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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.415.6406" ID-GBIF-Dataset="aefb1cac-93e1-4a01-8735-f8ba09948390" ID-PMC="PMC4089821" ID-Pensoft-Pub="1313-2970-415-81" ID-PubMed="25009425" ID-ZBK="372DF48DD1634742AABF5D7E4913050C" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-415-81" ModsDocOrigin="ZooKeys 415" ModsDocTitle="A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant &amp; Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina)" checkinTime="1451245791699" checkinUser="pensoft" docAuthor="Kaminski, Marcin Jan" docDate="2014" docId="BEB3E9E2EEC2729A140589B9DE697FF1" docLanguage="en" docName="ZooKeys 415: 81-132" docOrigin="ZooKeys 415" docSource="http://dx.doi.org/10.3897/zookeys.415.6406" docTitle="Ectateus crenatus Fairmaire 1897" docType="treatment" docVersion="3" lastPageNumber="92" masterDocId="FF91FFC18F381D1915336F07FFCA965D" masterDocTitle="A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant &amp; Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina)" masterLastPageNumber="132" masterPageNumber="81" pageNumber="90" updateTime="1668158650439" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant &amp; Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaminski, Marcin Jan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>415</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>132</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.415.6406</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.415.6406</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-415-81</mods:identifier>
<mods:identifier type="ZBK">372DF48DD1634742AABF5D7E4913050C</mods:identifier>
<mods:identifier type="ZooBank">372DF48DD1634742AABF5D7E4913050C</mods:identifier>
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<treatment ID-GBIF-Taxon="152053540" LSID="urn:lsid:plazi:treatment:BEB3E9E2EEC2729A140589B9DE697FF1" httpUri="http://treatment.plazi.org/id/BEB3E9E2EEC2729A140589B9DE697FF1" lastPageId="11" lastPageNumber="92" pageId="9" pageNumber="90">
<subSubSection pageId="9" pageNumber="90" type="nomenclature">
<paragraph pageId="9" pageNumber="90">
<taxonomicName LSID="http://species-id.net/wiki/Ectateus_crenatus" authority="Fairmaire, 1897" authorityName="Fairmaire" authorityYear="1897" class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus crenatus" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="crenatus">Ectateus crenatus (Fairmaire, 1897)</taxonomicName>
Figs 2, 8, 12, 34-36, 41, 46
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="90" type="reference_group">
<paragraph pageId="9" pageNumber="90">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus crenatus" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="crenatus">Selinus crenatus</taxonomicName>
Fairmaire, 1897: 121. -
<bibRefCitation author="Gebien, H" journalOrPublisher="Berlin" pageId="40" pageNumber="121" pagination="167 - 354" title="W. Junk, S. Schenkling, Coleopterorum Catalogus" year="1910">Gebien 1910</bibRefCitation>
: 277,
<bibRefCitation author="Gebien, H" journalOrPublisher="Mitteilungen der Muncher Entomologischen Gesellschaft" pageId="41" pageNumber="122" title="Katalog der Tenebrioniden. Teil II" volume="28" year="1938">1938</bibRefCitation>
: 297.
</paragraph>
<paragraph pageId="9" pageNumber="90">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus crenatus" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="crenatus">Ectateus crenatus</taxonomicName>
(Fairmaire, 1897). -
<bibRefCitation author="Koch, C" journalOrPublisher="Bruxelles" pageId="42" pageNumber="123" title="Exploration du Parc National de l'Upemba. II. Tenebrionidae (Coleoptera, Polyphaga), Opatrinae, First part: Platynotini, Litoborini and Loensini" year="1956">Koch 1956</bibRefCitation>
: 235,
<bibRefCitation author="Iwan, D" journalOrPublisher="Genus" pageId="41" pageNumber="122" pagination="219 - 323" title="Catalogue of the World Platynotini (Coleoptera: Tenebrionidae)." volume="13" year="2002 b">Iwan 2002b</bibRefCitation>
: 265;
<bibRefCitation author="Iwan, D" journalOrPublisher="Annales Zoologici" pageId="42" pageNumber="123" pagination="725 - 731" title="Platymedvedevia, a new genus of Ectateus group from Tropical Africa (Coleoptera: Tenebrionidae: Platynotina)." volume="57" year="2007">Iwan and Banaszkiewicz 2007</bibRefCitation>
: 728.
</paragraph>
<paragraph pageId="9" pageNumber="90">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus monardi" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="monardi">Selinus monardi</taxonomicName>
Kaszab, 1951: 2 (syn. nov.)
</paragraph>
<paragraph pageId="9" pageNumber="90">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus latipennis" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="latipennis">Ectateus latipennis</taxonomicName>
Koch, 1956: 234 (syn. nov.). -
<bibRefCitation author="Iwan, D" journalOrPublisher="Annales Zoologici" pageId="41" pageNumber="122" pagination="1 - 149" title="Generic classification of the tribe Platynotini (Coleoptera: Tenebrionidae), with notes on phylogeny." volume="52" year="2002 a">Iwan 2002a</bibRefCitation>
: 67,
<bibRefCitation author="Iwan, D" journalOrPublisher="Genus" pageId="41" pageNumber="122" pagination="219 - 323" title="Catalogue of the World Platynotini (Coleoptera: Tenebrionidae)." volume="13" year="2002 b">2002b</bibRefCitation>
: 266.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="90" type="notes">
<paragraph pageId="9" pageNumber="90">Notes.</paragraph>
<paragraph pageId="9" pageNumber="90">
While describing
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus latipennis" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="latipennis">Ectateus latipennis</taxonomicName>
, Koch has noted that types of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus crenatus" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="crenatus">Ectateus crenatus</taxonomicName>
were unknown to him. The characters used by Koch to separate those two species (body size, pronotum structure) were based only on the
<bibRefCitation author="Fairmaire, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="40" pageNumber="121" pagination="109 - 152" title="Coleopteres nouveaux de l'Afrique intertropicale et australe." volume="66" year="1897">Fairmaire (1897)</bibRefCitation>
description. During the examination of available material I have not found any consistent morphological characters to separate those two species. Therefore, I propose to consider
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus latipennis" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="latipennis">Ectateus latipennis</taxonomicName>
as a synonim of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus crenatus" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="crenatus">Ectateus crenatus</taxonomicName>
.
</paragraph>
<paragraph pageId="9" pageNumber="90">
The examination of the type material representing
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus monardi" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="monardi">Selinus monardi</taxonomicName>
resulted in similar conclusions - there are no consistent morphological characters to separate it from
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus crenatus" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="crenatus">Ectateus crenatus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90" type="studied material">
<paragraph pageId="9" pageNumber="90">Studied material.</paragraph>
<paragraph lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90">
Three specimens with
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
labels are available.
<bibRefCitation author="Fairmaire, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="40" pageNumber="121" pagination="109 - 152" title="Coleopteres nouveaux de l'Afrique intertropicale et australe." volume="66" year="1897">Fairmaire (1897)</bibRefCitation>
do not specify the number of individuals on which he described this species. Lectotype designation is needed to fix the taxonomic status of the genus and the species. Lectotype designated here, male (MNHN):
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
, &quot;Museum Paris, 1906, Coll. L. Fairmaire&quot;, &quot;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus crenatus" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="crenatus">
<pageBreakToken pageId="10" pageNumber="91" start="start">Selinus</pageBreakToken>
crenatus
</taxonomicName>
&quot;; Paralectotypes, male (MNHN), female (MNHN):
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
, &quot;Museum Paris, 1906, Coll. L. Fairmaire&quot;, &quot;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus crenatus" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="crenatus">Selinus crenatus</taxonomicName>
Farim 1896, Congo&quot;, female (MNHN): &quot;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus crenatus" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="crenatus">Selinus crenatus</taxonomicName>
Fm n. sp.&quot;,
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
, &quot;Museum Paris, 1906, Coll. L. Fairmaire&quot;,
<normalizedToken originalValue="“Congo”">&quot;Congo&quot;</normalizedToken>
. Other material: male (MRAC):
<normalizedToken originalValue="“Musée">&quot;Musee</normalizedToken>
du Congo, Barumbu - VIII-1925, (J.
<normalizedToken originalValue="Ghesquière">Ghesquiere</normalizedToken>
), S.A.R. Prince
<normalizedToken originalValue="Léopold”">Leopold&quot;</normalizedToken>
(Holotype of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus latipennis" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="latipennis">Ectateus latipennis</taxonomicName>
Koch, 1956), female (RBINS): &quot;Ibembo, Itimbiri&quot; (Allotype of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Ectateus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ectateus latipennis" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="latipennis">Ectateus latipennis</taxonomicName>
Koch, 1956), male (MNHN): &quot;Kamerun, Joko&quot; (Holotype of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus monardi" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="monardi">Selinus monardi</taxonomicName>
Kaszab, 1951), male and female (MNHN): &quot;Joko, Kamerun&quot; (Paratype and allotype of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selinus monardi" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="monardi">Selinus monardi</taxonomicName>
Kaszab, 1951), male and female (MNHN): &quot;Cameroun, Dr. G. Nonveiller&quot;, male and female (MNHN):
<normalizedToken originalValue="“Muséum">&quot;Museum</normalizedToken>
Paris, Cameroun, B. de
<normalizedToken originalValue="Miré”">Mire&quot;</normalizedToken>
, 3 males and 2 females (MNHN):
<normalizedToken originalValue="“Ogodué">&quot;Ogodue</normalizedToken>
,
<normalizedToken originalValue="Lambaréné">Lambarene</normalizedToken>
, R. Ellenberger 1913&quot;, male (MNHN): &quot;Congo
<normalizedToken originalValue="Français">Francais</normalizedToken>
, region
<normalizedToken originalValue="D’">D'</normalizedToken>
ouesso, Bassin
<normalizedToken originalValue="N`Gokko-Sanga">N'Gokko-Sanga</normalizedToken>
, Dr. J. Gravot 1906&quot;, male (MNHN): &quot;Congo
<normalizedToken originalValue="Français">Francais</normalizedToken>
, Talagouga
<normalizedToken originalValue="Prés">Pres</normalizedToken>
<normalizedToken originalValue="N`Jolé">N'Jole</normalizedToken>
, legit R. Ellenberger, E. Haug 1906&quot;, male and female (MNHN):
<normalizedToken originalValue="“Ogooué">&quot;Ogooue</normalizedToken>
, Sam-Kita&quot;, 3 females (MNHN): &quot;Congo,
<normalizedToken originalValue="Ogooué">Ogooue</normalizedToken>
, Sam Kita, R. Ellenberger 1910&quot;, 2 males and 2 females (MNHN): &quot;Gabon, Libreville et env.&quot;, male and female (MNB): &quot;Span. Guinea, Nkolentangan, XI. 07-V.08, G.
<normalizedToken originalValue="Teßmann">Tessmann</normalizedToken>
S.G.&quot;, female (MNHN): &quot;Benito, Congo Franc.&quot;, female (MNHN): &quot;Gabon, Tholon&quot;, male and female (MNHN): &quot;XII.1970, Mbalmayo, CAMEROUN, Mbarga leg.&quot;, female (MNHN): &quot;Congo belge Centr., Kassai, Edm. Taymans, 1904&quot;, male (MHNG): &quot;CAMEROUN VII.83, Etoubi Assok,
<normalizedToken originalValue="à">a</normalizedToken>
la
<normalizedToken originalValue="lumière">lumiere</normalizedToken>
, F. Notari&quot;, female (MNB): &quot;Neu-Kamerun, No. 3337-52, Tessmann S.G.&quot;, female (MRAC): &quot;Coll. Mus. Congo, Mayumbe: Terr. Tshela,
<normalizedToken originalValue="rég">reg</normalizedToken>
. de Mabuba VI-1958, Dr R. Laurent&quot;, male (MRAC): &quot;Mus. Roy. Afr. Centr.,
<normalizedToken originalValue="Guinée">Guinee</normalizedToken>
Esp: Bata,
<normalizedToken originalValue="Rév">Rev</normalizedToken>
. P. Basilio&quot;, male (MNHN): &quot;Kuilu, Fr. Congo., Mocquerys, 1892&quot;, male (RBINS): &quot;Chutes de Samlia, Riv N. Gamie, Mocquereys&quot;.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="92" pageId="10" pageNumber="91" type="redescription">
<paragraph pageId="10" pageNumber="91">Redescription.</paragraph>
<paragraph pageId="10" pageNumber="91">Habitus as in Fig. 46. Body length = 7.0-9.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1-1.2; length ratio elytra / the middle of pronotum = 2.4-2.6).</paragraph>
<paragraph pageId="10" pageNumber="91">Dorsal side of head shiny, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 5.7-6.5). Mentum with median part narrow. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0-1.2). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.1-1.3). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8-3.0).</paragraph>
<paragraph pageId="10" pageNumber="91">Pronotal disc transverse (middle of pronotum length / width ratio = 0.5-0.6); shiny, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotal disc sinusoidal. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded. Pronotal hypomera dull; without punctures.</paragraph>
<paragraph pageId="10" pageNumber="91">Elytra oblong (elytra length / width ratio = 1.1-1.2). Elytral striae with coarse punctures (Fig. 8); intervals convex, with transverse sculpture and fine puncturation (Fig. 8). Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; situated in a depression.</paragraph>
<paragraph lastPageId="11" lastPageNumber="92" pageId="10" pageNumber="91">
Intercoxal process protruding towards mesoventrite, peaked at the apex, slightly saddle-like. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of
<pageBreakToken pageId="11" pageNumber="92" start="start">metaventrite</pageBreakToken>
= 2.1-2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).
</paragraph>
<paragraph pageId="11" pageNumber="92">Male legs. Protarsi slightly narrow. Protibiae as in Fig. 34. Mesotibiae and mesofemorae with large denticle (Figs 35-36). Metafemorae with an hair fringe. Female legs simple.</paragraph>
<paragraph pageId="11" pageNumber="92">Male genitalia. Parameres narrowest in the half of their length; length equal to 0.2 of the rest of aedeagal tegmen. Clavae hook-shaped. Female genitalia. Paraproct equal to coxites. Bursa copulatrix without sclerites. Spermatheca with narrow ducts.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="92" type="distribution">
<paragraph pageId="11" pageNumber="92">Distribution.</paragraph>
<paragraph pageId="11" pageNumber="92">This species has been collected in the following ecoregions of Central Africa (Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Gabonese Republic, Republic of the Congo): Atlantic Equatorial coastal forests, Northern Congolian forest-savanna mosaic, Northwestern Congolian lowland forests, Southern Congolian forest-savanna mosaic, Western Congolian forest-savanna mosaic (Fig. 41).</paragraph>
</subSubSection>
</treatment>
</document>
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