436 lines
52 KiB
XML
436 lines
52 KiB
XML
<document ID-CLB-Dataset="24226" ID-DOI="http://dx.doi.org/10.3897/zookeys.907.36563" ID-GBIF-Dataset="69422977-aa4d-4fe3-a450-8ef54d6fa905" ID-PMC="PMC7002455" ID-Pensoft-Pub="1313-2970-907-1" ID-Pensoft-UUID="9920267E73CF5E00B644DED1F101D965" ID-PubMed="32063727" ID-ZooBank="DBF339E5EBBC461994388359C769473F" ModsDocID="1313-2970-907-1" checkinTime="1580398733216" checkinUser="pensoft" docAuthor="Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. & Nuss, Matthias" docDate="2020" docId="32794D663A525E4F937B1F932F002558" docLanguage="en" docName="ZooKeys 907: 1-99" docOrigin="ZooKeys 907" docSource="http://dx.doi.org/10.3897/zookeys.907.36563" docTitle="Hoploscopa Meyrick 1886" docType="treatment" docVersion="6" id="9920267E73CF5E00B644DED1F101D965" lastPageNumber="1" masterDocId="9920267E73CF5E00B644DED1F101D965" masterDocTitle="Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding" masterLastPageNumber="99" masterPageNumber="1" pageNumber="1" updateTime="1732652479166" updateUser="ExternalLinkService">
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<mods:title id="34013AEED0D9A81ED8338D10A74C6222">Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding</mods:title>
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<mods:namePart id="B5FCC3F4EFEA53A22865BA5A552CC585">Leger, Theo</mods:namePart>
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<mods:namePart id="70A7A7504B5BBD3FFCD65CF645718D3B">Kehlmaier, Christian</mods:namePart>
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<mods:namePart id="F21008BF285EC802E850E9435F93C6CA">Vairappan, Charles S.</mods:namePart>
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<mods:namePart id="1B054348A64A757FDDE31DDE4EFBC549">Nuss, Matthias</mods:namePart>
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<mods:date id="D1E9F58142F16E39675D0534C4E77D5C">2020</mods:date>
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<mods:number id="965806F03020682468942F0511E0C696">907</mods:number>
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<mods:url id="EF32FDF1F6BC5F9F1194F7C72BD840D4">http://dx.doi.org/10.3897/zookeys.907.36563</mods:url>
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<mods:identifier id="7EED4FD094809EBE169987262F23702C" type="DOI">http://dx.doi.org/10.3897/zookeys.907.36563</mods:identifier>
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<treatment id="32794D663A525E4F937B1F932F002558" ID-GBIF-Taxon="161926034" LSID="urn:lsid:plazi:treatment:32794D663A525E4F937B1F932F002558" httpUri="http://treatment.plazi.org/id/32794D663A525E4F937B1F932F002558" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Crambidae" scope_order="Lepidoptera">
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<subSubSection id="2CE2ABCD879A7985E99D31622BA917F7" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="18B25885836859FAE44615FDC4FF01BF" pageId="0" pageNumber="1">
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<taxonomicName id="78C36A669E5A10258E2A80AF9F75EAF0" ID-CoL="92BRZ" LSID="32794D66-3A52-5E4F-937B-1F932F002558" authority="Meyrick, 1886" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Hoploscopa Meyrick, 1886</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="CC2DD3D85FC42B67466B56272AF0759F" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="FBBA187012DB7B306E19CCAB9300C843" pageId="0" pageNumber="1">
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<taxonomicName id="D4C314C12B06D912D105AF5D30AC2F13" class="Insecta" family="Crambidae" genus="Haploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Haploscopa</taxonomicName>
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Hampson, 1897: 223.
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</paragraph>
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<paragraph id="CD36C096D39858BE3AC19E5F561F9C07" pageId="0" pageNumber="1">
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<taxonomicName id="8D85D49F2DDBFED6A73481D7BAB80FCC" class="Insecta" family="Crambidae" genus="Syncrotaula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syncrotaula" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Syncrotaula</taxonomicName>
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Meyrick, 1933: 378. Type species
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<taxonomicName id="2633135685E12514F8D2D67A28FDEA44" class="Insecta" family="Crambidae" genus="Eudorina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eudorina aurantiacalis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aurantiacalis">Eudorina aurantiacalis</taxonomicName>
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Snellen, 1895, by subsequent designation (for
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<taxonomicName id="BEE1BEE4EDE265F668561DC3D0FC39D1" class="Insecta" family="Crambidae" genus="Eudorina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eudorina" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eudorina</taxonomicName>
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Snellen, 1895) by Joannis, 1930.
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</paragraph>
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</subSubSection>
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<subSubSection id="BCE06E74097E7634160EDE6F4BD5565B" pageId="0" pageNumber="1" type="type species">
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<paragraph id="6EB4E96C27EEF5D540FD5B21941ACF78" pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph id="64ABEAEFB6B9E5FE4E011D564D2AD434" pageId="0" pageNumber="1">
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<taxonomicName id="7DF67D79AE3108012979DBD635B16CB2" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa astrapias" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="astrapias">
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<emphasis id="414ED87B65EA7302174EF311170FA822" italics="true" pageId="0" pageNumber="1">Hoploscopa astrapias</emphasis>
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</taxonomicName>
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Meyrick, 1886, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="B9E909152F04F5B8577A165577885F86" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="B0EAAF47D7A5DAC7B9A3473F47BBC359" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="6A0178B62AE5D05BDB2414A92B80B68C" pageId="0" pageNumber="1">
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<taxonomicName id="50BA03F0569BD60067A3FB04AD225DB1" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="15332F6364EB7E5AD64E943E8B9C15E3" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
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</taxonomicName>
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displays brown to dark brown forewings, often bearing pale yellow-, yellow-, or red-coloured median and postmedian diagonal stripes. In male genitalia, the uncus is well developed, the gnathos forms a ribbon-like structure, often with a posterior projection, and the vinculum bears laterally a pair of coremata. In female genitalia, the antrum is short, often sclerotised, and the corpus bursae bears a thorn, often with a basal sclerotisation.
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<taxonomicName id="A1B8BCCAC28AB92CE7F78A85F51D528C" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="D2636B920FA31675AA79034A75712BDA" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
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</taxonomicName>
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is morphologically very similar to
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<taxonomicName id="6EEAD367461CDC3BFA607BC86DE8E392" authorityName="Turner" authorityYear="1915" class="Insecta" family="Crambidae" genus="Perimeceta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perimeceta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="42CE9B5DB3CE8A7B98B7583C03C5E98B" italics="true" pageId="0" pageNumber="1">Perimeceta</emphasis>
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</taxonomicName>
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. The forewings of
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<taxonomicName id="E7E0D56D88D063B3E0874BDD0859DF47" authorityName="Turner" authorityYear="1915" class="Insecta" family="Crambidae" genus="Perimeceta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perimeceta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="08147684E5099B403F44521DCDF45AB4" italics="true" pageId="0" pageNumber="1">Perimeceta</emphasis>
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</taxonomicName>
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are slightly larger, display a yellow to brown ground colour, with one basal elliptic and one postmedian lunule-shaped white spot. In male genitalia,
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<taxonomicName id="D01F914451C8A01DAA66F0AA1218AA39" authorityName="Turner" authorityYear="1915" class="Insecta" family="Crambidae" genus="Perimeceta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perimeceta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="8303BFF3949953A57057875E7A6A9728" italics="true" pageId="0" pageNumber="1">Perimeceta</emphasis>
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</taxonomicName>
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shows a spade-shaped uncus, the gnathos arms connect shortly after arising and are expanded posteriorly into an elongated tip, the valva shows a conspicuously rounded ventral margin, and is apically narrowed into a tip, while the ventral margin is more or less straight in
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<taxonomicName id="AEEA86D50D667B37BF832D77B3B87046" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="AEB59C792CF1A8055531799785F178AA" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="A9511743E1112AA14733B61C7D72D16B" authorityName="Turner" authorityYear="1915" class="Insecta" family="Crambidae" genus="Perimeceta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perimeceta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="130E124528AC794DD12DF80FCDC2D7EA" italics="true" pageId="0" pageNumber="1">Perimeceta</emphasis>
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</taxonomicName>
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exhibits female genitalia similar to those of
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<taxonomicName id="3CA9A5D1FEBD88D0227887BE7760DAD9" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="BE0C439651DD56D3E4AF5C4140907F9E" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
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</taxonomicName>
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with a slender ductus bursae and a rounded corpus bursa bearing a thorn. However, the latter displays a membranous antrum in female genitalia, while it is sclerotised in most species of
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<taxonomicName id="73DE0B6887FE566D9230C18C6C90D258" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="4F7F7E4EECB4534B6FDCE6F7550C5202" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="171AB94EBEE4F284AD82B7CE368EA229" pageId="0" pageNumber="1" type="description">
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<paragraph id="6E2EE55C86CF3F7B64AEA2EE823D472C" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="69255D6C3B3294033052ECE9FEBD3014" pageId="0" pageNumber="1">
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<emphasis id="10FE8FD22B559EB70E93842702DE7FEC" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Antennae dorsally with pale yellow to brown scales. Ocelli absent. Frons slightly produced, rounded. Proboscis basally white to brown scaled. Maxillary palpi brown to dark brown, first segment pale yellow, inner side brown to pale yellow. Labial palpi porrect, 2-2.9
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<normalizedToken id="D6A5255DFFEAE8B3F88CE703864D1269" originalValue="×">x</normalizedToken>
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diameter of compound eye, brown, white to pale yellow at base, inner side brown to pale yellow.
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</paragraph>
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<paragraph id="4F936E750345C6F53F47B00BE1C51F1B" pageId="0" pageNumber="1">
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<emphasis id="31293A3477FA3596C71A55D6F653AA77" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
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(Figs
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<figureCitation id="6CADEBE181D2E6E23FB33B5F3F62C60A" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Habitus, venation and forewing pattern in Hoploscopa. 1 Habitus of Hoploscopa matheae sp. nov., holotype, ♀, Malaysia, Sabah, Kundasang, Kinabalu Mt. Lodge veranda, 6 ° 0 ' 42.15 " N, 116 ° 32 ' 3.63 " E, 1570 m (T. Leger & R. Mally) (genitalia on slide TL 315 ♀) 2 wing venation of Hoploscopa species: H. jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Prov., nr Bulolo, Mt Susu Nat. Res., Araucaria For., 975 m, 27 - 28 Aug 1983 (S. Miller) (wing preparation TL 706) 3 nomenclature used for description of forewing pattern of Hoploscopa spp. Abbreviations: bp (basal patch), bs (basal streak), cf (costal field), dds (distal discoidal stigma), mcp (median cubital patch), mds (median discoidal stigma), mdp (median dorsal patch), pds (proximal discoidal stigma), pma (postmedian area), pml (postmedian line), pmp (postmedian patch), sbf (subterminal field), sbl (subterminal line)." figureDoi="10.3897/zookeys.907.36563.figures1-3" httpUri="https://binary.pensoft.net/fig/375756" pageId="0" pageNumber="1">1</figureCitation>
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-
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<figureCitation id="EA696ACCC9DF16765D973526FDBC0AA1" captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies & C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies & C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey & K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. & Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel & R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">45</figureCitation>
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). Collar white to pale yellow. Frenulum simple in ♂, triple in ♀. Forewing length 7-13 mm, 2.4-2.9
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<normalizedToken id="736702FDA9937FE39997ECC490596D64" originalValue="×">x</normalizedToken>
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maximal width, females slightly larger than males. Wing venation (Fig.
|
||
<figureCitation id="7F04AE5C1927499591CFA0287C686AF5" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Habitus, venation and forewing pattern in Hoploscopa. 1 Habitus of Hoploscopa matheae sp. nov., holotype, ♀, Malaysia, Sabah, Kundasang, Kinabalu Mt. Lodge veranda, 6 ° 0 ' 42.15 " N, 116 ° 32 ' 3.63 " E, 1570 m (T. Leger & R. Mally) (genitalia on slide TL 315 ♀) 2 wing venation of Hoploscopa species: H. jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Prov., nr Bulolo, Mt Susu Nat. Res., Araucaria For., 975 m, 27 - 28 Aug 1983 (S. Miller) (wing preparation TL 706) 3 nomenclature used for description of forewing pattern of Hoploscopa spp. Abbreviations: bp (basal patch), bs (basal streak), cf (costal field), dds (distal discoidal stigma), mcp (median cubital patch), mds (median discoidal stigma), mdp (median dorsal patch), pds (proximal discoidal stigma), pma (postmedian area), pml (postmedian line), pmp (postmedian patch), sbf (subterminal field), sbl (subterminal line)." figureDoi="10.3897/zookeys.907.36563.figures1-3" httpUri="https://binary.pensoft.net/fig/375756" pageId="0" pageNumber="1">2</figureCitation>
|
||
): R1 not connected to Sc; R2, R3 and R4 stalked together; R5 free, arising from upper corner of cell; M1, M2 not stalked; M3 arising from lower corner of cell; CuA1 arising below lower corner; CuA2 at distal 1/3 of cell; 1A+2A arising from cell base; Hindwing Sc+R1 connected to Rs at distal 1/3; M1 connected to Sc+R1 by short vein; cell closed; M2, M3 and CuA1 arising at lower angle of cell; CuA2 arising at middle of cell; 1A, 2A, 3A free. Forewing ground colour brown to dark brown, markings white, yellow or red colour. The pattern elements can be described as follows (Fig.
|
||
<figureCitation id="08652ED9A21484C8AD88FF053EDC70A2" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Habitus, venation and forewing pattern in Hoploscopa. 1 Habitus of Hoploscopa matheae sp. nov., holotype, ♀, Malaysia, Sabah, Kundasang, Kinabalu Mt. Lodge veranda, 6 ° 0 ' 42.15 " N, 116 ° 32 ' 3.63 " E, 1570 m (T. Leger & R. Mally) (genitalia on slide TL 315 ♀) 2 wing venation of Hoploscopa species: H. jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Prov., nr Bulolo, Mt Susu Nat. Res., Araucaria For., 975 m, 27 - 28 Aug 1983 (S. Miller) (wing preparation TL 706) 3 nomenclature used for description of forewing pattern of Hoploscopa spp. Abbreviations: bp (basal patch), bs (basal streak), cf (costal field), dds (distal discoidal stigma), mcp (median cubital patch), mds (median discoidal stigma), mdp (median dorsal patch), pds (proximal discoidal stigma), pma (postmedian area), pml (postmedian line), pmp (postmedian patch), sbf (subterminal field), sbl (subterminal line)." figureDoi="10.3897/zookeys.907.36563.figures1-3" httpUri="https://binary.pensoft.net/fig/375756" pageId="0" pageNumber="1">3</figureCitation>
|
||
):
|
||
</paragraph>
|
||
<caption id="7858AC319ECDAD05984583767043CE6E" doi="10.3897/zookeys.907.36563.figures1-3" httpUri="https://binary.pensoft.net/fig/375756" pageId="0" pageNumber="1" start="Figures 1–3" startId="F1">
|
||
<paragraph id="31BF54D671CAAC2FE614B4EAD7C7FBA0" pageId="0" pageNumber="1">
|
||
<emphasis id="8496AB982D1A2FFB1B0EBBB21B4B5387" bold="true" pageId="0" pageNumber="1">Figures 1-3.</emphasis>
|
||
Habitus, venation and forewing pattern in
|
||
<taxonomicName id="37B39701C82A6E88843B366D9EBDA2D0" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6B577A83CA00B6874279E2CAB9EA45C8" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B2772EBA240A9863C9DD20F64EE98465" bold="true" pageId="0" pageNumber="1">1</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="6FC84A1AF93A0A35A296FB8DADF61BAA" authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa matheae" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="matheae">
|
||
<emphasis id="FB418DBBEC8AE1C5AB11A64F1ED01BE9" italics="true" pageId="0" pageNumber="1">Hoploscopa matheae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♀, Malaysia, Sabah, Kundasang, Kinabalu Mt. Lodge veranda,
|
||
<geoCoordinate id="325DD8F0FB30ED74A1251041E126FA6D" degrees="6" direction="north" minutes="0" orientation="latitude" precision="1" seconds="42.15" value="6.0117083">6°0'42.15"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C6BEE4DE869B890D923D0FF6AB017F84" degrees="116" direction="east" minutes="32" orientation="longitude" precision="1" seconds="3.63" value="116.53434">116°32'3.63"E</geoCoordinate>
|
||
, 1570 m (T.
|
||
<normalizedToken id="AAF08053EE4391E29316B72D78B29508" originalValue="Léger">Leger</normalizedToken>
|
||
& R. Mally) (genitalia on slide TL315 ♀)
|
||
<emphasis id="B7BD72651156F7A7857671FA88313274" bold="true" pageId="0" pageNumber="1">2</emphasis>
|
||
wing venation of
|
||
<taxonomicName id="A9BFF0DBF9584376E7AE8D01ED6CDC8A" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2E18236DD6C75483B62BEE0C9CFEE177" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
species:
|
||
<taxonomicName id="023645DC746316293D10F73C2B9A0C66" lsidName="H. jubata" pageId="0" pageNumber="1" rank="species" species="jubata">
|
||
<emphasis id="3C45D74289B32E1404A32E99382251E5" italics="true" pageId="0" pageNumber="1">H. jubata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♂, Papua New Guinea, Morobe Prov., nr Bulolo, Mt Susu Nat. Res.,
|
||
<taxonomicName id="6230541A4AAE7D29882ACBBE9DB820D6" class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="FC7C66B2A61F5B1093EABFAB88505A62" italics="true" pageId="0" pageNumber="1">Araucaria</emphasis>
|
||
</taxonomicName>
|
||
For., 975 m, 27-28 Aug 1983 (S. Miller) (wing preparation TL706)
|
||
<emphasis id="17DE7CDB7AB032498D3C74865605FA3B" bold="true" pageId="0" pageNumber="1">3</emphasis>
|
||
nomenclature used for description of forewing pattern of
|
||
<taxonomicName id="61F0692E1375332C48B187B5018343B7" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EE5325E4DBC5EF26FCAFDBFCD3F89E31" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
spp. Abbreviations: bp (basal patch), bs (basal streak), cf (costal field), dds (distal discoidal stigma), mcp (median cubital patch), mds (median discoidal stigma), mdp (median dorsal patch), pds (proximal discoidal stigma), pma (postmedian area), pml (postmedian line), pmp (postmedian patch), sbf (subterminal field), sbl (subterminal line).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6610A40B4A1DE1F051618DD3689E8B2D" pageId="0" pageNumber="1">
|
||
<emphasis id="44092EBDC0ED7F7548826F08F0F082BC" bold="true" pageId="0" pageNumber="1">cf</emphasis>
|
||
costal field stretched along costa up to postmedian area;
|
||
</paragraph>
|
||
<paragraph id="CF2C313E51FE371AF1554486E522AA31" pageId="0" pageNumber="1">
|
||
<emphasis id="939E6D2E8490590DC7323A7D7DC43128" bold="true" pageId="0" pageNumber="1">bs</emphasis>
|
||
basal streak small, weakly or not marked;
|
||
</paragraph>
|
||
<paragraph id="381475042A10DBD1B48EC1A79E79D543" pageId="0" pageNumber="1">
|
||
<emphasis id="52943A6405C0FC475CB10228B28746C2" bold="true" pageId="0" pageNumber="1">bp</emphasis>
|
||
basal patch often quadrangular, ill-defined;
|
||
</paragraph>
|
||
<paragraph id="B41C7B5DE2048CF5720EBD4693D22D1A" pageId="0" pageNumber="1">
|
||
<emphasis id="42D01163BF78FD72BA2EC382A6704810" bold="true" pageId="0" pageNumber="1">pds</emphasis>
|
||
and
|
||
<emphasis id="FEB7F40B87F3A3FDE820A7A4B5217F0A" bold="true" pageId="0" pageNumber="1">dds</emphasis>
|
||
proximal and distal discoidal stigmata along costal field;
|
||
</paragraph>
|
||
<paragraph id="991E38CAF6073CABC6D502BB85696D99" pageId="0" pageNumber="1">
|
||
<emphasis id="1B2A78512FBEE2B644EF70228C102173" bold="true" pageId="0" pageNumber="1">mds</emphasis>
|
||
the median discoidal stigma trapezoid, with costal edge longer than dorsal one;
|
||
</paragraph>
|
||
<paragraph id="743FC945E99A01DAD3DE6407A54B4B4A" pageId="0" pageNumber="1">
|
||
<emphasis id="6D2FEF5CC50ADEC93455221D35CC1953" bold="true" pageId="0" pageNumber="1">mcp</emphasis>
|
||
and
|
||
<emphasis id="696E753503F05F113A6A6552A47F4F63" bold="true" pageId="0" pageNumber="1">mdp</emphasis>
|
||
median cubital patch and median dorsal patch in line with median discoidal stigma, often forming together a Y-shape pattern;
|
||
</paragraph>
|
||
<paragraph id="93A8266398E0AA224BF8BCC9AEB8F5DF" pageId="0" pageNumber="1">
|
||
<emphasis id="8F7CC688122D16305C163D74B3042604" bold="true" pageId="0" pageNumber="1">pmp</emphasis>
|
||
postmedian patch roughly triangular, narrowing at costa;
|
||
</paragraph>
|
||
<paragraph id="8E50F17CDBF79523EEA4EAF9B1020F34" pageId="0" pageNumber="1">
|
||
<emphasis id="B15EC7CBDBCF7EE53F5E25D566D54E47" bold="true" pageId="0" pageNumber="1">pml</emphasis>
|
||
postmedian line running from costa to middle of dorsum, often barely marked, mostly visible close to costa;
|
||
</paragraph>
|
||
<paragraph id="F7796A23769B27A9FFEA7FDAA7231520" pageId="0" pageNumber="1">
|
||
<emphasis id="91474D66A7D02A0340C29380BCBA6DF1" bold="true" pageId="0" pageNumber="1">pma</emphasis>
|
||
postmedian area often suffused to a various extend with white, yellow or red;
|
||
</paragraph>
|
||
<paragraph id="5AC0207AEF3CADF0010ADDA5733A1AFA" pageId="0" pageNumber="1">
|
||
<emphasis id="654B541A80E13F2491A424B2267530B2" bold="true" pageId="0" pageNumber="1">sbl</emphasis>
|
||
subterminal line running from costa close to apex towards dorsum, often incurved inwards at CuA2;
|
||
</paragraph>
|
||
<paragraph id="F1CF543550A1F63C7014AB470C518D90" pageId="0" pageNumber="1">
|
||
<emphasis id="F67A29A449CDD1A67DFE47DAC75DC001" bold="true" pageId="0" pageNumber="1">sbf</emphasis>
|
||
subterminal field more or less suffused with brown, red or yellow.
|
||
</paragraph>
|
||
<paragraph id="50238A4040BD1C65825E7DF73281C02A" pageId="0" pageNumber="1">The margin is brown, in some species with spots, fringe unicoloured or chequered.</paragraph>
|
||
<paragraph id="0B8A54A3F309A9FBB1C5E10640512088" pageId="0" pageNumber="1">Hindwing upper side pale yellow to pale brown; underside pale yellow, with brown markings on costa and subterminal line toward costa; males of some species with androconial organ on the upper side at dorsum, consisting of upright scales along CuA2 and 1A veins and a protruded margin between CuA2 and 1A, bearing a patch of greyish scales. Forelegs brown to dark brown. Midlegs with femur brown, tibia and tarsi often pale yellow, speckled with brown. Hindlegs pale yellow on inner side, pale yellow to brown on outer side, tarsi bronze to pale brown.</paragraph>
|
||
<paragraph id="C6D2141AB9278029932A5BC4BB03FD9C" pageId="0" pageNumber="1">
|
||
<emphasis id="07FF70A162574047F72572D4AC17021E" bold="true" italics="true" pageId="0" pageNumber="1">Tympanal organs</emphasis>
|
||
(Figs
|
||
<figureCitation id="E8D87364DD9BE55E26728FC5AD1ECF64" captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">83-84</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DC5917FC51E5ECF46915EB31E8E34539" pageId="0" pageNumber="1">
|
||
<emphasis id="07C2105E09CE570740E49910ADA409E4" bold="true" pageId="0" pageNumber="1">tb</emphasis>
|
||
Tympanic bridge deeply divided in the centre;
|
||
</paragraph>
|
||
<paragraph id="D79491C3EEED0DC1165FF57B7119D1B6" pageId="0" pageNumber="1">
|
||
<emphasis id="4DCC2F63CF163CA3EF170FE4B33B3FD1" bold="true" pageId="0" pageNumber="1">td</emphasis>
|
||
Tympanic drum (Fig.
|
||
<figureCitation id="839AD9757A1B04336DC3800070A37C66" captionStart="Figures 46–51" captionStartId="F5" captionText="Figures 46 - 51. Male genitalia features of Hoploscopa. For each figure left above: genitalia without phallus; left below: phallus; right above: gnathos drawing; right below: juxta drawing. 46 Hoploscopa albipuncta sp. nov., paratype, TL 336 ♂ (genitalia without phallus), TL 635 ♂ (phallus) 47 Hoploscopa matheae sp. nov., paratype, TL 730 ♂ 48 Hoploscopa sepanggi sp. nov., paratype, TL 641 ♂ 49 Hoploscopa cynodonta sp. nov., paratype, TL 531 ♂ 50 Hoploscopa parvimacula sp. nov., holotype, TL 510 ♂ 51 Hoploscopa kinabaluensis sp. nov., paratype, TL 342 ♂. Abbreviations: c (cornutus), de (ductus ejaculatorius), g (gnathos), jx (juxta), s (saccus), ta (tegumen arm), v (valva), vc (vinculum), uc (uncus)." figureDoi="10.3897/zookeys.907.36563.figures46-51" httpUri="https://binary.pensoft.net/fig/375760" pageId="0" pageNumber="1">50</figureCitation>
|
||
) ovoid, antero-ventrally semi-closed, posteriorly not reaching transverse ridge;
|
||
</paragraph>
|
||
<paragraph id="BEA2AB86B3014918DC4BFE2366BC2859" pageId="0" pageNumber="1">
|
||
<emphasis id="5CB97727DDDC3BC97FDF32B02517C2EC" bold="true" pageId="0" pageNumber="1">tdp</emphasis>
|
||
Tympanic depressions broad, opened ventrally;
|
||
</paragraph>
|
||
<paragraph id="8549CF3579E8FF1EA76C3B883191B383" pageId="0" pageNumber="1">
|
||
<emphasis id="83DAD75A25908565DCFD99A5D71FAEA0" bold="true" pageId="0" pageNumber="1">tm</emphasis>
|
||
Tympanum subtriangular;
|
||
</paragraph>
|
||
<paragraph id="EB63427D207054D562DE944891711AAC" pageId="0" pageNumber="1">
|
||
<emphasis id="2664BA36FD87656FAD99A26269C06E31" bold="true" pageId="0" pageNumber="1">tr</emphasis>
|
||
Transverse ridge (
|
||
<normalizedToken id="61177669D1C464338DEDF8E09C48B852" originalValue="“”">""</normalizedToken>
|
||
) not sclerotised.
|
||
</paragraph>
|
||
<paragraph id="E3F1608C7E33B3B7DC09162803753E90" pageId="0" pageNumber="1">
|
||
<emphasis id="E6233C885C9774CF473619FA8981DB5D" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen.</emphasis>
|
||
Pale brown to brown. In males, sternum A8 more or less broadly indented, in some species with short, rounded lateral projections (Figs
|
||
<figureCitation id="4A334F6A085831BAD67451373C8019B8" captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">85</figureCitation>
|
||
,
|
||
<figureCitation id="789544BE3C8C21870FF657BDFD897FAE" captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">86</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="81F2B3A8E50CDD53F7604BACD461BB89" pageId="0" pageNumber="1">
|
||
<emphasis id="2FBFCB279F02D423B15B6EC75C12F3C1" bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
|
||
(Figs
|
||
<figureCitation id="DDAACA4F31BEAD70F33BD4EC03A0FD67" captionStart="Figures 46–51" captionStartId="F5" captionText="Figures 46 - 51. Male genitalia features of Hoploscopa. For each figure left above: genitalia without phallus; left below: phallus; right above: gnathos drawing; right below: juxta drawing. 46 Hoploscopa albipuncta sp. nov., paratype, TL 336 ♂ (genitalia without phallus), TL 635 ♂ (phallus) 47 Hoploscopa matheae sp. nov., paratype, TL 730 ♂ 48 Hoploscopa sepanggi sp. nov., paratype, TL 641 ♂ 49 Hoploscopa cynodonta sp. nov., paratype, TL 531 ♂ 50 Hoploscopa parvimacula sp. nov., holotype, TL 510 ♂ 51 Hoploscopa kinabaluensis sp. nov., paratype, TL 342 ♂. Abbreviations: c (cornutus), de (ductus ejaculatorius), g (gnathos), jx (juxta), s (saccus), ta (tegumen arm), v (valva), vc (vinculum), uc (uncus)." figureDoi="10.3897/zookeys.907.36563.figures46-51" httpUri="https://binary.pensoft.net/fig/375760" pageId="0" pageNumber="1">46</figureCitation>
|
||
-
|
||
<figureCitation id="CAEBE800945F874DA9BA65A713E6CBFC" captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">82</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C6DEB98760ECA420E13D03B3ED5D6E28" pageId="0" pageNumber="1">
|
||
<emphasis id="4EC6B6584EAB58D6B75CEF5B88796D9B" bold="true" pageId="0" pageNumber="1">c</emphasis>
|
||
cornutus on phallus elongated, flat, spatula-shaped, except otherwise mentioned;
|
||
</paragraph>
|
||
<paragraph id="ABB7537D5B8D8CBA4A79442BB7A1A859" pageId="0" pageNumber="1">
|
||
<emphasis id="5F8B3336CA8A246B13A40B7B9CEC1CF6" bold="true" pageId="0" pageNumber="1">de</emphasis>
|
||
ductus ejaculatorius;
|
||
</paragraph>
|
||
<paragraph id="D7CB8A2232224BD5BB61952650C6882E" pageId="0" pageNumber="1">
|
||
<emphasis id="8E7128147129253ADE5F1B2671EF6198" bold="true" pageId="0" pageNumber="1">g</emphasis>
|
||
Gnathos arms (Fig.
|
||
<figureCitation id="0CCB89CA26744CA90773DFF04164026C" captionStart="Figures 52–57" captionStartId="F6" captionText="Figures 52 - 57. Male genitalia features of Hoploscopa. 52 Hoploscopa luteomacula Nuss, holotype, GU prep. Nuss 744 ♂ 53 Hoploscopa obliqua, holotype, Pyralidae Brit. Mus. Slide N ° BMNH 20252 ♂ (phallus with juxta attached) 54 Hoploscopa gombongi sp. nov., holotype, TL 653 ♂ 55 Hoploscopa tonsepi sp. nov., paratype, TL 661 ♂ 56 Hoploscopa niveofascia sp. nov., holotype, TL 442 ♂ 57 Hoploscopa marijoweissae sp. nov., holotype, TL 710 ♂." figureDoi="10.3897/zookeys.907.36563.figures52-57" httpUri="https://binary.pensoft.net/fig/375761" pageId="0" pageNumber="1">54</figureCitation>
|
||
) originating from the dorsal part of tegumen, forming ring, with or without dorso-median extension;
|
||
</paragraph>
|
||
<paragraph id="D8DCF820961F7087BC724ACADF226699" pageId="0" pageNumber="1">
|
||
<emphasis id="31F2007EBD79F8F0DBA7007436E705A5" bold="true" pageId="0" pageNumber="1">jx</emphasis>
|
||
Juxta basally wide with anterior margin rounded to quadrangular, narrowing at basal 1/4;
|
||
</paragraph>
|
||
<paragraph id="A571818DB4C6591CB01299E22E3FB608" pageId="0" pageNumber="1">
|
||
<emphasis id="F0ED1E83646C2F1C12A865D9AF986B0D" bold="true" pageId="0" pageNumber="1">s</emphasis>
|
||
saccus triangular to quadrangular, pointing dorsally;
|
||
</paragraph>
|
||
<paragraph id="5B202A8B0141C783A73B95B04030E264" pageId="0" pageNumber="1">
|
||
<emphasis id="A9520513B9FEF3E3C3CF508FE8FE03E3" bold="true" pageId="0" pageNumber="1">ta</emphasis>
|
||
Tegumen arms anteriorly sclerotised, with edge marked, posteriorly membranous; dorsally fused into a bridge of various width, articulated or fused to uncus base;
|
||
</paragraph>
|
||
<paragraph id="1B3C6F067E1F6423D96F6BE15D06CB35" pageId="0" pageNumber="1">
|
||
<emphasis id="638F984CCCACDC2045E0F2D671A7106B" bold="true" pageId="0" pageNumber="1">uc</emphasis>
|
||
Uncus well-developed, densely haired apically;
|
||
</paragraph>
|
||
<paragraph id="678C840507AAB0214DB13A2F2FD469AB" pageId="0" pageNumber="1">
|
||
<emphasis id="AC6AE954DF40EC019E8DF8B0FF54AD7C" bold="true" pageId="0" pageNumber="1">v</emphasis>
|
||
Valva elongated, hairy, sclerotised, with strongly sclerotised dorsal edge;
|
||
</paragraph>
|
||
<paragraph id="BD6FE5CB05CC6A2221D0BFEEAF6C525C" pageId="0" pageNumber="1">
|
||
<emphasis id="81FCADAC846BE89117BF3C536A9005AA" bold="true" pageId="0" pageNumber="1">vc</emphasis>
|
||
Vinculum U-shaped in posterior view, dorso-laterally on each side with anterior projection bearing brush-shaped coremata.
|
||
</paragraph>
|
||
<paragraph id="6EF4E5A7B5ACDBDC6BEBD696F5075911" pageId="0" pageNumber="1">
|
||
<emphasis id="BB3278BD228E5893F39BAC02F09D52F8" bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
|
||
(Figs
|
||
<figureCitation id="92D2AC9430A0B1B17858A6631CD3FE26" captionStart="Figures 87–90" captionStartId="F12" captionText="Figures 87 - 90. Female genitalia features of Hoploscopa. 87 Hoploscopa albipuncta sp. nov., paratype, TL 609 ♀ 88 Hoploscopa matheae sp. nov., paratype, TL 599 ♀ 89 Hoploscopa sepanggi sp. nov., paratype, TL 547 ♀ 90 Hoploscopa cynodonta sp. nov., paratype, TL 625 ♀. Abbreviations: aa (anterior apophyses), at (antrum), cb (corpus bursae), cs (corpus sclerotization), db (ductus bursae), ds (ductus seminalis), pa (posterior apophyses), pp (papillae anales), segm. VIII (segment VIII), t (thorn)." figureDoi="10.3897/zookeys.907.36563.figures87-90" httpUri="https://binary.pensoft.net/fig/375767" pageId="0" pageNumber="1">87</figureCitation>
|
||
-
|
||
<figureCitation id="A57F66DFF0371BC61D3E4AEFB421B904" captionStart="Figures 119–122" captionStartId="F20" captionText="Figures 119 - 122. Female genitalia features of Hoploscopa. 119 Hoploscopa boleta sp. nov., paratype, TL 448 ♀ 120 Hoploscopa pseudometacrossa sp. nov., paratype, TL 435 ♀ 121 Hoploscopa metacrossa, paratype, TL 446 ♀ 122 Hoploscopa jubata sp. nov., paratype, TL 445 ♀." figureDoi="10.3897/zookeys.907.36563.figures119-122" httpUri="https://binary.pensoft.net/fig/375775" pageId="0" pageNumber="1">122</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3F0756D6BD3A6EFACCCB5248FA596890" pageId="0" pageNumber="1">
|
||
<emphasis id="7079103D945BAE5B9BD1A03C6E34D925" bold="true" pageId="0" pageNumber="1">aa</emphasis>
|
||
Anterior apophyses (Fig.
|
||
<figureCitation id="BE6C6EE810C80DC05EC0B6E86F4B2DA4" captionStart="Figures 87–90" captionStartId="F12" captionText="Figures 87 - 90. Female genitalia features of Hoploscopa. 87 Hoploscopa albipuncta sp. nov., paratype, TL 609 ♀ 88 Hoploscopa matheae sp. nov., paratype, TL 599 ♀ 89 Hoploscopa sepanggi sp. nov., paratype, TL 547 ♀ 90 Hoploscopa cynodonta sp. nov., paratype, TL 625 ♀. Abbreviations: aa (anterior apophyses), at (antrum), cb (corpus bursae), cs (corpus sclerotization), db (ductus bursae), ds (ductus seminalis), pa (posterior apophyses), pp (papillae anales), segm. VIII (segment VIII), t (thorn)." figureDoi="10.3897/zookeys.907.36563.figures87-90" httpUri="https://binary.pensoft.net/fig/375767" pageId="0" pageNumber="1">88</figureCitation>
|
||
) bluntly angled at posterior 1/3, bent ventrad;
|
||
</paragraph>
|
||
<paragraph id="9759913E65D1CA5ECA09A7642DF52B36" pageId="0" pageNumber="1">
|
||
<emphasis id="439F2A447BDB2DBFCC16998FE239E8E7" bold="true" pageId="0" pageNumber="1">at</emphasis>
|
||
Antrum sclerotised or membranous;
|
||
</paragraph>
|
||
<paragraph id="3D14E7393CD03F49BBE71DAF98474EC7" pageId="0" pageNumber="1">
|
||
<emphasis id="E8585C0DA2A4519B879C65C08B15CD93" bold="true" pageId="0" pageNumber="1">cb</emphasis>
|
||
Corpus bursae globular or ovoid, its membrane with a reticulate structure;
|
||
</paragraph>
|
||
<paragraph id="FE64493928A52A6C668C97ADB7FA1D14" pageId="0" pageNumber="1">
|
||
<emphasis id="90FE14CED68FFCB5B2B4A4CF107D9CC5" bold="true" pageId="0" pageNumber="1">cs</emphasis>
|
||
Broad sclerotisation of the corpus bursae;
|
||
</paragraph>
|
||
<paragraph id="7903FFB43AC3CE4E7591766571A3275A" pageId="0" pageNumber="1">
|
||
<emphasis id="29C8F7714F002B51E4B7C9EB526EC743" bold="true" pageId="0" pageNumber="1">db</emphasis>
|
||
Ductus bursae membranous with longitudinal wrinkles, variable in length and shape;
|
||
</paragraph>
|
||
<paragraph id="58EBE3FD71793ADB82B8CB79ED67BF48" pageId="0" pageNumber="1">
|
||
<emphasis id="A661F86A523D6210D927579C8359C7D0" bold="true" pageId="0" pageNumber="1">pa</emphasis>
|
||
Posterior apophyses slender, straight, directed posterad except otherwise mentioned;
|
||
</paragraph>
|
||
<paragraph id="749ADC6DF33FA369C8D6D142379D367E" pageId="0" pageNumber="1">
|
||
<emphasis id="6D067C149F6FD1BF3D93A3B9F09A031E" bold="true" pageId="0" pageNumber="1">pp</emphasis>
|
||
Papillae anales thin in lateral view, dorsally and ventrally connected, setose;
|
||
</paragraph>
|
||
<paragraph id="C155EA330BB87F92AB01DBD8995A4228" pageId="0" pageNumber="1">
|
||
<emphasis id="16370946AAD54C7D1853BECC1B57BA47" bold="true" pageId="0" pageNumber="1">segm. VIII</emphasis>
|
||
Segment VIII faintly sclerotised, ventrally membranous; setae scattered across segment, more densely concentrated on posterior margin;
|
||
</paragraph>
|
||
<paragraph id="287D9DE89B4ED3E1A909E81C69E2AFA3" pageId="0" pageNumber="1">
|
||
<emphasis id="A9E105157D10092C80F943F002C1DCE6" bold="true" pageId="0" pageNumber="1">t</emphasis>
|
||
thorn on corpus bursae large, sclerotised, inwardly projecting, its base slighly extending outwards into a small sclerotised protuberance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="86D507CAEA723F4F9A2F2DA459F7D73B" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="C3112CD146987240A38D1621C0B8C7BB" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="5EDA78B51184AA9F471F58AB7A147A01" pageId="0" pageNumber="1">
|
||
<taxonomicName id="071BA8C5665CB681BC3250EB7C88F7AB" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4326C336F8527CDF478D4FFCD9D43CE9" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
is found from the Malay Peninsula and Sumatra in the West to the Samoa Islands in the East, as well as from the North of Thailand to Vanuatu and Fiji in the South. It is absent from Queensland (Australia) and New Caledonia. It is predominantly found in tropical mountain forests, with only few species encountered in the lowlands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5D9318B9A079458EECF7B270F4720993" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="050CBF6D19B1EDBB0760E312556ED641" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="490BC300A4D4BF1F81F35D5B92A56FC1" pageId="0" pageNumber="1">
|
||
Host plant data is available for five
|
||
<taxonomicName id="9177717C2A51E7ECE9B6C19FA1733D11" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="83608D3AE69FFC5EE321DBEBCA821DD1" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
species, all feeding on ferns. The larvae of
|
||
<taxonomicName id="5F1D989B163431129AFD7581547250BB" lsidName="H. gombongi" pageId="0" pageNumber="1" rank="species" species="gombongi">
|
||
<emphasis id="D8694CFEA7E26D880626CBAA4052AF63" italics="true" pageId="0" pageNumber="1">H. gombongi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="A952ACC6C5159305F112ABD7709623A3" lsidName="H. obliqua" pageId="0" pageNumber="1" rank="species" species="obliqua">
|
||
<emphasis id="628FE1B1873A2258831A4C01C321DF88" italics="true" pageId="0" pageNumber="1">H. obliqua</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2A62BE0968DF2FEA96F97147097F3E9C" lsidName="H. tonsepi" pageId="0" pageNumber="1" rank="species" species="tonsepi">
|
||
<emphasis id="941E11455240914419911966B2533CB5" italics="true" pageId="0" pageNumber="1">H. tonsepi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Papua New Guinea are reported from
|
||
<taxonomicName id="DD7E7D33D209D0A67A2AEA18BEC45E66" class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diplazium esculentum" order="Polypodiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="esculentum">
|
||
<emphasis id="B7B0DEE81B073CF3688A4A2577FF367A" italics="true" pageId="0" pageNumber="1">Diplazium esculentum</emphasis>
|
||
</taxonomicName>
|
||
(Retzius in Retzius &
|
||
<normalizedToken id="6A4349E72B287DC96C99961C765E773F" originalValue="König">Koenig</normalizedToken>
|
||
, 1791) Swartz, 1803 (
|
||
<taxonomicName id="9588EB97AA56E7D8761A9D3B032FD3E5" class="Polypodiopsida" family="Athyriaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="family">Athyriaceae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="4B07D026751EED4F75511C46AAA8D735" DOI="https://doi.org/10.4289/0013-8797.117.2.247" author="Miller, SE" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="247 - 250" refId="B35" refString="Miller, SE, Rosati, ME, Gewa, B, Novotny, V, Weiblen, GD, Hebert, PD, 2015. DNA Barcodes of Lepidoptera Reared from Yawan, Papua New Guinea. Proceedings of the Entomological Society of Washington 117: 247 - 250, DOI: https://doi.org/10.4289/0013-8797.117.2.247" title="DNA Barcodes of Lepidoptera Reared from Yawan, Papua New Guinea." url="https://doi.org/10.4289/0013-8797.117.2.247" volume="117" year="2015">Miller et al. 2015</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7B43142C32A1719AF94CD303703325E7" author="Mally, R" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="100 - 108" refId="B32" refString="Mally, R, Leger, T, Vairappan, CS, Sutton, S, Nuss, M, 2017. Discovery of another fern-feeding group of moths: the larvae of Hoploscopini (Insecta: Lepidoptera: Pyraloidea) from Borneo. Raffles Bulletin of Zoology 65: 100 - 108" title="Discovery of another fern-feeding group of moths: the larvae of Hoploscopini (Insecta: Lepidoptera: Pyraloidea) from Borneo." volume="65" year="2017">Mally et al. 2017</bibRefCitation>
|
||
). Another undescribed
|
||
<taxonomicName id="FD532020F319AABF710A0172D90FC637" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1448CDE55CA57A0E693944C03303DD01" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
species (sample USNM_ENT_00739239) from Papua New Guinea is reported from
|
||
<taxonomicName id="1A9F16C9A826DC3094982D507AD44950" authorityName="Holtt" authorityYear="1974" baseAuthorityName="L." class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sphaerostephanos unitus" order="Polypodiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="unitus">
|
||
<emphasis id="8EAAB91AAEE89F13D70DC2B85BA642EE" italics="true" pageId="0" pageNumber="1">Sphaerostephanos unitus</emphasis>
|
||
</taxonomicName>
|
||
(Linnaeus, 1759) Holttum, 1794 (
|
||
<taxonomicName id="CF173D5B6C6544F8D847118CCC559391" class="Polypodiopsida" family="Thelypteridaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="family">Thelypteridaceae</taxonomicName>
|
||
) (ibid). Lastly, one undescribed
|
||
<taxonomicName id="2AE7B7398685B9ECF9D65E4CE7482556" authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A94FF84CDBB205234069488017EE4D53" italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
species from Borneo is reported from the fern
|
||
<taxonomicName id="8CCD74F2F851E13A12489B907EF44931" class="Polypodiopsida" family="Gleicheniaceae" genus="Dicranopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dicranopteris linearis" order="Gleicheniales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="linearis">
|
||
<emphasis id="6A47DBC8271A8944315030176CBB90B3" italics="true" pageId="0" pageNumber="1">Dicranopteris linearis</emphasis>
|
||
</taxonomicName>
|
||
(Burman, 1768) Underwood, 1907 (
|
||
<taxonomicName id="FE83FECFBB92BF98F1FF20D8C5D0F061" class="Polypodiopsida" family="Gleicheniaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Gleicheniales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="family">Gleicheniaceae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="8DFDDA8819AFA40E9D65CD61F112A95B" author="Mally, R" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="100 - 108" refId="B32" refString="Mally, R, Leger, T, Vairappan, CS, Sutton, S, Nuss, M, 2017. Discovery of another fern-feeding group of moths: the larvae of Hoploscopini (Insecta: Lepidoptera: Pyraloidea) from Borneo. Raffles Bulletin of Zoology 65: 100 - 108" title="Discovery of another fern-feeding group of moths: the larvae of Hoploscopini (Insecta: Lepidoptera: Pyraloidea) from Borneo." volume="65" year="2017">Mally et al. 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |