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<document id="F53DCD2C4ACA0A2CB0E5E71FF4F7EDF3" ID-CLB-Dataset="24286" ID-DOI="10.11646/zootaxa.4728.2.2" ID-GBIF-Dataset="a1c40b73-0c11-4863-a188-412c33523e02" ID-ISSN="1175-5326" ID-Zenodo-Dep="3621117" ID-ZooBank="AE4F8089-7D2E-4ABA-AA09-CD504705788A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579677041620" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Withers, Phil" docDate="2020" docId="784D878B601E21078FF103E1FE4A887E" docLanguage="en" docName="zootaxa.4728.2.2.pdf" docOrigin="Zootaxa 4728 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoiriella anchoriformis Salamanna 1975" docType="treatment" docVersion="6" lastPageNumber="201" masterDocId="8474FFF3600C21158F66006DFFB28F20" masterDocTitle="The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)" masterLastPageNumber="210" masterPageNumber="183" pageNumber="201" updateTime="1698778572713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="153B1A0300CB0DC99EDFBCAF03FE1AA4">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<mods:namePart id="66218F168022CA6FDA286CF4CD6766E5">Wagner, Rüdiger</mods:namePart>
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<mods:title id="16C3A7A385AF69FDE4EE0580136D6DF3">Zootaxa</mods:title>
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<mods:date id="7D5EAF3CDDCF1915EDD1EB0AF058C1BD">2020</mods:date>
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<treatment id="784D878B601E21078FF103E1FE4A887E" ID-GBIF-Taxon="161827855" LSID="urn:lsid:plazi:treatment:784D878B601E21078FF103E1FE4A887E" httpUri="http://treatment.plazi.org/id/784D878B601E21078FF103E1FE4A887E" lastPageNumber="201" pageId="18" pageNumber="201">
<subSubSection id="B8FE6516601E21078FF103E1FD0F8C86" box="[151,701,908,935]" pageId="18" pageNumber="201" type="nomenclature">
<paragraph id="F05B369D601E21078FF103E1FD0F8C86" blockId="18.[151,701,908,971]" box="[151,701,908,935]" pageId="18" pageNumber="201">
<heading id="AB1381F1601E21078FF103E1FD0F8C86" bold="true" box="[151,701,908,935]" fontSize="11" level="1" pageId="18" pageNumber="201" reason="1">
<taxonomicName id="37E44D1E601E21078FF103E1FD0F8C86" ID-CoL="7CHTB" authority="Salamanna, 1975" authorityName="Salamanna" authorityYear="1975" box="[151,701,908,935]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="201" phylum="Arthropoda" rank="species" species="anchoriformis">
<emphasis id="C290EA8F601E21078FF103E1FD0F8C86" bold="true" box="[151,701,908,935]" pageId="18" pageNumber="201">
<emphasis id="C290EA8F601E21078FF103E1FE528C87" bold="true" box="[151,480,908,935]" italics="true" pageId="18" pageNumber="201">Tonnoiriella anchoriformis</emphasis>
<bibRefCitation id="94754B6C601E21078E8E03E1FD0F8C86" author="Salamanna, G." box="[488,701,908,934]" pageId="18" pageNumber="201" pagination="193 - 214" refId="ref20365" refString="Salamanna, G. (1975) Psychodidae Psychodinae della Puglia e della Basilicata con descrizione di due nuove species (Diptera, Nematocera). Entomologia, Bari, 11, 193 - 214." type="journal article" year="1975">Salamanna, 1975</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601E21078FF103DDFE9C8CEB" box="[151,302,944,971]" pageId="18" pageNumber="201" type="description">
<paragraph id="F05B369D601E21078FF103DDFE9C8CEB" blockId="18.[151,701,908,971]" box="[151,302,944,971]" pageId="18" pageNumber="201">
(
<figureCitation id="68DF2A18601E21078FF903DDFE948CEB" box="[159,294,944,971]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="18" pageNumber="201">Figs. 38–40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601E21078FF10398FC028B49" pageId="18" pageNumber="201" type="reference_group">
<paragraph id="F05B369D601E21078FF10398FC348B2D" blockId="18.[151,1437,1012,1129]" box="[151,902,1012,1037]" pageId="18" pageNumber="201">
<taxonomicName id="37E44D1E601E21078FF10398FDD08B2D" ID-CoL="7CHTB" authority="Salamanna, 1975" authorityName="Salamanna" authorityYear="1975" box="[151,610,1012,1037]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="201" phylum="Arthropoda" rank="species" species="anchoriformis">
<emphasis id="C290EA8F601E21078FF10398FE188B2D" box="[151,426,1013,1037]" italics="true" pageId="18" pageNumber="201">Tonnoiriella anchoriformis</emphasis>
<bibRefCitation id="94754B6C601E21078ED60399FDD08B2D" author="Salamanna, G." box="[432,610,1012,1037]" pageId="18" pageNumber="201" pagination="193 - 214" refId="ref20365" refString="Salamanna, G. (1975) Psychodidae Psychodinae della Puglia e della Basilicata con descrizione di due nuove species (Diptera, Nematocera). Entomologia, Bari, 11, 193 - 214." type="journal article" year="1975">Salamanna, 1975</bibRefCitation>
</taxonomicName>
: Entomologia, Bari, 11: 208.
</paragraph>
<paragraph id="F05B369D601E21078FF1047EFEE98B6A" blockId="18.[151,1437,1012,1129]" pageId="18" pageNumber="201">
<taxonomicName id="37E44D1E601E21078FF1047EFE488B0B" ID-CoL="7CHTL" authority="Withers, 1997" authorityName="Withers" authorityYear="1997" box="[151,506,1043,1068]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="201" phylum="Arthropoda" rank="species" species="disneyi">
<emphasis id="C290EA8F601E21078FF1047EFED08B0B" box="[151,354,1043,1067]" italics="true" pageId="18" pageNumber="201">Tonnoiriella disneyi</emphasis>
<bibRefCitation id="94754B6C601E21078E0E047EFE488B0B" author="Withers, P." box="[360,506,1043,1068]" pageId="18" pageNumber="201" pagination="61 - 64" refId="ref21053" refString="Withers, P. (1997) Tonnoiriella disneyi sp. n. and Threticus balkaneoalpinus, new to Britain (Diptera, Psychodidae). Dipterists Digest, 4, 61 - 64." type="journal article" year="1997">Withers, 1997</bibRefCitation>
</taxonomicName>
: Dipterists Digest 1997 (4): 61;
<bibRefCitation id="94754B6C601E21078C24047EFC058B0B" author="Jezek, J." box="[834,951,1043,1068]" pageId="18" pageNumber="201" pagination="7 - 18" refId="ref20095" refString="Jezek, J. (1999) Rare and new Palaearctic Tonnoiriella - species (Diptera, Psychodidae). Casopis Narodniho muzea Rada prirodovedna, 168 (1 - 4), 7 - 18." type="journal article" year="1999">Ježek 1999</bibRefCitation>
: Časopis Národniho muzea Řada přirodovědná, 168: 17 (syn.).
</paragraph>
<paragraph id="F05B369D601E21078FF1043CFC028B49" blockId="18.[151,1437,1012,1129]" box="[151,944,1104,1129]" pageId="18" pageNumber="201">
<taxonomicName id="37E44D1E601E21078FF1043CFE5B8B49" ID-CoL="7CHTW" authority="Wagner, 1993" authorityName="Wagner" authorityYear="1993" box="[151,489,1105,1129]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="201" phylum="Arthropoda" rank="species" species="holmi">
<emphasis id="C290EA8F601E21078FF1043CFEE68B49" box="[151,340,1105,1129]" italics="true" pageId="18" pageNumber="201">Tonnoiriella holmi</emphasis>
<bibRefCitation id="94754B6C601E21078E3F043CFE5B8B49" author="Wagner, R." box="[345,489,1105,1129]" pageId="18" pageNumber="201" pagination="401 - 416" refId="ref20909" refString="Wagner, R. (1993) Einige neue europaische Psychodiden. Entomofauna, 14 (24), 401 - 416." type="journal article" year="1993">Wagner, 1993</bibRefCitation>
</taxonomicName>
: Entomofauna 14 (24): 406,
<emphasis id="C290EA8F601E21078C6A043CFC028B49" bold="true" box="[780,944,1105,1129]" pageId="18" pageNumber="201">new synonymy.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601E21078FF104FBFA958BF6" pageId="18" pageNumber="201" type="materials_examined">
<paragraph id="F05B369D601E21078FF104FBFA958BF6" blockId="18.[151,1437,1174,1886]" pageId="18" pageNumber="201">
<emphasis id="C290EA8F601E21078FF104FBFF4D8B91" bold="true" box="[151,255,1174,1201]" pageId="18" pageNumber="201">Material</emphasis>
:
<materialsCitation id="408C3CC0601E21078E6804FAFBF28B91" ID-GBIF-Occurrence="2556505406" box="[270,1088,1174,1202]" collectingDate="1985-05-17" country="Germany" location="upper Schierenseebach" pageId="18" pageNumber="201" specimenCount="1" specimenCount-male="1" stateProvince="Schleswig-Holstein">
<specimenCount id="E6E2FD14601E21078E6804FAFE838B91" box="[270,305,1175,1201]" pageId="18" pageNumber="201" type="male">1♂</specimenCount>
<collectingCountry id="88F3760D601E21078E5E04FAFE178B91" box="[312,421,1175,1201]" name="Germany" pageId="18" pageNumber="201">Germany</collectingCountry>
:
<collectingRegion id="3220F87F601E21078EC904FAFD3F8B91" box="[431,653,1175,1201]" country="Germany" name="Schleswig-Holstein" pageId="18" pageNumber="201">Schleswig-Holstein</collectingRegion>
,
<location id="F53B6046601E21078DFE04F5FC2F8B91" LSID="urn:lsid:plazi:treatment:784D878B601E21078FF103E1FE4A887E:F53B6046601E21078DFE04F5FC2F8B91" box="[664,925,1175,1202]" country="Germany" name="upper Schierenseebach" pageId="18" pageNumber="201" stateProvince="Schleswig-Holstein">upper Schierenseebach</location>
,
<date id="845A105D601E21078CCE04FAFBF28B91" box="[936,1088,1174,1202]" pageId="18" pageNumber="201" value="1985-05-17">
<collectingDate id="941EE9B5601E21078CCE04FAFBF28B91" box="[936,1088,1174,1202]" pageId="18" pageNumber="201" value="1985-05-17">17 May 1985</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="408C3CC0601E21078B2A04FAFD308BF5" ID-GBIF-Occurrence="2556505414" collectingDate="1985-05-22" country="Germany" location="lower Schierenseebach" pageId="18" pageNumber="201" specimenCount="1" specimenCount-male="1">
<specimenCount id="E6E2FD14601E21078B2A04FAFBDD8B91" box="[1100,1135,1175,1201]" pageId="18" pageNumber="201" type="male">1♂</specimenCount>
<collectingCountry id="88F3760D601E21078B1004FAFB518B91" box="[1142,1251,1175,1201]" name="Germany" pageId="18" pageNumber="201">Germany</collectingCountry>
: Schleswig-Hol- stein,
<location id="F53B6046601E21078FBC04D6FE6D8BF5" LSID="urn:lsid:plazi:treatment:784D878B601E21078FF103E1FE4A887E:F53B6046601E21078FBC04D6FE6D8BF5" box="[218,479,1211,1237]" country="Germany" name="lower Schierenseebach" pageId="18" pageNumber="201">lower Schierenseebach</location>
,
<date id="845A105D601E21078E8F04D6FD308BF5" box="[489,642,1210,1238]" pageId="18" pageNumber="201" value="1985-05-22">
<collectingDate id="941EE9B5601E21078E8F04D6FD308BF5" box="[489,642,1210,1238]" pageId="18" pageNumber="201" value="1985-05-22">22 May 1985</collectingDate>
</date>
</materialsCitation>
, all leg. Holm (
<typeStatus id="2F5F883F601E21078C4804D6FC208BF5" box="[814,914,1211,1237]" pageId="18" pageNumber="201" type="holotype">holotype</typeStatus>
and
<typeStatus id="2F5F883F601E21078CAF04D1FB9B8BF6" box="[969,1065,1212,1238]" pageId="18" pageNumber="201" type="paratype">paratype</typeStatus>
of
<taxonomicName id="37E44D1E601E21078B2804D6FAAD8BF5" authorityName="Wagner" authorityYear="1993" box="[1102,1311,1211,1237]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="201" phylum="Arthropoda" rank="species" species="holmi">
<emphasis id="C290EA8F601E21078B2804D6FAAD8BF5" box="[1102,1311,1211,1237]" italics="true" pageId="18" pageNumber="201">Tonnoiriella holmi</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601E21078FA104B3FCBC89ED" pageId="18" pageNumber="201" type="description">
<paragraph id="F05B369D601E21078FA104B3FD5F8AA9" blockId="18.[151,1437,1174,1886]" pageId="18" pageNumber="201">
<emphasis id="C290EA8F601E21078FA104B3FEE38BD9" bold="true" box="[199,337,1246,1273]" pageId="18" pageNumber="201">Description</emphasis>
: Head round, eyebridge of 3 facet rows; distance between eyes 4.7 facet diameters; with postocular bristles. Palpus segments, absolute length: 0.072-0.076-0.080-0.152; relative length: 36-38-40-76. Antenna with scape, pedicel and 5 flagellomeres, distal segments missing: absolute length 0.044-0.052-0.040-0.040-0.038-0.036- 0.040- mm; relative length 22-26-20-20-19-18-20-; distal flagellomeres missing; flagellomere 5 and following with a pair of digitiform curved ascoids in the apical third.
</paragraph>
<paragraph id="F05B369D601E21078FA105FEFC708A8E" blockId="18.[151,1437,1174,1886]" box="[199,962,1426,1454]" pageId="18" pageNumber="201">
Wing 2.83 times longer than wide; length
<quantity id="371C9B78601E21078DF905FFFCB78A8E" box="[671,773,1426,1454]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="18" pageNumber="201" unit="mm" value="2.05">2.05 mm</quantity>
, width
<quantity id="371C9B78601E21078C3105FEFC0C8A8E" box="[855,958,1427,1454]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="18" pageNumber="201" unit="mm" value="0.72">0.72 mm</quantity>
.
</paragraph>
<paragraph id="F05B369D601E21078FA105DAFAA28939" blockId="18.[151,1437,1174,1886]" pageId="18" pageNumber="201">
Male terminalia with hypandrium with broad median linguiform enlargement, tip almost reaching tip of gonocoxites (
<figureCitation id="68DF2A18601E21078F9305B6FEFB8AD5" box="[245,329,1499,1525]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="18" pageNumber="201">Fig. 38</figureCitation>
). Gonocoxites as long as their basal width, with 1 long seta near apical end on inner edge. Gonostylus with basal swelling as long as the gonocoxite incised near tip with spearhead appearance (
<figureCitation id="68DF2A18601E21078BC90592FAB68939" box="[1199,1284,1535,1561]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="18" pageNumber="201">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="F05B369D601E21078FA10649FC5489A5" blockId="18.[151,1437,1174,1886]" pageId="18" pageNumber="201">Parameres not clearly identified, small but seemingly not different from other species. Aedeagus with ventral lobe of the basiphallus short straight; distiphallus sclerite with long basal arm, cross part slightly bent, thin. Dorsal lobe of basiphallus is broad, straight, and symmetric, no dorsal distiphallus.</paragraph>
<paragraph id="F05B369D601E21078FA106E2FCBC89ED" blockId="18.[151,1437,1174,1886]" pageId="18" pageNumber="201">
Epandrium rhomboid short; epandrial processes basally bulky distally thinner with 3 feathered retinacula, 1 long and 2 short (
<figureCitation id="68DF2A18601E21078E3A06DEFE0289ED" box="[348,432,1715,1741]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="18" pageNumber="201">Fig. 40</figureCitation>
). Hypoproct oval, microsetose.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601E21078FA106BBFD0589D1" box="[199,695,1750,1778]" pageId="18" pageNumber="201" type="distribution">
<paragraph id="F05B369D601E21078FA106BBFD0589D1" blockId="18.[151,1437,1174,1886]" box="[199,695,1750,1778]" pageId="18" pageNumber="201">
<emphasis id="C290EA8F601E21078FA106BBFEEB89D1" bold="true" box="[199,345,1750,1777]" pageId="18" pageNumber="201">Distribution</emphasis>
:
<collectingCountry id="88F3760D601E21078E0E06BAFE2E89D1" box="[360,412,1751,1777]" name="Italy" pageId="18" pageNumber="201">Italy</collectingCountry>
,
<collectingCountry id="88F3760D601E21078ECE06BAFDA389D1" box="[424,529,1751,1777]" name="Germany" pageId="18" pageNumber="201">Germany</collectingCountry>
,
<collectingCountry id="88F3760D601E21078D7A06BAFD0089D1" box="[540,690,1751,1777]" name="United Kingdom" pageId="18" pageNumber="201">Great Britain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601E21078FA10696FE4A887E" pageId="18" pageNumber="201" type="discussion">
<paragraph id="F05B369D601E21078FA10696FE4A887E" blockId="18.[151,1437,1174,1886]" pageId="18" pageNumber="201">
<emphasis id="C290EA8F601E21078FA10696FE868835" bold="true" box="[199,308,1787,1813]" pageId="18" pageNumber="201">Remarks</emphasis>
: This well defined species is widely distributed in Western Europe. However, it is rarely collected which finally caused the repeated descriptions from
<collectingCountry id="88F3760D601E21078DB10772FC8D8819" box="[727,831,1823,1849]" name="Germany" pageId="18" pageNumber="201">Germany</collectingCountry>
and
<collectingCountry id="88F3760D601E21078C150772FC668819" box="[883,980,1823,1849]" name="United Kingdom" pageId="18" pageNumber="201">England</collectingCountry>
. German specimens were collected along a slow running lowland stream.
</paragraph>
</subSubSection>
</treatment>
</document>
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