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<document id="CAED09817E2C892BA0C5D94D1F2A4989" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFDDFFC33C2C60AAFEE17304" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechepaphiopsis uniporosa Deuve and Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="415" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="413" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="5361D80B5B76E0A3D44341D48C80C412">
<mods:title id="EB068AB7FAC1FAE43D86B5F3F62C2A54">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:name id="83A008EB5BCE56AA74918305EEC4341D" type="personal">
<mods:role id="48A926CCC49791E54AD0F0BEFEC03450">
<mods:roleTerm id="8539E0AE7D7D378C61B4312473173ADA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="30BB27F8ECFE981BC6B4B37617BFDFA2">Deuve, Thierry</mods:namePart>
<mods:affiliation id="9549DB5A661B98C9F36573AEF0A6DEBB">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
<mods:name id="499A34BB94A022DAF69F8DA72C770CF1" type="personal">
<mods:role id="608D1A93F49BD1E177DB7241F2963510">
<mods:roleTerm id="20990D8E4CB3DCD17A43DBF49C275F7A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CDC0DEAFB53EFB30411F07B75E1ADEEB">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="10DAF552503634D27B92C62E68803E82">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="4F33AFBF786C1FE2E9E2B07505A9D665" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="D5F15433D8A30E2E02B02B1651D35DA4" type="personal">
<mods:role id="14AF1B8B8D6E0FF5BAD7E8E8BA5E492C">
<mods:roleTerm id="434219FF2EF87FA676EE4EE391DF28E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B69A2D87620FFF59598D7142AC747D6D">Liang, Hongbin</mods:namePart>
<mods:affiliation id="D92BA4F25A266DED95D061692640B1BB">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:typeOfResource id="335E05B488195F1629D42B622BECFE0D">text</mods:typeOfResource>
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<mods:title id="78551B56AD04A19204869F268A00B807">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="ED8BB980F5FFBAF1577BB7B19EB78C48">
<mods:date id="763E6AE3A9D6EACF157AABB734EFF749">2016</mods:date>
<mods:detail id="4248DF56AD97DE6357601610ECA5FFF4" type="pubDate">
<mods:number id="D0B6849260603EA98B6E2C00FDB72756">2016-10-14</mods:number>
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<mods:number id="8E03EE8B50AC2887E451826B1B6B8195">63</mods:number>
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<treatment id="03D6879DFFDDFFC33C2C60AAFEE17304" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304" httpUri="http://treatment.plazi.org/id/03D6879DFFDDFFC33C2C60AAFEE17304" lastPageId="74" lastPageNumber="415" pageId="72" pageNumber="413">
<subSubSection id="C3656500FFDDFFC13C2C60AAFD2F77A3" box="[123,737,219,243]" pageId="72" pageNumber="413" type="nomenclature">
<paragraph id="8BC0368BFFDDFFC13C2C60AAFD2F77A3" blockId="72.[123,737,219,273]" box="[123,737,219,243]" pageId="72" pageNumber="413">
<heading id="D08881E7FFDDFFC13C2C60AAFD2F77A3" box="[123,737,219,243]" fontSize="10" level="2" pageId="72" pageNumber="413" reason="4">
<emphasis id="B90BEA99FFDDFFC13C2C60AAFD2F77A3" bold="true" box="[123,737,219,243]" pageId="72" pageNumber="413">
<taxonomicName id="4C7F4D08FFDDFFC13C2C60AAFDB077A3" ID-CoL="8MLL2" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,638,219,243]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDDFFC13C2C60AAFE7C77A3" bold="true" box="[123,434,219,243]" italics="true" pageId="72" pageNumber="413">Trechepaphiopsis uniporosa</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFDDFFC13EDC60AAFD2F77A3" box="[651,737,219,243]" pageId="72" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC13C2C608AFEB37641" box="[123,381,251,273]" pageId="72" pageNumber="413" type="description">
<paragraph id="8BC0368BFFDDFFC13C2C608AFEB37641" blockId="72.[123,737,219,273]" box="[123,381,251,273]" pageId="72" pageNumber="413">
(
<figureCitation id="13442A0EFFDDFFC13CD5608AFF157641" box="[130,219,251,273]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="72" pageNumber="413">Figs. 16f</figureCitation>
,
<figureCitation id="13442A0EFFDDFFC13CB2608AFF317640" box="[229,255,251,272]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="72" pageNumber="413">28</figureCitation>
,
<figureCitation id="13442A0EFFDDFFC13D5D608AFEE07640" box="[266,302,251,272]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="72" pageNumber="413">37a</figureCitation>
,
<figureCitation id="13442A0EFFDDFFC13D6E608AFEB97641" box="[313,375,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="72" pageNumber="413">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC13CFC6155FC46758C" pageId="72" pageNumber="413" type="materials_examined">
<paragraph id="8BC0368BFFDDFFC13CFC6155FBD0756C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<emphasis id="B90BEA99FFDDFFC13CFC6155FE97766A" bold="true" box="[171,345,292,316]" pageId="72" pageNumber="413">
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<smallCapsWord id="8D26A057FFDDFFC13CBB6156FE97766A" baselines="309" box="[236,345,295,314]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="72" pageNumber="413">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFDDFFC13DD56155FF4A76EC" collectingDate="1998-10-15" collectingDateMax="1998-10-18" collectingDateMin="1998-10-15" collectorName="C. E. Griswold &amp; C. Ferraris &amp; C. - L. Long" country="China" location="Nujiang Prefecture" municipality="Gaoligongshan Mountains" pageId="72" pageNumber="413" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFDDFFC13DD56155FE26766C" box="[386,488,292,316]" pageId="72" pageNumber="413" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1001923”/ “
<collectingCountry id="F368761BFFDDFFC13FAB6155FB9B766C" box="[1020,1109,292,316]" name="China" pageId="72" pageNumber="413">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDDFFC138366155FEC4760C" country="China" name="Yunnan" pageId="72" pageNumber="413">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFDDFFC13D426135FDFD760C" box="[277,563,324,348]" pageId="72" pageNumber="413">Gaoligongshan Mountains</collectingMunicipality>
,
<location id="8EA06050FFDDFFC13E696135FCC5760C" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDDFFC13E696135FCC5760C" box="[574,779,324,348]" country="China" municipality="Gaoligongshan Mountains" name="Nujiang Prefecture" pageId="72" pageNumber="413" stateProvince="Yunnan">Nujiang Prefecture</location>
, 9 [actually 9.3] km
<locationDeviation id="9A722FA9FFDDFFC13FBD6135FB5C760C" box="[1002,1170,324,348]" location="Pianma" pageId="72" pageNumber="413">ESE of Pianma</locationDeviation>
, 25°59.6N/ 98°37.6E.”/ “2450 [actually 2460-2470] m,
<date id="FFC1104BFFDDFFC13E816115FC61762C" box="[726,943,356,380]" pageId="72" pageNumber="413" value="1998-10-15" valueMax="1998-10-18" valueMin="1998-10-15">
<collectingDate id="EF85E9A3FFDDFFC13E816115FC61762C" box="[726,943,356,380]" pageId="72" pageNumber="413" value="1998-10-15" valueMax="1998-10-18" valueMin="1998-10-15">15-18 October 1998</collectingDate>
</date>
, Stop #98-118D D.H.
<taxonomicName id="4C7F4D08FFDDFFC13C2C61F5FF3976CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,388,412]" pageId="72" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFDDFFC13D5E61F5FE6376CC" box="[265,429,388,412]" pageId="72" pageNumber="413">C.E. Griswold</collectorName>
,
<collectorName id="268A535DFFDDFFC13DE861F5FDF776CC" box="[447,569,388,412]" pageId="72" pageNumber="413">C. Ferraris</collectorName>
&amp;
<collectorName id="268A535DFFDDFFC13E3C61F5FD2776CC" box="[619,745,388,412]" pageId="72" pageNumber="413">C.-L. Long</collectorName>
collectors”/ “IMAGE” [green label]/ “
</materialsCitation>
<typeStatus id="54C48829FFDDFFC13CD361D5FEDD76EC" box="[132,275,420,444]" pageId="72" pageNumber="413" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDDFFC13D4D61D5FD3776EC" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[282,761,420,444]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDDFFC13D4D61D5FD8676EC" box="[282,584,420,444]" italics="true" pageId="72" pageNumber="413">Trechepaphiopsis uniporosa</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFDDFFC13F5361D5FC9676EB" box="[772,856,420,443]" pageId="72" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label].
<materialsCitation id="3B173CD6FFDDFFC13C2C61B5FBD7756C" collectionCode="CAS, IOZ, MNHN" country="China" pageId="72" pageNumber="413" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFDDFFC13C2C61B5FF2D768C" box="[123,227,452,476]" pageId="72" pageNumber="413" type="paratype">Paratypes</typeStatus>
(a total of 5):
<specimenCount id="9D79FD02FFDDFFC13D2861B5FE1C768C" box="[383,466,452,476]" count="3" pageId="72" pageNumber="413" type="male">3 males</specimenCount>
and
<specimenCount id="9D79FD02FFDDFFC13E5061B5FDA1768C" box="[519,623,452,476]" count="2" pageId="72" pageNumber="413" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFDDFFC13ECB61B5FD1C768C" box="[668,722,452,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="72" pageNumber="413" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFDDFFC13E8A61B5FCC2768C" box="[733,780,452,476]" pageId="72" pageNumber="413">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFDDFFC13F4F61B5FCA5768C" box="[792,875,452,476]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="72" pageNumber="413">MNHN</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFDDFFC1386661B5FB5F768C" box="[1073,1169,452,476]" pageId="72" pageNumber="413" type="holotype">holotype</typeStatus>
, except first label “CASENT 1001920” to “CASENT 1001922” and “CASENT 1001924” to “CASENT 1001925”, respectively. All
<typeStatus id="54C48829FFDDFFC13E606274FD53754D" box="[567,669,517,541]" pageId="72" pageNumber="413" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFDDFFC138596275FB5B754C" box="[1038,1173,516,540]" pageId="72" pageNumber="413" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDDFFC13C2C6255FD94756C" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,602,548,572]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDDFFC13C2C6255FE66756C" box="[123,424,548,572]" italics="true" pageId="72" pageNumber="413">Trechepaphiopsis uniporosa</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFDDFFC13E326254FD76756C" box="[613,696,549,572]" pageId="72" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC6235FD29752C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<materialsCitation id="3B173CD6FFDDFFC13CFC6235FD2D752C" country="China" county="Lushui County" elevation="2465" location="Pianma Township" municipality="Gaoligong Shan" pageId="72" pageNumber="413" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54C48829FFDDFFC13CFC6235FF2A750B" box="[171,228,580,604]" pageId="72" pageNumber="413">TYPE</typeStatus>
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</emphasis>
.—
<collectingCountry id="F368761BFFDDFFC13DD46235FE0B750C" box="[387,453,580,604]" name="China" pageId="72" pageNumber="413">China</collectingCountry>
,
<collectingRegion id="49BBF869FFDDFFC13D856234FDE8750C" box="[466,550,581,604]" country="China" name="Yunnan" pageId="72" pageNumber="413">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFDDFFC13E636235FD28750C" box="[564,742,580,604]" pageId="72" pageNumber="413">Gaoligong Shan</collectingMunicipality>
,
<collectingCounty id="62A14E07FFDDFFC13EA46235FC58750C" box="[755,918,580,604]" pageId="72" pageNumber="413">Lushui County</collectingCounty>
,
<location id="8EA06050FFDDFFC13FF46235FBA9750C" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDDFFC13FF46235FBA9750C" box="[931,1127,580,604]" country="China" county="Lushui County" municipality="Gaoligong Shan" name="Pianma Township" pageId="72" pageNumber="413" stateProvince="Yunnan">Pianma Township</location>
,
<locationDeviation id="9A722FA9FFDDFFC138236235FE85752C" location="Pianma" pageId="72" pageNumber="413">
<quantity id="4C879B6EFFDDFFC138236235FF53752C" metricMagnitude="3" metricUnit="m" metricValue="9.3" pageId="72" pageNumber="413" unit="km" value="9.3">9.3 km</quantity>
ESE of Pianma
</locationDeviation>
, 25.99363°/ 98.66651°,
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<elevation id="0052D1B8FFDDFFC13E076215FD2D752C" box="[592,739,612,636]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="72" pageNumber="413" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">2460-2470 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC62F5FC46758C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
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.— The species epithet,
<taxonomicName id="4C7F4D08FFDDFFC13EA662F4FC9275CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[753,860,645,668]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa">
<emphasis id="B90BEA99FFDDFFC13EA662F4FC9275CC" box="[753,860,645,668]" italics="true" pageId="72" pageNumber="413">uniporosa</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B90BEA99FFDDFFC13D6662D4FEAD75EC" box="[305,355,677,700]" italics="true" pageId="72" pageNumber="413">unus</emphasis>
, meaning one, and
<emphasis id="B90BEA99FFDDFFC13E6062D4FDBA75EC" box="[567,628,677,700]" italics="true" pageId="72" pageNumber="413">porus</emphasis>
, meaning pore or hole. The name refers to the single discal setiferous pore found on the elytra of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC13CFC6295FF23734C" pageId="72" pageNumber="413" type="diagnosis">
<paragraph id="8BC0368BFFDDFFC13CFC6295FF23734C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
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<emphasis id="B90BEA99FFDDFFC13CFC6295FEEE75AB" bold="true" box="[171,288,740,764]" pageId="72" pageNumber="413">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFFDDFFC13E106295FD6D75AC" box="[583,675,740,764]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="72" pageNumber="413">Fig. 28a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
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), apterous; body color light brown, slightly iridescent; eyes small; tempora distinctly convex, sparsely pubescent; pronotum transverse (ratio PW/PL = 1.34), with basal angles obtuse and rounded; elytra convex, with stria 1 deeply impressed and finely punctate, other striae more faintly impressed and lateral striae effaced; recurrent stria terminated anteriorly in presumed location of interval 6; only one discal setiferous pore present, inserted at anterior one-third next to stria 3; preapical seta present, inserted next to stria 3; median lobe of male aedeagus (
<figureCitation id="13442A0EFFDDFFC13F9563B5FBD0748C" box="[962,1054,964,988]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="72" pageNumber="413">Fig. 28b</figureCitation>
) with apex slender but bluntly rounded and slightly recurved dorsally, endophallus with a faintly sclerotized scaly area.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC33CFC6454FEE17304" lastPageId="74" lastPageNumber="415" pageId="72" pageNumber="413" type="description">
<paragraph id="8BC0368BFFDDFFC13CFC6454FCDA730D" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
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<emphasis id="B90BEA99FFDDFFC13CFC6454FEF4736B" bold="true" box="[171,314,1061,1085]" pageId="72" pageNumber="413">DESCRIPTION</emphasis>
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.— Size medium BL =
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. Body color light brown, appendages concolorous, except palpi paler, dorsum shiny, slightly iridescent.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC6414FBE972CD" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<materialsCitation id="3B173CD6FFDDFFC13CFC6414FBED72CD" location="Antennae" municipality="Submentum" pageId="72" pageNumber="413" specimenCount="16" stateProvince="Chin">
Head. Moderate in size, short; eyes small, not projected, their diameter about equal to length of tempora but their convexity less than that of tempora, the latter markedly convex, inflated and sparsely pubescent. Frontal furrows thin and linear, deeply impressed, arcuate, continuous posteriorly to hind margins of tempora. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave or emarginate. Mandibles short and slender, right mandible as in
<figureCitation id="13442A0EFFDDFFC13F756574FCB2724D" box="[802,892,1285,1309]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="72" pageNumber="413">
Fig.
<specimenCount id="9D79FD02FFDDFFC13F026574FCB2724D" box="[853,892,1285,1309]" count="16" pageId="72" pageNumber="413" type="generic">16f</specimenCount>
</figureCitation>
.
<collectingRegion id="49BBF869FFDDFFC13FDF6574FC2C724C" box="[904,994,1285,1308]" country="Myanmar" name="Chin" pageId="72" pageNumber="413">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFDDFFC13CA66554FE85726C" box="[241,331,1317,1340]" country="Myanmar" name="Chin" pageId="72" pageNumber="413">Mentum</collectingRegion>
with medial tooth apically truncate, about one-half the length of the lateral lobes.
<collectingMunicipality id="6BA4ACF1FFDDFFC13C956534FE8D720D" box="[194,323,1349,1373]" pageId="72" pageNumber="413">Submentum</collectingMunicipality>
with six setae anteriorly.
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rather short, not quite extended posteriorly to basal one-fourth of the elytra, only 2.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC65D7FDFA71ED" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">Pronotum. Distinctly transverse (ratio PW/PL = 1.34), with greatest width near anterior one-third, only slightly narrowed posteriorly; lateral margins rounded, slightly more so anteriorly, slightly straighten just anterior to basal angles, the latter obtuse and rounded. Disc smooth and convex, median longitudinal impression finely impressed, but continuous between anterior and posterior margins; basal foveae shallow, faintly impressed; median basal area reduced, faintly delimit- ed, slightly punctate in some specimens. Basal margin nearly rectilinear. Lateral pronotal borders moderately slender, regular, narrowly and regularly reflexed, lateral grooves narrow but distinctly impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
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28.
<taxonomicName id="4C7F4D08FFDCFFC03D5B6648FE33711C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[268,509,1593,1612]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="414" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDCFFC03D5B6648FE33711C" box="[268,509,1593,1612]" italics="true" pageId="73" pageNumber="414">Trechepaphiopsis uniporosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFDCFFC03E556648FD89711C" box="[514,583,1593,1612]" pageId="73" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFDCFFC0387C6622FB587135" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1067,1174,1619,1637]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="414" phylum="Arthropoda" rank="species" species="uniporosa">
<emphasis id="B90BEA99FFDCFFC0387C6622FB587135" box="[1067,1174,1619,1637]" italics="true" pageId="73" pageNumber="414">T. uniporosa</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFDFFFC33CFC60AAFC287683" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">Elytra. Ovoid, only slightly more narrowed anteriorly than posteriorly, humeri distinct but rounded. Disc convex, striae faintly impressed and finely punctate, the medial three or four striae distinctly impressed, the more lateral striae more or less effaced, but perceptible if only by the presence of fine punctures seen as rows of brown dots visible through the integument. Parascutellar striole present. Recurrent stria abruptly terminated anteriorly in presumed location of interval 6. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Only one discal setiferous pore present, inserted slightly anterior to middle next to stria 3. Preapical seta present on interval 3 near stria 3, closer to elytra apical margin that to sutural margin.</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC61ADFE9E7543" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC626AFB857503" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC622AFF7F7584" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFDFFFC33DA9622AFD907523" box="[510,606,603,627]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="74" pageNumber="415">Fig. 28b</figureCitation>
) with sagittal aileron very small, shaft moderately broad basally, expanded near mid-length, then gradually narrowed apically to a thin, bluntly round- ed apex, the latter slightly recurved dorsally; endophallus with a small scaly sclerotized area (
<figureCitation id="13442A0EFFDFFFC3383D62EAFF6B7584" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="74" pageNumber="415">Fig. 28c</figureCitation>
).
</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC62ADFE7A7424" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">
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</emphasis>
.— Members of this species have been found in mixed broadleaf evergreen/ deciduous forest at elevations ranging from
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and collected by sifting leaf litter on the forest floor (
<figureCitation id="13442A0EFFDFFFC33D2C636DFE147464" box="[379,474,796,820]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="74" pageNumber="415">Fig. 37a</figureCitation>
). Specimens of
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<emphasis id="B90BEA99FFDFFFC33EDA636DFC677463" box="[653,937,796,820]" italics="true" pageId="74" pageNumber="415">Trechepaphiosis unisetosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFDFFFC33FB4636DFF297404" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="415" phylum="Arthropoda" rank="species" species="gaoligong" status="sp. nov.">
<emphasis id="B90BEA99FFDFFFC33FB4636DFF297404" italics="true" pageId="74" pageNumber="415">Trechepaphiama gaoligong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFDFFFC33CB8634DFEF07403" box="[239,318,828,851]" pageId="74" pageNumber="415" rank="species">sp. nov</taxonomicNameLabel>
were collected in the same sifted leaf litter samples and are therefore syntopic with
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<emphasis id="B90BEA99FFDFFFC33CE6632DFEF97423" box="[177,311,860,883]" italics="true" pageId="74" pageNumber="415">T. uniporosa</emphasis>
</taxonomicName>
at this site.
</paragraph>
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<materialsCitation id="3B173CD6FFDFFFC33F4F630AFE257484" location="We" pageId="74" pageNumber="415" specimenCount="12" specimenCount-female="2" specimenCount-male="4" stateProvince="Shan">
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.—
<figureCitation id="13442A0EFFDFFFC33F28630DFC1174C4" box="[895,991,892,916]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="74" pageNumber="415">Fig. 28d</figureCitation>
.
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examined a total of
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(
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and
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) from the
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locality on the western slope of the southcentral part of the Gaoligong
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in
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(see
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material above for exact collection data). This locality is in
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4
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.
</paragraph>
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.— This species currently is known only from western Lushui County in the southcentral part of the Gaoligong Shan region, western
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,
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.
</paragraph>
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</treatment>
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