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<document id="1011058EB5D3612D1B45D9528BC43335" ID-CLB-Dataset="27120" ID-DOI="10.11646/zootaxa.4582.1.1" ID-GBIF-Dataset="c79b6327-48fb-47bf-b17b-d20d2de295e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2643724" ID-ZooBank="9DC61794-4BD7-4F6D-BB8C-84D9855C8151" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555512297075" checkinUser="plazi" docAuthor="Mbo, Zingisile &amp; Haddad, Charles R." docDate="2019" docId="B74F87AB9E7E400053B3CC8F66A9982B" docLanguage="en" docName="zootaxa.4582.1.1.pdf.imf" docOrigin="Zootaxa 4582 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Drassodella tolkieni Mbo &amp; Haddad 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="34" masterDocId="4B76FFD39E6140215324C93E62409D53" masterDocTitle="A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="32" updateTime="1698721989978" updateUser="ExternalLinkService">
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<mods:title id="3DF975887FE0AC4CDE9BEE012A2BBC82">A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae)</mods:title>
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<mods:namePart id="EF49758815E882DA9F225D083786CEE2">Mbo, Zingisile</mods:namePart>
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<mods:namePart id="DD534F32241C676A8D1A65C0921BD0AC">Haddad, Charles R.</mods:namePart>
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<mods:title id="9D20C4F2CF392001C6FD96B50783AA35">Zootaxa</mods:title>
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<mods:date id="5095B91D61DD7B4FB6D6B1833DF4060E">2019</mods:date>
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<mods:number id="8B899EA3A392B37C3798F4F9F0E6C54B">2019-04-10</mods:number>
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<mods:number id="B1F305B069E66ED1031050EA86EE7A23">4582</mods:number>
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<mods:classification id="430239631E019668A60DDFF9EE9A490C">journal article</mods:classification>
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<mods:identifier id="5043093C745F63E8983DA3AF3F8AB2CA" type="DOI">10.11646/zootaxa.4582.1.1</mods:identifier>
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<treatment id="B74F87AB9E7E400053B3CC8F66A9982B" ID-GBIF-Taxon="156208929" LSID="urn:lsid:plazi:treatment:B74F87AB9E7E400053B3CC8F66A9982B" httpUri="http://treatment.plazi.org/id/B74F87AB9E7E400053B3CC8F66A9982B" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
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<paragraph id="3F5936BD9E7E403E53B3CC8F63AD9898" blockId="31.[151,878,1457,1517]" box="[151,493,1457,1483]" pageId="31" pageNumber="32">
<heading id="641181D19E7E403E53B3CC8F63AD9898" bold="true" box="[151,493,1457,1483]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="0D92EAAF9E7E403E53B3CC8F63AD9898" bold="true" box="[151,493,1457,1483]" pageId="31" pageNumber="32">
<taxonomicName id="F8E64D3E9E7E403E53B3CC8F63C89898" ID-CoL="37LL3" authority="Mbo &amp; Haddad, 2019" authorityName="Mbo &amp; Haddad" authorityYear="2019" box="[151,392,1457,1483]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
<emphasis id="0D92EAAF9E7E403E53B3CC8F63C89898" bold="true" box="[151,392,1457,1483]" italics="true" pageId="31" pageNumber="32">Drassodella tolkieni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16A157D49E7E403E52B4CC8F63AD9898" box="[400,493,1457,1483]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="77FC65369E7E403E53B3CCEA612E98BE" box="[151,878,1492,1517]" pageId="31" pageNumber="32" type="description">
<paragraph id="3F5936BD9E7E403E53B3CCEA612E98BE" blockId="31.[151,878,1457,1517]" box="[151,878,1492,1517]" pageId="31" pageNumber="32">
<figureCitation id="A7DD2A389E7E403E53B3CCEA62AC98BF" box="[151,236,1492,1517]" captionStart="FIGURES 18–26" captionStartId="7.[151,264,1551,1573]" captionTargetBox="[162,1424,517,1529]" captionTargetId="figure@7.[162,1425,517,1529]" captionTargetPageId="7" captionText="FIGURES 18–26. Scanning electron microscope photographs of Drassodella guttata sp. nov. female (18–21, 24–26) and male (23), and D. tolkieni sp. nov. female (22): 18 posterior half of carapace, dorsal view, indicating fovea, erect and feathery setae, and rebordered concave posterior margin; 19 eye region, dorsal view; 20 feathery setae on posterior carapace margin and coxa IV; 21 same, feathery setae enlarged; 22 chelicerae, ventral view; 23 chelicerae teeth; 24 endites and labium, ventral view; 25 serrula on endites, ventral view; 26 sternum. Abbreviations: CES—promarginal escort seta; ES—erect setae; FS—feathery setae; MHT—maxillar hair tuft." httpUri="https://zenodo.org/record/2643734/files/figure.png" pageId="31" pageNumber="32">Figs 22</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E53DCCCEA630098BF" box="[248,320,1492,1516]" captionStart="FIGURES 27–38" captionStartId="8.[151,264,1589,1611]" captionTargetBox="[158,1429,193,1568]" captionTargetId="figure@8.[158,1429,193,1568]" captionTargetPageId="8" captionText="FIGURES 27–38. Scanning electron microscope photographs of male (27, 28, 35–38) and female (29–31, 34) Drassodella guttata sp. nov. and female D. tolkieni sp. nov. (32, 33). 27 femur I, curved long ventral setae, lateral view; 28 femur IV, curved long ventral setae, lateral view; 29 leg II, patellar indentation, retrolateral view; 30 same, detail of lyriform organ; 31 lyriform organ at distal end of tibia II, retrolateral view; 32 metatarsus III with spine, feathery and chemosensory setae, and trichobothria, dorsal view; 33 distal end of metatarsus III, dorsal view; 34 metatarsus III, scopula, prolateral view; 35 distal end of tarsus I, dorsal view; 36 tarsus III, trichobothrium; 37 tarsus I, chemosensory seta; 38 same, tarsal organ. Abbreviations: CS—chemosensory setae; FS—feathery setae; MS—metatarsal stopper; SP—spine; SS—slit sensilla; TC—tarsal claws; TOtarsal organ; TR—trichobothria." httpUri="https://zenodo.org/record/2643736/files/figure.png" pageId="31" pageNumber="32">32, 33</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E5268CCEA63A998BF" box="[332,489,1492,1517]" captionStart="FIGURES 39–50" captionStartId="9.