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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.161.53384" ID-GBIF-Dataset="8121676d-d9dc-4a3d-8325-b821656da6a9" ID-PMC="PMC7508921" ID-Pensoft-Pub="1314-2003-161-27" ID-Pensoft-UUID="35A869D2676E5D0686AFAC667D799E7B" ID-PubMed="33005088" ModsDocID="1314-2003-161-27" checkinTime="1600207728933" checkinUser="pensoft" docAuthor="Tomasello, Salvatore &amp; Konowalik, Kamil" docDate="2020" docId="B74B4568D4D25C3FB19D446C390B7445" docLanguage="en" docName="PhytoKeys 161: 27-40" docOrigin="PhytoKeys 161" docSource="http://dx.doi.org/10.3897/phytokeys.161.53384" docTitle="Leucanthemopsis alpina subsp. vranicae Tomasello &amp; Konowalik 2020, subsp. nov." docType="treatment" docVersion="4" id="35A869D2676E5D0686AFAC667D799E7B" lastPageNumber="27" masterDocId="35A869D2676E5D0686AFAC667D799E7B" masterDocTitle="On the Leucanthemopsis alpina (L.) Heywood growing in the Illyrian region" masterLastPageNumber="40" masterPageNumber="27" pageNumber="27" updateTime="1668140045980" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>On the Leucanthemopsis alpina (L.) Heywood growing in the Illyrian region</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tomasello, Salvatore</mods:namePart>
<mods:affiliation>Department of Systematics, Biodiversity and Evolution of Plants (with Herbarium), Albrecht-von-Haller Institute for Plant Sciences, Untere Karspuele 2, 37073, Georg-August University, Goettingen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5201-1156</mods:nameIdentifier>
<mods:nameIdentifier type="email">salvatore.tomasello@uni-goettingen.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Konowalik, Kamil</mods:namePart>
<mods:affiliation>Institute of Biology, Wroclaw University of Environmental and Life Sciences, Kozuchowska 5 b, Wroclaw, Poland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4821-0608</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>161</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>40</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.161.53384</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.161.53384</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-161-27</mods:identifier>
<mods:identifier type="Pensoft-UUID">35A869D2676E5D0686AFAC667D799E7B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="167613832" LSID="urn:lsid:plazi:treatment:B74B4568D4D25C3FB19D446C390B7445" httpUri="http://treatment.plazi.org/id/B74B4568D4D25C3FB19D446C390B7445" lastPageNumber="27" pageId="0" pageNumber="27">
<subSubSection pageId="0" pageNumber="27" type="nomenclature">
<paragraph pageId="0" pageNumber="27">
<taxonomicName LSID="B74B4568-D4D2-5C3F-B19D-446C390B7445" authority="Tomasello &amp; Konowalik" authorityName="Tomasello &amp; Konowalik" authorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucanthemopsis alpina subsp. vranicae" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" status="subsp. nov." subSpecies="vranicae">Leucanthemopsis alpina subsp. vranicae Tomasello &amp; Konowalik</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="27">subsp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="description">
<paragraph pageId="0" pageNumber="27">Description.</paragraph>
<paragraph pageId="0" pageNumber="27">Perennial plant, subscapose hemicryptophyte ~ 8-10 cm. Basal leaves pinnatilobate 20-30 mm long and 4-9 mm wide, spatulate with ovate lamina, moderately pilose abaxially, leaflets 5 (-7), 1.5 times longer than width of leaf rachis. Cauline bracts 3 or 4, linear, 3-5 mm long. Inflorescences scapiform, capitula solitary. Involucral bracts green with black margins, pilose over midrib, margin ciliate. Corolla of ray florets white throughout anthesis. Diploid.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Tomasello &amp; Konowalik" authorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucanthemopsis alpina subsp. vranicae" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="vranicae">Leucanthemopsis alpina subsp. vranicae</taxonomicName>
differs from subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. cuneifolia" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="cuneifolia">cuneifolia</taxonomicName>
in possessing abaxially pilose leaves (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A Herbarium voucher (WRSL: WR GN 064834) of the type material from the Leucanthemopsis alpina subsp. vranicae, collected in 2018 by K. Konowalik in the Vranica Mts. B Detailed pictures of the involucral bract C abaxial face of a basal leaf." figureDoi="10.3897/phytokeys.161.53384.figure3" httpUri="https://binary.pensoft.net/fig/452673" pageId="0" pageNumber="27">3C</figureCitation>
). In
<taxonomicName lsidName="L. alpina subsp. cuneifolia" pageId="0" pageNumber="27" rank="subSpecies" species="alpina" subSpecies="cuneifolia">L. alpina subsp. cuneifolia</taxonomicName>
, leaves are glabrous on both surfaces (a few diploid populations from the area surrounding the Dolomites possess abaxially pilose leaves, but are usually smaller plants [up to ca. 5 cm] than those in the Vranica Mts). In
<taxonomicName lsidName="L. alpina subsp. vranicae" pageId="0" pageNumber="27" rank="subSpecies" species="alpina" subSpecies="vranicae">L. alpina subsp. vranicae</taxonomicName>
, the green midribs of the involucral bracts are pilose (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A Herbarium voucher (WRSL: WR GN 064834) of the type material from the Leucanthemopsis alpina subsp. vranicae, collected in 2018 by K. Konowalik in the Vranica Mts. B Detailed pictures of the involucral bract C abaxial face of a basal leaf." figureDoi="10.3897/phytokeys.161.53384.figure3" httpUri="https://binary.pensoft.net/fig/452673" pageId="0" pageNumber="27">3B</figureCitation>
), but are glabrous in subsp.
<taxonomicName lsidName="subsp. cuneifolia" pageId="0" pageNumber="27" rank="subSpecies" species="alpina" subSpecies="cuneifolia">cuneifolia</taxonomicName>
.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucanthemopsis alpina subsp. tatrae" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="tatrae">Leucanthemopsis alpina subsp. tatrae</taxonomicName>
leaves are less incised (leaflets up to 1.5 times longer than the width of the leaf rachis), which are tomentose on both surfaces (according to
<bibRefCitation author="Vierhapper, F" journalOrPublisher="Magyar Botanikai Lapok" pageId="0" pageNumber="27" pagination="13 - 35" refId="B32" refString="Vierhapper, F, 1914. Chrysanthemum alpinum forma tatrae. Magyar Botanikai Lapok 13: 13 - 35" title="Chrysanthemum alpinum forma tatrae." volume="13" year="1914">Vierhapper 1914</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="distribution">
<paragraph pageId="0" pageNumber="27">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="27">
Krstac Mt and between Devetaci Mt and Nadkrstac Mt. (Vranica Mts). Bosnia and Herzegovina. Grows solitary or forms clumps on siliceous substrates (gravel and stones) in xerophytic alpine meadows above
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus mugo" order="Pinales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="mugo">
<emphasis italics="true" pageId="0" pageNumber="27">Pinus mugo</emphasis>
</taxonomicName>
shrubs. 2000-2100 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="materials_examined">
<paragraph pageId="0" pageNumber="27">Type.</paragraph>
<paragraph pageId="0" pageNumber="27">
Bosnia and Herzegovina, Central Bosnia Canton, between municipalities of Gornji Vakuf-Uskoplje and Fojnica, close to the village Prokos at the
<normalizedToken originalValue="Prokoško">Prokosko</normalizedToken>
jezero lake, Vranica Planina Mountains, Krstac Mt;
<geoCoordinate degrees="43.954833" direction="north" orientation="latitude" precision="1" value="43.954834">43.954833N</geoCoordinate>
,
<geoCoordinate degrees="17.736917" direction="east" orientation="longitude" precision="1" value="17.736917">17.736917E</geoCoordinate>
; alt. 2036 m; 7 Aug 2018;
<emphasis italics="true" pageId="0" pageNumber="27">K. Konowalik 20180807-02</emphasis>
; (holotype: WRSL [WR GN 064834]; paratypes: WU [WU0043817], SARA [SARA43310, SARA43311, SARA43312]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="conservation status">
<paragraph pageId="0" pageNumber="27">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Tomasello &amp; Konowalik" authorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucanthemopsis alpina subsp. vranicae" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="vranicae">Leucanthemopsis alpina subsp. vranicae</taxonomicName>
is a point endemic with a very small range (less than 2.5 km2) which meets the IUCN Red List criteria for critically endangered species (CR B2a, IUCN Standards and Petition Committe 2019). Though its population size is not known, it is expected to be low (i.e. during the survey in 2018, only ca. 25 flowering plants were seen in both accessible stands). Therefore, it would also meet criterion D (very small and restricted population, IUCN Standards and Petition Committe 2019). As the subspecies is confined to an alpine habitat, which is heavily restricted in the Vranica Mts, it may face extinction in case of any stochastic event or effects of anthropogenic climate warming. Though, a long time span of collections (130 years) and certain features of those mountains (e.g. microhabitats, inaccessible slopes) may indicate that it will manage to persist there.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="identification key to subspecies of leucanthemopsis alpina (modified from tomasello and oberprieler 2017)">
<paragraph pageId="0" pageNumber="27">
Identification key to subspecies of
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucanthemopsis alpina" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis italics="true" pageId="0" pageNumber="27">Leucanthemopsis alpina</emphasis>
</taxonomicName>
(modified from
<bibRefCitation DOI="https://doi.org/10.1093/botlinnean/bow009" author="Tomasello, S" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="27" pagination="211 - 235" refId="B31" refString="Tomasello, S, Oberprieler, C, 2017. Frozen ploidies - A phylogeographic analysis of the Leucanthemopsis alpina polyploid complex (Compositae, Anthemideae). Botanical Journal of the Linnean Society 183 (2): 211 - 235, DOI: https://doi.org/10.1093/botlinnean/bow009" title="Frozen ploidies - A phylogeographic analysis of the Leucanthemopsis alpina polyploid complex (Compositae, Anthemideae)." url="https://doi.org/10.1093/botlinnean/bow009" volume="183" year="2017">Tomasello and Oberprieler 2017</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="27">
<table pageId="0" pageNumber="27">
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">Dwarf, caespitose and subscapose perennial; leaves palmatifid, villose-tomentose, ca. 10 mm long; leaf segments not spaced; scape short, with 0 (or 1) linear cauline bract; corolla of ray florets reddish during anthesis</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. tomentosa" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="tomentosa">tomentosa</taxonomicName>
(Loisel.) Heywood
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">Plants without a dwarf habit; leaves pinnatisect or pinnatilobate; scape usually with 1 or more cauline bracts</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaves pinnatisect, segments&gt; 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as width of leaf rachis
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaves pinnatilobate, segments as long or &lt;2
<normalizedToken originalValue="×">x</normalizedToken>
as long as width of leaf
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">Plants sericeous-tomentose; corolla of ray florets becoming reddish during anthesis</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. pseudotomentosa" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="pseudotomentosa">pseudotomentosa</taxonomicName>
(Fiori) Tomasello &amp; Oberpr.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">Plants glabrous or moderately pilose, never sericeous-tomentose; corolla of ray florets white or reddish</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaflets narrow, 3.0-4.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, 2.5-4.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of leaf rachis; corolla of ray florets sometimes becoming reddish after anthesis
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. pyrenaica" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="pyrenaica">pyrenaica</taxonomicName>
(Vierh.) Tomasello &amp; Oberpr.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaflets broader, up to 2.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of leaf rachis; corolla of ray florets always remaining white
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">Plants generally minute (5-10 cm), moderately pilose; leaves 15-20(-25) mm long with 5 to 7 segments; scape with 2 to 4 cauline bracts</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. minima" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="minima">minima</taxonomicName>
(Vill.) Holub
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">Plants larger (8-15 cm), glabrous; leaves 20-30 mm long, usually with 7 to 9 segments; scape with 1 to 3 cauline bracts</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. alpina" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="alpina">alpina</taxonomicName>
(L.) Heywood
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaves pubescent on both surfaces; leaflets 1.0-1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of leaf rachis
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. tatrae" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="tatrae">tatrae</taxonomicName>
(Vierh.) Holub
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaves glabrous or pilose abaxially; leaflets 1.5-2.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of leaf rachis
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaves mostly glabrous; leaflets 1.5-2.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than rachis width; green midribs of involucral bracts glabrous
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. cuneifolia" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="cuneifolia">cuneifolia</taxonomicName>
(Murr) Tomasello &amp; Oberpr.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="27">
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
Leaves moderately pilose abaxially; leaflets 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of leaf rachis; green midribs of involucral bracts pilose
</td>
<td colspan="1" pageId="0" pageNumber="27" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="27">
subsp.
<taxonomicName authorityName="Tomasello &amp; Konowalik" authorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. vranicae" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="alpina" subSpecies="vranicae">vranicae</taxonomicName>
Tomasello &amp; Konowalik
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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