<mods:titleid="4CC6C200FE712C120E6507FAB71693CB">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
<mods:affiliationid="D077DDE1EA91B0150CBD18A145C79488">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:affiliationid="4A4F8ED08E01E01A7B980CA21215487C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
<mods:affiliationid="16ED7FE862044C70251DE9D0F7003161">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
<mods:affiliationid="CBD21506C33AC49D98BFBAF19C99D408">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
<mods:affiliationid="8855ED3403269C2D234EE19DB9CE765C">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
<mods:affiliationid="F289821E144ADA5C569D4A94BB5E80CF">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:affiliationid="D5991FF6154A5E9A435ABF5E7EB596B4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<figureCitationid="06426C839D8D2D40EF1A3A4A8039E14A"captionStart="Figure 17"captionStartId="F17"captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)."figureDoi="10.3897/zookeys.1142.90097.figure17"httpUri="https://binary.pensoft.net/fig/799875"pageId="0"pageNumber="1">Figs 17A</figureCitation>
<figureCitationid="EDBFBE11A3285043E115077DB22F0ECE"captionStart="Figure 18"captionStartId="F18"captionText="Figure 18. Distribution map of Rhiostoma housei - species group."figureDoi="10.3897/zookeys.1142.90097.figure18"httpUri="https://binary.pensoft.net/fig/799876"pageId="0"pageNumber="1">, 18</figureCitation>
<figureCitationid="4EDCB25D3F1345F4780C119944060091"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">, 19</figureCitation>
<figureCitationid="0D0F253064D5B638A77509FB783B661F"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Drawing of general anatomy of Rhiostoma housei specimen CUMZ 3982 from Saraburi Province, Thailand. A female B male. Abbreviations: bc, bursa copulatrix;; cm, columellar muscle; ct, cephalic tentacle; dg, digestive gland; es, eyespot; f, foot; fc, faeces; int, intestine; lc, lung cavity; mc, mantle collar; od, oviduct; op, operculum; ov, ovary; p, external penis; pg, prostate gland; sg, sperm groove; sn, snout; te, testis; ut, uterus; vd, vas deferens; vg, vaginal groove."figureDoi="10.3897/zookeys.1142.90097.figure20"httpUri="https://binary.pensoft.net/fig/799878"pageId="0"pageNumber="1">, 20</figureCitation>
<figureCitationid="3507F58B2E14BE1AE62144F294892261"captionStart="Figure 32"captionStartId="F32"captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai."figureDoi="10.3897/zookeys.1142.90097.figure32"httpUri="https://binary.pensoft.net/fig/799890"pageId="0"pageNumber="1">, 32A</figureCitation>
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<taxonomicNameid="4B2B7FFA0A1944E65B43F630429B3D34"authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit"authorityYear="2023"class="Gastropoda"family="Cyclophoridae"genus="Rhiostoma"higherTaxonomySource="treatment-meta"kingdom="Animalia"lsidName="Rhiostoma"order="Architaenioglossa"pageId="0"pageNumber="1"phylum="Mollusca"rank="genus">Rhiostoma</taxonomicName>
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This species was described from specimens from the Haines collection (AMNH). The unique name-bearing type was not designated; however, at least two specimens were illustrated in the original description. The specimen identical to the illustration (shape of breathing device) and measurements in
<bibRefCitationid="918DE16FCB44503C1079ACD42750FDCC"DOI="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x"author="Haines, WA"journalOrPublisher="Nature and Life in Southeast Asia, Kyoto"pageId="0"pageNumber="1"refId="B54"refString="Haines, WA, 1855. Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158. https://doi.org/10.1111/j.1749-6632.1858.tb00358.x"title="Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158."url="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x"year="1855">Haines (1855</bibRefCitation>
<figureCitationid="6C98E49CA31FE7464BB9E63B9276D311"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19A, B</figureCitation>
<figureCitationid="CE445DE56609E32417F941F7EEB161C8"captionStart="Figure 7"captionStartId="F7"captionText="Figure 7. A-C Internal anatomy of Rhiostoma housei specimen CUMZ 3982 from Tham Sriwilai, Saraburi, A inside view of lung cavity shows an arrangement of the kidney, circulatory system and the terminal part of the digestive tract B male genitalia and C female genitalia D spermatophore of Rhiostoma samuiense specimen CUMZ 4713 from Kanchanadit, Surat Thani with its cross sections showing spermatophore shape in each position. Abbreviations: an, anus; bc, bursa copulatrix; fc, faeces; go, genital opening; h, heart; kd, kidney; mc, mantle collar; od, oviduct; pg, prostate gland; re, rectum; sr, seminal receptacle; ut, uterus; v, vein; vd, vas deferens."figureDoi="10.3897/zookeys.1142.90097.figure7"httpUri="https://binary.pensoft.net/fig/799865"pageId="0"pageNumber="1">7A-C</figureCitation>
<figureCitationid="41041582FA4282D07ECB5EE8AC0DB2DE"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19D</figureCitation>
<figureCitationid="154C16225DBC7C4D5646D1D40791DEAC"captionStart="Figure 32"captionStartId="F32"captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai."figureDoi="10.3897/zookeys.1142.90097.figure32"httpUri="https://binary.pensoft.net/fig/799890"pageId="0"pageNumber="1">32A</figureCitation>
<figureCitationid="A952C043FEB75B9608F857D65EB09A4B"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19E</figureCitation>
<locationid="181AF75D22508CB0BB8E8430244AE702"LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:181AF75D22508CB0BB8E8430244AE702"country="Thailand"county="East Siam"municipality="East Siam"name="Sakaerat Environmental Research Station"stateProvince="Saraburi">Sakaerat Environmental Research Station</location>
<figureCitationid="D193FBC77D4A38C4E5B207FB2D9D8D09"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19F</figureCitation>
<locationid="4A21BB501F97840176D365B9116B9D6F"LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:4A21BB501F97840176D365B9116B9D6F"country="Thailand"county="East Siam"municipality="East Siam"name="Tham Na Kwang"stateProvince="Saraburi">Tham Na Kwang</location>
<locationid="865DDE916C2BFBDC42304B33904FE650"LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:865DDE916C2BFBDC42304B33904FE650"country="Thailand"county="East Siam"municipality="East Siam"name="Pa La Au Waterfall"stateProvince="Saraburi">Pa La Au Waterfall</location>
<figureCitationid="0CFD7E1E875262E7AAD7653ADFA7C257"captionStart="Figure 17"captionStartId="F17"captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)."figureDoi="10.3897/zookeys.1142.90097.figure17"httpUri="https://binary.pensoft.net/fig/799875"pageId="0"pageNumber="1">17A</figureCitation>
<locationid="F42B3B25FEEB327418C3E6483928E409"LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:F42B3B25FEEB327418C3E6483928E409"country="Thailand"county="East Siam"municipality="East Siam"name="Saun Wiweg Bureau of Monks"stateProvince="Saraburi">Saun Wiweg Bureau of Monks</location>
<figureCitationid="C2D2F1294345C8E82193243EEBEF95B5"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19C</figureCitation>
<paragraphid="C9EA6C10161BB24237B1488A54C15AE0"pageId="0"pageNumber="1">Shell medium to large, thick, and depressed. Detached whorl longer than aperture width, curved, and descending. Breathing device straight with complete tube. Shell usually with dark brown zigzag patterns.</paragraph>
Shell medium to large, cW 22.1-26.4 mm, cH 12.1-13.8 mm, thick, and sub-discoidal to discoidal shape; detached-whorl length 8.0-14.5 mm. Apex acute with dark colouration; spire slightly elevated. Whorls 5 to 6, convex and increasing regularly; suture wide and deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thin corneous, and transparent to thick with brownish colour. Shell colour whitish with dark brown zigzag pattern; dorsal shell surface with darker colour pattern than ventral surface; with narrow brownish spiral band on periphery (sometimes absent). Detached whorl of medium length, approximately the same as apertural width. Peristome circular and double; lip thickened, slightly expanded and multi-layered. Aperture opened sub-laterally. Breathing device tubular and its tip usually attached to preceding whorl; outer lip forms a long, curved or straight, closed tube; inner lip with small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, tall cup-shaped, and multispiral (Fig.
<figureCitationid="FDE6835EDF60A2323E592CB95BD52105"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19</figureCitation>
Taenioglossate radula arranged in inverted V-shaped rows. Central tooth with well-developed central cusp and two lateral cusps on each side tapering in size; central cusp large with blunt tip; lateral cusps with triangular shape and pointed tip. Lateral teeth consisting of four cusps; central cusp large, elongate, and flanked by pointed tips of two small inner cusps and one outer cusp. Inner and outer marginal teeth each composed of three cusps; central cusp with large and dull tip, flanked by small pointed tips of one inner cusp and one outer cusp (Fig.
<figureCitationid="257B5EF330B4FD63FEE7ACF58BCDF366"captionStart="Figure 32"captionStartId="F32"captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai."figureDoi="10.3897/zookeys.1142.90097.figure32"httpUri="https://binary.pensoft.net/fig/799890"pageId="0"pageNumber="1">32A</figureCitation>
<figureCitationid="A81D1846FE7595920D2A52C13E41FE07"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19A, B</figureCitation>
has a wide distribution ranging from central to peninsular Thailand. The southernmost limits are around the Isthmus of Kra (Pratiew, Chumphon Province), and the northern limit is approximately in Nakhon Sawan Province (Fig.
<figureCitationid="7957971FF961498F60EBE9F61FDB99DC"captionStart="Figure 18"captionStartId="F18"captionText="Figure 18. Distribution map of Rhiostoma housei - species group."figureDoi="10.3897/zookeys.1142.90097.figure18"httpUri="https://binary.pensoft.net/fig/799876"pageId="0"pageNumber="1">18</figureCitation>
<bibRefCitationid="ED07E872CDEDBF94F630057A5CE9459D"DOI="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x"author="Haines, WA"journalOrPublisher="Nature and Life in Southeast Asia, Kyoto"pageId="0"pageNumber="1"refId="B54"refString="Haines, WA, 1855. Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158. https://doi.org/10.1111/j.1749-6632.1858.tb00358.x"title="Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158."url="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x"year="1855">Haines (1855</bibRefCitation>
<bibRefCitationid="9B4F7E4ACCEC12AA887D28B73E5C8281"author="Moellendorff, von OF"journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft"pageId="0"pageNumber="1"pagination="349 - 398"refId="B95"refString="Moellendorff, von OF, 1885. Materialien zur fauna von China (Die Auriculaceen, Nachtraege und Berichtigungen). Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 12: 349 - 398"title="Materialien zur fauna von China (Die Auriculaceen, Nachtraege und Berichtigungen)."volume="12"year="1885">
<bibRefCitationid="F6816FE962F9AD0DE91CA18DF5555DE1"author="Ancey, C-F"journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille"pageId="0"pageNumber="1"pagination="125 - 150"refId="B3"refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150"title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region."volume="1"year="1898">Ancey 1898</bibRefCitation>
). However, these records seem ambiguous because the type specimens have never been examined, were very limited in the number of specimens, and many species have a similar shell morphology to
. Moreover, the species delimitation by previous authors is probably different from the concept of the type specimens. For example, this species has been reported in Chiangdao, Chiangmai, northern Thailand (sensu
<bibRefCitationid="CF7D801B7159DA1DB84CA6FBCA369E9C"author="Solem, A"journalOrPublisher="Sapolia Zoologica Musei Hauniensis"pageId="0"pageNumber="1"pagination="1 - 110"refId="B155"refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110"title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae."volume="24"year="1966">Solem 1966</bibRefCitation>
<bibRefCitationid="33B7DD9109BDC542FCB6BA7946922889"DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4"author="Do, DS"journalOrPublisher="Ruthenica"pageId="0"pageNumber="1"pagination="165 - 172"refId="B36"refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4"title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species."url="https://doi.org/10.35885/ruthenica.2021.30(3).4"volume="30"year="2020 b">Do et al. (2020b)</bibRefCitation>
" populations from Northern Vietnam in Dien Bien, Hao Binh, and Son La provinces. The figured specimen from Dien Bien Province shows the distance ~800 km away from the central population of this species from central Thailand. This specimen (
<bibRefCitationid="6BF78CA7ED0B70C4725D2BACF72DD768"DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4"author="Do, DS"journalOrPublisher="Ruthenica"pageId="0"pageNumber="1"pagination="165 - 172"refId="B36"refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4"title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species."url="https://doi.org/10.35885/ruthenica.2021.30(3).4"volume="30"year="2020 b">Do et al. 2020b</bibRefCitation>
The specimens collected from southern Thailand (Chumporn, Phetchaburi, and Prachuapkhirikhan provinces) tended to have longer detached whorls, longer breathing devices, and brighter shell colour than the populations from central Thailand (typical). However, the shell colour pattern, detached whorl, and breathing device structures are identical to the type specimens (Fig.
<figureCitationid="973B01544155923AD71C97F9B26BF431"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan."figureDoi="10.3897/zookeys.1142.90097.figure19"httpUri="https://binary.pensoft.net/fig/799877"pageId="0"pageNumber="1">19A, B</figureCitation>
