,1904]" pageId="25" pageNumber="364">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="BAC04D3CFFBAFFE9FF5EF8C8FA947A65" authority="(Mayotte)" baseAuthorityName="Mayotte" box="[1094,1338,1880,1904]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9FF5EF8C8FB6C7A65" box="[1094,1218,1880,1904]" italics="true" pageId="25" pageNumber="364">P. brevipes</emphasis>
|
||
(
|
||
<collectingCountry id="05D7762FFFBAFFE9FFCAF8C9FA9D7A65" box="[1234,1331,1880,1904]" name="Mayotte" pageId="25" pageNumber="364">Mayotte</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
is about
|
||
<quantity id="BA389B5AFFBAFFE9FB8FF8EDFF577A80" box="[151,249,1916,1941]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="364" unit="km" value="1000.0">1000 km</quantity>
|
||
southeast of
|
||
<collectingCountry id="05D7762FFFBAFFE9FA94F8EDFE5C7A80" box="[396,498,1916,1941]" name="Tanzania" pageId="25" pageNumber="364">Zanzibar</collectingCountry>
|
||
. As such, the species should now be known as
|
||
<taxonomicName id="BAC04D3CFFBAFFE9F8E2F8EDFF757AAC" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" class="Malacostraca" family="Aphanodactylidae" genus="Gandoa" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="364" phylum="Arthropoda" rank="species" species="brevipes">
|
||
<emphasis id="4FB4EAADFFBAFFE9F8E2F8EDFB177A81" box="[1018,1209,1916,1940]" italics="true" pageId="25" pageNumber="364">Gandoa brevipes</emphasis>
|
||
(H. Milne Edwards, 1853)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |