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<document id="2C761E0576B785D367A93F8A14153700" ID-DOI="10.1093/zoolinnean/zlad078" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240120" ID-ZooBank="5D6C7EBF-E568-4100-9364-2DD357003878" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1716361325736" checkinUser="plazi" docAuthor="Marceniuk, Alexandre Pires, Oliveira, Claudio &amp; Ferraris Jr, Carl J." docDate="2024" docId="03F32B77FFCAFF8F0943FD37FDEB4B75" docLanguage="en" docName="zlad078.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad078" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Paracinetodus Marceniuk &amp; Oliveira &amp; Ferraris Jr 2024, gen. nov." docType="treatment" docVersion="3" lastPageNumber="472" masterDocId="FFCA530FFFE7FFA10D44FFD6FFAB4A1E" masterDocTitle="A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data" masterLastPageNumber="476" masterPageNumber="426" pageNumber="471" updateTime="1716560620312" updateUser="guilherme">
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<mods:title id="1685329A712D5872E494685A388165AF">A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data</mods:title>
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<mods:namePart id="1D435938FD141FA02FD4BA9194406035">Marceniuk, Alexandre Pires</mods:namePart>
<mods:affiliation id="14428E5ED632DB6BD28F62B3759E305D">Programa de Pós-Graduação em Ciências Biológicas, Universidade Federal da Paraíba, conjunto Presidente Castelo Branco III, João Pessoa, PB 58050 - 000, Brazil &amp; Programa de Pós-Graduação em Ecologia e Conservação, Universidade Estadual da Paraíba, Campina Grande, PB 58429 - 500, Brazil &amp; Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
<mods:nameIdentifier id="4C8218D4BB8D36E61C0516B47B7AC27C" type="email">a_marceniuk@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="27A61C5A2F9820A9F87900F5253604A7">Oliveira, Claudio</mods:namePart>
<mods:affiliation id="E73878E9ECFE0FFFCE40A371067A752F">Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
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<mods:namePart id="56ACFB646CAAAA0D91BC9263657744BA">Ferraris Jr, Carl J.</mods:namePart>
<mods:affiliation id="936821E22B94B76000CE5311DCB6B5E5">545 NE Floral Pl., Portland, OR 97232, USA</mods:affiliation>
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<mods:title id="ADD2B7428D93AB75E009A365E55F44F6">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="8F3BE37C9E401BE7A7ABFB38D054452A">2024</mods:date>
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<mods:number id="25949A84CE92A52B50C3BC97A3432DDA">2023-08-21</mods:number>
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<treatment id="03F32B77FFCAFF8F0943FD37FDEB4B75" LSID="urn:lsid:plazi:treatment:03F32B77FFCAFF8F0943FD37FDEB4B75" httpUri="http://treatment.plazi.org/id/03F32B77FFCAFF8F0943FD37FDEB4B75" lastPageId="46" lastPageNumber="472" pageId="45" pageNumber="471">
<subSubSection id="C340C9EAFFCAFF8C0943FD37FB5F48E2" box="[1031,1268,737,764]" pageId="45" pageNumber="471" type="nomenclature">
<paragraph id="8BE59A61FFCAFF8C0943FD37FB5F48E2" blockId="45.[1031,1268,737,764]" box="[1031,1268,737,764]" pageId="45" pageNumber="471">
<heading id="D0AD2D0DFFCAFF8C0943FD37FB5F48E2" bold="true" box="[1031,1268,737,764]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="471" reason="2">
<emphasis id="B92E4673FFCAFF8C0943FD37FB5F48E2" bold="true" box="[1031,1268,737,764]" pageId="45" pageNumber="471">
<taxonomicName id="4C5AE1E2FFCAFF8C0943FD37FB3E48E2" authority="Marceniuk &amp; Oliveira &amp; Ferraris Jr, 2024" authorityName="Marceniuk &amp; Oliveira &amp; Ferraris Jr" authorityYear="2024" box="[1031,1173,737,764]" family="Ariidae" genus="Paracinetodus" kingdom="Animalia" order="Siluriformes" pageId="45" pageNumber="471" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92E4673FFCAFF8C0943FD37FB3E48E2" bold="true" box="[1031,1173,737,764]" italics="true" pageId="45" pageNumber="471">Paracinetodus</emphasis>
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<taxonomicNameLabel id="A21DFB08FFCAFF8C09DDFD34FB5F48E2" box="[1177,1268,738,764]" pageId="45" pageNumber="471" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C340C9EAFFCAFF8C095CFCD8FB694961" pageId="45" pageNumber="471" type="description">
<paragraph id="8BE59A61FFCAFF8C095CFCD8FB484936" blockId="45.[1048,1251,782,809]" box="[1048,1251,782,809]" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C095CFCD8FB484936" bold="true" box="[1048,1251,782,809]" pageId="45" pageNumber="471">
(
<figureCitation id="136186E4FFCAFF8C0967FCD8FBD74936" box="[1059,1148,782,808]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[129,194,1805,1829]" captionStartId-1="3.[825,890,1803,1827]" captionStartId-2="4.[113,178,1813,1837]" captionTargetBox-0="[152,776,143,1771]" captionTargetBox-1="[843,1439,143,1771]" captionTargetBox-2="[151,708,146,1775]" captionTargetId-0="figure-8@3.[152,744,149,1755]" captionTargetId-1="figure-61@3.[851,1439,143,1771]" captionTargetId-2="figure-643@4.[149,717,144,1776]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-1="Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-2="Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." figureDoi-1="http://doi.org/10.5281/zenodo.11240122" figureDoi-2="http://doi.org/10.5281/zenodo.11240126" httpUri-1="https://zenodo.org/record/11240122/files/figure.png" httpUri-2="https://zenodo.org/record/11240126/files/figure.png" pageId="45" pageNumber="471">Figs 1–3</figureCitation>
,
<figureCitation id="136186E4FFCAFF8C09C3FCD8FB184936" box="[1159,1203,782,808]" captionStart="Figure 38" captionStartId="38.[117,182,1915,1939]" captionTargetBox="[247,1333,150,1886]" captionTargetId="figure-7@38.[245,1333,144,1888]" captionTargetPageId="38" captionText="Figure 38. Representative skulls, in dorsal view, and distribution of the subtribe Doiichthyina.A, Brustiarius nox, AUM 47488. B, Paracinetodus carinatus, AUM 47550.C, Cochlefelis spatula, AUM 50296. D, Nedystoma dayi, AUM 47500. E, Aceroichthys dioctes, AUM 47507. F, Pachyula crassilabris, AUM 47509. G, Pararius proximus, AUM 47486.H, Potamosilurus velutinus, AUM 47489.* marine, ** brackish waters, *** freshwater." figureDoi="http://doi.org/10.5281/zenodo.11240197" httpUri="https://zenodo.org/record/11240197/files/figure.png" pageId="45" pageNumber="471">38B</figureCitation>
,
<figureCitation id="136186E4FFCAFF8C09F9FCD8FB724936" box="[1213,1241,782,808]" captionStart="Figure 50" captionStartId="46.[113,178,1308,1332]" captionTargetBox="[114,1458,794,1281]" captionTargetId="figure-256@46.[114,1458,794,1281]" captionTargetPageId="46" captionText="Figure 50. Paracinetodus carinatus, photograph by Ricardo Betancur-R., Papua New Guinea and northern Australia, type species of the genus." figureDoi="http://doi.org/10.5281/zenodo.11240221" httpUri="https://zenodo.org/record/11240221/files/figure.png" pageId="45" pageNumber="471">50</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BE59A61FFCAFF8C0E7DFC9EFB694961" blockId="45.[825,1474,840,895]" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C0E7DFC9EFB8D4941" box="[825,1062,840,864]" italics="true" pageId="45" pageNumber="471">ZooBank registration:</emphasis>
<uri id="FFCB9663FFCAFF8C0902FC9EFB174961" pageId="45" pageNumber="471">
urn:lsid:zoobank.org:act:
<uuid id="FFFCA0B4FFCAFF8C0808FC9EFB174961" pageId="45" pageNumber="471">01D775F1- CA33-42B6-AE05-AC1D3E9DBE6C</uuid>
</uri>
.
</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFCAFF8C0E7DFC77FB7449A7" box="[825,1247,928,953]" pageId="45" pageNumber="471" type="type_taxon">
<paragraph id="8BE59A61FFCAFF8C0E7DFC77FB7449A7" blockId="45.[825,1247,928,953]" box="[825,1247,928,953]" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C0E7DFC77FC1E49A6" box="[825,949,929,952]" italics="true" pageId="45" pageNumber="471">
<typeStatus id="54E124C3FFCAFF8C0E7DFC77FCC249A6" box="[825,873,929,952]" pageId="45" pageNumber="471">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C5AE1E2FFCAFF8C0EFBFC77FB7049A7" authority="Weber, 1913" authorityName="Weber" authorityYear="1913" box="[959,1243,928,953]" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="45" pageNumber="471" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B92E4673FFCAFF8C0EFBFC77FBF849A6" box="[959,1107,929,952]" italics="true" pageId="45" pageNumber="471">Arius carinatus</emphasis>
<bibRefCitation id="EFCBE790FFCAFF8C091CFC76FB7049A7" author="Weber M &amp; de Beaufort LF" box="[1112,1243,928,953]" pageId="45" pageNumber="471" refId="ref32046" refString="Weber M, de Beaufort LF. The Fishes of the Indo-Australian Archipelago. II. Malacopterygii, Myctophoidea, Ostariophysi: I Siluroidea. Leiden: EJ Brill, 1913." type="book" year="1913">Weber, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFCAFF8C090BFC0FFABE4CB6" pageId="45" pageNumber="471" type="diagnosis">
<paragraph id="8BE59A61FFCAFF8C090BFC0FFB0749EC" blockId="45.[825,1474,985,1704]" box="[1103,1196,985,1010]" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C090BFC0FFB0749EC" box="[1103,1196,985,1010]" italics="true" pageId="45" pageNumber="471">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE59A61FFCAFF8C0E7DFC28FABE4CB6" blockId="45.[825,1474,985,1704]" pageId="45" pageNumber="471">Lateral horn of lateral ethmoid compressed and spatulate (11, 0&gt; 1); extrascapular subtriangular (37, 1&gt; 2); epioccipital contacting small narrow area of diagonal crest associated with neural spine of fourth vertebra (44, 2&gt; 0); ventral tip of subvertebral process acute (80, 0&gt; 2); basioccipital lateral process absent with anterior and posterior portions equally developed (83, 1&gt; 0); transcapular process at right angle to body axis (86, 0&gt; 1); transcapular process short and thick (87, 0&gt; 2); space between transcapular process and otic capsule small (89, 1&gt; 2); premaxilla narrow and long, length two to three times its width (120, 1&gt; 2); bony blade connecting posterolateral processes of urohyal absent (159, 0&gt; 1); posterolateral processes of urohyal two-thirds as long as distal portion of bone (162, 0&gt; 2); transverse crest associated with neural spine of fourth vertebra high (200, 0&gt; 1); median crest associated with neural spine of fourth vertebra high (202, 0&gt; 1); Müllerian ramus bony blade evident in more than one-half length (206, 0&gt; 2); ventral surfaces of parapophyses of fifth and sixth vertebrae conspicuously concave (212, 0&gt; 1); seventh vertebra free from ventral superficial ossification (214, 0&gt; 1); posterior process of cleithrum very long (224, 1&gt; 2); second dorsal cleithral process dorsally directed and parallel to first dorsal process (226, 0&gt; 1).</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFCAFF8C090FF917FA1E4D00" pageId="45" pageNumber="471" type="etymology">
<paragraph id="8BE59A61FFCAFF8C090FF917FB1A4CC5" blockId="45.[825,1474,1729,1823]" box="[1099,1201,1729,1755]" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C090FF917FB1A4CC5" box="[1099,1201,1729,1755]" italics="true" pageId="45" pageNumber="471">Etymology</emphasis>
</paragraph>
<paragraph id="8BE59A61FFCAFF8C0E7DF931FA1E4D00" blockId="45.[825,1474,1729,1823]" pageId="45" pageNumber="471">
Derived from the Greek ‘para’ for ‘near’, highlighting its morphological similarity with the genus
<emphasis id="B92E4673FFCAFF8C09C5F8D0FB434D00" box="[1153,1256,1798,1822]" italics="true" pageId="45" pageNumber="471">
<taxonomicName id="4C5AE1E2FFCAFF8C09C5F8D0FB4F4D00" authorityName="Ogilby" authorityYear="1898" box="[1153,1252,1798,1822]" family="Ariidae" genus="Cinetodus" kingdom="Animalia" order="Siluriformes" pageId="45" pageNumber="471" phylum="Chordata" rank="genus">Cinetodus</taxonomicName>
.
</emphasis>
Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFCAFF8C0971F8E8FB074D63" pageId="45" pageNumber="471" type="taxon_list">
<paragraph id="8BE59A61FFCAFF8C0971F8E8FB6D4D46" blockId="45.[825,1222,1854,1917]" box="[1077,1222,1854,1880]" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C0971F8E8FB6D4D46" box="[1077,1222,1854,1880]" italics="true" pageId="45" pageNumber="471">Included species</emphasis>
</paragraph>
<paragraph id="8BE59A61FFCAFF8C0E7DF8B2FB074D63" blockId="45.[825,1222,1854,1917]" box="[825,1196,1892,1917]" pageId="45" pageNumber="471">
<taxonomicName id="4C5AE1E2FFCAFF8C0E7DF8B2FB034D63" authority="Weber, 1913" authorityName="Weber" authorityYear="1913" box="[825,1192,1892,1917]" family="Ariidae" genus="Paracinetodus" kingdom="Animalia" order="Siluriformes" pageId="45" pageNumber="471" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B92E4673FFCAFF8C0E7DF8B2FB8B4D62" box="[825,1056,1892,1916]" italics="true" pageId="45" pageNumber="471">Paracinetodus carinatus</emphasis>
<bibRefCitation id="EFCBE790FFCAFF8C0961F8B2FB034D63" author="Weber M &amp; de Beaufort LF" box="[1061,1192,1892,1917]" pageId="45" pageNumber="471" refId="ref32046" refString="Weber M, de Beaufort LF. The Fishes of the Indo-Australian Archipelago. II. Malacopterygii, Myctophoidea, Ostariophysi: I Siluroidea. Leiden: EJ Brill, 1913." type="book" year="1913">Weber, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFCAFF8F0E7DF848FE4B4AB4" lastPageId="46" lastPageNumber="472" pageId="45" pageNumber="471" type="distribution">
<paragraph id="8BE59A61FFCAFF8F0E7DF848FE4B4AB4" blockId="45.[825,1474,1950,1974]" lastBlockId="46.[113,763,144,363]" lastPageId="46" lastPageNumber="472" pageId="45" pageNumber="471">
<emphasis id="B92E4673FFCAFF8C0E7DF848FBB64DA8" box="[825,1053,1950,1974]" italics="true" pageId="45" pageNumber="471">Habitat and distribution:</emphasis>
Freshwater,southern New
<collectingCountry id="F34DDAF1FFCAFF8C085AF848FACE4DA8" box="[1310,1381,1950,1974]" name="Guinea-Bissau" pageId="45" pageNumber="471">Guinea</collectingCountry>
(
<figureCitation id="136186E4FFCAFF8C0836F848FA194DA8" box="[1394,1458,1950,1974]" captionStart="Figure 38" captionStartId="38.[117,182,1915,1939]" captionTargetBox="[247,1333,150,1886]" captionTargetId="figure-7@38.[245,1333,144,1888]" captionTargetPageId="38" captionText="Figure 38. Representative skulls, in dorsal view, and distribution of the subtribe Doiichthyina.A, Brustiarius nox, AUM 47488. B, Paracinetodus carinatus, AUM 47550.C, Cochlefelis spatula, AUM 50296. D, Nedystoma dayi, AUM 47500. E, Aceroichthys dioctes, AUM 47507. F, Pachyula crassilabris, AUM 47509. G, Pararius proximus, AUM 47486.H, Potamosilurus velutinus, AUM 47489.* marine, ** brackish waters, *** freshwater." figureDoi="http://doi.org/10.5281/zenodo.11240197" httpUri="https://zenodo.org/record/11240197/files/figure.png" pageId="45" pageNumber="471">Fig.38</figureCitation>
).
<emphasis id="B92E4673FFC9FF8F0CC8FF46FE4B4AB4" box="[396,480,144,170]" italics="true" pageId="46" pageNumber="472">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFC9FF8F0D35FF61FDEB4B75" pageId="46" pageNumber="472" type="discussion">
<paragraph id="8BE59A61FFC9FF8F0D35FF61FDEB4B75" blockId="46.[113,763,144,363]" pageId="46" pageNumber="472">
<taxonomicName id="4C5AE1E2FFC9FF8F0D35FF61FEA34AD0" ID-CoL="GQTH" authorityName="Weber" authorityYear="1913" box="[113,264,183,206]" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="472" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B92E4673FFC9FF8F0D35FF61FEA34AD0" box="[113,264,183,206]" italics="true" pageId="46" pageNumber="472">Arius carinatus</emphasis>
</taxonomicName>
was included in
<taxonomicName id="4C5AE1E2FFC9FF8F0CFEFF60FDB04AD0" ID-CoL="3PZ5" authorityName="Ogilby" authorityYear="1898" box="[442,539,182,206]" family="Ariidae" genus="Cinetodus" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="472" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFC9FF8F0CFEFF60FDB04AD0" box="[442,539,182,206]" italics="true" pageId="46" pageNumber="472">Cinetodus</emphasis>
</taxonomicName>
in previous morphological studies (
<bibRefCitation id="EFCBE790FFC9FF8F0C5FFF00FE084AF0" author="Kailola PJ" box="[283,419,214,238]" pageId="46" pageNumber="472" pagination="87 - 166" refId="ref30669" refString="Kailola PJ. A phylogenetic exploration of the catfish family Ariidae (Otophysi: Siluriformes). The Beagle, Records of Museum Art Galleries of North Territory 2004; 20: 87 - 166." type="journal article" year="2004">Kailola 2004</bibRefCitation>
,
<bibRefCitation id="EFCBE790FFC9FF8F0CF6FF00FD5F4AF0" author="Marceniuk AP &amp; Menezes NA" box="[434,756,214,238]" pageId="46" pageNumber="472" pagination="1 - 126" refId="ref30826" refString="Marceniuk AP, Menezes NA. Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa 2007; 1416: 1 - 126." type="journal article" year="2007">Marceniuk and Menezes 2007</bibRefCitation>
,
<bibRefCitation id="EFCBE790FFC9FF8F0D35FF23FEE74B13" author="Marceniuk AP &amp; Menezes NA &amp; Britto MR" box="[113,332,245,269]" pageId="46" pageNumber="472" pagination="534 - 669" refId="ref30859" refString="Marceniuk AP, Menezes NA, Britto MR. Phylogenetic analysis of the family Ariidae (Ostariophysi: Siluriformes), with a hypothesis on the monophyly and relationships of the genera. Zoological Journal of the Linnean Society 2012; 165: 534 - 669. https: // doi. org / 10.1111 / j. 1096 - 3642.2012.00822. x" type="journal article" year="2012">
Marceniuk
<emphasis id="B92E4673FFC9FF8F0DA1FF20FEB84B13" box="[229,275,245,269]" italics="true" pageId="46" pageNumber="472">et al.</emphasis>
2012
</bibRefCitation>
). The results of the total-evidence analysis place the species in a lineage distinct from that of
<taxonomicName id="4C5AE1E2FFC9FF8F0F22FEC2FD6C4B32" ID-CoL="3PZ5" authorityName="Ogilby" authorityYear="1898" box="[614,711,276,300]" family="Ariidae" genus="Cinetodus" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="472" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFC9FF8F0F22FEC2FD6C4B32" box="[614,711,276,300]" italics="true" pageId="46" pageNumber="472">Cinetodus</emphasis>
</taxonomicName>
, corroborating a previous molecular result (Betancur-R. 2009), and requiring the establishment of the new genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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