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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.744.22970" ID-GBIF-Dataset="b39e3520-4b7d-46aa-bdc8-d4c1e8cd9b1b" ID-PMC="PMC5904516" ID-Pensoft-Pub="1313-2970-744-19" ID-PubMed="29670442" ID-ZBK="0676B41BF28B42068B88FC39B0F2A196" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-744-19" ModsDocOrigin="ZooKeys 744" ModsDocTitle="The genus Paleosepharia Laboissière, 1936 in Taiwan: review and nomenclatural changes (Coleoptera, Chrysomelidae, Galerucinae)" checkinTime="1521539696622" checkinUser="pensoft" docAuthor="Lee, Chi-Feng" docDate="2018" docId="49A92C5B17D1CA5E7422FFE208022009" docLanguage="en" docName="ZooKeys 744: 19-41" docOrigin="ZooKeys 744" docSource="http://dx.doi.org/10.3897/zookeys.744.22970" docTitle="Paleosepharia excavata" docType="treatment" docVersion="4" lastPageNumber="27" masterDocId="8563FFB6FF92812D59793616FFF06A69" masterDocTitle="The genus Paleosepharia Laboissiere, 1936 in Taiwan: review and nomenclatural changes (Coleoptera, Chrysomelidae, Galerucinae)" masterLastPageNumber="41" masterPageNumber="19" pageNumber="23" updateTime="1668165581208" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The genus Paleosepharia Laboissiere, 1936 in Taiwan: review and nomenclatural changes (Coleoptera, Chrysomelidae, Galerucinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lee, Chi-Feng</mods:namePart>
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<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>744</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.744.22970</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.744.22970</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-744-19</mods:identifier>
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<treatment ID-GBIF-Taxon="142359515" LSID="urn:lsid:plazi:treatment:49A92C5B17D1CA5E7422FFE208022009" httpUri="http://treatment.plazi.org/id/49A92C5B17D1CA5E7422FFE208022009" lastPageId="8" lastPageNumber="27" pageId="4" pageNumber="23">
<subSubSection pageId="4" pageNumber="23" type="nomenclature">
<paragraph pageId="4" pageNumber="23">
<taxonomicName authority="Chujo" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia excavata" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="excavata">
Paleosepharia excavata (
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
)
</taxonomicName>
Figs 1
<normalizedToken originalValue="DF">D-F</normalizedToken>
, 3
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="24" type="reference_group">
<paragraph pageId="5" pageNumber="24">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta excavata" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="excavata">
<pageBreakToken pageId="5" pageNumber="24" start="start">Monolepta</pageBreakToken>
excavata
</taxonomicName>
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
, 1938 (Taiwan: Ilan, Nantou, Hualien): 144;
<bibRefCitation author="Chujo, M" journalOrPublisher="The Philippine Journal of Science" pageId="20" pageNumber="39" pagination="1 - 239" title="A taxonomic study on the Chrysomelidae (Insecta: Coleoptera) from Formosa Part XI. Subfamily Galerucinae." volume="91" year="1962">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
1962
</bibRefCitation>
: 114 (redescription);
<bibRefCitation author="Chujo, M" journalOrPublisher="Special Bulletin of Lepidopterological Society Japan" pageId="20" pageNumber="39" pagination="88 - 104" title="Chrysomelid-beetles of Formosa (I)." volume="1" year="1965">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
1965
</bibRefCitation>
: 96 (Nantou);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Esakia" pageId="20" pageNumber="39" pagination="1 - 68" title="Motes on the Chrysomelidae from Taiwan II." volume="7" year="1969">Kimoto 1969</bibRefCitation>
: 47 (Taichung, Nantou);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="20" pageNumber="39" pagination="53 - 62" title="The Chrysomelidae (Insecta: Coleoptera) collected by the Nagoya University Scientific Expedition to Taiwan in 1984." volume="35" year="1986">Kimoto 1986</bibRefCitation>
: 58 (Nantou);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="20" pageNumber="39" pagination="237 - 272" title="The Taiwanese Chrysomelidae (Insecta: Coleoptera) collected by Dr. Kintaro Baba, on the occasion of his entomological survey in 1983 and 1986." volume="38" year="1989">Kimoto 1989</bibRefCitation>
: 254 (Taitung);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="21" pageNumber="40" pagination="1 - 27" title="The Taiwanese Chrysomelidae (Insecta: Coleoptera) collected by Dr. Kintaro Baba, on the occasion of his entomological survey in 1987, 1988 and 1989." volume="40" year="1991">Kimoto 1991</bibRefCitation>
: 14 (Taoyuan);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Bulletin of the Institute of Comparative Studies of International Cultures and Societies" pageId="21" pageNumber="40" pagination="1 - 152" title="Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera)." volume="16" year="1996">Kimoto and Chu 1996</bibRefCitation>
: 79 (catalogue);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="21" pageNumber="40" title="Leaf beetles (Chrysomelidae) of Taiwan." year="1997">Kimoto and Takizawa 1997</bibRefCitation>
: 385 (catalogue);
<bibRefCitation author="Lee, C-F" journalOrPublisher="Sishou-Hills Insect Observation Network and Taiwan Agricultural Research Institute, COA, New Taipei City and Taichung City" pageId="21" pageNumber="40" title="The Chrysomelidae of Taiwan 2." year="2010">Lee and Cheng 2010</bibRefCitation>
: 101 (food plants);
<bibRefCitation author="Yang, X-K" journalOrPublisher="Science Press, Beijing" pageId="21" pageNumber="40" title="Chinese leaf beetles." year="2015">Yang et al. 2015</bibRefCitation>
: 268 (catalogue).
</paragraph>
<paragraph pageId="5" pageNumber="24">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia excavata" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="excavata">Paleosepharia excavata</taxonomicName>
:
<bibRefCitation pageId="5" pageNumber="24">Gressitt and Kimoto 1963</bibRefCitation>
: 646;
<bibRefCitation pageId="5" pageNumber="24">Beenen 2010</bibRefCitation>
: 485 (catalogue).
</paragraph>
<paragraph pageId="5" pageNumber="24">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia polychroma" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="polychroma">Paleosepharia polychroma</taxonomicName>
<normalizedToken originalValue="Laboissière">Laboissiere</normalizedToken>
, 1938: 8 (China: Jiangsu, Jiangxi);
<bibRefCitation pageId="5" pageNumber="24">Gressitt and Kimoto 1963</bibRefCitation>
: 646 (as synonym of
<taxonomicName family="Chrysomelidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. excavata" order="Coleoptera" pageId="5" pageNumber="24" rank="species" species="excavata">P. excavata</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="24" type="type material">
<paragraph pageId="5" pageNumber="24">Type material.</paragraph>
<paragraph pageId="5" pageNumber="24">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta excavata" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="excavata">Monolepta excavata</taxonomicName>
. Lecotype ♂ (TARI), here designated, labeled: &quot;Taiheizan [= Taipingshan,
<normalizedToken originalValue="太平山">太平山</normalizedToken>
] / 23-V-1931 / Col. R. Takahashi [p, w] // CO / Type [p, y, circular label with yellow border] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Monolepta</taxonomicName>
/
<taxonomicName lsidName="excavata" pageId="5" pageNumber="24" rank="species" species="excavata">excavata</taxonomicName>
/
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] // DET. M. CHUJO [p, w] // 2193 [p, w]&quot;. Paralectotypes. 1♂ (TARI): &quot;Horisha [= Puli,
<normalizedToken originalValue="埔里">埔里</normalizedToken>
] / 12/V/1913 [h] / Col. M. Maki [p, w] // CO / Type [p, y, circular label with yellow border] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Monolepta</taxonomicName>
/
<taxonomicName lsidName="excavata" pageId="5" pageNumber="24" rank="species" species="excavata">excavata</taxonomicName>
/
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] // DET. M. CHUJO [p, w] // 2194 [p, w]&quot;; 1♂, 1♀ (TARI): &quot;Kuaru [h] [unknown] / FORMOSA [p] / 20.VI.1937 [h] / COL. M. CHUJO [p, w] // CO / Type [p, y, circular label with yellow border] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Monolepta</taxonomicName>
/
<taxonomicName lsidName="excavata" pageId="5" pageNumber="24" rank="species" species="excavata">excavata</taxonomicName>
/
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] // DET. M. CHUJO [p, w] // No. 1640 or 1346 [p, w]&quot;; 1♀ (TARI): &quot;Formosa / Musha, [= Wushe,
<normalizedToken originalValue="霧社">霧社</normalizedToken>
] 1919 / V.18-VI.15. / T. Okuni [p, w] // CO / Type [p, y, circular label with yellow border] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Monolepta</taxonomicName>
/
<taxonomicName lsidName="excavata" pageId="5" pageNumber="24" rank="species" species="excavata">excavata</taxonomicName>
/
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] // DET. M. CHUJO [p, w] // 1396 [p, w]&quot;; 1♀ (TARI): &quot;18/IV/1910 / Kanmon [h] [= Kuangman,
<normalizedToken originalValue="關門">關門</normalizedToken>
] / Col. I. Nitobe [p, w] // CO / Type [p, y, circular label with yellow border] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Monolepta</taxonomicName>
/
<taxonomicName lsidName="excavata" pageId="5" pageNumber="24" rank="species" species="excavata">excavata</taxonomicName>
/
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] // DET. M. CHUJO [p, w] // 1973 [p, w]&quot;; 1♀ (SDEI): &quot;Formosa / Karenko [= Hualien,
<normalizedToken originalValue="花蓮">花蓮</normalizedToken>
], -19. / VII 20-VIII 4. / T. Okuni [p, w] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Monolepta" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Monolepta</taxonomicName>
/
<taxonomicName lsidName="excavata" pageId="5" pageNumber="24" rank="species" species="excavata">excavata</taxonomicName>
/
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] // DET. M. CHUJO [p, w] // 1973 [p, w] // Syntypus [p, r]&quot;.
</paragraph>
<paragraph pageId="5" pageNumber="24">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia polychroma" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="polychroma">Paleosepharia polychroma</taxonomicName>
. The syntypes at the Institute Royal des Sciences Naturelles de Belgique, Bruxelles, and the Naturhistoriska Riksmuseet, Stockholm were not studied.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">Paleosepharia</taxonomicName>
<taxonomicName lsidName="polychroma" pageId="5" pageNumber="24" rank="species" species="polychroma">polychroma</taxonomicName>
could be a distinct species because of slight difference of color patterns on the elytra between both species. Correctly assessing the status of this species requires further study.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="26" pageId="5" pageNumber="24" type="materials_examined">
<paragraph pageId="5" pageNumber="24">Other material examined</paragraph>
<paragraph lastPageId="7" lastPageNumber="26" pageId="5" pageNumber="24">
(n = 72). Chiayi: 1♂ (TARI), Alishan [阿里山], 5-9.VII.1981, leg. L. Y. Chou &amp; S. C. Lin; Hsinchu: 1♂ (TARI), Kuanwu [觀霧], 6.IV.2010, leg. L.-H. Sun; 1♂ (TARI), same locality, 30.IV.2010, leg. M.-H. Tsou; 1♀ (TARI), Litungshan [李棟山], 6.VI.2010, leg. Y.-L. Lin; 1♂ (TARI), Mamei [馬美], 4.V.2008, leg. S.-F. Yu; 3♀♀ (TARI), same locality, 18.V.2008, leg. M.-H. Tsou; 1♂ (TARI), Talu trail [大鹿林道], 19.VI.2010, leg. Y.-L. Lin; 1♀ (TARI), same locality, 11.VII.2016, leg. H. Lee; Hualien: 1♂, 1♀ (TARI), Hsinpaiyang [新白陽], 15-20.X.2007, leg. Y.-F. Hsu; 1♀ (TARI), Huitouwan [迴頭灣], 10.VII.2007, leg. C.-F. Lee; 1♂, 1♀ (TARI), Wanjung [萬榮], 11.VI.2011, leg. M.-H. Tsou; Ilan: 1♀ (TARI), Chinyang [金洋], 23.X.2011, leg. C.-H. Hsieh; 1♀ (NMNS), Tsuifenghu [翠峰湖], 21.VI.1992, leg. C. Y. Li; 1♂ (NMNS), Taipingshan [太平山], 8.III.1989, leg. K. W. Huang; 1♂, 3♀♀ (TARI), same locality, 17.VIII.2007, leg. Y.-C. Chang; Nantou: 1♂ (NMNS), Aowanta [奧萬大], 9.IX.2008, leg. C. C. Lai; 1♂ (TARI), Fenghuangshan [鳳凰山], 10.V.2010, leg. Y.-T. Wang; 1♀ (TARI), Hoshe [和社], 23.I.2014, leg. H.-T. Yeh; 1♂ (TARI), Huakang [華岡], 13.IX.2010, leg. C.-F. Lee; 1♀ (TARI), Meifeng [梅峰], 20-22.VI.1979, leg. K. S. Lin &amp; B. H. Chen; 1♀ (NMNS), same locality, 24.IX.1997, leg. W. T. Yang; 1♂ (NMNS), Oiwake [= Tsuifeng,
<normalizedToken originalValue="翠"></normalizedToken>
<pageBreakToken pageId="6" pageNumber="25" start="start">
<normalizedToken originalValue="峰"></normalizedToken>
</pageBreakToken>
]-Tattaka [= Sungkang,
<normalizedToken originalValue="松崗">松崗</normalizedToken>
], 24.VI.1961, leg. T. Shirozu; 1♂ (TARI), Tsuifeng [翠峰], 21.VI.1979, leg. K. S. Lin &amp; B. H. Chen; 1♂ (TARI), same locality, 25-27.VI.1981, leg. K. S. Lin &amp; W. S. Tang; 1♂ (TARI), Tungfu [同富], 8.V.2011, leg. C.-
<pageBreakToken pageId="7" pageNumber="26" start="start">F</pageBreakToken>
. Lee; Taichung: 3♂♂, 3♀♀ (TARI), Anmashan [鞍馬山], 21.IV.2010, leg. C.-F. Lee; Taitung: 1♂ (TARI), Hsiangyang [向陽], 9.V.2013, leg. J.-C. Chen; 1♂ (TARI), Lichia [利嘉], 19.V.2009, leg. U. Ong; 3♀♀ (TARI), same locality, 15.VII.2014, leg. Y.-T. Chung; 1♀ (TARI), Litao [利稻], 23.VI.2010, leg. M.-H. Tsou; 4♀♀ (TARI), Liyuan [栗園], 29.III.2011, leg. C.-F. Lee; 1♀ (TARI), same locality, 24.VII.2013, leg. C.-F. Lee; 1♂ (TARI), same locality, 24.II.2014, leg. J.-C. Chen; 1♂ (TARI), same locality, 14.III.2014, leg. W.-C. Huang; 2♀♀, same locality, 28.III.2014, leg. W.-C. Huang; 1♀ (TARI), Motien [摩天], 23.VI.2010, leg. M.-H. Tsou; 1♀ (TARI), Wulu [霧鹿], 24.VI.2010, leg. M.-H. Tsou; 1♀ (TARI), same locality, 5.X.2010, leg. M.-H. Tsou; 1♂, 5♀♀ (TARI), same locality, 29.III.2011, leg. C.-F. Lee; 1♂ (NMNS), Yenping [延平], 30.VII.1992, leg. W. T. Yang; Taoyuan: 3♂♂, 1♀ (TARI), Hsuanyuan [萱源], 16.XII2008, leg. S.-F. Yu; 1♀ (TARI), Lalashan [拉拉山], 10.X.2009, leg. H. Lee; 2♂♂, 6♀♀ (TARI), Paling [巴陵], 8.XI.2009, leg. M.-H. Tsou; 1♀ (TARI), Tungyanshan [東眼山], 15.V.2010, leg. H. Lee; 1♂ (TARI), same locality, 5.XI.2011, leg. H. Lee.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="26" type="diagnosis">
<paragraph pageId="7" pageNumber="26">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="26">
Adults of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia excavata" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="excavata">Paleosepharia excavata</taxonomicName>
are similar to those of
<taxonomicName lsidName="P. formosana" pageId="7" pageNumber="26" rank="species" species="formosana">P. formosana</taxonomicName>
and
<taxonomicName lsidName="P. yasumatsui" pageId="7" pageNumber="26" rank="species" species="yasumatsui">P. yasumatsui</taxonomicName>
in possessing two transverse black bands on yellow elytra, but this species is easily recognized by its slender, indistinctly margined transverse black bands (broad and distinctly margined bands in others). Males of these species are also similar in possessing a broader penis (less than 6.0 times longer than wide; more than 6.5 times in other species), bifurcate apex of tectum (acute apex in other species), and lacking a pair of elongate and apically curved spiculae (such spiculae present in other species). Males of
<taxonomicName lsidName="P. excavata" pageId="7" pageNumber="26" rank="species" species="excavata">P. excavata</taxonomicName>
differ in possessing a shallow notch in the apex of the tectum (deep notch in apex of tectum in
<taxonomicName lsidName="P. yasumatsui" pageId="7" pageNumber="26" rank="species" species="yasumatsui">P. yasumatsui</taxonomicName>
), recurved apex of penis (not recurved in
<taxonomicName lsidName="P. yasumatsui" pageId="7" pageNumber="26" rank="species" species="yasumatsui">P. yasumatsui</taxonomicName>
), short hooked spiculae 1/2 lengths of longer spiculae (1/6 lengths of longer spiculae in
<taxonomicName lsidName="P. formosana" pageId="7" pageNumber="26" rank="species" species="formosana">P. formosana</taxonomicName>
, 4/5 lengths of longer spiculae in
<taxonomicName lsidName="P. yasumatsui" pageId="7" pageNumber="26" rank="species" species="yasumatsui">P. yasumatsui</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26" type="description">
<paragraph pageId="7" pageNumber="26">Description.</paragraph>
<paragraph lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26">
Males. Length 4.8-6.1 mm, width 2.5-3.0 mm. General color reddish brown (Fig. 1
<normalizedToken originalValue="DE">D-E</normalizedToken>
); antennae, scutellum, tibiae, and tarsi black; elytra yellow, with slender black stripes along each lateral margin and suture extending from base to apex, and two transverse, slender, indistinctly defined black bands at basal 1/3 and 2/3, usually abbreviated near suture and lateral margins. Antenna (Fig. 3A) filiform, ratio of length of antennomeres I to XI 1.0: 0.4: 0.6: 0.9: 0.9: 0.9: 0.8: 0.8: 0.8: 0.7: 0.8; ratio of length to width from antennomere I-XI 4.6: 2.1: 3.1: 4.8: 4.8: 5.6: 5.4: 5.5: 5.5: 5.0: 5.7. Pronotum 1.58-1.59 times wider than long; lateral margins slightly curved, basal margin slightly curved, apical margin slightly concave; disc with lateral fovea and dense minute punctures. Elytra parallel-sided; 1.49-1.56 times longer than wide; with one pair of oblique dark depressions at suture behind scutellum; disc with dense, minute punctures; apex truncate. Penis (Fig. 3
<normalizedToken originalValue="CE">C-E</normalizedToken>
) elongate, 4.5 times longer than wide; parallel-sided, abruptly widened at apical 1/3, apically tapering; tectum elongate from middle to near apex, basally broadened, apex bifurcate; moderately recurved at basal 1/4 in lateral view; ventral surface with lateral areas membranous, apical membranous area short. Endophallic spiculae complex (Fig. 3F) with longest pair directed anteriorly, one pair of hooked spiculae 1/2 lengths of the longest spiculae;
<pageBreakToken pageId="8" pageNumber="27" start="start">one</pageBreakToken>
pair of longitudinal rows of hooked setae starting from bases of longest spiculae, smaller near base; one row of elongate, apically tapering setae covering surface behind hook-like setae; and one paired cluster of short, hooked setae near base.
</paragraph>
<caption pageId="8" pageNumber="27">
<paragraph pageId="8" pageNumber="27">
Figures 3. Diagnostic characters of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paleosepharia excavata" order="Coleoptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="excavata">Paleosepharia excavata</taxonomicName>
(
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
). A Antenna, male B Antenna, female C Penis, dorsal view D Penis, lateral view E Penis, ventral view F Endophallic spiculae, hooked spiculae removed at right side G Abdominal ventrite VIII H Bursa sclerites, left sclerite in lateral view, right sclerite in dorsal view I Spermatheca J Gonocoxae.
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="27">Females. Length 5.4-6.3 mm, width 2.8-3.3 mm. Similar to male (Fig. 1F); but elytra lacking oblique depressions, anterior transverse black band extending to suture; ratio of length of antennomeres I to XI (Fig. 3B) 1.0: 0.3: 0.6: 0.8: 0.7: 0.7: 0.8: 0.8: 0.8: 0.7: 0.8; ratio of length to width from antennomere I to XI 4.1: 1.9: 3.3: 4.5: 4.4: 4.4: 5.1: 4.9: 5.5: 4.8: 6.0. Gonocoxae (Fig. 3J) slender, tightly conjunct from apical 1/3 to base; each gonocoxa with six or seven setae from apical 1/6 to apex, apex truncate. Ventrite VIII (Fig. 3G) weakly sclerotized medially, with several short setae at apex, and two rows of long setae at sides, spiculum extremely elongate. Spermathecal receptaculum (Fig. 3I) swollen; pump extremely slender and curved; sclerotized spermathecal duct slender. Bursa sclerites (Fig. 3H) elongate and apically tapering, with stout teeth along lateral margin at base; slightly curved in lateral view.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="27" type="food plant">
<paragraph pageId="8" pageNumber="27">Food plant.</paragraph>
<paragraph pageId="8" pageNumber="27">
<taxonomicName family="Betulaceae" lsidName="" pageId="8" pageNumber="27" rank="family">Betulaceae</taxonomicName>
:
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus formosana" order="Fagales" pageId="8" pageNumber="27" phylum="Tracheophyta" rank="species" species="formosana">Alnus formosana</taxonomicName>
(Burkill ex Forbes &amp; Hemsl.) Makino (
<bibRefCitation author="Lee, C-F" journalOrPublisher="Sishou-Hills Insect Observation Network and Taiwan Agricultural Research Institute, COA, New Taipei City and Taichung City" pageId="21" pageNumber="40" title="The Chrysomelidae of Taiwan 2." year="2010">Lee and Cheng 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="27" type="distribution">
<paragraph pageId="8" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="27">Taiwan, China.</paragraph>
</subSubSection>
</treatment>
</document>