242 lines
20 KiB
XML
242 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="1D5B716EC5CB5397A1659BC9C8255F0B" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina dorri subsp. isanensis Jirapatrasilp 2022, ssp. nov." docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="199838999" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B" httpUri="http://treatment.plazi.org/id/1D5B716EC5CB5397A1659BC9C8255F0B" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="1D5B716E-C5CB-5397-A165-9BC9C8255F0B" authority="Jirapatrasilp" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina dorri subsp. isanensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="Dorri" status="ssp. nov." subSpecies="isanensis">Pupina dorri isanensis Jirapatrasilp</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">ssp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">Figs 36K, L</figureCitation>
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<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure39" httpUri="https://binary.pensoft.net/fig/735400" pageId="0" pageNumber="1">, 39E, F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 12140/1 (Figs
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36K</figureCitation>
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,
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<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure39" httpUri="https://binary.pensoft.net/fig/735400" pageId="0" pageNumber="1">39E</figureCitation>
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), 31 Aug. 2020, coll. C. Sutcharit, P. Jirapatrasilp, A. Pholyotha. Measurement: shell height 6.6 mm, shell width 4.6 mm and
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<normalizedToken originalValue="5½">51/2</normalizedToken>
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whorls.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 12140/2 (22 shells) and NHMUK 20210337 (3 shells), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Pa Pha Ya Temple, Suwannakhuha District, Nong Bua Lam Phu Province, Thailand,
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<geoCoordinate degrees="17" direction="north" minutes="37" orientation="latitude" precision="1" seconds="38.8" value="17.627445">17°37'38.8"N</geoCoordinate>
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,
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<geoCoordinate degrees="102" direction="east" minutes="10" orientation="longitude" precision="1" seconds="13.7" value="102.17047">102°10'13.7"E</geoCoordinate>
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, 250 m amsl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2007-10-15" collectorName="Figs" location="Na Wang District" municipality="Nong Bua Lam Phu Province" specimenCount="1">
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CUMZ 12137 (1 shell;
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<collectorName>Figs</collectorName>
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36L</figureCitation>
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,
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<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure39" httpUri="https://binary.pensoft.net/fig/735400" pageId="0" pageNumber="1">39F</figureCitation>
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) from Khao Wang Pha,
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:E93DDB9D345C751B0BFC5A62F9144728" municipality="Nong Bua Lam Phu Province" name="Na Wang District">Na Wang District</location>
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,
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<collectingMunicipality>Nong Bua Lam Phu Province</collectingMunicipality>
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,
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<collectingDate value="2007-10-15">15 Oct. 2007</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2007-10-15" county="Nong Bua Lam Phu Province" location="Pa Jor Cave" municipality="Na Wang District" specimenCount="1">
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CUMZ 12138 (1 shell) from
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:2D28654533D89CBDC4FAAEC7B05C40D0" county="Nong Bua Lam Phu Province" municipality="Na Wang District" name="Pa Jor Cave">Pa Jor Cave</location>
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,
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<collectingMunicipality>Na Wang District</collectingMunicipality>
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,
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<collectingCounty>Nong Bua Lam Phu Province</collectingCounty>
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,
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<collectingDate value="2007-10-15">15 Oct. 2007</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-08-31" collectorName="Tham Suwannakhuha Temple" location="Suwannakhuha District" municipality="Nong Bua Lam Phu Province" specimenCount="1">
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CUMZ 12139 (9 shells) from
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<collectorName>Tham Suwannakhuha Temple</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:59A3B16627A6DACFFE6B6AC70D5E40B6" municipality="Nong Bua Lam Phu Province" name="Suwannakhuha District">Suwannakhuha District</location>
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,
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<collectingMunicipality>Nong Bua Lam Phu Province</collectingMunicipality>
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,
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<collectingDate value="2020-08-31">31 Aug. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-08-31" county="Nong Bua Lam Phu Province" location="Namtok Thao To Forest" municipality="Mueang Nong Bua Lam Phu District" specimenCount="1">
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CUMZ 12141 (2 shells) from
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:DE23C23CEA71AC910AC94D5A0D768342" county="Nong Bua Lam Phu Province" municipality="Mueang Nong Bua Lam Phu District" name="Namtok Thao To Forest">Namtok Thao To Forest</location>
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Park,
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<collectingMunicipality>Mueang Nong Bua Lam Phu District</collectingMunicipality>
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,
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<collectingCounty>Nong Bua Lam Phu Province</collectingCounty>
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,
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<collectingDate value="2020-08-31">31 Aug. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-09-01" collectorName="Phu Pha Lom" location="Mueang Loei District" municipality="Loei Province" specimenCount="1">
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CUMZ 12142 (3 shells and
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<specimenCount type="generic">1 specimen</specimenCount>
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in ethanol) from
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<collectorName>Phu Pha Lom</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:55B18CDB07CD41BF4F16E9A44D7F6E0E" municipality="Loei Province" name="Mueang Loei District">Mueang Loei District</location>
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,
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<collectingMunicipality>Loei Province</collectingMunicipality>
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,
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<collectingDate value="2020-09-01">1 Sept. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-09-02" collectorName="Hin Pha Ngam Park" location="Nong Hin District" municipality="Loei Province" specimenCount="1">
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CUMZ 12143 (
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<specimenCount type="generic">1 specimen</specimenCount>
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in ethanol) from
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<collectorName>Hin Pha Ngam Park</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:C2401E5B90FE7FA1199C2B3BD1DFCF5D" municipality="Loei Province" name="Nong Hin District">Nong Hin District</location>
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,
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<collectingMunicipality>Loei Province</collectingMunicipality>
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,
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<collectingDate value="2020-09-02">2 Sept. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-07-21" county="Khon Kaen Province" location="Phraya Nakharaj Cave" municipality="Phu Pha Man District" specimenCount="1">
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CUMZ 12144 (1 shell) from
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:67E285BDD6E12ED410106652C6D4E082" county="Khon Kaen Province" municipality="Phu Pha Man District" name="Phraya Nakharaj Cave">Phraya Nakharaj Cave</location>
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,
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<collectingMunicipality>Phu Pha Man District</collectingMunicipality>
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,
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<collectingCounty>Khon Kaen Province</collectingCounty>
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,
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<collectingDate value="2020-07-21">21 July 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2020-08-30" collectorName="Phu Thong Thep Nimit Temple" location="Nong Saeng District" municipality="Udon Thani Province" specimenCount="1">
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CUMZ 12145 (
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<specimenCount type="generic">1 specimen</specimenCount>
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in ethanol) from
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<collectorName>Phu Thong Thep Nimit Temple</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:8A0BBAB5FA466B934DE29172C6CC20D1" municipality="Udon Thani Province" name="Nong Saeng District">Nong Saeng District</location>
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,
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<collectingMunicipality>Udon Thani Province</collectingMunicipality>
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,
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<collectingDate value="2020-08-30">30 Aug. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2009-09-15" county="Chanthaburi Province" location="Khao Wong Cave" municipality="Kaeng Hang Maeo District" specimenCount="1">
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CUMZ 12170 (1 shell) from
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<location LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:82DE7C824E0DA7FE3720F53B43B74F47" county="Chanthaburi Province" municipality="Kaeng Hang Maeo District" name="Khao Wong Cave">Khao Wong Cave</location>
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,
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<collectingMunicipality>Kaeng Hang Maeo District</collectingMunicipality>
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,
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<collectingCounty>Chanthaburi Province</collectingCounty>
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,
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<collectingDate value="2009-09-15">15 Sept. 2009</collectingDate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate-fusiform; last whorl ca. 70% of shell height. Apertural lip thickened but not expanded. Parietal tooth triangular, thickened, covering posterior canal, approaching but not extending beyond the outer margin of apertural lip; columellar tooth fin-shaped, thickened, located next to slit-like anterior canal.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">This new subspecies is slightly different from the nominotypical subspecies in having the apertural lip, and parietal and columellar teeth more thickened.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell height 6.0-6.6 mm; shell width 4.2-4.6 mm. Shell ovate-fusiform, solid, semi-transparent, grey to pale brown, devoid of prominent sculpture on glazed smooth surface. Apex obtuse. Growth lines on shell surface inconspicuous. Whorls
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<normalizedToken originalValue="5½">51/2</normalizedToken>
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, last whorl large (ca. 70% of shell height). Spire angle ca. 75-80°, slightly extended. Sutures slightly impressed, but shallow. Aperture circular; lip thickened (ca. 0.2 mm wide and 0.3-0.4 mm thick) with paler colour, not expanded. Parietal callus not sharply defined and somewhat thickened with paler colour. Peristome interrupted by two canals; posterior canal slit-like ca. 0.7 mm long; anterior canal slit-like continuing horizontally ca. 0.8-0.9 mm. Parietal tooth triangular, thickened (ca. 0.7 mm long, 0.5 mm at its widest and 0.3 mm thick), covering posterior canal, approaching but not extending beyond the outer margin of apertural lip. Columellar tooth thickened (ca. 1.0 mm long, 0.3 mm at its widest and 0.3 mm thick), fin-shaped. Umbilicus closed. Operculum round, thin, flat, multispiral, whitish to pale yellow, opaque corneous with smooth edge.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specific epithet refers to the Thai name
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<normalizedToken originalValue="“Isan”">"Isan"</normalizedToken>
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for the northeastern region of Thailand, where this new subspecies is mainly distributed.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Northeastern and eastern Thailand.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">Although the collecting localities of this new subspecies are ca. 600 km from the known occurrence of the nominotypical subspecies, DNA data and morphometric analyses are required to demonstrate whether these Thai specimens are distinct from the Vietnamese specimens and should be elevated to specific status.</paragraph>
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</subSubSection>
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</treatment>
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</document> |