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<mods:title id="C6C77EAAAFF8F53318814A8DE637E8D2">Amphibians of Togo: taxonomy, distribution and conservation status</mods:title>
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<mods:affiliation id="CDABAFEAD54D389D99121F6C9FEFD1B5">Laboratory of Ecology and Ecotoxicology, Faculty of Sciences, University of Lomé, BP 6057 Lomé, Togo, BP 1515, Lomé (Togo) and Institute for Development, Ecology, Conservation and Cooperation, Rome (Italy) gsegniagbeto @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="1CBB38AF4D290FCF2AF8C68F9F05E0B3">Luca, Mark-Oliver Rödel</mods:namePart>
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<mods:namePart id="A4B8037BDEB3F6EC4F8C46B64FA8046C">Dubois, Alain</mods:namePart>
<mods:affiliation id="2A5346A12AD53A1CADEAC88153F3325A">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE-PSL, Université des Antilles, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr</mods:affiliation>
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<taxonomicName id="4C584D31FFA5FFF3AA8EFB2EFD61FB60" ID-CoL="3NQTY" authority="Gunther, 1859" authorityName="Gunther" authorityYear="1859" box="[253,654,1171,1199]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="17" pageNumber="646" phylum="Chordata" rank="species" species="guttulatus">
<emphasis id="B92CEAA0FFA5FFF3AA8EFB2EFE09FB61" bold="true" box="[253,486,1171,1198]" italics="true" pageId="17" pageNumber="646">Hyperolius guttulatus</emphasis>
Günther, 1859
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. —
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<collectingCountry id="F34F7622FFA5FFF3ABE3FB70FE2AFB2A" box="[400,453,1229,1253]" name="Togo" pageId="17" pageNumber="646">Togo</collectingCountry>
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,
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;Kloto; MNHN- RA-1993.6106, MNHN-RA-1993.6107, MNHN-RA-1993.6110-6118, MNHN-RA-1993.6108, MNHN- RA-1993.6109.
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. — Medium sized tree frog (larger than most other Togolese tree frogs) (SVL
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,
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), with moderately slender but fairly robust body, wider anteriorly than posteriorly. Head slightly longer than broad (HW 29-34% SVL, HL 33-36% SVL). Snout rounded.Tympanum distinct (TYD 2-5% SVL). Tibia short (TL 44-53% SVL). Webbing large, leaving up to two phalanges free; webbing formula: I ½ 1 ½ II ½ 1 ½ III ½ 1 IV 1 ½ ½ V. Skin smooth on the snout and strongly shagreen on the back and flanks; belly and chest granular; some warts at the corner of the mouth.
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. — The dorsal colouration of females is extremely variable, but fades in alcohol. Males are mostly uniformly greenish. For colour descriptions and figures see
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,
<bibRefCitation id="EFC94B43FFA5FFF3A8FBF914FD54F90E" author="SCHIOTZ A." box="[648,699,1705,1729]" pageId="17" pageNumber="646" pagination="1 - 346" refId="ref38284" refString="SCHIOTZ A. 1967. - The treefrogs (Rhacophoridae) of West Africa. Spolia zoologica Musei haunienses 25: 1 - 346." type="journal article" year="1967">1967</bibRefCitation>
,
<bibRefCitation id="EFC94B43FFA5FFF3A8B5F914FD15F90E" author="SCHIOTZ A." box="[710,762,1705,1729]" pageId="17" pageNumber="646" refId="ref38309" refString="SCHIOTZ A. 1999. - Treefrogs of Africa. Chimaira, ̿ rankfurt am Main, 352 p." type="book" year="1999">1999</bibRefCitation>
),
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,
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,
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Kouamé
<emphasis id="B92CEAA0FFA5FFF3AB9CF979FDF4F914" box="[495,539,1731,1755]" italics="true" pageId="17" pageNumber="646">et al.</emphasis>
(2015)
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and
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. This species differs from other large and robust
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<emphasis id="B92CEAA0FFA5FFF3A890F962FF24F8DF" italics="true" pageId="17" pageNumber="646">Hyperolius</emphasis>
species
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by a dark canthal stripe.
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. — Males have a subgular, bluish-green vocal sac with a roundish gland. Females are distinctly larger.
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. —
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<emphasis id="B92CEAA0FFA5FFF3ABBBF8C0FD62F85A" box="[456,653,1917,1941]" italics="true" pageId="17" pageNumber="646">Hyperolius guttulatus</emphasis>
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is essentially found in swampy forests. Our localities represent the typical habitat of the species according to
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,
<bibRefCitation id="EFC94B43FFA5FFF3A837F80FFD2EF806" author="AMIET J. - L." box="[580,705,1969,1994]" pageId="17" pageNumber="646" pagination="30 - 107" refId="ref32271" refString="AMIET J. - L. 1975. - Ecologie et distribution des Amphibiens Anoures de la region de Nkongsamba (Cameroun). Annales de la Faculte des Sciences Yaounde 20: 30 - 107." type="journal article" year="1975">Amiet (1975</bibRefCitation>
,
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) and
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.
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