[151,264,1768,1790]" captionTargetBox="[161,1426,399,1747]" captionTargetId="figure@9.[161,1426,399,1747]" captionTargetPageId="9" captionText="FIGURES 39–50. Scanning electron microscope photographs of female (39, 41, 44, 46, 48–50) and male (43) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (40, 42, 45, 47): 39 tarsus III, scopula, ventrolateral view; 40 tarsus II, claws and scopulate setae, lateral view; 41 detail of scopulate setae, ventral view; 42 same, lateral view; 43 tarsus III, detail of tarsal claws and associated setae, arrow indicating tenant setae forming claw tuft; 44 dorsal abdominal feathery setae, arrow indicating white patch composed of dense modified feathery setae; 45 ventral abdominal feathery setae; 46, 47 spinnerets, general view; 48 left ALS, ventral view; 49 left PMS, ventral view; 50 left PLS, ventral view. Abbreviations as follows: Acaciniform gland spigot(s); Cy—cylindrical gland spigot(s); maAmp—major ampullate gland spigot(s); miAmp—minor ampullate gland spigot(s); nu—nubbin(s); Pi—piriform gland spigot(s)." httpUri="https://zenodo.org/record/2643738/files/figure.png" pageId="31" pageNumber="32">40, 42, 45, 47</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E52D0CCEA607C98BE" box="[500,572,1492,1517]" captionStart="FIGURES 51–59" captionStartId="10.[151,264,1740,1762]" captionTargetBox="[157,1429,700,1719]" captionTargetId="figure@10.[157,1429,700,1719]" captionTargetPageId="10" captionText="FIGURES 51–59. Scanning electron microscope photographs of female (51, 52) and male (55–59) Drassodella guttata sp. nov., and female D. tolkieni sp. nov. (53, 54): 51 epigyne, slightly ventrolateral view; 52 detail of epigynal muscle sigilla; 53 epigyne, dorsal view; 54 detail of glandular pores on spermathecal head; 55 right palp, prolateral view; 56 same, distal end of palp and cymbial scopula; 57 left palp, ventral view; 58 detail of distal end of tegulum, right palp, prolateral view; 59 same, left palp, ventral view. Abbreviations: AH—anterior hood; CD—copulatory duct; CO—copulatory opening; CS—cymbial scopula; DP—dorsal embolic prong; EM—embolus; EMS—epigynal muscle sigilla; LH—lateral hood; MA—median apophysis; MSmedian septum; ST—spermathecae; VP—ventral embolic prong." httpUri="https://zenodo.org/record/2643740/files/figure.png" pageId="31" pageNumber="32">53, 54</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E516CCCEA602698BE" box="[584,614,1492,1517]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="31" pageNumber="32">67</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E5156CCEA60D098BF" box="[626,656,1492,1516]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="31" pageNumber="32">83</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E51B8CCEA614098BE" box="[668,768,1492,1517]" captionStart="FIGURES 93–110" captionStartId="17.[151,264,1792,1814]" captionTargetBox="[160,1427,195,1767]" captionTargetId="figure@17.[160,1427,195,1770]" captionTargetPageId="17" captionText="FIGURES 93–110. Dorsal habitus photographs of Drassodella species in the melana species group: 93, 94 female and male of D. flava sp. nov.; 95, 96 female and male of D. guttata sp. nov.; 97 female of D. lotzi sp. nov.; 98 female of D. maculata sp. nov.; 99, 100 female and male of D. melana Tucker, 1923; 101, 102 female and male of D. montana sp. nov.; 103, 104 female and male of D. tenebrosa Lawrence, 1938; 105, 106 female and male of D. tolkieni sp. nov.; 107, 108 female and male of D. transversa sp. nov.; 109, 110 female and male of D. venda sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643746/files/figure.png" pageId="31" pageNumber="32">105, 106</figureCitation>
,
<figureCitation id="A7DD2A389E7E403E5028CCEA612E98BE" box="[780,878,1492,1517]" captionStart="FIGURES 141–145" captionStartId="32.[151,264,1311,1333]" captionTargetBox="[169,1426,200,1290]" captionTargetId="figure@32.[161,1426,200,1290]" captionTargetPageId="32" captionText="FIGURES 141–145. Genitalic morphology of Drassodella tolkieni sp. nov.: 141 female epigyne, ventral view; 142 same, dorsal view; 143 male palp, prolateral view; 144 same, ventral view; 145 same, retrolateral view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643768/files/figure.png" pageId="31" pageNumber="32">141–145</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="77FC65369E7E403E53B3CF2267479BF0" pageId="31" pageNumber="32" type="etymology">
<paragraph id="3F5936BD9E7E403E53B3CF2267479BF0" blockId="31.[151,1436,1564,2023]" pageId="31" pageNumber="32">
<emphasis id="0D92EAAF9E7E403E53B3CF22635E9B66" bold="true" box="[151,286,1564,1589]" pageId="31" pageNumber="32">Etymology.</emphasis>
Named after John Ronald Reuel Tolkien, who was born in Bloemfontein,
<collectingRegion id="FD22F85F9E7E403E574CCF23669B9B67" box="[1128,1243,1564,1588]" country="South Africa" name="Free State" pageId="31" pageNumber="32">Free State</collectingRegion>
,
<collectingCountry id="47F1762D9E7E403E57CCCF2267389B66" box="[1256,1400,1564,1589]" name="South Africa" pageId="31" pageNumber="32">South Africa</collectingCountry>
on
<date id="4B58107D9E7E403E53B3CF7D630D9B08" box="[151,333,1599,1627]" pageId="31" pageNumber="32" value="1892-01-03">
3
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January 1892
</date>
and died on
<date id="4B58107D9E7E403E52F8CF7D60F89B08" box="[476,696,1599,1627]" pageId="31" pageNumber="32" value="1973-09-02">
2
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September 1973
</date>
. He is internationally recognised for his authorship of The Hobbit, The Lord of the Rings and The Silmarillion, amongst other works. His fictional “Middle Earth” is believed to have been inspired in part by the exceptional natural scenery of Hogsback, the
<typeStatus id="E05D881F9E7E403E50EACFB561BE9BF0" box="[974,1022,1675,1699]" pageId="31" pageNumber="32">type</typeStatus>
locality of this species.
</paragraph>
</subSubSection>
<subSubSection id="77FC65369E7E403E53E3CF9066F19A04" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="3F5936BD9E7E403E53E3CF9066F19A04" blockId="31.[151,1436,1564,2023]" pageId="31" pageNumber="32">
<emphasis id="0D92EAAF9E7E403E53E3CF9063019B94" bold="true" box="[199,321,1710,1735]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
Females of
<taxonomicName id="F8E64D3E9E7E403E52F7CF91600B9B94" authority="Mbo &amp; Haddad, 2019" authorityName="Mbo &amp; Haddad" authorityYear="2019" box="[467,587,1710,1735]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
<emphasis id="0D92EAAF9E7E403E52F7CF91600B9B94" box="[467,587,1710,1735]" italics="true" pageId="31" pageNumber="32">D. tolkieni</emphasis>
</taxonomicName>
<emphasis id="0D92EAAF9E7E403E5171CF9060F19B94" bold="true" box="[597,689,1710,1735]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="16A157D49E7E403E5171CF9060F19B94" box="[597,689,1710,1735]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from others in the
<taxonomicName id="F8E64D3E9E7E403E57ACCF9167409B94" authorityName="Tucker" authorityYear="1923" box="[1160,1280,1710,1735]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="melana">
<emphasis id="0D92EAAF9E7E403E57ACCF9167409B94" box="[1160,1280,1710,1735]" italics="true" pageId="31" pageNumber="32">D. melana</emphasis>
</taxonomicName>
group by the distinctly concave posterior margin of the anterior hood (
<figureCitation id="A7DD2A389E7E403E5033CFEC61289BB8" box="[791,872,1746,1771]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="31" pageNumber="32">Fig. 67</figureCitation>
) and the chevron markings in the posterior half of the abdomen (
<figureCitation id="A7DD2A389E7E403E521ECFC863DF9A5C" box="[314,415,1782,1807]" captionStart="FIGURES 93–110" captionStartId="17.[151,264,1792,1814]" captionTargetBox="[160,1427,195,1767]" captionTargetId="figure@17.[160,1427,195,1770]" captionTargetPageId="17" captionText="FIGURES 93–110. Dorsal habitus photographs of Drassodella species in the melana species group: 93, 94 female and male of D. flava sp. nov.; 95, 96 female and male of D. guttata sp. nov.; 97 female of D. lotzi sp. nov.; 98 female of D. maculata sp. nov.; 99, 100 female and male of D. melana Tucker, 1923; 101, 102 female and male of D. montana sp. nov.; 103, 104 female and male of D. tenebrosa Lawrence, 1938; 105, 106 female and male of D. tolkieni sp. nov.; 107, 108 female and male of D. transversa sp. nov.; 109, 110 female and male of D. venda sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643746/files/figure.png" pageId="31" pageNumber="32">Fig. 105</figureCitation>
). Males of
<taxonomicName id="F8E64D3E9E7E403E5107CFC960DB9A5C" authority="Mbo &amp; Haddad, 2019" authorityName="Mbo &amp; Haddad" authorityYear="2019" box="[547,667,1782,1807]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
<emphasis id="0D92EAAF9E7E403E5107CFC960DB9A5C" box="[547,667,1782,1807]" italics="true" pageId="31" pageNumber="32">D. tolkieni</emphasis>
</taxonomicName>
<emphasis id="0D92EAAF9E7E403E5180CFC861409A5C" bold="true" box="[676,768,1782,1807]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="16A157D49E7E403E5180CFC861409A5C" box="[676,768,1782,1807]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share with males of
<taxonomicName id="F8E64D3E9E7E403E50D7CFC9661B9A5C" authority="Mbo &amp; Haddad, 2019" authorityName="Mbo &amp; Haddad" authorityYear="2019" box="[1011,1115,1782,1807]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="venda" status="sp. nov.">
<emphasis id="0D92EAAF9E7E403E50D7CFC9661B9A5C" box="[1011,1115,1782,1807]" italics="true" pageId="31" pageNumber="32">D. venda</emphasis>
</taxonomicName>
<emphasis id="0D92EAAF9E7E403E5740CFC866FF9A5C" bold="true" box="[1124,1215,1782,1807]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="16A157D49E7E403E5740CFC866FF9A5C" box="[1124,1215,1782,1807]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the dorsal embolic prong that is larger than the ventral prong, but can be distinguished by the shorter RTA with a slightly curved tip, while the RTA is longer and triangular in
<taxonomicName id="F8E64D3E9E7E403E514CCE01608E9A04" authority="Mbo &amp; Haddad, 2019" authorityName="Mbo &amp; Haddad" authorityYear="2019" box="[616,718,1854,1879]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="venda" status="sp. nov.">
<emphasis id="0D92EAAF9E7E403E514CCE01608E9A04" box="[616,718,1854,1879]" italics="true" pageId="31" pageNumber="32">D. venda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16A157D49E7E403E51F0CE00616E9A05" box="[724,814,1854,1879]" pageId="31" pageNumber="32" rank="species">
<emphasis id="0D92EAAF9E7E403E51F0CE0061679A04" bold="true" box="[724,807,1854,1879]" pageId="31" pageNumber="32">sp. nov</emphasis>
.
</taxonomicNameLabel>
(compare
<figureCitation id="A7DD2A389E7E403E5081CE0066439A04" box="[933,1027,1854,1879]" captionStart="FIGURES 141–145" captionStartId="32.[151,264,1311,1333]" captionTargetBox="[169,1426,200,1290]" captionTargetId="figure@32.[161,1426,200,1290]" captionTargetPageId="32" captionText="FIGURES 141–145. Genitalic morphology of Drassodella tolkieni sp. nov.: 141 female epigyne, ventral view; 142 same, dorsal view; 143 male palp, prolateral view; 144 same, ventral view; 145 same, retrolateral view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643768/files/figure.png" pageId="31" pageNumber="32">Fig. 145</figureCitation>
with
<figureCitation id="A7DD2A389E7E403E5767CE0066E49A04" box="[1091,1188,1854,1879]" captionStart="FIGURES 151–155" captionStartId="36.[151,264,1287,1309]" captionTargetBox="[165,1413,205,1259]" captionTargetId="figure@36.[161,1426,193,1266]" captionTargetPageId="36" captionText="FIGURES 151–155. Genitalic morphology of Drassodella venda sp. nov.: 151 female epigyne, ventral view; 152 same, dorsal view; 153 male palp, prolateral view; 154 same, ventral view; 155 same, retrolateral view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643772/files/figure.png" pageId="31" pageNumber="32">Fig. 155</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="77FC65369E7E400053E3CE5C604A992E" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="description">
<paragraph id="3F5936BD9E7E403E53E3CE5C628F9AB4" blockId="31.[151,1436,1564,2023]" pageId="31" pageNumber="32">
<emphasis id="0D92EAAF9E7E403E53E3CE5C61D89A28" bold="true" box="[199,920,1890,1915]" pageId="31" pageNumber="32">
Description. Female (
<typeStatus id="E05D881F9E7E403E52ECCE5C60789A28" box="[456,568,1890,1915]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
, Hogsback,
<bibRefCitation id="5B774B4C9E7E403E51EFCE5C61099A28" author="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R." box="[715,841,1890,1915]" pageId="31" pageNumber="32" refId="ref44234" refString="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R. (2013) Spiders of the Savanna Biome. University of Venda, Thohoyandou, and Agricultural Research Council, Pretoria, 134 pp." type="book" year="2013">NCA 2013</bibRefCitation>
/4616).
</emphasis>
Measurements: CL 1.92, CW 2.24, AL 3.44, AW 2.24, TL 6.60 (4.90–6.60), FL 0.46, SL 1.64, SW 1.47,
<collectingCountry id="47F1762D9E7E403E5076CEB961399ACC" box="[850,889,1927,1951]" name="Switzerland" pageId="31" pageNumber="32">CH</collectingCountry>
0.12, AME-AME 0.07, AME-ALE 0.02, ALE- ALE 0.29, PME-PME 0.10, PME-PLE 0.09, PLE-PLE 0.48, PERW 0.60, MOQAW 0.23, MOQPW 0.44, MOQL 0.28.
</paragraph>
<caption id="6B9966359E41400153B3CC2167159807" httpUri="https://zenodo.org/record/2643768/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,264,1311,1333]" targetBox="[169,1426,200,1290]" targetPageId="32">
<paragraph id="3F5936BD9E41400153B3CC2167159807" blockId="32.[151,1436,1311,1364]" pageId="32" pageNumber="33">
<emphasis id="0D92EAAF9E41400153B3CC2163309866" bold="true" box="[151,368,1311,1333]" pageId="32" pageNumber="33">FIGURES 141–145.</emphasis>
Genitalic morphology of
<taxonomicName id="F8E64D3E9E41400151A7CC2161149866" authority="Mbo &amp; Haddad, 2019" authorityName="Mbo &amp; Haddad" authorityYear="2019" box="[643,852,1311,1333]" class="Arachnida" family="Gallieniellidae" genus="Drassodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tolkieni" status="sp. nov.">
<emphasis id="0D92EAAF9E41400151A7CC2161149866" box="[643,852,1311,1333]" italics="true" pageId="32" pageNumber="33">Drassodella tolkieni</emphasis>
</taxonomicName>
<emphasis id="0D92EAAF9E414001507ACC2161F29866" bold="true" box="[862,946,1311,1333]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="16A157D49E414001507ACC2161F29866" box="[862,946,1311,1333]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 141 female epigyne, ventral view; 142 same, dorsal view; 143 male palp, prolateral view; 144 same, ventral view; 145 same, retrolateral view. Scale bars = 0.25 mm.
</paragraph>
</caption>
<paragraph id="3F5936BD9E41400153E3CC9B663898B2" blockId="32.[151,1437,1445,2010]" pageId="32" pageNumber="33">Length of leg segments: I 3.00 + 1.60 + 2.64 + 2.40 + 1.58 = 11.22, II 2.60 + 1.54 + 2.18 + 2.16 + 1.44 = 9.90, III 2.52 + 1.38 + 1.76 + 2.24 + 1.18 = 9.08, IV 3.52 + 1.70 + 2.72 + 3.56 + 1.24 = 12.74.</paragraph>
<paragraph id="3F5936BD9E41400053E3CCD362929C4F" blockId="32.[151,1437,1445,2010]" lastBlockId="33.[151,1437,151,1400]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
General appearance as in
<figureCitation id="A7DD2A389E41400152C8CCD3600D9B55" box="[492,589,1517,1542]" captionStart="FIGURES 93–110" captionStartId="17.[151,264,1792,1814]" captionTargetBox="[160,1427,195,1767]" captionTargetId="figure@17.[160,1427,195,1770]" captionTargetPageId="17" captionText="FIGURES 93–110. Dorsal habitus photographs of Drassodella species in the melana species group: 93, 94 female and male of D. flava sp. nov.; 95, 96 female and male of D. guttata sp. nov.; 97 female of D. lotzi sp. nov.; 98 female of D. maculata sp. nov.; 99, 100 female and male of D. melana Tucker, 1923; 101, 102 female and male of D. montana sp. nov.; 103, 104 female and male of D. tenebrosa Lawrence, 1938; 105, 106 female and male of D. tolkieni sp. nov.; 107, 108 female and male of D. transversa sp. nov.; 109, 110 female and male of D. venda sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643746/files/figure.png" pageId="32" pageNumber="33">Fig. 105</figureCitation>
. Carapace orange-brown, with black mottling and scattered white feathery setae; upsilon marking indistinct. Clypeus height twice AME diameter; AME smaller than ALE; AME separated by distance slightly larger than their diameter, AME separated from ALE by distance equal to 0.33 AME diameter; PME equal to PLE; PME separated by distance equal to 1.66 their diameter, PME separated from PLE by distance equal to 1.50 PME diameter. Chelicerae brown; promargin with three teeth, median tooth largest, proximal tooth smallest; retromargin with two teeth, distal tooth smaller than proximal tooth. Endites brown, with mediolateral mottling, pale at maxillar hair tuft. Labium brown, as long as wide. Sternum brown, with dark brown mottling on surface. Legs with dark brown proximal segments, distal segments orange. Leg spination: femora: I do 2, II do 2, III pl 1 do 2 rl 2, IV do 3; patellae: spineless; tibiae: I &amp; II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 2 rl 2 plv 2 rlv 2 vt 2; metatarsi: I &amp; II spineless, III pl 4 rl 4 plv 2 rlv 2 vt 2, IV pl 4 rl 4 plv 2 rlv 2 vt 2; palpal spination: femora do 3; patellae do 1; tibia pl 3 do 3 rl 3; tarsi pl 1 do 2 rl 2. Abdomen dark brown; dorsum with white patch anteriorly, stretching to half abdomen length, with four white lateral markings, posterior half with faint dark grey chevrons, white marking on anal tubercle (
<figureCitation id="A7DD2A389E41400151DFCEA361209AE5" box="[763,864,1949,1974]" captionStart="FIGURES 93–110" captionStartId="17.[151,264,1792,1814]" captionTargetBox="[160,1427,195,1767]" captionTargetId="figure@17.[160,1427,195,1770]" captionTargetPageId="17" captionText="FIGURES 93–110. Dorsal habitus photographs of Drassodella species in the melana species group: 93, 94 female and male of D. flava sp. nov.; 95, 96 female and male of D. guttata sp. nov.; 97 female of D. lotzi sp. nov.; 98 female of D. maculata sp. nov.; 99, 100 female and male of D. melana Tucker, 1923; 101, 102 female and male of D. montana sp. nov.; 103, 104 female and male of D. tenebrosa Lawrence, 1938; 105, 106 female and male of D. tolkieni sp. nov.; 107, 108 female and male of D. transversa sp. nov.; 109, 110 female and male of D. venda sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643746/files/figure.png" pageId="32" pageNumber="33">Fig. 105</figureCitation>
); lateral sclerites above booklungs narrow; venter brown; lateral sclerites small, post-epigastric sclerites absent; double rows of tiny sclerites from booklungs to spinnerets. Spinnerets cream. Epigyne with deep anterior hood, anterior margin convex, posterior margin strongly concave, lateral ends tapering abruptly to sharp point; lateral hoods curved, close to copulatory openings; ST II yellow, connecting ducts and ST I yellow-brown; ST I and II similar in size, ST II elongate-oval (
<figureCitation id="A7DD2A389E4040005620C9E1671E9DAB" box="[1284,1374,223,248]" captionStart="FIGURES 60–77" captionStartId="12.[151,264,1529,1551]" captionTargetBox="[156,1422,224,1505]" captionTargetId="figure@12.[156,1422,223,1505]" captionTargetPageId="12" captionText="FIGURES 60–77. Female epigynes of Drassodella species in the D. melana species group (60–68) and D. salisburyi species group (69–77) in dorsal view: 60 D. flava sp. nov.; 61 D. guttata sp. nov.; 62 D. lotzi sp. nov.; 63 D. maculata sp. nov.; 64 D. melana Tucker, 1923; 65 D. montana sp. nov.; 66 D. tenebrosa Lawrence, 1938; 67 D. tolkieni sp. nov.; 68 D. transversa sp. nov.; 69 D. venda sp. nov.; 70 D. amatola sp. nov.; 71 D. aurostriata sp. nov.; 72 D. purcelli Tucker, 1923; 73 D. quinquelabecula Tucker, 1923; 74 D. salisburyi Hewitt, 1916; 75 D. septemmaculata (Strand, 1909); 76 D. trilineata sp. nov.; 77 D. vasivulva Tucker, 1923. Abbreviations: AH—anterior hood; FD—fertilisation duct; MS—median septum; STspermatheca(e). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643742/files/figure.png" pageId="33" pageNumber="34">Figs 67</figureCitation>
,
<figureCitation id="A7DD2A389E404000564FC9DE62859C4F" captionStart="FIGURES 141–145" captionStartId="32.[151,264,1311,1333]" captionTargetBox="[169,1426,200,1290]" captionTargetId="figure@32.[161,1426,200,1290]" captionTargetPageId="32" captionText="FIGURES 141–145. Genitalic morphology of Drassodella tolkieni sp. nov.: 141 female epigyne, ventral view; 142 same, dorsal view; 143 male palp, prolateral view; 144 same, ventral view; 145 same, retrolateral view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643768/files/figure.png" pageId="33" pageNumber="34">141, 142</figureCitation>
).
</paragraph>
<paragraph id="3F5936BD9E40400053E3C819674D9CDB" blockId="33.[151,1437,151,1400]" pageId="33" pageNumber="34">
<emphasis id="0D92EAAF9E40400053E3C81960AA9C13" bold="true" box="[199,746,295,320]" pageId="33" pageNumber="34">
Male (
<typeStatus id="E05D881F9E4040005231C81963C59C13" box="[277,389,295,320]" pageId="33" pageNumber="34" type="paratype">Paratype</typeStatus>
, Hogsback,
<bibRefCitation id="5B774B4C9E404000513FC81960DB9C13" author="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R." box="[539,667,295,320]" pageId="33" pageNumber="34" refId="ref44234" refString="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R. (2013) Spiders of the Savanna Biome. University of Venda, Thohoyandou, and Agricultural Research Council, Pretoria, 134 pp." type="book" year="2013">NCA 2013</bibRefCitation>
/4541).
</emphasis>
Measurements: CL 1.78, CW 1.40, AL 1.96, AW 1.08, TL 3.84 (3.80–5.24), FL 0.21, SL 1.00, SW 0.80,
<collectingCountry id="47F1762D9E40400051BAC87260859C37" box="[670,709,332,356]" name="Switzerland" pageId="33" pageNumber="34">CH</collectingCountry>
0.10, AME-AME 0.05, AME-ALE 0.01, ALE-ALE 0.17, PME- PME 0.06, PME-PLE 0.08, PLE-PLE 0.33, PERW 0.38, MOQAW 0.14, MOQPW 0.16, MOQL 0.14.
</paragraph>
<paragraph id="3F5936BD9E40400053E3C8AA662A9C83" blockId="33.[151,1437,151,1400]" pageId="33" pageNumber="34">Length of leg segments: I 1.28 + 0.62 + 0.98 + 0.96 + 0.96 = 4.80, II 1.18 + 0.56 + 0.88 + 0.98 + 0.72 = 4.32, III 1.04 + 0.48 + 0.76 + 1.04 + 0.66 = 3.98, IV 1.68 + 0.64 + 1.24 + 1.76 + 0.82 = 6.14.</paragraph>
<paragraph id="3F5936BD9E40400053E3C8E2604A992E" blockId="33.[151,1437,151,1400]" pageId="33" pageNumber="34">
General appearance as in
<figureCitation id="A7DD2A389E40400052DEC8E2601F9CA6" box="[506,607,476,501]" captionStart="FIGURES 93–110" captionStartId="17.[151,264,1792,1814]" captionTargetBox="[160,1427,195,1767]" captionTargetId="figure@17.[160,1427,195,1770]" captionTargetPageId="17" captionText="FIGURES 93–110. Dorsal habitus photographs of Drassodella species in the melana species group: 93, 94 female and male of D. flava sp. nov.; 95, 96 female and male of D. guttata sp. nov.; 97 female of D. lotzi sp. nov.; 98 female of D. maculata sp. nov.; 99, 100 female and male of D. melana Tucker, 1923; 101, 102 female and male of D. montana sp. nov.; 103, 104 female and male of D. tenebrosa Lawrence, 1938; 105, 106 female and male of D. tolkieni sp. nov.; 107, 108 female and male of D. transversa sp. nov.; 109, 110 female and male of D. venda sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643746/files/figure.png" pageId="33" pageNumber="34">Fig. 106</figureCitation>
. Carapace uniformly dark brown, with radiating black mottling; white feathery setae scattered; upsilon marking indistinct but thick white line composed of feathery setae diverging behind fovea to posterior margin. Clypeus height equal to 1.43 times AME diameter; AME smaller than ALE; AME separated by distance slightly smaller than their diameter, AME almost touching ALE; PME equal to PLE; PME separated by distance equal to 1.16 their diameter, PME separated from PLE by distance equal to 1.33 PME diameter. Chelicerae brown; promargin with three teeth, median tooth largest, proximal tooth smallest; retromargin with two teeth, distal tooth smaller than proximal tooth. Endites brown, pale at maxillar hair tuft. Labium brown, slightly longer than wide. Sternum brown. Legs brown to dark-brown. Leg spination: femora: I pl 1 do 2, II do 2, III do 3, IV pl 1 do 2 rl 1; patellae: spineless; tibiae: I &amp; II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 2 rl 2 plv 2 rlv 2 vt 2; metatarsi: I &amp; II spineless, III pl 4 do 1 rl 4 plv 2 rlv 2 vt 2, IV pl 4 rl 4 plv 2 rlv 2 vt 2; palpal spination: femora do 3; patellae do 1; tibiae spineless; tarsi do 1. Abdomen dark-brown, markings similar to female, but with less dense white feathery setae (
<figureCitation id="A7DD2A389E40400051F2CA59617B9ED3" box="[726,827,871,896]" captionStart="FIGURES 93–110" captionStartId="17.[151,264,1792,1814]" captionTargetBox="[160,1427,195,1767]" captionTargetId="figure@17.[160,1427,195,1770]" captionTargetPageId="17" captionText="FIGURES 93–110. Dorsal habitus photographs of Drassodella species in the melana species group: 93, 94 female and male of D. flava sp. nov.; 95, 96 female and male of D. guttata sp. nov.; 97 female of D. lotzi sp. nov.; 98 female of D. maculata sp. nov.; 99, 100 female and male of D. melana Tucker, 1923; 101, 102 female and male of D. montana sp. nov.; 103, 104 female and male of D. tenebrosa Lawrence, 1938; 105, 106 female and male of D. tolkieni sp. nov.; 107, 108 female and male of D. transversa sp. nov.; 109, 110 female and male of D. venda sp. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2643746/files/figure.png" pageId="33" pageNumber="34">Fig. 106</figureCitation>
); dorsal scutum and anterior marginal sigilla absent; venter creamy-yellow; laterally with two large distinct oval sclerites; two pairs of tiny sclerites running from epigastric furrow to spinnerets; small ante- and post-epigastric sclerites present. Spinnerets cream. Palp yellowbrown; RTA short, protruding retrolaterally, not obscured in ventral view by retrolateral heel of cymbium, tip slightly curved; in retrolateral view subtriangular, with broad base and slightly curved tip; cymbium broad, 1.60 times longer than wide, distal end broadly rounded; ventral and dorsal embolic prongs spike-like, dorsal prong larger in prolateral view; dorsal prong distinctly bent away from tegulum in ventral view; median apophysis stout, hook-shaped (
<figureCitation id="A7DD2A389E4040005211CD5A63CC992F" box="[309,396,1124,1149]" captionStart="FIGURES 78–92" captionStartId="16.[151,264,1460,1482]" captionTargetBox="[157,1428,190,1448]" captionTargetId="figure@16.[157,1429,190,1448]" captionTargetPageId="16" captionText="FIGURES 78–92. Male palps of Drassodella species of the D. melana species group (78–85) and D. salisburyi species group (86–92) in ventral view: 78 D. flava sp. nov.; 79 D. guttata sp. nov.; 80 D. melana Tucker, 1923; 81 D. montana sp. nov.; 82 D. tenebrosa Lawrence, 1923; 83 D. tolkieni sp. nov.; 84 D. transversa sp. nov.; 85 D. venda sp. nov.; 86 D. amatola sp. nov.; 87 D. aurostriata sp. nov.; 88 D. baviaans sp. nov.; 89 D. quinquelabecula Tucker, 1923; 90 D. salisburyi Hewitt, 1916; 91 D. septemmaculata (Strand, 1909); 92 D. vasivulva Tucker, 1923. Abbreviations as follows: DP—dorsal embolic prong; EMembolus; MA—median apophysis; RTA—retrolateral tibial apophysis; VP—ventral embolic prong. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643744/files/figure.png" pageId="33" pageNumber="34">Figs 83</figureCitation>
,
<figureCitation id="A7DD2A389E40400052B3CD5A63BE992E" box="[407,510,1124,1149]" captionStart="FIGURES 141–145" captionStartId="32.[151,264,1311,1333]" captionTargetBox="[169,1426,200,1290]" captionTargetId="figure@32.[161,1426,200,1290]" captionTargetPageId="32" captionText="FIGURES 141–145. Genitalic morphology of Drassodella tolkieni sp. nov.: 141 female epigyne, ventral view; 142 same, dorsal view; 143 male palp, prolateral view; 144 same, ventral view; 145 same, retrolateral view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2643768/files/figure.png" pageId="33" pageNumber="34">143–145</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="77FC65369E40400053E3CDB9670A9806" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="3F5936BD9E40400053E3CDB9609599BB" blockId="33.[151,1437,151,1400]" pageId="33" pageNumber="34">
<emphasis id="0D92EAAF9E40400053E3CDB9633999F3" bold="true" box="[199,377,1159,1184]" pageId="33" pageNumber="34">Type material.</emphasis>
<materialsCitation id="8F8E3CE09E40400052A0CDB9609599BB" ID-GBIF-Occurrence="2236308158" collectingDate="2013-03-25" collectingDateMax="2013-03-30" collectingDateMin="2013-03-25" collectionCode="NCA" collectorName="C. Haddad &amp; J. Neethling" country="South Africa" county="Amatola Mountains" elevation="1210" latitude="-32.62785" location="Amatola Forestry Company" longLatPrecision="1" longitude="26.97185" municipality="Hogsback" pageId="33" pageNumber="34" specimenCode="NCA 2016" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape" typeStatus="holotype">
<typeStatus id="E05D881F9E40400052A0CDB963AC99F3" box="[388,492,1159,1184]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
<specimenCount id="29E0FD349E40400052D3CDB6604E99CC" box="[503,526,1160,1183]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
:
<emphasis id="0D92EAAF9E4040005104CDB9614399F3" bold="true" box="[544,771,1159,1184]" pageId="33" pageNumber="34">
<collectingCountry id="47F1762D9E4040005104CDB960BE99F3" box="[544,766,1159,1184]" name="South Africa" pageId="33" pageNumber="34">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="0D92EAAF9E404000502BCDB7665C99F3" box="[783,1052,1160,1184]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="FD22F85F9E404000502BCDB7665C99F3" box="[783,1052,1160,1184]" country="South Africa" name="Eastern Cape" pageId="33" pageNumber="34">Eastern Cape Province</collectingRegion>
</emphasis>
:
<collectingCounty id="D6384E319E404000570BCDB9675699F3" box="[1071,1302,1159,1184]" pageId="33" pageNumber="34">Amatola Mountains</collectingCounty>
,
<collectingMunicipality id="DF3DACC79E4040005601CDB967D899F3" box="[1317,1432,1159,1184]" pageId="33" pageNumber="34">Hogsback</collectingMunicipality>
,
<location id="3A3960669E40400053B3CD92639F9997" LSID="urn:lsid:plazi:treatment:B74F87AB9E7E400053B3CC8F66A9982B:3A3960669E40400053B3CD92639F9997" box="[151,479,1196,1221]" country="South Africa" county="Amatola Mountains" latitude="-32.62785" longLatPrecision="1" longitude="26.97185" municipality="Hogsback" name="Amatola Forestry Company" pageId="33" pageNumber="34" stateProvince="Eastern Cape">Amatola Forestry Company</location>
,
<geoCoordinate id="5AD2507A9E40400052D6CD92603D9996" box="[498,637,1196,1221]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="-32.62785">32°37.671'S</geoCoordinate>
,
<geoCoordinate id="5AD2507A9E40400051B4CD92615C9996" box="[656,796,1196,1221]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="26.97185">26°58.311'E</geoCoordinate>
,
<elevation id="B4CBD18E9E404000500BCD92618F9996" box="[815,975,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="33" pageNumber="34" unit="m" value="1210.0">
<quantity id="F81E9B589E404000500BCD9261CC9997" box="[815,908,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="33" pageNumber="34" unit="m" value="1210.0">1210 m</quantity>
a.s.l.
</elevation>
,
<date id="4B58107D9E40400050C7CD9266CE9996" box="[995,1166,1196,1221]" pageId="33" pageNumber="34" value="2013-03-25" valueMax="2013-03-30" valueMin="2013-03-25">
<collectingDate id="5B1CE9959E40400050C7CD9266CE9996" box="[995,1166,1196,1221]" pageId="33" pageNumber="34" value="2013-03-25" valueMax="2013-03-30" valueMin="2013-03-25">25–30.III.2013</collectingDate>
</date>
, leg.
<collectorName id="9213536B9E40400057FCCD9267189996" box="[1240,1368,1196,1221]" pageId="33" pageNumber="34">C. Haddad</collectorName>
&amp;
<collectorName id="9213536B9E40400056AECD93634599BB" pageId="33" pageNumber="34">J. Neethling</collectorName>
(
<collectingMethod id="E6A74EAA9E4040005230CDF163D899B4" box="[276,408,1231,1256]" pageId="33" pageNumber="34">pitfall traps</collectingMethod>
, fynbos) (
<specimenCode id="6F409EC69E404000512CCDEE60C699BB" box="[520,646,1231,1256]" collectionCode="NCA" pageId="33" pageNumber="34">NCA 2016</specimenCode>
/2819).
</materialsCitation>
</paragraph>
<paragraph id="3F5936BD9E40400053E3CDCB670A9806" blockId="33.[151,1437,151,1400]" pageId="33" pageNumber="34">
<typeStatus id="E05D881F9E40400053E3CDCB6379985E" box="[199,313,1269,1293]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<emphasis id="0D92EAAF9E4040005267CDCA6063985E" bold="true" box="[323,547,1268,1293]" pageId="33" pageNumber="34">
<collectingCountry id="47F1762D9E4040005267CDCA605D985E" box="[323,541,1268,1293]" name="South Africa" pageId="33" pageNumber="34">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="0D92EAAF9E404000510ECDCB616F985F" box="[554,815,1269,1292]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="FD22F85F9E404000510ECDCB616F985F" box="[554,815,1269,1292]" country="South Africa" name="Eastern Cape" pageId="33" pageNumber="34">Eastern Cape Province</collectingRegion>
</emphasis>
: Same data as
<typeStatus id="E05D881F9E40400050FCCDCA667D985E" box="[984,1085,1268,1293]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
,
<specimenCount id="29E0FD349E404000576DCDCA662C985E" box="[1097,1132,1268,1293]" pageId="33" pageNumber="34" type="male">5♂</specimenCount>
(NCA 2016/2823),
<specimenCount id="29E0FD349E404000566BCDCA6732985E" box="[1359,1394,1268,1293]" pageId="33" pageNumber="34" type="male">6♂</specimenCount>
<specimenCount id="29E0FD349E404000565DCDCA67DC985E" box="[1401,1436,1268,1293]" pageId="33" pageNumber="34" type="female">5♀</specimenCount>
(
<bibRefCitation id="5B774B4C9E40400053BACC2663259863" author="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R." box="[158,357,1303,1328]" pageId="33" pageNumber="34" refId="ref44234" refString="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R. (2013) Spiders of the Savanna Biome. University of Venda, Thohoyandou, and Agricultural Research Council, Pretoria, 134 pp." type="book" year="2013">NCA 2013/4541</bibRefCitation>
); Amatola Mountains, Hogsback, Amatola Forestry Company, Robinson Dam,
<geoCoordinate id="5AD2507A9E4040005628CC2667D89863" box="[1292,1432,1304,1328]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="-32.621315">32°37.279'S</geoCoordinate>
,
<geoCoordinate id="5AD2507A9E40400053B3CC0263649806" box="[151,292,1340,1365]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="26.969933">26°58.196'E</geoCoordinate>
,
<quantity id="F81E9B589E404000520BCC0263C39807" box="[303,387,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="33" pageNumber="34" unit="m" value="1190.0">1190 m</quantity>
a.s.l., leg. C. Haddad &amp; J. Neethling (pitfall traps, fynbos),
<specimenCount id="29E0FD349E4040005705CC0266049807" box="[1057,1092,1340,1364]" pageId="33" pageNumber="34" type="male">4♂</specimenCount>
<specimenCount id="29E0FD349E404000576FCC02662E9807" box="[1099,1134,1340,1364]" pageId="33" pageNumber="34" type="female">1♀</specimenCount>
(
<bibRefCitation id="5B774B4C9E4040005759CC02677E9806" author="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R." box="[1149,1342,1340,1365]" pageId="33" pageNumber="34" refId="ref44234" refString="Dippenaar-Schoeman, A. S. &amp; Foord, S. &amp; Haddad, C. R. (2013) Spiders of the Savanna Biome. University of Venda, Thohoyandou, and Agricultural Research Council, Pretoria, 134 pp." type="book" year="2013">NCA 2013/4616</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="77FC65369E40400053E3CC6166A9982B" box="[199,1257,1375,1400]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="3F5936BD9E40400053E3CC6166A9982B" blockId="33.[151,1437,151,1400]" box="[199,1257,1375,1400]" pageId="33" pageNumber="34">
<emphasis id="0D92EAAF9E40400053E3CC616320982B" bold="true" box="[199,352,1375,1400]" pageId="33" pageNumber="34">Distribution.</emphasis>
Known only from the
<typeStatus id="E05D881F9E4040005144CC5E60D0982B" box="[608,656,1376,1400]" pageId="33" pageNumber="34">type</typeStatus>
locality in the
<collectingRegion id="FD22F85F9E404000501CCC5E618F982B" box="[824,975,1376,1400]" country="South Africa" name="Eastern Cape" pageId="33" pageNumber="34">Eastern Cape</collectingRegion>
,
<collectingCountry id="47F1762D9E40400050FFCC61662A982B" box="[987,1130,1375,1400]" name="South Africa" pageId="33" pageNumber="34">South Africa</collectingCountry>
(
<figureCitation id="A7DD2A389E404000575ECC61669C982B" box="[1146,1244,1375,1400]" captionStart="FIGURE 156" captionStartId="37.[151,250,1578,1600]" captionTargetBox="[161,1426,591,1556]" captionTargetId="figure@37.[161,1426,591,1556]" captionTargetPageId="37" captionText="FIGURE 156. Distribution of Drassodella tolkieni sp. nov. (circle), D. transversa sp. nov. (stars) and D. venda sp. nov. (squares) in southern Africa." httpUri="https://zenodo.org/record/2643774/files/figure.png" pageId="33" pageNumber="34">Fig. 156</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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