545 lines
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545 lines
72 KiB
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<document ID-CLB-Dataset="51924" ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-GBIF-Dataset="c2e00414-9633-4890-ab72-cb2aa3e9d174" ID-PMC="PMC10208988" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-PubMed="37250203" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="03122B5FADC55EFE8BC5CF4838D12522" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma housei" docType="treatment" docVersion="5" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1732652259105" updateUser="ExternalLinkService">
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<mods:title id="4CC6C200FE712C120E6507FAB71693CB">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:namePart id="0BB9A1FFB52E83A06FC09F45ED22D853">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="D077DDE1EA91B0150CBD18A145C79488">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="33D0BFDAA3C15F990E50FF1AB0291EAD">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="4A4F8ED08E01E01A7B980CA21215487C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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<mods:namePart id="22CF55124D1AB3084849DB21E5A774F0">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="16ED7FE862044C70251DE9D0F7003161">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:namePart id="DBC6E1627CEB16B231A6976DF24CFC56">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="CBD21506C33AC49D98BFBAF19C99D408">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="5F6885A7E84C6E8A1C72ECB30796323E">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="8855ED3403269C2D234EE19DB9CE765C">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="FF3934F045FCC7D6D834B116302CB9EF">Panha, Somsak</mods:namePart>
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<mods:affiliation id="F289821E144ADA5C569D4A94BB5E80CF">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="5316E7CDB2BA054A8AA67ABFB0981281" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="A7F372884727FAB635EFA93F284D3113">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="D5991FF6154A5E9A435ABF5E7EB596B4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="CD82D373380C21E7BD6506576F6D2E22" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:date id="BC5B70FF8C81BDA3E7D2BE2A326E0183">2023</mods:date>
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<mods:number id="13A1456BCED1073C30F665EA603AB1E0">1142</mods:number>
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<mods:identifier id="45C709C1AD2B769BB8E6EB3A606E4586" type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
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<treatment id="03122B5FADC55EFE8BC5CF4838D12522" ID-GBIF-Taxon="209102274" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522" httpUri="http://treatment.plazi.org/id/03122B5FADC55EFE8BC5CF4838D12522" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
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<subSubSection id="4AEFC8C7593C722BFAF9880A62F8C599" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="E222EACBE4EA02180E2444748DA61746" pageId="0" pageNumber="1">
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4.
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<taxonomicName id="E36C7E76A911A4D47C71D50F9BA23759" ID-CoL="7SZX2" LSID="03122B5F-ADC5-5EFE-8BC5-CF4838D12522" authority="(Haines, 1855)" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei (Haines, 1855)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="36D0A534C2FE908EFA1BBAAD6CF81371" pageId="0" pageNumber="1" type="description">
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<paragraph id="502A3821C0BCAAE6FB8FB552F47A9A34" pageId="0" pageNumber="1">
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<figureCitation id="06426C839D8D2D40EF1A3A4A8039E14A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)." figureDoi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1">Figs 17A</figureCitation>
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<figureCitation id="EDBFBE11A3285043E115077DB22F0ECE" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">, 18</figureCitation>
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<figureCitation id="4EDCB25D3F1345F4780C119944060091" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">, 19</figureCitation>
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<figureCitation id="0D0F253064D5B638A77509FB783B661F" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawing of general anatomy of Rhiostoma housei specimen CUMZ 3982 from Saraburi Province, Thailand. A female B male. Abbreviations: bc, bursa copulatrix;; cm, columellar muscle; ct, cephalic tentacle; dg, digestive gland; es, eyespot; f, foot; fc, faeces; int, intestine; lc, lung cavity; mc, mantle collar; od, oviduct; op, operculum; ov, ovary; p, external penis; pg, prostate gland; sg, sperm groove; sn, snout; te, testis; ut, uterus; vd, vas deferens; vg, vaginal groove." figureDoi="10.3897/zookeys.1142.90097.figure20" httpUri="https://binary.pensoft.net/fig/799878" pageId="0" pageNumber="1">, 20</figureCitation>
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<figureCitation id="3507F58B2E14BE1AE62144F294892261" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">, 32A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="CD4463CD2E346DA229A266C46215DD1D" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="B10FC32573B5757618A309855301D9DE" pageId="0" pageNumber="1">
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||
<taxonomicName id="7D737CAB5FBDDB4231CAA5B968E02FC5" authorityName="Haines" authorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Cyclostoma housei</taxonomicName>
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Haines, 1855: 157, pl. 5, figs 12-15. Type locality: Siam [= Thailand].
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<bibRefCitation id="38B1564DE57BC91373E257F9428E1924" DOI="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" author="Boyko, CB" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 158" refId="B24" refString="Boyko, CB, Cordeiro, JR, 2001. Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1 - 158, DOI: https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" title="Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia)." url="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" volume="262" year="2001">Boyko and Cordeiro 2001</bibRefCitation>
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: 52.
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</paragraph>
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<paragraph id="4B9CE667B9AE3AA405EA5EFDCA800F51" pageId="0" pageNumber="1">
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<taxonomicName id="62733DE4312635F4A0004C58D3F2DADC" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Pterocyclos housei</taxonomicName>
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-
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<bibRefCitation id="F00D9103110157075563D028C818B9B5" DOI="https://doi.org/10.5962/bhl.title.11217" author="Pfeiffer, L" journalOrPublisher="Supplementum primum, Volume 2. Cassellis: Theodori Fischeri" pageId="0" pageNumber="1" refId="B133" refString="Pfeiffer, L, 1858. Monographia Pneumonopomorum viventium. Supplementum primum, Volume 2. Cassellis: Theodori Fischeri, DOI: https://doi.org/10.5962/bhl.title.11217" title="Monographia Pneumonopomorum viventium." url="https://doi.org/10.5962/bhl.title.11217" year="1858">Pfeiffer 1858</bibRefCitation>
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: 29.
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<bibRefCitation id="846E6B15D03F5AC30838DAFC61127841" author="Reeve, LA" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" refId="B145" refString="Reeve, LA, 1863. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 14, Pterocyclos, plates 1-5. Lovell Reeve and Co., London." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 14, Pterocyclos, plates 1 - 5. Lovell Reeve and Co., London." year="1863">Reeve 1863</bibRefCitation>
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:
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<taxonomicName id="8C7D5BCA1D793EFACB02BCA61FE64D44" authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pterocyclos</taxonomicName>
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pl. 4, species 21.
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</paragraph>
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<paragraph id="EBEF949C8B00BAEEA466D22531B952AA" pageId="0" pageNumber="1">
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<taxonomicName id="BF776190D64872CBA7CCD19186A55CC1" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei</taxonomicName>
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-
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<bibRefCitation id="25D2BD61737A15DEF0A316A04342F27C" DOI="https://doi.org/10.1080/00222936008697183" author="Benson, WH" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" refId="B14" refString="Benson, WH, 1860. On Clostophis and Rhiostoma, new Burmese genera of land shells. Annals and Magazine of Natural History, Series 3 5(26): 95-97. https://doi.org/10.1080/00222936008697183" title="On Clostophis and Rhiostoma, new Burmese genera of land shells. Annals and Magazine of Natural History, Series 3 5 (26): 95 - 97." url="https://doi.org/10.1080/00222936008697183" year="1860">Benson 1860</bibRefCitation>
|
||
: 97.
|
||
<bibRefCitation id="B4337A97210FA93AA8FEF0D1DAD147ED" author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="115 - 117" refId="B134" refString="Pfeiffer, L, 1862a. Description of eight new species of Cyclostomacea from the collection of H. Cumming. Esq. Proceedings of the Zoological Society of London 30: 115 - 117" title="Description of eight new species of Cyclostomacea from the collection of H. Cumming. Esq." volume="30" year="1862 a">Pfeiffer 1862a</bibRefCitation>
|
||
: 117, pl. 12, fig. 9.
|
||
<bibRefCitation id="55374CB96F626C6C5A1DF1B46DEADC95" DOI="https://doi.org/10.5962/bhl.title.10621" author="Pfeiffer, L" journalOrPublisher="Supplementum Secundum, Volume 3. Cassellis: Theodori Fischeri" pageId="0" pageNumber="1" refId="B136" refString="Pfeiffer, L, 1865. Monographia Pneumonopomorum viventium. Supplementum Secundum, Volume 3. Cassellis: Theodori Fischeri, DOI: https://doi.org/10.5962/bhl.title.10621" title="Monographia Pneumonopomorum viventium." url="https://doi.org/10.5962/bhl.title.10621" year="1865">Pfeiffer 1865</bibRefCitation>
|
||
: 38.
|
||
<bibRefCitation id="6B14DC4F824E00542410E25A9958EB00" author="Martens, E von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B91" refString="Martens, E von, 1867. Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." title="Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." year="1867">Martens 1867</bibRefCitation>
|
||
: 63.
|
||
<bibRefCitation id="CF71AB300D57368EF12F673FAC07D8D8" DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Extrait du Bulletin de la Societe ďHistoire Naturelle ďAutun" pageId="0" pageNumber="1" pagination="1 - 193" refId="B40" refString="Fischer, P, 1891. Catalogue et distribution geographique des mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Extrait du Bulletin de la Societe ďHistoire Naturelle ďAutun 4: 1 - 193, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" volume="4" year="1891">Fischer 1891</bibRefCitation>
|
||
: 101.
|
||
<bibRefCitation id="AC4D5B4B7844647D82574E096A4FDB6B" author="Moellendorff, OF von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="146 - 156" refId="B98" refString="Moellendorff, OF von, 1894. On a collection of land-shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London 62: 146 - 156" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." volume="62" year="1894">
|
||
<normalizedToken id="D29B0D05B3FE01EA435B9CAF0FA1837E" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
1894
|
||
</bibRefCitation>
|
||
: 152. Kobelt and
|
||
<normalizedToken id="A073973CD7AC554536766591C1C2E80F" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
1897: 115.
|
||
<bibRefCitation id="19424715403393685CE7A1C7911E5D38" author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 663" refId="B71" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 663" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
|
||
: 178.
|
||
<bibRefCitation id="946D9EAFB31C876BA3D1907CC653CECF" author="Fischer, H" editor="Pavie, A" journalOrPublisher="Ernest Leroux, Editeur, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B42" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour. In: Pavie, A, Ed., Mission Pavie, Indo-Chine 1879-1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III. Ernest Leroux, Editeur, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour." volumeTitle="Mission Pavie, Indo-Chine 1879 - 1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
|
||
: 427. Kobelt 1911: 757, 758, pl. 110, figs 8-10, pl. 113, fig. 2.
|
||
<bibRefCitation id="5474C62C1423620BEF6609DE578ECC8D" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
|
||
: 174.
|
||
<bibRefCitation id="DF72227FEF4C98AB49C4449025566A04" author="Rees, WJ" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="55 - 67" refId="B144" refString="Rees, WJ, 1964. A review of breathing devices in land operculated snail. Proceedings of the Malacological Society of London 36: 55 - 67" title="A review of breathing devices in land operculated snail." volume="36" year="1964">Rees 1964</bibRefCitation>
|
||
: 67, pl. 4, fig. 15.
|
||
<bibRefCitation id="48CEDBFBE35F12449000044C5C64030A" author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
||
: 11.
|
||
<bibRefCitation id="F6305E72710C83099A7490B32B5629CC" author="Fischer, P-H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B41" refString="Fischer, P-H, 1973. Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches." volume="90" year="1973">Fischer 1973</bibRefCitation>
|
||
: 47.
|
||
<bibRefCitation id="D22D88606EA987E4B8CBC9C977382861" author="Abbott, RT" journalOrPublisher=", Melbourne, Florida" pageId="0" pageNumber="1" refId="B1" refString="Abbott, RT, 1989. Compendium of Landshells. American Malacologists Inc., Melbourne, Florida" title="Compendium of Landshells. American Malacologists Inc." year="1989">Abbott 1989</bibRefCitation>
|
||
: 29, 42, with 3 figures.
|
||
<bibRefCitation id="61578B9C981596F140BCAC47AD9413AF" author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan 2001</bibRefCitation>
|
||
: 41-46, figs 4.10-4.12.
|
||
<bibRefCitation id="44BE7B8D529295702885E54C37009CA3" author="Kongim, B" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="13 - 20" refId="B78" refString="Kongim, B, Sutcharit, C, Tongkerd, P, Panha, S, 2013b. Karyotype of the snorkel snails genera Pterocyclos and Rhiostoma (Prosobranchia: Cyclophoridae). The Raffles Bulletin of Zoology 61: 13 - 20" title="Karyotype of the snorkel snails genera Pterocyclos and Rhiostoma (Prosobranchia: Cyclophoridae)." volume="61" year="2013 b">Kongim et al. 2013b</bibRefCitation>
|
||
: 16, fig. 2f, g.
|
||
</paragraph>
|
||
<paragraph id="532565E824ACC6BDE891CA1EB952A416" pageId="0" pageNumber="1">
|
||
<taxonomicName id="98619C4D927ED35429EC25F44CF814D1" class="Gastropoda" family="Cyclophoridae" genus="Rhinostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhinostoma</taxonomicName>
|
||
[sic]
|
||
<taxonomicName id="6C29ED99326FD87C916E1177AE7F3C1D" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhinostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhinostoma housei</taxonomicName>
|
||
-
|
||
<bibRefCitation id="1F5BC41127284F5B74A2C7319E4CFD24" author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" pagination="111 - 128" refId="B53" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia, Kyoto 4: 111 - 128" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe 1965</bibRefCitation>
|
||
: 128 (in part), pl. 2, fig. 10.
|
||
</paragraph>
|
||
<paragraph id="1A96DFABAFA0F2C7CBEE6BDBD2ED6458" pageId="0" pageNumber="1">
|
||
<taxonomicName id="3BDAF19DC1B55D6A9B497E16699FA1A0" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos (Spiraculum) housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei" subGenus="Spiraculum">Pterocyclos (Spiraculum) housei</taxonomicName>
|
||
-
|
||
<bibRefCitation id="35C0A1FA47ADE50B27A4BDD9B5CFF24D" author="Martens, E von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="6 - 18" refId="B90" refString="Martens, E von, 1860. On the Mollusca of Siam. Proceedings of the Zoological Society of London 28: 6 - 18" title="On the Mollusca of Siam." volume="28" year="1860">Martens 1860</bibRefCitation>
|
||
: 10.
|
||
</paragraph>
|
||
<paragraph id="3C07117B32A1DB100B6237335B538A52" pageId="0" pageNumber="1">
|
||
<taxonomicName id="9FC892851198E9796477294B97C4BB60" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterocyclus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pterocyclus</taxonomicName>
|
||
[sic] (
|
||
<taxonomicName id="4B2B7FFA0A1944E65B43F630429B3D34" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
|
||
)
|
||
<taxonomicName id="2A65A5824FDE92A4721A4E98EC0F697F" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei</taxonomicName>
|
||
-
|
||
<bibRefCitation id="2A8F258B647C775154F7B0DCC6765239" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Calcutta, Trustees" pageId="0" pageNumber="1" refId="B114" refString="Nevill, G, 1878. Hand list of Mollusca in the Indian Museum, Calcutta. Calcutta, Trustees, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
|
||
: 263.
|
||
</paragraph>
|
||
<paragraph id="C4F0CC57087274D57FB70BF330FBCF39" pageId="0" pageNumber="1">
|
||
<taxonomicName id="57229AB54FEA6D92D1C654BEBC8FAE75" authorityName="Haines" authorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma (Pterocyclos) housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei" subGenus="Pterocyclos">Cyclostoma (Pterocyclos) housei</taxonomicName>
|
||
-
|
||
<bibRefCitation id="FD50D30B27A8E3C5E97C9A33BCCFA1B1" DOI="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" author="Boyko, CB" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 158" refId="B24" refString="Boyko, CB, Cordeiro, JR, 2001. Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1 - 158, DOI: https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" title="Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia)." url="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" volume="262" year="2001">Boyko and Cordeiro 2001</bibRefCitation>
|
||
: 52.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC7DA02DA3D1874507CC7A9F8288C1BB" pageId="0" pageNumber="1" type="type material">
|
||
<paragraph id="57A0D702D83CC89C2F0FC407B36B20FD" pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph id="8A4E59B58BF29204132596A8AAE68802" pageId="0" pageNumber="1">
|
||
This species was described from specimens from the Haines collection (AMNH). The unique name-bearing type was not designated; however, at least two specimens were illustrated in the original description. The specimen identical to the illustration (shape of breathing device) and measurements in
|
||
<bibRefCitation id="918DE16FCB44503C1079ACD42750FDCC" DOI="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" author="Haines, WA" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" refId="B54" refString="Haines, WA, 1855. Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158. https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" title="Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158." url="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" year="1855">Haines (1855</bibRefCitation>
|
||
: figs 12, 13) is figured herein,
|
||
<emphasis id="3B37E6611E3DCAE144F68C9B70C540A5" bold="true" italics="true" pageId="0" pageNumber="1">syntype</emphasis>
|
||
AMNH 42923 (5 shells; Fig.
|
||
<figureCitation id="6C98E49CA31FE7464BB9E63B9276D311" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19A, B</figureCitation>
|
||
) from Siam [Thailand].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19BC2A9D6195337D225690AD06F61349" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="9724AB0E9A49FE3E583ECDA356CE80E1" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
||
<paragraph id="A01B43467CA18E7E553EFF7CDC1F9D16" pageId="0" pageNumber="1">
|
||
<materialsCitation id="55C5C26746F2B840125297BEC57AE41B" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" county="East Siam" location="East Siam" municipality="East Siam" specimenCount="1" stateProvince="Saraburi">
|
||
<emphasis id="34C240D135C48FE73D115E449EAEB4D0" bold="true" pageId="0" pageNumber="1">
|
||
<collectingCountry id="F948459746F947B0BE508D3108088F19" name="Thailand">Thailand</collectingCountry>
|
||
</emphasis>
|
||
: Boong Tuey,
|
||
<collectingCounty id="782AF1FB3EBECC356D543FD23D713B48">East Siam</collectingCounty>
|
||
: RBINS 659990, 659991; Pak Chong,
|
||
<collectingMunicipality id="36DE507703C5F8D2838BDC390DCF5DEF">East Siam</collectingMunicipality>
|
||
: RBINS 659978, 659979, 659980; Lam Ton Lang,
|
||
<location id="09A020DC20452C1AB7A3BDD344678012" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:09A020DC20452C1AB7A3BDD344678012" country="Thailand" county="East Siam" municipality="East Siam" name="East Siam" stateProvince="Saraburi">East Siam</location>
|
||
: RBINS 659982, 659983, 659984. Siam: NHMW 75000/E/23126 (1 shell). Samui, Siam: NHMW Rusnov (1 shell). Tham Phet Tham Thong, Takhli, Nakhonsawan: CUMZ 4385. Pukae Botanical Garden, Chaloem Phra Kiat,
|
||
<collectingRegion id="B6610FAFDEF2B67D322F8597D674738D" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4742. Wat Tham Srivilai, Chaloem Phra Kiat,
|
||
<collectingRegion id="9617A98D19875D2838F20773B2EB4D42" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 3982 (Figs
|
||
<figureCitation id="CE445DE56609E32417F941F7EEB161C8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-C Internal anatomy of Rhiostoma housei specimen CUMZ 3982 from Tham Sriwilai, Saraburi, A inside view of lung cavity shows an arrangement of the kidney, circulatory system and the terminal part of the digestive tract B male genitalia and C female genitalia D spermatophore of Rhiostoma samuiense specimen CUMZ 4713 from Kanchanadit, Surat Thani with its cross sections showing spermatophore shape in each position. Abbreviations: an, anus; bc, bursa copulatrix; fc, faeces; go, genital opening; h, heart; kd, kidney; mc, mantle collar; od, oviduct; pg, prostate gland; re, rectum; sr, seminal receptacle; ut, uterus; v, vein; vd, vas deferens." figureDoi="10.3897/zookeys.1142.90097.figure7" httpUri="https://binary.pensoft.net/fig/799865" pageId="0" pageNumber="1">7A-C</figureCitation>
|
||
,
|
||
<figureCitation id="41041582FA4282D07ECB5EE8AC0DB2DE" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19D</figureCitation>
|
||
,
|
||
<figureCitation id="364C877D43311DF68A14183F2787812E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawing of general anatomy of Rhiostoma housei specimen CUMZ 3982 from Saraburi Province, Thailand. A female B male. Abbreviations: bc, bursa copulatrix;; cm, columellar muscle; ct, cephalic tentacle; dg, digestive gland; es, eyespot; f, foot; fc, faeces; int, intestine; lc, lung cavity; mc, mantle collar; od, oviduct; op, operculum; ov, ovary; p, external penis; pg, prostate gland; sg, sperm groove; sn, snout; te, testis; ut, uterus; vd, vas deferens; vg, vaginal groove." figureDoi="10.3897/zookeys.1142.90097.figure20" httpUri="https://binary.pensoft.net/fig/799878" pageId="0" pageNumber="1">20</figureCitation>
|
||
,
|
||
<figureCitation id="154C16225DBC7C4D5646D1D40791DEAC" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32A</figureCitation>
|
||
), 4449. Kaeng Khoi,
|
||
<collectingRegion id="256F5DC206CB35E98291CF58F9B1B43B" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4315.
|
||
<location id="481EF7EA020CEF01EC7556F3C157DBA8" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:481EF7EA020CEF01EC7556F3C157DBA8" country="Thailand" county="East Siam" municipality="East Siam" name="Ban Tha Sao Village" stateProvince="Saraburi">Ban Tha Sao Village</location>
|
||
, Kaeng Khoi,
|
||
<collectingRegion id="D81835B5559185B45103B828C018F2A0" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 10135. Wat Tham Phra Phothisat, Kaeng Khoi,
|
||
<collectingRegion id="776686224B655D06F33EC4BD13A4E1F4" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 10026, 10158. Chet Sao Noi Waterfall, Muaklek,
|
||
<collectingRegion id="47AB1B0B71A88FC6B155532948664575" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4732. Muaklek,
|
||
<collectingRegion id="5C07D852997F387E7EF862186025304C" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4390, 4460. Wat Tham Dao Khao Kaew, Muaklek,
|
||
<collectingRegion id="4D0B5B2239AB41AF6AB8DBF06C90B2D2" country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 3802, 3934, 4389, 4760, 4842, 4897, 10131, 10133. Khao Sompod, Chai Ba Dan,
|
||
<collectingRegion id="0CAAD172B98C5B116AAAD8BA22E5697E" country="Thailand" name="Lop Buri">Lopburi</collectingRegion>
|
||
: CUMZ 3932, 3936, 3937. Wat Tham Promlok, Chai Ba Dan,
|
||
<collectingRegion id="A027CE3E94F7D402D9542D0A73BB8F42" country="Thailand" name="Lop Buri">Lopburi</collectingRegion>
|
||
: CUMZ 10128. Sub Langka, Lamsonthi,
|
||
<collectingRegion id="1F235A5B64980F72AFFD695FDDEDE45F" country="Thailand" name="Lop Buri">Lopburi</collectingRegion>
|
||
: CUMZ 4736. Khao Look Chang, Pak Chong,
|
||
<collectingRegion id="74F1E2F0BF0BA90F4DC8A2B15FCC1200" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 3805, 3981, 4316, 4318, 4320, 4379, 4384 (Fig.
|
||
<figureCitation id="A952C043FEB75B9608F857D65EB09A4B" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19E</figureCitation>
|
||
), 4454. The way to Khao Yai, Pak Chong,
|
||
<collectingRegion id="07BDEFFF96EBEE8E6541FA345E163218" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 4454. Wat Tham Prommachan Thammaram, Pak Chong,
|
||
<collectingRegion id="974EBE883272033F7C2CDF779451CA62" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10160. Wat Tham Sub Muet, Pak Chong,
|
||
<collectingRegion id="5B5BF3291978DF4F43CC756E2695D87F" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10132. Wat Tham Thep Nimit, Pak Chong,
|
||
<collectingRegion id="A7B39B5A5EED5F72F1F8580C38E18478" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10126. Wat Thepphitak Punnaram, Pak Chong,
|
||
<collectingRegion id="22C89C897B0AC4CBADA73D78047DAE3C" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 4321, 4416. Sakaerat Biosphere, Pak Thong Chai,
|
||
<collectingRegion id="E95681D583E7584D13CE9310400BD1DA" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 4391.
|
||
<location id="181AF75D22508CB0BB8E8430244AE702" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:181AF75D22508CB0BB8E8430244AE702" country="Thailand" county="East Siam" municipality="East Siam" name="Sakaerat Environmental Research Station" stateProvince="Saraburi">Sakaerat Environmental Research Station</location>
|
||
, Wang Nam Khiao,
|
||
<collectingRegion id="D01E5217F2335FF1D3B743B0D34DAD88" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10134. Ban Bueng, Ban Bueng,
|
||
<collectingRegion id="5311B97548273779642E4A003BBFABB5" country="Thailand" name="Chon Buri">Chonburi</collectingRegion>
|
||
: CUMZ 4312. Khao Chakan, Srakeo: CUMZ 10156. Khao Nang Panturat, Cha Am,
|
||
<collectingRegion id="A7545AC9E109299FDD358EBBBB8052BB" country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 4879 (Fig.
|
||
<figureCitation id="D193FBC77D4A38C4E5B207FB2D9D8D09" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19F</figureCitation>
|
||
), 10129, 10130, 10164, 10165.
|
||
<location id="0CAE6503CB067FBB1C0B54954B8704AE" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:0CAE6503CB067FBB1C0B54954B8704AE" country="Thailand" county="East Siam" municipality="East Siam" name="Khao Nang Panturat Nature Reserve" stateProvince="Saraburi">Khao Nang Panturat Nature Reserve</location>
|
||
, Cha Am,
|
||
<collectingRegion id="D3E78E7BE7CB99DB1403D8316840E2FE" country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 4480.
|
||
<location id="4A21BB501F97840176D365B9116B9D6F" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:4A21BB501F97840176D365B9116B9D6F" country="Thailand" county="East Siam" municipality="East Siam" name="Tham Na Kwang" stateProvince="Saraburi">Tham Na Kwang</location>
|
||
, Cha Am,
|
||
<collectingRegion id="AAF82DA1F39D2A3431973D71741CDB73" country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 3988, 4452, 4719, 10162. 30 km. before
|
||
<location id="865DDE916C2BFBDC42304B33904FE650" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:865DDE916C2BFBDC42304B33904FE650" country="Thailand" county="East Siam" municipality="East Siam" name="Pa La Au Waterfall" stateProvince="Saraburi">Pa La Au Waterfall</location>
|
||
, Hua Hin,
|
||
<collectingRegion id="FC21498A1181CBB81E1DA4E6969D53E7" country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 3827, 4740. Kaeng Kachan,
|
||
<collectingRegion id="5E04D9081236139245DB53525051FB88" country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 4409, 4412. Khao Lom Muak, Muang,
|
||
<collectingRegion id="7C22760DCC0AC1DA6E8B756FDFA7117E" country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 3870, 3987 (Fig.
|
||
<figureCitation id="0CFD7E1E875262E7AAD7653ADFA7C257" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)." figureDoi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1">17A</figureCitation>
|
||
), 4322, 10163. Bang Poo, Sam Roi Yod,
|
||
<collectingRegion id="EBD73F83A05D22988E2FDB9B220DDC9F" country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 4414, 4417, 4429, 4430.
|
||
<location id="F42B3B25FEEB327418C3E6483928E409" LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:F42B3B25FEEB327418C3E6483928E409" country="Thailand" county="East Siam" municipality="East Siam" name="Saun Wiweg Bureau of Monks" stateProvince="Saraburi">Saun Wiweg Bureau of Monks</location>
|
||
, Sam Roi Yod,
|
||
<collectingRegion id="29CDD471FCFED49AD6DCD4694871FE82" country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 10127. Ao Noi,
|
||
<collectingRegion id="875C26BCC1998D0A75062FE93AA47D81" country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 3868. Tham Pissadarn, Tha Sae,
|
||
<collectingRegion id="7D9F953A69D7A1EBD041FB0AC14F8871" country="Thailand" name="Chumphon">Chumphon</collectingRegion>
|
||
: CUMZ 4383 (Fig.
|
||
<figureCitation id="C2D2F1294345C8E82193243EEBEF95B5" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19C</figureCitation>
|
||
), 4453, 4851
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="780FEBFC96ACD8EB2E9B3BCA98BD1A58" doi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1" start="Figure 17" startId="F17">
|
||
<paragraph id="BF170C7AC3732EA485F78C14AA8F8E83" pageId="0" pageNumber="1">
|
||
<emphasis id="584CC66BF2AC4B6BA56802FDBB754564" bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
|
||
Live snails of the species group II
|
||
<emphasis id="771257226CCD66A1357580888C5437AB" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName id="C1B7EAB0D62E8B9EED622297864295D5" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis id="DF10682E75DD0F3CCC11A7D7E08F8D1A" italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 3987 from Ao Manow,
|
||
<collectingRegion id="124755D59D0180F1788C6F6E1269C61C" country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
<emphasis id="FE43EDFF2A767A0C5134555F31C3AAFD" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
|
||
<taxonomicName id="C877D82AA04FB7C745FDD963E3BA003B" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||
<emphasis id="4A4B663A61603CAF9EB728F2E70CD327" italics="true" pageId="0" pageNumber="1">Rhiostoma hainesi</emphasis>
|
||
</taxonomicName>
|
||
from Khao Soi Dao,
|
||
<collectingRegion id="B85CB9E64C38BB4F4C93E981285E79C3" country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
<emphasis id="D592DF177DF02C4D97C393B5A171B41B" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
without breathing device and
|
||
<emphasis id="0609120BD63E628281E4431692EE99C9" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
with long breathing device
|
||
<emphasis id="85A0FD74AEBB7F2EC130CBC0C719108C" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName id="51DCC540ACFF3CF26EC998FBCFB62B2F" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma jalorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jalorensis">
|
||
<emphasis id="C6B5FAF997FF90DCC5C2A1909060AA5F" italics="true" pageId="0" pageNumber="1">Rhiostoma jalorensis</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 3994 from Wat Tham Sue,
|
||
<collectingRegion id="5D824020D888FA71CF7E73C16060E53B" country="Thailand" name="Krabi">Krabi</collectingRegion>
|
||
. All figures are not to scale. Photographs: P. Tongkerd (
|
||
<emphasis id="D0098132B7EEAB058AA5DFBC65D51E9C" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="43800A87D76D7E4D5CC57E1E1F7E4B1C" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="EB051FD5FC9224C55F0B507C12F8C191" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="C9EA6C10161BB24237B1488A54C15AE0" pageId="0" pageNumber="1">Shell medium to large, thick, and depressed. Detached whorl longer than aperture width, curved, and descending. Breathing device straight with complete tube. Shell usually with dark brown zigzag patterns.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EC4756DC1131480BC146D5263DF7E537" pageId="0" pageNumber="1" type="differential diagnosis">
|
||
<paragraph id="1960E9ED0B2F752718143AADD23DE361" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||
<paragraph id="31CD65A38FDE9D7902D2D4A631C89E12" pageId="0" pageNumber="1">
|
||
<taxonomicName id="8109D60431B1AE63607B2A6CB5ECA4BD" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis id="591C56747EAC63D0D0A852028955DBFE" italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="1401C34F35A843412F3A1CA293D7F2BB" lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
|
||
<emphasis id="B1681FBBC8800694081238B6D7E15C8F" italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in usually having a stout detached whorl, generally a curved tubular breathing device, and a faded colour pattern. In contrast,
|
||
<taxonomicName id="343BEB1007AAFE43BFFE920AB60C1807" lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
|
||
<emphasis id="D9D6B4DE9D147614774B8A9450E4CD28" italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has a slender detached whorl, generally a straight tubular breathing device, and a brownish zigzag colour pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2EA1BF220F7947074320318E461C6D68" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="2C5B536911BAFA19C52613CD8347C209" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="06830A6C5E4E89B7F4EA9694BA397EDB" pageId="0" pageNumber="1">
|
||
<emphasis id="8AC8C80796E32EB72C3ECA6896B1A3E7" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="7E38C9F19624D1BCC6F8DD0FCF77D2FF" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shell medium to large, cW 22.1-26.4 mm, cH 12.1-13.8 mm, thick, and sub-discoidal to discoidal shape; detached-whorl length 8.0-14.5 mm. Apex acute with dark colouration; spire slightly elevated. Whorls 5 to 6, convex and increasing regularly; suture wide and deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thin corneous, and transparent to thick with brownish colour. Shell colour whitish with dark brown zigzag pattern; dorsal shell surface with darker colour pattern than ventral surface; with narrow brownish spiral band on periphery (sometimes absent). Detached whorl of medium length, approximately the same as apertural width. Peristome circular and double; lip thickened, slightly expanded and multi-layered. Aperture opened sub-laterally. Breathing device tubular and its tip usually attached to preceding whorl; outer lip forms a long, curved or straight, closed tube; inner lip with small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, tall cup-shaped, and multispiral (Fig.
|
||
<figureCitation id="FDE6835EDF60A2323E592CB95BD52105" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AB80FBFAC6378708DC4B6D8AC4289BE6" pageId="0" pageNumber="1">
|
||
<emphasis id="64489BAF859EA2F5EB2BC698A81833DA" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="CF405DB855C3B98A78F1B07AB1083DBA" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||
.
|
||
</emphasis>
|
||
Taenioglossate radula arranged in inverted V-shaped rows. Central tooth with well-developed central cusp and two lateral cusps on each side tapering in size; central cusp large with blunt tip; lateral cusps with triangular shape and pointed tip. Lateral teeth consisting of four cusps; central cusp large, elongate, and flanked by pointed tips of two small inner cusps and one outer cusp. Inner and outer marginal teeth each composed of three cusps; central cusp with large and dull tip, flanked by small pointed tips of one inner cusp and one outer cusp (Fig.
|
||
<figureCitation id="257B5EF330B4FD63FEE7ACF58BCDF366" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35EB9C795DC7B33D8142A47C30C52E95" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="CC1A80E00C2720C0044FD379B5D5198A" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="4F978FBE3D0C71348704B6A121F7B88A" pageId="0" pageNumber="1">
|
||
The type locality was given as a broad location of
|
||
<normalizedToken id="916465904C3AD429355B6D13C9D453BE" originalValue="“Siam”">"Siam"</normalizedToken>
|
||
; however, the specimens from central Thailand are the most similar to the syntype illustrated herein (Fig.
|
||
<figureCitation id="A81D1846FE7595920D2A52C13E41FE07" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19A, B</figureCitation>
|
||
). In this study,
|
||
<taxonomicName id="CA3AE8BDD7660A75C21EEC2AA0F99B38" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="30B1591D7F76A88AA57289D02CAC92E2" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
has a wide distribution ranging from central to peninsular Thailand. The southernmost limits are around the Isthmus of Kra (Pratiew, Chumphon Province), and the northern limit is approximately in Nakhon Sawan Province (Fig.
