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<mods:title id="DA692BEE1128FDBE6D45E79DBCC1D8B5">Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)</mods:title>
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<mods:namePart id="D58F9B2BD6E51BF0F03A0859EF97AE59">Han, Hui-Lin</mods:namePart>
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<mods:affiliation id="A58C309B9B8AA3A94096634E775665AE">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:namePart id="4265DFA9C152C09B744F39ABF79867AF">Kononenko, Vladimir S.</mods:namePart>
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<mods:affiliation id="8E6043BF9049966871C586621BCCD5B0">Northeast Asia Biodiversity Research Center, Northeast Forestry University, Harbin 15004, China</mods:affiliation>
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<mods:namePart id="EF419877D8F0E77AB2C547905D52E6CB">Pan, Zhao-Hui</mods:namePart>
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<treatment id="C0854133FC925C6C8C7B4FA428762C5B" ID-GBIF-Taxon="212973837" LSID="urn:lsid:plazi:treatment:C0854133FC925C6C8C7B4FA428762C5B" httpUri="http://treatment.plazi.org/id/C0854133FC925C6C8C7B4FA428762C5B" lastPageNumber="35" pageId="0" pageNumber="35" scope_family="Noctuidae" scope_order="Lepidoptera" scope_subFamily="Noctuinae">
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<taxonomicName id="1B4DECD45BE4E6AA45385A38CD3E388E" ID-CoL="6M3G6" LSID="C0854133-FC92-5C6C-8C7B-4FA428762C5B" authority="Moore, 1882" authorityName="Moore" authorityYear="1882" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa cuprina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cuprina">Hermonassa cuprina Moore, 1882</taxonomicName>
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<paragraph id="B882A9C9DB2A9AEFE1E5D41FA96F36C3" pageId="0" pageNumber="35">
<figureCitation id="BFEFBE96ACB513ED2BE38AF33AED40CF" captionStart="Figures 2834" captionStartId="F4" captionText="Figures 28 - 34. Hermonassa spp., adults (28 - 30), male (31 - 32) and female (33 - 34) genitalia 28 H. brunneocuprina sp. nov., ♂, holotype 29 Ditto, ♀, paratype 30 H. cuprina ¸ Nepal (after Sugi 1995) 31 H. brunneocuprina sp. nov., holotype, genit. prep. GB- 283 - 1 32 H. cuprina (after Sugi 1995) 33 H. brunneocuprina sp. nov., paratype, genit. prep. GB- 322 - 2 34 H. cuprina, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures28-34" httpUri="https://binary.pensoft.net/fig/903266" pageId="0" pageNumber="35">Figs 30</figureCitation>
<figureCitation id="1C4E38A8F12EC7B2D0ABD43A38CC6C2B" captionStart="Figures 2834" captionStartId="F4" captionText="Figures 28 - 34. Hermonassa spp., adults (28 - 30), male (31 - 32) and female (33 - 34) genitalia 28 H. brunneocuprina sp. nov., ♂, holotype 29 Ditto, ♀, paratype 30 H. cuprina ¸ Nepal (after Sugi 1995) 31 H. brunneocuprina sp. nov., holotype, genit. prep. GB- 283 - 1 32 H. cuprina (after Sugi 1995) 33 H. brunneocuprina sp. nov., paratype, genit. prep. GB- 322 - 2 34 H. cuprina, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures28-34" httpUri="https://binary.pensoft.net/fig/903266" pageId="0" pageNumber="35">, 32</figureCitation>
<figureCitation id="09852711B6998DBE13D4E913DFEAAE0D" captionStart="Figures 2834" captionStartId="F4" captionText="Figures 28 - 34. Hermonassa spp., adults (28 - 30), male (31 - 32) and female (33 - 34) genitalia 28 H. brunneocuprina sp. nov., ♂, holotype 29 Ditto, ♀, paratype 30 H. cuprina ¸ Nepal (after Sugi 1995) 31 H. brunneocuprina sp. nov., holotype, genit. prep. GB- 283 - 1 32 H. cuprina (after Sugi 1995) 33 H. brunneocuprina sp. nov., paratype, genit. prep. GB- 322 - 2 34 H. cuprina, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures28-34" httpUri="https://binary.pensoft.net/fig/903266" pageId="0" pageNumber="35">, 34</figureCitation>
</paragraph>
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<subSubSection id="785E665A4866DB10A8D965048B632196" pageId="0" pageNumber="35" type="reference_group">
<paragraph id="FE52415B03B55CD2A198EB6002317A2F" pageId="0" pageNumber="35">
<taxonomicName id="67E618E8C16EE02001C446A4CB7838BD" authorityName="Moore" authorityYear="1882" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa cuprina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cuprina">Hermonassa cuprina</taxonomicName>
Moore, 1882, Descriptions of new Indian Lepidopterous Insects from the Collection of the Late Mr. W.S. Atkinson.
<taxonomicName id="52ED70CB598DDB8AB3BFD6E74D6403D8" authorityName="Walker" authorityYear="1865" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="subOrder" subOrder="Heterocera">Heterocera</taxonomicName>
(continued) (
<taxonomicName id="2CBA6421BDDC8F2B4307E4D0D6D4FDD0" authorityName="Herrich-Schaffer" authorityYear="1847" class="Insecta" family="Cymatophoridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="family">Cymatophoridae</taxonomicName>
-
<taxonomicName id="EB101F27BB486140F2B93CB2EA5D3A06" class="Insecta" family="Herminiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="family">Herminiidae</taxonomicName>
). Part II: 120;
<bibRefCitation id="D48E0329CF6568C96DE94424599802F4" author="Hampson, GF" journalOrPublisher="Alfred Kernen - Verlag, Stuttgart" pageId="0" pageNumber="35" refId="B10" refString="Hampson, GF, 1903. Catalogue of the Lepidoptera Phalaenae in the British Museum. Vol. 4 (Heliothidinae, Noctuinae). London. Taylor and Francis, [xiv +] 689 pp." title="Catalogue of the Lepidoptera Phalaenae in the British Museum. Vol. 4 (Heliothidinae, Noctuinae). London. Taylor and Francis, [xiv +] 689 pp." year="1903">Hampson 1903</bibRefCitation>
: 357;
<bibRefCitation id="9EA4ECA2A9DB95B1555A2129F36A352A" author="Boursin, C" journalOrPublisher="Zeitschrift der Wiener Entomologischen Gesellschaft" pageId="0" pageNumber="35" pagination="21 - 38" refId="B2" refString="Boursin, C, 1967. Die neuen Hermonassa Walk.-arten aus Dr. H. Hӧne 's China ausbeuten. Zeitschrift der Wiener Entomologischen Gesellschaft 52: 21 - 38" title="Die neuen Hermonassa Walk. - arten aus Dr. H. Hӧne ' s China ausbeuten." volume="52" year="1967">Boursin 1967</bibRefCitation>
: 36; Poole, 1989: 502;
<bibRefCitation id="644DD25D4B6FB870F9A80E68FE4E11D8" author="Sugi, S" editor="Haruta, T" journalOrPublisher="Tinea 14 (suppl. 2)" pageId="0" pageNumber="35" pagination="90 - 109" refId="B25" refString="Sugi, S, 1995. Noctuidae: Noctuinae, Hermonassa. In: Haruta, T, Ed., Moths of Nepal IV. Tinea 14 (suppl. 2): 90 - 109" title="Noctuidae: Noctuinae, Hermonassa." volumeTitle="Moths of Nepal IV." year="1995">Sugi 1995</bibRefCitation>
: 90, pl. 117, fig. 1, genit. fig. 691(♂), 721 (♀);
<bibRefCitation id="10CDBC9AF4607F461D81D0F03A7C7DAE" author="Krusek, K" journalOrPublisher="Agrotinae), Vol. 13 (3). Institut Zoologique de L'Academie des sciences de l'USSR (N. S. 15). Moskva - Leningrad" pageId="0" pageNumber="35" refId="B18" refString="Krusek, K, Behounek, G, 1996a. Noctuidae - Typen 1 (Lepidoptera, Noctuidae). Selbstverlag: Krusek, Prag, Vol. 1, pls 1-114." title="Noctuidae - Typen 1 (Lepidoptera, Noctuidae). Selbstverlag: Krusek, Prag, Vol. 1, pls 1 - 114." year="1996 a">Krusek and Behounek 1996a</bibRefCitation>
: pl. 50, fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="B1323B50C7BDE083506F662FEFD15D2E" pageId="0" pageNumber="35" type="type material">
<paragraph id="AC055A33AA53F52B5B712CA3FC358651" pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph id="86905D1C9AAB214D70221867FF09B9CE" pageId="0" pageNumber="35">
<emphasis id="EAEAE2F1FBD81376A51E20E4CBE32180" bold="true" italics="true" pageId="0" pageNumber="35">Syntype(s)</emphasis>
: India, Prov. West Bengal, Darjeeling, Deposited in NHM London, United Kingdom and MNHU, Berlin, Germany), not examined.
</paragraph>
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<subSubSection id="EEE983D0125C062FCFA99FD5EE783151" pageId="0" pageNumber="35" type="distribution">
<paragraph id="133968E29D72A72070F3DAF2194DA351" pageId="0" pageNumber="35">Distribution and biology.</paragraph>
<paragraph id="031120BD09FB991058B6B9D7BF41E1A4" pageId="0" pageNumber="35">North India: Prov. Sikkim, Prov. West Bengal, Darjiling [Darjeeling].</paragraph>
</subSubSection>
<subSubSection id="222892EBFFF642750920D9D9B537CE0D" pageId="0" pageNumber="35" type="remarks">
<paragraph id="BDC2145B57E0631907CE8EBFDCC4D854" pageId="0" pageNumber="35">Remarks.</paragraph>
<paragraph id="29397113BEB4CEABD5D1E4628412752D" pageId="0" pageNumber="35">
<taxonomicName id="CB89BCDE1CC58818DC9E00A3B6A17485" authorityName="Moore" authorityYear="1882" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa cuprina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cuprina">
<emphasis id="1E22A103C2954F12F4D9409292D62CA3" italics="true" pageId="0" pageNumber="35">Hermonassa cuprina</emphasis>
</taxonomicName>
has not yet been found in China. The syntypes of this species are deposited in NHM UK and MNHU (
<bibRefCitation id="94309C69D706E0170CAE11176B77883C" author="Moore, F" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="0" pageNumber="35" refId="B21" refString="Moore, F, 1882. Descriptions of new Indian Lepidopterous Insects from the Collection of the Late Mr. W.S. Atkinson. Heterocera (continued) (Cymatophoridae-Herminiidae). Part II. Taylor and Francis, London, 89-198. [pls 4-6]" title="Descriptions of new Indian Lepidopterous Insects from the Collection of the Late Mr. W. S. Atkinson. Heterocera (continued) (Cymatophoridae-Herminiidae). Part II. Taylor and Francis, London, 89 - 198. [pls 4 - 6]" year="1882">Moore 1882</bibRefCitation>
;
<bibRefCitation id="26C9524F5CB1944F1890A9B604F3A565" author="Poole, RW" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="35" refId="B23" refString="Poole, RW, 1989. Noctuidae. Lepidopterorum Catalogues (New Series). Fascicle 118, pt 1-3. E.J. Brill, Leiden" title="Noctuidae. Lepidopterorum Catalogues (New Series). Fascicle 118, pt 1 - 3. E. J." year="1989">Poole 1989</bibRefCitation>
). The lectotype is not designated. For the illustration of
<taxonomicName id="AC68772093B0FA9DF688F001C8BD863B" class="Insecta" family="Noctuidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="H. cuprina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cuprina">
<emphasis id="87177554FEE7B28AFE8910B17FE09F42" italics="true" pageId="0" pageNumber="35">H. cuprina</emphasis>
</taxonomicName>
(Figs
<figureCitation id="F7F8CA8D5160D5A8529F3D7FD10CC37D" captionStart="Figures 2834" captionStartId="F4" captionText="Figures 28 - 34. Hermonassa spp., adults (28 - 30), male (31 - 32) and female (33 - 34) genitalia 28 H. brunneocuprina sp. nov., ♂, holotype 29 Ditto, ♀, paratype 30 H. cuprina ¸ Nepal (after Sugi 1995) 31 H. brunneocuprina sp. nov., holotype, genit. prep. GB- 283 - 1 32 H. cuprina (after Sugi 1995) 33 H. brunneocuprina sp. nov., paratype, genit. prep. GB- 322 - 2 34 H. cuprina, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures28-34" httpUri="https://binary.pensoft.net/fig/903266" pageId="0" pageNumber="35">30</figureCitation>
,
<figureCitation id="D62D86D5AB9567584FF88F1D273A62A2" captionStart="Figures 2834" captionStartId="F4" captionText="Figures 28 - 34. Hermonassa spp., adults (28 - 30), male (31 - 32) and female (33 - 34) genitalia 28 H. brunneocuprina sp. nov., ♂, holotype 29 Ditto, ♀, paratype 30 H. cuprina ¸ Nepal (after Sugi 1995) 31 H. brunneocuprina sp. nov., holotype, genit. prep. GB- 283 - 1 32 H. cuprina (after Sugi 1995) 33 H. brunneocuprina sp. nov., paratype, genit. prep. GB- 322 - 2 34 H. cuprina, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures28-34" httpUri="https://binary.pensoft.net/fig/903266" pageId="0" pageNumber="35">32</figureCitation>
,
<figureCitation id="52D04C472950106424065778D884F5AB" captionStart="Figures 2834" captionStartId="F4" captionText="Figures 28 - 34. Hermonassa spp., adults (28 - 30), male (31 - 32) and female (33 - 34) genitalia 28 H. brunneocuprina sp. nov., ♂, holotype 29 Ditto, ♀, paratype 30 H. cuprina ¸ Nepal (after Sugi 1995) 31 H. brunneocuprina sp. nov., holotype, genit. prep. GB- 283 - 1 32 H. cuprina (after Sugi 1995) 33 H. brunneocuprina sp. nov., paratype, genit. prep. GB- 322 - 2 34 H. cuprina, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures28-34" httpUri="https://binary.pensoft.net/fig/903266" pageId="0" pageNumber="35">34</figureCitation>
) we follow the treatment and illustrations in
<bibRefCitation id="26EA6EA29A0745F6894620DD028978D3" author="Sugi, S" editor="Haruta, T" journalOrPublisher="Tinea 14 (suppl. 2)" pageId="0" pageNumber="35" pagination="90 - 109" refId="B25" refString="Sugi, S, 1995. Noctuidae: Noctuinae, Hermonassa. In: Haruta, T, Ed., Moths of Nepal IV. Tinea 14 (suppl. 2): 90 - 109" title="Noctuidae: Noctuinae, Hermonassa." volumeTitle="Moths of Nepal IV." year="1995">Sugi (1995)</bibRefCitation>
of this species from north India.
</paragraph>
</subSubSection>
<subSubSection id="DA4D2CEB6D164D1EE5955C75AA0E43E0" pageId="0" pageNumber="35" type="the dictyota species group">
<paragraph id="DDA9C3CD9A9D12382312EA5F00F7A5CC" pageId="0" pageNumber="35">
The
<taxonomicName id="8973BAD49D3A01355291F6196E480C41" authorityName="Boursin" authorityYear="1967" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa dictyota" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="dictyota">
<emphasis id="E6CE231976069B9B7EB632956981DCC6" italics="true" pageId="0" pageNumber="35">Hermonassa dictyota</emphasis>
</taxonomicName>
species group
</paragraph>
<paragraph id="E45469625D224BF8CA68A7E6D2C05185" pageId="0" pageNumber="35">
The
<taxonomicName id="C135DFDC93E9758101C287BCED146664" authorityName="Boursin" authorityYear="1967" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa dictyota" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="dictyota">
<emphasis id="2CD3229E6903FE70B2AE942677955186" italics="true" pageId="0" pageNumber="35">Hermonassa dictyota</emphasis>
</taxonomicName>
species group externally can be characterized by the relatively large size (wingspan 38-40 mm), the robust habitus and the reticulate forewing pattern atypical for most
<taxonomicName id="20C110401F31FE99F5B7BD3DDC64049C" authorityName="Walker" authorityYear="1865" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="F1652492B3179821B077238B14C4B040" italics="true" pageId="0" pageNumber="35">Hermonassa</emphasis>
</taxonomicName>
spp. The male genitalia of these species are rather uniform and characterized by having an apically swollen or spatulate uncus, anchor-like, apically hooked juxta, moderate harpe, variable in length saccular extension and the presence of a pollex at the apical part of the valva. In the female genitalia, the antevaginal plate is large, with a deep cut in the centre, a sclerotised antrum, the relatively short ductus bursae and the sclerotised proximal part of the corpus bursae. The
<taxonomicName id="4938B60A67FF4AFC850A873ED16BDB27" authorityName="Boursin" authorityYear="1967" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa dictyota" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="dictyota">
<emphasis id="801643646A7F3F0E5BCD15B4FAB28EFB" italics="true" pageId="0" pageNumber="35">Hermonassa dictyota</emphasis>
</taxonomicName>
species group comprises the following four species:
<taxonomicName id="7EA5AEBEA43B88CC71FCAF58F029AF06" class="Insecta" family="Noctuidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="H. dictyota" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="dictyota">
<emphasis id="778A315646CB19E556A4D5363E5CDAD6" italics="true" pageId="0" pageNumber="35">H. dictyota</emphasis>
</taxonomicName>
,
<taxonomicName id="4FEE65548A77A45C3C952CC23974BAC6" class="Insecta" family="Noctuidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="H. legraini" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="legraini">
<emphasis id="AAAB7AD73267845D93BA4AD8A64FD51E" italics="true" pageId="0" pageNumber="35">H. legraini</emphasis>
</taxonomicName>
,
<taxonomicName id="5FD3FCF476AC5651CE0C85EAF762F06C" class="Insecta" family="Noctuidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="H. yixincheni" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="yixincheni">
<emphasis id="F155C2AA850748049FAF7FD9BF16D86B" italics="true" pageId="0" pageNumber="35">H. yixincheni</emphasis>
</taxonomicName>
, and
<taxonomicName id="FB2EE3D50E4E5B1378BDCC779E71FF1A" class="Insecta" family="Noctuidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="H. albimacula" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="albimacula">
<emphasis id="311F7B670C154956FC82662B023C271B" italics="true" pageId="0" pageNumber="35">H. albimacula</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
</treatment>
</document>