155 lines
19 KiB
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155 lines
19 KiB
XML
<document id="74FCCA8A8912899F7B5DF339F07D164E" ID-DOI="10.5281/zenodo.197988" ID-GBIF-Dataset="a3004a0d-e2e8-4d06-9bda-d8f7fec9db2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="197988" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460283465716" checkinUser="plazi" docAuthor="Hippa, Heikki" docDate="2010" docId="494687A6AB5AFFA0FF3B51C177DEB0F7" docLanguage="en" docName="zt02618p060.pdf" docOrigin="Zootaxa 2618" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paramanota grandaeva Hippa, 2010, sp. n." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="B57FFFDEAB5DFFA8FFAC51567414B37D" masterDocTitle="Review of the genus Paramanota Tuomikoski (Diptera, Mycetophilidae), with the description of new fossil and recent species" masterLastPageNumber="60" masterPageNumber="47" pageNumber="54" updateTime="1698249876373" updateUser="plazi">
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<mods:title id="EE7C102A16EF8F839696D65D6D764280">Review of the genus Paramanota Tuomikoski (Diptera, Mycetophilidae), with the description of new fossil and recent species</mods:title>
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<mods:namePart id="875474E5FF6F0CA7C793E48E970006CC">Hippa, Heikki</mods:namePart>
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<mods:title id="62328518AF80CD91B9FBF6EF290177F5">Zootaxa</mods:title>
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<mods:part id="FB06CC5AB3D8F28A3C45976ED5D74023">
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<mods:date id="76719CC928C6A45C01FBBE60FED92A1E">2010</mods:date>
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<mods:detail id="252FEBFE84ABEE3EBE5ABC5C7466A1B8" type="volume">
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<mods:number id="DB55700E22C51BC81518E32DD9B4A9B4">2618</mods:number>
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<mods:identifier id="DFF2F64688177908F0D2EED8F6F95578" type="DOI">10.5281/zenodo.197988</mods:identifier>
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<mods:identifier id="EBDA69B96DD06C8849C5351A2A5D84CF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AC3E0D117FC014DD6EB3CCE96DD0BB9B" type="Zenodo-Dep">197988</mods:identifier>
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<treatment id="494687A6AB5AFFA0FF3B51C177DEB0F7" ID-DOI="http://doi.org/10.5281/zenodo.6205272" ID-GBIF-Taxon="119397436" ID-Zenodo-Dep="6205272" LSID="urn:lsid:plazi:treatment:494687A6AB5AFFA0FF3B51C177DEB0F7" httpUri="http://treatment.plazi.org/id/494687A6AB5AFFA0FF3B51C177DEB0F7" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
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<subSubSection id="89F5653BAB5AFFA0FF3B51C17598B291" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54" type="nomenclature">
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<paragraph id="C15036B0AB5AFFAFFF3B51C175E8B3CC" blockId="7.[151,508,151,212]" box="[151,508,151,177]" pageId="7" pageNumber="54">
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<heading id="9A1881DCAB5AFFAFFF3B51C175E8B3CC" bold="true" box="[151,508,151,177]" fontSize="11" level="1" pageId="7" pageNumber="54" reason="1">
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<emphasis id="F39BEAA2AB5AFFAFFF3B51C175E8B3CC" bold="true" box="[151,508,151,177]" pageId="7" pageNumber="54">
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<taxonomicName id="06EF4D33AB5AFFAFFF3B51C175A7B3CC" ID-CoL="8D6XH" box="[151,435,151,177]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="grandaeva" status="sp. nov.">
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<emphasis id="F39BEAA2AB5AFFAFFF3B51C175A7B3CC" bold="true" box="[151,435,151,177]" italics="true" pageId="7" pageNumber="54">Paramanota grandaeva</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E8A857D9AB5AFFAFFE1651C175E8B3CC" box="[442,508,151,177]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C15036B0AB5AFFAFFF3B51EC756EB3A9" blockId="7.[151,508,151,212]" box="[151,378,186,212]" pageId="7" pageNumber="54">
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<figureCitation id="59D42A35AB5AFFAFFF3B51EC74F4B3A9" box="[151,224,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,255,1909,1933]" captionTargetBox="[227,1370,898,1908]" captionTargetId="figure@2.[213,1370,853,1931]" captionTargetPageId="2" captionText="FIGURE 1. Wing, dorsal view (A and C – F) and ventral view (B). A, B. Paramanota grandaeva sp n. (holotype). C. P. orientalis Tuomikoski (Thailand). D, F. Paramanota furcillata sp. n. (D holotype, F paratype). E. P. bifalx sp. n. (holotype). Scale 1.0 mm." httpUri="https://zenodo.org/record/197989/files/figure.png" pageId="7" pageNumber="54">Figs 1</figureCitation>
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A, B, 4 A, B
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</paragraph>
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<caption id="95906638AB5AFFAFFF3B57A57170B456" httpUri="https://zenodo.org/record/197992/files/figure.png" pageId="7" pageNumber="54" targetBox="[175,1378,261,1757]" targetPageId="7">
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<paragraph id="C15036B0AB5AFFAFFF3B57A57170B456" blockId="7.[151,1435,1779,1835]" pageId="7" pageNumber="54">
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<emphasis id="F39BEAA2AB5AFFAFFF3B57A5750DB476" bold="true" box="[151,281,1779,1803]" pageId="7" pageNumber="54">FIGURE 4.</emphasis>
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<taxonomicName id="06EF4D33AB5AFFAFFE8D57A2760BB476" box="[289,543,1780,1803]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="grandaeva" status="sp. nov.">
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<emphasis id="F39BEAA2AB5AFFAFFE8D57A2760BB476" box="[289,543,1780,1803]" italics="true" pageId="7" pageNumber="54">Paramanota grandaeva</emphasis>
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</taxonomicName>
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<emphasis id="F39BEAA2AB5AFFAFFD8B57A57677B476" bold="true" box="[551,611,1779,1803]" pageId="7" pageNumber="54">
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<taxonomicNameLabel id="E8A857D9AB5AFFAFFD8B57A57677B476" box="[551,611,1779,1803]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(holotype).
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<emphasis id="F39BEAA2AB5AFFAFFD4457A576EFB476" bold="true" box="[744,763,1779,1803]" pageId="7" pageNumber="54">A</emphasis>
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. Head, oblique frontal view.
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<emphasis id="F39BEAA2AB5AFFAFFB9A57A5705CB476" bold="true" box="[1078,1096,1779,1803]" pageId="7" pageNumber="54">B</emphasis>
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. Basal part of abdomen, lateroventral view. Scale 1.0 mm. cx 3 = hind coxa, f 2 = middle femur, st 2–st 3 = sternites 2–3, tg 1–tg 4 = tergites 1–4.
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</paragraph>
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</caption>
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<paragraph id="C15036B0AB5AFFA0FF3B562270A9B2B7" blockId="7.[151,1436,1908,2038]" lastBlockId="8.[151,1437,152,906]" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
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Female. The single specimen is in amber which is partly milky so that many characters are invisible.
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<emphasis id="F39BEAA2AB5AFFAFFA9256227180B4F3" bold="true" box="[1342,1428,1908,1934]" pageId="7" pageNumber="54">Colour</emphasis>
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. Pale brown, face dark brown, finer setae and trichia pale brown, the thicker ones dark brown to black.
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<emphasis id="F39BEAA2AB5AFFAFFAFE56C07188B4CD" bold="true" box="[1362,1436,1942,1968]" pageId="7" pageNumber="54">Head,</emphasis>
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<figureCitation id="59D42A35AB5AFFAFFF3B56EC74C8B4A9" box="[151,220,1978,2004]" captionStart="FIGURE 4" captionStartId="7.[151,255,1779,1803]" captionTargetBox="[175,1378,261,1757]" captionTargetId="figure@7.[167,1434,237,1771]" captionTargetPageId="7" captionText="FIGURE 4. Paramanota grandaeva sp. n. (holotype). A. Head, oblique frontal view. B. Basal part of abdomen, lateroventral view. Scale 1.0 mm. cx 3 = hind coxa, f 2 = middle femur, st 2 – st 3 = sternites 2 – 3, tg 1 – tg 4 = tergites 1 – 4." httpUri="https://zenodo.org/record/197992/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
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A: medial part of eyes and the ocelli not visible, the curvature of the mesial margin of eye suggests the presence of an eye bridge. Antenna,
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<figureCitation id="59D42A35AB5AFFAFFDF3568A76BCB48B" box="[607,680,2012,2038]" captionStart="FIGURE 4" captionStartId="7.[151,255,1779,1803]" captionTargetBox="[175,1378,261,1757]" captionTargetId="figure@7.[167,1434,237,1771]" captionTargetPageId="7" captionText="FIGURE 4. Paramanota grandaeva sp. n. (holotype). A. Head, oblique frontal view. B. Basal part of abdomen, lateroventral view. Scale 1.0 mm. cx 3 = hind coxa, f 2 = middle femur, st 2 – st 3 = sternites 2 – 3, tg 1 – tg 4 = tergites 1 – 4." httpUri="https://zenodo.org/record/197992/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
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A: setae on scapus and pedicellus unusually long. Maxillary palpus,
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<figureCitation id="59D42A35AB55FFA0FF5051CE755CB3CF" box="[252,328,152,178]" captionStart="FIGURE 4" captionStartId="7.[151,255,1779,1803]" captionTargetBox="[175,1378,261,1757]" captionTargetId="figure@7.[167,1434,237,1771]" captionTargetPageId="7" captionText="FIGURE 4. Paramanota grandaeva sp. n. (holotype). A. Head, oblique frontal view. B. Basal part of abdomen, lateroventral view. Scale 1.0 mm. cx 3 = hind coxa, f 2 = middle femur, st 2 – st 3 = sternites 2 – 3, tg 1 – tg 4 = tergites 1 – 4." httpUri="https://zenodo.org/record/197992/files/figure.png" pageId="8" pageNumber="55">Fig. 4</figureCitation>
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A: basal segmentation fairly indistinguishable, no sensory pit on the antepenultimate palpomere, ultimate palpomere twice as long as the penultimate one. Number of strong postocular setae 7.
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<emphasis id="F39BEAA2AB55FFA0FF3B518B74E1B38A" bold="true" box="[151,245,221,247]" pageId="8" pageNumber="55">Thorax</emphasis>
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. Similar to fig. 8 a in
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<bibRefCitation id="A57E4B41AB55FFA0FDAF518B76FCB38A" box="[515,744,221,247]" pageId="8" pageNumber="55" refString="Hippa, H., Jaschhof, M. & Vilkamaa, P. (2004) Phylogeny of the Manotinae, with a review of Eumanota Edwards, Paramanota Tuomikoski and Promanota Tuomikoski (Diptera Mycetophilidae). Studia dipterologica, 11 (2), 405 - 428." type="journal article">
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Hippa
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<emphasis id="F39BEAA2AB55FFA0FDF8518B7684B38B" box="[596,656,221,246]" italics="true" pageId="8" pageNumber="55">et al.</emphasis>
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(2004)
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</bibRefCitation>
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except for prothoracic pleura which has longer setae, the longest ones being as long as the long postocular setae.
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<emphasis id="F39BEAA2AB55FFA0FCE051A9779DB264" bold="true" box="[844,905,255,281]" pageId="8" pageNumber="55">Legs</emphasis>
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. Front tibia is seen in ventral view and the characters apically on the prolateral side are not visible.
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<emphasis id="F39BEAA2AB55FFA0FC9C5074776FB241" bold="true" box="[816,891,290,316]" pageId="8" pageNumber="55">Wing,</emphasis>
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<figureCitation id="59D42A35AB55FFA0FC2F507477D8B241" box="[899,972,290,316]" captionStart="FIGURE 1" captionStartId="2.[151,255,1909,1933]" captionTargetBox="[227,1370,898,1908]" captionTargetId="figure@2.[213,1370,853,1931]" captionTargetPageId="2" captionText="FIGURE 1. Wing, dorsal view (A and C – F) and ventral view (B). A, B. Paramanota grandaeva sp n. (holotype). C. P. orientalis Tuomikoski (Thailand). D, F. Paramanota furcillata sp. n. (D holotype, F paratype). E. P. bifalx sp. n. (holotype). Scale 1.0 mm." httpUri="https://zenodo.org/record/197989/files/figure.png" pageId="8" pageNumber="55">Figs 1</figureCitation>
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A, B: Sc unusually strong, as strong as R1, Rs not observed with certainty. Wing length
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<quantity id="06179B55AB55FFA0FD6B50137731B222" box="[711,805,325,351]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="55" unit="mm" value="3.2">3.2 mm</quantity>
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.
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<emphasis id="F39BEAA2AB55FFA0FC83501377A4B222" bold="true" box="[815,944,325,351]" pageId="8" pageNumber="55">Abdomen,</emphasis>
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<figureCitation id="59D42A35AB55FFA0FC1B501377EFB222" box="[951,1019,325,351]" captionStart="FIGURE 4" captionStartId="7.[151,255,1779,1803]" captionTargetBox="[175,1378,261,1757]" captionTargetId="figure@7.[167,1434,237,1771]" captionTargetPageId="7" captionText="FIGURE 4. Paramanota grandaeva sp. n. (holotype). A. Head, oblique frontal view. B. Basal part of abdomen, lateroventral view. Scale 1.0 mm. cx 3 = hind coxa, f 2 = middle femur, st 2 – st 3 = sternites 2 – 3, tg 1 – tg 4 = tergites 1 – 4." httpUri="https://zenodo.org/record/197992/files/figure.png" pageId="8" pageNumber="55">Fig. 4</figureCitation>
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B: sternite 1 large, sloping posteroventrad, its posterior part pushed over the base of sternite 2, the medial part (anterior to the broken line in
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<figureCitation id="59D42A35AB55FFA0FAC1503C74B2B2DA" captionStart="FIGURE 4" captionStartId="7.[151,255,1779,1803]" captionTargetBox="[175,1378,261,1757]" captionTargetId="figure@7.[167,1434,237,1771]" captionTargetPageId="7" captionText="FIGURE 4. Paramanota grandaeva sp. n. (holotype). A. Head, oblique frontal view. B. Basal part of abdomen, lateroventral view. Scale 1.0 mm. cx 3 = hind coxa, f 2 = middle femur, st 2 – st 3 = sternites 2 – 3, tg 1 – tg 4 = tergites 1 – 4." httpUri="https://zenodo.org/record/197992/files/figure.png" pageId="8" pageNumber="55">Fig. 4</figureCitation>
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B) seems to be membranous but it is not fully visible because of the milkyness. Apical part of abdomen not very visible, similar to fig. 11 d in
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<bibRefCitation id="A57E4B41AB55FFA0FD8050E6771AB2B7" box="[556,782,432,458]" pageId="8" pageNumber="55" refString="Hippa, H., Jaschhof, M. & Vilkamaa, P. (2004) Phylogeny of the Manotinae, with a review of Eumanota Edwards, Paramanota Tuomikoski and Promanota Tuomikoski (Diptera Mycetophilidae). Studia dipterologica, 11 (2), 405 - 428." type="journal article">
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Hippa
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<emphasis id="F39BEAA2AB55FFA0FDD050E676A2B2B4" box="[636,694,432,457]" italics="true" pageId="8" pageNumber="55">et al.</emphasis>
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(2004)
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</bibRefCitation>
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, the apical cercomere slightly larger.