|
||
<figureCitation id="7957971FF961498F60EBE9F61FDB99DC" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="12D276008D8A2DD4C46C23739E9F632D" doi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1" start="Figure 18" startId="F18">
|
||
<paragraph id="03C69B89EEE53F64600D2C5790395E61" pageId="0" pageNumber="1">
|
||
<emphasis id="CB04ACB4BCE3ECD656C165D0EBE9BC65" bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="2EC54E43C35E7BF6B86EE57615A1608E" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis id="4581D810639C2E44BDCEDF5CD7219B2D" italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
-species group.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EE30D3BEE5216D88B57D8FC15441D1B" doi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1" start="Figure 19" startId="F19">
|
||
<paragraph id="1173133A0781CDE5CCEC276C4E9213AE" pageId="0" pageNumber="1">
|
||
<emphasis id="D88D4D5C562DBD1F26172DF0E34163EE" bold="true" pageId="0" pageNumber="1">Figure 19.</emphasis>
|
||
Shell of
|
||
<taxonomicName id="F790C3F3D807E5D7B36C7C869CF850E5" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis id="250BE063373A3AC4DA5ED8398141B1BD" italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="ED60CEAB7FDBA240A9D2C56EBFE7CA12" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
syntype AMNH 42923 from Siam
|
||
<emphasis id="F0E22CFFFB0D9D96316874F78044A99A" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
figured specimen in
|
||
<bibRefCitation id="ED07E872CDEDBF94F630057A5CE9459D" DOI="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" author="Haines, WA" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" refId="B54" refString="Haines, WA, 1855. Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158. https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" title="Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158." url="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" year="1855">Haines (1855</bibRefCitation>
|
||
: figs 12, 13) anb
|
||
<emphasis id="CD70729E2295EE04811A7DFC2384C8B0" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
other specimen from the same syntype lot
|
||
<emphasis id="3332432040E535B8C87774327ED93535" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
specimens CUMZ 4383 from Tham Phitsadan, Chumphon
|
||
<emphasis id="7E9450E163B4E79D219366F60B04C357" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
specimen CUMZ 3982 from Tham Sriwilai, Saraburi
|
||
<emphasis id="17803761EB8C3A976707036DB182E785" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima
|
||
<emphasis id="8960AA9E869C277032BC47018BCE9F3C" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3C7B1F3CFDAA9F5171DAD19DF2D064FC" doi="10.3897/zookeys.1142.90097.figure20" httpUri="https://binary.pensoft.net/fig/799878" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
|
||
<paragraph id="1D63D5668598EF2644AEBDB8EF74D12B" pageId="0" pageNumber="1">
|
||
<emphasis id="0885A02887BBF392C0FA3D35ADE83646" bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
|
||
Drawing of general anatomy of
|
||
<taxonomicName id="0945E80A96669CDFDEF2E319DF9F98BB" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis id="4A6D39FE65B0261CACEF383BB71F2D75" italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 3982 from Saraburi Province, Thailand.
|
||
<emphasis id="8FCB5C06E1C27F50F05444DD65E81266" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
female
|
||
<emphasis id="A8DCD69EF60ACE498A3D431035FB70B7" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
male. Abbreviations:
|
||
<emphasis id="0B8E9E9A49FE30656D58F2901CD8CC42" bold="true" pageId="0" pageNumber="1">bc</emphasis>
|
||
, bursa copulatrix;;
|
||
<emphasis id="BEB08B1CAE8B639088BDE229C40D252A" bold="true" pageId="0" pageNumber="1">cm</emphasis>
|
||
, columellar muscle;
|
||
<emphasis id="24F99A5CE3FF1EEC97BE1D4DE8220DA1" bold="true" pageId="0" pageNumber="1">ct</emphasis>
|
||
, cephalic tentacle;
|
||
<emphasis id="6A2138D14F975D7AC56774B16F1A6244" bold="true" pageId="0" pageNumber="1">dg</emphasis>
|
||
, digestive gland;
|
||
<emphasis id="BE3A4E9F226CB058B03B6A488984E30D" bold="true" pageId="0" pageNumber="1">es</emphasis>
|
||
, eyespot;
|
||
<emphasis id="09D978ABF518AE6F5ED5E16D8478F76B" bold="true" pageId="0" pageNumber="1">f</emphasis>
|
||
, foot;
|
||
<emphasis id="D3C663DF33A5AAC8684D199E8F43EF4F" bold="true" pageId="0" pageNumber="1">fc</emphasis>
|
||
, faeces;
|
||
<emphasis id="A744BA86D58D15EB4D8A47249BACC227" bold="true" pageId="0" pageNumber="1">int</emphasis>
|
||
, intestine;
|
||
<emphasis id="682074D9C5EE51F7C2D45271AC03A426" bold="true" pageId="0" pageNumber="1">lc</emphasis>
|
||
, lung cavity;
|
||
<emphasis id="A817A70B10334C507360E427566FC955" bold="true" pageId="0" pageNumber="1">mc</emphasis>
|
||
, mantle collar;
|
||
<emphasis id="3F5118805DD2E3249A98E90E012E5534" bold="true" pageId="0" pageNumber="1">od</emphasis>
|
||
, oviduct;
|
||
<emphasis id="1FEF08F491EAF81DA96126615FB1C50C" bold="true" pageId="0" pageNumber="1">op</emphasis>
|
||
, operculum;
|
||
<emphasis id="9AF889C257D214EC0FAE4C3904D71A17" bold="true" pageId="0" pageNumber="1">ov</emphasis>
|
||
, ovary;
|
||
<emphasis id="131ABA9449EDB1A82CFF310B72A0A833" bold="true" pageId="0" pageNumber="1">p</emphasis>
|
||
, external penis;
|
||
<emphasis id="680A0A1FEE5680135CC6867AEC24B4C0" bold="true" pageId="0" pageNumber="1">pg</emphasis>
|
||
, prostate gland;
|
||
<emphasis id="3B8554BE273A9A430DAF64E672B100D2" bold="true" pageId="0" pageNumber="1">sg</emphasis>
|
||
, sperm groove;
|
||
<emphasis id="8F2C5C13DCE1164AD558808CC33DF0F5" bold="true" pageId="0" pageNumber="1">sn</emphasis>
|
||
, snout;
|
||
<emphasis id="20CA9BF797D24BD57BEDB813DDB4440B" bold="true" pageId="0" pageNumber="1">te</emphasis>
|
||
, testis;
|
||
<emphasis id="58B7F25E79890BE00DB7F1DA9BBA3210" bold="true" pageId="0" pageNumber="1">ut</emphasis>
|
||
, uterus;
|
||
<emphasis id="9467E90882AB25EE1A9DEA7620E55AD5" bold="true" pageId="0" pageNumber="1">vd</emphasis>
|
||
, vas deferens;
|
||
<emphasis id="155DB1135DB3D9716E0BAC107A9C23EB" bold="true" pageId="0" pageNumber="1">vg</emphasis>
|
||
, vaginal groove.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="E31109D6F5773D47B3DAE3B69A040461" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="7C745D84F7163A7DB1252EC0EEC4C401" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="4C355798FF1F72B8F21DC7CD4D72E336" pageId="0" pageNumber="1">
|
||
Previous literature recorded
|
||
<taxonomicName id="9095FF5F700E76A9FED28F289A4B4A06" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="70E319FC99FE4DFC1048B01D78F537C7" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
from several places in Laos, Cambodia, and Vietnam [Tonkin] (
|
||
<bibRefCitation id="9B4F7E4ACCEC12AA887D28B73E5C8281" author="Moellendorff, von OF" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="349 - 398" refId="B95" refString="Moellendorff, von OF, 1885. Materialien zur fauna von China (Die Auriculaceen, Nachtraege und Berichtigungen). Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 12: 349 - 398" title="Materialien zur fauna von China (Die Auriculaceen, Nachtraege und Berichtigungen)." volume="12" year="1885">
|
||
<normalizedToken id="74BFD8099656BB22F7C9755E71320959" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
1885
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F6816FE962F9AD0DE91CA18DF5555DE1" author="Ancey, C-F" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille" pageId="0" pageNumber="1" pagination="125 - 150" refId="B3" refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150" title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region." volume="1" year="1898">Ancey 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8E493ED8F09597985E6017204590121B" author="Fischer, P-H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B41" refString="Fischer, P-H, 1973. Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches." volume="90" year="1973">Fischer 1973</bibRefCitation>
|
||
). However, these records seem ambiguous because the type specimens have never been examined, were very limited in the number of specimens, and many species have a similar shell morphology to
|
||
<taxonomicName id="70EEDB20026B75476BA5878B9560FFC7" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="7BDA875AD4827F2806D3531634D2343B" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the species delimitation by previous authors is probably different from the concept of the type specimens. For example, this species has been reported in Chiangdao, Chiangmai, northern Thailand (sensu
|
||
<bibRefCitation id="CF7D801B7159DA1DB84CA6FBCA369E9C" author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
||
). We collected many specimens from several populations in the same area and examined
|
||
<normalizedToken id="E2C22E879EA71AB3538988B2175F5F4F" originalValue="Solem’s">Solem's</normalizedToken>
|
||
specimens in ZMUC. Due to the clear differences in shell characters, we recognise these populations as distinct species (
|
||
<taxonomicName id="7A4E20A868A08B1EAE3CF9E0C86CEFFA" lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
|
||
<emphasis id="4648B7987D7DAF9742E933E5CC863284" italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.). For that reason, the distribution range of
|
||
<taxonomicName id="A0DE5EB188273068D5C1C4E481B3638A" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="66A9CE25AF8FE13E30472FAB90B5936D" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
, especially from Laos, Cambodia, and Vietnam, needs to be re-examined with new specimen collections with more precise locality.
|
||
</paragraph>
|
||
<paragraph id="FB9A6AE840C546B08C2DC15CE8AC0C8A" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="33B7DD9109BDC542FCB6BA7946922889" DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. (2020b)</bibRefCitation>
|
||
reported several "
|
||
<taxonomicName id="FF40CBD8F27D7D279173B21BA2F00B41" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="03C34C1B71422B2E8C615D38ABE10931" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
" populations from Northern Vietnam in Dien Bien, Hao Binh, and Son La provinces. The figured specimen from Dien Bien Province shows the distance ~800 km away from the central population of this species from central Thailand. This specimen (
|
||
<bibRefCitation id="6BF78CA7ED0B70C4725D2BACF72DD768" DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
|
||
: fig. 6b, c) had a depressed spire, and a curved, tubular breathing device, and is probably
|
||
<taxonomicName id="CCAC691DDBC2232B8263362EFBC5665C" lsidName="R. simplicilabre" pageId="0" pageNumber="1" rank="species" species="simplicilabre">
|
||
<emphasis id="39C06375F85DF64139CAB2113ACD3DC6" italics="true" pageId="0" pageNumber="1">R. simplicilabre</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="94BB726A8059E74D06853CFA2A945F9A" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis id="7DF310137DFB47EF347AC46AF4227052" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B4728A138B348EEFFC47B7608020104" pageId="0" pageNumber="1">
|
||
The specimens collected from southern Thailand (Chumporn, Phetchaburi, and Prachuapkhirikhan provinces) tended to have longer detached whorls, longer breathing devices, and brighter shell colour than the populations from central Thailand (typical). However, the shell colour pattern, detached whorl, and breathing device structures are identical to the type specimens (Fig.
|
||
<figureCitation id="973B01544155923AD71C97F9B26BF431" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19A, B</figureCitation>
|
||
). Therefore, these are attributed to morphological variation across the wide distribution range of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |