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<document id="10D6A8C8ACA51128DF8F16714D2D79FB" ID-DOI="10.5252/zoosystema2024v46a25" ID-ISSN="1638-9387" ID-Zenodo-Dep="13907152" ID-ZooBank="D4AC1F89-AC34-43C4-9761-3F2015A02265" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728463297646" checkinUser="plazi" docAuthor="Segniagbeto, Gabriel Hoinsoudé, Mark-Oliver, Annemarie Ohler, Luca, Mark-Oliver Rödel, Alain, Luca Luiselli &amp; Dubois, Alain" docDate="2024" docId="03F187A4FFAEFFF8A9B0FC48FBE5F829" docLanguage="en" docName="zoosystema.2024.46.25.pdf" docOrigin="Zoosystema 46 (25)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a25.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Xenopus tropicalis Agote" docType="treatment" docVersion="2" lastPageNumber="655" masterDocId="FFC8FFDCFFB4FFE2AA73FFBDFFEFFFCF" masterDocTitle="Amphibians of Togo: taxonomy, distribution and conservation status" masterLastPageNumber="670" masterPageNumber="631" pageNumber="655" updateTime="1728463900544" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="461827AE283BC81231F83BB2CE791401">Amphibians of Togo: taxonomy, distribution and conservation status</mods:title>
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<mods:namePart id="167548620D4C8F6A1A835423420029F9">Segniagbeto, Gabriel Hoinsoudé</mods:namePart>
<mods:affiliation id="FE1EF967A8A30F329D07EE1173FD8D62">Laboratory of Ecology and Ecotoxicology, Faculty of Sciences, University of Lomé, BP 6057 Lomé, Togo, BP 1515, Lomé (Togo) and Institute for Development, Ecology, Conservation and Cooperation, Rome (Italy) gsegniagbeto @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="3F252AB6E3EF87DE41A4CE894D390F82">Mark-Oliver, Annemarie Ohler</mods:namePart>
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<mods:namePart id="8782D82E8D43ABE53BAE381C68E10997">Luca, Mark-Oliver Rödel</mods:namePart>
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<mods:namePart id="AA8F49725FA4F9DB350477E3D9134240">Alain, Luca Luiselli</mods:namePart>
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<mods:namePart id="1126D79EDEA762C82352D9404D1040E0">Dubois, Alain</mods:namePart>
<mods:affiliation id="6678095E2ACF7FC40E6E148A0C05D16F">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE-PSL, Université des Antilles, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr</mods:affiliation>
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<mods:date id="1F1EC9E4B1CF3EA7C09342E59C2279FA">2024</mods:date>
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<taxonomicName id="4C584D31FFAEFFF8A9B0FC48FAF6FBC1" ID-CoL="5CBCN" authority="(Gray, 1864)" authorityName="Agote" baseAuthorityName="Gray" baseAuthorityYear="1864" box="[963,1305,1012,1039]" class="Amphibia" family="Pipidae" genus="Xenopus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="655" phylum="Chordata" rank="species" species="tropicalis">
<emphasis id="B92CEAA0FFAEFFF8A9B0FC48FB68FBC0" bold="true" box="[963,1159,1012,1039]" italics="true" pageId="26" pageNumber="655">Xenopus tropicalis</emphasis>
(Gray, 1864)
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<paragraph id="8BE736B2FFAEFFF8AE46FBA9FB49FBE1" blockId="26.[963,1305,1012,1070]" box="[1077,1190,1044,1070]" pageId="26" pageNumber="655">
(
<figureCitation id="13632A37FFAEFFF8AE4EFBA9FB71FBE1" box="[1085,1182,1044,1070]" captionStart="FIG" captionStartId="28.[132,143,1789,1807]" captionTargetBox="[132,1455,215,1752]" captionTargetId="figure-209@28.[698,1267,756,1300]" captionTargetPageId="28" captionText="FIG. 12. — Representatives of anuran species from Togo in life: A, Phrynomantis microps Peters, 1875 (Tsévié); B, Xenopus fischbergi Evans, Carter, Greenbaum, Gvoždík,Kelley,McLaughlin, Pauwels,Portik, Stanley, Tinsley,Tobias &amp; Blackburn, 2015 (Siou);C, Xenopus tropicalis (Gray, 1864) (Agoté); D, Ptychadena bibroni (Hallowell, 1845) (Kovié); E, Ptychadena mascareniensis (Duméril &amp; Bibron, 1841) (Kovié); F, Ptychadena oxyrhynchus (Smith, 1849) (Tchamba)," figureDoi="http://doi.org/10.5281/zenodo.13907186" httpUri="https://zenodo.org/record/13907186/files/figure.png" pageId="26" pageNumber="655">Fig. 12C</figureCitation>
)
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<subSubSection id="C3426539FFAEFFF8A95EFBEFFC36FB4C" pageId="26" pageNumber="655" type="materials_examined">
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. —
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<collectingCountry id="F34F7622FFAEFFF8AE37FBECFB95FBA6" box="[1092,1146,1105,1129]" name="Togo" pageId="26" pageNumber="655">Togo</collectingCountry>
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<specimenCount id="9D5EFD3BFFAEFFF8AEEAFBECFB2DFBA3" box="[1177,1218,1105,1132]" count="4" pageId="26" pageNumber="655" type="female">4 ♀</specimenCount>
; Agoté; MNHN-RA-2006.2172-2175.
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. — Medium to large frog (SVL
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), with flattened body. Snout rounded. Head very small, wider than long (HW 26-29% SVL; HL 21-26% SVL). Very small eyes and eyelids (EL
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diameter); a very small subocular tentacle, about one third of eye dimeter. Tympanum indistinct. Hind leg rather short (TL 36-50% SVL). Webbing complete, no free phalanges. Four horny claws, three at the toes I, II, III and one covering the metatarsal tubercle. Skin smooth, lateral line system with 18-20 tubercles from eyelid to rear part of flanks, also on neck region and border of lower jaw; belly smooth.
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. — The dorsal colouration is dark brown or black depending on the habitat. The dorsal part of forearm, thigh and tibia is also dark. The belly is lighter.
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. — Differences between males and females are little marked (
<bibRefCitation id="EFC94B43FFAEFFF8A996F9FEFB07F994" author="ARNOULT J. &amp; LAMOTTE M." box="[997,1256,1603,1627]" pageId="26" pageNumber="655" pagination="270 - 306" refId="ref32423" refString="ARNOULT J. &amp; LAMOTTE M. 1968. - Les Pipidae de l' Ouest africain et du Cameroun. Bulletin de l'Institut Fondamental d'Afrique Noire (A) 30: 270 - 306." type="journal article" year="1968">Arnoult &amp; Lamotte 1968</bibRefCitation>
). Males are thinner and smaller than females, the outer sides of their fingers carry black nuptial pads and their dermal lobes above vent are larger than in females. The head is narrower in males than in females.
</paragraph>
<paragraph id="8BE736B2FFAEFFF8A95EF97BFBE5F829" blockId="26.[811,1457,1734,2022]" pageId="26" pageNumber="655">
HABITATS AND DISTRIBUTION. —
<taxonomicName id="4C584D31FFAEFFF8AE1FF97AFAE0F911" authorityName="Agote" baseAuthorityName="Gray" baseAuthorityYear="1864" box="[1132,1295,1734,1758]" class="Amphibia" family="Pipidae" genus="Xenopus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="655" phylum="Chordata" rank="species" species="tropicalis">
<emphasis id="B92CEAA0FFAEFFF8AE1FF97AFAE0F911" box="[1132,1295,1734,1758]" italics="true" pageId="26" pageNumber="655">Xenopus tropicalis</emphasis>
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is a forest species that enters the savannah zone along gallery forests (
<bibRefCitation id="EFC94B43FFAEFFF8AF59F95DFA4FF936" author="RODEL M. - O." box="[1322,1440,1760,1785]" pageId="26" pageNumber="655" refId="ref37146" refString="RODEL M. - O. 2000. - Herpetofauna of West Africa. Vol 1. Amphibians of theWest African savanna. ̿ rankfurt am Main, Chimaira, 335 p." type="book" year="2000">Rödel 2000</bibRefCitation>
). It is most frequent in the ecological zone IV (Kpélé and Badou regions). It lives in ponds or abandoned wells where it hides in the mud during the dry season. It is widespread in humid forests and savannahs with forest areas or forest galleries in Kpélé and Akposso.
<specimenCount id="9D5EFD3BFFAEFFF8A9E2F8D8FB8CF8B2" box="[913,1123,1893,1917]" count="59" pageId="26" pageNumber="655" type="generic">Fifty-nine specimens</specimenCount>
of this species were collected by
<bibRefCitation id="EFC94B43FFAEFFF8A95DF83DFC24F858" author="BOURGAT R." box="[814,971,1919,1944]" pageId="26" pageNumber="655" pagination="597 - 624" refId="ref33022" refString="BOURGAT R. 1979. - Trematodes d'amphibiens du Togo. Bulletin du Museum national d'Histoire naturelle (4 e serie) 3: 597 - 624." type="journal article" year="1979">Bourgat (1979)</bibRefCitation>
in Kandé (Kanté), Lamakara (
<collectingRegion id="499CF850FFAEFFF8AF75F8C2FAD5F858" box="[1286,1338,1919,1943]" country="Togo" name="Kara" pageId="26" pageNumber="655">Kara</collectingRegion>
), Niamtougou, Sotougoua (Sotouboua), Badou and Klouto. The occurrence of this species in the ecological zones I, II, III and IV is known from these specimens.
</paragraph>
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