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<mods:title id="28FA13E65727DDFF58F2AFD6152909D8">Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph</mods:title>
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<mods:namePart id="FAAAFBB2A6FB0E76A56E831C2EEA2BBC">Nakayama, Naohide</mods:namePart>
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<treatment id="B711B23FFF3486F1DA3FC4C3FCEE7A5F" ID-DOI="http://doi.org/10.5281/zenodo.6455200" ID-GBIF-Taxon="194588037" ID-Zenodo-Dep="6455200" LSID="urn:lsid:plazi:treatment:B711B23FFF3486F1DA3FC4C3FCEE7A5F" httpUri="http://treatment.plazi.org/id/B711B23FFF3486F1DA3FC4C3FCEE7A5F" lastPageId="227" lastPageNumber="228" pageId="226" pageNumber="227">
<subSubSection id="77A250A2FF3486F0DA3FC4C3FAEA7A1D" pageId="226" pageNumber="227" type="nomenclature">
<paragraph id="3F070329FF3486F0DA3FC4C3FB027A30" blockId="226.[810,1293,1771,1832]" box="[810,1253,1771,1798]" pageId="226" pageNumber="227">
<heading id="644FB445FF3486F0DA3FC4C3FB027A30" box="[810,1253,1771,1798]" fontSize="11" level="2" pageId="226" pageNumber="227" reason="2">
<emphasis id="0DCCDF3BFF3486F0DA3FC4C3FB027A30" bold="true" box="[810,1253,1771,1798]" pageId="226" pageNumber="227">
Genus
<taxonomicName id="F8B878AAFF3486F0DA94C4C3FB027A30" ID-CoL="59FH" authority="Iwamoto, 1974" authorityName="Iwamoto" authorityYear="1974" box="[897,1253,1771,1798]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="226" pageNumber="227" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF3486F0DA94C4C3FBC17A30" bold="true" box="[897,1062,1771,1797]" italics="true" pageId="226" pageNumber="227">Kuronezumia</emphasis>
Iwamoto, 1974
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="3F070329FF3486F0DA3FC526FAEA7A1D" blockId="226.[810,1293,1771,1832]" box="[810,1293,1805,1832]" pageId="226" pageNumber="227">[Japanese name: Suruga-nezumidara-zoku]</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF3486F0DA3FC57EFC3A7A9B" pageId="226" pageNumber="227" type="description">
<paragraph id="3F070329FF3486F0DA3FC57EFC3A7A9B" blockId="226.[810,1452,1877,1966]" pageId="226" pageNumber="227">
<taxonomicName id="F8B878AAFF3486F0DA3FC57EFB767A5E" authority="Iwamoto, 1974: 509" authorityName="Iwamoto" authorityPageNumber="509" authorityYear="1974" box="[810,1169,1877,1899]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="226" pageNumber="227" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF3486F0DA3FC57EFC4D7A5E" box="[810,938,1878,1899]" italics="true" pageId="226" pageNumber="227">Kuronezumia</emphasis>
Iwamoto, 1974:509
</taxonomicName>
[
<typeStatus id="E003BD8BFF3486F0DDBAC57EFB3C7A5E" box="[1199,1243,1878,1899]" pageId="226" pageNumber="227">type</typeStatus>
species:
<taxonomicName id="F8B878AAFF3486F0DC42C57EFAF47AB9" authority="Iwamoto, 1974" authorityName="Iwamoto" authorityYear="1974" class="Actinopterygii" family="Macrouridae" genus="Nezumia" kingdom="Animalia" order="Gadiformes" pageId="226" pageNumber="227" phylum="Chordata" rank="species" species="bubonis" subGenus="Kuronezumia">
<emphasis id="0DCCDF3BFF3486F0DC42C57EFBB37AB9" italics="true" pageId="226" pageNumber="227">Nezumia (Kuronezumia) bubonis</emphasis>
Iwamoto, 1974
</taxonomicName>
, by original designation].
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF3486F1DA3FC5F1FE827E81" lastPageId="227" lastPageNumber="228" pageId="226" pageNumber="227" type="diagnosis">
<paragraph id="3F070329FF3486F1DA3FC5F1FE827E81" blockId="226.[810,1451,2009,2070]" lastBlockId="227.[136,778,173,1458]" lastPageId="227" lastPageNumber="228" pageId="226" pageNumber="227">
<emphasis id="0DCCDF3BFF3486F0DA3FC5F1FC417AC6" bold="true" box="[810,934,2009,2035]" pageId="226" pageNumber="227">Diagnosis.</emphasis>
Anus separated from anal-fin origin, closer to pelvic-fin bases; periproct teardrop-shaped, moderately developed. Small dermal window of light organ between pelvic-fin bases. Infraorbital ridge not connected with preopercular ridge, separated by distinct gap. Second spinous ray of first dorsal fin finely serrated along its leading edge. Pelvic fin inserted below or anterior to pectoral-fin base. Mouth moderately large, usually subterminal. Chin barbel well developed. Teeth small, in broad bands in both jaws; premaxillary band ending anterior to posterior edge of maxillary process. Tip and lateral angles of snout lacking prominent scutes in most species. Suborbital region almost vertical, without coarsely modified scales. Head fully scaled, except for narrow naked area above upper lip. Body scales not deciduous, closely adhered to body surfaces; scale spinules needle-like, densely scattered over exposed portion or tightly packed subparallel to convergent rows; buttresses of spinules rudimentary; reticulate structure developed on posterior parts of unexposed portion. Grooved lateral line present. Cephalic sensory pores present or absent. Branchiostegal rays 7. Body dark to swarthy, lacking silvery reflection when fresh. [Modified from
<bibRefCitation id="5B297ED8FF3586F1D829C15FFD947EA4" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[316,627,887,914]" pageId="227" pageNumber="228" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto(1992)</bibRefCitation>
and
<bibRefCitation id="5B297ED8FF3586F1DBB1C150FEBC7E81" author="Iwamoto, T. &amp; Graham, K. J." pageId="227" pageNumber="228" pagination="407 - 509" refId="ref288409" refString="Iwamoto, T. &amp; Graham, K. J. (2001) Grenadiers (families Bathygadidae and Macrouridae, Gadiformes, Pisces) of New South Wales, Australia. Proceedings of the California Academy of Sciences, 52, 407 - 509." type="journal article" year="2001">Iwamoto &amp; Graham (2001)</bibRefCitation>
.]
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF3586F1D9A8C193FDB47884" pageId="227" pageNumber="228" type="discussion">
<paragraph id="3F070329FF3586F1D9A8C193FDB47884" blockId="227.[136,778,173,1458]" pageId="227" pageNumber="228">
<emphasis id="0DCCDF3BFF3586F1D9A8C193FED67EE0" bold="true" box="[189,305,955,981]" pageId="227" pageNumber="228">Remarks.</emphasis>
<taxonomicName id="F8B878AAFF3586F1D828C194FE347EE0" authorityName="Iwamoto" authorityYear="1974" box="[317,467,956,981]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF3586F1D828C194FE347EE0" box="[317,467,956,981]" italics="true" pageId="227" pageNumber="228">Kuronezumia</emphasis>
</taxonomicName>
was originally established as a subgenus of
<taxonomicName id="F8B878AAFF3586F1D849C1F6FE597EC2" authorityName="Jordan in Jordan &amp; Starks" authorityYear="1904" box="[348,446,990,1015]" class="Actinopterygii" family="Macrouridae" genus="Nezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF3586F1D849C1F6FE597EC2" box="[348,446,990,1015]" italics="true" pageId="227" pageNumber="228">Nezumia</emphasis>
</taxonomicName>
<collectingCountry id="47AF43B9FF3586F1D8DEC1F5FDF17EC2" box="[459,534,989,1015]" name="Jordan" pageId="227" pageNumber="228">Jordan</collectingCountry>
<emphasis id="0DCCDF3BFF3586F1DB37C1F6FDDF7EC2" box="[546,568,990,1015]" italics="true" pageId="227" pageNumber="228">in</emphasis>
<bibRefCitation id="5B297ED8FF3586F1DB51C1F5FF24792C" author="Jordan, D. S. &amp; Starks, E. C." pageId="227" pageNumber="228" pagination="577 - 630" refId="ref290133" refString="Jordan, D. S. &amp; Starks, E. C. (1904) List of fishes dredged by the steamer Albatross off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the U. S. Fish Commission, 22, 577 - 630, pls. 1 - 8." type="journal article" year="1904">
<collectingCountry id="47AF43B9FF3586F1DB51C1F5FD687EC2" box="[580,655,989,1015]" name="Jordan" pageId="227" pageNumber="228">Jordan</collectingCountry>
&amp; Starks, 1904
</bibRefCitation>
, but
<bibRefCitation id="5B297ED8FF3586F1D9EDC1D7FDD6792C" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[248,561,1023,1050]" pageId="227" pageNumber="228" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto (1992)</bibRefCitation>
elevated this taxon to full generic rank.
<bibRefCitation id="5B297ED8FF3586F1D87AC609FD79790E" author="Shcherbachev, Y. N. &amp; Sazonov, Y. I. &amp; Iwamoto, T." box="[367,670,1057,1083]" pageId="227" pageNumber="228" pagination="97 - 108" refId="ref299815" refString="Shcherbachev, Y. N., Sazonov, Y. I. &amp; Iwamoto, T. (1992) Synopsis of the grenadier genus Kuronezumia (Pisces: Gadiformes: Macrouridae), with description of a new species. Proceedings of the California Academy of Sciences, 48, 97 - 108." type="journal article" year="1992">
Shcherbachev
<emphasis id="0DCCDF3BFF3586F1DB06C60AFDAB790E" box="[531,588,1057,1083]" italics="true" pageId="227" pageNumber="228">et al.</emphasis>
(1992)
</bibRefCitation>
provided a synopsis for all the then-known species (seven species), viz.,
<taxonomicName id="F8B878AAFF3586F1D9D6C64EFE11794A" authority="Iwamoto, 1974" authorityName="Iwamoto" authorityYear="1974" box="[195,502,1125,1152]" class="Actinopterygii" family="Macrouridae" genus="Nezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="bubonis">
<emphasis id="0DCCDF3BFF3586F1D9D6C64EFEA7794A" box="[195,320,1125,1151]" italics="true" pageId="227" pageNumber="228">K. bubonis</emphasis>
Iwamoto, 1974
</taxonomicName>
(
<typeStatus id="E003BD8BFF3586F1DB1DC64DFDDD794A" box="[520,570,1125,1151]" pageId="227" pageNumber="228">type</typeStatus>
species);
<taxonomicName id="F8B878AAFF3586F1DBA4C64EFE4E7997" authority="(Gilbert &amp; Hubbs, 1916)" baseAuthorityName="Gilbert &amp; Hubbs" baseAuthorityYear="1916" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="dara">
<emphasis id="0DCCDF3BFF3586F1DBA4C64EFCED794A" box="[689,778,1125,1151]" italics="true" pageId="227" pageNumber="228">K. dara</emphasis>
(
<bibRefCitation id="5B297ED8FF3586F1D985C6AFFE467997" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[144,417,1159,1186]" pageId="227" pageNumber="228" pagination="135 - 214" refId="ref285931" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1916) Report on the Japanese macrourid fishes collected by the United States fisheries steamer &quot; Albatross &quot; in 1906, with a synopsis of the genera. Proceedings of the United States National Museum, 51, 135 - 214, pls. 8 - 11. http: // dx. doi. org / 10.5479 / si. 00963801.51 - 2149.135" type="journal article" year="1916">Gilbert &amp; Hubbs, 1916</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="F8B878AAFF3586F1D8ADC6A0FD067997" authority="(Barnard, 1925)" baseAuthorityName="Barnard" baseAuthorityYear="1925" box="[440,737,1159,1186]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="leonis">
<emphasis id="0DCCDF3BFF3586F1D8ADC6A0FDF97994" box="[440,542,1159,1185]" italics="true" pageId="227" pageNumber="228">K. leonis</emphasis>
(
<bibRefCitation id="5B297ED8FF3586F1DB25C6AFFD3E7997" author="Barnard, K. H." box="[560,729,1159,1186]" pageId="227" pageNumber="228" pagination="498 - 504" refId="ref282128" refString="Barnard, K. H. (1925) Descriptions of new species of marine fishes from S. Africa. Annals and Magazine of Natural History, Series 9, 15, 498 - 504. http: // dx. doi. org / 10.1080 / 00222932508633240" type="journal article" year="1925">Barnard, 1925</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="F8B878AAFF3586F1DBE5C6A0FEEC79F6" baseAuthorityName="Smith &amp; Radcliffe in Radcliffe" baseAuthorityYear="1912" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="macronema">
<emphasis id="0DCCDF3BFF3586F1DBE5C6A0FEEC79F6" italics="true" pageId="227" pageNumber="228">K. macronema</emphasis>
</taxonomicName>
(Smith &amp; Radcliffe
<emphasis id="0DCCDF3BFF3586F1DB14C682FDF079F6" box="[513,535,1194,1219]" italics="true" pageId="227" pageNumber="228">in</emphasis>
<bibRefCitation id="5B297ED8FF3586F1DB37C681FD3F79F6" author="Radcliffe, L." box="[546,728,1193,1219]" pageId="227" pageNumber="228" pagination="105 - 140" refId="ref298180" refString="Radcliffe, L. (1912) Description of a new family, two new genera, and twenty-nine new species of anacanthine fishes from the Philippine Islands and contiguous waters. Proceedings of the United States National Museum, 43, 105 - 140, pls. 22 - 31. http: // dx. doi. org / 10.5479 / si. 00963801.43 - 1924.105" type="journal article" year="1912">Radcliffe, 1912</bibRefCitation>
);
<taxonomicName id="F8B878AAFF3586F1DBE5C682FD3979D0" authority="Shcherbachev, Sazonov &amp; Iwamoto, 1992" authorityName="Shcherbachev, Sazonov &amp; Iwamoto" authorityYear="1992" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="paepkei">
<emphasis id="0DCCDF3BFF3586F1DBE5C682FF3879D0" italics="true" pageId="227" pageNumber="228">K. paepkei</emphasis>
<bibRefCitation id="5B297ED8FF3586F1D9F9C6E3FD3979D0" author="Shcherbachev, Y. N. &amp; Sazonov, Y. I. &amp; Iwamoto, T." box="[236,734,1227,1254]" pageId="227" pageNumber="228" pagination="97 - 108" refId="ref299815" refString="Shcherbachev, Y. N., Sazonov, Y. I. &amp; Iwamoto, T. (1992) Synopsis of the grenadier genus Kuronezumia (Pisces: Gadiformes: Macrouridae), with description of a new species. Proceedings of the California Academy of Sciences, 48, 97 - 108." type="journal article" year="1992">Shcherbachev, Sazonov &amp; Iwamoto, 1992</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="F8B878AAFF3586F1DBE5C6E4FDCC7832" authority="Sazonov &amp; Iwamoto, 1992" authorityName="Sazonov &amp; Iwamoto" authorityYear="1992" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="pallida">
<emphasis id="0DCCDF3BFF3586F1DBE5C6E4FF307832" italics="true" pageId="227" pageNumber="228">K. pallida</emphasis>
<bibRefCitation id="5B297ED8FF3586F1D9F0C6C5FDCC7832" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[229,555,1261,1288]" pageId="227" pageNumber="228" pagination="27 - 95" refId="ref298977" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48, 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto, 1992</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="F8B878AAFF3586F1DB61C6C6FCED7832" authorityName="Iwamoto" authorityYear="1974" box="[628,778,1262,1287]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF3586F1DB61C6C6FCED7832" box="[628,778,1262,1287]" italics="true" pageId="227" pageNumber="228">Kuronezumia</emphasis>
</taxonomicName>
sp. [this undescribed species was initially recognized by Iwamoto (1974), but still remains to be described]. Two species are distributed in Japanese waters, one of which is new to science [different from the one reported by Iwamoto (1974)] and is described below.
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF3586F1D99DC7D6FCEE7A5F" pageId="227" pageNumber="228" type="key">
<paragraph id="3F070329FF3586F1D99DC7D6FEB17B0F" blockId="227.[136,777,1533,1594]" pageId="227" pageNumber="228">
<emphasis id="0DCCDF3BFF3586F1D99DC7D6FEB17B0F" bold="true" pageId="227" pageNumber="228">
Key to species of
<taxonomicName id="F8B878AAFF3586F1D86FC7D5FDC27B22" ID-CoL="59FH" authorityName="Iwamoto" authorityYear="1974" box="[378,549,1533,1559]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF3586F1D86FC7D5FDC27B22" bold="true" box="[378,549,1533,1559]" italics="true" pageId="227" pageNumber="228">Kuronezumia</emphasis>
</taxonomicName>
from
<collectingCountry id="47AF43B9FF3586F1DB6BC7D6FD297B2D" box="[638,718,1534,1560]" name="Japan" pageId="227" pageNumber="228">Japan</collectingCountry>
and adjacent waters
</emphasis>
</paragraph>
<keyStep id="844C1B8CFF3586F1D99DC44FFCEE7A5F" pageId="227" pageNumber="228">
<key id="4B295418FF3586F1D99DC44FFCEE7A5F" pageId="227" pageNumber="228">
<paragraph id="3F070329FF3586F1D99DC44FFCE57BD6" blockId="227.[136,778,1639,1899]" pageId="227" pageNumber="228">
<keyLead id="8449A01CFF3586F1D99DC44FFCE57BD6" pageId="227" pageNumber="228">
<emphasis id="0DCCDF3BFF3586F1D99DC44FFF477B49" bold="true" box="[136,160,1639,1660]" pageId="227" pageNumber="228">1a</emphasis>
Pelvic-fin rays 1112; posterior half of mandibular rami scaled; branchiostegal membranes naked; no open pores along cephalic sensory canals; oral cavity blackish............... ....................................................................................
<taxonomicName id="F8B878AAFF3586F1DBACC4E5FCE57BD6" ID-CoL="3RG6K" baseAuthorityName="Gilbert &amp; Hubbs" baseAuthorityYear="1916" box="[697,770,1741,1763]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="dara">
<emphasis id="0DCCDF3BFF3586F1DBACC4E5FCE57BD6" bold="true" box="[697,770,1741,1763]" italics="true" pageId="227" pageNumber="228">K. dara</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3F070329FF3586F1D99DC4C7FCEE7A5F" blockId="227.[136,778,1639,1899]" pageId="227" pageNumber="228">
<keyLead id="8449A01CFF3586F1D99DC4C7FCEE7A5F" pageId="227" pageNumber="228">
<emphasis id="0DCCDF3BFF3586F1D99DC4C7FF467A31" bold="true" box="[136,161,1775,1796]" pageId="227" pageNumber="228">1b</emphasis>
Pelvic-fin rays 15; mandibular rami almost entirely naked except for posterior margins; scaly patches present on lower portions of branchiostegal rays; cephalic sensory pores present; oral cavity pale ..............................
<emphasis id="0DCCDF3BFF3586F1DB48C57EFCEE7A5F" bold="true" box="[605,777,1877,1899]" pageId="227" pageNumber="228">
<taxonomicName id="F8B878AAFF3586F1DB48C57EFD547A5E" box="[605,691,1877,1899]" class="Actinopterygii" family="Macrouridae" genus="Kuronezumia" kingdom="Animalia" order="Gadiformes" pageId="227" pageNumber="228" phylum="Chordata" rank="species" species="endoi" status="sp. nov.">
<emphasis id="0DCCDF3BFF3586F1DB48C57EFD547A5E" bold="true" box="[605,691,1877,1899]" italics="true" pageId="227" pageNumber="228">K. endoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16FF6240FF3586F1DBAEC57DFCEE7A5F" box="[699,777,1877,1898]" pageId="227" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>