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</paragraph>
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<paragraph id="C15036B0AB55FFA0FF6A50847598B291" blockId="8.[151,1437,152,906]" box="[198,396,466,492]" pageId="8" pageNumber="55">Male. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="89F5653BAB55FFA0FF6A50AE7140B188" pageId="8" pageNumber="55" type="discussion">
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<paragraph id="C15036B0AB55FFA0FF6A50AE7140B188" blockId="8.[151,1437,152,906]" pageId="8" pageNumber="55">
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<emphasis id="F39BEAA2AB55FFA0FF6A50AE7546B16F" bold="true" box="[198,338,504,530]" pageId="8" pageNumber="55">Discussion.</emphasis>
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Three of the key characters of
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<taxonomicName id="06EF4D33AB55FFA0FD1250AE7744B16C" box="[702,848,504,529]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F39BEAA2AB55FFA0FD1250AE7744B16C" box="[702,848,504,529]" italics="true" pageId="8" pageNumber="55">Paramanota</emphasis>
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</taxonomicName>
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cannot be seen in the
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<typeStatus id="1E548812AB55FFA0FBF950AE70AFB16F" box="[1109,1211,504,530]" pageId="8" pageNumber="55" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="06EF4D33AB55FFA0FB4F50AE7169B16C" box="[1251,1405,504,529]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="grandaeva">
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<emphasis id="F39BEAA2AB55FFA0FB4F50AE7169B16C" box="[1251,1405,504,529]" italics="true" pageId="8" pageNumber="55">P. grandaeva</emphasis>
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</taxonomicName>
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: i. e. the complete eye bridge, medially divided anterior ocellus and the transverse comb-like row of strong setae prolaterally at the apex of front tibia. The two first characters are obscured by the milkyness of the amber, the latter because both of the front tibiae are visible in ventral aspect. Otherwise the species is quite similar to the recent
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<taxonomicName id="06EF4D33AB55FFA0FF4553D87594B1DA" box="[233,384,654,679]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F39BEAA2AB55FFA0FF4553D87594B1DA" box="[233,384,654,679]" italics="true" pageId="8" pageNumber="55">Paramanota</emphasis>
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</taxonomicName>
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and does not differ more than can be expected between closely related species. When compared with all the recent
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<emphasis id="F39BEAA2AB55FFA0FE5F53E57726B1B1" box="[499,818,691,716]" italics="true" pageId="8" pageNumber="55">
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<taxonomicName id="06EF4D33AB55FFA0FE5F53E5769EB1B1" box="[499,650,691,716]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">Paramanota</taxonomicName>
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,
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<taxonomicName id="06EF4D33AB55FFA0FD3A53E57726B1B1" box="[662,818,691,716]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="grandaeva">P. grandaeva</taxonomicName>
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</emphasis>
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has h and Sc more strongly sclerotized, equal to R1 instead of being conspicuously weaker, and Sc is slightly longer. This is supposedly true for both sexes.
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</paragraph>
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</subSubSection>
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<subSubSection id="89F5653BAB55FFA0FF6A525677DEB0F7" pageId="8" pageNumber="55" type="etymology">
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<paragraph id="C15036B0AB55FFA0FF6A525674F2B042" blockId="8.[151,1437,152,906]" pageId="8" pageNumber="55">
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<emphasis id="F39BEAA2AB55FFA0FF6A52567547B067" bold="true" box="[198,339,768,794]" pageId="8" pageNumber="55">Etymology.</emphasis>
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The name is Latin,
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<taxonomicName id="06EF4D33AB55FFA0FDE3525776D8B067" box="[591,716,769,794]" class="Insecta" family="Mycetophilidae" genus="Paramanota" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="grandaeva">
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<emphasis id="F39BEAA2AB55FFA0FDE3525776D8B067" box="[591,716,769,794]" italics="true" pageId="8" pageNumber="55">grandaeva</emphasis>
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</taxonomicName>
|
||
, ancient, referring to the discovery of the species in Baltic amber.
|
||
</paragraph>
|
||
<paragraph id="C15036B0AB55FFA0FF6A521C77DEB0F7" blockId="8.[151,1437,152,906]" pageId="8" pageNumber="55">
|
||
<emphasis id="F39BEAA2AB55FFA0FF6A521C7500B019" bold="true" box="[198,276,842,868]" pageId="8" pageNumber="55">
|
||
<typeStatus id="1E548812AB55FFA0FF6A521C7519B019" box="[198,269,842,868]" pageId="8" pageNumber="55">Types</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="F39BEAA2AB55FFA0FEB2521D759DB019" box="[286,393,843,868]" italics="true" pageId="8" pageNumber="55">
|
||
<typeStatus id="1E548812AB55FFA0FEB2521D759DB019" box="[286,393,843,868]" pageId="8" pageNumber="55" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
. Female, Baltic amber; Eocene: Lutetian;
|
||
<collectingCountry id="B9F87620AB55FFA0FCD0521D77C3B018" box="[892,983,843,869]" name="Zambia" pageId="8" pageNumber="55">Sambia</collectingCountry>
|
||
, former East Prussia; ex coll. Albertus University Königsberg. Labelled GZG.BST.02724 (G4393) (in GZG).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |