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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1039.64740" ID-GBIF-Dataset="8fe0041e-40a6-4fd0-a674-948f3f919703" ID-PMC="PMC8159918" ID-Pensoft-Pub="1313-2970-1039-1" ID-Pensoft-UUID="FA2622A854585C59A7E8182A4E66C6DF" ID-PubMed="34084064" ID-ZooBank="3487C3570FAC4907A5C0C0A0C801C6E3" ModsDocID="1313-2970-1039-1" checkinTime="1621541808324" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2021" docId="C81A269EDA7D58E88E6432C578A9AEE6" docLanguage="en" docName="ZooKeys 1039: 1-108" docOrigin="ZooKeys 1039" docPubDate="2021-05-20" docSource="http://dx.doi.org/10.3897/zookeys.1039.64740" docTitle="Pyrrhalta semifulva" docType="treatment" docVersion="4" id="FA2622A854585C59A7E8182A4E66C6DF" lastPageNumber="1" masterDocId="FA2622A854585C59A7E8182A4E66C6DF" masterDocTitle="Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei" masterLastPageNumber="108" masterPageNumber="1" pageNumber="1" updateTime="1668152740594" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:affiliation>Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4358-7211</mods:nameIdentifier>
<mods:affiliation>Mendel University in Brno, Department of Zoology, Fisheries, Hydrobiology and Apiculture, Zemĕdĕlska 1, 613 00, Brno, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1039</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>108</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1039.64740</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1039.64740</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1039-1</mods:identifier>
<mods:identifier type="ZooBank">3487C3570FAC4907A5C0C0A0C801C6E3</mods:identifier>
<mods:identifier type="Pensoft-UUID">FA2622A854585C59A7E8182A4E66C6DF</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182248271" LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6" httpUri="http://treatment.plazi.org/id/C81A269EDA7D58E88E6432C578A9AEE6" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="C81A269E-DA7D-58E8-8E64-32C578A9AEE6" authority="(Jacoby, 1885)" baseAuthorityName="Jacoby" baseAuthorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">Pyrrhalta semifulva (Jacoby, 1885)</taxonomicName>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Field photographs of adults of Pyrrhalta species on host plants A P. tsoui Bezdek &amp; Lee B P. formosanensis sp. nov. C P. semifulva (Jacoby) D P. ishiharai Kimoto." figureDoi="10.3897/zookeys.1039.64740.figure26" httpUri="https://binary.pensoft.net/fig/545556" pageId="0" pageNumber="1">Figs 26C</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitus of Pyrrhalta formosanensis sp. nov. and P. semifulva (Jacoby) A P. formosanensis sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D P. semifulva, from Taiwan, female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure27" httpUri="https://binary.pensoft.net/fig/545557" pageId="0" pageNumber="1">, 27D-F</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">, 29</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Habitus of Pyrrhalta semifulva (Jacoby) from Japan A female, color variation, dorsal view B ditto, ventral view C Female, color variation, dorsal view D ditto, ventral view E female, color variation, dorsal view F male, color variation, dorsal view." figureDoi="10.3897/zookeys.1039.64740.figure30" httpUri="https://binary.pensoft.net/fig/545560" pageId="0" pageNumber="1">, 30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Chrysomelidae" genus="Gallerucella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Gallerucella semifulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="semifulva">Gallerucella semifulva</taxonomicName>
Jacoby, 1885: 745 (Japan: Kiga).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Lochmaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lochmaea (Tricholochmaea) semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva" subGenus="Tricholochmaea">Lochmaea (Tricholochmaea) semifulva</taxonomicName>
:
<bibRefCitation author="Laboissiere, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="1" refId="B46" refString="Laboissiere, V, 1932. Galerucini de la collection du Museum National d'Histoire naturelle recueillis dans l'Himalaya par le Dr. J. Harmand. Bulletin du Museum National d'Histoire Naturelle (2)4: 960-970." title="Galerucini de la collection du Museum National d'Histoire naturelle recueillis dans l'Himalaya par le Dr. J. Harmand. Bulletin du Museum National d'Histoire Naturelle (2) 4: 960 - 970." year="1932">
<normalizedToken originalValue="Laboissière">Laboissiere</normalizedToken>
1932
</bibRefCitation>
: 964;
<bibRefCitation author="Ogloblin, DA" journalOrPublisher="Moskva-Leningrad, Izdatel'stvo Akademii Nauk SSSR" pageId="0" pageNumber="1" refId="B72" refString="Ogloblin, DA, 1936. Listoedy, Galerucinae, Fauna SSR. Nasekomye Zhestkokrylye, n. s. 8, 26 (1). Moskva-Leningrad, Izdatel'stvo Akademii Nauk SSSR" title="Listoedy, Galerucinae, Fauna SSR. Nasekomye Zhestkokrylye, n. s. 8, 26 (1)." year="1936">Ogloblin 1936</bibRefCitation>
: 91 (redescription);
<bibRefCitation author="Chujo, M" journalOrPublisher="Kontyu" pageId="0" pageNumber="1" pagination="106 - 125" refId="B10" refString="Chujo, M, 1940. Beitrag zur Chrysomeliden-Fauna der Insel Shikoku (Japan) II. Kontyu 14: 106 - 125" title="Beitrag zur Chrysomeliden-Fauna der Insel Shikoku (Japan) II." volume="14" year="1940">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
1940
</bibRefCitation>
: 112 (Japan: Kyushu, Shikoku);
<bibRefCitation author="Chujo, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="1" pagination="51 - 62" refId="B11" refString="Chujo, M, 1954. Chrysomelid-beetles from Shikoku, Japan (III). Transactions of the Shikoku Entomological Society 4: 51 - 62" title="Chrysomelid-beetles from Shikoku, Japan (III)." volume="4" year="1954">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
1954
</bibRefCitation>
: 57 (Japan: Shikoku);
<bibRefCitation author="Wilcox, JA" journalOrPublisher="Junk, ' s-Gravenhage" pageId="0" pageNumber="1" refId="B92" refString="Wilcox, JA, 1971. Chrysomelidae: Galerucinae (Oidini, Galerucini, Metacyclini, Sermylini). In: Wilcox JA (Ed.) Coleopterorum Catalogus Supplementa. Pars 78 (1), (2nd edn.). W. Junk, 's-Gravenhage" title="Chrysomelidae: Galerucinae (Oidini, Galerucini, Metacyclini, Sermylini). In: Wilcox JA (Ed.) Coleopterorum Catalogus Supplementa. Pars 78 (1), (2 nd edn.). W." year="1971">Wilcox 1971</bibRefCitation>
: 82 (catalogue).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Tricholochmaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricholochmaea semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">Tricholochmaea semifulva</taxonomicName>
:
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
&amp; Kimoto 1961: 169 (catalogue);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kontyu" pageId="0" pageNumber="1" pagination="371 - 377" refId="B24" refString="Kimoto, S, 1964b. A revisional note on the type specimens of Japanese Chrysomelidae preserved in the museums of Europe and the United States. II. Kontyu 32: 371 - 377" title="A revisional note on the type specimens of Japanese Chrysomelidae preserved in the museums of Europe and the United States. II." volume="32" year="1964 b">Kimoto 1964b</bibRefCitation>
: 373 (deposition of type specimens);
<bibRefCitation author="Lopatin, IK" journalOrPublisher="Mantis, Olsztyn" pageId="0" pageNumber="1" refId="B55" refString="Lopatin, IK, Aleksandrovich, OR, Konstantinov, AS, 2004. Check list of leaf-beetle Chrysomelidae (Coleoptera) of the eastern Europe and northern Asia. Mantis, Olsztyn" title="Check list of leaf-beetle Chrysomelidae (Coleoptera) of the eastern Europe and northern Asia." year="2004">Lopatin et al. 2004</bibRefCitation>
: 130 (catalogue);
<bibRefCitation author="Beenen, R" editor="Loebl, I" journalOrPublisher="6). Chrysomeloidea. Apollo Books, Stenstrup" pageId="0" pageNumber="1" pagination="443 - 491" publicationUrl="https://www.cabdirect.org/cabdirect/abstract/20123189777" refId="B3" refString="Beenen, R, 2010. Galerucinae. In: Loebl, I, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera (Vol. 6). Chrysomeloidea. Apollo Books, Stenstrup: 443 - 491, https://www.cabdirect.org/cabdirect/abstract/20123189777" title="Galerucinae." url="https://www.cabdirect.org/cabdirect/abstract/20123189777" volumeTitle="Catalogue of Palaearctic Coleoptera (Vol." year="2010">Beenen 2010</bibRefCitation>
: 455 (catalogue).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Jacoby" baseAuthorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">Pyrrhalta semifulva</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.5109/22724" author="Kimoto, S" journalOrPublisher="Journal of the Faculty of Agriculture, Kyushu University" pageId="0" pageNumber="1" pagination="287 - 308" refId="B23" refString="Kimoto, S, 1964a. The Chrysomelidae of Japan and the Ryukyu islands, VI. Subfamily Galerucinae I. Journal of the Faculty of Agriculture, Kyushu University 13: 287 - 308, DOI: https://doi.org/10.5109/22724" title="The Chrysomelidae of Japan and the Ryukyu islands, VI. Subfamily Galerucinae I." url="https://doi.org/10.5109/22724" volume="13" year="1964 a">Kimoto 1964a</bibRefCitation>
: 299;
<bibRefCitation author="Kimoto, S" journalOrPublisher="Bulletin of the Osaka Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 26" publicationUrl="http://www.mus-nh.city.osaka.jp/publication/bulletin/bulletin/25/25-001.pdf" refId="B40" refString="Kimoto, S, Hiura, I, 1971. A list of the chrysomelid specimens preserved in the Osaka Museum of Natural History, III (Insecta: Coleoptera). Bulletin of the Osaka Museum of Natural History 25: 1 - 26, http://www.mus-nh.city.osaka.jp/publication/bulletin/bulletin/25/25-001.pdf" title="A list of the chrysomelid specimens preserved in the Osaka Museum of Natural History, III (Insecta: Coleoptera)." url="http://www.mus-nh.city.osaka.jp/publication/bulletin/bulletin/25/25-001.pdf" volume="25" year="1971">Kimoto and Hiura 1971</bibRefCitation>
: 15 (Japan);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="1" pagination="467 - 577" refId="B38" refString="Kimoto, S, Gressitt, JL, 1966. The Chrysomelidae of the Ryukyu Archipelago. Pacific Insects 8: 467 - 577" title="The Chrysomelidae of the Ryukyu Archipelago." volume="8" year="1966">Kimoto and Gressitt 1966</bibRefCitation>
: 476 (key), 520 (Ryukyus);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Entomological Review of Japan" pageId="0" pageNumber="1" pagination="21 - 26" publicationUrl="http://coleoptera.sakura.ne.jp/ERJ/ERJ26-1974.pdf" refId="B26" refString="Kimoto, S, 1974. Notes on the Chrysomelidae from Taiwan, VII. Entomological Review of Japan 26: 21 - 26, http://coleoptera.sakura.ne.jp/ERJ/ERJ26-1974.pdf" title="Notes on the Chrysomelidae from Taiwan, VII." url="http://coleoptera.sakura.ne.jp/ERJ/ERJ26-1974.pdf" volume="26" year="1974">Kimoto 1974</bibRefCitation>
: 24 (Taiwan);
<bibRefCitation author="Nakane, T" journalOrPublisher="Melanges de Philosophie et de Mathematique de la Societe Royal de Turin 3 [1762 - 1765]" pageId="0" pageNumber="1" refId="B67" refString="Nakane, T, 1984. On the Coleoptera occurring in Yaku-shima Island. In: Conservation Reports of the Yaku-shima Wilderness Area, Kyushu, Japan. Nature Conservation Bureau, Environment Agency of Japan, Tokyo, 587-631." title="On the Coleoptera occurring in Yaku-shima Island. In: Conservation Reports of the Yaku-shima Wilderness Area, Kyushu, Japan. Nature Conservation Bureau, Environment Agency of Japan, Tokyo, 587 - 631." year="1984">Nakane 1984</bibRefCitation>
: 626 (Japan);
<bibRefCitation author="Kimoto, S" journalOrPublisher="The Coleopterists' Association of Japan, Tokyo" pageId="0" pageNumber="1" refId="B30" refString="Kimoto, S, 1985. Family Chrysomelidae (Galerucinae). Check-list of Coleoptera of Japan, No. 28. The Coleopterists' Association of Japan, Tokyo" title="Family Chrysomelidae (Galerucinae). Check-list of Coleoptera of Japan, No. 28." year="1985">Kimoto 1985</bibRefCitation>
: 4 (catalogue);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="0" pageNumber="1" pagination="53 - 62" refId="B31" refString="Kimoto, S, 1986. The Chrysomelidae (Insecta: Coleoptera) collected by the Nagoya University Scientific Expedition to Taiwan in 1984. Kurume University Journal 35: 53 - 62" title="The Chrysomelidae (Insecta: Coleoptera) collected by the Nagoya University Scientific Expedition to Taiwan in 1984." volume="35" year="1986">Kimoto 1986</bibRefCitation>
: 56 (additional records in Taiwan);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="0" pageNumber="1" pagination="183 - 194" refId="B32" refString="Kimoto, S, 1987. The Chrysomelidae (Insecta: Coleoptera) collected by the Nagoya University Scientific Expedition to Taiwan in 1986. Kurume University Journal 36: 183 - 194" title="The Chrysomelidae (Insecta: Coleoptera) collected by the Nagoya University Scientific Expedition to Taiwan in 1986." volume="36" year="1987">Kimoto 1987</bibRefCitation>
: 188 (additional records in Taiwan);
<bibRefCitation author="Medvedev, LN" journalOrPublisher="AN SSSR, Moscow" pageId="0" pageNumber="1" refId="B62" refString="Medvedev, LN, Roginskaya, EY, 1988. Catalogue of the Host Plants of USSR Leaf-Beetles. AN SSSR, Moscow" title="Catalogue of the Host Plants of USSR Leaf-Beetles." year="1988">Medvedev and Roginskaya 1988</bibRefCitation>
: 116 (host plants);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="0" pageNumber="1" pagination="237 - 272" refId="B33" refString="Kimoto, S, 1989a. The Taiwanese Chrysomelidae (Insecta: Coleoptera) collected by Dr. Kintaro Baba, on the occasion of his entomological survey in 1983 and 1986. Kurume University Journal 38: 237 - 272" title="The Taiwanese Chrysomelidae (Insecta: Coleoptera) collected by Dr. Kintaro Baba, on the occasion of his entomological survey in 1983 and 1986." volume="38" year="1989 a">Kimoto 1989a</bibRefCitation>
: 268 (additional records in Taiwan);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Kurume University Journal" pageId="0" pageNumber="1" pagination="1 - 27" refId="B35" refString="Kimoto, S, 1991. The Taiwanese Chrysomelidae (Insecta: Coleoptera) collected by Dr. Kintaro Baba, on the occasion of his entomological survey in 1987, 1988 and 1989. Kurume University Journal 40: 1 - 27" title="The Taiwanese Chrysomelidae (Insecta: Coleoptera) collected by Dr. Kintaro Baba, on the occasion of his entomological survey in 1987, 1988 and 1989." volume="40" year="1991">Kimoto 1991</bibRefCitation>
: 9 (additional records in Taiwan);
<bibRefCitation author="Li, J-K" journalOrPublisher="Second Edition. Volumn Four, Angiosperms Dicotyledons [Diapensiaceae - Compositae]. Editorial Committee of the Flora of Taiwan, Department of Botany, National Taiwan University, Second Edition, Taipei" pageId="0" pageNumber="1" publicationUrl="https://www.cabdirect.org/cabdirect/abstract/19941106780" refId="B54" refString="Li, J-K, 1992. The Coleoptera fauna of northeast China. Education Publishing House, Jilin, 2 + 6 + 205 pp. https://www.cabdirect.org/cabdirect/abstract/19941106780" title="The Coleoptera fauna of northeast China. Education Publishing House, Jilin, 2 + 6 + 205 pp." url="https://www.cabdirect.org/cabdirect/abstract/19941106780" year="1992">Li 1992</bibRefCitation>
: 184 (China: Liaoning);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="0" pageNumber="1" refId="B41" refString="Kimoto, S, Takizawa, H, 1994. Leaf beetles (Chrysomelidae) of Japan. Tokai University Press, Tokyo" title="Leaf beetles (Chrysomelidae) of Japan." year="1994">Kimoto and Takizawa 1994</bibRefCitation>
: 234 (key), 307 (Japan);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Bulletin of the Institute of Comparative Studies of International Cultures and Societies" pageId="0" pageNumber="1" pagination="1 - 152" refId="B39" refString="Kimoto, S, Chu, Y-I, 1996. Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera). Bulletin of the Institute of Comparative Studies of International Cultures and Societies 16: 1 - 152" title="Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera)." volume="16" year="1996">Kimoto and Chu 1996</bibRefCitation>
: 57 (catalogue);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="0" pageNumber="1" refId="B42" refString="Kimoto, S, Takizawa, H, 1997. Leaf beetles (Chrysomelidae) of Taiwan. Tokai University Press, Tokyo" title="Leaf beetles (Chrysomelidae) of Taiwan." year="1997">Kimoto and Takizawa 1997</bibRefCitation>
: 300 (key), 374;
<bibRefCitation author="Wang, J-S" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="0" pageNumber="1" refId="B84" refString="Wang, J-S, Yang, X-K, 1998. Fauna of Chrysomelidae of Wuyishan Nature Reserve in China. China Forestry Publishing House, Beijing" title="Fauna of Chrysomelidae of Wuyishan Nature Reserve in China." year="1998">Wang and Yang 1998</bibRefCitation>
: 68 (China: Fujian);
<bibRefCitation author="Yang, X-K" editor="Huang, P-K" journalOrPublisher="6). Fuzhou: Fujian Science &amp; Technology Press" pageId="0" pageNumber="1" pagination="621 - 663" refId="B96" refString="Yang, X-K, 2002. Chrysomelidae: Galerucinae. In: Huang, P-K, Ed., Fauna of insects in Fujian province of China (Vol. 6). Fuzhou: Fujian Science &amp; Technology Press: 621 - 663" title="Chrysomelidae: Galerucinae." volumeTitle="Fauna of insects in Fujian province of China (Vol." year="2002">Yang 2002</bibRefCitation>
: 628 (China: Fujian);
<bibRefCitation author="Xue, HJ" journalOrPublisher="Junk, ' s-Gravenhage" pageId="0" pageNumber="1" publicationUrl="https://www.cabdirect.org/cabdirect/abstract/20103331643" refId="B93" refString="Xue, HJ, Yang, X-K, 2010. Species catalogue of Pyrrhalta Joannis (Coleoptera: Chrysomelidae: Galerucinae) of the world. Entomotaxonomia 32 (supplement): 119-136. https://www.cabdirect.org/cabdirect/abstract/20103331643" title="Species catalogue of Pyrrhalta Joannis (Coleoptera: Chrysomelidae: Galerucinae) of the world. Entomotaxonomia 32 (supplement): 119 - 136." url="https://www.cabdirect.org/cabdirect/abstract/20103331643" year="2010">Xue and Yang 2010</bibRefCitation>
: 129 (catalogue);
<bibRefCitation author="Yang, X" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B98" refString="Yang, X, Ge, S, Nie, R, Ruan, Y, Li, W, 2015. Chinese leaf beetles. Science Press, Beijing" title="Chinese leaf beetles." year="2015">Yang et al. 2015</bibRefCitation>
: 120 (catalogue);
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.720.13015" author="Matsumura, Y" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="77 - 89" refId="B56" refString="Matsumura, Y, Suenaga, H, Kamimura, Y, Gorb, SN, 2017. Traumatic mating by hand saw-like spines on the internal sac in Pyrrhalta maculicollis (Coleoptera, Chrysomelidae, Galerucinae). ZooKeys 720: 77 - 89, DOI: https://doi.org/10.3897/zookeys.720.13015" title="Traumatic mating by hand saw-like spines on the internal sac in Pyrrhalta maculicollis (Coleoptera, Chrysomelidae, Galerucinae)." url="https://doi.org/10.3897/zookeys.720.13015" volume="720" year="2017">Matsumura et al. 2017</bibRefCitation>
: 85 (female reproductive system).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Jacoby" baseAuthorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta (Tricholochmaea) semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva" subGenus="Tricholochmaea">Pyrrhalta (Tricholochmaea) semifulva</taxonomicName>
:
<bibRefCitation author="Wilcox, JA" journalOrPublisher="Bulletin of the New York State Museum of Sciences Service" pageId="0" pageNumber="1" pagination="1 - 160" refId="B91" refString="Wilcox, JA, 1965. A synopsis of the north American Galerucinae (Coleoptera: Chrysomelidae). Bulletin of the New York State Museum of Sciences Service 400: 1 - 160" title="A synopsis of the north American Galerucinae (Coleoptera: Chrysomelidae)." volume="400" year="1965">Wilcox 1965</bibRefCitation>
: 38;
<bibRefCitation author="Medvedev, LN" editor="Ler, PA" journalOrPublisher="Tom III Zhestkokrylye, ili zhuki. Chast' 2. Nauka, Sankt Petersburg" pageId="0" pageNumber="1" pagination="533 - 602" refId="B58" refString="Medvedev, LN, 1992. 105. Sem. Chrysomeloidae - Listoedy. In: Ler, PA, Ed., Opredelitel Nasekomykh Dalnego Vostoka SSSR v Shestikh Tomakh. Tom III Zhestkokrylye, ili zhuki. Chast' 2. Nauka, Sankt Petersburg: 533 - 602" title="105. Sem. Chrysomeloidae - Listoedy." volumeTitle="Opredelitel Nasekomykh Dalnego Vostoka SSSR v Shestikh Tomakh." year="1992">Medvedev 1992</bibRefCitation>
: 580 (key);
<bibRefCitation author="Medvedev, LN" journalOrPublisher="Izdatelstvo Irkutskogo Universiteta, Irkutsk" pageId="0" pageNumber="1" refId="B61" refString="Medvedev, LN, Dubeshko, LN, 1992. Opredelitel Listoedov Sibiri. Izdatelstvo Irkutskogo Universiteta, Irkutsk" title="Opredelitel Listoedov Sibiri." year="1992">Medvedev and Dubeshko 1992</bibRefCitation>
: 133 (key);
<bibRefCitation author="Medvedev, LN" journalOrPublisher="Euroasian Entomological Journal" pageId="0" pageNumber="1" pagination="137 - 143" publicationUrl="http://www.eco.nsc.ru/EEJ_contents/2006_5_2.pdf" refId="B59" refString="Medvedev, LN, 2006. Contribution to the fauna of leaf-beetles (Coleoptera, Chrysomelidae) of Amur Oblast. Euroasian Entomological Journal 5: 137 - 143, http://www.eco.nsc.ru/EEJ_contents/2006_5_2.pdf" title="Contribution to the fauna of leaf-beetles (Coleoptera, Chrysomelidae) of Amur Oblast." url="http://www.eco.nsc.ru/EEJ_contents/2006_5_2.pdf" volume="5" year="2006">Medvedev 2006</bibRefCitation>
: 141 (Russia: Far East).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Chrysomelidae" genus="Gallerucella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Gallerucella modesta" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="modesta">Gallerucella modesta</taxonomicName>
Jacoby, 1885: 745 (Japan: Nikko). Synonymized by
<bibRefCitation author="Chujo, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="1" pagination="51 - 62" refId="B11" refString="Chujo, M, 1954. Chrysomelid-beetles from Shikoku, Japan (III). Transactions of the Shikoku Entomological Society 4: 51 - 62" title="Chrysomelid-beetles from Shikoku, Japan (III)." volume="4" year="1954">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
1954
</bibRefCitation>
: 57.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Lochmaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lochmaea (Tricholochmaea) modesta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="modesta" subGenus="Tricholochmaea">Lochmaea (Tricholochmaea) modesta</taxonomicName>
:
<bibRefCitation author="Chujo, M" journalOrPublisher="Kontyu" pageId="0" pageNumber="1" pagination="106 - 125" refId="B10" refString="Chujo, M, 1940. Beitrag zur Chrysomeliden-Fauna der Insel Shikoku (Japan) II. Kontyu 14: 106 - 125" title="Beitrag zur Chrysomeliden-Fauna der Insel Shikoku (Japan) II." volume="14" year="1940">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
1940
</bibRefCitation>
: 112 (Japan: Shikoku).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Chrysomelidae" genus="Gallerucella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Gallerucella signaticeps" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="signaticeps">Gallerucella signaticeps</taxonomicName>
Weise, 1887: 191 (Vladivostok). Synonymized by
<bibRefCitation author="Ogloblin, DA" journalOrPublisher="Moskva-Leningrad, Izdatel'stvo Akademii Nauk SSSR" pageId="0" pageNumber="1" refId="B72" refString="Ogloblin, DA, 1936. Listoedy, Galerucinae, Fauna SSR. Nasekomye Zhestkokrylye, n. s. 8, 26 (1). Moskva-Leningrad, Izdatel'stvo Akademii Nauk SSSR" title="Listoedy, Galerucinae, Fauna SSR. Nasekomye Zhestkokrylye, n. s. 8, 26 (1)." year="1936">Ogloblin 1936</bibRefCitation>
: 91.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Lochmaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lochmaea japonica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="japonica">Lochmaea japonica</taxonomicName>
Weise, 1922: 67 (Japan: Honshu). Synonymized by
<bibRefCitation author="Laboissiere, V" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="1" refId="B46" refString="Laboissiere, V, 1932. Galerucini de la collection du Museum National d'Histoire naturelle recueillis dans l'Himalaya par le Dr. J. Harmand. Bulletin du Museum National d'Histoire Naturelle (2)4: 960-970." title="Galerucini de la collection du Museum National d'Histoire naturelle recueillis dans l'Himalaya par le Dr. J. Harmand. Bulletin du Museum National d'Histoire Naturelle (2) 4: 960 - 970." year="1932">
<normalizedToken originalValue="Laboissière">Laboissiere</normalizedToken>
1932
</bibRefCitation>
: 964.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Chrysomelidae" genus="Gallerucella" kingdom="Animalia" lsidName="Gallerucella semifulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Gallerucella semifulva</emphasis>
</taxonomicName>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
♀ (NHMUK, here designated): &quot;Kiga [h, w, underside of card glued with specimen] // Type / H.T. [p, w, circle label with red border] // Japan. / G. Lewis. / 1910-320. [p, w] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Galeruca</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Galeruca semifulva</emphasis>
</taxonomicName>
Jac [h, b] // SYN- / TYPE [p, w, circle label with blue border]&quot;. Paralectotypes. 1♂ (NHMUK): &quot;Kiga [h, w, underside of card glued with specimen] // Japan / Lewis [h, w] // Jacoby Coll. / 1909-28a. [p, w] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">Galeruca semifulva</taxonomicName>
Jac [h, b]&quot;; 1♂ (NHMUK): &quot;Kiga [h, w, underside of card glued with specimen] // Japan. / G. Lewis. / 1910-320. [p, w]&quot;; 1♀ (NHMUK): &quot;Japan. / G. Lewis. / 1910-320. [p, w]&quot;; 1 (TARI, sex undetermined): &quot;KIGA [h] / JAPAN [p] / 18.III.1880 [h] / Col. G. LEWIS [p, w] // CO / Type [p, circle label with yellow letters border] //
<taxonomicName family="Chrysomelidae" genus="Gallerucella" kingdom="Animalia" lsidName="Gallerucella" order="Coleoptera" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Gallerucella</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Galeruca semifulva</emphasis>
</taxonomicName>
/ Jacoby [h] / DET. M. CHUJO [p, w] //
<taxonomicName family="Chrysomelidae" genus="Gallerucella" kingdom="Animalia" lsidName="Gallerucella" order="Coleoptera" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Gallerucella</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Galeruca semifulva</emphasis>
</taxonomicName>
Jac. [h] / Det. T. Shiraki [p, w] // 1934 [p, w]&quot;; 1 (MCZC, sex undetermined): &quot;Japan / Lewis [h, w] // 1st Jacoby / Coll. [p, w] // Type. / Sp. figured. [p, w] // Type [p] / 17878 [h, r]&quot;. Since types much be collected from
<normalizedToken originalValue="“Kiga”">&quot;Kiga&quot;</normalizedToken>
based on
<bibRefCitation author="Jacoby, M" journalOrPublisher="Proceedings of the Scientific Meetings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="719 - 755" publicationUrl="https://zslpublications.onlinelibrary.wiley.com/doi/abs/10.1111/j.1096-3642.1885.tb02918.x" refId="B20" refString="Jacoby, M, 1885. Descriptions of the phytophagous Coleoptera of Japan obtained by Mr. George Lewis during his second journey, from February 1880 to September 1881. Part II. Halticinae and Galerucinae. Proceedings of the Scientific Meetings of the Zoological Society of London 1885: 719 - 755, https://zslpublications.onlinelibrary.wiley.com/doi/abs/10.1111/j.1096-3642.1885.tb02918.x" title="Descriptions of the phytophagous Coleoptera of Japan obtained by Mr. George Lewis during his second journey, from February 1880 to September 1881. Part II. Halticinae and Galerucinae." url="https://zslpublications.onlinelibrary.wiley.com/doi/abs/10.1111/j.1096-3642.1885.tb02918.x" volume="1885" year="1885">Jacoby (1885)</bibRefCitation>
. Those specimens with different localities but with type labels should not be regarded as type series, including 1 (NHMUK, sex undetermined): &quot;marshes / nagasaki (h, w) // Japan. / G. Lewis. / 1910-320. [p, w] // SYN- / TYPE [p, w, circle label with blue border]&quot;; 1♂ (NHMUK): &quot;Subashiri [h, w, underside of card glued with specimen] // Japan. / G. Lewis. / 1910-320. [p, w] // SYN- / TYPE [p, w, circle label with blue border]&quot;; 1 (TARI, sex undetermined): &quot;Ipongi [h] / JAPAN [p] / III.1881 [h] / Col. G. LEWIS [p] // CO / Type [p, circle label with yellow letters border] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galerucella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galerucella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Galerucella</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Galeruca semifulva</emphasis>
</taxonomicName>
/ Jacoby [h] / DET. M. CHUJO [p, w]&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Chrysomelidae" genus="Gallerucella" kingdom="Animalia" lsidName="Gallerucella modesta" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="modesta">
<emphasis italics="true" pageId="0" pageNumber="1">Gallerucella modesta</emphasis>
</taxonomicName>
. Lectotype (sex undetermined, NHMUK, here designated): &quot;Nikko [h, w, underside of card glued with specimen] // Type / H.T. [p, w, circle label with red border] // Japan. / G. Lewis. / 1910-320. [p, w] // Nikko. [p, w] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galerucella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galerucella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Galerucella</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galerucella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galerucella modesta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="modesta">
<emphasis italics="true" pageId="0" pageNumber="1">Galerucella modesta</emphasis>
</taxonomicName>
Jac. [h, b]&quot;. Paralectotypes. 1 (NHMUK, sex undetermined): &quot;Nikko. [p, w] // Japan. / G. Lewis. / 1910-320. [p, w] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galerucella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galerucella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Galerucella</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galerucella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galerucella modesta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="modesta">
<emphasis italics="true" pageId="0" pageNumber="1">Galerucella modesta</emphasis>
</taxonomicName>
/ Jac [h, w] //
<taxonomicName authorityName="Weise" authorityYear="1883" class="Insecta" family="Chrysomelidae" genus="Lochmaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lochmaea" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lochmaea</emphasis>
</taxonomicName>
/ (
<taxonomicName authorityName="Laboissiere" authorityYear="1932" class="Insecta" family="Chrysomelidae" genus="Tricholochmaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricholochmaea" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Tricholochmaea</emphasis>
</taxonomicName>
)
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galeruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galeruca semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Galeruca semifulva</emphasis>
</taxonomicName>
Jacob. [h] / D. Ogloblin det. [p, w]&quot;; 1 (MCZC, sex undetermined): &quot;Japan / Lewis [h, w] / 1st Jacoby / Coll. [p, w] //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Galerucella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Galerucella modesta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="modesta">Galerucella modesta</taxonomicName>
Jac. [h, b] // Type [p] / 17877 [h, r]&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2011-06-18" collectorName="H. Suenaga" country="Japan" location="Nopporo" municipality="Etetsu-shi" specimenCount="1" specimenCount-female="1" stateProvince="Hokkaido">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Japan">Japan</collectingCountry>
.
</emphasis>
<collectingRegion country="Japan" name="Hokkaido">Hokkaido</collectingRegion>
:
<specimenCount type="female">1♀</specimenCount>
(HSC),
<collectingMunicipality>Etetsu-shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:208843D7054B965ACC33E53D72370346" country="Japan" municipality="Etetsu-shi" name="Nopporo" stateProvince="Hokkaido">Nopporo</location>
,
<collectingDate value="2011-06-18">18.VI.2011</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2011-08-29" collectorName="H. Suenaga" country="Japan" location="Lake Utonai-ko" municipality="Tomakomai-shi" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
(HSC),
<collectingMunicipality>Tomakomai-shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:1A2BDDE2959C4BFAAD01A037486458C1" country="Japan" municipality="Tomakomai-shi" name="Lake Utonai-ko">Lake Utonai-ko</location>
,
<collectingDate value="2011-08-29">29.VIII.2011</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(HSC), same but with
<normalizedToken originalValue="“22.V.2012”">
&quot;
<date value="2012-05-22">22.V.2012</date>
&quot;
</normalizedToken>
; Honshu:
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2016-06-10" collectorName="S. Sejima" country="Japan" county="Nikaho-shi" location="Hokodate" municipality="Chokaisan" specimenCount="1" stateProvince="Akita Pref.">
<collectingRegion country="Japan" name="Akita">Akita Pref.</collectingRegion>
,
<collectingCounty>Nikaho-shi</collectingCounty>
,
<normalizedToken originalValue="Chôkaisan">
<collectingMunicipality>Chokaisan</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:C42704F072999A7CC2D1DC53583B62B8" country="Japan" county="Nikaho-shi" municipality="Chokaisan" name="Hokodate" stateProvince="Akita Pref.">Hokodate</location>
,
<collectingDate value="2016-06-10">10.VI.2016</collectingDate>
, leg.
<collectorName>S. Sejima</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(TARI),
<materialsCitation collectingDate="1934-06-29" collectorName="F. Watanabe" country="Japan" location="Aomori Pref." specimenCount="1" stateProvince="Aomori Pref.">
<collectingRegion country="Japan" name="Aomori">Aomori Pref.</collectingRegion>
,
<collectingDate value="1934-06-29">29.VI. 1934</collectingDate>
, leg.
<collectorName>F. Watanabe</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(TARI),
<materialsCitation collectingDate="1933-06-01" collectorName="A. Fukuda" country="Japan" location="Hatinohe" specimenCount="1" stateProvince="Aomori Pref.">
<collectingRegion country="Japan" name="Aomori">Aomori Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:8B0B78796981846C10D981FF42FB006C" country="Japan" name="Hatinohe" stateProvince="Aomori Pref.">Hatinohe</location>
,
<collectingDate value="1933-06-01">1.VI.1933</collectingDate>
, leg.
<collectorName>A. Fukuda</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(KMNH),
<materialsCitation collectingDate="1963-05-03" collectorName="H. Sasaji" country="Japan" location="Mt. Monju" specimenCount="1" stateProvince="Fukui Pref.">
<collectingRegion country="Japan" name="Fukui">Fukui Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:A22E9221F9CC5C8AE8E89C80CB29121F" country="Japan" name="Mt. Monju" stateProvince="Fukui Pref.">Mt. Monju</location>
,
<collectingDate value="1963-05-03">3.V.1963</collectingDate>
, leg.
<collectorName>H. Sasaji</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(NMNS),
<materialsCitation collectingDate="1946-04-02" collectingDateMax="1947-05-06" collectingDateMin="1946-04-02" collectorName="K. Ohbayashi" country="Japan" location="Gifu Pref." specimenCount="1" specimenCount-male="1" stateProvince="Gifu Pref.">
<collectingRegion country="Japan" name="Gifu">Gifu Pref.</collectingRegion>
,
<collectingDate value="1946-04-02">2.IV.1946</collectingDate>
;
<specimenCount type="male">1♂</specimenCount>
(NMNS), same locality,
<collectingDate value="1947-05-06">6.V.1947</collectingDate>
, leg.
<collectorName>K. Ohbayashi</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(NMNS),
<materialsCitation collectingDate="1956-04-15" collectorName="K. &amp; M. Ohbayashi" country="Japan" location="Suhara" specimenCount="1" stateProvince="Gifu Pref.">
<collectingRegion country="Japan" name="Gifu">Gifu Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:7719F2AAB5B887350A96B67DFF12285B" country="Japan" name="Suhara" stateProvince="Gifu Pref.">Suhara</location>
,
<collectingDate value="1956-04-15">15.IV.1956</collectingDate>
, leg.
<collectorName>K. &amp; M. Ohbayashi</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1955-05-05" collectorName="K. Ohbayashi" country="Japan" location="Suhara" specimenCount="1" specimenCount-female="1" stateProvince="Gifu">
<specimenCount type="female">1♀</specimenCount>
(NMNS), same locality,
<collectingDate value="1955-05-05">5.V.1955</collectingDate>
, leg.
<collectorName>K. Ohbayashi</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1956-05-13" collectingDateMax="1957-05-03" collectingDateMin="1956-05-13" collectorName="N. Ohbayashi" country="Japan" location="Suhara" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Gifu">
<specimenCount type="male">1♂</specimenCount>
(NMNS), same but with
<normalizedToken originalValue="“13.V.1956”">
&quot;
<collectingDate value="1956-05-13">13.V.1956</collectingDate>
&quot;
</normalizedToken>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(NMNS), same but with
<normalizedToken originalValue="“15.V.1956”">
&quot;
<collectingDate value="1956-05-15">15.V.1956</collectingDate>
&quot;
</normalizedToken>
;
<specimenCount type="male">1♂</specimenCount>
(NMNS), same locality,
<collectingDate value="1957-05-03">3.V.1957</collectingDate>
, leg.
<collectorName>N. Ohbayashi</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1957-05-19" country="Japan" specimenCount="3" specimenCount-male="3">
<specimenCount type="male">1♂</specimenCount>
(NMNS), same but with
<normalizedToken originalValue="“19.V.1957”">
&quot;
<collectingDate value="1957-05-19">19.V.1957</collectingDate>
&quot;
</normalizedToken>
;
<specimenCount type="male">1♂</specimenCount>
(NMNS), same but with &quot;
<specimenCount type="male">1♂</specimenCount>
(NMNS)
</materialsCitation>
,
<materialsCitation collectingDate="1941-06-11" collectorName="K. Ohbayashi" country="Japan" location="Tanigumi" specimenCount="1" stateProvince="Gifu Pref.">
<collectingRegion country="Japan" name="Gifu">Gifu Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:F768DDC0CBD7C1FA29AB7E440C7CADEE" country="Japan" name="Tanigumi" stateProvince="Gifu Pref.">Tanigumi</location>
,
<collectingDate value="1941-06-11">11.VI.1941</collectingDate>
, leg.
<collectorName>K. Ohbayashi</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2008-06-06" collectorName="S. Sejima" country="Japan" location="Fujiwara" municipality="Minakami-machi" specimenCount="1" stateProvince="Gunma Pref.">
<collectingRegion country="Japan" name="Gunma">Gunma Pref.</collectingRegion>
,
<collectingMunicipality>Minakami-machi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:86C19398DE7C8D774B9A2D2FB88D84EF" country="Japan" municipality="Minakami-machi" name="Fujiwara" stateProvince="Gunma Pref.">Fujiwara</location>
,
<collectingDate value="2008-06-06">6.VI.2008</collectingDate>
, leg.
<collectorName>S. Sejima</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2010-06-14" collectorName="H. Suenaga" country="Japan" location="Mt. Mushiki-toge" municipality="Akioota-cho" specimenCount="1" stateProvince="Hiroshima Pref.">
<collectingRegion country="Japan" name="Hiroshima">Hiroshima Pref.</collectingRegion>
,
<collectingMunicipality>Akioota-cho</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:0552D8D5478BDBAE3CC7DD3E7659704B" country="Japan" municipality="Akioota-cho" name="Mt. Mushiki-toge" stateProvince="Hiroshima Pref.">Mt. Mushiki-toge</location>
,
<collectingDate value="2010-06-14">14.VI.2010</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="female">2♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2009-06-13" collectorName="H. Suenaga" country="Japan" location="Takano" municipality="Takano-cho" specimenCount="1" stateProvince="Hiroshima Pref.">
<collectingRegion country="Japan" name="Hiroshima">Hiroshima Pref.</collectingRegion>
,
<collectingMunicipality>Takano-cho</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:CC49DAE8BACB7D64D6B8336037021291" country="Japan" municipality="Takano-cho" name="Takano" stateProvince="Hiroshima Pref.">Takano</location>
,
<collectingDate value="2009-06-13">13.VI.2009</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="male">8♂</specimenCount>
,
<specimenCount type="female">5♀</specimenCount>
(NMNS),
<materialsCitation collectingDate="1964-05-04" collectorName="M. - H. Chujo" country="Japan" location="Mt. Oginosen" specimenCount="1" stateProvince="Hyogo Pref.">
<collectingRegion country="Japan" name="Hyogo">Hyogo Pref.</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:3CA0EFA7DF9100796420966C89FA4A2C" country="Japan" name="Mt. Oginosen" stateProvince="Hyogo Pref.">Mt. Oginosen</location>
,
<collectingDate value="1964-05-04">4.V.1964</collectingDate>
, leg.
<collectorName>
M.-H.
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1965-05-02" collectorName="Y. Ohira" country="Japan" location="Mt. Oginosen" specimenCount="12" specimenCount-female="4" specimenCount-male="8" stateProvince="Hyogo">
<specimenCount type="male">8♂</specimenCount>
,
<specimenCount type="female">4♀</specimenCount>
(TARI), same locality,
<collectingDate value="1965-05-02">2.V.1965</collectingDate>
, leg.
<collectorName>Y. Ohira</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(HSC),
<materialsCitation collectingDate="2012-04-18" collectorName="H. Suenaga" country="Japan" location="Junmu-ji" municipality="Zushi-shi" specimenCount="1" specimenCount-female="1" stateProvince="Kanagawa Pref.">
<collectingRegion country="Japan" name="Kanagawa">Kanagawa Pref.</collectingRegion>
,
<collectingMunicipality>Zushi-shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:70C8C2545532CC59E3186AF43C5B7B4F" country="Japan" municipality="Zushi-shi" name="Junmu-ji" stateProvince="Kanagawa Pref.">Junmu-ji</location>
,
<collectingDate value="2012-04-18">18.IV.2012</collectingDate>
,
<collectorName>H. Suenaga</collectorName>
;
<specimenCount type="female">1♀</specimenCount>
(TARI)
</materialsCitation>
,
<materialsCitation collectingDate="1935-05-12" collectorName="S. Miyamoto" country="Japan" location="Koganezawa" specimenCount="1" stateProvince="Nagano Pref.">
<collectingRegion country="Japan" name="Nagano">Nagano Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:183EFE9BC9CEFA2E8644830202327BAB" country="Japan" name="Koganezawa" stateProvince="Nagano Pref.">Koganezawa</location>
,
<collectingDate value="1935-05-12">12.V.1935</collectingDate>
, leg.
<collectorName>S. Miyamoto</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(KMNH),
<materialsCitation collectingDate="1973-07-27" collectorName="S. Kimoto" country="Japan" location="Misuzuko" specimenCount="1" stateProvince="Nagano Pref.">
<collectingRegion country="Japan" name="Nagano">Nagano Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:7741E9E6884D573FF6ED12F0F30C7ABF" country="Japan" name="Misuzuko" stateProvince="Nagano Pref.">Misuzuko</location>
,
<collectingDate value="1973-07-27">27.VII.1973</collectingDate>
, leg.
<collectorName>S. Kimoto</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(KMNH),
<materialsCitation collectingDate="1951-07-10" collectorName="M. Takahashi" country="Japan" location="Wada" specimenCount="1" stateProvince="Nagano Pref.">
<collectingRegion country="Japan" name="Nagano">Nagano Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:ADCEEB5AA669D2FB16E226DF454C0F4D" country="Japan" name="Wada" stateProvince="Nagano Pref.">Wada</location>
,
<collectingDate value="1951-07-10">10.VII.1951</collectingDate>
, leg.
<collectorName>M. Takahashi</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(TARI),
<materialsCitation collectingDate="1931-07-25" collectorName="K. Obayashi" country="Japan" location="Yarisawa" specimenCount="1" stateProvince="Nagano Pref.">
<collectingRegion country="Japan" name="Nagano">Nagano Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:70CB03B285057934ACC785A6CE90D739" country="Japan" name="Yarisawa" stateProvince="Nagano Pref.">Yarisawa</location>
,
<collectingDate value="1931-07-25">25.VII.1931</collectingDate>
, leg.
<collectorName>K. Obayashi</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(TARI),
<materialsCitation collectingDate="1937-05-22" collectorName="K. Baba" country="Japan" location="Suizu" municipality="Sado" specimenCount="1" stateProvince="Niigata Pref.">
<collectingRegion country="Japan" name="Niigata">Niigata Pref.</collectingRegion>
,
<collectingMunicipality>Sado</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:F9729247294358B838C223F18287ABC2" country="Japan" municipality="Sado" name="Suizu" stateProvince="Niigata Pref.">Suizu</location>
,
<collectingDate value="1937-05-22">22.V.1937</collectingDate>
, leg.
<collectorName>K. Baba</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2006-05-04" collectorName="H. Suenaga" country="Japan" location="Kagamino-cho" municipality="Tomata-Gun" specimenCount="1" stateProvince="Okayama Pref.">
<collectingRegion country="Japan" name="Okayama">Okayama Pref.</collectingRegion>
,
<collectingMunicipality>Tomata-Gun</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:0EDA516207717F55D1198EABDEA9765D" country="Japan" municipality="Tomata-Gun" name="Kagamino-cho" stateProvince="Okayama Pref.">Kagamino-cho</location>
,
<collectingDate value="2006-05-04">4.V.2006</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC), same but with
<normalizedToken originalValue="“Shiraka-keikoku”">&quot;Shiraka-keikoku&quot;</normalizedToken>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2006-07-02" collectorName="S. Sejima" country="Japan" location="Toyanagakoma" municipality="Niimi-shi" specimenCount="1" stateProvince="Okayama Pref.">
<collectingRegion country="Japan" name="Okayama">Okayama Pref.</collectingRegion>
,
<collectingMunicipality>Niimi-shi</collectingMunicipality>
,
<normalizedToken originalValue="Toyanagakôma">
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:3994F33DEEC1FABE113D2495DBB9B2BE" country="Japan" municipality="Niimi-shi" name="Toyanagakoma" stateProvince="Okayama Pref.">Toyanagakoma</location>
</normalizedToken>
,
<collectingDate value="2006-07-02">2.VII.2006</collectingDate>
, leg.
<collectorName>S. Sejima</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(TARI),
<materialsCitation collectingDate="1940-06-16" collectorName="Tn. Nakane" country="Japan" location="Sanno-Toge" municipality="Nikko" specimenCount="1" stateProvince="Tochigi Pref.">
<collectingRegion country="Japan" name="Tochigi">Tochigi Pref.</collectingRegion>
,
<collectingMunicipality>Nikko</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:87DAAB1A80DFC6C5E6447B6857DB1689" country="Japan" municipality="Nikko" name="Sanno-Toge" stateProvince="Tochigi Pref.">Sanno-Toge</location>
,
<collectingDate value="1940-06-16">16.VI.1940</collectingDate>
, leg.
<collectorName>Tn. Nakane</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Kyushu" specimenCount="2" specimenCount-male="2">
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:E50891DEA0BACE0DCE886A353EB6B74B" country="Japan" name="Kyushu">Kyushu</location>
:
<specimenCount type="male">2♂</specimenCount>
(TARI)
</materialsCitation>
,
<materialsCitation collectingDate="1934-07-07" collectorName="K. Yamauchi" country="Japan" location="Mt. Hiko" specimenCount="1" stateProvince="Fukuoka Pref.">
<collectingRegion country="Japan" name="Fukuoka">Fukuoka Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:B47302024BF484B4A520E6C0A171E88C" country="Japan" name="Mt. Hiko" stateProvince="Fukuoka Pref.">Mt. Hiko</location>
,
<collectingDate value="1934-07-07">7.VII.1934</collectingDate>
, leg.
<collectorName>K. Yamauchi</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1941-07-14" collectorName="M. Chujo" country="Japan" location="Mt. Hiko" specimenCount="1" specimenCount-female="1" stateProvince="Fukuoka">
<specimenCount type="female">1♀</specimenCount>
(TARI), same locality,
<collectingDate value="1941-07-14">14.VII.1941</collectingDate>
, leg.
<collectorName>M. Chujo</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(TARI),
<materialsCitation collectingDate="1939-07-16" collectorName="Y. Takemura" country="Japan" location="Kirishima" specimenCount="1" stateProvince="Kagoshima Pref.">
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:F4AD80D22B3454E3560D59AE9653879F" country="Japan" name="Kirishima" stateProvince="Kagoshima Pref.">Kirishima</location>
,
<collectingDate value="1939-07-16">16.VII.1939</collectingDate>
, leg.
<collectorName>Y. Takemura</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2013-05-06" collectorName="H. Suenaga" country="Japan" location="Sata-misaki" municipality="Minamioosumi-machi" specimenCount="1" stateProvince="Kagoshima Pref.">
<collectingRegion country="Japan" name="Kagoshima">Kagoshima Pref.</collectingRegion>
,
<collectingMunicipality>Minamioosumi-machi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:D460D0149DBF7B820F94B1114227E297" country="Japan" municipality="Minamioosumi-machi" name="Sata-misaki" stateProvince="Kagoshima Pref.">Sata-misaki</location>
,
<collectingDate value="2013-05-06">6.V.2013</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2013-05-05" collectorName="H. Suenaga" country="Japan" location="Hakuso" municipality="Kamizue-cho" specimenCount="1" stateProvince="Oita Pref.">
<collectingRegion country="Japan" name="Oita">Oita Pref.</collectingRegion>
,
<collectingMunicipality>Kamizue-cho</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:52A9B8282871B3F861EA841C84C8D8D3" country="Japan" municipality="Kamizue-cho" name="Hakuso" stateProvince="Oita Pref.">Hakuso</location>
,
<collectingDate value="2013-05-05">5.V.2013</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2006-06-11" collectorName="S. Sejima" country="Japan" location="Kurodake" municipality="Yufu-shi" specimenCount="1" stateProvince="Oita Pref.">
<collectingRegion country="Japan" name="Oita">Oita Pref.</collectingRegion>
,
<collectingMunicipality>Yufu-shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:4EFC4F518BFFB68CEB12B6FDEC10EC1E" country="Japan" municipality="Yufu-shi" name="Kurodake" stateProvince="Oita Pref.">Kurodake</location>
,
<collectingDate value="2006-06-11">11.VI.2006</collectingDate>
, leg.
<collectorName>S. Sejima</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Shikoku" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:2207B5CB0BD45027B13F6443C065D0B1" country="Japan" name="Shikoku">Shikoku</location>
:
<specimenCount type="female">1♀</specimenCount>
(HSC)
</materialsCitation>
,
<materialsCitation collectingDate="2009-06-28" collectorName="H. Suenaga" country="Japan" county="Kumakogen-cho" location="Tsuchigoya" municipality="Mt. Ishizuchi" specimenCount="1" stateProvince="Ehime Pref.">
<collectingRegion country="Japan" name="Ehime">Ehime Pref.</collectingRegion>
,
<collectingCounty>Kumakogen-cho</collectingCounty>
,
<collectingMunicipality>Mt. Ishizuchi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:7D0B52AF7D1647372CA1EDA7C29B44CD" country="Japan" county="Kumakogen-cho" municipality="Mt. Ishizuchi" name="Tsuchigoya" stateProvince="Ehime Pref.">Tsuchigoya</location>
,
<collectingDate value="2009-06-28">28.VI.2009</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(HSC),
<materialsCitation collectingDate="2009-05-23" collectorName="H. Suenaga" country="Japan" location="Mt. Saraga-mine" municipality="Kumakogen-cho" specimenCount="1" stateProvince="Ehime Pref.">
<collectingRegion country="Japan" name="Ehime">Ehime Pref.</collectingRegion>
,
<collectingMunicipality>Kumakogen-cho</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:F424D610288318D73D00A5EC53A75A9F" country="Japan" municipality="Kumakogen-cho" name="Mt. Saraga-mine" stateProvince="Ehime Pref.">Mt. Saraga-mine</location>
,
<collectingDate value="2009-05-23">23.V.2009</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(HSC),
<materialsCitation collectingDate="2007-07-16" collectorName="H. Suenaga" country="Japan" location="Omogokei" municipality="Kumakogen-cho" specimenCount="1" stateProvince="Ehime Pref.">
<collectingRegion country="Japan" name="Ehime">Ehime Pref.</collectingRegion>
,
<collectingMunicipality>Kumakogen-cho</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:645462AC410390DC1C6587645A92BC75" country="Japan" municipality="Kumakogen-cho" name="Omogokei" stateProvince="Ehime Pref.">Omogokei</location>
,
<collectingDate value="2007-07-16">16.VII.2007</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2007-05-27" collectorName="T. Ichiyanagi" country="Japan" location="Komenono" municipality="Matsuyama-shi" specimenCount="1" stateProvince="Ehime Pref.">
<collectingRegion country="Japan" name="Ehime">Ehime Pref.</collectingRegion>
,
<collectingMunicipality>Matsuyama-shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:90DA4A99BA6980104DF984461BC7CCE4" country="Japan" municipality="Matsuyama-shi" name="Komenono" stateProvince="Ehime Pref.">Komenono</location>
,
<collectingDate value="2007-05-27">27.V.2007</collectingDate>
, leg.
<collectorName>T. Ichiyanagi</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2007-05-26" collectorName="H. Suenaga" country="Japan" location="Komenono" municipality="Matsuyama-shi" specimenCount="1" specimenCount-female="1" stateProvince="Ehime">
<specimenCount type="female">1♀</specimenCount>
(HSC), same locality,
<collectingDate value="2007-05-26">26.V.2007</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<specimenCount type="female">3♀♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2010-05-02" collectorName="K. Hashimoto" country="Japan" county="Matsuyama-shi" location="Ishitegawa" municipality="Shukuno" specimenCount="1" stateProvince="Ehime Pref.">
<collectingRegion country="Japan" name="Ehime">Ehime Pref.</collectingRegion>
,
<collectingCounty>Matsuyama-shi</collectingCounty>
,
<collectingMunicipality>Shukuno</collectingMunicipality>
, near the dame of
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:FE52AD90E6E2975A0BEC3DAA20A9AB29" country="Japan" county="Matsuyama-shi" municipality="Shukuno" name="Ishitegawa" stateProvince="Ehime Pref.">Ishitegawa</location>
,
<collectingDate value="2010-05-02">2.V.2010</collectingDate>
, leg.
<collectorName>K. Hashimoto</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2007-05-07" collectorName="S. Sejima" country="Japan" county="Uwajima-shi" location="Yatsurayama" municipality="Onigajozan" specimenCount="1" stateProvince="Ehime Pref.">
<collectingRegion country="Japan" name="Ehime">Ehime Pref.</collectingRegion>
,
<collectingCounty>Uwajima-shi</collectingCounty>
,
<normalizedToken originalValue="Onigajôzan">
<collectingMunicipality>Onigajozan</collectingMunicipality>
</normalizedToken>
to
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:BDA45FF06A60AB293BFE5A1DE5C9DF28" country="Japan" county="Uwajima-shi" municipality="Onigajozan" name="Yatsurayama" stateProvince="Ehime Pref.">Yatsurayama</location>
,
<collectingDate value="2007-05-07">7.V.2007</collectingDate>
, leg.
<collectorName>S. Sejima</collectorName>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="2007-07-29" collectorName="H. Suenaga" country="Japan" county="Mannou-cho" location="Mt. Daisenzan" municipality="Nakadouri" specimenCount="1" stateProvince="Kagawa Pref.">
<collectingRegion country="Japan" name="Kagawa">Kagawa Pref.</collectingRegion>
,
<collectingCounty>Mannou-cho</collectingCounty>
,
<collectingMunicipality>Nakadouri</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:073E00EDAC8A476908AC696C6EEFAB60" country="Japan" county="Mannou-cho" municipality="Nakadouri" name="Mt. Daisenzan" stateProvince="Kagawa Pref.">Mt. Daisenzan</location>
,
<collectingDate value="2007-07-29">29.VII.2007</collectingDate>
, leg.
<collectorName>H. Suenaga</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Kooti-Ken, I. Okubo" country="Japan" location="Kochi Pref." specimenCount="2" specimenCount-female="2" stateProvince="Kochi Pref.">
<specimenCount type="female">2♀♀</specimenCount>
(TARI),
<collectorName>Kooti-Ken</collectorName>
(=
<collectingRegion country="Japan" name="Kochi">Kochi Pref.</collectingRegion>
), leg.
<collectorName>I. Okubo</collectorName>
</materialsCitation>
;
<specimenCount type="female">1♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“26.V.1935”">
&quot;
<date value="1935-05-26">26.V.1935</date>
&quot;
</normalizedToken>
;
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
(TARI), same but with
<normalizedToken originalValue="“24.VII.1936”">
&quot;
<date value="1936-07-24">24.VII.1936</date>
&quot;
</normalizedToken>
;
<specimenCount type="female">1♀</specimenCount>
(HSC),
<materialsCitation collectingDate="1987-05-18" collectorName="S. Mano" country="Japan" location="Mt. Kotsu-zan" municipality="Yoshinokawa-shi" specimenCount="1" stateProvince="Tokushima Pref.">
<collectingRegion country="Japan" name="Tokushima">Tokushima Pref.</collectingRegion>
,
<collectingMunicipality>Yoshinokawa-shi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:A533263F76D872C78708196E146A9FD4" country="Japan" municipality="Yoshinokawa-shi" name="Mt. Kotsu-zan" stateProvince="Tokushima Pref.">Mt. Kotsu-zan</location>
,
<collectingDate value="1987-05-18">18.V.1987</collectingDate>
, leg.
<collectorName>S. Mano</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1965-07-06" collectorName="S. Kimoto" country="Japan" determinerName="S. Kimoto" location="Alishan" municipality="Taiwan. Chiayi" specimenCount="1" specimenCount-female="1">
<collectingMunicipality>Taiwan. Chiayi</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
(KMNH),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:F369AC3FB1159118E55F43F62517A4C5" country="Japan" municipality="Taiwan. Chiayi" name="Alishan">Alishan</location>
(阿里山),
<collectingDate value="1965-07-06">6.VII.1965</collectingDate>
, leg.
<collectorName>S. Kimoto</collectorName>
, det.
<determinerName>S. Kimoto</determinerName>
</materialsCitation>
,
<materialsCitation collectingDate="2010-07-10" collectorName="M. - H. Tsou" country="Japan" location="Lupi" municipality="Hsinchu" specimenCount="1" specimenCount-female="1">
1974;
<collectingMunicipality>Hsinchu</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:8FD13C1809A74D7EC1A62CFEF10DD8AC" country="Japan" municipality="Hsinchu" name="Lupi">Lupi</location>
(魯壁),
<collectingDate value="2010-07-10">10.VII.2010</collectingDate>
, leg.
<collectorName>M.-H. Tsou</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2018-08-04" collectorName="H. - F. Lu" country="Japan" location="Hahuan Cross-Ridge" municipality="Hualien" specimenCount="1" specimenCount-female="1">
<collectingMunicipality>Hualien</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:5B7BCFE5085CA151305344CDFD50F494" country="Japan" municipality="Hualien" name="Hahuan Cross-Ridge">Hahuan Cross-Ridge</location>
(合歡越嶺古道),
<collectingDate value="2018-08-04">4.VIII.2018</collectingDate>
, leg.
<collectorName>H.-F. Lu</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2009-04-28" collectorName="M. - H. Tsou" country="Japan" location="Ssuyuan" municipality="Ilan" specimenCount="1" specimenCount-female="1">
<collectingMunicipality>Ilan</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:B01373FA108B9FC1EDAA6F7FAA416A97" country="Japan" municipality="Ilan" name="Ssuyuan">Ssuyuan</location>
(思源),
<collectingDate value="2009-04-28">28.IV.2009</collectingDate>
, leg.
<collectorName>M.-H. Tsou</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1985-05" collectorName="W. L. Chen, Nagoya Univ. Col." country="Japan" determinerName="S. Kimoto" location="Liukuei" municipality="Kaohsiung" specimenCount="1" specimenCount-female="1">
<collectingMunicipality>Kaohsiung</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
(KMNH),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:312E14595B915FBBBA02811894F1101E" country="Japan" municipality="Kaohsiung" name="Liukuei">Liukuei</location>
(六龜),
<collectingDate value="1985-05">V.1985</collectingDate>
, leg.
<collectorName>W. L. Chen</collectorName>
,
<collectorName>Nagoya Univ. Col.</collectorName>
, det.
<determinerName>S. Kimoto</determinerName>
</materialsCitation>
,
<materialsCitation collectingDate="1990-05-11" collectorName="C. C. Chiang" country="Japan" location="Shanlinchi" municipality="Nantou" specimenCount="1" specimenCount-male="1">
1987;
<collectingMunicipality>Nantou</collectingMunicipality>
:
<specimenCount type="male">1♂</specimenCount>
(NMNS),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:D1C0D57DBE085331886B26292B7118D3" country="Japan" municipality="Nantou" name="Shanlinchi">Shanlinchi</location>
(杉林溪),
<collectingDate value="1990-05-11">11.V.1990</collectingDate>
, leg.
<collectorName>C. C. Chiang</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2015-04-18" collectorName="B. - X. Guo" country="Japan" location="Sungkang" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:5EB7757A70C950DFE0B5860D3D2DF750" country="Japan" name="Sungkang">Sungkang</location>
(松崗),
<collectingDate value="2015-04-18">18.IV.2015</collectingDate>
, leg.
<collectorName>B.-X. Guo</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1984-09-12" collectingDateMax="1984-09-14" collectingDateMin="1984-09-12" collectorName="K. S. Lin, S. C. Lin" country="Japan" location="Tsuifeng" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
(TARI),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:2C1C9C7C47D3C596C053B409A5E43569" country="Japan" name="Tsuifeng">Tsuifeng</location>
(翠峰),
<collectingDate value="1984-09-12" valueMax="1984-09-14" valueMin="1984-09-12">12-14.IX.1984</collectingDate>
, leg.
<collectorName>K. S. Lin</collectorName>
and
<collectorName>S. C. Lin</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1982-05-07" collectorName="N. Ohbayashi." country="Japan" location="Lalashan" municipality="Taoyuan" specimenCount="1" specimenCount-female="1">
<collectingMunicipality>Taoyuan</collectingMunicipality>
:
<specimenCount type="female">1♀</specimenCount>
(KMNH),
<location LSID="urn:lsid:plazi:treatment:C81A269EDA7D58E88E6432C578A9AEE6:1FB0726B56AAD58415DE2BBC9043F873" country="Japan" municipality="Taoyuan" name="Lalashan">Lalashan</location>
(拉拉山),
<collectingDate value="1982-05-07">7.V.1982</collectingDate>
, leg.
<collectorName>N. Ohbayashi.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Length 4.3-5.4 mm, width 2.4-3.0 mm. Body color (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitus of Pyrrhalta formosanensis sp. nov. and P. semifulva (Jacoby) A P. formosanensis sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D P. semifulva, from Taiwan, female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure27" httpUri="https://binary.pensoft.net/fig/545557" pageId="0" pageNumber="1">27D-F</figureCitation>
) reddish brown; head (including antennae), scutellum, thoracic ventrites, and legs black. Eyes small, interocular space 2.50-2.86
<normalizedToken originalValue="×">x</normalizedToken>
diameter of eye. Antennae filiform in males (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29A</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.6: 0.6: 0.6: 0.6: 0.6: 0.5: 0.5: 0.5: 0.5: 0.8, length to width ratios of antennomeres I-XI 3.1: 2.6: 2.2: 2.1: 2.3: 2.1: 2.0: 2.0: 1.9: 1.9: 3.2; similar in females (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.5: 0.6: 0.5: 0.5: 0.6: 0.5: 0.5: 0.5: 0.5: 0.7, length to width ratios of antennomeres I-XI 3.1: 1.9: 2.2: 1.9: 1.8: 2.0: 1.8: 1.8: 1.9: 1.8: 2.5. Pronotum and elytra convex. Pronotum 2.2-2.3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, disc with reticulate microsculpture; with dense, extremely coarse punctures and extremely short pubescence, with transverse ridge near apical margin that curves downwards near antero-lateral corners, no punctures or pubescence above ridge but coarse punctures present on antero-lateral corners; with median longitudinal and lateral depressions; lateral margins moderately rounded, apical margin slightly concave, basal margin straight; anterior and posterior setiferous punctures slightly erect. Elytra elongate and broad, parallel-sided, 1.4-1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc smooth, with extremely coarse, dense punctures and extremely short pubescence. Apical spur of tibia of middle leg absent, tarsomere I not modified in males. Aedeagus (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29C, D</figureCitation>
) extremely slender in dorsal view, 7.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, sides asymmetric, parallel-sided, apex truncate, curved near apex; strongly curved near base in lateral view, apex narrowly rounded; ostium not covered membrane; two endophallic sclerites elongate, several fine teeth on apex of primary endophallic sclerite, 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, secondary sclerite a little shorter, 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as primary endophallic sclerite, apex acute. Sclerotized gonocoxae (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29K</figureCitation>
) stout and cylindrical, gonocoxae separated, disk with several longer setae mixed with dense, shorter setae. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29G</figureCitation>
) transverse; disc with three layers of different lengths of setae on apical area, shortest setae along apical margin, longest setae a slightly before apex, intermediate setae further from apex; spiculum short. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29H</figureCitation>
) slightly swollen; pump short and strongly curved; sclerotized proximal spermathecal duct wide and short. Apical margin of abdominal ventrite V slightly concave medially, with deep depression at middle in males (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29J</figureCitation>
); only slightly concave in females (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29I</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
Diagnostic characters of
<taxonomicName baseAuthorityName="Jacoby" baseAuthorityYear="1885" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta semifulva</emphasis>
</taxonomicName>
(Jacoby)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna, male
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna, female
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, typical form, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
ditto, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
ditto, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
ermatheca
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
abdominal ventrite V, female
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
abdominal ventrite V, male
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
gonocoxae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
Aedeagi of many individuals have apically tapering apices and look straight in lateral view (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29E, F</figureCitation>
). Japanese populations display great color variation. Some individuals (Hokkaido) have the entire reddish brown bodies but one black spot is present on the vertex, and five apical antennomeres are darkened. Some are similar to the previous ones, but the head is black except mouth parts (Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Habitus of Pyrrhalta semifulva (Jacoby) from Japan A female, color variation, dorsal view B ditto, ventral view C Female, color variation, dorsal view D ditto, ventral view E female, color variation, dorsal view F male, color variation, dorsal view." figureDoi="10.3897/zookeys.1039.64740.figure30" httpUri="https://binary.pensoft.net/fig/545560" pageId="0" pageNumber="1">30A, B</figureCitation>
). Some are similar the previous ones, but the pronotum has one black spot at center, without a clear margin; legs are reddish brown but outer sides of tibiae and entire tarsi are dark brown, antennae and the scutellum are blackish brown (Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Habitus of Pyrrhalta semifulva (Jacoby) from Japan A female, color variation, dorsal view B ditto, ventral view C Female, color variation, dorsal view D ditto, ventral view E female, color variation, dorsal view F male, color variation, dorsal view." figureDoi="10.3897/zookeys.1039.64740.figure30" httpUri="https://binary.pensoft.net/fig/545560" pageId="0" pageNumber="1">30C, D</figureCitation>
). Some specimens are similar to Taiwanese populations but with different degrees of darkness on the pronotum (Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Habitus of Pyrrhalta semifulva (Jacoby) from Japan A female, color variation, dorsal view B ditto, ventral view C Female, color variation, dorsal view D ditto, ventral view E female, color variation, dorsal view F male, color variation, dorsal view." figureDoi="10.3897/zookeys.1039.64740.figure30" httpUri="https://binary.pensoft.net/fig/545560" pageId="0" pageNumber="1">30E, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1039.64740.figure30" httpUri="https://binary.pensoft.net/fig/545560" pageId="0" pageNumber="1" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 30.</emphasis>
Habitus of
<taxonomicName baseAuthorityName="Jacoby" baseAuthorityYear="1885" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta semifulva</emphasis>
</taxonomicName>
(Jacoby) from Japan
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
female, color variation, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
ditto, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Female, color variation, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
ditto, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
female, color variation, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
male, color variation, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Jacoby" baseAuthorityYear="1885" class="Insecta" family="Ericaceae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta semifulva" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">Pyrrhalta semifulva</emphasis>
</taxonomicName>
(Jacoby) and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. may be separated from others within the species group by the reddish brown bodies (Figs
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitus of Pyrrhalta formosanensis sp. nov. and P. semifulva (Jacoby) A P. formosanensis sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D P. semifulva, from Taiwan, female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure27" httpUri="https://binary.pensoft.net/fig/545557" pageId="0" pageNumber="1">27</figureCitation>
,
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Habitus of Pyrrhalta semifulva (Jacoby) from Japan A female, color variation, dorsal view B ditto, ventral view C Female, color variation, dorsal view D ditto, ventral view E female, color variation, dorsal view F male, color variation, dorsal view." figureDoi="10.3897/zookeys.1039.64740.figure30" httpUri="https://binary.pensoft.net/fig/545560" pageId="0" pageNumber="1">30</figureCitation>
) and short, cylindrical gonocoxae with dense setae (Figs
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28J</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29K</figureCitation>
). Taiwanese populations of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. semifulva" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">P. semifulva</emphasis>
</taxonomicName>
differ from
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov. by the black scutellum, legs, and thoracic ventrites (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitus of Pyrrhalta formosanensis sp. nov. and P. semifulva (Jacoby) A P. formosanensis sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D P. semifulva, from Taiwan, female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure27" httpUri="https://binary.pensoft.net/fig/545557" pageId="0" pageNumber="1">27E</figureCitation>
) (reddish brown scutellum, legs, and thoracic ventrites (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitus of Pyrrhalta formosanensis sp. nov. and P. semifulva (Jacoby) A P. formosanensis sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D P. semifulva, from Taiwan, female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure27" httpUri="https://binary.pensoft.net/fig/545557" pageId="0" pageNumber="1">27B</figureCitation>
) in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov.), normal abdominal ventrite V (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29I</figureCitation>
), and unmodified mesotarsomere I of middle leg in males (modified abdominal ventrite V (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28K</figureCitation>
) and mesotarsomere of middle leg (Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Diagnostic characters of Pyrrhalta formosanensis sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D ditto, left-side view E ditto, ventral view F ditto, right-side view G apex of tibia of middle leg, male H abdominal ventrite VIII I spermatheca J gonocoxae K abdominal ventrite V, male L abdominal ventrite V, female M tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure28" httpUri="https://binary.pensoft.net/fig/545558" pageId="0" pageNumber="1">28M</figureCitation>
in males of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. formosanensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="formosanensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. formosanensis</emphasis>
</taxonomicName>
sp. nov.). In males of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. semifulva" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="semifulva">
<emphasis italics="true" pageId="0" pageNumber="1">P. semifulva</emphasis>
</taxonomicName>
, the elongate and apically curved aedeagus is similar to that of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. discalis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="discalis">
<emphasis italics="true" pageId="0" pageNumber="1">P. discalis</emphasis>
</taxonomicName>
Gressitt and Kimoto, but differs in the relatively longer secondary endophallic sclerite, 0.9x as long as primary endophallic sclerite (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Diagnostic characters of Pyrrhalta semifulva (Jacoby) A antenna, male B antenna, female C aedeagus, typical form, dorsal view D ditto, lateral view E aedeagus, variation (endophallic sclerites omitted), dorsal view apex of tibia of middle leg, male F ditto, lateral view G abdominal ventrite VIII H ermatheca I abdominal ventrite V, female J abdominal ventrite V, male K gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure29" httpUri="https://binary.pensoft.net/fig/545559" pageId="0" pageNumber="1">29C, D</figureCitation>
) (vs. relatively shorter secondary endophallic sclerite, 0.6x as long as primary endophallic sclerite in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. discalis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="discalis">
<emphasis italics="true" pageId="0" pageNumber="1">P. discalis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Diagnostic characters of Pyrrhalta discalis Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E abdominal ventrite VIII F spermatheca G abdominal ventrite V, female H abdominal ventrite V, male I gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure32" httpUri="https://binary.pensoft.net/fig/545562" pageId="0" pageNumber="1">32C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="food plants">
<paragraph pageId="0" pageNumber="1">Food plants.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Rosaceae</taxonomicName>
:
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Prunus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Prunus jamasakura" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="jamasakura">
<emphasis italics="true" pageId="0" pageNumber="1">Prunus jamasakura</emphasis>
</taxonomicName>
Sieb., ex Koidz.,
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. yedoensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="yedoensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. yedoensis</emphasis>
</taxonomicName>
Matsum., and
<taxonomicName class="Dicotyledoneae" family="Hamamelidaceae" genus="Sorbus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sorbus japonica" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="japonica">
<emphasis italics="true" pageId="0" pageNumber="1">Sorbus japonica</emphasis>
</taxonomicName>
(Decne.) Hedl.;
<taxonomicName class="Dicotyledoneae" family="Hamamelidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Hamamelidaceae</taxonomicName>
:
<taxonomicName authorityName="Makino" authorityYear="1901" class="Dicotyledoneae" family="Hamamelidaceae" genus="Corylopsis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Corylopsis gotoana" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="gotoana">
<emphasis italics="true" pageId="0" pageNumber="1">Corylopsis gotoana</emphasis>
</taxonomicName>
Makino, (
<bibRefCitation author="Chujo, M" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="1" pagination="117 - 202" refId="B13" refString="Chujo, M, Kimoto, S, 1961. Systematic catalog of Japanese Chrysomelidae (Coleoptera). Pacific Insects 3: 117 - 202" title="Systematic catalog of Japanese Chrysomelidae (Coleoptera)." volume="3" year="1961">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
and Kimoto 1961
</bibRefCitation>
). adults of Taiwanese populations feed on flowers of
<taxonomicName authorityName="Hayata" authorityYear="1911" class="Dicotyledoneae" family="Rosaceae" genus="Photinia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Photinia niitakayamensis" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="niitakayamensis">
<emphasis italics="true" pageId="0" pageNumber="1">Photinia niitakayamensis</emphasis>
</taxonomicName>
Hayata (
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Rosaceae</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Field photographs of adults of Pyrrhalta species on host plants A P. tsoui Bezdek &amp; Lee B P. formosanensis sp. nov. C P. semifulva (Jacoby) D P. ishiharai Kimoto." figureDoi="10.3897/zookeys.1039.64740.figure26" httpUri="https://binary.pensoft.net/fig/545556" pageId="0" pageNumber="1">26C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Japan, Russian, Taiwan. The species is widespread at mid-altitudes (1,500-2,500 m) in Taiwan.</paragraph>
</subSubSection>
</treatment>
</document>
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