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<mods:title id="E1551BD074F1EC40346354D1B1C88C8B">The Genus Grasshoffia with the Description of a New Deep-water Species from the Northern Philippines (Octocorallia: Pennatulacea: Virgulariidae)</mods:title>
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<mods:namePart id="944019D798CF2D32B3A9721C30DC40D7">Williams, Gary C.</mods:namePart>
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<mods:date id="F634A69BF73C3CCAE091115CD56D07F3">2017</mods:date>
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<taxonomicName id="4C354D327302FFA3FF8FFA6FC0CFFA4C" ID-CoL="3HBCV" authority="Williams" authorityName="Williams" authorityYear="2017" box="[123,490,1510,1534]" class="Anthozoa" family="Virgulariidae" genus="Grasshoffia" kingdom="Animalia" order="Pennatulacea" pageId="2" pageNumber="97" phylum="Cnidaria" rank="species" species="profundica" status="sp. nov.">
<emphasis id="B941EAA37302FFA3FF8FFA6FC058FA4C" bold="true" box="[123,381,1510,1534]" italics="true" pageId="2" pageNumber="97">Grasshoffia profundica</emphasis>
Williams
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,
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<figureCitation id="130E2A347302FFA3FF8FF98EC1CAF9AE" box="[123,239,1543,1565]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStart-4="FIGURE" captionStartId-0="6.[171,183,1548,1567]" captionStartId-1="7.[171,183,1569,1588]" captionStartId-2="8.[171,183,1669,1688]" captionStartId-3="9.[171,183,1671,1690]" captionStartId-4="10.[171,183,1654,1673]" captionTargetBox-0="[120,1176,216,1530]" captionTargetBox-1="[125,1170,220,1546]" captionTargetBox-2="[167,1130,216,1650]" captionTargetBox-3="[129,1167,216,1651]" captionTargetBox-4="[120,1176,216,1637]" captionTargetId-0="figure-60@6.[120,1176,216,1530]" captionTargetId-1="figure-67@7.[120,1176,216,1551]" captionTargetId-2="figure-60@8.[167,1130,216,1650]" captionTargetId-3="figure-67@9.[129,1167,216,1651]" captionTargetId-4="figure-60@10.[120,1176,216,1637]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIGURE 3. Grasshoffia profundica n. sp. External morphology. A. Holotype (CAS 207514). B. Paratype (CAS 207515); scale bar for A and B = 40 mm. C. Detail of holotype rachis. D. Detail of holotype rachis. E. Detail of paratype rachis; scale bar for C-E = 10 mm. F. Detail of rachis of paratype showing broadly ovate polyp leaves with narrow proximal region attached to the rachis; scale bar = 5.0 mm. G. Diagram of a single autozooid, pinnules not shown; scale bar = 0.5 mm." captionText-1="FIGURE 4. Grasshoffia profundica n. sp. Scanning electron micrographs of axial morphology. A. Transverse section of axis from the distal region of the rachis showing arrangement of wedge-shaped sections of calcareous material radiating outward from the central core; scale bar = 0.3 mm. B. Enlarged detail of the central region from Fig. 2A showing concentration of organic matter in the central core and several elongated, tadpole-shaped channels surrounding the core; scale bar = 0.2 mm. C. Longitudinal external view of portion of axis from the distal region of the rachis showing uniformly smooth surface; scale bar = 0.5 mm. D. Enlarged detail of axial surface (from Fig. 2C) showing short, slit-shaped orifices; scale bar = 0.1 mm." captionText-2="FIGURE 5. Grasshoffia profundica n. sp. Scanning electron micrographs of sclerites from polyps and polyp leaves. Scale bar = 0.03 mm." captionText-3="FIGURE 6. Grasshoffia profundica n. sp. Scanning electron micrographs of sclerites from polyps and polyp leaves. Scale bar = 0.03 mm." captionText-4="FIGURE 7. Grasshoffia profundica n. sp. Scanning electron micrographs of polyp sclerites from polyps and polyp leaves. Scale bars = 0.02 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13155410" figureDoi-1="http://doi.org/10.5281/zenodo.13155412" figureDoi-2="http://doi.org/10.5281/zenodo.13155414" figureDoi-3="http://doi.org/10.5281/zenodo.13155416" figureDoi-4="http://doi.org/10.5281/zenodo.13155418" httpUri-0="https://zenodo.org/record/13155410/files/figure.png" httpUri-1="https://zenodo.org/record/13155412/files/figure.png" httpUri-2="https://zenodo.org/record/13155414/files/figure.png" httpUri-3="https://zenodo.org/record/13155416/files/figure.png" httpUri-4="https://zenodo.org/record/13155418/files/figure.png" pageId="2" pageNumber="97">Figures 3-7.</figureCitation>
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.—
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.
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.—
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.
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<collectionCode id="ED24AE747302FFA3FF05F9F9C045F934" box="[241,352,1648,1672]" pageId="2" pageNumber="97">LOCALITY</collectionCode>
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(Fig. 1).—
<location id="8EEA606A7302FFA3FE18F9F9C35CF93A" LSID="urn:lsid:plazi:treatment:039C87A77302FFA2FF8FFA6FC57AFE67:8EEA606A7302FFA3FE18F9F9C35CF93A" box="[492,633,1648,1672]" country="Philippines" latitude="13.764167" longLatPrecision="1" longitude="120.85167" municipality="southwestern Luzon" name="Balayan Bay" pageId="2" pageNumber="97">Balayan Bay</location>
,
<collectingMunicipality id="6BEEACCB7302FFA3FD79F9F9C24EF935" box="[653,875,1648,1672]" pageId="2" pageNumber="97">southwestern Luzon</collectingMunicipality>
,
<collectingCountry id="F32276217302FFA3FC8BF9F9C2D2F93A" box="[895,1015,1648,1672]" name="Philippines" pageId="2" pageNumber="97">Philippines</collectingCountry>
(
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,
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to
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,
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); station number HEPD-010;
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depth
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; 30
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</paragraph>
<paragraph id="8B8A36B17303FFA2FF8FFF52C388FE81" blockId="3.[123,1174,219,470]" pageId="3" pageNumber="98">
May 2011; G.C. Williams on board the research ship of the Philippine Bureau of Fisheries and Aquatic Resources,
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M/
<collectionCode id="ED24AE747303FFA2FE85FF75C0A4FEA1" box="[369,385,252,275]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="98" type="Museum">V</collectionCode>
DA-BFAR
</emphasis>
, during the 2011
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Hearst Philippine Biodiversity Expedition; two partial specimens collected by beam trawl.
</paragraph>
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<emphasis id="B941EAA37303FFA2FEB1FEB5C0F8FEE1" bold="true" box="[325,477,316,340]" pageId="3" pageNumber="98">DISTRIBUTION</emphasis>
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.— Collected from muddy sea bottom in a north to south linear transect in the eastern part of Balayan Bay (Fig. 1), which represents the only known occurrence of the species.
</paragraph>
</subSubSection>
<subSubSection id="C32F653A7303FFA2FF5FFE17C57AFE67" pageId="3" pageNumber="98" type="etymology">
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<emphasis id="B941EAA37303FFA2FF5FFE17C010FE06" bold="true" box="[171,309,414,438]" pageId="3" pageNumber="98">ETYMOLOGY</emphasis>
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.— The specific epithet is derived from the Latin
<emphasis id="B941EAA37303FFA2FC86FE14C2F8FE07" box="[882,989,413,437]" italics="true" pageId="3" pageNumber="98">profundus</emphasis>
(deep or of the depths), and the suffix
<emphasis id="B941EAA37303FFA2FE86FE36C08FFE64" box="[370,426,447,470]" italics="true" pageId="3" pageNumber="98">icus</emphasis>
(belonging to), referring to the deep-sea habitat of the
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.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1B50F970415A8A2359F1D69B4A656191">The Genus Grasshoffia with the Description of a New Deep-water Species from the Northern Philippines (Octocorallia: Pennatulacea: Virgulariidae)</mods:title>
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,
<figureCitation id="13442A0EFF83FF9F3C8B663BFED47130" box="[220,282,1610,1632]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="22" pageNumber="363">4648</figureCitation>
)
</paragraph>
</subSubSection>
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<treatmentCitation id="0ADE109AFF83FF9F3C2C661CFE1871D2" author="UENO, S. I." box="[123,470,1645,1666]" page="182" pageId="22" pageNumber="363" year="1997">
<taxonomicName id="4C7F4D08FF83FF9F3C2C661CFE1871D2" ID-CoL="8LNL7" authority="Ueno, 1997: 182" authorityName="Ueno" authorityPageNumber="182" authorityYear="1997" box="[123,470,1645,1666]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90BEA99FF83FF9F3C2C661CFEFF71D2" box="[123,305,1645,1666]" italics="true" pageId="22" pageNumber="363">Stevensius minutus</emphasis>
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<bibRefCitation id="EFEE4B7AFF83FF9F3D6F661CFE1871D2" author="UENO, S. I." box="[312,470,1645,1666]" pageId="22" pageNumber="363" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno, 1997:182</bibRefCitation>
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</taxonomicName>
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.
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, a female, in NMST. Type locality:
<collectingCountry id="F368761BFF83FF9F3FC2661CFC1F71D3" box="[917,977,1645,1667]" name="China" pageId="22" pageNumber="363">China</collectingCountry>
,
<collectingRegion id="49BBF869FF83FF9F3F8D661CFBE871D2" box="[986,1062,1645,1666]" country="China" name="Yunnan" pageId="22" pageNumber="363">Yunnan</collectingRegion>
,
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</paragraph>
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<materialsCitation id="3B173CD6FF83FF9F3CFC66FAFDA771EC" county="Tengchong County" elevation="2430" location="Dabei" pageId="22" pageNumber="363" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF83FF9F3CFC66FAFF1371F1" box="[171,221,1675,1697]" country="Myanmar" name="Shan" pageId="22" pageNumber="363">Shan</collectingRegion>
,
<collectingCounty id="62A14E07FF83FF9F3CB066FAFE6E71F0" box="[231,416,1675,1697]" pageId="22" pageNumber="363">Tengchong County</collectingCounty>
,
<location id="8EA06050FF83FF9F3DFE66FAFE2B71F1" LSID="urn:lsid:plazi:treatment:03D6879DFF83FF913C2C665BFBE0758A:8EA06050FF83FF9F3DFE66FAFE2B71F1" box="[425,485,1675,1697]" county="Tengchong County" name="Dabei" pageId="22" pageNumber="363" stateProvince="Shan">Dabei</location>
,
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<elevation id="0052D1B8FF83FF9F3DB866FAFDF271F0" box="[495,572,1675,1696]" metricMagnitude="3" metricUnit="m" metricValue="2.43" pageId="22" pageNumber="363" unit="m" value="2430.0">2430 m</elevation>
</quantity>
.
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<emphasis id="B90BEA99FF83FF9F3C2C66D9FE9671ED" box="[123,344,1704,1725]" italics="true" pageId="22" pageNumber="363">Minutotrechus minutus</emphasis>
(Uéno)
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<emphasis id="B90BEA99FF83FF9F3D8866D9FDA771EC" bold="true" box="[479,617,1704,1726]" pageId="22" pageNumber="363">COMBINATION</emphasis>
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.
</paragraph>
</subSubSection>
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11.
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<emphasis id="B90BEA99FF82FF9E3D426653FE107164" box="[277,478,1570,1589]" italics="true" pageId="23" pageNumber="364">Minutotrechus minutus</emphasis>
(Uéno)
</taxonomicName>
; a. Dorsal habitus (paratype). scale line = 1.0 mm; b. Map of locality records (red circle) for
<taxonomicName id="4C7F4D08FF82FF9E3D12664DFE68711E" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Ueno" baseAuthorityYear="1997" box="[325,422,1596,1614]" class="Insecta" family="Carabidae" genus="Minutotrechus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="364" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90BEA99FF82FF9E3D12664DFE68711E" box="[325,422,1596,1614]" italics="true" pageId="23" pageNumber="364">M. minutus</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a = 0.5mm, b = 100 km.
</paragraph>
</caption>
<subSubSection id="C3656500FF8DFF913CFC60AAFC9A7664" pageId="24" pageNumber="365" type="description">
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.— We have not had an opportunity to study the
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of this species, but we have examined a
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female deposited in IOZ. Features noted below are based on our examination of that
<typeStatus id="54C48829FF8DFF913DF7616DFE327664" box="[416,508,284,308]" pageId="24" pageNumber="365" type="paratype">paratype</typeStatus>
and Uénos orginal description.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8DFF913CFC614DFBE0758A" pageId="24" pageNumber="365" type="diagnosis">
<paragraph id="8BC0368BFF8DFF913CFC614DFE0D76C5" blockId="24.[123,1174,219,730]" pageId="24" pageNumber="365">
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<emphasis id="B90BEA99FF8DFF913CFC614DFEEE7603" bold="true" box="[171,288,316,340]" pageId="24" pageNumber="365">DIAGNOSIS</emphasis>
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.— Adults of this species (
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), the only known species in this new genus, can be distinguished from those of all other species in the region by the combination of character states noted in the generic diagnosis.
</paragraph>
<paragraph id="8BC0368BFF8DFF913CFC61EFFDA77547" blockId="24.[123,1174,219,730]" pageId="24" pageNumber="365">
<emphasis id="B90BEA99FF8DFF913CFC61EFFE6B76E4" bold="true" box="[171,421,414,438]" pageId="24" pageNumber="365">
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<smallCapsWord id="8D26A057FF8DFF913D4661D0FE6B76E4" baselines="431" box="[273,421,417,436]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="24" pageNumber="365">DISTRIBUTION</smallCapsWord>
</emphasis>
.— According to
<bibRefCitation id="EFEE4B7AFF8DFF913E3461EFFD2376E6" author="UENO, S. I." box="[611,749,414,438]" pageId="24" pageNumber="365" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno (1997)</bibRefCitation>
specimens of the
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series were collected at an elevation of
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in a dense
<taxonomicName id="4C7F4D08FF8DFF913E0261CFFD3D7686" box="[597,755,446,470]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="24" pageNumber="365" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FF8DFF913E0261CFFD3D7686" box="[597,755,446,470]" italics="true" pageId="24" pageNumber="365">Rhododendron</emphasis>
</taxonomicName>
forest by sifting moist leaf litter accumulations on the ground. He also noted that many specimens of “
<taxonomicName id="4C7F4D08FF8DFF913F1461AEFBF376A7" authority="Ueno" authorityName="Ueno" authorityYear="1997" box="[835,1085,479,503]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="species" species="asetosus">
<emphasis id="B90BEA99FF8DFF913F1461AEFC3676A6" box="[835,1016,479,503]" italics="true" pageId="24" pageNumber="365">Trechus asetosus</emphasis>
Uéno
</taxonomicName>
” (1997) were also collected in the same litter samples.
</paragraph>
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<emphasis id="B90BEA99FF8DFF913CFC626EFD4C7566" bold="true" box="[171,642,543,567]" pageId="24" pageNumber="365">
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</emphasis>
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<emphasis id="B90BEA99FF8DFF913EDA626EFCDC7566" bold="true" box="[653,786,543,567]" pageId="24" pageNumber="365">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FF8DFF913F4A626EFC397528" county="Tengchong County" location="This" pageId="24" pageNumber="365" specimenCount="6" specimenCount-female="6" stateProvince="Shan">
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<collectingRegion id="49BBF869FF8DFF913F4A626EFC967566" box="[797,856,543,567]" country="Myanmar" name="Shan" pageId="24" pageNumber="365">SHAN</collectingRegion>
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.—
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.
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species is known only from the
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specimens of the
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series collected at the
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locality, in
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, high on the western slope of the southern part of the Gaoligong
</materialsCitation>
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in
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6
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.
</paragraph>
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</emphasis>
.—
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This species currently is known only from the
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locality in the southern part of the
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, in western
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,
<collectingCountry id="F368761BFF8DFF913FB062B3FBE4758A" box="[999,1066,706,730]" name="China" pageId="24" pageNumber="365">China</collectingCountry>
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.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,204 @@
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<mods:title id="E834973F5D7B24EB5536C3582552FDD5">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="580C83F1638387EF3FFDB8073EC8E4C9">Deuve, Thierry</mods:namePart>
<mods:affiliation id="185E21496B01434AF3701256F9B18FAD">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="95B9F20806F567F31C1CFEDC9EBAAE9B">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="4DDAF67468502848EF356356D5875FB9">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="3F8072EDD1EBE29563C4305448A544C8">Liang, Hongbin</mods:namePart>
<mods:affiliation id="E1BB77C981AEEAD92F2D9D5496611D5B">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FF83FF9F3D1861FAFC0F76F3" bold="true" box="[335,961,395,419]" pageId="22" pageNumber="363">
Genus
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<emphasis id="B90BEA99FF83FF9F3DC861FAFD8C76F3" bold="true" box="[415,578,395,419]" italics="true" pageId="22" pageNumber="363">Minutotrechus</emphasis>
Deuve
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and Kavanaugh,
<taxonomicNameLabel id="A23857E2FF83FF9F3F0B61FAFC0F76F3" box="[860,961,395,419]" pageId="22" pageNumber="363" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF83FF9F3CFC61C2FDE2757C" pageId="22" pageNumber="363" type="type_taxon">
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.—
<taxonomicName id="4C7F4D08FF83FF9F3D3F61C5FD75769B" ID-CoL="8LNL7" authority="Ueno, 1997" authorityName="Ueno" authorityYear="1997" box="[360,699,435,459]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90BEA99FF83FF9F3D3F61C5FDFC769B" box="[360,562,436,459]" italics="true" pageId="22" pageNumber="363">Stevensius minutus</emphasis>
<bibRefCitation id="EFEE4B7AFF83FF9F3E6D61C2FD75769B" author="UENO, S. I." box="[570,699,435,459]" pageId="22" pageNumber="363" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno, 1997</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0368BFF83FF9F3CFC61A2FDE2757C" blockId="22.[123,1174,435,1549]" pageId="22" pageNumber="363">
<emphasis id="B90BEA99FF83FF9F3CFC61A2FDFC76BA" bold="true" box="[171,562,467,491]" pageId="22" pageNumber="363">
<smallCapsWord id="8D26A057FF83FF9F3CFC61A2FEE076BA" baselines="485,485" box="[171,302,467,491]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="22" pageNumber="363">DERIVATION</smallCapsWord>
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<smallCapsWord id="8D26A057FF83FF9F3D0E61A6FE5076BA" baselines="485" box="[345,414,471,490]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="genus" pageId="22" pageNumber="363">GENUS</smallCapsWord>
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.— The genus group name (masculine) is a combination of the Latin adjective,
<taxonomicName id="4C7F4D08FF83FF9F3D3C6185FE71755B" ID-CoL="8LNL7" authorityName="Ueno" authorityYear="1997" box="[363,447,500,523]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90BEA99FF83FF9F3D3C6185FE71755B" box="[363,447,500,523]" italics="true" pageId="22" pageNumber="363">minutus</emphasis>
</taxonomicName>
, meaning very small, and the generic name,
<taxonomicName id="4C7F4D08FF83FF9F3FCC6182FC20755B" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[923,1006,499,523]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3FCC6182FC20755B" box="[923,1006,499,523]" italics="true" pageId="22" pageNumber="363">Trechus</emphasis>
</taxonomicName>
, in reference to the small size of members of this genus.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF83FF9F3CFC6245FBAE743C" pageId="22" pageNumber="363" type="diagnosis">
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<smallCapsWord id="8D26A057FF83FF9F3CFC6245FEEE751A" baselines="582,581" box="[171,288,564,588]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="22" pageNumber="363">
<emphasis id="B90BEA99FF83FF9F3CFC6245FEEE751A" bold="true" box="[171,288,564,588]" pageId="22" pageNumber="363">DIAGNOSIS</emphasis>
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.— Adults of this genus (
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) can be recognized by the following combination of character states: size small (BL =
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), apterous, body color brown to black; head large with small but protruding eyes, their diameter shorter than length of tempora; mandibles short, obtusely bidentate, mentum and submentum at least partial fused, labial suture partially perceptible, mentum with medial tooth truncate; pronotum small, cordate, narrow (ratio PW/PL = 1.25), very convex, globulose, glabrous, basal angles small and subrectangular, slightly obtuse, with basal margin broadly lobate, basal area convex, both midlateral and basolateral setae present; elytra ovoid and markedly convex, with discal striae 2 to 8 striae effaced, stria 1 deeply impressed and punctate, both anterior and middle discal setae present, preapical seta absent, lateral groove abruptly terminated anteriorly at humerus; legs short, protibiae without longitudinal furrows.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF83FF9F3CFC6305FEE1715D" pageId="22" pageNumber="363" type="discussion">
<paragraph id="8BC0368BFF83FF9F3CFC6305FB4F72FD" blockId="22.[123,1174,435,1549]" pageId="22" pageNumber="363">
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<emphasis id="B90BEA99FF83FF9F3CFC6305FEE674DA" bold="true" box="[171,296,884,908]" pageId="22" pageNumber="363">COMMENTS</emphasis>
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.— This new genus is known from only
<specimenCount id="9D79FD02FF83FF9F3E8F6305FC8974DC" box="[728,839,884,908]" count="6" pageId="22" pageNumber="363" type="female">six female</specimenCount>
specimens that were originally assigned by
<bibRefCitation id="EFEE4B7AFF83FF9F3D5163E5FE5C74FC" author="UENO, S. I." box="[262,402,916,940]" pageId="22" pageNumber="363" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno (1997)</bibRefCitation>
to genus
<taxonomicName id="4C7F4D08FF83FF9F3E5363E5FCD174FC" authority="Jeannel (1923)" authorityName="Jeannel" authorityYear="1923" box="[516,799,916,940]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3E5363E5FDBD74FB" box="[516,627,916,939]" italics="true" pageId="22" pageNumber="363">Stevensius</emphasis>
<bibRefCitation id="EFEE4B7AFF83FF9F3E2A63E5FCD174FC" author="JEANNEL, R." box="[637,799,916,940]" pageId="22" pageNumber="363" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel (1923)</bibRefCitation>
</taxonomicName>
of the eastern Himalayan region. However, they can be distinguished from members of that genus by their smaller head size, protibiae without longitudinal furrows, pronotum more cordate and with basal angles smaller and basal margin broadly lobate and elytra with the lateral groove abruptly terminated anteriorly at humerus.
<taxonomicName id="4C7F4D08FF83FF9F3C2C6465FED8737C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,278,1044,1068]" class="Insecta" family="Carabidae" genus="Minutotrechus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3C2C6465FED8737C" box="[123,278,1044,1068]" italics="true" pageId="22" pageNumber="363">Minutotrechus</emphasis>
</taxonomicName>
appears to be more closely related to
<taxonomicName id="4C7F4D08FF83FF9F3EE66465FC10737C" authority="Deuve (2005)" authorityName="Deuve" authorityYear="2005" box="[689,990,1044,1068]" class="Insecta" family="Carabidae" genus="Hubeitrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3EE66465FC8E737C" box="[689,832,1044,1068]" italics="true" pageId="22" pageNumber="363">Hubeitrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF83FF9F3F106464FC10737C" author="DEUVE, T." box="[839,990,1044,1068]" pageId="22" pageNumber="363" pagination="305 - 317" refId="ref58974" refString="DEUVE, T. 2005. Descriptions de nouveaux Trechini de la Chine centrale et du plateau Tibetain (Coleoptera, Trechidae). Coleopteres 11: 305 - 317." type="journal article" year="2005">Deuve (2005)</bibRefCitation>
</taxonomicName>
, but its members can be distinguished from those of the latter in having the mentum and submentum at least partial fused, the mandibular teeth short and obtuse, the pronotum with the median basal area more convex and basal margin broadly lobate and without margination, and lateral groove abruptly terminated at the humerus. Because no male specimens of
<taxonomicName id="4C7F4D08FF83FF9F3E9364E5FC9173FC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[708,863,1172,1196]" class="Insecta" family="Carabidae" genus="Minutotrechus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3E9364E5FC9173FC" box="[708,863,1172,1196]" italics="true" pageId="22" pageNumber="363">Minutotrechus</emphasis>
</taxonomicName>
have been collected to date, we do not know if male protarsomeres 1 and 2 are elongate as in males of
<taxonomicName id="4C7F4D08FF83FF9F3FF364C5FBFD739C" authorityName="Deuve" authorityYear="2005" box="[932,1075,1204,1228]" class="Insecta" family="Carabidae" genus="Hubeitrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3FF364C5FBFD739C" box="[932,1075,1204,1228]" italics="true" pageId="22" pageNumber="363">Hubeitrechus</emphasis>
</taxonomicName>
or broad as in
<taxonomicName id="4C7F4D08FF83FF9F3CE164A4FEEB73BC" authorityName="Jeannel" authorityYear="1923" box="[182,293,1237,1260]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3CE164A4FEEB73BC" box="[182,293,1237,1260]" italics="true" pageId="22" pageNumber="363">Stevensius</emphasis>
</taxonomicName>
males. Members of this new genus can also be compared with those of
<taxonomicName id="4C7F4D08FF83FF9F386164A4FE7E725C" authority="Belousov and Kabak (2016)" authorityName="Belousov and Kabak" authorityYear="2016" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F386164A4FB5B73BC" box="[1078,1173,1237,1260]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
Belousov and Kabak (2016)
</taxonomicName>
, from which they differ in having the right mandible obtusely bifid (tridentate in
<taxonomicName id="4C7F4D08FF83FF9F3D5C6564FEA4727C" authorityName="Belousov and Kabak" authorityYear="2016" box="[267,362,1301,1324]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3D5C6564FEA4727C" box="[267,362,1301,1324]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
</taxonomicName>
members), the protibiae without longitudinal furrows (longitudinal furrows present in
<taxonomicName id="4C7F4D08FF83FF9F3CA66544FE9E721C" authorityName="Belousov and Kabak" authorityYear="2016" box="[241,336,1333,1356]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3CA66544FE9E721C" box="[241,336,1333,1356]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
</taxonomicName>
members), the mentum and submentum at least partial fused (not fused in
<taxonomicName id="4C7F4D08FF83FF9F3C2C6524FF14723C" authorityName="Belousov and Kabak" authorityYear="2016" box="[123,218,1365,1388]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3C2C6524FF14723C" box="[123,218,1365,1388]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
</taxonomicName>
members), elytra with only two discal setae (three or more discal setae present in
<taxonomicName id="4C7F4D08FF83FF9F3C2C6504FF1472DC" authorityName="Belousov and Kabak" authorityYear="2016" box="[123,218,1397,1420]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3C2C6504FF1472DC" box="[123,218,1397,1420]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
</taxonomicName>
members), the preapical seta absent (present in
<taxonomicName id="4C7F4D08FF83FF9F3E8F6504FCF972DC" authorityName="Belousov and Kabak" authorityYear="2016" box="[728,823,1397,1420]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3E8F6504FCF972DC" box="[728,823,1397,1420]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
</taxonomicName>
members) and the lateral groove abruptly terminated at humerus (gradually narrowed anterior to humerus in
<taxonomicName id="4C7F4D08FF83FF9F3FFF65E4FBC972FC" authorityName="Belousov and Kabak" authorityYear="2016" box="[936,1031,1429,1452]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF83FF9F3FFF65E4FBC972FC" box="[936,1031,1429,1452]" italics="true" pageId="22" pageNumber="363">Uenoites</emphasis>
</taxonomicName>
members).
</paragraph>
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<emphasis id="B90BEA99FF83FF9F3CFC65C4FE3C729B" bold="true" box="[171,498,1461,1485]" pageId="22" pageNumber="363">
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</emphasis>
.—
<materialsCitation id="3B173CD6FF83FF9F3E4C65C4FEE4715D" country="China" location="Gaoligong Shan region" pageId="22" pageNumber="363" specimenCount="1" stateProvince="Yunnan">
This genus currently is known only from the
<typeStatus id="54C48829FF83FF9F385C65C4FBF7729D" box="[1035,1081,1461,1485]" pageId="22" pageNumber="363">type</typeStatus>
species, which is known only from the southern part of the
<location id="8EA06050FF83FF9F3EEA65A4FC7272BD" LSID="urn:lsid:plazi:treatment:03D6879DFF83FF9F3D1861FAFEE1715D:8EA06050FF83FF9F3EEA65A4FC7272BD" box="[701,956,1493,1517]" country="China" name="Gaoligong Shan region" pageId="22" pageNumber="363" stateProvince="Yunnan">Gaoligong Shan region</location>
of western
<collectingRegion id="49BBF869FF83FF9F381365A4FF13715D" country="China" name="Yunnan" pageId="22" pageNumber="363">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF83FF9F3CB06584FEE4715D" box="[231,298,1525,1549]" name="China" pageId="22" pageNumber="363">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,516 @@
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<mods:title id="E1C386A139A8D77D48C0E65A10A2D288">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="04DA6FB3C1FDE48C050D985EEEF68F7D">Deuve, Thierry</mods:namePart>
<mods:affiliation id="E51447FA0A141C32D50EB69E604A3A9F">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="FE9445BAE6B6C88B078591969D16FCE3">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="D474D037B8AB17B077C732C1027DBB80">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="41AF75B304C8954245E3E6DE836ED67C" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:name id="D81141F529E8ED8EB0B365D0FB831A9B" type="personal">
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<mods:namePart id="9C9EF6AC89D384FF7E2352ACACEC6B08">Liang, Hongbin</mods:namePart>
<mods:affiliation id="A78FDB9079B2994B8421CB41C4F66627">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FF87FF9B3C2C60AAFCDD77A3" bold="true" box="[123,787,219,243]" pageId="18" pageNumber="359">
<taxonomicName id="4C7F4D08FF87FF9B3C2C60AAFE3F77A3" ID-CoL="8MGQP" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,497,219,243]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF87FF9B3C2C60AAFE6D77A3" bold="true" box="[123,419,219,243]" italics="true" pageId="18" pageNumber="359">Agonotrechus xiaoheishan</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FF87FF9B3EEA60AAFCDD77A3" box="[701,787,219,243]" pageId="18" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3656500FF87FF9B3C2C608AFEA77641" box="[123,361,251,273]" pageId="18" pageNumber="359" type="description">
<paragraph id="8BC0368BFF87FF9B3C2C608AFEA77641" blockId="18.[123,787,219,273]" box="[123,361,251,273]" pageId="18" pageNumber="359">
(
<figureCitation id="13442A0EFF87FF9B3CD5608AFF097640" box="[130,199,251,273]" captionStart="FIGURE" captionStartId="19.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1578]" captionTargetId="figure-15@19.[120,1176,216,1579]" captionTargetPageId="19" captionText="FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A. xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="18" pageNumber="359">Figs. 9</figureCitation>
,
<figureCitation id="13442A0EFF87FF9B3C86608AFF387641" box="[209,246,251,273]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="18" pageNumber="359">16a</figureCitation>
,
<figureCitation id="13442A0EFF87FF9B3D57608AFED47640" box="[256,282,251,272]" captionStart="FIGURE" captionStartId="106.[171,183,895,914]" captionTargetBox="[120,1176,216,881]" captionTargetId="figure-12@106.[120,1176,215,882]" captionTargetPageId="106" captionText="FIGURE 44. Photograph of habitat for trechine species in the Gaoligong Shan region. Longling County, Longjiang Township, Xiaoheishan Forest Reserve, elevation 2020 m; habitat in which specimens of Agonotrechus xiaoheishan sp. nov. and Trechus indicus Putzeys were collected. Photo by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155374" httpUri="https://zenodo.org/record/13155374/files/figure.png" pageId="18" pageNumber="359">44</figureCitation>
,
<figureCitation id="13442A0EFF87FF9B3D72608AFEAD7641" box="[293,355,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="18" pageNumber="359">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF87FF9B3CFC6155FCB774ED" pageId="18" pageNumber="359" type="materials_examined">
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</emphasis>
.—
<materialsCitation id="3B173CD6FF87FF9B3DD46155FF2376EC" collectingDate="2005-05-28" collectorName="H. B. Liang &amp; H. M. Yan &amp; K. J. Gao" country="China" county="Longling County" elevation="2067" latitude="24.83671" location="Longjiang Township" longLatPrecision="1" longitude="98.76185" pageId="18" pageNumber="359" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FF87FF9B3DD46155FE27766C" box="[387,489,292,316]" pageId="18" pageNumber="359" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1036866”/ “
<collectingCountry id="F368761BFF87FF9B3FAA6155FB98766C" box="[1021,1110,292,316]" name="China" pageId="18" pageNumber="359">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FF87FF9B3CEC6135FE4B760C" box="[187,389,324,348]" pageId="18" pageNumber="359">Longling County</collectingCounty>
,
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,
<taxonomicName id="4C7F4D08FF87FF9B3EF56135FBD5760C" authority="Forest Reserve, N" authorityName="Forest Reserve, N" authorityYear="2067" box="[674,1051,324,348]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
Xiaoheishan Forest Reserve,
<geoCoordinate id="EE4B504CFF87FF9B38506135FBD5760C" box="[1031,1051,324,348]" degrees="24.83671" direction="north" orientation="latitude" pageId="18" pageNumber="359" precision="1" value="24.83671">N</geoCoordinate>
</taxonomicName>
24.83671°/
<geoCoordinate id="EE4B504CFF87FF9B3C2C6115FEC5762C" box="[123,267,356,380]" degrees="098.76185" direction="east" orientation="longitude" pageId="18" pageNumber="359" precision="1" value="98.76185">E098.76185°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFF87FF9B3D666115FE48762B" box="[305,390,356,380]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="18" pageNumber="359" unit="m" value="2067.0">
<elevation id="0052D1B8FF87FF9B3D666115FE48762B" box="[305,390,356,380]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="18" pageNumber="359" unit="m" value="2067.0">2067 m</elevation>
</quantity>
,
<date id="FFC1104BFF87FF9B3DC66115FDEF762C" box="[401,545,356,380]" pageId="18" pageNumber="359" value="2005-05-28">
<collectingDate id="EF85E9A3FF87FF9B3DC66115FDEF762C" box="[401,545,356,380]" pageId="18" pageNumber="359" value="2005-05-28">28 May 2005</collectingDate>
</date>
, Stop# HBL-05-19,
<collectorName id="268A535DFF87FF9B3EAD6115FCBA762C" box="[762,884,356,380]" pageId="18" pageNumber="359">H.B. Liang</collectorName>
,
<collectorName id="268A535DFF87FF9B3F296115FC28762B" box="[894,998,356,379]" pageId="18" pageNumber="359">H.M. Yan</collectorName>
&amp;
<collectorName id="268A535DFF87FF9B385F6115FBAB762C" box="[1032,1125,356,380]" pageId="18" pageNumber="359">K.J. Gao</collectorName>
collectors”/ “
<typeStatus id="54C48829FF87FF9B3CB361F5FEBD76CC" box="[228,371,388,412]" pageId="18" pageNumber="359" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FF87FF9B3D2E61F5FD1176CB" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[377,735,388,412]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF87FF9B3D2E61F5FD5D76CC" box="[377,659,388,412]" italics="true" pageId="18" pageNumber="359">Agonotrechus xiaoheishan</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FF87FF9B3FD261F5FC1976CB" box="[901,983,388,411]" pageId="18" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FF87FF9B3CAF61D5FCEC754C" collectionCode="CAS, IOZ, MNHN" country="China" pageId="18" pageNumber="359" specimenCount="9" specimenCount-female="7" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FF87FF9B3CAF61D5FEAE76EC" box="[248,352,420,444]" pageId="18" pageNumber="359" type="paratype">Paratypes</typeStatus>
(a total of 13):
<specimenCount id="9D79FD02FF87FF9B3E5B61D5FD9176EC" box="[524,607,420,444]" count="2" pageId="18" pageNumber="359" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FF87FF9B3EC161D5FD3076EC" box="[662,766,420,444]" count="7" pageId="18" pageNumber="359" type="female">7 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF87FF9B3F7C61D5FCAF76EC" box="[811,865,420,444]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="359" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF87FF9B3F3A61D5FC5276EC" box="[877,924,420,444]" pageId="18" pageNumber="359">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF87FF9B3FFF61D5FC3576EC" box="[936,1019,420,444]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="359">MNHN</collectionCode>
) labeled same as
<typeStatus id="54C48829FF87FF9B3CCD61B5FF39768C" box="[154,247,452,476]" pageId="18" pageNumber="359" type="holotype">holotype</typeStatus>
except first label “CASENT 1036874”, “CASENT 1036875”, “CASENT 1036867”, “CASENT 1036868”, “CASENT 1036869”, “CASENT 1036870”, “CASENT 1036871”, “CASENT 1036872” and “CASENT 1036873”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FF87FF9B3F646275FEFA75CC" collectingDate="2005-05-28" collectionCode="CAS" collectorName="H. B. Liang &amp; D. Z. Dong &amp; J. L. Yang" country="China" county="Longling County" elevation="2020" latitude="24.82888" location="Longjiang Township" longLatPrecision="1" longitude="98.76001" pageId="18" pageNumber="359" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF87FF9B3F646275FC4F754C" box="[819,897,516,540]" count="1" pageId="18" pageNumber="359" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FF87FF9B3F976275FBED754C" box="[960,1059,516,540]" count="1" pageId="18" pageNumber="359" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF87FF9B380E6275FB40754C" box="[1113,1166,516,540]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="359" type="Museum">CAS</collectionCode>
) labeled “CASENT 1036863” and “CASENT 1036864”, respectively/ “
<collectingCountry id="F368761BFF87FF9B3F266255FC04756C" box="[881,970,548,572]" name="China" pageId="18" pageNumber="359">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF87FF9B3F836254FBE6756C" box="[980,1064,549,572]" country="China" name="Yunnan" pageId="18" pageNumber="359">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF87FF9B38636255FF05750C" pageId="18" pageNumber="359">Longling County</collectingCounty>
,
<location id="8EA06050FF87FF9B3C8F6235FE74750C" LSID="urn:lsid:plazi:treatment:03D6879DFF87FF9F3C2C60AAFBE07623:8EA06050FF87FF9B3C8F6235FE74750C" box="[216,442,580,604]" country="China" county="Longling County" latitude="24.82888" longLatPrecision="1" longitude="98.76001" name="Longjiang Township" pageId="18" pageNumber="359" stateProvince="Yunnan">Longjiang Township</location>
,
<taxonomicName id="4C7F4D08FF87FF9B3D906235FC10750C" authority="Forest Reserve, Guchengshan, 2020" authorityName="Forest Reserve, Guchengshan" authorityYear="2020" box="[455,990,580,604]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
Xiaoheishan Forest Reserve, Guchengshan,
<quantity id="4C879B6EFF87FF9B3FFE6235FC10750C" box="[937,990,580,604]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="18" pageNumber="359" unit="m" value="2020.0">
<elevation id="0052D1B8FF87FF9B3FFE6235FC10750C" box="[937,990,580,604]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="18" pageNumber="359" unit="m" value="2020.0">2020</elevation>
</quantity>
</taxonomicName>
m,
<geoCoordinate id="EE4B504CFF87FF9B385B6235FB43750C" box="[1036,1165,580,604]" degrees="24.82888" direction="north" orientation="latitude" pageId="18" pageNumber="359" precision="1" value="24.82888">N24.82888°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF87FF9B3C2C6215FEC4752C" box="[123,266,612,636]" degrees="098.76001" direction="east" orientation="longitude" pageId="18" pageNumber="359" precision="1" value="98.76001">E098.76001°</geoCoordinate>
,”/ “
<date id="FFC1104BFF87FF9B3D666215FE0F752C" box="[305,449,612,637]" pageId="18" pageNumber="359" value="2005-05-28">
<collectingDate id="EF85E9A3FF87FF9B3D666215FE0F752C" box="[305,449,612,637]" pageId="18" pageNumber="359" value="2005-05-28">28 May 2005</collectingDate>
</date>
, Stop# 2005-033B, D.H.
<taxonomicName id="4C7F4D08FF87FF9B3E846215FC81752C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[723,847,612,636]" pageId="18" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFF87FF9B3F0E6214FC1C752C" box="[857,978,612,636]" pageId="18" pageNumber="359">H.B. Liang</collectorName>
,
<collectorName id="268A535DFF87FF9B3F8C6214FB83752D" box="[987,1101,613,637]" pageId="18" pageNumber="359">D.Z. Dong</collectorName>
&amp;
<collectorName id="268A535DFF87FF9B383A6214FF7C75CD" pageId="18" pageNumber="359">J.L. Yang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF87FF9B3D1462F5FD78746C" collectingDate="2005-05-28" collectionCode="CAS" collectorName="C. E. Griswold &amp; H. B. Liang &amp; D. Z. Dong &amp; H. M. Yan &amp; K. J. Guo" country="China" county="Longling County" elevation="2020" latitude="24.82888" location="Longjiang Township" longLatPrecision="1" longitude="98.76001" pageId="18" pageNumber="359" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF87FF9B3D1462F5FE6175CC" box="[323,431,644,668]" count="2" pageId="18" pageNumber="359" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF87FF9B3DB462F5FDD675CC" box="[483,536,644,668]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="359" type="Museum">CAS</collectionCode>
) labeled “CASENT 1031915” and “CASENT 1031916”, respectively/ “
<collectingCountry id="F368761BFF87FF9B3D4B62D5FEBB75EC" box="[284,373,676,700]" name="China" pageId="18" pageNumber="359">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF87FF9B3DDF62D4FE1275EC" box="[392,476,677,700]" country="China" name="Yunnan" pageId="18" pageNumber="359">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF87FF9B3DB862D5FD7E75EC" box="[495,688,676,700]" pageId="18" pageNumber="359">Longling County</collectingCounty>
,
<location id="8EA06050FF87FF9B3E9462D5FC6575EC" LSID="urn:lsid:plazi:treatment:03D6879DFF87FF9F3C2C60AAFBE07623:8EA06050FF87FF9B3E9462D5FC6575EC" box="[707,939,676,700]" country="China" county="Longling County" latitude="24.82888" longLatPrecision="1" longitude="98.76001" name="Longjiang Township" pageId="18" pageNumber="359" stateProvince="Yunnan">Longjiang Township</location>
,
<taxonomicName id="4C7F4D08FF87FF9B3FE962D5FE7A758C" authority="Forest Reserve, Guchengshan, 2020" authorityName="Forest Reserve, Guchengshan" authorityYear="2020" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
Xiaoheishan Forest Reserve, Guchengshan,
<quantity id="4C879B6EFF87FF9B3D2862B5FE7A758C" box="[383,436,708,732]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="18" pageNumber="359" unit="m" value="2020.0">
<elevation id="0052D1B8FF87FF9B3D2862B5FE7A758C" box="[383,436,708,732]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="18" pageNumber="359" unit="m" value="2020.0">2020</elevation>
</quantity>
</taxonomicName>
m,
<geoCoordinate id="EE4B504CFF87FF9B3D8862B5FDAE758C" box="[479,608,708,732]" degrees="24.82888" direction="north" orientation="latitude" pageId="18" pageNumber="359" precision="1" value="24.82888">N24.82888°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF87FF9B3E2762B5FCCE758C" box="[624,768,708,732]" degrees="098.76001" direction="east" orientation="longitude" pageId="18" pageNumber="359" precision="1" value="98.76001">E098.76001°</geoCoordinate>
,”/ “
<date id="FFC1104BFF87FF9B3F7F62B5FC73758C" box="[808,957,708,733]" pageId="18" pageNumber="359" value="2005-05-28">
<collectingDate id="EF85E9A3FF87FF9B3F7F62B5FC73758C" box="[808,957,708,733]" pageId="18" pageNumber="359" value="2005-05-28">28 May 2005</collectingDate>
</date>
, Stop# 2005-033C, D.H.
<taxonomicName id="4C7F4D08FF87FF9B3CE06295FEFD75AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[183,307,740,764]" pageId="18" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFF87FF9B3D176295FE1075AC" box="[320,478,740,764]" pageId="18" pageNumber="359">C.E. Griswold</collectorName>
,
<collectorName id="268A535DFF87FF9B3DBC6294FDA975AC" box="[491,615,740,764]" pageId="18" pageNumber="359">H.B. Liang</collectorName>
,
<collectorName id="268A535DFF87FF9B3E246294FD2575AD" box="[627,747,741,765]" pageId="18" pageNumber="359">D.Z. Dong</collectorName>
,
<collectorName id="268A535DFF87FF9B3EAF6294FCAC75AC" box="[760,866,741,764]" pageId="18" pageNumber="359">H.M. Yan</collectorName>
&amp;
<collectorName id="268A535DFF87FF9B3FD06294FC2775AC" box="[903,1001,740,764]" pageId="18" pageNumber="359">K.J. Guo</collectorName>
collectors”. All
<typeStatus id="54C48829FF87FF9B3C2C6374FF2F744D" box="[123,225,773,797]" pageId="18" pageNumber="359" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FF87FF9B3E356375FD27744C" box="[610,745,772,796]" pageId="18" pageNumber="359" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FF87FF9B3EAE6375FBBC744C" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[761,1138,772,796]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF87FF9B3EAE6375FBD2744C" box="[761,1052,772,796]" italics="true" pageId="18" pageNumber="359">Agonotrechus xiaoheishan</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FF87FF9B3D556354FE98746C" box="[258,342,805,828]" pageId="18" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF87FF9B3CFC6335FD4E742C" blockId="18.[123,1174,292,1725]" pageId="18" pageNumber="359">
<materialsCitation id="3B173CD6FF87FF9B3CFC6335FDB2742C" country="China" county="Longling County" elevation="2067" latitude="24.83671" location="Longjiang Township" longLatPrecision="1" longitude="98.76185" pageId="18" pageNumber="359" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FF87FF9B3CFC6335FE99740B" bold="true" box="[171,343,836,860]" pageId="18" pageNumber="359">
<smallCapsWord id="8D26A057FF87FF9B3CFC6335FF2A740B" baselines="854,854" box="[171,228,836,860]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="18" pageNumber="359">
<typeStatus id="54C48829FF87FF9B3CFC6335FF2A740B" box="[171,228,836,860]" pageId="18" pageNumber="359">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FF87FF9B3CBB6339FE99740B" baselines="854" box="[236,343,840,859]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="18" pageNumber="359">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFF87FF9B3D286335FE0F740C" box="[383,449,836,860]" name="China" pageId="18" pageNumber="359">China</collectingCountry>
,
<collectingRegion id="49BBF869FF87FF9B3D9A6334FDEF740C" box="[461,545,837,860]" country="China" name="Yunnan" pageId="18" pageNumber="359">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF87FF9B3E7B6335FD2B740D" box="[556,741,836,861]" pageId="18" pageNumber="359">Longling County</collectingCounty>
,
<location id="8EA06050FF87FF9B3EA76335FC1E740C" LSID="urn:lsid:plazi:treatment:03D6879DFF87FF9F3C2C60AAFBE07623:8EA06050FF87FF9B3EA76335FC1E740C" box="[752,976,836,860]" country="China" county="Longling County" latitude="24.83671" longLatPrecision="1" longitude="98.76185" name="Longjiang Township" pageId="18" pageNumber="359" stateProvince="Yunnan">Longjiang Township</location>
,
<taxonomicName id="4C7F4D08FF87FF9B3F8C6335FED6742C" authority="Forest Reserve, N" authorityName="Forest Reserve, N" authorityYear="2067" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
Xiaoheishan Forest Reserve,
<geoCoordinate id="EE4B504CFF87FF9B3D536315FED6742C" box="[260,280,868,892]" degrees="24.83671" direction="north" orientation="latitude" pageId="18" pageNumber="359" precision="1" value="24.83671">N</geoCoordinate>
</taxonomicName>
24.83671°/
<geoCoordinate id="EE4B504CFF87FF9B3DD96315FDD2742C" box="[398,540,868,892]" degrees="098.76185" direction="east" orientation="longitude" pageId="18" pageNumber="359" precision="1" value="98.76185">E098.76185°</geoCoordinate>
,
<quantity id="4C879B6EFF87FF9B3E706315FDB2742C" box="[551,636,868,892]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="18" pageNumber="359" unit="m" value="2067.0">
<elevation id="0052D1B8FF87FF9B3E706315FDB2742C" box="[551,636,868,892]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="18" pageNumber="359" unit="m" value="2067.0">2067 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF87FF9B3CFC63F4FCB774ED" blockId="18.[123,1174,292,1725]" pageId="18" pageNumber="359">
<emphasis id="B90BEA99FF87FF9B3CFC63F4FE3B74CB" bold="true" box="[171,501,901,925]" pageId="18" pageNumber="359">
<smallCapsWord id="8D26A057FF87FF9B3CFC63F4FEE074CB" baselines="919,918" box="[171,302,901,925]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="18" pageNumber="359">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FF87FF9B3D6063F9FE9D74CB" baselines="918" box="[311,339,904,923]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="18" pageNumber="359">OF</smallCapsWord>
<smallCapsWord id="8D26A057FF87FF9B3D0C63F9FE6074CB" baselines="918" box="[347,430,904,923]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="18" pageNumber="359">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FF87FF9B3DEF63F9FE3B74CB" baselines="918" box="[440,501,904,923]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="18" pageNumber="359">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FF87FF9B3F5663F4FC4C74CD" authorityName="Forest Reserve, N" authorityYear="2067" box="[769,898,901,925]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF87FF9B3F5663F4FC4C74CD" box="[769,898,901,925]" italics="true" pageId="18" pageNumber="359">xiaoheishan</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the area in which the
<typeStatus id="54C48829FF87FF9B3E2B63D4FD1774ED" box="[636,729,933,957]" pageId="18" pageNumber="359" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF87FF9B3CFC63B4FCD4726D" pageId="18" pageNumber="359" type="diagnosis">
<paragraph id="8BC0368BFF87FF9B3CFC63B4FCD4726D" blockId="18.[123,1174,292,1725]" pageId="18" pageNumber="359">
<smallCapsWord id="8D26A057FF87FF9B3CFC63B4FEEE748B" baselines="983,982" box="[171,288,965,989]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="18" pageNumber="359">
<emphasis id="B90BEA99FF87FF9B3CFC63B4FEEE748B" bold="true" box="[171,288,965,989]" pageId="18" pageNumber="359">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFF87FF9B3E1B63B4FD55748D" box="[588,667,965,989]" captionStart="FIGURE" captionStartId="19.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1578]" captionTargetId="figure-15@19.[120,1176,216,1579]" captionTargetPageId="19" captionText="FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A. xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="18" pageNumber="359">Fig. 9a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
<quantity id="4C879B6EFF87FF9B38006395FF1C734D" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="18" pageNumber="359" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.3 to 5.7 mm</quantity>
), fully-winged, dorsum piceous to reddish brown and distinctly iridescent; head slender, eyes only moderately projected; clypeus with four setae; mentum and submentum fused, submentum with six setae; mandibles with two or three small setae along the dorsolateral margin of the scrobe; pronotum small and narrow (ratio PW/PL = 1.21), basal angles subrectangular, basal flattened area restricted; midlateral seta inserted at anterior one-fourth, basolateral seta at hind angle; elytra with all discal striae evident, deeply impressed and punctate, however striae 1 to 4 effaced or nearly so near base, parascutellar striole rather long, recurrent stria continuous anteriorly with stria 5, intervals convex, only a single discal seta (anterior seta) present, inserted at basal one-eighth of elytra in stria 3; median lobe of aedeagus of male (
<figureCitation id="13442A0EFF87FF9B3EAF6574FC8A724D" box="[760,836,1285,1309]" captionStart="FIGURE" captionStartId="19.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1578]" captionTargetId="figure-15@19.[120,1176,216,1579]" captionTargetPageId="19" captionText="FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A. xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="18" pageNumber="359">Fig. 9b</figureCitation>
) short, with apex bent ventrally and bluntly pointed, endophallus with a spoon-shaped sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF87FF9D3CFC6534FDF57348" lastPageId="20" lastPageNumber="361" pageId="18" pageNumber="359" type="description">
<paragraph id="8BC0368BFF87FF9B3CFC6534FF6D72CD" blockId="18.[123,1174,292,1725]" pageId="18" pageNumber="359">
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<emphasis id="B90BEA99FF87FF9B3CFC6534FEF4720B" bold="true" box="[171,314,1349,1373]" pageId="18" pageNumber="359">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size medium, BL =
<quantity id="4C879B6EFF87FF9B3E166534FD11720D" box="[577,735,1349,1373]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="18" pageNumber="359" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.3 to 5.7 mm</quantity>
. Color of dorsum piceous, shiny, iridescent, femora concolorous, tibiae, tarsi, antennae and mandibles paler reddish tan, palpi yellowish tan.
</paragraph>
<paragraph id="8BC0368BFF87FF9D3CFC65D4FCD27604" blockId="18.[123,1174,292,1725]" lastBlockId="20.[123,1174,219,1693]" lastPageId="20" lastPageNumber="361" pageId="18" pageNumber="359">
Head. Relatively slender, eyes only moderately projected, but nonetheless convex and with diameter twice length of tempora. Tempora not or only slightly convex, joined to neck region at ca. 120° angle. Frons slightly flattened, with two pairs of supraorbital setae, frontal furrows linear, distinct between the eyes, but effaced posterior to insertion of second supraorbital seta. Clypeus with four setae. Labrum slightly widened apically, apical margin slightly concave. Mandibles sharp, the right mandible (
<figureCitation id="13442A0EFF87FF9B3D7C6634FE46710D" box="[299,392,1605,1629]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="18" pageNumber="359">Fig. 16a</figureCitation>
) bidentate with the anterior tooth spaced well forward of the posterior tooth and the left mandible with only a small subtriangular process, also with two or three small setae along the dorsolateral margin of the scrobe.
<collectingRegion id="49BBF869FF87FF9B3EF666F7FD3571CD" box="[673,763,1670,1693]" country="Myanmar" name="Chin" pageId="18" pageNumber="359">Mentum</collectingRegion>
and submentum fused.
<collectingRegion id="49BBF869FF87FF9B385466F7FB9371CD" box="[1027,1117,1670,1693]" country="Myanmar" name="Chin" pageId="18" pageNumber="359">Mentum</collectingRegion>
with medial tooth rather broad, with the apex either obtuse or truncate, less than one-half as long as lateral lobes. Submentum with six setae subapically, gula broad. Genae with a single seta ventrally on each side. Antennae long and slender, extended to (in females) or slightly beyond (in males) the middle of the elytra, with five or six antennomeres beyond the pronotal base; antennonmeres 3 and 4 virtually the same length and antennomere 2 slightly shorter.
</paragraph>
<caption id="DF006603FF86FF9A3CFC664DFDF471D2" ID-DOI="http://doi.org/10.5281/zenodo.13155299" ID-Zenodo-Dep="13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="19" pageNumber="360" startId="19.[171,183,1596,1615]" targetBox="[120,1176,216,1578]" targetPageId="19" targetType="figure">
<paragraph id="8BC0368BFF86FF9A3CFC664DFDF471D2" blockId="19.[123,1174,1595,1666]" pageId="19" pageNumber="360">
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9.
<taxonomicName id="4C7F4D08FF86FF9A3D53664AFE26711E" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[260,488,1595,1614]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="species" species="xiaoheishan" status="sp. nov.">
<emphasis id="B90BEA99FF86FF9A3D53664AFE26711E" box="[260,488,1595,1614]" italics="true" pageId="19" pageNumber="360">Agonotrechus xiaoheishan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF86FF9A3DB9664DFDFA711F" box="[494,564,1596,1615]" pageId="19" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FF86FF9A3F2B6624FC327138" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[892,1020,1621,1640]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF86FF9A3F2B6624FC327138" box="[892,1020,1621,1640]" italics="true" pageId="19" pageNumber="360">A. xiaoheishan</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF81FF9D3CFC612DFCC27505" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">Pronotum. Relatively small, not or only slightly narrowed basally, only slightly transverse, ratio PW/PL = 1.21, widest at anterior one-third; lateral margins rectilinear in basal half, not sinuate except for a very slight and short inflexion just anterior to the basal angle, which is subrectangular and sharp; pronotal disc smooth, glabrous, moderately convex; median longitudinal impression very fine; basal foveae deep, basal flattened area rather small, delimited laterally by short but deep and oblique furrows. Lateral explanation slender and moderately reflexed in anterior two-thirds, then progressively broader in basal one-third without. Midlateral pair of setae inserted at anterior one-fifth and basolateral pair inserted at hind angles.</paragraph>
<paragraph id="8BC0368BFF81FF9D3CFC622FFC6974C7" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">Elytra. Convex, large and broad, especially in relation to pronotum, elytral silhouette ovoid, about equally narrowed apically and basally, humeri evident but rounded. All discal striae evident, regular, deeply impressed and punctate, however striae 1 to 4 effaced or nearly so near base, parascutellar striole rather long, recurrent stria continuous anteriorly with discal stria 5, intervals convex, only a single discal seta (anterior seta) present, inserted at basal one-eighth near stria 3. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Anterior discal seta present, inserted at basal one-eighth of elytra in stria 3, which is effaced anterior to that point. Middle discal seta absent. Preapical seta present on interval 3 near stria 2 opposite the anterior edge of the subapical sinuation. Umbilicate setal series with setae of humeral group equidistant for each other and those of median group both inserted posterior to middle of elytra.</paragraph>
<paragraph id="8BC0368BFF81FF9D3CFC63D1FDFF7488" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">Legs. Slender but only moderately long. Protibiae furrowed. Male protarsi with tarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFF81FF9D3CFC6391FDF57348" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFF81FF9D3DAE6391FD8474A8" box="[505,586,992,1016]" captionStart="FIGURE" captionStartId="19.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1578]" captionTargetId="figure-15@19.[120,1176,216,1579]" captionTargetPageId="19" captionText="FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A. xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="20" pageNumber="361">Fig. 9b</figureCitation>
) short, with apex bent ventrally and bluntly pointed, endophallus with a spoon-shaped sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF81FF9F3CFC6451FBE07623" lastPageId="22" lastPageNumber="363" pageId="20" pageNumber="361" type="discussion">
<paragraph id="8BC0368BFF81FF9D3CFC6451FDA473AA" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">
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<emphasis id="B90BEA99FF81FF9D3CFC6451FEE67367" bold="true" box="[171,296,1056,1080]" pageId="20" pageNumber="361">COMMENTS</emphasis>
</smallCapsWord>
.— Members of this new species are morphologically similar to those of
<taxonomicName id="4C7F4D08FF81FF9D3C2C6430FD877309" authority="Bates (1892)" authorityName="Bates" authorityYear="1892" box="[123,585,1089,1113]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="361" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="B90BEA99FF81FF9D3C2C6430FE507309" box="[123,414,1089,1113]" italics="true" pageId="20" pageNumber="361">Agonotrechus birmanicus</emphasis>
<bibRefCitation id="EFEE4B7AFF81FF9D3DF86433FD877309" author="BATES, H. W." box="[431,585,1089,1113]" pageId="20" pageNumber="361" pagination="267 - 428" refId="ref58154" refString="BATES, H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428" type="journal article" year="1892">Bates (1892)</bibRefCitation>
</taxonomicName>
, described from
<collectingRegion id="49BBF869FF81FF9D3F496430FC7B7309" box="[798,949,1089,1113]" country="Myanmar" name="Kachin" pageId="20" pageNumber="361">Kachin State</collectingRegion>
in
<collectingCountry id="F368761BFF81FF9D3FBB6433FB97730A" box="[1004,1113,1090,1114]" name="Myanmar" pageId="20" pageNumber="361">Myanmar</collectingCountry>
, and
<taxonomicName id="4C7F4D08FF81FF9D3C2C6410FDD57329" authority="Ueno (1986)" authorityName="Ueno" authorityYear="1986" box="[123,539,1121,1145]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="361" phylum="Arthropoda" rank="species" species="tenuicollis">
<emphasis id="B90BEA99FF81FF9D3C2C6410FE487329" box="[123,390,1121,1145]" italics="true" pageId="20" pageNumber="361">Agonotrechus tenuicollis</emphasis>
<bibRefCitation id="EFEE4B7AFF81FF9D3DD96410FDD57329" author="UENO, S. I." box="[398,539,1121,1145]" pageId="20" pageNumber="361" pagination="83 - 92" refId="ref61144" refString="UENO, S. I. 1986. A new Agonotrechus from Nepal with notes on its congeners. Bulletin of the National Science Museum, Tokyo, Series A 12: 83 - 92." type="journal article" year="1986">Uéno (1986)</bibRefCitation>
</taxonomicName>
, described from eastern
<collectingCountry id="F368761BFF81FF9D3F7F6410FCA57329" box="[808,875,1121,1145]" name="Nepal" pageId="20" pageNumber="361">Nepal</collectingCountry>
. However, they can be distinguished from members of both of these other species by the following features: pronotum with anterior margin straight, not concave, anterior angles only faintly projected and more broadly rounded, and lateral borders more slender anteriorly and at middle; and apex of the median lobe of the male aedeagus narrower and more curved.
</paragraph>
<paragraph id="8BC0368BFF81FF9D3CFC6572FC01722B" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">
<emphasis id="B90BEA99FF81FF9D3CFC6572FE6A7249" bold="true" box="[171,420,1283,1307]" pageId="20" pageNumber="361">
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<smallCapsWord id="8D26A057FF81FF9D3D476577FE6A7249" baselines="1300" box="[272,420,1286,1305]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="20" pageNumber="361">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in and under rotting logs in dark, closed-canopy broadleaf evergreen forest (
<figureCitation id="13442A0EFF81FF9D3EC36552FD2F726B" box="[660,737,1315,1339]" captionStart="FIGURE" captionStartId="106.[171,183,895,914]" captionTargetBox="[120,1176,216,881]" captionTargetId="figure-12@106.[120,1176,215,882]" captionTargetPageId="106" captionText="FIGURE 44. Photograph of habitat for trechine species in the Gaoligong Shan region. Longling County, Longjiang Township, Xiaoheishan Forest Reserve, elevation 2020 m; habitat in which specimens of Agonotrechus xiaoheishan sp. nov. and Trechus indicus Putzeys were collected. Photo by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155374" httpUri="https://zenodo.org/record/13155374/files/figure.png" pageId="20" pageNumber="361">Fig. 44</figureCitation>
) at elevations ranging from
<quantity id="4C879B6EFF81FF9D385D6552FF5D720A" metricMagnitude="3" metricUnit="m" metricValue="2.0435" metricValueMax="2.0669999999999997" metricValueMin="2.02" pageId="20" pageNumber="361" unit="m" value="2043.5" valueMax="2067.0" valueMin="2020.0">2020 to 2067 m</quantity>
. Although members of no other
<taxonomicName id="4C7F4D08FF81FF9D3DBE6532FD1E720B" box="[489,720,1347,1371]" pageId="20" pageNumber="361">
<emphasis id="B90BEA99FF81FF9D3DBE6532FDB2720B" box="[489,636,1347,1371]" italics="true" pageId="20" pageNumber="361">Agonotrechus</emphasis>
species
</taxonomicName>
have been found syntopic with those of
<taxonomicName id="4C7F4D08FF81FF9D38296532FF32722B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="361" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF81FF9D38296532FF32722B" italics="true" pageId="20" pageNumber="361">A. xiaoheishan</emphasis>
</taxonomicName>
, specimens of
<taxonomicName id="4C7F4D08FF81FF9D3DC96512FD8B722B" authorityName="Putzeys Size" authorityYear="1870" box="[414,581,1379,1403]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="361" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FF81FF9D3DC96512FD8B722B" box="[414,581,1379,1403]" italics="true" pageId="20" pageNumber="361">Trechus indicus</emphasis>
</taxonomicName>
were collected in the same samples.
</paragraph>
<paragraph id="8BC0368BFF81FF9D3CFC65F2FC02728C" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">
<emphasis id="B90BEA99FF81FF9D3CFC65F2FDBA72CA" bold="true" box="[171,628,1411,1435]" pageId="20" pageNumber="361">
<smallCapsWord id="8D26A057FF81FF9D3CFC65F2FE9872CA" baselines="1429,1429" box="[171,342,1411,1435]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="20" pageNumber="361">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF81FF9D3D0C65F6FE2172CA" baselines="1429" box="[347,495,1415,1434]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="20" pageNumber="361">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FF81FF9D3DA265F6FD8A72CA" baselines="1429" box="[501,580,1415,1434]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="20" pageNumber="361">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FF81FF9D3E1D65F6FDBA72CA" baselines="1429" box="[586,628,1415,1434]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="20" pageNumber="361">THE</smallCapsWord>
</emphasis>
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<emphasis id="B90BEA99FF81FF9D3E2D65F2FD3172CA" bold="true" box="[634,767,1411,1435]" pageId="20" pageNumber="361">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FF81FF9D3F5265F2FEAC728C" location="We" pageId="20" pageNumber="361" specimenCount="28" specimenCount-female="10" specimenCount-male="4" stateProvince="Shan">
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<emphasis id="B90BEA99FF81FF9D3F5265F2FC8E72CA" bold="true" box="[773,832,1411,1435]" pageId="20" pageNumber="361">
<collectingRegion id="49BBF869FF81FF9D3F5265F2FC8E72CA" box="[773,832,1411,1435]" country="Myanmar" name="Shan" pageId="20" pageNumber="361">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFF81FF9D3F3F65F2FC7A72CB" box="[872,948,1411,1435]" captionStart="FIGURE" captionStartId="19.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1578]" captionTargetId="figure-15@19.[120,1176,216,1579]" captionTargetPageId="19" captionText="FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A. xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="20" pageNumber="361">Fig. 9c</figureCitation>
.
<location id="8EA06050FF81FF9D3FEA65F5FC2E72CB" LSID="urn:lsid:plazi:treatment:03D6879DFF87FF9F3C2C60AAFBE07623:8EA06050FF81FF9D3FEA65F5FC2E72CB" box="[957,992,1412,1435]" name="We" pageId="20" pageNumber="361" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FF81FF9D3CCD65D5FEE272EC" box="[154,300,1444,1468]" count="14" pageId="20" pageNumber="361" type="generic">14 specimens</specimenCount>
(
<specimenCount id="9D79FD02FF81FF9D3D6A65D5FE5C72EC" box="[317,402,1444,1468]" count="4" pageId="20" pageNumber="361" type="male">4 males</specimenCount>
and
<specimenCount id="9D79FD02FF81FF9D3D9C65D5FD8B72EC" box="[459,581,1444,1468]" count="10" pageId="20" pageNumber="361" type="female">10 females</specimenCount>
), all from
<taxonomicName id="4C7F4D08FF81FF9D3EED65D5FC2472EB" authority="Forest Reserve" authorityName="Forest Reserve, N" authorityYear="2067" box="[698,1002,1444,1468]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="361" phylum="Arthropoda" rank="species" species="xiaoheishan">Xiaoheishan Forest Reserve</taxonomicName>
in the southern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FF81FF9D3D3E65B5FC07728C" box="[361,969,1476,1500]" location="Shan" pageId="20" pageNumber="361" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF81FF9D3D3E65B5FE50728C" box="[361,414,1476,1500]" country="Myanmar" name="Shan" pageId="20" pageNumber="361">Shan</collectingRegion>
(see
<typeStatus id="54C48829FF81FF9D3D8065B5FDC2728C" box="[471,524,1476,1500]" pageId="20" pageNumber="361">Type</typeStatus>
material above for exact collection data)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF81FF9D3CFC6595FCD7710D" blockId="20.[123,1174,219,1693]" pageId="20" pageNumber="361">
This species was recorded only from near the top of the western slope in the southern part of the study area (Core Area 6). Its known geographical range does not overlap with that of any other
<taxonomicName id="4C7F4D08FF81FF9D3C2C6655FEAB716C" box="[123,357,1572,1596]" pageId="20" pageNumber="361">
<emphasis id="B90BEA99FF81FF9D3C2C6655FEC0716C" box="[123,270,1572,1596]" italics="true" pageId="20" pageNumber="361">Agonotrechus</emphasis>
species
</taxonomicName>
, although
<taxonomicName id="4C7F4D08FF81FF9D3D816654FD94716C" authorityName="Ueno Size" authorityYear="1999" box="[470,602,1573,1596]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="361" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B90BEA99FF81FF9D3D816654FD94716C" box="[470,602,1573,1596]" italics="true" pageId="20" pageNumber="361">A. yunnanus</emphasis>
</taxonomicName>
has been recorded from the adjacent Core Area 7 on the eastern slope of the Gaoligong
<collectingRegion id="49BBF869FF81FF9D3DA56634FDE4710D" box="[498,554,1605,1629]" country="Myanmar" name="Shan" pageId="20" pageNumber="361">Shan</collectingRegion>
,
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to the north.
</paragraph>
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</emphasis>
.—
<materialsCitation id="3B173CD6FF81FF9D3E296614FBC871CD" country="China" location="Gaoligong Shan" pageId="20" pageNumber="361" specimenCount="1" stateProvince="Yunnan">
This species currently is known only from the
<typeStatus id="54C48829FF81FF9D38306614FB5B712D" box="[1127,1173,1637,1661]" pageId="20" pageNumber="361">type</typeStatus>
area in the southern part of the
<location id="8EA06050FF81FF9D3D9B66F4FDB271CD" LSID="urn:lsid:plazi:treatment:03D6879DFF87FF9F3C2C60AAFBE07623:8EA06050FF81FF9D3D9B66F4FDB271CD" box="[460,636,1669,1693]" country="China" name="Gaoligong Shan" pageId="20" pageNumber="361" stateProvince="Yunnan">Gaoligong Shan</location>
, in western
<collectingRegion id="49BBF869FF81FF9D3EAA66F7FC7671CD" box="[765,952,1669,1693]" country="China" name="Yunnan" pageId="20" pageNumber="361">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF81FF9D3F9466F4FBC871CD" box="[963,1030,1669,1693]" name="China" pageId="20" pageNumber="361">China</collectingCountry>
</materialsCitation>
.
</paragraph>
<caption id="DF006603FF80FF9C3EEC6658FCF3713E" ID-DOI="http://doi.org/10.5281/zenodo.13155301" ID-Zenodo-Dep="13155301" httpUri="https://zenodo.org/record/13155301/files/figure.png" pageId="21" pageNumber="362" startId="21.[699,711,1577,1596]" targetBox="[123,1176,216,1694]" targetPageId="21" targetType="figure">
<paragraph id="8BC0368BFF80FF9C3EEC6658FB597105" blockId="21.[651,1175,1576,1647]" pageId="21" pageNumber="362">
<smallCapsWord id="8D26A057FF80FF9C3EEC6658FD36716A" baselines="1591,1590" box="[699,760,1577,1596]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="21" pageNumber="362">FIGURE</smallCapsWord>
10.
<taxonomicName id="4C7F4D08FF80FF9C3F7B6659FBF1716B" authority="Ueno" authorityName="Ueno Size" authorityYear="1999" box="[812,1087,1576,1595]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="362" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B90BEA99FF80FF9C3F7B6659FBCF716B" box="[812,1025,1576,1595]" italics="true" pageId="21" pageNumber="362">Agonotrechus yunnanus</emphasis>
Uéno
</taxonomicName>
; Map of locality record (red circle) in the Gaoligong Shan region.
</paragraph>
<paragraph id="8BC0368BFF80FF9C3EDC662AFCF3713E" blockId="21.[651,1175,1576,1647]" box="[651,829,1627,1647]" pageId="21" pageNumber="362">Scale line = 100 km.</paragraph>
</caption>
<paragraph id="8BC0368BFF83FF9F3C2C60AAFC037663" blockId="22.[123,1174,219,371]" pageId="22" pageNumber="363">
Gaoligong
<collectingRegion id="49BBF869FF83FF9F3CA560AAFEE977A3" box="[242,295,219,243]" country="Myanmar" name="Shan" pageId="22" pageNumber="363">Shan</collectingRegion>
in the northern part of Core Area 7. This area is not within the geographical range of any other
<taxonomicName id="4C7F4D08FF83FF9F3D50608AFE3C7643" box="[263,498,251,275]" pageId="22" pageNumber="363">
<emphasis id="B90BEA99FF83FF9F3D50608AFE547643" box="[263,410,251,275]" italics="true" pageId="22" pageNumber="363">Agonotrechus</emphasis>
species
</taxonomicName>
, although
<taxonomicName id="4C7F4D08FF83FF9F3E33608AFCCA7643" authorityName="Forest Reserve, N" authorityYear="2067" box="[612,772,251,275]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="363" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FF83FF9F3E33608AFCCA7643" box="[612,772,251,275]" italics="true" pageId="22" pageNumber="363">A. xiaoheishan</emphasis>
</taxonomicName>
has been recorded from the adjacent Core Area 6 on the western slope of the Gaoligong
<collectingRegion id="49BBF869FF83FF9F3EF2616AFD137663" box="[677,733,283,307]" country="Myanmar" name="Shan" pageId="22" pageNumber="363">Shan</collectingRegion>
,
<quantity id="4C879B6EFF83FF9F3EBE616AFC8E7663" box="[745,832,283,307]" metricMagnitude="4" metricUnit="m" metricValue="1.33" pageId="22" pageNumber="363" unit="km" value="13.3">13.3 km</quantity>
to the south.
</paragraph>
<paragraph id="8BC0368BFF83FF9F3CFC614AFBE07623" blockId="22.[123,1174,219,371]" pageId="22" pageNumber="363">
<emphasis id="B90BEA99FF83FF9F3CFC614AFD997602" bold="true" box="[171,599,315,339]" pageId="22" pageNumber="363">
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</emphasis>
.—
<materialsCitation id="3B173CD6FF83FF9F3E29614AFBE47623" country="China" location="Gaoligong Shan" pageId="22" pageNumber="363" specimenCount="1" stateProvince="Yunnan">
This species currently is known only from the
<typeStatus id="54C48829FF83FF9F3830614DFB5B7604" box="[1127,1173,316,340]" pageId="22" pageNumber="363">type</typeStatus>
locality in the southern part of the
<location id="8EA06050FF83FF9F3DB8612AFD517623" LSID="urn:lsid:plazi:treatment:03D6879DFF87FF9F3C2C60AAFBE07623:8EA06050FF83FF9F3DB8612AFD517623" box="[495,671,347,371]" country="China" name="Gaoligong Shan" pageId="22" pageNumber="363" stateProvince="Yunnan">Gaoligong Shan</location>
, in western
<collectingRegion id="49BBF869FF83FF9F3F76612DFC127623" box="[801,988,347,371]" country="China" name="Yunnan" pageId="22" pageNumber="363">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF83FF9F3FB0612AFBE47623" box="[999,1066,347,371]" name="China" pageId="22" pageNumber="363">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,510 @@
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<mods:title id="E8DA025993F727F75AB2F257E7A2511A">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="71AF07CC1AB9F3D600AA3F05BF96CAE2">Deuve, Thierry</mods:namePart>
<mods:affiliation id="A0E97C8114D60CAF9A7C53D6C414B6E7">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="13D784D1688EAD482718CF1C813CDF81">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="3C50B1A80FC3860142E926293BCA1BCE">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="667BE2F1D2DB1F0A04A03CFDDEDDE12C" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:namePart id="6742F099E29C21E1A6B9A7F1AEAD7302">Liang, Hongbin</mods:namePart>
<mods:affiliation id="595879CF554D01BBAF7049D52967FB28">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FF88FF943C2C60AAFC9777A3" bold="true" box="[123,857,219,243]" pageId="29" pageNumber="370">
<taxonomicName id="4C7F4D08FF88FF943C2C60AAFD3877A3" ID-CoL="8MKHK" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,758,219,243]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="370" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF88FF943C2C60AAFEF977A3" bold="true" box="[123,311,219,243]" italics="true" pageId="29" pageNumber="370">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF88FF943D1060AAFE4F77A2" bold="true" box="[327,385,219,242]" italics="true" pageId="29" pageNumber="370">s. str.</emphasis>
)
<emphasis id="B90BEA99FF88FF943DC660AAFDE477A3" bold="true" box="[401,554,219,243]" italics="true" pageId="29" pageNumber="370">gongshanicus</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FF88FF943F5560AAFC9777A3" box="[770,857,219,243]" pageId="29" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF88FF943C2C608AFE9F7641" box="[123,337,251,273]" pageId="29" pageNumber="370" type="description">
<paragraph id="8BC0368BFF88FF943C2C608AFE9F7641" blockId="29.[123,857,219,273]" box="[123,337,251,273]" pageId="29" pageNumber="370">
(
<figureCitation id="13442A0EFF88FF943CD5608AFF1D7640" box="[130,211,251,273]" captionStart="FIGURE" captionStartId="30.[171,183,1582,1601]" captionTargetBox="[120,1176,221,1566]" captionTargetId="figure-11@30.[120,1176,221,1566]" captionTargetPageId="30" captionText="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="29" pageNumber="370">Figs. 13</figureCitation>
,
<figureCitation id="13442A0EFF88FF943C8A608AFECC7640" box="[221,258,251,272]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="29" pageNumber="370">39a</figureCitation>
,
<figureCitation id="13442A0EFF88FF943D5B608AFE847641" box="[268,330,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="29" pageNumber="370">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF88FF943CFC6155FD48736C" pageId="29" pageNumber="370" type="materials_examined">
<paragraph id="8BC0368BFF88FF943CFC6155FC8374CC" blockId="29.[123,1174,292,1725]" pageId="29" pageNumber="370">
<emphasis id="B90BEA99FF88FF943CFC6155FE94766A" bold="true" box="[171,346,292,316]" pageId="29" pageNumber="370">
<smallCapsWord id="8D26A057FF88FF943CFC6155FF2A766B" baselines="310,310" box="[171,228,292,316]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="29" pageNumber="370">TYPE</smallCapsWord>
<smallCapsWord id="8D26A057FF88FF943CBA6156FE94766A" baselines="309" box="[237,346,295,314]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="29" pageNumber="370">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FF88FF943DD46155FCD776EC" collectingDate="2006-08-20" collectorName="Y. Liu &amp; P. Hu &amp; D. Z. Dong &amp; J. Wang" country="China" county="Gongshan County" elevation="3950" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="29" pageNumber="370" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FF88FF943DD46155FE27766C" box="[387,489,292,316]" pageId="29" pageNumber="370" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1024868”/ “
<collectingCountry id="F368761BFF88FF943FAA6155FB98766C" box="[1021,1110,292,316]" name="China" pageId="29" pageNumber="370">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF88FF9438356155FF6A760B" country="China" name="Yunnan" pageId="29" pageNumber="370">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF88FF943CE76135FEBB760C" box="[176,373,324,348]" pageId="29" pageNumber="370">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF88FF943DD66135FD42760C" box="[385,652,324,348]" pageId="29" pageNumber="370">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF88FF943F706135FC2D760C" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF88FF943F706135FC2D760C" box="[807,995,324,348]" country="China" county="Gongshan County" latitude="27.98206" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="29" pageNumber="370" stateProvince="Yunnan">Kawakarpu Shan</location>
, on slope
<locationDeviation id="9A722FA9FF88FF9438046135FED7762C" location="Chukuai Lake" pageId="29" pageNumber="370">NE of Chukuai Lake</locationDeviation>
,
<quantity id="4C879B6EFF88FF943D7E6115FE4C762C" box="[297,386,356,380]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="29" pageNumber="370" unit="m" value="3950.0">
<elevation id="0052D1B8FF88FF943D7E6115FE4C762C" box="[297,386,356,380]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="29" pageNumber="370" unit="m" value="3950.0">3950 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFF88FF943DE76115FDE6762C" box="[432,552,356,380]" degrees="27. 98206" direction="north" orientation="latitude" pageId="29" pageNumber="370" precision="1" value="27.98206">N27.98206</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF88FF943E6B6115FD05762C" box="[572,715,356,380]" degrees="098.48027" direction="east" orientation="longitude" pageId="29" pageNumber="370" precision="1" value="98.48027">E098.48027°</geoCoordinate>
,
<date id="FFC1104BFF88FF943E8C6115FC5C762C" box="[731,914,356,380]" pageId="29" pageNumber="370" value="2006-08-20">
<collectingDate id="EF85E9A3FF88FF943E8C6115FC5C762C" box="[731,914,356,380]" pageId="29" pageNumber="370" value="2006-08-20">20 August 2006</collectingDate>
</date>
, Stop #DHK-2006-086
<collectorName id="268A535DFF88FF943C2C61F5FF0C76CC" box="[123,194,388,412]" pageId="29" pageNumber="370">Y. Liu</collectorName>
,
<collectorName id="268A535DFF88FF943C9861F5FEC076CB" box="[207,270,388,411]" pageId="29" pageNumber="370">P. Hu</collectorName>
,
<collectorName id="268A535DFF88FF943D4C61F5FE5A76CC" box="[283,404,388,412]" pageId="29" pageNumber="370">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFF88FF943DE861F5FDD876CC" box="[447,534,388,412]" pageId="29" pageNumber="370">J. Wang</collectorName>
collectors”/ “
<typeStatus id="54C48829FF88FF943EFB61F5FCF576CC" box="[684,827,388,412]" pageId="29" pageNumber="370" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FF88FF943F1261F5FE4876EC" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="370" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF88FF943F1261F5FC3776CC" box="[837,1017,388,412]" italics="true" pageId="29" pageNumber="370">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF88FF94385C61F4FB8876CC" box="[1035,1094,389,412]" italics="true" pageId="29" pageNumber="370">s. str.</emphasis>
)
<emphasis id="B90BEA99FF88FF94380061F4FF1676EC" italics="true" pageId="29" pageNumber="370">gongshanicus</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FF88FF943DC761D5FE2D76EB" box="[400,483,420,443]" pageId="29" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FF88FF943F7461D5FEB876AC" collectionCode="CAS, IOZ" country="China" pageId="29" pageNumber="370" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FF88FF943F7461D5FC4576EC" box="[803,907,420,444]" pageId="29" pageNumber="370" type="paratype">Paratypes</typeStatus>
(a total of 10):
<specimenCount id="9D79FD02FF88FF94387861D5FB5876EC" box="[1071,1174,420,444]" count="2" pageId="29" pageNumber="370" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF88FF943CF561B5FF16768C" box="[162,216,452,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="370" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF88FF943CB161B5FEDA768C" box="[230,276,452,476]" pageId="29" pageNumber="370">IOZ</collectionCode>
) labeled same as
<typeStatus id="54C48829FF88FF943D8C61B5FDF6768C" box="[475,568,452,476]" pageId="29" pageNumber="370" type="holotype">holotype</typeStatus>
except first label “CASENT 1024866” and “CASENT 1024867”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FF88FF943DD56195FC67750C" collectingDate="2006-08-21" collectionCode="IOZ, MNHN" collectorName="Y. Liu &amp; P. Hu &amp; J. Wang" country="China" county="Gongshan County" latitude="27.98631" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" pageId="29" pageNumber="370" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF88FF943DD56195FE0576AC" box="[386,459,484,508]" count="1" pageId="29" pageNumber="370" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FF88FF943E576195FD9076AC" box="[512,606,484,508]" count="1" pageId="29" pageNumber="370" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF88FF943EDC6195FD7476AC" box="[651,698,484,508]" pageId="29" pageNumber="370">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF88FF943E926195FCD676AC" box="[709,792,484,508]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="370">MNHN</collectionCode>
) labeled: “CASENT 1025160” and “CASENT 1025161”, respectively/ “
<collectingCountry id="F368761BFF88FF943E476275FDA7754C" box="[528,617,516,540]" name="China" pageId="29" pageNumber="370">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF88FF943E206274FD05754C" box="[631,715,517,540]" country="China" name="Yunnan" pageId="29" pageNumber="370">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF88FF943E8D6275FC6F754C" box="[730,929,516,540]" pageId="29" pageNumber="370">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF88FF943FF86275FF64756C" pageId="29" pageNumber="370">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF88FF943D696255FE36756C" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF88FF943D696255FE36756C" box="[318,504,548,572]" country="China" county="Gongshan County" latitude="27.98631" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="29" pageNumber="370" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FF88FF943E556255FC84756C" box="[514,842,548,572]" location="Chukuai Lake" pageId="29" pageNumber="370">
<quantity id="4C879B6EFF88FF943E556255FD94756C" box="[514,602,548,572]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="29" pageNumber="370" unit="km" value="0.75">0.75 km</quantity>
NW of Chukuai Lake
</locationDeviation>
,”/ “
<geoCoordinate id="EE4B504CFF88FF943F276255FC3D756C" box="[880,1011,548,572]" degrees="27. 98631" direction="north" orientation="latitude" pageId="29" pageNumber="370" precision="1" value="27.98631">N27.98631°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF88FF9438556255FB5F756C" box="[1026,1169,548,572]" degrees="098.47069" direction="east" orientation="longitude" pageId="29" pageNumber="370" precision="1" value="98.47069">E098.47069°</geoCoordinate>
,
<date id="FFC1104BFF88FF943C2C6235FEE8750C" box="[123,294,580,604]" pageId="29" pageNumber="370" value="2006-08-21">
<collectingDate id="EF85E9A3FF88FF943C2C6235FEE8750C" box="[123,294,580,604]" pageId="29" pageNumber="370" value="2006-08-21">21 August 2006</collectingDate>
</date>
, Stop #DHK-2006-095
<collectorName id="268A535DFF88FF943E736234FDA7750C" box="[548,617,580,604]" pageId="29" pageNumber="370">Y. Liu</collectorName>
,
<collectorName id="268A535DFF88FF943E236234FD7E750C" box="[628,688,581,604]" pageId="29" pageNumber="370">P. Hu</collectorName>
, &amp;
<collectorName id="268A535DFF88FF943E816234FCE5750D" box="[726,811,581,605]" pageId="29" pageNumber="370">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF88FF943FE46235FE0175AC" collectingDate="2006-08-19" collectingDateMax="2006-08-22" collectingDateMin="2006-08-19" collectionCode="CAS, IOZ" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu &amp; P. Hu &amp; J. Wang" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.44779" municipality="Bingzhongluo Township" pageId="29" pageNumber="370" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF88FF943FE46235FC35750C" box="[947,1019,580,604]" count="1" pageId="29" pageNumber="370" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FF88FF9438796235FB5B750C" box="[1070,1173,580,604]" count="3" pageId="29" pageNumber="370" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF88FF943CF56215FF16752C" box="[162,216,612,636]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="370" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF88FF943CB16215FEDA752C" box="[230,276,612,636]" pageId="29" pageNumber="370">IOZ</collectionCode>
) labeled “CASENT 1025935” and “CASENT 1025936” to “CASENT 1025938”, respectively/ “
<collectingCountry id="F368761BFF88FF943D4962F5FEB675CC" box="[286,376,644,668]" name="China" pageId="29" pageNumber="370">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF88FF943DDC62F4FE2E75CC" box="[395,480,645,668]" country="China" name="Yunnan" pageId="29" pageNumber="370">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF88FF943DA362F5FD0F75CC" box="[500,705,644,668]" pageId="29" pageNumber="370">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF88FF943E8262F5FC2775CC" box="[725,1001,644,668]" pageId="29" pageNumber="370">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF88FF943C2C62D5FEF875EC" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF88FF943C2C62D5FEF875EC" box="[123,310,676,700]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.44779" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="29" pageNumber="370" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FF88FF943D1562D5FDB375EC" box="[322,637,676,700]" location="Chukuai Lake" pageId="29" pageNumber="370">
<quantity id="4C879B6EFF88FF943D1562D5FE4075EC" box="[322,398,676,700]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="370" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite, “
<geoCoordinate id="EE4B504CFF88FF943F4F62D5FC5275EC" box="[792,924,676,700]" degrees="27.97686" direction="north" orientation="latitude" pageId="29" pageNumber="370" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF88FF943FFB62D5FBF475EC" box="[940,1082,676,700]" degrees="098.44779" direction="east" orientation="longitude" pageId="29" pageNumber="370" precision="1" value="98.44779">E098.44779°</geoCoordinate>
”/ “
<quantity id="4C879B6EFF88FF94380962D5FF5C758C" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="29" pageNumber="370" unit="m" value="3750.0">
<elevation id="0052D1B8FF88FF94380962D5FF5C758C" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="29" pageNumber="370" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFF88FF943CC962B5FEA1758C" box="[158,367,708,732]" pageId="29" pageNumber="370" value="2006-08-19" valueMax="2006-08-22" valueMin="2006-08-19">
<collectingDate id="EF85E9A3FF88FF943CC962B5FEA1758C" box="[158,367,708,732]" pageId="29" pageNumber="370" value="2006-08-19" valueMax="2006-08-22" valueMin="2006-08-19">19-22 August 2006</collectingDate>
</date>
, Stop #DHK-2006-095C D.H.
<taxonomicName id="4C7F4D08FF88FF943EE862B5FCF5758C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[703,827,708,732]" pageId="29" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFF88FF943F1062B5FC70758C" box="[839,958,708,732]" pageId="29" pageNumber="370">J.A. Miller</collectorName>
,
<collectorName id="268A535DFF88FF943F9D62B4FB8E758D" box="[970,1088,709,733]" pageId="29" pageNumber="370">D.Z. Dong</collectorName>
,
<collectorName id="268A535DFF88FF94381C62B4FB5F758C" box="[1099,1169,708,732]" pageId="29" pageNumber="370">Y. Liu</collectorName>
,
<collectorName id="268A535DFF88FF943C2C6294FF7375AC" box="[123,189,741,764]" pageId="29" pageNumber="370">P. Hu</collectorName>
, &amp;
<collectorName id="268A535DFF88FF943CA76294FE8275AD" box="[240,332,741,765]" pageId="29" pageNumber="370">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF88FF943DB76295FC8774CC" collectingDate="2006-08-19" collectionCode="CAS, MNHN" collectorName="J. A. Miller &amp; D. Z. Dong" country="China" county="Gongshan County" elevation="3745" latitude="27.98393" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" pageId="29" pageNumber="370" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF88FF943DB76295FDF675AC" box="[480,568,740,764]" count="2" pageId="29" pageNumber="370" type="male">2 males</specimenCount>
(in
<collectionCode id="ED6EAE4EFF88FF943E276295FD6875AC" box="[624,678,740,764]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="370" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF88FF943EEF6295FCC575AC" box="[696,779,740,764]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="370">MNHN</collectionCode>
) labeled “CASENT 1026203” and “CASENT 1026204”, respectively/ “
<collectingCountry id="F368761BFF88FF943E476375FDA7744C" box="[528,617,772,796]" name="China" pageId="29" pageNumber="370">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF88FF943E206374FD05744C" box="[631,715,773,796]" country="China" name="Yunnan" pageId="29" pageNumber="370">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF88FF943E8D6375FC6F744C" box="[730,929,772,796]" pageId="29" pageNumber="370">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF88FF943FF86375FF64746C" pageId="29" pageNumber="370">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF88FF943D166355FE35746C" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF88FF943D166355FE35746C" box="[321,507,804,828]" country="China" county="Gongshan County" latitude="27.98393" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="29" pageNumber="370" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FF88FF943E506355FC80746C" box="[519,846,804,828]" location="Chukuai Lake" pageId="29" pageNumber="370">
<quantity id="4C879B6EFF88FF943E506355FD9F746C" box="[519,593,804,828]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="370" unit="km" value="0.3">0.3 km</quantity>
NNE of Chukuai Lake
</locationDeviation>
, “
<geoCoordinate id="EE4B504CFF88FF943F346355FC28746C" box="[867,998,804,828]" degrees="27.98393" direction="north" orientation="latitude" pageId="29" pageNumber="370" precision="1" value="27.98393">N27.98393°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF88FF943FA16355FB4D746C" box="[1014,1155,804,828]" degrees="098.47491" direction="east" orientation="longitude" pageId="29" pageNumber="370" precision="1" value="98.47491">E098.47491°</geoCoordinate>
”/ “
<quantity id="4C879B6EFF88FF943CD36335FF14740C" box="[132,218,836,860]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="29" pageNumber="370" unit="m" value="3745.0">
<elevation id="0052D1B8FF88FF943CD36335FF14740C" box="[132,218,836,860]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="29" pageNumber="370" unit="m" value="3745.0">3745 m</elevation>
</quantity>
,
<date id="FFC1104BFF88FF943CB26335FE5C740C" box="[229,402,836,861]" pageId="29" pageNumber="370" value="2006-08-19">
<collectingDate id="EF85E9A3FF88FF943CB26335FE5C740C" box="[229,402,836,861]" pageId="29" pageNumber="370" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-081 D.H.
<taxonomicName id="4C7F4D08FF88FF943E9A6335FC87740C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[717,841,836,860]" pageId="29" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFF88FF943F036334FC04740C" box="[852,970,836,861]" pageId="29" pageNumber="370">J.A. Miller</collectorName>
, &amp;
<collectorName id="268A535DFF88FF943FA56334FBAB740D" box="[1010,1125,837,861]" pageId="29" pageNumber="370">D.Z. Dong</collectorName>
collectors”. All
<typeStatus id="54C48829FF88FF943D536314FEA4742D" box="[260,362,869,893]" pageId="29" pageNumber="370" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FF88FF943EEB6315FC8D742C" box="[700,835,868,892]" pageId="29" pageNumber="370" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FF88FF943F1D6315FE4774CC" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="370" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF88FF943F1D6315FC30742C" box="[842,1022,868,892]" italics="true" pageId="29" pageNumber="370">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF88FF9438596314FB86742C" box="[1038,1096,869,892]" italics="true" pageId="29" pageNumber="370">s. str.</emphasis>
)
<emphasis id="B90BEA99FF88FF9438006314FF1674CC" italics="true" pageId="29" pageNumber="370">gongshanicus</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FF88FF943DC263F4FE2674CC" box="[405,488,901,924]" pageId="29" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF88FF943CFC63D4FBCE748C" blockId="29.[123,1174,292,1725]" pageId="29" pageNumber="370">
<materialsCitation id="3B173CD6FF88FF943CFC63D4FC32748C" collectorName="Bingzhongluo Township, SW" country="China" county="Gongshan County" elevation="3950" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="29" pageNumber="370" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54C48829FF88FF943CFC63D4FF2A74EB" box="[171,228,933,957]" pageId="29" pageNumber="370">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FF88FF943CBB63D9FE9974EB" baselines="950" box="[236,343,936,955]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="29" pageNumber="370">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFF88FF943D2863D4FE0F74ED" box="[383,449,933,957]" name="China" pageId="29" pageNumber="370">China</collectingCountry>
,
<collectingRegion id="49BBF869FF88FF943D9A63D4FDEF74EC" box="[461,545,933,956]" country="China" name="Yunnan" pageId="29" pageNumber="370">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF88FF943E7A63D4FD3E74ED" box="[557,752,933,957]" pageId="29" pageNumber="370">Gongshan County</collectingCounty>
,
<collectorName id="268A535DFF88FF943EAC63D4FBF674ED" box="[763,1080,933,957]" pageId="29" pageNumber="370">
<collectingMunicipality id="6BA4ACF1FF88FF943EAC63D4FBCB74ED" box="[763,1029,933,957]" pageId="29" pageNumber="370">Bingzhongluo Township</collectingMunicipality>
, SW
</collectorName>
slope of
<location id="8EA06050FF88FF943C2C63B4FEFB748D" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF88FF943C2C63B4FEFB748D" box="[123,309,965,989]" country="China" county="Gongshan County" latitude="27.98206" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="29" pageNumber="370" stateProvince="Yunnan">Kawakarpu Shan</location>
, on slope
<locationDeviation id="9A722FA9FF88FF943DF563B4FD4A748D" box="[418,644,965,989]" location="Chukuai Lake" pageId="29" pageNumber="370">NE of Chukuai Lake</locationDeviation>
,
<geoCoordinate id="EE4B504CFF88FF943ED863B4FCC6748C" box="[655,776,964,989]" degrees="27. 98206" direction="north" orientation="latitude" pageId="29" pageNumber="370" precision="1" value="27.98206">N27.98206</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF88FF943F4763B5FC53748C" box="[784,925,964,988]" degrees="098.48027" direction="east" orientation="longitude" pageId="29" pageNumber="370" precision="1" value="98.48027">E098.48027°</geoCoordinate>
,
<quantity id="4C879B6EFF88FF943FFF63B5FC32748C" box="[936,1020,964,988]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="29" pageNumber="370" unit="m" value="3950.0">
<elevation id="0052D1B8FF88FF943FFF63B5FC32748C" box="[936,1020,964,988]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="29" pageNumber="370" unit="m" value="3950.0">3950 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF88FF943CFC6394FD48736C" blockId="29.[123,1174,292,1725]" pageId="29" pageNumber="370">
<emphasis id="B90BEA99FF88FF943CFC6394FE2274AB" bold="true" box="[171,492,997,1021]" pageId="29" pageNumber="370">
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<smallCapsWord id="8D26A057FF88FF943D636399FE9E74AB" baselines="1014" box="[308,336,1000,1019]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="29" pageNumber="370">OF</smallCapsWord>
<smallCapsWord id="8D26A057FF88FF943D026399FE6674AB" baselines="1014" box="[341,424,1000,1019]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="29" pageNumber="370">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FF88FF943DF86399FE2274AB" baselines="1014" box="[431,492,1000,1019]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="29" pageNumber="370">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FF88FF943EBB6394FCB174AD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[748,895,997,1021]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="370" phylum="Arthropoda" rank="species" species="gongshanicus">
<emphasis id="B90BEA99FF88FF943EBB6394FCB174AD" box="[748,895,997,1021]" italics="true" pageId="29" pageNumber="370">gongshanicus</emphasis>
</taxonomicName>
, is derived from the name of the county (Xian) in which the
<typeStatus id="54C48829FF88FF943DBD6474FD89734D" box="[490,583,1029,1053]" pageId="29" pageNumber="370" type="holotype">holotype</typeStatus>
was collected, Gongshan, and the Latin adjectival suffix,
<emphasis id="B90BEA99FF88FF943CF16454FF14736C" box="[166,218,1061,1084]" italics="true" pageId="29" pageNumber="370">-icus</emphasis>
, meaning belonging to or pertaining to.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF88FF943CFC6434FBCF724D" pageId="29" pageNumber="370" type="diagnosis">
<paragraph id="8BC0368BFF88FF943CFC6434FBCF724D" blockId="29.[123,1174,292,1725]" pageId="29" pageNumber="370">
<smallCapsWord id="8D26A057FF88FF943CFC6434FEEE730B" baselines="1111,1110" box="[171,288,1093,1117]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="29" pageNumber="370">
<emphasis id="B90BEA99FF88FF943CFC6434FEEE730B" bold="true" box="[171,288,1093,1117]" pageId="29" pageNumber="370">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFF88FF943E106434FD6D730D" box="[583,675,1093,1117]" captionStart="FIGURE" captionStartId="30.[171,183,1582,1601]" captionTargetBox="[120,1176,221,1566]" captionTargetId="figure-11@30.[120,1176,221,1566]" captionTargetPageId="30" captionText="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="29" pageNumber="370">Fig. 13a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: same features as members of
<taxonomicName id="4C7F4D08FF88FF943CCF64F4FED173CD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[152,287,1157,1181]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="370" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF88FF943CCF64F4FED173CD" box="[152,287,1157,1181]" italics="true" pageId="29" pageNumber="370">Q. griswoldi</emphasis>
</taxonomicName>
except, size smaller (BL =
<quantity id="4C879B6EFF88FF943E1F64F4FD2D73CD" box="[584,739,1157,1181]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="29" pageNumber="370" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.5 to 3.8 mm</quantity>
), body form short, with elytra, in particular, shorter and more oval, less elongate; digitiform projections of pronotal basal angles slightly more divergent laterally; elytral recurrent stria slightly deeper impressed, median lobe of male aedeagus (
<figureCitation id="13442A0EFF88FF943CBB6494FE8673AD" box="[236,328,1253,1277]" captionStart="FIGURE" captionStartId="30.[171,183,1582,1601]" captionTargetBox="[120,1176,221,1566]" captionTargetId="figure-11@30.[120,1176,221,1566]" captionTargetPageId="30" captionText="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="29" pageNumber="370">Fig. 13b</figureCitation>
) markedly different, with shaft thin and straighter, abruptly bent basally, apex short and rectangular, endophallus with a single apically acuminate internal sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF88FF963CFC6554FC5F7265" lastPageId="31" lastPageNumber="372" pageId="29" pageNumber="370" type="description">
<paragraph id="8BC0368BFF88FF943CFC6554FD16720D" blockId="29.[123,1174,292,1725]" pageId="29" pageNumber="370">
<smallCapsWord id="8D26A057FF88FF943CFC6554FEF4726B" baselines="1335,1334" box="[171,314,1317,1341]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="29" pageNumber="370">
<emphasis id="B90BEA99FF88FF943CFC6554FEF4726B" bold="true" box="[171,314,1317,1341]" pageId="29" pageNumber="370">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size smaller, BL =
<quantity id="4C879B6EFF88FF943E636554FD7B726C" box="[564,693,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="29" pageNumber="370" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.5-3.8 mm</quantity>
. Body color dark, reddish-piceous, antennae and legs reddish tan, palpi and tarsi paler, yellowish tan.
</paragraph>
<paragraph id="8BC0368BFF88FF943CFC6514FBDE710D" blockId="29.[123,1174,292,1725]" pageId="29" pageNumber="370">
Head. Moderate in size; eyes small and convex, their convexity greater than and their diameter about as long as tempora, the latter moderately convex and glabrous. Frons not flattened; frontal furrows deep, rounded, slightly attenuated posterior to the eyes. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave.
<collectingRegion id="49BBF869FF88FF943EF565B7FD32728D" box="[674,764,1478,1501]" country="Myanmar" name="Chin" pageId="29" pageNumber="370">Mentum</collectingRegion>
and submentum fused.
<collectingRegion id="49BBF869FF88FF94385465B7FB93728D" box="[1027,1117,1478,1501]" country="Myanmar" name="Chin" pageId="29" pageNumber="370">Mentum</collectingRegion>
with medial tooth short, wide, bifid, less than one-half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae slightly shorter, with only two antennomeres extended posteriorly beyond basal pronotal margin, antennomeres slightly broadened, antennomere 3 slightly longer than antennomeres 2 or 4.
</paragraph>
<paragraph id="8BC0368BFF88FF963CFC6617FEC376E3" blockId="29.[123,1174,292,1725]" lastBlockId="31.[123,1174,219,1333]" lastPageId="31" lastPageNumber="372" pageId="29" pageNumber="370">Pronotum. Cordate (ratio PW/PL = 1.15), markedly narrowed posteriorly, greatest width near anterior one-third, lateral margins rounded anteriorly, then straightened posteriorly just anterior to basal angles, the latter prolonged as slender, slightly divergent digitiform processes. Disc markedly convex, smooth and glabrous, median longitudinal impression only superficially impressed; basal foveae distinct but small and round; median basal area smooth and markedly transverse, delimited anteriorly by a faint transverse impression; basal margin convex and rounded. Lateral border of pronotum rudimentary, extremely slender, distinctly defined only in anterior one-third to one-half, effaced in posterior one-half to two thirds. Single midlateral setae on each side inserted at anterior one-third; single basolateral seta on each side, inserted on basal angle at base of digitform process.</paragraph>
<caption id="DF006603FF8BFF973CFC665FFD547124" ID-DOI="http://doi.org/10.5281/zenodo.13155307" ID-Zenodo-Dep="13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="30" pageNumber="371" startId="30.[171,183,1582,1601]" targetBox="[120,1176,221,1566]" targetPageId="30" targetType="figure">
<paragraph id="8BC0368BFF8BFF973CFC665FFD547124" blockId="30.[123,1173,1582,1652]" pageId="30" pageNumber="371">
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13.
<taxonomicName id="4C7F4D08FF8BFF973D5A665FFD967111" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[269,600,1582,1601]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="371" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF8BFF973D5A665FFE537111" box="[269,413,1582,1601]" italics="true" pageId="30" pageNumber="371">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8BFF973DFD665FFE167110" box="[426,472,1582,1600]" italics="true" pageId="30" pageNumber="371">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8BFF973DB4665FFD967111" box="[483,600,1582,1601]" italics="true" pageId="30" pageNumber="371">gongshanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF8BFF973E09665FFD6D7111" box="[606,675,1582,1601]" pageId="30" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FF8BFF973F866639FBAE710A" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[977,1120,1607,1627]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="371" phylum="Arthropoda" rank="species" species="gongshanicus">
<emphasis id="B90BEA99FF8BFF973F866639FBAE710A" box="[977,1120,1607,1627]" italics="true" pageId="30" pageNumber="371">Q. gongshanicus</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF8AFF963CFC61CAFDDB75C4" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">Elytra. Short, ovoid, very slightly tear-shaped, narrower anteriorly than posteriorly, humeri effaced. Disc markedly convex and smooth. All discal striae effaced, except for a faintly impressed parascutellar striole and a short and shallow recurrent stria. Basal setiferous pore present. Three discal setae present, aligned along presumed track of stria 3. Preapicale seta absent, two or three apicoangular setules present. Umbilicate setal series with setae of humeral group equidistance from each other, with the first slightly more medially inserted than the others, setae of median group inserted distinctly posterior to middle.</paragraph>
<paragraph id="8BC0368BFF8AFF963CFC62EDFB477584" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">Legs. Slightly short but slender. Protibiae with longitudinal furrow, glabrous or sparely pubescent apically on anterior surface. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFF8AFF963CFC62ADFB857444" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFF8AFF963CFC636DFF017424" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFF8AFF963DA5636DFD817464" box="[498,591,796,820]" captionStart="FIGURE" captionStartId="30.[171,183,1582,1601]" captionTargetBox="[120,1176,221,1566]" captionTargetId="figure-11@30.[120,1176,221,1566]" captionTargetPageId="30" captionText="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="31" pageNumber="372">Fig. 13b</figureCitation>
) with shaft with shaft thin and straight, abruptly bent basally, apex short and rectangular, endophallus with a single long, apically acuminate internal sclerite.
</paragraph>
<paragraph id="8BC0368BFF8AFF963CFC630DFEF37324" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">
<emphasis id="B90BEA99FF8AFF963CFC630DFE6A74C3" bold="true" box="[171,420,892,916]" pageId="31" pageNumber="372">
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</emphasis>
.— Members of this species have been found in a variety of microhabitats in the alpine zone on the southwest slope of Kawakarpu
<collectingRegion id="49BBF869FF8AFF963F4063EDFC8174E4" box="[791,847,924,948]" country="Myanmar" name="Shan" pageId="31" pageNumber="372">Shan</collectingRegion>
, at elevations ranging
<quantity id="4C879B6EFF8AFF96381463EDFF1C7484" metricMagnitude="3" metricUnit="m" metricValue="3.8475" metricValueMax="3.95" metricValueMin="3.745" pageId="31" pageNumber="372" unit="m" value="3847.5" valueMax="3950.0" valueMin="3745.0">3745 to 3950 m</quantity>
. Specimens were collected from under stones in moist meadows, on tundra slopes and ridges with sparse to thick herbaceous vegetation, at the edges of small streams and seeps from talus slopes They were also collected in pitfall traps placed at the edges of
<taxonomicName id="4C7F4D08FF8AFF963FC9638DFBF27344" box="[926,1084,1020,1044]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="31" pageNumber="372" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FF8AFF963FC9638DFBF27344" box="[926,1084,1020,1044]" italics="true" pageId="31" pageNumber="372">Rhododendron</emphasis>
</taxonomicName>
thickets up to two meters tall. Members of this species have been found syntopic with specimens of
<taxonomicName id="4C7F4D08FF8AFF96381C646CFE357304" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="372" phylum="Arthropoda" rank="species" species="balli" status="sp. nov." subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FF8AFF96381C646CFF207304" italics="true" pageId="31" pageNumber="372">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8AFF963CAB644DFE727304" box="[252,444,1084,1108]" italics="true" pageId="31" pageNumber="372">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FF8AFF963D9D644DFE357304" box="[458,507,1084,1108]" italics="true" pageId="31" pageNumber="372">balli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF8AFF963E56644CFD9B7304" box="[513,597,1085,1108]" pageId="31" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF8AFF963E08644DFC027304" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[607,972,1084,1109]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="372" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8AFF963E08644DFCDD7304" box="[607,787,1084,1108]" italics="true" pageId="31" pageNumber="372">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8AFF963F76644CFC977304" box="[801,857,1085,1108]" italics="true" pageId="31" pageNumber="372">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8AFF963F30644DFC027304" box="[871,972,1084,1108]" italics="true" pageId="31" pageNumber="372">griswoldi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FF8AFF963FA9644DFF2D7324" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="372" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FF8AFF963FA9644DFF2D7324" italics="true" pageId="31" pageNumber="372">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF8AFF963CBD642CFEF37324" box="[234,317,1117,1140]" pageId="31" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFF8AFF963CFC640DFC1973A5" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">
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<smallCapsWord id="8D26A057FF8AFF963D3764F1FE3A73C3" baselines="1166" box="[352,500,1152,1171]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="31" pageNumber="372">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FF8AFF963DAA64F1FD8273C3" baselines="1166" box="[509,588,1152,1171]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="31" pageNumber="372">WITHIN</smallCapsWord>
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</emphasis>
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<emphasis id="B90BEA99FF8AFF963EDD640DFCC173C3" bold="true" box="[650,783,1148,1172]" pageId="31" pageNumber="372">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FF8AFF963F4E640DFBF073E5" location="We" pageId="31" pageNumber="372" specimenCount="22" specimenCount-female="7" specimenCount-male="4" stateProvince="Shan">
<smallCapsWord id="8D26A057FF8AFF963F4E640DFC9A73C3" baselines="1166,1166" box="[793,852,1148,1172]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="31" pageNumber="372">
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<collectingRegion id="49BBF869FF8AFF963F4E640DFC9A73C3" box="[793,852,1148,1172]" country="Myanmar" name="Shan" pageId="31" pageNumber="372">SHAN</collectingRegion>
</emphasis>
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.—
<figureCitation id="13442A0EFF8AFF963FD7640DFC1373C5" box="[896,989,1148,1173]" captionStart="FIGURE" captionStartId="30.[171,183,1582,1601]" captionTargetBox="[120,1176,221,1566]" captionTargetId="figure-11@30.[120,1176,221,1566]" captionTargetPageId="30" captionText="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="31" pageNumber="372">Fig. 13c</figureCitation>
.
<location id="8EA06050FF8AFF963FBC640CFBC073C4" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF8AFF963FBC640CFBC073C4" box="[1003,1038,1149,1172]" name="We" pageId="31" pageNumber="372" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FF8AFF963C9964ECFE9073E5" box="[206,350,1181,1205]" count="11" pageId="31" pageNumber="372" type="generic">11 specimens</specimenCount>
(
<specimenCount id="9D79FD02FF8AFF963D3B64ECFE0E73E5" box="[364,448,1181,1205]" count="4" pageId="31" pageNumber="372" type="male">4 males</specimenCount>
and
<specimenCount id="9D79FD02FF8AFF963DA364ECFD9173E5" box="[500,607,1181,1205]" count="7" pageId="31" pageNumber="372" type="female">7 females</specimenCount>
), all from the southwest slope of Kawakarpu
</materialsCitation>
<collectingRegion id="49BBF869FF8AFF96381264ECFBB473E5" box="[1093,1146,1181,1205]" country="Myanmar" name="Shan" pageId="31" pageNumber="372">Shan</collectingRegion>
in the northern part of the Gaoligong
<materialsCitation id="3B173CD6FF8AFF963DA564CCFC1D73A5" county="Gongshan County" location="Core Area" municipality="Bingzhongluo Township" pageId="31" pageNumber="372" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF8AFF963DA564CCFDE47385" box="[498,554,1213,1237]" country="Myanmar" name="Shan" pageId="31" pageNumber="372">Shan</collectingRegion>
, in
<collectingMunicipality id="6BA4ACF1FF8AFF963E0664CCFC957385" box="[593,859,1213,1237]" pageId="31" pageNumber="372">Bingzhongluo Township</collectingMunicipality>
,
<collectingCounty id="62A14E07FF8AFF963F3164CCFBE67385" box="[870,1064,1213,1237]" pageId="31" pageNumber="372">Gongshan County</collectingCounty>
(see
<typeStatus id="54C48829FF8AFF96383764CCFB5B7385" box="[1120,1173,1213,1237]" pageId="31" pageNumber="372">Type</typeStatus>
material above for exact collection data). These localities are all in
<location id="8EA06050FF8AFF963F1964ACFC7273A5" LSID="urn:lsid:plazi:treatment:03D6879DFF88FF963C2C60AAFC5F7265:8EA06050FF8AFF963F1964ACFC7273A5" box="[846,956,1245,1269]" county="Gongshan County" municipality="Bingzhongluo Township" name="Core Area" pageId="31" pageNumber="372" stateProvince="Shan">Core Area</location>
2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF8AFF963CFC648DFC5F7265" blockId="31.[123,1174,219,1333]" pageId="31" pageNumber="372">
<emphasis id="B90BEA99FF8AFF963CFC648DFDAF7243" bold="true" box="[171,609,1276,1300]" pageId="31" pageNumber="372">
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<smallCapsWord id="8D26A057FF8AFF963D4A6571FE0A7243" baselines="1294" box="[285,452,1280,1299]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="31" pageNumber="372">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF8AFF963D9A6571FDAF7243" baselines="1294" box="[461,609,1280,1299]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="31" pageNumber="372">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FF8AFF963ED3656CFCF17265" box="[644,831,1309,1333]" country="China" name="Yunnan" pageId="31" pageNumber="372">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF8AFF963F1D656CFC437265" box="[842,909,1309,1333]" name="China" pageId="31" pageNumber="372">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,105 @@
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<mods:title id="A7D7C51344BCA7FBEF93CF23638ACF7F">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="8BAA007E6A7C386DD09E8F86CF898E75">Deuve, Thierry</mods:namePart>
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<mods:affiliation id="8C14096514EDF85DA53E271E9EC47734">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="6AB1E8C704454762509A554D6AC1BCA6">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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Subgenus
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<emphasis id="B90BEA99FF8CFF903E4B6191FD1676A8" bold="true" box="[540,728,480,504]" italics="true" pageId="25" pageNumber="366">Queinnectrechus</emphasis>
Deuve, 1992
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<treatmentCitation id="0ADE109AFF8CFF903D8E627DFCFA7572" author="DEUVE, T." box="[473,820,524,546]" page="354" pageId="25" pageNumber="366" year="1992">
<taxonomicName id="4C7F4D08FF8CFF903D8E627DFCFA7572" ID-CoL="8KYP9" authority="Deuve, 1992 a: 354" authorityName="Deuve" authorityPageNumber="354" authorityYear="1992" box="[473,820,524,546]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8CFF903D8E627DFDB57571" box="[473,635,524,545]" italics="true" pageId="25" pageNumber="366">Queinnectrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF8CFF903ED6627CFCFA7572" author="DEUVE, T." box="[641,820,524,546]" pageId="25" pageNumber="366" refId="ref58815" refString="DEUVE, T. 1992 a. Un nouveau genre de Trechinae des montagnes du Sichuan (Coleoptera, Trechidae). Bulletin de la Societe Entomologique de France 96 [1991]: 354." type="journal volume" year="1992">Deuve, 1992a:354</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8CFF903CFC6244FCA6747E" pageId="25" pageNumber="366" type="diagnosis">
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<emphasis id="B90BEA99FF8CFF903CFC6244FEEE751C" bold="true" box="[171,288,565,589]" pageId="25" pageNumber="366">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this subgenus (
<figureCitation id="13442A0EFF8CFF903E086244FD09751D" box="[607,711,565,589]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="25" pageNumber="366">Figs. 12a</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903E816244FD33751D" box="[726,765,565,589]" captionStart="FIGURE" captionStartId="30.[171,183,1582,1601]" captionTargetBox="[120,1176,221,1566]" captionTargetId="figure-11@30.[120,1176,221,1566]" captionTargetPageId="30" captionText="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155307" httpUri="https://zenodo.org/record/13155307/files/figure.png" pageId="25" pageNumber="366">13a</figureCitation>
) can be recognized by the following combination of character states: size moderate (BL =
<quantity id="4C879B6EFF8CFF903EEE6224FC9C753E" box="[697,850,597,622]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.8" metricValueMin="3.5" pageId="25" pageNumber="366" unit="mm" value="4.15" valueMax="4.8" valueMin="3.5">3.5 to 4.8 mm</quantity>
), apterous, body dark, reddish brown to piceous, surface micaceous; pronotum with basal angles prolonged posteriorly as slender digitiform processes, with two setae (midlateral and basolateral) present on each side; elytra markedly convex, inflated, slightly tear-shaped, humeri effaced, elytral discal striae effaced, with two (in most members) or three (in a few members) discal setae present, aligned on interval 3 near stria 3, preapical seta absent from most members.
</paragraph>
<paragraph id="8BC0368BFF8CFF903CFC6364FCA6747E" blockId="25.[123,1174,565,814]" box="[171,872,789,814]" pageId="25" pageNumber="366">
<emphasis id="B90BEA99FF8CFF903CFC6364FE3F747C" bold="true" box="[171,497,789,813]" pageId="25" pageNumber="366">
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</emphasis>
.— Same as for genus (see above).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,657 @@
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<mods:title id="25358D0B5D18A0D541AD091D7EA67194">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="EB57460AAB0D5B472900450C99A03B5D">Deuve, Thierry</mods:namePart>
<mods:affiliation id="E62D56929F0807C8ADB32BDECA39970B">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="AB57C5E8ED9B3969FAF2FA296D8B0C22">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="910E2572C24543349F68B1DDDB3D98E6">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="A6820F4950169F31486A50CE45711D9D" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="1050D20A069F39A3A94DB8DF5AFC3A64" type="personal">
<mods:role id="27B471146B3DD77AC2C97A0F1556E6FC">
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<mods:namePart id="443C038F7B28DCD170016C9343027466">Liang, Hongbin</mods:namePart>
<mods:affiliation id="5DC634AEB9168B63EE7F2E3DC06F25CD">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:date id="DD4A6671C2000F94E55A5CF1FDBBB9DC">2016</mods:date>
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<heading id="D08881E7FF8CFF903C2C644AFCA87304" box="[123,870,1083,1108]" fontSize="10" level="2" pageId="25" pageNumber="366" reason="4">
<emphasis id="B90BEA99FF8CFF903C2C644AFCA87304" bold="true" box="[123,870,1083,1108]" pageId="25" pageNumber="366">
<taxonomicName id="4C7F4D08FF8CFF903C2C644AFD8A7304" ID-CoL="8MCS6" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,580,1083,1108]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="366" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8CFF903C2C644AFEF97303" bold="true" box="[123,311,1083,1107]" italics="true" pageId="25" pageNumber="366">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8CFF903D10644DFE4F7303" bold="true" box="[327,385,1084,1107]" italics="true" pageId="25" pageNumber="366">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8CFF903DC6644AFE387303" bold="true" box="[401,502,1083,1107]" italics="true" pageId="25" pageNumber="366">griswoldi</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FF8CFF903F47644DFCA87304" box="[784,870,1084,1108]" pageId="25" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8CFF903C2C642DFE2D7321" box="[123,483,1115,1138]" pageId="25" pageNumber="366" type="description">
<paragraph id="8BC0368BFF8CFF903C2C642DFE2D7321" blockId="25.[123,870,1083,1138]" box="[123,483,1115,1138]" pageId="25" pageNumber="366">
(
<figureCitation id="13442A0EFF8CFF903CD5642DFF1D7321" box="[130,211,1116,1138]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="25" pageNumber="366">Figs. 12</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903C8A642DFECD7322" box="[221,259,1116,1138]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="25" pageNumber="366">37b</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903D59642DFEAD7322" box="[270,355,1116,1138]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="25" pageNumber="366">38a, 38b</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903D39642DFE5A7322" box="[366,404,1116,1138]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="25" pageNumber="366">39b</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903DC9642AFE127321" box="[414,476,1115,1137]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="25" pageNumber="366">4648</figureCitation>
)
</paragraph>
</subSubSection>
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</emphasis>
.—
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<typeStatus id="54C48829FF8CFF903DD464F5FE2773CC" box="[387,489,1156,1180]" pageId="25" pageNumber="366" type="holotype">Holotype</typeStatus>
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,
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,
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SE of Heipu Yakou
</locationDeviation>
in valley below tunnel,
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/
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,”/ “
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,
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</date>
, Stop #DHK-2006-073 D.H.
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&amp;
<collectorName id="268A535DFF8CFF903C2C6494FF2173AD" box="[123,239,1253,1277]" pageId="25" pageNumber="366">J.A. Miller</collectorName>
collectors”/ “
<typeStatus id="54C48829FF8CFF903D286494FDC073AD" box="[383,526,1253,1277]" pageId="25" pageNumber="366" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FF8CFF903E426494FC1E73AC" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[533,976,1253,1277]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="366" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8CFF903E426494FD0773AD" box="[533,713,1253,1277]" italics="true" pageId="25" pageNumber="366">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8CFF903E8F6494FCDE73AC" box="[728,784,1253,1276]" italics="true" pageId="25" pageNumber="366">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8CFF903F496494FC4D73AD" box="[798,899,1253,1277]" italics="true" pageId="25" pageNumber="366">griswoldi</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FF8CFF9038206494FF66724C" pageId="25" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FF8CFF903DB86574FBC4720D" collectionCode="CAS, IOZ, MNHN" country="China" pageId="25" pageNumber="366" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FF8CFF903DB86574FD99724D" box="[495,599,1285,1309]" pageId="25" pageNumber="366" type="paratype">Paratypes</typeStatus>
(a total of 20):
<specimenCount id="9D79FD02FF8CFF903F556575FC9B724D" box="[770,853,1284,1309]" count="5" pageId="25" pageNumber="366" type="male">5 males</specimenCount>
and
<specimenCount id="9D79FD02FF8CFF903FDC6574FC3D724D" box="[907,1011,1285,1309]" count="3" pageId="25" pageNumber="366" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF9038776574FB98724D" box="[1056,1110,1285,1309]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF9038356574FB5F724D" box="[1122,1169,1285,1309]" pageId="25" pageNumber="366">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903C2C6554FF00726D" box="[123,206,1317,1341]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="366">MNHN</collectionCode>
) labeled same as
<typeStatus id="54C48829FF8CFF903DDD6554FE29726D" box="[394,487,1317,1341]" pageId="25" pageNumber="366" type="holotype">holotype</typeStatus>
except first label “CASENT 1026333”, “CASENT 1026335” to “CASENT 1026338” and “CASENT 1026330” to “CASENT 1026332”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF9038426534FC3972ED" collectingDate="2002-10-01" collectingDateMax="2002-10-02" collectingDateMin="2002-10-01" collectionCode="IOZ" collectorName="P. E. Marek" country="China" county="Gongshan County" elevation="3370" latitude="27.77032" location="Cikai Township" longLatPrecision="1" longitude="98.44661" pageId="25" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF9038426534FBBC720D" box="[1045,1138,1349,1373]" count="1" pageId="25" pageNumber="366" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903C2C6514FF67722D" box="[123,169,1381,1405]" pageId="25" pageNumber="366">IOZ</collectionCode>
) labeled “CASENT 1010344”/ “
<collectingCountry id="F368761BFF8CFF903E476514FDA7722D" box="[528,617,1381,1405]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF903E216514FD04722C" box="[630,714,1381,1404]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903E8F6514FC50722D" box="[728,926,1381,1405]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<location id="8EA06050FF8CFF903FFC6514FB94722D" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903FFC6514FB94722D" box="[939,1114,1381,1405]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44661" name="Cikai Township" pageId="25" pageNumber="366" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FF8CFF9038306514FDA372CD" location="Gongshan on Dulong Valley Road" pageId="25" pageNumber="366">
<quantity id="4C879B6EFF8CFF9038306514FF5372CD" metricMagnitude="4" metricUnit="m" metricValue="5.26" pageId="25" pageNumber="366" unit="km" value="52.6">52.6 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFF8CFF903E2B65F4FCDF72CD" box="[636,785,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="25" pageNumber="366" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FF8CFF903E2B65F4FCDF72CD" box="[636,785,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="25" pageNumber="366" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFF8CFF903F6A65F4FC0F72CD" box="[829,961,1413,1437]" degrees="27.77032" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903F8365F4FBAD72CD" box="[980,1123,1413,1437]" degrees="098.44661" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.44661">E098.44661°</geoCoordinate>
,
<date id="FFC1104BFF8CFF90382565F4FEDC72ED" pageId="25" pageNumber="366" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">
<collectingDate id="EF85E9A3FF8CFF90382565F4FEDC72ED" pageId="25" pageNumber="366" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">1-2 October 2002</collectingDate>
</date>
, Stop #DHK-2002-034, D.H.
<taxonomicName id="4C7F4D08FF8CFF903E0A65D4FD1872ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[605,726,1445,1469]" pageId="25" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFF8CFF903EA965D4FCB972ED" box="[766,887,1445,1469]" pageId="25" pageNumber="366">P.E. Marek</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF90385365D4FC62716D" collectingDate="2006-08-20" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; P. Hu &amp; D. Z. Dong &amp; J. Wang" country="China" county="Gongshan County" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="25" pageNumber="366" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF90385365D4FB8172ED" box="[1028,1103,1445,1469]" count="1" pageId="25" pageNumber="366" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FF8CFF9038DF65D4FF00728D" count="3" pageId="25" pageNumber="366" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903D5365B4FEF4728D" box="[260,314,1477,1501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903D1D65B4FEB6728D" box="[330,376,1477,1501]" pageId="25" pageNumber="366">IOZ</collectionCode>
) labeled “CASENT 1024862” and “CASENT 1024863” to “CASENT 1024865”, respectively/ “
<collectingCountry id="F368761BFF8CFF903DDD6594FE2D72AD" box="[394,483,1509,1533]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF903DB96597FD8C72AD" box="[494,578,1510,1533]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903E1A6594FCDE72AD" box="[589,784,1509,1533]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF8CFF903F4D6594FBEA72AD" box="[794,1060,1509,1533]" pageId="25" pageNumber="366">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF8CFF903CC06674FE9F714D" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903CC06674FE9F714D" box="[151,337,1541,1565]" country="China" county="Gongshan County" latitude="27.98206" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="25" pageNumber="366" stateProvince="Yunnan">Kawakarpu Shan</location>
, on slope
<locationDeviation id="9A722FA9FF8CFF903DEC6677FD54714D" box="[443,666,1541,1565]" location="Chukuai Lake" pageId="25" pageNumber="366">NE of Chukuai Lake</locationDeviation>
, 3950,”/”
<geoCoordinate id="EE4B504CFF8CFF903EA96674FC4F714D" box="[766,897,1541,1565]" degrees="27. 98206" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.98206">N27.98206°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903FD86674FBD0714D" box="[911,1054,1541,1565]" degrees="098.48027" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.48027">E098.48027°</geoCoordinate>
,
<date id="FFC1104BFF8CFF90387E6674FF7D716D" pageId="25" pageNumber="366" value="2006-08-20">
<collectingDate id="EF85E9A3FF8CFF90387E6674FF7D716D" pageId="25" pageNumber="366" value="2006-08-20">20 August 2006</collectingDate>
</date>
, Stop #DHK-2006-086
<collectorName id="268A535DFF8CFF903DF96657FE3D716D" box="[430,499,1573,1597]" pageId="25" pageNumber="366">Y. Liu</collectorName>
,
<collectorName id="268A535DFF8CFF903DAB6657FDF6716D" box="[508,568,1574,1597]" pageId="25" pageNumber="366">P. Hu</collectorName>
,
<collectorName id="268A535DFF8CFF903E156657FD79716E" box="[578,695,1574,1598]" pageId="25" pageNumber="366">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFF8CFF903E8C6657FCFE716E" box="[731,816,1574,1598]" pageId="25" pageNumber="366">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF903FE16654FC8071ED" collectingDate="2006-08-18" collectionCode="CAS, IOZ, MNHN" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="25" pageNumber="366" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF903FE16654FBC9716D" box="[950,1031,1573,1597]" count="3" pageId="25" pageNumber="366" type="male">3 males</specimenCount>
and
<specimenCount id="9D79FD02FF8CFF90386D6654FB58716D" box="[1082,1174,1573,1597]" count="1" pageId="25" pageNumber="366" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903CF36634FF14710D" box="[164,218,1605,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903CBD6634FED7710D" box="[234,281,1605,1629]" pageId="25" pageNumber="366">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903D7E6634FEB2710D" box="[297,380,1605,1629]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="366">MNHN</collectionCode>
) labeled “CASENT 1026813” to “CASENT 1026815” and “CASENT 1026815”, respectively/ “
<collectingCountry id="F368761BFF8CFF903DDD6614FE2D712D" box="[394,483,1637,1661]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF903DB96617FD8C712D" box="[494,578,1638,1661]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903E1A6614FCDE712D" box="[589,784,1637,1661]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF8CFF903F4D6614FBEA712D" box="[794,1060,1637,1661]" pageId="25" pageNumber="366">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF8CFF903CC066F4FE8371CD" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903CC066F4FE8371CD" box="[151,333,1669,1693]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="25" pageNumber="366" stateProvince="Yunnan">Kawakarpu Shan</location>
at
<location id="8EA06050FF8CFF903D3966F4FDC871CD" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903D3966F4FDC871CD" box="[366,518,1669,1693]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="25" pageNumber="366" stateProvince="Yunnan">Chukuai Lake</location>
,
<quantity id="4C879B6EFF8CFF903E4666F4FDAD71CD" box="[529,611,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="25" pageNumber="366" unit="m" value="3720.0">
<elevation id="0052D1B8FF8CFF903E4666F4FDAD71CD" box="[529,611,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="25" pageNumber="366" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFF8CFF903EDC66F4FCC071CD" box="[651,782,1669,1693]" degrees="27.98121" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903F4A66F4FC6271CD" box="[797,940,1669,1693]" degrees="098.47580" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.4758">E098.47580°</geoCoordinate>
,
<date id="FFC1104BFF8CFF903FE166F4FB9071CD" box="[950,1118,1669,1694]" pageId="25" pageNumber="366" value="2006-08-18">
<collectingDate id="EF85E9A3FF8CFF903FE166F4FB9071CD" box="[950,1118,1669,1694]" pageId="25" pageNumber="366" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-079
<collectorName id="268A535DFF8CFF903D6866D7FE7271ED" box="[319,444,1701,1726]" pageId="25" pageNumber="366">J.A. Miller</collectorName>
,
<collectorName id="268A535DFF8CFF903D9966D7FD8571EE" box="[462,587,1702,1726]" pageId="25" pageNumber="366">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFF8CFF903E2866D7FD0771ED" box="[639,713,1701,1725]" pageId="25" pageNumber="366">Y. Liu</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF903F3766D4FB5B71ED" box="[864,1173,1701,1725]" collectionCode="CAS" country="China" pageId="25" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF903F3766D4FC0A71ED" box="[864,964,1701,1725]" count="1" pageId="25" pageNumber="366" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903FA966D4FBFD71ED" box="[1022,1075,1701,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
) labeled
</materialsCitation>
</paragraph>
<caption id="DF006603FF8FFF933CFC6640FD547126" ID-DOI="http://doi.org/10.5281/zenodo.13155305" ID-Zenodo-Dep="13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="26" pageNumber="367" startId="26.[171,183,1585,1604]" targetBox="[147,1176,216,1569]" targetPageId="26" targetType="figure">
<paragraph id="8BC0368BFF8FFF933CFC6640FD547126" blockId="26.[123,1173,1584,1655]" pageId="26" pageNumber="367">
<smallCapsWord id="8D26A057FF8FFF933CFC6640FF267112" baselines="1599,1598" box="[171,232,1585,1604]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="26" pageNumber="367">FIGURE</smallCapsWord>
12.
<taxonomicName id="4C7F4D08FF8FFF933D446641FD8C7113" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[275,578,1584,1603]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="367" phylum="Arthropoda" rank="species" species="griswoldi" status="sp. nov.">
<emphasis id="B90BEA99FF8FFF933D446641FE6D7113" box="[275,419,1584,1603]" italics="true" pageId="26" pageNumber="367">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8FFF933DE56640FE2D7113" box="[434,483,1585,1603]" italics="true" pageId="26" pageNumber="367">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8FFF933DA66641FD8C7113" box="[497,578,1584,1603]" italics="true" pageId="26" pageNumber="367">griswoldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF8FFF933E1D6640FD5C7114" box="[586,658,1585,1604]" pageId="26" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FF8FFF933FB8663BFB92710D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1007,1116,1610,1629]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="367" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8FFF933FB8663BFB92710D" box="[1007,1116,1610,1629]" italics="true" pageId="26" pageNumber="367">Q. griswoldi</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF8EFF923C2C60AAFCEA7684" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<materialsCitation id="3B173CD6FF8EFF923CD360AAFCD17684" collectingDate="2006-08-18" collectingDateMax="2006-08-19" collectingDateMin="2006-08-18" collectionCode="IOZ" collectorName="Y. Liu &amp; All" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="27" pageNumber="368" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
CASENT 1025813”/ “
<collectingCountry id="F368761BFF8EFF923D2360AAFE0377A3" box="[372,461,219,243]" name="China" pageId="27" pageNumber="368">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8EFF923D8060AAFDE577A2" box="[471,555,219,242]" country="China" name="Yunnan" pageId="27" pageNumber="368">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8EFF923E6260AAFD3977A3" box="[565,759,219,243]" pageId="27" pageNumber="368">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF8EFF923F5660AAFBC777A3" box="[769,1033,219,243]" pageId="27" pageNumber="368">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF8EFF923C2C608AFEFF7643" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923C2C608AFEFF7643" box="[123,305,251,275]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="27" pageNumber="368" stateProvince="Yunnan">Kawakarpu Shan</location>
<locationDeviation id="9A722FA9FF8EFF923D60608AFDAB7643" box="[311,613,251,275]" location="Chukuai Lake" pageId="27" pageNumber="368">
<quantity id="4C879B6EFF8EFF923D60608AFE4E7643" box="[311,384,251,275]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="368" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/, “
<geoCoordinate id="EE4B504CFF8EFF923F43608AFC597643" box="[788,919,251,275]" degrees="27.97686" direction="north" orientation="latitude" pageId="27" pageNumber="368" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8EFF923FF1608AFBFA7643" box="[934,1076,251,275]" degrees="098.47799" direction="east" orientation="longitude" pageId="27" pageNumber="368" precision="1" value="98.47799">E098.47799°</geoCoordinate>
,
<quantity id="4C879B6EFF8EFF923868608AFB5F7643" box="[1087,1169,251,275]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="27" pageNumber="368" unit="m" value="3750.0">
<elevation id="0052D1B8FF8EFF923868608AFB5F7643" box="[1087,1169,251,275]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="27" pageNumber="368" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFF8EFF923C2C616DFEE47663" box="[123,298,283,308]" pageId="27" pageNumber="368" value="2006-08-18">
<collectingDate id="EF85E9A3FF8EFF923C2C616DFEE47663" box="[123,298,283,308]" pageId="27" pageNumber="368" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-078 D.H.
<taxonomicName id="4C7F4D08FF8EFF923E39616AFD297663" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[622,743,283,307]" pageId="27" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”;
<specimenCount id="9D79FD02FF8EFF923F3B616DFC197663" box="[876,983,283,308]" count="2" pageId="27" pageNumber="368" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8EFF92385E616DFBF97664" box="[1033,1079,284,308]" pageId="27" pageNumber="368">IOZ</collectionCode>
) labeled “CASENT 1025836” and “CASENT 1025837”, respectively / “
<collectingCountry id="F368761BFF8EFF923F48614DFCB67604" box="[799,888,316,340]" name="China" pageId="27" pageNumber="368">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8EFF923FD5614DFC187603" box="[898,982,316,339]" country="China" name="Yunnan" pageId="27" pageNumber="368">Yunnan</collectingRegion>
,
<location id="8EA06050FF8EFF923FB7614DFF5F7624" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923FB7614DFF5F7624" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Gongshan County" pageId="27" pageNumber="368" stateProvince="Yunnan">Gongshan County</location>
,
<location id="8EA06050FF8EFF923CCC612DFE6B7624" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923CCC612DFE6B7624" box="[155,421,348,372]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Bingzhongluo Township" pageId="27" pageNumber="368" stateProvince="Yunnan">Bingzhongluo Township</location>
, SW slope of Kawakarpu Shan,
<locationDeviation id="9A722FA9FF8EFF923EA9612DFBFF7624" box="[766,1073,348,372]" location="Chukuai Lake" pageId="27" pageNumber="368">
<quantity id="4C879B6EFF8EFF923EA9612DFC877624" box="[766,841,348,372]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="368" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/ “
<geoCoordinate id="EE4B504CFF8EFF923C9D610DFE8076C4" box="[202,334,380,404]" degrees="27.97686" direction="north" orientation="latitude" pageId="27" pageNumber="368" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8EFF923D0A610DFE2276C4" box="[349,492,380,404]" degrees="098.44779" direction="east" orientation="longitude" pageId="27" pageNumber="368" precision="1" value="98.44779">E098.44779°</geoCoordinate>
,
<quantity id="4C879B6EFF8EFF923DA0610DFD8576C4" box="[503,587,380,404]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="27" pageNumber="368" unit="m" value="3750.0">
<elevation id="0052D1B8FF8EFF923DA0610DFD8576C4" box="[503,587,380,404]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="27" pageNumber="368" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFF8EFF923E00610DFCCD76C4" box="[599,771,380,404]" pageId="27" pageNumber="368" value="2006-08-19">
<collectingDate id="EF85E9A3FF8EFF923E00610DFCCD76C4" box="[599,771,380,404]" pageId="27" pageNumber="368" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-082
<collectorName id="268A535DFF8EFF923854610CFB8876C4" box="[1027,1094,380,404]" pageId="27" pageNumber="368">Y. Liu</collectorName>
collector”.
<collectorName id="268A535DFF8EFF923CE661EDFF1D76E4" box="[177,211,412,436]" pageId="27" pageNumber="368">All</collectorName>
<typeStatus id="54C48829FF8EFF923C8C61ECFE8F76E5" box="[219,321,413,437]" pageId="27" pageNumber="368" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FF8EFF923EC361EDFCD576E4" box="[660,795,412,436]" pageId="27" pageNumber="368" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FF8EFF923F7461EDFF0F7684" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="368" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8EFF923F7461EDFC1976E4" box="[803,983,412,436]" italics="true" pageId="27" pageNumber="368">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8EFF923FB061ECFBEF76E4" box="[999,1057,413,436]" italics="true" pageId="27" pageNumber="368">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8EFF92386661EDFB5876E4" box="[1073,1174,412,436]" italics="true" pageId="27" pageNumber="368">griswoldi</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FF8EFF923D3C61CCFE717684" box="[363,447,445,468]" pageId="27" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC61ACFC807545" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<materialsCitation id="3B173CD6FF8EFF923CFC61ACFC847545" country="China" county="Gongshan County" elevation="3270" latitude="27.77437" location="Cikai Township" longLatPrecision="1" longitude="98.44793" pageId="27" pageNumber="368" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FF8EFF923CFC61ACFE9676A3" bold="true" box="[171,344,477,501]" pageId="27" pageNumber="368">
<smallCapsWord id="8D26A057FF8EFF923CFC61ACFF2A76A4" baselines="495,495" box="[171,228,477,501]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="368">
<typeStatus id="54C48829FF8EFF923CFC61ACFF2A76A4" box="[171,228,477,501]" pageId="27" pageNumber="368">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FF8EFF923CBA6191FE9676A3" baselines="494" box="[237,344,480,499]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="368">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFF8EFF923DD661ACFE0D76A5" box="[385,451,477,501]" name="China" pageId="27" pageNumber="368">China</collectingCountry>
,
<collectingRegion id="49BBF869FF8EFF923D8761ACFDEA76A4" box="[464,548,477,500]" country="China" name="Yunnan" pageId="27" pageNumber="368">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8EFF923E6761ACFD3B76A5" box="[560,757,477,501]" pageId="27" pageNumber="368">Gongshan County</collectingCounty>
,
<location id="8EA06050FF8EFF923F5661ACFC6076A5" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923F5661ACFC6076A5" box="[769,942,477,501]" country="China" county="Gongshan County" latitude="27.77437" longLatPrecision="1" longitude="98.44793" name="Cikai Township" pageId="27" pageNumber="368" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FF8EFF923FED61ACFF0C7545" location="Heipu Yakou" pageId="27" pageNumber="368">
<quantity id="4C879B6EFF8EFF923FED61ACFBC876A5" box="[954,1030,477,501]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="368" unit="km" value="0.1">0.1 km</quantity>
SE of Heipu Yakou
</locationDeviation>
, in valley below tunnel,
<geoCoordinate id="EE4B504CFF8EFF923D9C618CFD827545" box="[459,588,509,533]" degrees="27.77437" direction="north" orientation="latitude" pageId="27" pageNumber="368" precision="1" value="27.77437">N27.77437°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8EFF923E0B618CFD247545" box="[604,746,509,533]" degrees="098.44793" direction="east" orientation="longitude" pageId="27" pageNumber="368" precision="1" value="98.44793">E098.44793°</geoCoordinate>
,
<quantity id="4C879B6EFF8EFF923EA1618CFC847545" box="[758,842,509,533]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="27" pageNumber="368" unit="m" value="3270.0">
<elevation id="0052D1B8FF8EFF923EA1618CFC847545" box="[758,842,509,533]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="27" pageNumber="368" unit="m" value="3270.0">3270 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC626CFD4375E6" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<emphasis id="B90BEA99FF8EFF923CFC626CFE217564" bold="true" box="[171,495,541,565]" pageId="27" pageNumber="368">
<smallCapsWord id="8D26A057FF8EFF923CFC626CFEE07564" baselines="559,559" box="[171,302,541,565]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="27" pageNumber="368">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FF8EFF923D626250FE9F7564" baselines="559" box="[309,337,545,564]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="27" pageNumber="368">OF</smallCapsWord>
<smallCapsWord id="8D26A057FF8EFF923D006250FE647564" baselines="559" box="[343,426,545,564]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="27" pageNumber="368">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FF8EFF923DE56250FE217564" baselines="559" box="[434,495,545,564]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="27" pageNumber="368">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FF8EFF923EA3626CFC977565" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[756,857,541,565]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="368" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8EFF923EA3626CFC977565" box="[756,857,541,565]" italics="true" pageId="27" pageNumber="368">griswoldi</emphasis>
</taxonomicName>
, is the Latinized form (in the genitive case) of the surname of Charles E. Griswold, now Curator Emeritus and former Schlinger Chair of Arachnology at the
<collectingRegion id="49BBF869FF8EFF923DE0622FFDED7526" box="[439,547,606,630]" country="United States of America" name="California" pageId="27" pageNumber="368">California</collectingRegion>
Academy of Sciences, who participated in several of the expeditions to the study area and helped collect many carabid specimens for this project. We are pleased to name this elegant species in his honor.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8EFF923CFC62CFFD917488" pageId="27" pageNumber="368" type="diagnosis">
<paragraph id="8BC0368BFF8EFF923CFC62CFFD917488" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<smallCapsWord id="8D26A057FF8EFF923CFC62CFFEEE7585" baselines="720,720" box="[171,288,702,726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="27" pageNumber="368">
<emphasis id="B90BEA99FF8EFF923CFC62CFFEEE7585" bold="true" box="[171,288,702,726]" pageId="27" pageNumber="368">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFF8EFF923E1062CFFD6D7587" box="[583,675,702,727]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="27" pageNumber="368">Fig. 12a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
<quantity id="4C879B6EFF8EFF92380062AEFF1F7447" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="27" pageNumber="368" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">4.3 to 4.8 mm</quantity>
), body dark reddish brown, very shiny; eyes small and convex, tempora slightly convex; pronotum markedly narrowed posteriorly, markedly convex, globulose, narrowly cordate, disc smooth, basal angles with distict digitorm projections, each side with a single midlateral and basolateral seta; elytra markedly convex, tear-shaped, discal striae effaced, with three discal setae in a row along the presumed location of stria 3, preapical seta absent; median lobe of male aedeagus (
<figureCitation id="13442A0EFF8EFF923CD463D1FF1074E8" box="[131,222,928,952]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="27" pageNumber="368">Fig. 12b</figureCitation>
) large, elongate, broadest at mid-shaft, with apex long and recurved dorsally, endophallus with internal sclerites acuminate apically.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8EFF953CFC6391FC797526" lastPageId="28" lastPageNumber="369" pageId="27" pageNumber="368" type="description">
<paragraph id="8BC0368BFF8EFF923CFC6391FF107369" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<smallCapsWord id="8D26A057FF8EFF923CFC6391FEF474A7" baselines="1010,1010" box="[171,314,992,1016]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="27" pageNumber="368">
<emphasis id="B90BEA99FF8EFF923CFC6391FEF474A7" bold="true" box="[171,314,992,1016]" pageId="27" pageNumber="368">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size medium, BL =
<quantity id="4C879B6EFF8EFF923E146390FD2C74A8" box="[579,738,993,1017]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="27" pageNumber="368" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">4.3 to 4.8 mm</quantity>
. Color of body, antennae and legs dark reddish brown, palpi paler, yellowish tan, body surface very shiny, micaceous, smooth and glabrous.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC6430FBDE72CB" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
Head. Moderate in size, eyes small and convex, their convexity greater than and their diameter about as long as tempora, the latter only slightly convex and glabrous. Frons with frontal furrows deep, rounded, and not interrupted posteriorly; two suporaorbital setae present, the anterior inserted opposite midpoint of eye, the posterior inserted in postocular groove. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible tridentate. Left mandible with a small, minutely tridentate process.
<collectingRegion id="49BBF869FF8EFF923EFF6492FCCC73AA" box="[680,770,1251,1274]" country="Myanmar" name="Chin" pageId="27" pageNumber="368">Mentum</collectingRegion>
and submentum fused.
<collectingRegion id="49BBF869FF8EFF9238536492FB9073AA" box="[1028,1118,1251,1274]" country="Myanmar" name="Chin" pageId="27" pageNumber="368">Mentum</collectingRegion>
with medial tooth wide, bifid or truncate, one half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae of moderate length, extended posteriorly almost to or slightly beyond basal one-fourth of elytra, with 3.5 antennomeres in females and 4.5 antennomeres in males extended beyond basal pronotal margin, antennomeres slightly broadened, antennomere 3 slightly longer than antennomeres 2 or 4.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC65D5FE6371ED" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">Pronotum. Shape narrowly cordiform, only slightly transverse (ratio PW/PL = 1.1), widest at the anterior one-fourth, markedly narrowed posteriorly, lateral margins distinctly sinuate anterior to basal angles, the latter extended posteriorly as slender, pointed, digitiform processes. Disc markedly convex, globulose, smooth and glabrous, with median longitudinal impression very faintly impressed or effaced, basal fovea small and smooth; median basal area smooth; basal margin slightly convex and rounded. Lateral border of pronotum slender, distinctly defined only in anteri- or one-third to one-half, effaced in posterior one-half to two thirds. Single midlateral setae on each side inserted at anterior one-third; single basolateral seta on each side, inserted on basal angle at base of digitform process.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC60AAFEE776E4" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">Elytra. Elytral silhouette slightly tear-shaped, with humeri effaced, disc markedly convex, smooth and glabrous. All discal striae effaced except for faintly impressed parascutellar striole and a short, faintly impressed recurrent stria. Basal setiferous pore present. Three discal setae present, aligned along presumed track of stria 3. Preapicale seta absent, one or two apicoangular setules present. Umbilicate setal series with setae of humeral group equidistance from each other, with the first slightly more medially inserted than the others, setae of median group inserted distinctly posterior to middle.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC61CCFBFB76A5" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">Legs. Moderately long but slender. Protibiae with longitudinal furrow, sparely pubescent apically on anterior surface. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC618CFBE07565" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, females with two pairs.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC624CFC797526" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFF89FF953DB8624CFD847505" box="[495,586,573,597]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="28" pageNumber="369">Fig. 12b</figureCitation>
) large, elongate, broadest at mid-shaft, with apex long and recurved dorsally; endophallus with internal sclerites acuminate apically.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF89FF953CFC620FFC5F712D" pageId="28" pageNumber="369" type="discussion">
<paragraph id="8BC0368BFF89FF953CFC620FFD917407" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
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<emphasis id="B90BEA99FF89FF953CFC620FFEE675C4" bold="true" box="[171,296,638,662]" pageId="28" pageNumber="369">COMMENTS</emphasis>
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.— Based on similarities in the form of the aedeagus of males, this species is closely related to
<taxonomicName id="4C7F4D08FF89FF953D5362EFFDBE75E6" authority="Deuve" authorityName="Deuve" box="[260,624,670,694]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953D5362EFFDEC75E6" box="[260,546,670,694]" italics="true" pageId="28" pageNumber="369">Queinnectrechus jiuhecola</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh (
<bibRefCitation id="EFEE4B7AFF89FF953F4B62EFFC2E75E6" author="DEUVE, T. &amp; D. KAVANAUGH &amp; H. B. LIANG" box="[796,992,670,694]" pageId="28" pageNumber="369" pagination="171 - 178" refId="ref59155" refString="DEUVE, T., D. KAVANAUGH, AND H. B. LIANG. 2015. Trois Trechini nouveaux du Mont Laojun, pres de Lijiang, dans le Yunnan, Chine (Coleoptera, Caraboidea). Coleopteres 21: 171 - 178." type="journal article" year="2015">Deuve et al. 2015</bibRefCitation>
), described from Lijiang County, northwestern
<collectingRegion id="49BBF869FF89FF953D9562CEFDD97586" box="[450,535,703,726]" country="China" name="Yunnan" pageId="28" pageNumber="369">Yunnan</collectingRegion>
. However, members of this new species are distinguished from those of the latter by their much smaller size (BL =
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in
<taxonomicName id="4C7F4D08FF89FF953FCE62AEFBD375A7" box="[921,1053,735,759]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953FCE62AEFBD375A7" box="[921,1053,735,759]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
members), more elongate body form, more convex elytra, discal striae fully effaced (at least three medial striae evident in
<taxonomicName id="4C7F4D08FF89FF953D45636EFE567467" box="[274,408,799,823]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953D45636EFE567467" box="[274,408,799,823]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
members) and, most significantly, preapical seta absent (present near stria 2 subapically in
<taxonomicName id="4C7F4D08FF89FF953D366331FE287407" box="[353,486,831,855]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953D366331FE287407" box="[353,486,831,855]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
members).
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC6311FEB573C9" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
In a recent paper, Belousov and Kabak (2016) established a new genus,
<taxonomicName id="4C7F4D08FF89FF953F9F6311FBE97427" authorityName="Belousov and Kabak" authorityYear="2016" box="[968,1063,864,887]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF953F9F6311FBE97427" box="[968,1063,864,887]" italics="true" pageId="28" pageNumber="369">Uenoites</emphasis>
</taxonomicName>
, in which they placed
<taxonomicName id="4C7F4D08FF89FF953CA963F1FE4C74C8" box="[254,386,896,920]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953CA963F1FE4C74C8" box="[254,386,896,920]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
, as well as three other species previously included in
<taxonomicName id="4C7F4D08FF89FF953F9C63F1FBF474C8" authorityName="Jeannel" authorityYear="1935" box="[971,1082,896,920]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF953F9C63F1FBF474C8" box="[971,1082,896,920]" italics="true" pageId="28" pageNumber="369">Kozlovites</emphasis>
</taxonomicName>
or
<taxonomicName id="4C7F4D08FF89FF95383763F1FEB174E8" authority="Ueno (1995)" authorityName="Ueno" authorityYear="1995" class="Insecta" family="Carabidae" genus="Deuveotrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF95383763F1FF2074E8" italics="true" pageId="28" pageNumber="369">Deuveotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF89FF953CA263D1FEB174E8" author="UENO, S. I." box="[245,383,928,952]" pageId="28" pageNumber="369" pagination="13 - 25" refId="ref61208" refString="UENO, S. I. 1995. The Trechinae (Coleoptera, Carabidae) from Northern Vietnam. I. Two new species from Mt. Tam Dao. Bulletin of the National Science Museum, Ser. A (Zool.) 21: 13 - 25." type="journal article" year="1995">Uéno (1995)</bibRefCitation>
</taxonomicName>
, based on the presence of a preapical seta, more convex tempora, and the median lobe of the male aedeagus not markedly hooked apically. However, all these features are plesiomorphic among trechines, which is problematic for demonstrating phylogenetic affinity of the included taxa. Clearly, phylogenetic relationships among these groups of species remain unresolved. As noted above, the genitalia of
<taxonomicName id="4C7F4D08FF89FF953E716450FD627369" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[550,684,1057,1081]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF89FF953E716450FD627369" box="[550,684,1057,1081]" italics="true" pageId="28" pageNumber="369">Q. griswoldi</emphasis>
</taxonomicName>
males are very similar to those of
<taxonomicName id="4C7F4D08FF89FF95384A6450FF5E7309" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF95384A6450FF5E7309" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
males, wherease those of the new species described below,
<taxonomicName id="4C7F4D08FF89FF953F776430FBA17309" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[800,1135,1089,1113]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953F776430FBA17309" box="[800,1135,1089,1113]" italics="true" pageId="28" pageNumber="369">Queinnectrechus gongshanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF9538206433FF657329" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
, are of a very different form, although both of the new species described here are members of genus
<taxonomicName id="4C7F4D08FF89FF953C9764F0FEBA73C9" authorityName="Deuve" authorityYear="1992" box="[192,372,1153,1177]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF953C9764F0FEBA73C9" box="[192,372,1153,1177]" italics="true" pageId="28" pageNumber="369">Queinnectrechus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC64D3FBA672CB" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953CFC64D3FE6A73E8" bold="true" box="[171,420,1186,1210]" pageId="28" pageNumber="369">
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<smallCapsWord id="8D26A057FF89FF953D4764D4FE6A73E8" baselines="1203" box="[272,420,1189,1208]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="28" pageNumber="369">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in a variety of microhabitats in the alpine zone near the crest of the Gaoligong
<collectingRegion id="49BBF869FF89FF953E8D64B3FCC1738A" box="[730,783,1218,1242]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">Shan</collectingRegion>
and the eastern slope, at elevations ranging from
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. Specimens were collected under stones in moist meadows and on tundra slopes and ridges with sparse to thick herbaceous vegetation (
<figureCitation id="13442A0EFF89FF953F346573FC05724A" box="[867,971,1282,1306]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="28" pageNumber="369">Figs. 37b</figureCitation>
,
<figureCitation id="13442A0EFF89FF953F806572FBF8724A" box="[983,1078,1282,1307]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="28" pageNumber="369">38a, 38b</figureCitation>
,
<figureCitation id="13442A0EFF89FF9538146573FBA0724A" box="[1091,1134,1282,1306]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="28" pageNumber="369">39b</figureCitation>
), at the edges of small streams and seeps from talus slopes, and at the edges of
<taxonomicName id="4C7F4D08FF89FF953FC96552FBF2726B" box="[926,1084,1315,1339]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="28" pageNumber="369" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FF89FF953FC96552FBF2726B" box="[926,1084,1315,1339]" italics="true" pageId="28" pageNumber="369">Rhododendron</emphasis>
</taxonomicName>
thickets up to
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tall. Members of this species have been found syntopic with specimens of
<taxonomicName id="4C7F4D08FF89FF9538716532FE16722B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="balli" status="sp. nov." subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FF89FF9538716532FF07722B" italics="true" pageId="28" pageNumber="369">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF89FF953C8F6512FE56722B" box="[216,408,1379,1403]" italics="true" pageId="28" pageNumber="369">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FF89FF953DF06512FE16722B" box="[423,472,1379,1403]" italics="true" pageId="28" pageNumber="369">balli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953D886515FDFA722B" box="[479,564,1380,1403]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF89FF953E696512FC12722B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[574,988,1379,1403]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953E696512FD3C722B" box="[574,754,1379,1403]" italics="true" pageId="28" pageNumber="369">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF89FF953F556515FCF4722B" box="[770,826,1380,1403]" italics="true" pageId="28" pageNumber="369">s. str.</emphasis>
)
<emphasis id="B90BEA99FF89FF953F1E6512FC12722B" box="[841,988,1379,1403]" italics="true" pageId="28" pageNumber="369">gongshanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953FB56515FBF9722B" box="[994,1079,1380,1403]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF89FF9538156512FED672CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FF89FF9538156512FED672CB" italics="true" pageId="28" pageNumber="369">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953D4865F5FEBB72CB" box="[287,373,1412,1435]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF89FF953DD765F2FDF772CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[384,569,1411,1435]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="qiqiensis" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953DD765F2FDF772CB" box="[384,569,1411,1435]" italics="true" pageId="28" pageNumber="369">Trechus qiqiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953E1665F5FD5A72CB" box="[577,660,1412,1435]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C7F4D08FF89FF953E9F65F2FBC372CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[712,1037,1411,1435]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="monochaeta" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953E9F65F2FBC372CB" box="[712,1037,1411,1435]" italics="true" pageId="28" pageNumber="369">Trechepaphiopsis monochaeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF95384265F5FBA672CB" box="[1045,1128,1412,1435]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC65D2FF04716D" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953CFC65D2FD4E72EA" bold="true" box="[171,640,1443,1467]" pageId="28" pageNumber="369">
<smallCapsWord id="8D26A057FF89FF953CFC65D2FE9872EA" baselines="1461,1461" box="[171,342,1443,1467]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="28" pageNumber="369">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953D0865D6FE3D72EA" baselines="1461" box="[351,499,1447,1466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="28" pageNumber="369">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953DAA65D6FD8272EA" baselines="1461" box="[509,588,1447,1466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="28" pageNumber="369">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953E0165D6FD4E72EA" baselines="1461" box="[598,640,1447,1466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="28" pageNumber="369">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FF89FF953EDD65D2FCC172EA" baselines="1461,1461" box="[650,783,1443,1467]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953EDD65D2FCC172EA" bold="true" box="[650,783,1443,1467]" pageId="28" pageNumber="369">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FF89FF953F4E65D2FB9D728C" location="We" pageId="28" pageNumber="369" specimenCount="42" specimenCount-female="11" specimenCount-male="10" stateProvince="Shan">
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<emphasis id="B90BEA99FF89FF953F4E65D2FC9A72EA" bold="true" box="[793,852,1443,1467]" pageId="28" pageNumber="369">
<collectingRegion id="49BBF869FF89FF953F4E65D2FC9A72EA" box="[793,852,1443,1467]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">SHAN</collectingRegion>
</emphasis>
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.—
<figureCitation id="13442A0EFF89FF953FD765D5FC1072EC" box="[896,990,1444,1468]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="28" pageNumber="369">Fig. 12c</figureCitation>
.
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examined a total of
<specimenCount id="9D79FD02FF89FF953C9965B5FE90728C" box="[206,350,1476,1500]" count="21" pageId="28" pageNumber="369" type="generic">21 specimens</specimenCount>
(
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and
<specimenCount id="9D79FD02FF89FF953E5665B5FDB6728C" box="[513,632,1476,1500]" count="11" pageId="28" pageNumber="369" type="female">11 females</specimenCount>
), all from the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FF89FF95380D65B5FC31714C" county="Gongshan County" location="Cikai Townships" municipality="Bingzhongluo" pageId="28" pageNumber="369" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF89FF95380D65B5FB5C728C" box="[1114,1170,1476,1500]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">Shan</collectingRegion>
, in
<collectingMunicipality id="6BA4ACF1FF89FF953CC06595FEE072AC" box="[151,302,1508,1532]" pageId="28" pageNumber="369">Bingzhongluo</collectingMunicipality>
and
<location id="8EA06050FF89FF953D336595FDD572AC" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF89FF953D336595FDD572AC" box="[356,539,1508,1532]" county="Gongshan County" municipality="Bingzhongluo" name="Cikai Townships" pageId="28" pageNumber="369" stateProvince="Shan">Cikai Townships</location>
,
<collectingCounty id="62A14E07FF89FF953E716595FD2772AC" box="[550,745,1508,1532]" pageId="28" pageNumber="369">Gongshan County</collectingCounty>
(see
<typeStatus id="54C48829FF89FF953F756594FC9972AD" box="[802,855,1509,1533]" pageId="28" pageNumber="369">Type</typeStatus>
material above for exact collection data). These localities are all on the crest or eastern slope of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FF89FF9538506675FF08716D" location="Core Area" pageId="28" pageNumber="369" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF89FF9538506675FBF2714C" box="[1031,1084,1540,1564]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">Shan</collectingRegion>
in
<location id="8EA06050FF89FF9538356675FF61716D" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF89FF9538356675FF61716D" name="Core Area" pageId="28" pageNumber="369" stateProvince="Shan">Core Area</location>
2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC6635FC5F712D" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953CFC6635FDAF710B" bold="true" box="[171,609,1604,1628]" pageId="28" pageNumber="369">
<smallCapsWord id="8D26A057FF89FF953CFC6635FEDC710B" baselines="1622,1622" box="[171,274,1604,1628]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="28" pageNumber="369">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953D4A6639FE0A710B" baselines="1622" box="[285,452,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="28" pageNumber="369">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953D9A6639FDAF710B" baselines="1622" box="[461,609,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="28" pageNumber="369">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FF89FF953ED36614FCF1712D" box="[644,831,1637,1661]" country="China" name="Yunnan" pageId="28" pageNumber="369">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF89FF953F1D6614FC43712D" box="[842,909,1637,1661]" name="China" pageId="28" pageNumber="369">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,194 @@
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<mods:title id="61BB2B74664EF635435DA8580788856C">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="BB3B1AA4A6D09E366F1F65CE7D65E182">Deuve, Thierry</mods:namePart>
<mods:affiliation id="2FB34DCAEE4C98750CDF628541C4086C">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="A8B055855495E8B33AF947A9566E6575">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="ED05F5773DAD53A42CB6CD09BBDF1471">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="9F2C19464D3896CEB97D9A37C0ADC2F8">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<paragraph id="8BC0368BFF8DFF913DEE6373FC99744B" blockId="24.[441,855,770,795]" box="[441,855,770,795]" pageId="24" pageNumber="365">
<heading id="D08881E7FF8DFF913DEE6373FC99744B" box="[441,855,770,795]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="365" reason="2">
<emphasis id="B90BEA99FF8DFF913DEE6373FC99744B" bold="true" box="[441,855,770,795]" pageId="24" pageNumber="365">
Genus
<taxonomicName id="4C7F4D08FF8DFF913E5E6373FC99744B" ID-CoL="8KYP9" authority="Deuve, 1992" authorityName="Deuve" authorityYear="1992" box="[521,855,770,795]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913E5E6373FD0B744A" bold="true" box="[521,709,770,794]" italics="true" pageId="24" pageNumber="365">Queinnectrechus</emphasis>
Deuve, 1992
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8DFF913D8F6341FCF77416" box="[472,825,816,838]" pageId="24" pageNumber="365" type="reference_group">
<paragraph id="8BC0368BFF8DFF913D8F6341FCF77416" blockId="24.[472,825,816,838]" box="[472,825,816,838]" pageId="24" pageNumber="365">
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<treatmentCitation id="0ADE109AFF8DFF913D8F6341FCFB7416" author="DEUVE, T." box="[472,821,816,838]" page="354" pageId="24" pageNumber="365" year="1992">
<taxonomicName id="4C7F4D08FF8DFF913D8F6341FCFB7416" ID-CoL="8KYP9" authority="Deuve, 1992 b: 354" authorityName="Deuve" authorityPageNumber="354" authorityYear="1992" box="[472,821,816,838]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913D8F6341FDB47415" box="[472,634,816,837]" italics="true" pageId="24" pageNumber="365">Queinnectrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF8DFF913ED76341FCFB7416" author="DEUVE, T." box="[640,821,816,838]" pageId="24" pageNumber="365" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve, 1992b:354</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8DFF913CFC632BFCF77422" box="[171,825,858,882]" pageId="24" pageNumber="365" type="type_taxon">
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<emphasis id="B90BEA99FF8DFF913CFC632BFEF17420" bold="true" box="[171,319,858,882]" pageId="24" pageNumber="365">
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<typeStatus id="54C48829FF8DFF913CFC632BFF2A7420" box="[171,228,858,882]" pageId="24" pageNumber="365">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FF8DFF913CBB632CFEF17420" baselines="875" box="[236,319,861,880]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="24" pageNumber="365">SPECIES</smallCapsWord>
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.—
<taxonomicName id="4C7F4D08FF8DFF913D3F632BFCF77422" ID-CoL="8MD2H" authority="Deuve, 1992 a" authorityName="Deuve" authorityYear="1992" box="[360,825,858,882]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="species" species="excentricus">
<emphasis id="B90BEA99FF8DFF913D3F632BFD527421" box="[360,668,858,882]" italics="true" pageId="24" pageNumber="365">Queinnectrechus excentricus</emphasis>
<bibRefCitation id="EFEE4B7AFF8DFF913EF3632BFCF77422" author="DEUVE, T." box="[676,825,858,882]" pageId="24" pageNumber="365" refId="ref58815" refString="DEUVE, T. 1992 a. Un nouveau genre de Trechinae des montagnes du Sichuan (Coleoptera, Trechidae). Bulletin de la Societe Entomologique de France 96 [1991]: 354." type="journal volume" year="1992">Deuve, 1992a</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8DFF913CFC630BFD0073A7" pageId="24" pageNumber="365" type="diagnosis">
<paragraph id="8BC0368BFF8DFF913CFC630BFD0073A7" blockId="24.[123,1174,858,1725]" pageId="24" pageNumber="365">
<smallCapsWord id="8D26A057FF8DFF913CFC630BFEEE74C1" baselines="908,908" box="[171,288,890,914]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="24" pageNumber="365">
<emphasis id="B90BEA99FF8DFF913CFC630BFEEE74C1" bold="true" box="[171,288,890,914]" pageId="24" pageNumber="365">DIAGNOSIS</emphasis>
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.— Adults of this genus (
<figureCitation id="13442A0EFF8DFF913E65630BFD7D74C2" box="[562,691,890,914]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="26.[171,183,1585,1604]" captionStartId-1="30.[171,183,1582,1601]" captionStartId-2="33.[171,183,1577,1596]" captionTargetBox-0="[147,1176,216,1569]" captionTargetBox-1="[120,1176,221,1566]" captionTargetBox-2="[144,1175,216,1563]" captionTargetId-0="figure-11@26.[120,1176,216,1569]" captionTargetId-1="figure-11@30.[120,1176,221,1566]" captionTargetId-2="figure-15@33.[120,1175,216,1563]" captionTargetPageId-0="26" captionTargetPageId-1="30" captionTargetPageId-2="33" captionText-0="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." captionText-1="FIGURE 13. Queinnectrechus (s. str.) gongshanicus sp. nov.; a. Dorsal habitus (CASENT1024868). b. Median lobe of aedeagus of male (CASENT1024868), left lateral aspect. c. Map of locality records (red circles) for Q. gongshanicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." captionText-2="FIGURE 14. Queinnectrechus (Gaoligongtrechus) balli sp. nov.; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for Q. balli in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi-0="http://doi.org/10.5281/zenodo.13155305" figureDoi-1="http://doi.org/10.5281/zenodo.13155307" figureDoi-2="http://doi.org/10.5281/zenodo.13155309" httpUri-0="https://zenodo.org/record/13155305/files/figure.png" httpUri-1="https://zenodo.org/record/13155307/files/figure.png" httpUri-2="https://zenodo.org/record/13155309/files/figure.png" pageId="24" pageNumber="365">Figs. 1214</figureCitation>
) can be recognized by the following combination of character states: size moderate (BL =
<quantity id="4C879B6EFF8DFF913E2463EBFCC474E3" box="[627,778,922,947]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.8" metricValueMin="3.5" pageId="24" pageNumber="365" unit="mm" value="4.15" valueMax="4.8" valueMin="3.5">3.5 to 4.8 mm</quantity>
), apterous, body dark, reddish brown to piceous, surface micaceous; head with small eyes, mentum and submentum fused; right mandible tridentate, the premolar tooth distinct but joined with the retinaculum (see
<bibRefCitation id="EFEE4B7AFF8DFF91385763ADFB5C74A3" author="DEUVE, T." box="[1024,1170,987,1011]" pageId="24" pageNumber="365" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve 1992b</bibRefCitation>
,
<figureCitation id="13442A0EFF8DFF913C2C638DFF057344" box="[123,203,1020,1044]" captionStart="FIGURE" captionStartId="60.[171,183,1578,1597]" captionTargetBox="[132,1176,216,1563]" captionTargetId="figure-12@60.[120,1176,216,1563]" captionTargetPageId="60" captionText="FIGURE 23. Trechus gongshanensis sp. nov.; a. Dorsal habitus (CASENT1001929). b. Median lobe of aedeagus of male (CASENT1001929, left lateral aspect. c. Map of locality record s(red circles) for T. gongshanensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155328" httpUri="https://zenodo.org/record/13155328/files/figure.png" pageId="24" pageNumber="365">Fig. 23</figureCitation>
); pronotum cordiform and markedly convex, lateral margination effaced posteriorly, with two setae (midlateral and basolateral) present on each side; elytra markedly convex, inflated, slightly tear-shaped, humeri effaced, elytral discal striae absent or vestigial, with two (in most members) or three (in a few members) discal setae present, aligned on interval 3 near stria 3, preapical seta absent from most members, in some of these members inserted forward in a subdiscal position, in very few placed in typical trechine position nearer elytral apex and next to stria 2; protibiae furrowed; abdominal ventrites IV to VI glabrous except for a single pair of paramedial setae; endophallus of male aedeagus with two sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8DFF903CFC648EFE867644" lastPageId="25" lastPageNumber="366" pageId="24" pageNumber="365" type="discussion">
<paragraph id="8BC0368BFF8DFF913CFC648EFEB8710C" blockId="24.[123,1174,858,1725]" pageId="24" pageNumber="365">
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<emphasis id="B90BEA99FF8DFF913CFC648EFEE67246" bold="true" box="[171,296,1279,1303]" pageId="24" pageNumber="365">COMMENTS</emphasis>
</smallCapsWord>
.— Two new genera closely related to
<taxonomicName id="4C7F4D08FF8DFF913EAC6571FC617248" authorityName="Deuve" authorityYear="1992" box="[763,943,1280,1304]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913EAC6571FC617248" box="[763,943,1280,1304]" italics="true" pageId="24" pageNumber="365">Queinnectrechus</emphasis>
</taxonomicName>
have recently been described. Members of
<taxonomicName id="4C7F4D08FF8DFF913D216551FC877268" authority="Belousov and Kabak (2003)" authorityName="Belousov and Kabak" authorityYear="2003" box="[374,841,1312,1336]" class="Insecta" family="Carabidae" genus="Dactylotrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913D216551FDD67268" box="[374,536,1312,1336]" italics="true" pageId="24" pageNumber="365">Dactylotrechus</emphasis>
Belousov and Kabak (2003)
</taxonomicName>
are distinguished by the supernumerary setae present on the external margins of the pronotum and on more lateral areas of the elytral disc. Members of
<taxonomicName id="4C7F4D08FF8DFF913DDC6510FC817229" authority="Belousov and Kabak (2014 a)" authorityName="Belousov and Kabak" authorityYear="2014" box="[395,847,1377,1401]" class="Insecta" family="Carabidae" genus="Puertrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913DDC6510FDC27229" box="[395,524,1377,1401]" italics="true" pageId="24" pageNumber="365">Puertrechus</emphasis>
Belousov and Kabak (2014a)
</taxonomicName>
are distinguished by the presence of a single discal setae subbasally on interval 5. Members of both taxa have a preapical seta inserted near stria 2, a plesiomorphic feature among trechines. Taxonomic limits and phylogenetic relationships among the “genera”
<taxonomicName id="4C7F4D08FF8DFF913DBD65B2FD97728A" authorityName="Jeannel" authorityYear="1923" box="[490,601,1475,1498]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913DBD65B2FD97728A" box="[490,601,1475,1498]" italics="true" pageId="24" pageNumber="365">Stevensius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FF8DFF913E3065B3FCB1728A" authority="Jeannel (1935)" authorityName="Jeannel" authorityYear="1935" box="[615,895,1474,1498]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913E3065B3FD18728A" box="[615,726,1474,1498]" italics="true" pageId="24" pageNumber="365">Kozlovites</emphasis>
<bibRefCitation id="EFEE4B7AFF8DFF913E8965B3FCB1728A" author="JEANNEL, R." box="[734,895,1474,1498]" pageId="24" pageNumber="365" pagination="273 - 282" refId="ref59618" refString="JEANNEL, R. 1935. Sur quelques Trechinae de l'Asie Centrale. Revue Francaise d'Entomologie 1: 273 - 282." type="journal article" year="1935">Jeannel (1935)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FF8DFF913FDC65B3FBF1728A" authorityName="Deuve" authorityYear="1992" box="[907,1087,1474,1498]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913FDC65B3FBF1728A" box="[907,1087,1474,1498]" italics="true" pageId="24" pageNumber="365">Queinnectrechus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FF8DFF91381A65B2FF1372AB" authorityName="Belousov and Kabak" authorityYear="2003" class="Insecta" family="Carabidae" genus="Dactylotrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF91381A65B2FF1372AB" italics="true" pageId="24" pageNumber="365">Dactylotrechus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FF8DFF913CB96592FEA172AB" authorityName="Belousov and Kabak" authorityYear="2014" box="[238,367,1507,1531]" class="Insecta" family="Carabidae" genus="Puertrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913CB96592FEA172AB" box="[238,367,1507,1531]" italics="true" pageId="24" pageNumber="365">Puertrechus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FF8DFF913DD66592FD7A72AB" authority="Ueno (2000)" authorityName="Ueno" authorityYear="2000" box="[385,692,1507,1531]" class="Insecta" family="Carabidae" genus="Sinotrechiama" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913DD66592FDD272AB" box="[385,540,1507,1531]" italics="true" pageId="24" pageNumber="365">Sinotrechiama</emphasis>
Uéno (2000)
</taxonomicName>
,
<emphasis id="B90BEA99FF8DFF913E946592FCD972AA" box="[707,791,1507,1530]" italics="true" pageId="24" pageNumber="365">Uenoite</emphasis>
s Belousov and Kabak (2016) and
<taxonomicName id="4C7F4D08FF8DFF913C2C6672FED8714B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,278,1539,1563]" class="Insecta" family="Carabidae" genus="Minutotrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF8DFF913C2C6672FED8714B" box="[123,278,1539,1563]" italics="true" pageId="24" pageNumber="365">Minutotrechus</emphasis>
</taxonomicName>
(describe above) are still poorly resolved. This is why we describe below a new taxon,
<taxonomicName id="4C7F4D08FF8DFF913C946655FE4D716C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[195,387,1572,1596]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="365" phylum="Arthropoda" rank="subGenus" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FF8DFF913C946655FE4D716C" box="[195,387,1572,1596]" italics="true" pageId="24" pageNumber="365">Gaoligongtrechus</emphasis>
</taxonomicName>
, provisionally with the rank of subgenus. This assignment can be changed as needed in the future.
</paragraph>
<paragraph id="8BC0368BFF8DFF903CFC6615FE867644" blockId="24.[123,1174,858,1725]" lastBlockId="25.[123,1173,219,276]" lastPageId="25" lastPageNumber="366" pageId="24" pageNumber="365">
<emphasis id="B90BEA99FF8DFF913CFC6615FE3D712B" bold="true" box="[171,499,1636,1660]" pageId="24" pageNumber="365">
<smallCapsWord id="8D26A057FF8DFF913CFC6615FE98712B" baselines="1654,1654" box="[171,342,1636,1660]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="24" pageNumber="365">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF8DFF913D086619FE3D712B" baselines="1654" box="[351,499,1640,1659]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="24" pageNumber="365">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This genus, which currently include 13 described species (Belousov and Kabak 2003, Casalle and Magrini 2009,
<bibRefCitation id="EFEE4B7AFF8DFF913E9B66F4FC9271CD" author="DEUVE, T." box="[716,860,1669,1693]" pageId="24" pageNumber="365" refId="ref58815" refString="DEUVE, T. 1992 a. Un nouveau genre de Trechinae des montagnes du Sichuan (Coleoptera, Trechidae). Bulletin de la Societe Entomologique de France 96 [1991]: 354." type="journal volume" year="1992">Deuve 1992a</bibRefCitation>
, and
<bibRefCitation id="EFEE4B7AFF8DFF913FC366F4FBD871CD" author="UENO, S. I." box="[916,1046,1669,1693]" pageId="24" pageNumber="365" pagination="61 - 68" refId="ref61351" refString="UENO, S. I. 1998 a. The trechine beetles (Coleoptera, Trechinae) from the Zhongdian area in northwestern Yunnan mainly collected by Ales Smetana. Elytra 26: 61 - 68." type="journal article" year="1998">Uéno 1998a</bibRefCitation>
and
<bibRefCitation id="EFEE4B7AFF8DFF91381D66F4FB4071CD" author="UENO, S. I." box="[1098,1166,1669,1693]" pageId="24" pageNumber="365" pagination="263 - 287" refId="ref61388" refString="UENO, S. I. 1998 b. The Trechinae (Coleoptera) from Mt. Gongga Shan and its vicinities, southwest China, with notes on the Epaphiopsis from Mt. Emei Shan. Elytra 26: 263 - 287." type="journal article" year="1998">1998b</bibRefCitation>
) is currently known from the Min
<collectingRegion id="49BBF869FF8DFF913DB266D4FDA971ED" box="[485,615,1701,1725]" country="Myanmar" name="Shan" pageId="24" pageNumber="365">Shan region</collectingRegion>
of northern
<collectingRegion id="49BBF869FF8DFF913EB866D4FC6271ED" box="[751,940,1701,1725]" country="China" name="Sichuan" pageId="24" pageNumber="365">Sichuan Province</collectingRegion>
southwest to western
<collectingRegion id="49BBF869FF8CFF903C2C60AAFEF977A3" box="[123,311,219,243]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan Province</collectingRegion>
. The Gaoligong
<collectingRegion id="49BBF869FF8CFF903DB960AAFDED77A3" box="[494,547,219,243]" country="Myanmar" name="Shan" pageId="25" pageNumber="366">Shan</collectingRegion>
forms the southwestern limit of the known distributional range of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,453 @@
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<mods:title id="0F4C4B082CE7E1F725530744177EE999">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="98DC3B21C63B5EF8B7F9B23210A6260C">Deuve, Thierry</mods:namePart>
<mods:affiliation id="18595DD1A812996BCE55B7EA933692C1">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="402135500A6F8F5605E99278542291B8">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="BA93F24C0FC67B882298D2A15D30A9C2">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="01088616A7B4E00C9B6CF64B9F26CA8E" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:namePart id="34E46F65E0EE4D1C33553C1C8B991274">Liang, Hongbin</mods:namePart>
<mods:affiliation id="F8CA5B4AEE908E849CBB2FFE35CAA68E">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FF92FF8E3C2C65A9FD4072A0" bold="true" box="[123,654,1496,1520]" pageId="7" pageNumber="348">
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<emphasis id="B90BEA99FF92FF8E3C2C65A9FEA872A0" bold="true" box="[123,358,1496,1520]" italics="true" pageId="7" pageNumber="348">Perileptus pusilloides</emphasis>
Deuve and Liang
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF92FF8E3E6F65A9FD4072A0" box="[568,654,1496,1520]" pageId="7" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3656500FF92FF8E3C2C6588FED8715F" box="[123,278,1529,1551]" pageId="7" pageNumber="348" type="description">
<paragraph id="8BC0368BFF92FF8E3C2C6588FED8715F" blockId="7.[123,654,1496,1551]" box="[123,278,1529,1551]" pageId="7" pageNumber="348">
(
<figureCitation id="13442A0EFF92FF8E3CD56588FF09715F" box="[130,199,1529,1551]" captionStart="FIGURE" captionStartId="10.[171,183,1609,1628]" captionTargetBox="[144,1176,216,1593]" captionTargetId="figure-11@10.[120,1176,216,1593]" captionTargetPageId="10" captionText="FIGURE 6. Perileptus pusilloides sp. nov.; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for P. pusilloides in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="7" pageNumber="348">Figs. 6</figureCitation>
,
<figureCitation id="13442A0EFF92FF8E3C866588FEC1715F" box="[209,271,1529,1551]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="7" pageNumber="348">4648</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="B90BEA99FF92FF8E3CFC6652FE947169" bold="true" box="[171,346,1571,1595]" pageId="7" pageNumber="348">
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.—
<materialsCitation id="3B173CD6FF92FF8E3DD46652FE6571ED" collectingDate="2006-06-02" collectorName="R. L. Brett &amp; D. Z. Dong" country="China" county="Tengchong County" elevation="1105" latitude="24.92346" location="Dengman village" longLatPrecision="1" longitude="98.38612" municipality="Hehua Township" pageId="7" pageNumber="348" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FF92FF8E3DD46652FE27716B" box="[387,489,1571,1595]" pageId="7" pageNumber="348" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1039259”/ “
<collectingCountry id="F368761BFF92FF8E3FAA6652FB98716B" box="[1021,1110,1571,1595]" name="China" pageId="7" pageNumber="348">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FF92FF8E3CE76632FEB0710C" box="[176,382,1603,1628]" pageId="7" pageNumber="348">Tengchong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF92FF8E3DDD6632FD8D710B" box="[394,579,1603,1627]" pageId="7" pageNumber="348">Hehua Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FF92FF8E3E186632FCD1710B" box="[591,799,1603,1628]" location="Hehua" pageId="7" pageNumber="348">
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S of Hehua
</locationDeviation>
at
<location id="8EA06050FF92FF8E3F156635FC35710B" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF92FF8E3F156635FC35710B" box="[834,1019,1603,1628]" country="China" county="Tengchong County" latitude="24.92346" longLatPrecision="1" longitude="98.38612" municipality="Hehua Township" name="Dengman village" pageId="7" pageNumber="348" stateProvince="Yunnan">Dengman village</location>
along
<location id="8EA06050FF92FF8E38106632FF78712C" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF92FF8E38106632FF78712C" country="China" county="Tengchong County" latitude="24.92346" longLatPrecision="1" longitude="98.38612" municipality="Hehua Township" name="Daying Jiang" pageId="7" pageNumber="348" stateProvince="Yunnan">Daying Jiang</location>
,
<geoCoordinate id="EE4B504CFF92FF8E3C976615FE8D712C" box="[192,323,1636,1660]" degrees="24.92346" direction="north" orientation="latitude" pageId="7" pageNumber="348" precision="1" value="24.92346">N24.92346°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF92FF8E3D1B6615FE02712C" box="[332,460,1636,1660]" degrees="98.38612" direction="east" orientation="longitude" pageId="7" pageNumber="348" precision="1" value="98.38612">E98.38612°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFF92FF8E3DA56615FD8B712C" box="[498,581,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="7" pageNumber="348" unit="m" value="1105.0">
<elevation id="0052D1B8FF92FF8E3DA56615FD8B712C" box="[498,581,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="7" pageNumber="348" unit="m" value="1105.0">1105 m</elevation>
</quantity>
,
<date id="FFC1104BFF92FF8E3E186615FD1C712C" box="[591,722,1636,1660]" pageId="7" pageNumber="348" value="2006-06-02">
<collectingDate id="EF85E9A3FF92FF8E3E186615FD1C712C" box="[591,722,1636,1660]" pageId="7" pageNumber="348" value="2006-06-02">2 June 2006</collectingDate>
</date>
, Stop # DHK-2006-053, D.H.
<taxonomicName id="4C7F4D08FF92FF8E38426615FB5F712C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1045,1169,1636,1660]" pageId="7" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFF92FF8E3C2C66F4FF2871CC" box="[123,230,1669,1692]" pageId="7" pageNumber="348">R.L. Brett</collectorName>
&amp;
<collectorName id="268A535DFF92FF8E3D5F66F4FEB571CD" box="[264,379,1669,1693]" pageId="7" pageNumber="348">D.Z. Dong</collectorName>
collectors”/ “
<typeStatus id="54C48829FF92FF8E3E5B66F4FD5571CD" box="[524,667,1669,1693]" pageId="7" pageNumber="348" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FF92FF8E3EF566F4FBF771CD" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[674,1081,1669,1693]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="pusilloides" status="sp. nov.">
<emphasis id="B90BEA99FF92FF8E3EF566F4FC4471CD" box="[674,906,1669,1693]" italics="true" pageId="7" pageNumber="348">Perileptus pusilloides</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FF92FF8E381366F4FB5871CC" box="[1092,1174,1669,1692]" pageId="7" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FF92FF8E3DE166D7FB5B71EE" box="[438,1173,1701,1726]" collectionCode="CAS, IOZ, MNHN" country="China" pageId="7" pageNumber="348" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FF92FF8E3DE166D7FDD071EE" box="[438,542,1702,1726]" pageId="7" pageNumber="348" type="paratype">Paratypes</typeStatus>
(a total of 4):
<specimenCount id="9D79FD02FF92FF8E3EE266D7FCC971ED" box="[693,775,1701,1726]" count="2" pageId="7" pageNumber="348" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FF92FF8E3F6D66D7FC5971ED" box="[826,919,1701,1726]" count="1" pageId="7" pageNumber="348" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF92FF8E3F9666D7FC3971EE" box="[961,1015,1702,1726]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="348" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF92FF8E385566D7FBFF71EE" box="[1026,1073,1702,1726]" pageId="7" pageNumber="348">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF92FF8E386C66D7FB4071EE" box="[1083,1166,1702,1726]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="348">MNHN</collectionCode>
)
</materialsCitation>
</paragraph>
<caption id="DF006603FF9DFF813CFC663CFE0671C3" ID-DOI="http://doi.org/10.5281/zenodo.13155291" ID-Zenodo-Dep="13155291" httpUri="https://zenodo.org/record/13155291/files/figure.png" pageId="8" pageNumber="349" startId="8.[171,183,1613,1632]" targetBox="[120,1176,224,1597]" targetPageId="8" targetType="figure">
<paragraph id="8BC0368BFF9DFF813CFC663CFE0671C3" blockId="8.[123,1175,1613,1683]" pageId="8" pageNumber="349">
<smallCapsWord id="8D26A057FF9DFF813CFC663CFF26710E" baselines="1627,1626" box="[171,232,1613,1632]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="8" pageNumber="349">FIGURE</smallCapsWord>
5.
<taxonomicName id="4C7F4D08FF9DFF813D50663CFE3A7130" authority="Jeannel" authorityName="Jeannel Body" authorityYear="1923" box="[263,500,1613,1632]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="imaicus">
<emphasis id="B90BEA99FF9DFF813D50663CFE67710F" box="[263,425,1613,1632]" italics="true" pageId="8" pageNumber="349">Perileptus imaicus</emphasis>
Jeannel
</taxonomicName>
; a. Dorsal habitus (CASENT1036287). b. Median lobe of aedeagus of male (CASENT10363020), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FF9DFF813F6B6616FC5C7129" authorityName="Jeannel Body" authorityYear="1923" box="[828,914,1639,1657]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="imaicus">
<emphasis id="B90BEA99FF9DFF813F6B6616FC5C7129" box="[828,914,1639,1657]" italics="true" pageId="8" pageNumber="349">P. imaicus</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF9CFF803C2C60AAFDFD76C4" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">
<materialsCitation id="3B173CD6FF9CFF803C2C60AAFDE176C4" collectingDate="2005-06-01" collectionCode="IOZ" collectorName="H. B. Liang" country="China" county="Longyang County" elevation="680" latitude="25.09065" location="Kunhong He" longLatPrecision="1" longitude="98.83721" municipality="Bawan Township" pageId="9" pageNumber="350" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
labeled same as
<typeStatus id="54C48829FF9CFF803D6460AAFE5E77A3" box="[307,400,219,243]" pageId="9" pageNumber="350" type="holotype">holotype</typeStatus>
except first label “CASENT 1039257” and “CASENT 1039260” and “CASENT 1039258”, respectively;
<specimenCount id="9D79FD02FF9CFF803E52608AFDA87643" box="[517,614,251,275]" count="1" pageId="9" pageNumber="350" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF9CFF803ECF608AFD087643" box="[664,710,251,275]" pageId="9" pageNumber="350">IOZ</collectionCode>
) labeled: “CASENT 1036294”/ “
<collectingCountry id="F368761BFF9CFF80386F608AFB5F7643" box="[1080,1169,251,275]" name="China" pageId="9" pageNumber="350">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF9CFF803C2C616DFF1D7663" box="[123,211,284,307]" country="China" name="Yunnan" pageId="9" pageNumber="350">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF9CFF803CBF616DFE737664" box="[232,445,284,308]" pageId="9" pageNumber="350">Longyang County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF9CFF803D84616DFD6F7663" box="[467,673,283,307]" pageId="9" pageNumber="350">Bawan Township</collectingMunicipality>
,
<location id="8EA06050FF9CFF803EE0616AFC9E7663" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF9CFF803EE0616AFC9E7663" box="[695,848,283,307]" country="China" county="Longyang County" latitude="25.09065" longLatPrecision="1" longitude="98.83721" municipality="Bawan Township" name="Kunhong He" pageId="9" pageNumber="350" stateProvince="Yunnan">Kunhong He</location>
at
<location id="8EA06050FF9CFF803FDF616AFC3F7663" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF9CFF803FDF616AFC3F7663" box="[904,1009,283,307]" country="China" county="Longyang County" latitude="25.09065" longLatPrecision="1" longitude="98.83721" municipality="Bawan Township" name="Bingmen" pageId="9" pageNumber="350" stateProvince="Yunnan">Bingmen</location>
,
<geoCoordinate id="EE4B504CFF9CFF803850616AFB427663" box="[1031,1164,283,307]" degrees="25.09065" direction="north" orientation="latitude" pageId="9" pageNumber="350" precision="1" value="25.09065">N25.09065°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF9CFF803C2C614DFF337604" box="[123,253,316,340]" degrees="98.83721" direction="east" orientation="longitude" pageId="9" pageNumber="350" precision="1" value="98.83721">E98.83721°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFF9CFF803D72614DFEA37603" box="[293,365,316,340]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="350" unit="m" value="680.0">
<elevation id="0052D1B8FF9CFF803D72614DFEA37603" box="[293,365,316,340]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="350" unit="m" value="680.0">680 m</elevation>
</quantity>
,
<date id="FFC1104BFF9CFF803D2E614DFE307603" box="[377,510,315,340]" pageId="9" pageNumber="350" value="2005-06-01">
<collectingDate id="EF85E9A3FF9CFF803D2E614DFE307603" box="[377,510,315,340]" pageId="9" pageNumber="350" value="2005-06-01">1 June 2005</collectingDate>
</date>
, Stop# 2005-039, D.H.
<taxonomicName id="4C7F4D08FF9CFF803EA8614DFCB67604" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[767,888,316,340]" pageId="9" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFF9CFF803FCB614DFBDD7604" box="[924,1043,316,340]" pageId="9" pageNumber="350">H.B. Liang</collectorName>
collectors”. All
<typeStatus id="54C48829FF9CFF803CF2612DFEC57624" box="[165,267,348,372]" pageId="9" pageNumber="350" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FF9CFF803E34612DFD247624" box="[611,746,348,372]" pageId="9" pageNumber="350" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FF9CFF803EA4612DFB5F7624" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[755,1169,348,372]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="pusilloides" status="sp. nov.">
<emphasis id="B90BEA99FF9CFF803EA4612DFC137624" box="[755,989,348,372]" italics="true" pageId="9" pageNumber="350">Perileptus pusilloides</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FF9CFF803C2C610CFF0076C4" box="[123,206,381,404]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF9CFF803CFC61EDFC6F7684" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">
<materialsCitation id="3B173CD6FF9CFF803CFC61EDFC537684" country="China" county="Tengchong County" elevation="1105" latitude="24.92346" location="Dengman village" longLatPrecision="1" longitude="98.38612" municipality="Hehua Township" pageId="9" pageNumber="350" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FF9CFF803CFC61EDFE9976E3" bold="true" box="[171,343,412,436]" pageId="9" pageNumber="350">
<smallCapsWord id="8D26A057FF9CFF803CFC61EDFF2A76E3" baselines="430,430" box="[171,228,412,436]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="9" pageNumber="350">
<typeStatus id="54C48829FF9CFF803CFC61EDFF2A76E3" box="[171,228,412,436]" pageId="9" pageNumber="350">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FF9CFF803CBB61D1FE9976E3" baselines="430" box="[236,343,416,435]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="9" pageNumber="350">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFF9CFF803DD761EDFE0C76E4" box="[384,450,412,436]" name="China" pageId="9" pageNumber="350">China</collectingCountry>
,
<collectingRegion id="49BBF869FF9CFF803D9A61ECFDEF76E4" box="[461,545,413,436]" country="China" name="Yunnan" pageId="9" pageNumber="350">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF9CFF803E7B61EDFD3476E4" box="[556,762,412,436]" pageId="9" pageNumber="350">Tengchong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF9CFF803F5261EDFC7076E4" box="[773,958,412,436]" pageId="9" pageNumber="350">Hehua Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FF9CFF803F9E61EDFB5876E4" box="[969,1174,412,436]" location="Hehua" pageId="9" pageNumber="350">
<quantity id="4C879B6EFF9CFF803F9E61EDFBDA76E4" box="[969,1044,412,436]" metricMagnitude="3" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="350" unit="km" value="5.4">5.4 km</quantity>
S of Hehua
</locationDeviation>
at
<location id="8EA06050FF9CFF803CC261CCFE837685" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF9CFF803CC261CCFE837685" box="[149,333,445,469]" country="China" county="Tengchong County" latitude="24.92346" longLatPrecision="1" longitude="98.38612" municipality="Hehua Township" name="Dengman village" pageId="9" pageNumber="350" stateProvince="Yunnan">Dengman village</location>
along
<location id="8EA06050FF9CFF803DC061CCFDE67685" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF9CFF803DC061CCFDE67685" box="[407,552,445,469]" country="China" county="Tengchong County" latitude="24.92346" longLatPrecision="1" longitude="98.38612" municipality="Hehua Township" name="Daying Jiang" pageId="9" pageNumber="350" stateProvince="Yunnan">Daying Jiang</location>
,
<geoCoordinate id="EE4B504CFF9CFF803E6461CDFD787684" box="[563,694,444,468]" degrees="24.92346" direction="north" orientation="latitude" pageId="9" pageNumber="350" precision="1" value="24.92346">N24.92346°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF9CFF803EE961CDFCF07684" box="[702,830,444,468]" degrees="98.38612" direction="east" orientation="longitude" pageId="9" pageNumber="350" precision="1" value="98.38612">E98.38612°</geoCoordinate>
,
<quantity id="4C879B6EFF9CFF803F1E61CDFC537684" box="[841,925,444,468]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="9" pageNumber="350" unit="m" value="1105.0">
<elevation id="0052D1B8FF9CFF803F1E61CDFC537684" box="[841,925,444,468]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="9" pageNumber="350" unit="m" value="1105.0">1105 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9CFF803CFC61ACFB9F7565" pageId="9" pageNumber="350" type="etymology">
<paragraph id="8BC0368BFF9CFF803CFC61ACFB9F7565" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">
<smallCapsWord id="8D26A057FF9CFF803CFC61ACFEFB76A3" baselines="495,494" box="[171,309,477,501]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Etymology" pageId="9" pageNumber="350">
<emphasis id="B90BEA99FF9CFF803CFC61ACFEFB76A3" bold="true" box="[171,309,477,501]" pageId="9" pageNumber="350">ETYMOLOGY</emphasis>
</smallCapsWord>
.— The species epithet,
<taxonomicName id="4C7F4D08FF9CFF803E1961ACFD0C76A5" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[590,706,477,501]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="pusilloides">
<emphasis id="B90BEA99FF9CFF803E1961ACFD0C76A5" box="[590,706,477,501]" italics="true" pageId="9" pageNumber="350">pusilloides</emphasis>
</taxonomicName>
, is a combination of the species epithet,
<emphasis id="B90BEA99FF9CFF803C2C618CFF1A7545" box="[123,212,509,533]" italics="true" pageId="9" pageNumber="350">
<taxonomicName id="4C7F4D08FF9CFF803C2C618CFF1F7545" box="[123,209,509,533]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="pusillus">pusillus</taxonomicName>
,
</emphasis>
and the Greek suffix,
<emphasis id="B90BEA99FF9CFF803D90618CFDC47545" box="[455,522,509,533]" italics="true" pageId="9" pageNumber="350">–ΕΙδής</emphasis>
(transliterated into Latin as
<emphasis id="B90BEA99FF9CFF803F6B618CFC4F7545" box="[828,897,509,533]" italics="true" pageId="9" pageNumber="350">oides</emphasis>
), meaning resembling, in reference to the similarity of members of this species to those of
<taxonomicName id="4C7F4D08FF9CFF803F61626FFB837565" authority="Jeannel (1923)" authorityName="Jeannel" authorityYear="1923" box="[822,1101,541,565]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9CFF803F61626FFC6A7565" box="[822,932,541,565]" italics="true" pageId="9" pageNumber="350">T. pusillus</emphasis>
<bibRefCitation id="EFEE4B7AFF9CFF803FFC626CFB837565" author="JEANNEL, R." box="[939,1101,541,565]" pageId="9" pageNumber="350" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel (1923)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9CFF803CFC624FFE8F7466" pageId="9" pageNumber="350" type="diagnosis">
<paragraph id="8BC0368BFF9CFF803CFC624FFE8F7466" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">
<smallCapsWord id="8D26A057FF9CFF803CFC624FFEEE7504" baselines="592,591" box="[171,288,574,598]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="9" pageNumber="350">
<emphasis id="B90BEA99FF9CFF803CFC624FFEEE7504" bold="true" box="[171,288,574,598]" pageId="9" pageNumber="350">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFF9CFF803E1B624CFD527505" box="[588,668,573,597]" captionStart="FIGURE" captionStartId="10.[171,183,1609,1628]" captionTargetBox="[144,1176,216,1593]" captionTargetId="figure-11@10.[120,1176,216,1593]" captionTargetPageId="10" captionText="FIGURE 6. Perileptus pusilloides sp. nov.; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for P. pusilloides in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="9" pageNumber="350">Fig. 6a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size very small (BL =
<quantity id="4C879B6EFF9CFF803823622FFF6675C6" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="350" unit="mm" value="2.3">2.3 mm</quantity>
), fully-winged; body color light yellowish brown, elytra concolorous throughout; microsculpture of elytra irregularly isodiametric and moderately impressed; dorsal pubescence dense and rather short; pronotum slightly transverse (ratio PW/PL = 1.32) only slightly convex, with lateral margins moderately rounded in anterior half, straightened just anterior to basal angles, the latter acute and sharp but small, median basal area rugulose; elytra alutaceous, distinctly flattened, striae finely punctate.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9CFF823CFC634EFBE077A3" lastPageId="11" lastPageNumber="352" pageId="9" pageNumber="350" type="description">
<paragraph id="8BC0368BFF9CFF803CFC634EFB9274C8" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">
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<emphasis id="B90BEA99FF9CFF803CFC634EFEF47406" bold="true" box="[171,314,831,855]" pageId="9" pageNumber="350">DESCRIPTION</emphasis>
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.— Size very small, BL =
<quantity id="4C879B6EFF9CFF803E02634EFD657407" box="[597,683,831,855]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="350" unit="mm" value="2.3">2.3 mm</quantity>
. Color of dorsum pale yellowish brown, head and pronotum slightly pale reddish in some specimens, appendages pale yellow, antennomeres 3 to 11 slightly darker yellowish brown; dorsal surface covered with dense but short pubescence.
</paragraph>
<paragraph id="8BC0368BFF9CFF803CFC63D1FEC87329" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">
Head. Relatively large, with eyes large and convex. Tempora very short. Frons flattened, nearly smooth, shiny but finely punctate; frontal furrows arcuate and deeply impressed to posterior margin of tempora; two pairs of supraorbital setae present and distinctly longer than setae of pubescence. Labrum broad with apical margin distinctly emarginate or concave. Mandibles small, slen- der.
<collectingRegion id="49BBF869FF9CFF803CFE6450FECD7368" box="[169,259,1057,1080]" country="Myanmar" name="Chin" pageId="9" pageNumber="350">Mentum</collectingRegion>
deeply concave, with median tooth broad and truncate. Submentum with a transverse row of ten setae anteriorly. Gula broad. Antennae of moderate length, antennomeres 2 and 3 of equal length.
</paragraph>
<paragraph id="8BC0368BFF9CFF803CFC64F3FC9A722B" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">Pronotum. Slightly transverse (ratio PW/PL = 1.32) and flattened, widest at anterior one-fourth, with lateral margins markedly rounded anteriorly, less so posteriorly, briefly and deeply sinuate just anterior to basal angles, the latter acute and sharp but small; basal margin projected posteriorly as a short, truncate lobe medial to lateral sinuations. Disc moderately punctate, flattened medially and slightly convex laterally; median basal area rugulose, only faintly defined anteriorly; median longitudinal impression distinct throughout, narrow anteriorly, widened toward base. Lateral margination slender throughout, slightly reflexed dorsally. One lateral seta (at anterior one-fifth) and one basolateral seta (on basal angle) present on each side.</paragraph>
<paragraph id="8BC0368BFF9CFF803CFC65F2FD88716C" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">Elytra. Alutaceous, moderately elongate, with humeri very distinct, rectangular but broadly rounded; elytral disc flattened, only moderately convex laterally; lateral margination slender and slightly reflexed dorsally; basal margin terminated medially at the origin of stria 5; striae 1 to 4 or 5 dinstinctly impressed and moderately punctate, striae 5 or 6 to 8 effaced. Recurrent stria absent. Umbilicate setal series with setae of humeral group equidistance from each other, those of the median group inserted posterior to middle.</paragraph>
<paragraph id="8BC0368BFF9CFF803CFC6634FC12710C" blockId="9.[123,1174,219,1725]" box="[171,988,1604,1629]" pageId="9" pageNumber="350">Legs. Short. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFF9CFF803CFC6614FF5371ED" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="350">Venter. Ventral surface of head and thoracic pleurae, prosternum, metasternum, metepisterna and abdominal ventrites punctate. Pubescence denser on abdominal ventrites than on thoracic venter.</paragraph>
<caption id="DF006603FF9FFF833CFC6638FDC471DE" ID-DOI="http://doi.org/10.5281/zenodo.13155293" ID-Zenodo-Dep="13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="10" pageNumber="351" startId="10.[171,183,1609,1628]" targetBox="[144,1176,216,1593]" targetPageId="10" targetType="figure">
<paragraph id="8BC0368BFF9FFF833CFC6638FDC471DE" blockId="10.[123,1174,1608,1679]" pageId="10" pageNumber="351">
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6.
<taxonomicName id="4C7F4D08FF9FFF833D546639FE73710B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[259,445,1608,1627]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="pusilloides" status="sp. nov.">
<emphasis id="B90BEA99FF9FFF833D546639FE73710B" box="[259,445,1608,1627]" italics="true" pageId="10" pageNumber="351">Perileptus pusilloides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF9FFF833D936638FDC7710C" box="[452,521,1609,1628]" pageId="10" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FF9FFF833F036612FC097125" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[852,967,1634,1653]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="pusilloides">
<emphasis id="B90BEA99FF9FFF833F036612FC097125" box="[852,967,1634,1653]" italics="true" pageId="10" pageNumber="351">P. pusilloides</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF9EFF823CFC60AAFBE077A3" blockId="11.[123,1174,219,1046]" box="[171,1070,219,243]" pageId="11" pageNumber="352">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFF9EFF823DB860AAFDF277A3" box="[495,572,219,243]" captionStart="FIGURE" captionStartId="10.[171,183,1609,1628]" captionTargetBox="[144,1176,216,1593]" captionTargetId="figure-11@10.[120,1176,216,1593]" captionTargetPageId="10" captionText="FIGURE 6. Perileptus pusilloides sp. nov.; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for P. pusilloides in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="11" pageNumber="352">Fig. 6b</figureCitation>
) with apex narrowly lobate, apically rounded.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9EFF823CFC608AFC197346" pageId="11" pageNumber="352" type="discussion">
<paragraph id="8BC0368BFF9EFF823CFC608AFEFB75C4" blockId="11.[123,1174,219,1046]" pageId="11" pageNumber="352">
<smallCapsWord id="8D26A057FF9EFF823CFC608AFEE67642" baselines="269,269" box="[171,296,251,275]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="11" pageNumber="352">
<emphasis id="B90BEA99FF9EFF823CFC608AFEE67642" bold="true" box="[171,296,251,275]" pageId="11" pageNumber="352">COMMENTS</emphasis>
</smallCapsWord>
.— Based on features of form and structure, this species appears to be closely relat- ed to
<taxonomicName id="4C7F4D08FF9EFF823CE2616DFEEC7663" box="[181,290,283,307]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9EFF823CE2616DFEEC7663" box="[181,290,283,307]" italics="true" pageId="11" pageNumber="352">P. pusillus</emphasis>
</taxonomicName>
, described from northern
<collectingCountry id="F368761BFF9EFF823E60616AFD5B7663" box="[567,661,283,307]" name="Vietnam" pageId="11" pageNumber="352">Vietnam</collectingCountry>
. Its members can be distinguished from those of the latter in having their elytra wider, less shiny, more alutaceous, and with more finely punctate striae, and the pronotum relatively larger, with a less smooth surface, and with the greatest pronotal width more anterior. The type series of
<taxonomicName id="4C7F4D08FF9EFF823E17610DFD6076C4" box="[576,686,380,404]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9EFF823E17610DFD6076C4" box="[576,686,380,404]" italics="true" pageId="11" pageNumber="352">P. pusillus</emphasis>
</taxonomicName>
originally consisted of ten
<typeStatus id="54C48829FF9EFF823F86610DFBFF76C4" box="[977,1073,380,404]" pageId="11" pageNumber="352" type="syntype">syntypes</typeStatus>
, seven of which are in the Jeannel Collection in MNHN, each with a “Type” label. Among these,
<specimenCount id="9D79FD02FF9EFF82387861EDFF7D7683" count="6" pageId="11" pageNumber="352" type="generic">six specimens</specimenCount>
match Jeannels description of
<taxonomicName id="4C7F4D08FF9EFF823E5661CCFDA37684" box="[513,621,444,468]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9EFF823E5661CCFDA37684" box="[513,621,444,468]" italics="true" pageId="11" pageNumber="352">P. pusillus</emphasis>
</taxonomicName>
very closely. The seventh specimen, a male, instead may be a member of our new species,
<taxonomicName id="4C7F4D08FF9EFF823E4861ACFD6076A4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[543,686,476,500]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusilloides">
<emphasis id="B90BEA99FF9EFF823E4861ACFD6076A4" box="[543,686,476,500]" italics="true" pageId="11" pageNumber="352">P. pusilloides</emphasis>
</taxonomicName>
. Consequently, in order to properly establish the identity to
<taxonomicName id="4C7F4D08FF9EFF823D4A618CFE437544" box="[285,397,508,532]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9EFF823D4A618CFE437544" box="[285,397,508,532]" italics="true" pageId="11" pageNumber="352">P. pusillus</emphasis>
</taxonomicName>
, we here designate a
<typeStatus id="54C48829FF9EFF823E29618DFD2E7544" box="[638,736,508,532]" pageId="11" pageNumber="352" type="lectotype">lectotype</typeStatus>
as follows: a male, in MNHN, labeled: “
<collectingRegion id="49BBF869FF9EFF823CD3626DFF3F7564" box="[132,241,540,564]" country="Vietnam" name="Hoa Binh" pageId="11" pageNumber="352">Hoa Binh</collectingRegion>
,
<collectingCountry id="F368761BFF9EFF823CAA626DFE847564" box="[253,330,540,564]" name="Vietnam" pageId="11" pageNumber="352">Tonkin</collectingCountry>
”/ “
<taxonomicName id="4C7F4D08FF9EFF823D27626DFD5A7564" authority="Jeannel" authorityName="Jeannel" box="[368,660,540,564]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9EFF823D27626DFDF77564" box="[368,569,540,564]" italics="true" pageId="11" pageNumber="352">Perileptus pusillus</emphasis>
Jeannel
</taxonomicName>
,
<typeStatus id="54C48829FF9EFF823EF7626DFCCB7564" box="[672,773,540,564]" pageId="11" pageNumber="352" type="lectotype">lectotype</typeStatus>
, design. 2016, ex coll. R. Jeannel, in coll. MNHN, Paris”. The remaining six MNHN
<typeStatus id="54C48829FF9EFF823ECA624CFD347505" box="[669,762,573,597]" pageId="11" pageNumber="352" type="syntype">syntypes</typeStatus>
are all
<typeStatus id="54C48829FF9EFF823F00624DFC3F7504" box="[855,1009,572,596]" pageId="11" pageNumber="352" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C7F4D08FF9EFF82384A624CFB417504" box="[1053,1167,572,596]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90BEA99FF9EFF82384A624CFB417504" box="[1053,1167,572,596]" italics="true" pageId="11" pageNumber="352">P. pusillus</emphasis>
</taxonomicName>
, including the lone possible male of
<taxonomicName id="4C7F4D08FF9EFF823DAC622CFD477524" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[507,649,604,628]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="pusilloides">
<emphasis id="B90BEA99FF9EFF823DAC622CFD477524" box="[507,649,604,628]" italics="true" pageId="11" pageNumber="352">P. pusilloides</emphasis>
</taxonomicName>
, which we are not including in the type series of the latter species.
</paragraph>
<paragraph id="8BC0368BFF9EFF823CFC62ECFD8A7405" blockId="11.[123,1174,219,1046]" pageId="11" pageNumber="352">
<emphasis id="B90BEA99FF9EFF823CFC62ECFE6D75E3" bold="true" box="[171,419,669,693]" pageId="11" pageNumber="352">
<smallCapsWord id="8D26A057FF9EFF823CFC62ECFEC775E3" baselines="687,686" box="[171,265,669,693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="11" pageNumber="352">HABITAT</smallCapsWord>
<smallCapsWord id="8D26A057FF9EFF823D5862D1FE6D75E3" baselines="686" box="[271,419,672,691]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="11" pageNumber="352">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in daytime on sandy flats, in gravel and under stones along the open, unshaded banks of small to large streams running through agricultural areas with subtropical crops at elevations ranging from
<quantity id="4C879B6EFF9EFF823FE462ACFB9D75A4" box="[947,1107,733,757]" metricMagnitude="2" metricUnit="m" metricValue="8.925" metricValueMax="11.05" metricValueMin="6.8" pageId="11" pageNumber="352" unit="m" value="892.5" valueMax="1105.0" valueMin="680.0">680 to 1105 m</quantity>
. Most specimens were driven from their hiding places by splashing the banks with water from the stream. At the collecting site in Longyang County (see below), members of this species were found syntopic with members of
<taxonomicName id="4C7F4D08FF9EFF823D22634CFDF37405" authorityName="Jeannel Body" authorityYear="1923" box="[373,573,829,853]" class="Insecta" family="Carabidae" genus="Perileptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="imaicus">
<emphasis id="B90BEA99FF9EFF823D22634CFDF37405" box="[373,573,829,853]" italics="true" pageId="11" pageNumber="352">Perileptus imaicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0368BFF9EFF823CFC632CFC287485" blockId="11.[123,1174,219,1046]" pageId="11" pageNumber="352">
<emphasis id="B90BEA99FF9EFF823CFC632CFDBA7424" bold="true" box="[171,628,861,885]" pageId="11" pageNumber="352">
<smallCapsWord id="8D26A057FF9EFF823CFC632CFE987424" baselines="879,879" box="[171,342,861,885]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="11" pageNumber="352">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF9EFF823D0C6310FE217424" baselines="879" box="[347,495,865,884]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="11" pageNumber="352">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FF9EFF823DA26310FD8A7424" baselines="879" box="[501,580,865,884]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="11" pageNumber="352">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FF9EFF823E1D6310FDBA7424" baselines="879" box="[586,628,865,884]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="11" pageNumber="352">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FF9EFF823E2D632CFD317424" baselines="879,879" box="[634,767,861,885]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="11" pageNumber="352">
<emphasis id="B90BEA99FF9EFF823E2D632CFD317424" bold="true" box="[634,767,861,885]" pageId="11" pageNumber="352">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FF9EFF823F52632CFE2074E5" location="We" pageId="11" pageNumber="352" specimenCount="10" specimenCount-female="2" specimenCount-male="3" stateProvince="Shan">
<smallCapsWord id="8D26A057FF9EFF823F52632CFC8E7424" baselines="879,879" box="[773,832,861,885]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="11" pageNumber="352">
<emphasis id="B90BEA99FF9EFF823F52632CFC8E7424" bold="true" box="[773,832,861,885]" pageId="11" pageNumber="352">
<collectingRegion id="49BBF869FF9EFF823F52632CFC8E7424" box="[773,832,861,885]" country="Myanmar" name="Shan" pageId="11" pageNumber="352">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFF9EFF823F3F632CFC7A7425" box="[872,948,861,885]" captionStart="FIGURE" captionStartId="10.[171,183,1609,1628]" captionTargetBox="[144,1176,216,1593]" captionTargetId="figure-11@10.[120,1176,216,1593]" captionTargetPageId="10" captionText="FIGURE 6. Perileptus pusilloides sp. nov.; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for P. pusilloides in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="11" pageNumber="352">Fig. 6c</figureCitation>
.
<location id="8EA06050FF9EFF823FEA632FFC2E7425" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF9EFF823FEA632FFC2E7425" box="[957,992,862,885]" name="We" pageId="11" pageNumber="352" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FF9EFF823CCF630CFED574C5" box="[152,283,893,917]" count="5" pageId="11" pageNumber="352" type="generic">5 specimens</specimenCount>
(
<specimenCount id="9D79FD02FF9EFF823D7C630CFEB374C5" box="[299,381,893,917]" count="3" pageId="11" pageNumber="352" type="male">3 males</specimenCount>
and
<specimenCount id="9D79FD02FF9EFF823DE6630CFDD474C5" box="[433,538,893,917]" count="2" pageId="11" pageNumber="352" type="female">2 females</specimenCount>
) from low elevations on both western and eastern slopes of the southern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FF9EFF823DA263ECFC2D7485" location="Longyang Counties" municipality="Tengchong" pageId="11" pageNumber="352" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF9EFF823DA263ECFDE474E5" box="[501,554,925,949]" country="Myanmar" name="Shan" pageId="11" pageNumber="352">Shan</collectingRegion>
in
<collectingMunicipality id="6BA4ACF1FF9EFF823E1963ECFD0A74E5" box="[590,708,925,949]" pageId="11" pageNumber="352">Tengchong</collectingMunicipality>
and
<location id="8EA06050FF9EFF823EAE63EFFC0174E5" LSID="urn:lsid:plazi:treatment:03D6879DFF92FF823C2C65A9FC197346:8EA06050FF9EFF823EAE63EFFC0174E5" box="[761,975,925,950]" municipality="Tengchong" name="Longyang Counties" pageId="11" pageNumber="352" stateProvince="Shan">Longyang Counties</location>
, (see
<typeStatus id="54C48829FF9EFF82385B63EFFB8F74E6" box="[1036,1089,926,950]" pageId="11" pageNumber="352">Type</typeStatus>
material above for exact collection data), which are in Core Areas 6 and 7, respectively
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF9EFF823CFC63ACFC197346" blockId="11.[123,1174,219,1046]" pageId="11" pageNumber="352">
<emphasis id="B90BEA99FF9EFF823CFC63ACFDAF74A4" bold="true" box="[171,609,989,1013]" pageId="11" pageNumber="352">
<smallCapsWord id="8D26A057FF9EFF823CFC63ACFEDC74A4" baselines="1007,1007" box="[171,274,989,1013]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="11" pageNumber="352">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FF9EFF823D4A6390FE0A74A4" baselines="1007" box="[285,452,993,1012]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="11" pageNumber="352">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF9EFF823D9A6390FDAF74A4" baselines="1007" box="[461,609,993,1012]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="11" pageNumber="352">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from the southern part of the Gaoligong Shan region in western
<collectingRegion id="49BBF869FF9EFF823E9D638FFC4B7346" box="[714,901,1022,1046]" country="China" name="Yunnan" pageId="11" pageNumber="352">Yunnan Province</collectingRegion>
,
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Genus
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.—
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.— Adults of this genus (
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,
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) can be recognized by the following combination of character states: size small to medium (BL = 2.0 to
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), fully winged, dorsal surface covered with more or less long pubescence; eyes large, convex, pubescent; frons flat, frontal furrows wide and deep, attenuated posteriorly; terminal palpomeres slender, attenuated apically; mentum free, not fused with submentum; submentum with 10 or 12 setae; antennal scape distinctly pubescent; pronotum narrowed basally, with base broadly projected posteriorly, median longitudinal furrow deeply and sharply defined; elytra elongate and flattened, recurrent stria indistinct; legs short, protibiae without longitudinal furrows.
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.—
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is a moderately diverse genus with about 50 described species arrayed in four subgenera (
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). It is represented in the Palearctic, Oriental, Afrotropical, Australian and Neotropical Regions. The study area is within the previously known range of this genus.
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<mods:number id="8F1771917689493919C83C7502D0CDAF">2016-10-14</mods:number>
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<mods:number id="6A433B534ED8D5A4EBD46957C7901656">63</mods:number>
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<mods:identifier id="CCA621FEA5A5AEEE8F9C45C1B8AA7D0F" type="DOI">10.5281/zenodo.13155283</mods:identifier>
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<treatment id="03D6879DFF99FF863C2C6624FC447147" LSID="urn:lsid:plazi:treatment:03D6879DFF99FF863C2C6624FC447147" httpUri="http://treatment.plazi.org/id/03D6879DFF99FF863C2C6624FC447147" lastPageId="15" lastPageNumber="356" pageId="12" pageNumber="353">
<subSubSection id="C3656500FF99FF853C2C6624FD1E713D" box="[123,720,1621,1645]" pageId="12" pageNumber="353" type="nomenclature">
<paragraph id="8BC0368BFF99FF853C2C6624FD1E713D" blockId="12.[123,720,1621,1676]" box="[123,720,1621,1645]" pageId="12" pageNumber="353">
<heading id="D08881E7FF99FF853C2C6624FD1E713D" box="[123,720,1621,1645]" fontSize="10" level="2" pageId="12" pageNumber="353" reason="4">
<emphasis id="B90BEA99FF99FF853C2C6624FD1E713D" bold="true" box="[123,720,1621,1645]" pageId="12" pageNumber="353">
<taxonomicName id="4C7F4D08FF99FF853C2C6624FDA0713D" ID-CoL="8MGKQ" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,622,1621,1645]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="species" species="fugongensis" status="sp. nov.">
<emphasis id="B90BEA99FF99FF853C2C6624FE6F713D" bold="true" box="[123,417,1621,1645]" italics="true" pageId="12" pageNumber="353">Agonotrechus fugongensis</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FF99FF853E2D6624FD1E713D" box="[634,720,1621,1645]" pageId="12" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3656500FF99FF853C2C6607FEBB71DC" box="[123,373,1654,1676]" pageId="12" pageNumber="353" type="description">
<paragraph id="8BC0368BFF99FF853C2C6607FEBB71DC" blockId="12.[123,720,1621,1676]" box="[123,373,1654,1676]" pageId="12" pageNumber="353">
(
<figureCitation id="13442A0EFF99FF853CD56607FF0971DB" box="[130,199,1654,1676]" captionStart="FIGURE" captionStartId="14.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1577]" captionTargetId="figure-11@14.[120,1176,216,1577]" captionTargetPageId="14" captionText="FIGURE 7. Agonotrechus fugongensis sp. nov.; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for A. fugongensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155295" httpUri="https://zenodo.org/record/13155295/files/figure.png" pageId="12" pageNumber="353">Figs. 7</figureCitation>
,
<figureCitation id="13442A0EFF99FF853C866607FF3971DC" box="[209,247,1654,1676]" captionStart="FIGURE" captionStartId="97.[173,185,1623,1642]" captionTargetBox="[142,1149,216,1606]" captionTargetId="figure-14@97.[142,1149,216,1606]" captionTargetPageId="97" captionText="FIGURE 35. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, Xiao Shangla He at Shuangla Village, elevation 1550 m; habitat in which specimens of Perileptus imaicus Jeannel were collected. b. Gongshan County, Qigi He, 9.9 airkm W of Cikai, elevation 2000 m; habitat in which specimens of Agonotrechus fugongensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155350" httpUri="https://zenodo.org/record/13155350/files/figure.png" pageId="12" pageNumber="353">35b</figureCitation>
,
<figureCitation id="13442A0EFF99FF853D556607FEE871DC" box="[258,294,1654,1676]" captionStart="FIGURE" captionStartId="98.[171,183,1623,1642]" captionTargetBox="[144,1152,216,1605]" captionTargetId="figure-11@98.[144,1152,216,1605]" captionTargetPageId="98" captionText="FIGURE 36. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Fugong County, Shiyueliang Township, 1 km E of Shibali on Shibali Road, elevation 2400 m; habitat in which specimens of Agonotrechus fugongensis sp. nov. were collected. b. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp, elevation 2770 m; habitat in which specimens of Agonotrechus wuyipeng Deuve were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155356" httpUri="https://zenodo.org/record/13155356/files/figure.png" pageId="12" pageNumber="353">36a</figureCitation>
,
<figureCitation id="13442A0EFF99FF853D666607FEA171DC" box="[305,367,1654,1676]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="12" pageNumber="353">4648</figureCitation>
)
</paragraph>
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<subSubSection id="C3656500FF99FF843CFC66D7FED175C6" lastPageId="13" lastPageNumber="354" pageId="12" pageNumber="353" type="materials_examined">
<paragraph id="8BC0368BFF99FF843CFC66D7FEAE76A5" blockId="12.[171,1174,1701,1726]" lastBlockId="13.[123,1174,219,1726]" lastPageId="13" lastPageNumber="354" pageId="12" pageNumber="353">
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.—
<materialsCitation id="3B173CD6FF99FF843DD466D4FE5F7604" collectingDate="2005-08-18" collectorName="D. Z. Dong" country="China" county="Fugong County" elevation="2415" lastPageId="13" lastPageNumber="354" latitude="27.16536" location="Lishadi Township" longLatPrecision="1" longitude="98.78003" pageId="12" pageNumber="353" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FF99FF853DD466D4FE2771ED" box="[387,489,1701,1725]" pageId="12" pageNumber="353" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1019979”/ “
<collectingCountry id="F368761BFF99FF853FAA66D7FB9871EE" box="[1021,1110,1702,1726]" name="China" pageId="12" pageNumber="353">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FF98FF843CE560AAFE9077A3" box="[178,350,219,243]" pageId="13" pageNumber="354">Fugong County</collectingCounty>
,
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,
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<quantity id="4C879B6EFF98FF843E6960AAFD6077A3" box="[574,686,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="13" pageNumber="354" unit="km" value="1.0" valueMax="2.0" valueMin="0.0">0 to 2 km</quantity>
E of Shibali on Shibali Road
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,
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to
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/”/ “
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,
<quantity id="4C879B6EFF98FF843E21608AFCC77643" box="[630,777,251,275]" metricMagnitude="3" metricUnit="m" metricValue="2.415" metricValueMax="2.53" metricValueMin="2.3" pageId="13" pageNumber="354" unit="m" value="2415.0" valueMax="2530.0" valueMin="2300.0">
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,
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<collectingDate id="EF85E9A3FF98FF843F42608AFC0D7643" box="[789,963,251,275]" pageId="13" pageNumber="354" value="2005-08-18">18 August 2005</collectingDate>
</date>
, Stop# DHK-2005- 096,
<collectorName id="268A535DFF98FF843CF8616DFEE97664" box="[175,295,284,308]" pageId="13" pageNumber="354">D. Z. Dong</collectorName>
collector”/ “
<typeStatus id="54C48829FF98FF843DFA616DFDF27664" box="[429,572,284,308]" pageId="13" pageNumber="354" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FF98FF843E15616AFBC67663" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[578,1032,283,308]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="fugongensis" status="sp. nov.">
<emphasis id="B90BEA99FF98FF843E15616AFC947664" box="[578,858,283,308]" italics="true" pageId="13" pageNumber="354">Agonotrechus fugongensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
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designated 2016” [red label]
</materialsCitation>
.
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<typeStatus id="54C48829FF98FF843DC9614DFDC87604" box="[414,518,316,340]" pageId="13" pageNumber="354" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="ED6EAE4EFF98FF843F52614DFCF47604" box="[773,826,316,340]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="354" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1006974”/ “
<collectingCountry id="F368761BFF98FF843CD3612DFF137624" box="[132,221,348,372]" name="China" pageId="13" pageNumber="354">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF98FF843CBD612DFEF07623" box="[234,318,348,371]" country="China" name="Yunnan" pageId="13" pageNumber="354">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF98FF843D1C612DFE327624" box="[331,508,348,372]" pageId="13" pageNumber="354">Gaoligong Shan</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF98FF843E5E612DFD167624" box="[521,728,348,372]" pageId="13" pageNumber="354">Nujiang Prefecture</collectingMunicipality>
,
<location id="8EA06050FF98FF843EB3612DFBE57623" LSID="urn:lsid:plazi:treatment:03D6879DFF99FF863C2C6624FC447147:8EA06050FF98FF843EB3612DFBE57623" box="[740,1067,348,372]" country="China" county="Gaoligong Shan" latitude="27.71542" longLatPrecision="1" longitude="98.56529" municipality="Nujiang Prefecture" name="Nujiang State Nature Reserve" pageId="13" pageNumber="354" stateProvince="Yunnan">Nujiang State Nature Reserve</location>
,
<location id="8EA06050FF98FF84386F612DFB5C7623" LSID="urn:lsid:plazi:treatment:03D6879DFF99FF863C2C6624FC447147:8EA06050FF98FF84386F612DFB5C7623" box="[1080,1170,348,372]" country="China" county="Gaoligong Shan" latitude="27.71542" longLatPrecision="1" longitude="98.56529" municipality="Nujiang Prefecture" name="Qigi He" pageId="13" pageNumber="354" stateProvince="Yunnan">Qigi He</location>
, 9.9 airkm
<locationDeviation id="9A722FA9FF98FF843CBA610CFE6E76C4" box="[237,416,380,404]" location="Gongshan" pageId="13" pageNumber="354">W of Gongshan</locationDeviation>
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/
<geoCoordinate id="EE4B504CFF98FF843E09610DFD2E76C4" box="[606,736,380,404]" degrees="98.56529" direction="east" orientation="longitude" pageId="13" pageNumber="354" precision="1" value="98.56529">E98.56529°</geoCoordinate>
,
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<elevation id="0052D1B8FF98FF843EB9610DFCF576C4" box="[750,827,380,404]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="354" unit="m" value="2000.0">2000m</elevation>
</quantity>
,
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<collectingDate id="EF85E9A3FF98FF843F1F610DFC3F76C4" box="[840,1009,380,404]" pageId="13" pageNumber="354" value="2000-07-09" valueMax="2000-07-14" valueMin="2000-07-09">9-14 July 2000</collectingDate>
</date>
, Stop#00-22A, D.H.
<taxonomicName id="4C7F4D08FF98FF843CEF61EDFEFA76E4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[184,308,412,436]" pageId="13" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFF98FF843D1461EDFE2C76E4" box="[323,482,412,436]" pageId="13" pageNumber="354">C.E. Griswold</collectorName>
, Liang H.-B., D. Ubick, &amp; Dong D.-Z. collectors”/ “IMAGE” [pale green label]/ “
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<taxonomicName id="4C7F4D08FF98FF843DB261CDFC767684" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[485,952,444,469]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="fugongensis" status="sp. nov.">
<emphasis id="B90BEA99FF98FF843DB261CDFCCE7685" box="[485,768,444,469]" italics="true" pageId="13" pageNumber="354">Agonotrechus fugongensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
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designated 2016” [yellow label].
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<materialsCitation id="3B173CD6FF98FF843CFC618CFDD27565" country="China" county="Fugong County" elevation="2415" location="Fugong County" pageId="13" pageNumber="354" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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.—
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,
<collectingRegion id="49BBF869FF98FF843D86618FFDEB7545" box="[465,549,510,533]" country="China" name="Yunnan" pageId="13" pageNumber="354">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF98FF843E65618FFD137545" box="[562,733,509,534]" pageId="13" pageNumber="354">Fugong County</collectingCounty>
,
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,
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<quantity id="4C879B6EFF98FF843FBE618CFB9B7545" box="[1001,1109,509,533]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="13" pageNumber="354" unit="km" value="1.0" valueMax="2.0" valueMin="0.0">0 to 2 km</quantity>
W of Shibali on Shibali Road
</locationDeviation>
,
<quantity id="4C879B6EFF98FF843DDE626CFDD27565" box="[393,540,541,565]" metricMagnitude="3" metricUnit="m" metricValue="2.415" metricValueMax="2.53" metricValueMin="2.3" pageId="13" pageNumber="354" unit="m" value="2415.0" valueMax="2530.0" valueMin="2300.0">
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.
</paragraph>
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.— The species epithet,
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<emphasis id="B90BEA99FF98FF843EA0624FFCB87506" box="[759,886,574,598]" italics="true" pageId="13" pageNumber="354">fugongensis</emphasis>
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, is derived from the name of the county (Xian) in which the
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was collected, Fugong, and the Latin suffix,
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, denoting place.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF98FF843CFC62EFFEDB7309" pageId="13" pageNumber="354" type="diagnosis">
<paragraph id="8BC0368BFF98FF843CFC62EFFEDB7309" blockId="13.[123,1174,219,1726]" pageId="13" pageNumber="354">
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<emphasis id="B90BEA99FF98FF843CFC62EFFEEE75E5" bold="true" box="[171,288,670,694]" pageId="13" pageNumber="354">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFF98FF843E1B62EFFD5575E6" box="[588,667,670,694]" captionStart="FIGURE" captionStartId="14.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1577]" captionTargetId="figure-11@14.[120,1176,216,1577]" captionTargetPageId="14" captionText="FIGURE 7. Agonotrechus fugongensis sp. nov.; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for A. fugongensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155295" httpUri="https://zenodo.org/record/13155295/files/figure.png" pageId="13" pageNumber="354">Fig. 7a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL = 5.8 to 6.0 mm), fully-winged, dorsum dark piceous to reddish brown, shiny, pronotum and elytra slightly iridescent; eyes convex, moderately projected; clypeus with four setae; mentum and submentum fused, submentum with six setae; pronotum small but transverse, ratio PW/PL = 1.30, median basal area smooth, framed by two deep, oblique furrows, lateral explanation rather narrow, broadened only posteriorly, lateral margins with a short sinuation anterior to the sharp, rectangular or slightly acute basal angles; elytra with all discal striae evident and finely punctate, striae 1 to 4 moderately impressed, striae 5 to 8 more faintly impressed, parascutellar striole rather long, recurrent stria deeply impressed but abruptly terminated anteriorly, not connected with stria 5, intervals slightly convex, two discal seta (anterior and middle setae) present in stria 3, the anterior seta inserted within basal one-sixth of elytra, the middle seta inserted near elytral middle, preapical seta also present; median lobe of aedeagus of male (
<figureCitation id="13442A0EFF98FF843E446450FDAE7369" box="[531,608,1057,1081]" captionStart="FIGURE" captionStartId="14.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1577]" captionTargetId="figure-11@14.[120,1176,216,1577]" captionTargetPageId="14" captionText="FIGURE 7. Agonotrechus fugongensis sp. nov.; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for A. fugongensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155295" httpUri="https://zenodo.org/record/13155295/files/figure.png" pageId="13" pageNumber="354">Fig. 7b</figureCitation>
) long and slender, endophallus with copulatory piece spoon-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF98FF863CFC6410FF687425" lastPageId="15" lastPageNumber="356" pageId="13" pageNumber="354" type="description">
<paragraph id="8BC0368BFF98FF843CFC6410FBC673C9" blockId="13.[123,1174,219,1726]" pageId="13" pageNumber="354">
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.— Size large, BL = 5.8 to 6.0 mm. Color of dorsum piceous to reddish brown, shiny, pronotum and elytra slightly iridescent, appendages paler, palpi yellowish tan.
</paragraph>
<paragraph id="8BC0368BFF98FF843CFC64D3FF41710D" blockId="13.[123,1174,219,1726]" pageId="13" pageNumber="354">
Head. Moderate in size, slightly elongate, eyes large, convex, moderately projected, their diameter more than two times length of tempora. Tempora short, only slightly convex and glabrous, joined to neck region at ca. 135° angle. Frons more or less flattened, with two pairs of supraorbital setae, frontal furrows distinct, impressed posterior to or beyond posterior supraorbital setae. Clypeus with four setae. Labrum with six setae, apical margin slightly concave. Mandibles slen- der; right mandible bidentate, the anterior tooth far forward and long, the basal tooth reduced, obtuse and blunt; left mandible with a very slender trifid process, formed from fusion of the retinaculum with premolar tooth. Palpi with apical palpomeres fusiform.
<collectingRegion id="49BBF869FF98FF843FD465F5FC1372CB" box="[899,989,1412,1435]" country="Myanmar" name="Chin" pageId="13" pageNumber="354">Mentum</collectingRegion>
and submentum fused.
<collectingRegion id="49BBF869FF98FF843C9565D5FED272EB" box="[194,284,1444,1467]" country="Myanmar" name="Chin" pageId="13" pageNumber="354">Mentum</collectingRegion>
bifossulate, divided into three sectors, a median and two lateral parts, separated by two deep, longitudinal furrows; medial tooth simple, subtriangular with blunt apex, half as long as lateral lobes. Submentum with six setae subapically, gula broad. Genae with a single ventral seta on each side. Antennae of moderate length, with only four antennomeres extended beyond the pronotal base; antennonmeres 3 and 4 virtually the same length and each longer than antennomere 2.
</paragraph>
<paragraph id="8BC0368BFF98FF863CFC6614FCDB7623" blockId="13.[123,1174,219,1726]" lastBlockId="15.[123,1174,219,1559]" lastPageId="15" lastPageNumber="356" pageId="13" pageNumber="354">Pronotum. Rather small, moderately narrow posteriorly, ratio PW/PL = 1.30, widest slightly anterior to middle, lateral margins with short sinuation just anterior to sharp, rectangular or slightly acute basal angles; pronotal disc convex, median longitudinal impression very fine and shallow, extended posteriorly to near basal margin, but not as close anteriorly to apical margin; basal foveae small vaguely delimited; median basal area smooth, delimited laterally by short but deep and oblique furrows; basal margin nearly straight, slightly sinuate. Lateral explanation moderately narrow and slightly reflexed, widened posteriorly but not flattened. Midlateral pair of setae inserted anterior to middle and basolateral pair inserted at hind angles.</paragraph>
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7.
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<emphasis id="B90BEA99FF9BFF873D53664AFE29711F" box="[260,487,1595,1615]" italics="true" pageId="14" pageNumber="355">Agonotrechus fugongensis</emphasis>
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; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for
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<emphasis id="B90BEA99FF9BFF873F2B6624FC327138" box="[892,1020,1621,1640]" italics="true" pageId="14" pageNumber="355">A. fugongensis</emphasis>
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in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF9AFF863CFC610AFEA975E4" blockId="15.[123,1174,219,1559]" pageId="15" pageNumber="356">Elytra. Moderately wide, elytral silhouette subovoid, humeri distinct but rounded, disc convex. All discal striae evident and finely punctate, striae 1 to 4 moderately impressed, striae 5 to 8 more faintly impressed, partially effaced, parascutellar striole rather long, recurrent stria deeply impressed but abruptly terminated anteriorly, not connected with discal stria 5, intervals slightly convex. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Two discal seta (anterior and middle setae) present in stria 3, the anterior seta inserted within basal one-sixth of elytra, the middle seta inserted near elytral middle. Preapical seta also present, inserted on interval 2 near stria 2, closer to sutural elytral margin than to apex. Umbilicate setal series with setae of humeral group equidistant for each other and those of median group both inserted posteri- or to middle of elytra.</paragraph>
<paragraph id="8BC0368BFF9AFF863CFC62CCFC9E75A4" blockId="15.[123,1174,219,1559]" pageId="15" pageNumber="356">Legs. Slender, moderately long. Protibiae furrowed, without anteroapical pubescence. Male protarsi with tarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFF9AFF863CFC628DFD247464" blockId="15.[123,1174,219,1559]" pageId="15" pageNumber="356">Abdomen. Ventrites each with a pair of paramedial setae, ventrite VII of males with a single pair of paramedial apical setae, of females with two pairs.</paragraph>
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Male aedeagus. Median lobe (
<figureCitation id="13442A0EFF9AFF863DBE634DFDFB7404" box="[489,565,828,852]" captionStart="FIGURE" captionStartId="14.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1577]" captionTargetId="figure-11@14.[120,1176,216,1577]" captionTargetPageId="14" captionText="FIGURE 7. Agonotrechus fugongensis sp. nov.; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for A. fugongensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155295" httpUri="https://zenodo.org/record/13155295/files/figure.png" pageId="15" pageNumber="356">Fig. 7b</figureCitation>
) long and slender, endophallus with a spoon-shaped sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9AFF863CFC630DFC447147" pageId="15" pageNumber="356" type="discussion">
<paragraph id="8BC0368BFF9AFF863CFC630DFE4274A5" blockId="15.[123,1174,219,1559]" pageId="15" pageNumber="356">
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<emphasis id="B90BEA99FF9AFF863CFC630DFEE674C3" bold="true" box="[171,296,892,916]" pageId="15" pageNumber="356">COMMENTS</emphasis>
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.— Males of this species are most similar to those of
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<emphasis id="B90BEA99FF9AFF863F0E630CFC1374C5" box="[857,989,893,917]" italics="true" pageId="15" pageNumber="356">A. wuyipeng</emphasis>
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in features of the aedeagus; but they are smaller, their pronota distinctly narrower, more slender, and with much narrower lateral margination, and the recurrent stria does not connect anteriorly with stria 5 as it does in
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<emphasis id="B90BEA99FF9AFF863CC063ACFED574A5" box="[151,283,989,1013]" italics="true" pageId="15" pageNumber="356">A. wuyipeng</emphasis>
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members.
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</emphasis>
.— The
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specimen of this species was found under a stone on moist substrate along a roadcut through an agricultural area formed in a large clearcut in what had been mixed broadleaf evergreen and conifer forest at an elevation somewhere between 2300 and
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(
<figureCitation id="13442A0EFF9AFF863C8A642CFEF57325" box="[221,315,1117,1141]" captionStart="FIGURE" captionStartId="98.[171,183,1623,1642]" captionTargetBox="[144,1152,216,1605]" captionTargetId="figure-11@98.[144,1152,216,1605]" captionTargetPageId="98" captionText="FIGURE 36. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Fugong County, Shiyueliang Township, 1 km E of Shibali on Shibali Road, elevation 2400 m; habitat in which specimens of Agonotrechus fugongensis sp. nov. were collected. b. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp, elevation 2770 m; habitat in which specimens of Agonotrechus wuyipeng Deuve were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155356" httpUri="https://zenodo.org/record/13155356/files/figure.png" pageId="15" pageNumber="356">Fig. 36a</figureCitation>
). The
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specimen was collected under stones on the shaded bank of the Qiqi River just above the Forestry station at Qiqi at an elevation of
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(
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). Members of no other
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<emphasis id="B90BEA99FF9AFF863D5E64EFFE5273E6" box="[265,412,1182,1206]" italics="true" pageId="15" pageNumber="356">Agonotrechus</emphasis>
</taxonomicName>
or other trechine species have been found syntopic with those of
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<emphasis id="B90BEA99FF9AFF863C2C64CFFED67386" box="[123,280,1214,1238]" italics="true" pageId="15" pageNumber="356">A. fugongensis</emphasis>
</taxonomicName>
.
</paragraph>
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.—
<figureCitation id="13442A0EFF9AFF863F3F64AFFC7A73A5" box="[872,948,1246,1270]" captionStart="FIGURE" captionStartId="14.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1577]" captionTargetId="figure-11@14.[120,1176,216,1577]" captionTargetPageId="14" captionText="FIGURE 7. Agonotrechus fugongensis sp. nov.; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for A. fugongensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155295" httpUri="https://zenodo.org/record/13155295/files/figure.png" pageId="15" pageNumber="356">Fig. 7c</figureCitation>
.
<location id="8EA06050FF9AFF863FEA64AFFC2E73A5" LSID="urn:lsid:plazi:treatment:03D6879DFF99FF863C2C6624FC447147:8EA06050FF9AFF863FEA64AFFC2E73A5" box="[957,992,1246,1269]" name="We" pageId="15" pageNumber="356" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FF9AFF863CC0648FFED77246" box="[151,281,1278,1302]" count="2" pageId="15" pageNumber="356" type="generic">2 specimens</specimenCount>
(
<specimenCount id="9D79FD02FF9AFF863D7D648FFEBE7246" box="[298,368,1278,1302]" count="1" pageId="15" pageNumber="356" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FF9AFF863DF5648FFDCE7246" box="[418,512,1278,1302]" count="1" pageId="15" pageNumber="356" type="female">1 female</specimenCount>
) from Fugong and Gongshan Counties, respectively (see
<typeStatus id="54C48829FF9AFF863836648FFB587246" box="[1121,1174,1278,1302]" pageId="15" pageNumber="356">Type</typeStatus>
material above for exact collection data)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF9AFF863CFC654FFB587286" blockId="15.[123,1174,219,1559]" pageId="15" pageNumber="356">
Specimens of this species were collected only in the northern half of the study area (Core Areas 2 and 3) and only on the eastern side of the mountain range. This distribution pattern may be an artifact of inadequate sampling on the western slope of the mountain range in the north, much of which is in
<collectingCountry id="F368761BFF9AFF863D4065EEFE4C72E7" box="[279,386,1439,1463]" name="Myanmar" pageId="15" pageNumber="356">Myanmar</collectingCountry>
. The geographical range of this species overlaps that of
<taxonomicName id="4C7F4D08FF9AFF863FB765EEFBAB72E6" box="[992,1125,1439,1462]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="wuyipeng">
<emphasis id="B90BEA99FF9AFF863FB765EEFBAB72E6" box="[992,1125,1439,1462]" italics="true" pageId="15" pageNumber="356">A. wuyipeng</emphasis>
</taxonomicName>
, but members of the latter species appear to occupy slightly higher elevations in the same general areas.
</paragraph>
<paragraph id="8BC0368BFF9AFF863CFC65AFFC447147" blockId="15.[123,1174,219,1559]" pageId="15" pageNumber="356">
<emphasis id="B90BEA99FF9AFF863CFC65AFFDAF72A5" bold="true" box="[171,609,1502,1526]" pageId="15" pageNumber="356">
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<smallCapsWord id="8D26A057FF9AFF863D4A6593FE0A72A5" baselines="1520" box="[285,452,1506,1525]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="15" pageNumber="356">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF9AFF863D9A6593FDAF72A5" baselines="1520" box="[461,609,1506,1525]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="15" pageNumber="356">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from the northern half of the Gaoligong Shan in western
<collectingRegion id="49BBF869FF9AFF863E29658EFCF67147" box="[638,824,1535,1559]" country="China" name="Yunnan" pageId="15" pageNumber="356">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF9AFF863F14658EFC487147" box="[835,902,1535,1559]" name="China" pageId="15" pageNumber="356">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,315 @@
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<mods:title id="F63A06AA7F9E8D640E7EEE81803376EE">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="74A0DDC8CFEDE8F8E6DD663B99BA863D">Deuve, Thierry</mods:namePart>
<mods:affiliation id="59D65C14996DC7197E240261F3FF8810">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="1E15394538BBBB0C969BAEE1B0A1795D">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="51728E8800ADC1D12B0BB713A4782846">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<taxonomicName id="4C7F4D08FF9AFF863C2C664FFDD97107" ID-CoL="5TT3D" authority="Deuve, 1992" authorityName="Deuve" authorityYear="1992" box="[123,535,1598,1623]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="wuyipeng">
<emphasis id="B90BEA99FF9AFF863C2C664FFDD97107" bold="true" box="[123,535,1598,1623]" pageId="15" pageNumber="356">
<emphasis id="B90BEA99FF9AFF863C2C664FFE4B7107" bold="true" box="[123,389,1598,1623]" italics="true" pageId="15" pageNumber="356">Agonotrechus wuyipeng</emphasis>
Deuve, 1992
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9AFF863C2C662EFE887125" box="[123,326,1631,1653]" pageId="15" pageNumber="356" type="description">
<paragraph id="8BC0368BFF9AFF863C2C662EFE887125" blockId="15.[123,535,1598,1653]" box="[123,326,1631,1653]" pageId="15" pageNumber="356">
(
<figureCitation id="13442A0EFF9AFF863CD5662EFF087124" box="[130,198,1631,1653]" captionStart="FIGURE" captionStartId="17.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1599]" captionTargetId="figure-14@17.[120,1176,216,1599]" captionTargetPageId="17" captionText="FIGURE 8. Agonotrechus wuyipeng Deuve; a. Dorsal habitus (CASENT1010854). b. Median lobe of aedeagus of male (CASENT1010854), left lateral aspect. c. Map of locality records (red circles) for A. wuyipeng in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155297" httpUri="https://zenodo.org/record/13155297/files/figure.png" pageId="15" pageNumber="356">Figs. 8</figureCitation>
,
<figureCitation id="13442A0EFF9AFF863C86662EFF367125" box="[209,248,1631,1653]" captionStart="FIGURE" captionStartId="98.[171,183,1623,1642]" captionTargetBox="[144,1152,216,1605]" captionTargetId="figure-11@98.[144,1152,216,1605]" captionTargetPageId="98" captionText="FIGURE 36. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Fugong County, Shiyueliang Township, 1 km E of Shibali on Shibali Road, elevation 2400 m; habitat in which specimens of Agonotrechus fugongensis sp. nov. were collected. b. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp, elevation 2770 m; habitat in which specimens of Agonotrechus wuyipeng Deuve were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155356" httpUri="https://zenodo.org/record/13155356/files/figure.png" pageId="15" pageNumber="356">36b</figureCitation>
,
<figureCitation id="13442A0EFF9AFF863D55662EFEF17125" box="[258,319,1631,1653]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStartId-0="106.[171,183,1674,1693]" captionStartId-1="107.[171,183,1629,1648]" captionStartId-2="108.[171,183,1359,1378]" captionStartId-3="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,974,1660]" captionTargetBox-1="[120,1176,245,1580]" captionTargetBox-2="[120,1176,228,1340]" captionTargetBox-3="[197,1099,216,1598]" captionTargetId-0="figure-14@106.[120,1176,974,1660]" captionTargetId-1="figure-14@107.[120,1176,216,1608]" captionTargetId-2="figure-12@108.[120,1176,228,1340]" captionTargetId-3="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="106" captionTargetPageId-1="107" captionTargetPageId-2="108" captionTargetPageId-3="109" captionText-0="FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km." captionText-1="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-2="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-3="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155376" figureDoi-1="http://doi.org/10.5281/zenodo.13155378" figureDoi-2="http://doi.org/10.5281/zenodo.13155380" figureDoi-3="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155376/files/figure.png" httpUri-1="https://zenodo.org/record/13155378/files/figure.png" httpUri-2="https://zenodo.org/record/13155380/files/figure.png" httpUri-3="https://zenodo.org/record/13155382/files/figure.png" pageId="15" pageNumber="356">4548</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF9AFF863C2C66FBFEA471ED" pageId="15" pageNumber="356" type="reference_group">
<paragraph id="8BC0368BFF9AFF863C2C66FBFEA471ED" blockId="15.[123,1174,1674,1726]" pageId="15" pageNumber="356">
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<treatmentCitation id="0ADE109AFF9AFF863C2C66FBFDD471F0" author="DEUVE, T." box="[123,538,1674,1696]" page="172" pageId="15" pageNumber="356" year="1992">
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<emphasis id="B90BEA99FF9AFF863C2C66FBFEAF71CF" box="[123,353,1674,1695]" italics="true" pageId="15" pageNumber="356">Agonotrechus wuyipeng</emphasis>
<materialsCitation id="3B173CD6FF9AFF863D3166FBFDD471F0" box="[358,538,1674,1696]" country="China" elevation="2500" location="Wuyipeng" municipality="Wolong" pageId="15" pageNumber="356" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<bibRefCitation id="EFEE4B7AFF9AFF863D3166FBFDD471F0" author="DEUVE, T." box="[358,538,1674,1696]" pageId="15" pageNumber="356" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve, 1992b:172</bibRefCitation>
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.
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, a male, in IOZ. Type locality:
<collectingCountry id="F368761BFF9AFF863FF066FBFC2D71F0" box="[935,995,1674,1696]" name="China" pageId="15" pageNumber="356">China</collectingCountry>
,
<collectingRegion id="49BBF869FF9AFF863FBC66FBFBF571F0" box="[1003,1083,1674,1696]" country="China" name="Sichuan" pageId="15" pageNumber="356">Sichuan</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FF9AFF86381366FBFB5C71F0" box="[1092,1170,1674,1696]" pageId="15" pageNumber="356">Wolong</collectingMunicipality>
,
<location id="8EA06050FF9AFF863CFC66D9FEDF71EE" LSID="urn:lsid:plazi:treatment:03D6879DFF9AFF993C2C664FFCFE710D:8EA06050FF9AFF863CFC66D9FEDF71EE" box="[171,273,1704,1726]" country="China" municipality="Wolong" name="Wuyipeng" pageId="15" pageNumber="356" stateProvince="Sichuan">Wuyipeng</location>
,
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<elevation id="0052D1B8FF9AFF863D4D66D9FEA971ED" box="[282,359,1704,1726]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="356" unit="m" value="2500.0">2500 m</elevation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="C3656500FF85FF993CFC60AAFBD17526" pageId="16" pageNumber="357" type="diagnosis">
<paragraph id="8BC0368BFF85FF993CFC60AAFBD17526" blockId="16.[123,1174,219,1629]" pageId="16" pageNumber="357">
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<emphasis id="B90BEA99FF85FF993CFC60AAFEEE77A1" bold="true" box="[171,288,219,243]" pageId="16" pageNumber="357">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFF85FF993E1B60AAFD5577A3" box="[588,667,219,243]" captionStart="FIGURE" captionStartId="17.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1599]" captionTargetId="figure-14@17.[120,1176,216,1599]" captionTargetPageId="17" captionText="FIGURE 8. Agonotrechus wuyipeng Deuve; a. Dorsal habitus (CASENT1010854). b. Median lobe of aedeagus of male (CASENT1010854), left lateral aspect. c. Map of locality records (red circles) for A. wuyipeng in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155297" httpUri="https://zenodo.org/record/13155297/files/figure.png" pageId="16" pageNumber="357">Fig. 8a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL =
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), fully-winged, dorsum piceous to reddish brown and distinctly iridescent; head slender, eyes markedly projected; clypeus with four or six setae; mentum and submentum incompletely fused, submentum with six or eight setae; mandibles with four to six small setae along the dorsolateral margin of the scrobe; pronotum transverse (ratio PW/PL = 1.30), with lateral explanation broader, especially basally, basal angles rectangular and sharp with a small apical tooth projected laterally; elytra broad, with all discal striae evident and finely punctate, striae 1 to 5 moderately impressed, striae 6 to 8 more faintly impressed, parascutellar striole rather long, recurrent stria continuous anteriorly with stria 5, intervals only faintly convex, two discal setae (anterior and middle setae) present, the anterior seta inserted within basal one-sixth of elytra in stria 3, the middle seta near elytral middle in stria 3; median lobe of aedeagus of male (
<figureCitation id="13442A0EFF85FF993F50624CFC997505" box="[775,855,573,597]" captionStart="FIGURE" captionStartId="17.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1599]" captionTargetId="figure-14@17.[120,1176,216,1599]" captionTargetPageId="17" captionText="FIGURE 8. Agonotrechus wuyipeng Deuve; a. Dorsal habitus (CASENT1010854). b. Median lobe of aedeagus of male (CASENT1010854), left lateral aspect. c. Map of locality records (red circles) for A. wuyipeng in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155297" httpUri="https://zenodo.org/record/13155297/files/figure.png" pageId="16" pageNumber="357">Fig. 8b</figureCitation>
) long and slender, with apex short, recurved dorsally and bluntly pointed, endophallus with a spoon-shaped sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF85FF993CFC620FFCFE710D" pageId="16" pageNumber="357" type="discussion">
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<emphasis id="B90BEA99FF85FF993CFC620FFEE675C4" bold="true" box="[171,296,638,662]" pageId="16" pageNumber="357">COMMENTS</emphasis>
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.— The polymorphism we observed in the number of setae on both the clypeus (four or six) and submentum (six or eight) among specimens from the Gaoligong
<collectingRegion id="49BBF869FF85FF993F8A62EFFBDC75E6" box="[989,1042,670,694]" country="Myanmar" name="Shan" pageId="16" pageNumber="357">Shan</collectingRegion>
populations was surprising.
<bibRefCitation id="EFEE4B7AFF85FF993D7A62CFFD967587" author="BELOUSOV, I. A. &amp; I. I. KABAK" box="[301,600,702,727]" pageId="16" pageNumber="357" pagination="15 - 86" refId="ref58192" refString="BELOUSOV, I. A., AND I. I. KABAK. 2003. New Trechini from China (Coleoptera, Carabidae). Tethys Entomological Research 8: 15 - 86." type="journal article" year="2003">Belousov &amp; Kabak (2003)</bibRefCitation>
described
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<emphasis id="B90BEA99FF85FF993E8162CFFC727586" box="[726,956,702,726]" italics="true" pageId="16" pageNumber="357">Agonotrechus dubius</emphasis>
</taxonomicName>
, based on a single female from
<collectingRegion id="49BBF869FF85FF993D5062AEFE7B75A7" box="[263,437,735,759]" country="China" name="Gansu" pageId="16" pageNumber="357">Gansu Province</collectingRegion>
, and noted similarities with
<taxonomicName id="4C7F4D08FF85FF993EB062AEFCA575A6" box="[743,875,735,758]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="wuyipeng">
<emphasis id="B90BEA99FF85FF993EB062AEFCA575A6" box="[743,875,735,758]" italics="true" pageId="16" pageNumber="357">A. wuyipeng</emphasis>
</taxonomicName>
. We have examined a male specimen from
<collectingRegion id="49BBF869FF85FF993D72628EFE2C7447" box="[293,482,767,791]" country="China" name="Shaanxi" pageId="16" pageNumber="357">Shaanxi Province</collectingRegion>
(Ningshan County, Huoditang Township,
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, collected by Matt Brantley on
<date id="FFC1104BFF85FF993D13636EFE057467" box="[324,459,799,823]" pageId="16" pageNumber="357" value="2005-07-09">9 July 2005</date>
) which shares features described for the
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of
<taxonomicName id="4C7F4D08FF85FF993871636EFB417467" authorityName="Belousov &amp; Kabak" authorityYear="2003" box="[1062,1167,799,823]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B90BEA99FF85FF993871636EFB417467" box="[1062,1167,799,823]" italics="true" pageId="16" pageNumber="357">A. dubius</emphasis>
</taxonomicName>
, including six setae on the submentum, which
<bibRefCitation id="EFEE4B7AFF85FF993E3F634EFC4B7407" author="BELOUSOV, I. A. &amp; I. I. KABAK" box="[616,901,831,855]" pageId="16" pageNumber="357" pagination="15 - 86" refId="ref58192" refString="BELOUSOV, I. A., AND I. I. KABAK. 2003. New Trechini from China (Coleoptera, Carabidae). Tethys Entomological Research 8: 15 - 86." type="journal article" year="2003">Belousov &amp; Kabak (2003)</bibRefCitation>
contrasted with the eight setae reported for
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<emphasis id="B90BEA99FF85FF993D6C6311FE717427" box="[315,447,864,887]" italics="true" pageId="16" pageNumber="357">A. wuyipeng</emphasis>
</taxonomicName>
members (
<bibRefCitation id="EFEE4B7AFF85FF993E626311FD077427" author="DEUVE, T." box="[565,713,863,888]" pageId="16" pageNumber="357" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve 1992b</bibRefCitation>
). Together with other similarities, the polymorphism in this feature found among Gaoligong
<collectingRegion id="49BBF869FF85FF993EC063F1FD0274C8" box="[663,716,896,920]" country="Myanmar" name="Shan" pageId="16" pageNumber="357">Shan</collectingRegion>
specimens suggests that these two species may be better treated as conspecific, with
<taxonomicName id="4C7F4D08FF85FF993E1B63D1FD7D74E8" authorityName="Belousov &amp; Kabak" authorityYear="2003" box="[588,691,928,952]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B90BEA99FF85FF993E1B63D1FD7D74E8" box="[588,691,928,952]" italics="true" pageId="16" pageNumber="357">A. dubius</emphasis>
</taxonomicName>
as a distinct subspecies, members of which have smaller size, relatively wider pronota, and more coarsely punctate elytral discal striae but are otherwise similar to members of the nominate form.
</paragraph>
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<emphasis id="B90BEA99FF85FF993CFC6470FE687347" bold="true" box="[171,422,1025,1049]" pageId="16" pageNumber="357">
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<smallCapsWord id="8D26A057FF85FF993D456475FE687347" baselines="1042" box="[274,422,1028,1047]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="16" pageNumber="357">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Specimens of this species were collected in daytime from under stones in shaded roadside and trailside areas with scattered grasses at elevations ranging from
<quantity id="4C879B6EFF85FF9938376450FF287309" metricMagnitude="3" metricUnit="m" metricValue="2.7285" metricValueMax="2.77" metricValueMin="2.687" pageId="16" pageNumber="357" unit="m" value="2728.5" valueMax="2770.0" valueMin="2687.0">2687 to 2770 m</quantity>
(
<figureCitation id="13442A0EFF85FF993CA46430FE9F7309" box="[243,337,1089,1113]" captionStart="FIGURE" captionStartId="98.[171,183,1623,1642]" captionTargetBox="[144,1152,216,1605]" captionTargetId="figure-11@98.[144,1152,216,1605]" captionTargetPageId="98" captionText="FIGURE 36. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Fugong County, Shiyueliang Township, 1 km E of Shibali on Shibali Road, elevation 2400 m; habitat in which specimens of Agonotrechus fugongensis sp. nov. were collected. b. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp, elevation 2770 m; habitat in which specimens of Agonotrechus wuyipeng Deuve were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155356" httpUri="https://zenodo.org/record/13155356/files/figure.png" pageId="16" pageNumber="357">Fig. 36b</figureCitation>
). Members of no other
<taxonomicName id="4C7F4D08FF85FF993E116430FD177309" authorityName="Jeannel" authorityYear="1923" box="[582,729,1089,1113]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF85FF993E116430FD177309" box="[582,729,1089,1113]" italics="true" pageId="16" pageNumber="357">Agonotrechus</emphasis>
</taxonomicName>
or other trechine species have been found syntopic with those of
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<emphasis id="B90BEA99FF85FF993D276413FE3B7329" box="[368,501,1122,1145]" italics="true" pageId="16" pageNumber="357">A. wuyipeng</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0368BFF85FF993CFC64F0FED9726A" blockId="16.[123,1174,219,1629]" pageId="16" pageNumber="357">
<emphasis id="B90BEA99FF85FF993CFC64F0FDBA73C8" bold="true" box="[171,628,1153,1177]" pageId="16" pageNumber="357">
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<smallCapsWord id="8D26A057FF85FF993D0C64F4FE2173C8" baselines="1171" box="[347,495,1157,1176]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="16" pageNumber="357">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FF85FF993DA264F4FD8A73C8" baselines="1171" box="[501,580,1157,1176]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="16" pageNumber="357">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FF85FF993E1D64F4FDBA73C8" baselines="1171" box="[586,628,1157,1176]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="16" pageNumber="357">THE</smallCapsWord>
</emphasis>
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<emphasis id="B90BEA99FF85FF993E2D64F0FD3173C8" bold="true" box="[634,767,1153,1177]" pageId="16" pageNumber="357">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FF85FF993F5264F0FDA973EA" location="We" pageId="16" pageNumber="357" specimenCount="20" specimenCount-female="3" specimenCount-male="7" stateProvince="Shan">
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<emphasis id="B90BEA99FF85FF993F5264F0FC8E73C8" bold="true" box="[773,832,1153,1177]" pageId="16" pageNumber="357">
<collectingRegion id="49BBF869FF85FF993F5264F0FC8E73C8" box="[773,832,1153,1177]" country="Myanmar" name="Shan" pageId="16" pageNumber="357">SHAN</collectingRegion>
</emphasis>
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.—
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.
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examined a total of
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(
<specimenCount id="9D79FD02FF85FF993D0764D3FE6573EA" box="[336,427,1186,1210]" count="7" pageId="16" pageNumber="357" type="male">7 males</specimenCount>
and
<specimenCount id="9D79FD02FF85FF993DB864D3FDAF73EA" box="[495,609,1186,1210]" count="3" pageId="16" pageNumber="357" type="female">3 females</specimenCount>
)
</materialsCitation>
<materialsCitation id="3B173CD6FF85FF993E2164D3FED9726A" collectingDate="2002-05-02" collectingDateMax="2004-05-01" collectingDateMin="2002-05-02" collectionCode="MNHN, CAS, IOZ" collectorName="H. B. Liang &amp; Qiqi Trail &amp; No &amp; Bridge &amp; W. D. Ba" county="Fugong County" elevation="2687" latitude="27.715" location="Gongshan County" longLatPrecision="1" longitude="98.50222" pageId="16" pageNumber="357" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="SHAN">
from the following localities:
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<collectingCounty id="62A14E07FF85FF993F8164D3FB4073EA" box="[982,1166,1186,1210]" pageId="16" pageNumber="357">Fugong County</collectingCounty>
:
</emphasis>
<taxonomicName id="4C7F4D08FF85FF993C2C64B3FEA7738A" authority="Township" authorityName="Township" box="[123,361,1218,1242]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="shiyueliang">Shiyueliang Township</taxonomicName>
(
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above Shibali on Yaping Road
</locationDeviation>
,
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<elevation id="0052D1B8FF85FF993F5364B3FC987389" box="[772,854,1218,1242]" metricMagnitude="3" metricUnit="m" metricValue="2.687" pageId="16" pageNumber="357" unit="m" value="2687.0">2687 m</elevation>
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,
<date id="FFC1104BFF85FF993F3664B3FC2A738A" box="[865,996,1218,1242]" pageId="16" pageNumber="357" value="2004-05-01">
<collectingDate id="EF85E9A3FF85FF993F3664B3FC2A738A" box="[865,996,1218,1242]" pageId="16" pageNumber="357" value="2004-05-01">1 May 2004</collectingDate>
</date>
,
<collectorName id="268A535DFF85FF993FB864B3FBAB738A" box="[1007,1125,1218,1242]" pageId="16" pageNumber="357">H.B. Liang</collectorName>
collector [
<specimenCount id="9D79FD02FF85FF993C866493FEF473AA" box="[209,314,1250,1274]" count="1" pageId="16" pageNumber="357" type="female">1 female</specimenCount>
;
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].
<emphasis id="B90BEA99FF85FF993D976493FD5573AA" bold="true" box="[448,667,1250,1274]" italics="true" pageId="16" pageNumber="357">
<location id="8EA06050FF85FF993D976493FD5A73AA" LSID="urn:lsid:plazi:treatment:03D6879DFF9AFF993C2C664FFCFE710D:8EA06050FF85FF993D976493FD5A73AA" box="[448,660,1250,1274]" county="Fugong County" latitude="27.715" longLatPrecision="1" longitude="98.50222" name="Gongshan County" pageId="16" pageNumber="357" stateProvince="SHAN">Gongshan County</location>
:
</emphasis>
<collectorName id="268A535DFF85FF993EFC6493FCD373AA" box="[683,797,1250,1274]" pageId="16" pageNumber="357">Qiqi Trail</collectorName>
at
<collectorName id="268A535DFF85FF993F066492FCBC73AA" box="[849,882,1251,1274]" pageId="16" pageNumber="357">No</collectorName>
12
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,
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/
<geoCoordinate id="EE4B504CFF85FF993C2C6573FF32724A" box="[123,252,1282,1306]" degrees="98.50222" direction="east" orientation="longitude" pageId="16" pageNumber="357" precision="1" value="98.50222">E98.50222°</geoCoordinate>
,
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</quantity>
,
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<collectingDate id="EF85E9A3FF85FF993D336572FE29724B" box="[356,487,1283,1307]" pageId="16" pageNumber="357" value="2002-05-02">2 May 2002</collectingDate>
</date>
,
<collectorName id="268A535DFF85FF993DA66572FDA6724A" box="[497,616,1282,1306]" pageId="16" pageNumber="357">H.B. Liang</collectorName>
&amp;
<collectorName id="268A535DFF85FF993EDE6572FD2B724A" box="[649,741,1283,1306]" pageId="16" pageNumber="357">W.D. Ba</collectorName>
collectors [
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and
<specimenCount id="9D79FD02FF85FF993FB16572FB9E724A" box="[998,1104,1282,1307]" count="2" pageId="16" pageNumber="357" type="female">2 females</specimenCount>
;
<collectionCode id="ED6EAE4EFF85FF99380C6572FB5F724B" box="[1115,1169,1283,1307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="357" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF85FF993C2C6552FF64726B" box="[123,170,1315,1339]" pageId="16" pageNumber="357">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF85FF993CE26552FEC3726A" box="[181,269,1315,1338]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="357">MNHN</collectionCode>
].
</materialsCitation>
</paragraph>
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Specimens of this species were collected only in the northern half of the study area (Core Areas 2 and 3) and only on the eastern side of the mountain range. This distribution pattern may be an artifact of inadequate sampling on the western slope of the mountain range in the north, much of which is in
<collectingCountry id="F368761BFF85FF993D4C65D5FE4872EC" box="[283,390,1444,1468]" name="Myanmar" pageId="16" pageNumber="357">Myanmar</collectingCountry>
. The geographical range of this species overlaps that of
<taxonomicName id="4C7F4D08FF85FF993FA765D5FB4072EC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1008,1166,1444,1468]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="fugongensis">
<emphasis id="B90BEA99FF85FF993FA765D5FB4072EC" box="[1008,1166,1444,1468]" italics="true" pageId="16" pageNumber="357">A. fugongensis</emphasis>
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, but members of the latter species appear to occupy slightly lower elevations in the same general areas.
</paragraph>
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</emphasis>
.—
<materialsCitation id="3B173CD6FF85FF993E286675FCE2710D" location="Gaoligong" pageId="16" pageNumber="357" specimenCount="1" stateProvince="Sichuan">
<figureCitation id="13442A0EFF85FF993E286675FD03714C" box="[639,717,1540,1564]" captionStart="FIGURE" captionStartId="106.[171,183,1674,1693]" captionTargetBox="[120,1176,974,1660]" captionTargetId="figure-14@106.[120,1176,974,1660]" captionTargetPageId="106" captionText="FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.13155376" httpUri="https://zenodo.org/record/13155376/files/figure.png" pageId="16" pageNumber="357">Fig. 45</figureCitation>
. This species currently is known only from the
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locality in northcentral
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and the northern half of the
<location id="8EA06050FF85FF993F396655FC11716C" LSID="urn:lsid:plazi:treatment:03D6879DFF9AFF993C2C664FFCFE710D:8EA06050FF85FF993F396655FC11716C" box="[878,991,1572,1596]" name="Gaoligong" pageId="16" pageNumber="357" stateProvince="Sichuan">Gaoligong</location>
<collectingRegion id="49BBF869FF85FF993FB06655FBD2716C" box="[999,1052,1572,1596]" country="Myanmar" name="Shan" pageId="16" pageNumber="357">Shan</collectingRegion>
in western
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, but it probably occurs in the intervening region as well
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.
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,218 @@
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<mods:title id="22FD7C69EFA84A9FB1F5A901E2D2E685">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:affiliation id="70A5006284F03E09E0B317054E639DF4">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="1DC80C2AD048F98883EFAF00DFB6E9A2">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FF9EFF823D95644FFC807306" bold="true" box="[450,846,1085,1110]" pageId="11" pageNumber="352">
Genus
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<emphasis id="B90BEA99FF9EFF823E45644CFD627305" bold="true" box="[530,684,1085,1109]" italics="true" pageId="11" pageNumber="352">Agonotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF9EFF823EE4644FFC807306" author="JEANNEL, R." box="[691,846,1086,1110]" pageId="11" pageNumber="352" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel, 1923</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<treatmentCitation id="0ADE109AFF9EFF823DBF641AFCEB73D0" author="JEANNEL, R." box="[488,805,1131,1153]" page="428" pageId="11" pageNumber="352" year="1923">
<taxonomicName id="4C7F4D08FF9EFF823DBF641AFCEB73D0" ID-CoL="8KV2Q" authority="Jeannel, 1923: 428" authorityName="Jeannel" authorityPageNumber="428" authorityYear="1923" box="[488,805,1131,1153]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF9EFF823DBF641AFDA273D0" box="[488,620,1131,1152]" italics="true" pageId="11" pageNumber="352">Agonotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF9EFF823E24641AFCEB73D0" author="JEANNEL, R." box="[627,805,1131,1153]" pageId="11" pageNumber="352" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel, 1923:428</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<treatmentCitation id="0ADE109AFF9EFF823DBA64FAFCD173F0" author="UENO, S. I." box="[493,799,1163,1184]" page="2" pageId="11" pageNumber="352" year="1981">
<taxonomicName id="4C7F4D08FF9EFF823DBA64FAFCD173F0" ID-CoL="8KY83" authority="Ueno, 1981: 2" authorityName="Ueno" authorityPageNumber="2" authorityYear="1981" box="[493,799,1163,1184]" class="Insecta" family="Carabidae" genus="Paragonotrechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF9EFF823DBA64FAFD5D73F0" box="[493,659,1163,1184]" italics="true" pageId="11" pageNumber="352">Paragonotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF9EFF823ECD64FAFCD173F0" author="UENO, S. I." box="[666,799,1163,1184]" pageId="11" pageNumber="352" pagination="1 - 10" refId="ref61106" refString="UENO, S. I. 1981. A remarkable new trechine beetle found in a superficial subterranean habitat near Tokyo, Central Japan. Journal of the Speleological Society of Japan 6: 1 - 10." type="journal article" year="1981">Uéno, 1981:2</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<treatmentCitation id="0ADE109AFF9EFF823DBE64DAFCEA7390" author="UENO, S. I." box="[489,804,1195,1216]" page="188" pageId="11" pageNumber="352" year="1977">
<taxonomicName id="4C7F4D08FF9EFF823DBE64DAFCEA7390" ID-CoL="8KVLF" authority="Ueno, 1977: 188" authorityName="Ueno" authorityPageNumber="188" authorityYear="1977" box="[489,804,1195,1216]" class="Insecta" family="Carabidae" genus="Bhutanotrechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF9EFF823DBE64DAFD4E7390" box="[489,640,1195,1216]" italics="true" pageId="11" pageNumber="352">Bhutanotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF9EFF823ED164DAFCEA7390" author="UENO, S. I." box="[646,804,1195,1216]" pageId="11" pageNumber="352" pagination="175 - 196" refId="ref61071" refString="UENO, S. I. 1977. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Carabidae Subfam. Trechinae. Entomologica Basiliensia 2: 175 - 196." type="journal article" year="1977">Uéno, 1977:188</bibRefCitation>
</taxonomicName>
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.
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.—
<taxonomicName id="4C7F4D08FF9EFF823D3F6495FD0A73AC" ID-CoL="7CNQS" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[360,708,1252,1276]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="birmanicus">
<emphasis id="B90BEA99FF9EFF823D3F6495FDF573AC" box="[360,571,1252,1276]" italics="true" pageId="11" pageNumber="352">Trechus birmanicus</emphasis>
<bibRefCitation id="EFEE4B7AFF9EFF823E156495FD0A73AC" author="BATES, H. W." box="[578,708,1252,1276]" pageId="11" pageNumber="352" pagination="267 - 428" refId="ref58154" refString="BATES, H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428" type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<emphasis id="B90BEA99FF9EFF823CFC6575FEEE724A" bold="true" box="[171,288,1284,1308]" pageId="11" pageNumber="352">DIAGNOSIS</emphasis>
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.— Adults of this genus (
<figureCitation id="13442A0EFF9EFF823E646575FD44724C" box="[563,650,1284,1308]" captionStart="FIGURE" captionStartId="14.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1577]" captionTargetId="figure-11@14.[120,1176,216,1577]" captionTargetPageId="14" captionText="FIGURE 7. Agonotrechus fugongensis sp. nov.; a. Dorsal habitus (CASENT1019979). b. Median lobe of aedeagus of male (CASENT1019979), left lateral aspect. c. Map of locality records (red circles) for A. fugongensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155295" httpUri="https://zenodo.org/record/13155295/files/figure.png" pageId="11" pageNumber="352">Figs. 7a</figureCitation>
<figureCitation id="13442A0EFF9EFF823ECE6575FD7C724C" box="[665,690,1284,1308]" captionStart="FIGURE" captionStartId="19.[171,183,1596,1615]" captionTargetBox="[120,1176,216,1578]" captionTargetId="figure-15@19.[120,1176,216,1579]" captionTargetPageId="19" captionText="FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A. xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155299" httpUri="https://zenodo.org/record/13155299/files/figure.png" pageId="11" pageNumber="352">9a</figureCitation>
) can be recognized by the following combination of character states: size medium to large for a trechine (BL = 4.5 to 7.0 mm) members of most species with full hindwings and large, convex eyes, but those of a few species apterous and with eyes reduced in size and/or convexity, some even microphthalmous or nearly anophthalmous; frons flattened, depressed; dorsolateral margin of mandibular scope with a row of small setae aligned as a sparse comb in some members (
<bibRefCitation id="EFEE4B7AFF9EFF823E0865D5FCBC72ED" author="BELOUSOV, I. A. &amp; I. I. KABAK" box="[607,882,1444,1469]" pageId="11" pageNumber="352" pagination="15 - 86" refId="ref58192" refString="BELOUSOV, I. A., AND I. I. KABAK. 2003. New Trechini from China (Coleoptera, Carabidae). Tethys Entomological Research 8: 15 - 86." type="journal article" year="2003">Belousov &amp; Kabak 2003</bibRefCitation>
); right mandible (
<figureCitation id="13442A0EFF9EFF82386465D5FB4072EC" box="[1075,1166,1444,1468]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="11" pageNumber="352">Fig. 16a</figureCitation>
) bidentate, the premolar tooth fused with the retinaculum, from which the anterior tooth is distinctly projected and in a forward position and the posterior tooth is absent or flush; left mandible with only a short, subconical, more or less trifid ridge; clypeus with six setae in most members; labium with anterior margin not or only slightly concave; mentum fused with submentum, incompletely in some members; submentum with six setae in most members, but some with eight setae; pronotum little narrowed basally, with lateral explanation distinctly broadened basally in many members; elytra with discal striae varied, from complete to effaced, punctate in many members, parascutellar striole very long in many members; anterior discal seta present in all members and inserted at the basal one-fifth or one-sixth of elytra, middle discal seta present or absent, preapical seta present at or near stria
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most members, absent from a very few members; legs slender, protibiae furrowed, the basal two protarsomeres dilated and toothed; abdominal ventrites glabrous except for usual paramedial setae; aedeagus with a spoon-shaped copulatory piece.
</paragraph>
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</emphasis>
.— Members of the first known
<taxonomicName id="4C7F4D08FF99FF853E82612FFC727626" box="[725,956,350,374]" pageId="12" pageNumber="353">
<emphasis id="B90BEA99FF99FF853E82612FFCA67626" box="[725,872,350,374]" italics="true" pageId="12" pageNumber="353">Agonotrechus</emphasis>
species
</taxonomicName>
had only the anteri- or discal seta of the elytron inserted at the basal one-fifth or one-sixth. Consequently, this feature was used by
<bibRefCitation id="EFEE4B7AFF99FF853D5F61EEFE6676E7" author="JEANNEL, R." box="[264,424,415,439]" pageId="12" pageNumber="353" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel (1923)</bibRefCitation>
to define his new genus. Members of
<taxonomicName id="4C7F4D08FF99FF853F1B61EEFB5B76E7" authority="Ueno (1981)" authorityName="Ueno" authorityYear="1981" box="[844,1173,415,439]" class="Insecta" family="Carabidae" genus="Paragonotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853F1B61EEFBCA76E7" box="[844,1028,415,439]" italics="true" pageId="12" pageNumber="353">Paragonotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF99FF85385B61EEFB5B76E7" author="UENO, S. I." box="[1036,1173,415,439]" pageId="12" pageNumber="353" pagination="1 - 10" refId="ref61106" refString="UENO, S. I. 1981. A remarkable new trechine beetle found in a superficial subterranean habitat near Tokyo, Central Japan. Journal of the Speleological Society of Japan 6: 1 - 10." type="journal article" year="1981">Uéno (1981)</bibRefCitation>
</taxonomicName>
are very similar and differ only in having a more slender body form and the parascutellar striole very long. Those of
<taxonomicName id="4C7F4D08FF99FF853D0E6191FD5A76A8" authority="Ueno (1977)" authorityName="Ueno" authorityYear="1977" box="[345,660,480,504]" class="Insecta" family="Carabidae" genus="Bhutanotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853D0E6191FDCF76A8" box="[345,513,480,504]" italics="true" pageId="12" pageNumber="353">Bhutanotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF99FF853E5E6191FD5A76A8" author="UENO, S. I." box="[521,660,480,504]" pageId="12" pageNumber="353" pagination="175 - 196" refId="ref61071" refString="UENO, S. I. 1977. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Carabidae Subfam. Trechinae. Entomologica Basiliensia 2: 175 - 196." type="journal article" year="1977">Uéno (1977)</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="F368761BFF99FF853F1B6191FC5276A8" box="[844,924,480,504]" name="Bhutan" pageId="12" pageNumber="353">Bhutan</collectingCountry>
, were distinguished by having the elytral discal striae effaced and a second elytral discal seta (the middle seta) present. The presence of the middle seta is a plesiotypic feature in
<taxonomicName id="4C7F4D08FF99FF853E936250FCEE7569" authorityName="Bonelli" authorityYear="1810" box="[708,800,545,569]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
, but species with members having two discal setae sometimes have been grouped together or assigned to genus
<taxonomicName id="4C7F4D08FF99FF853FBA6233FB5B750A" authorityName="Ueno" authorityYear="1977" box="[1005,1173,578,602]" class="Insecta" family="Carabidae" genus="Bhutanotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853FBA6233FB5B750A" box="[1005,1173,578,602]" italics="true" pageId="12" pageNumber="353">Bhutanotrechus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFEE4B7AFF99FF853CD56212FED8752B" author="DEUVE, T." box="[130,278,611,635]" pageId="12" pageNumber="353" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve 1992b</bibRefCitation>
,
<bibRefCitation id="EFEE4B7AFF99FF853D756212FE95752B" author="DEUVE, T." box="[290,347,611,635]" pageId="12" pageNumber="353" pagination="5 - 18" refId="ref58882" refString="DEUVE, T. 1995. Contribution a l'inventaire des Trechidae Trechinae de Chine et de Thailande (Coleoptera). Revue Francaise d'Entomologie (N. S.) 17: 5 - 18." type="journal article" year="1995">1995</bibRefCitation>
). Genus
<taxonomicName id="4C7F4D08FF99FF853DEC6212FD80752B" authorityName="Jeannel" authorityYear="1923" box="[443,590,611,635]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853DEC6212FD80752B" box="[443,590,611,635]" italics="true" pageId="12" pageNumber="353">Agonotrechus</emphasis>
</taxonomicName>
is really a homogenous group, despite the differences among its members in impression of the elytral discal striae, length of the parascutellar stiole, number of elytral discal setae and size of the eyes, differences in the last feature being associated with differential flight capability. There appears to be no reason to maintain
<taxonomicName id="4C7F4D08FF99FF853FFC62B4FBAD758D" authorityName="Ueno" authorityYear="1981" box="[939,1123,709,733]" class="Insecta" family="Carabidae" genus="Paragonotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853FFC62B4FBAD758D" box="[939,1123,709,733]" italics="true" pageId="12" pageNumber="353">Paragonotrechus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FF99FF853C2C6294FEED75AD" authorityName="Ueno" authorityYear="1977" box="[123,291,741,765]" class="Insecta" family="Carabidae" genus="Bhutanotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853C2C6294FEED75AD" box="[123,291,741,765]" italics="true" pageId="12" pageNumber="353">Bhutanotrechus</emphasis>
</taxonomicName>
as distinct genera.
</paragraph>
<paragraph id="8BC0368BFF99FF853CFC6377FD4B740F" blockId="12.[123,1174,219,1027]" pageId="12" pageNumber="353">
The bidentate dentition of the right mandible of
<taxonomicName id="4C7F4D08FF99FF853EE16377FC87744E" authorityName="Jeannel" authorityYear="1923" box="[694,841,774,798]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853EE16377FC87744E" box="[694,841,774,798]" italics="true" pageId="12" pageNumber="353">Agonotrechus</emphasis>
</taxonomicName>
, with the premolar tooth completely fused with retinaculum, distinguishes this group from members of the
<taxonomicName id="4C7F4D08FF99FF853FE06356FBE8746E" authorityName="Jeannel" authorityYear="1923" box="[951,1062,807,830]" class="Insecta" family="Carabidae" genus="Stevensius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF99FF853FE06356FBE8746E" box="[951,1062,807,830]" italics="true" pageId="12" pageNumber="353">Stevensius</emphasis>
</taxonomicName>
Complex, with which
<bibRefCitation id="EFEE4B7AFF99FF853CAD6336FE5C740F" author="JEANNEL, R." box="[250,402,839,863]" pageId="12" pageNumber="353" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel (1923</bibRefCitation>
,
<bibRefCitation id="EFEE4B7AFF99FF853DCA6336FE1A740F" author="JEANNEL, R." box="[413,468,839,863]" pageId="12" pageNumber="353" pagination="1 - 808" refId="ref59586" refString="JEANNEL, R. 1928. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres. (Troisieme livraison). L'Abeille 35: 1 - 808." type="journal article" year="1928">1928</bibRefCitation>
) had grouped it.
</paragraph>
<paragraph id="8BC0368BFF99FF853CFC6319FDD87353" blockId="12.[123,1174,219,1027]" pageId="12" pageNumber="353">
<emphasis id="B90BEA99FF99FF853CFC6319FE3F742E" bold="true" box="[171,497,872,896]" pageId="12" pageNumber="353">
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<smallCapsWord id="8D26A057FF99FF853D0A631AFE3F742E" baselines="889" box="[349,497,875,894]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="12" pageNumber="353">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This genus, which at present includes 16 species, is known from the southeastern part of the Palearctic Region and northern edge of the Oriental Region, from
<collectingCountry id="F368761BFF99FF853C2C63D8FF757491" box="[123,187,937,961]" name="Nepal" pageId="12" pageNumber="353">Nepal</collectingCountry>
eastward to southern Gansu Province in the north and
<collectingCountry id="F368761BFF99FF853F4163DBFC4E7492" box="[790,896,938,962]" name="Myanmar" pageId="12" pageNumber="353">Myanmar</collectingCountry>
,
<collectingCountry id="F368761BFF99FF853FDA63D8FC247491" box="[909,1002,937,961]" name="Vietnam" pageId="12" pageNumber="353">Vietnam</collectingCountry>
, and Hubei and Shaanxi Provinces; and one species has been described from
<collectingCountry id="F368761BFF99FF853F4E63BBFC9774B1" box="[793,857,970,993]" name="Japan" pageId="12" pageNumber="353">Japan</collectingCountry>
. The study area is within the previously known range of this genus.
</paragraph>
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<emphasis id="B90BEA99FFA3FFBF3C2C612AFF1A7623" bold="true" box="[123,212,347,371]" italics="true" pageId="54" pageNumber="395">Trechus</emphasis>
(
<emphasis id="B90BEA99FFA3FFBF3CB3612AFEF37623" bold="true" box="[228,317,347,371]" italics="true" pageId="54" pageNumber="395">Trechus</emphasis>
)
<emphasis id="B90BEA99FFA3FFBF3D19612AFE357623" bold="true" box="[334,507,347,371]" italics="true" pageId="54" pageNumber="395">pseudoqiqiensis</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFA3FFBF3E83612AFCE47623" box="[724,810,347,371]" pageId="54" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA3FFBF3C2C610AFE9C76C1" box="[123,338,379,401]" pageId="54" pageNumber="395" type="description">
<paragraph id="8BC0368BFFA3FFBF3C2C610AFE9C76C1" blockId="54.[123,810,347,401]" box="[123,338,379,401]" pageId="54" pageNumber="395">
(
<figureCitation id="13442A0EFFA3FFBF3CD5610AFF1D76C0" box="[130,211,379,401]" captionStart="FIGURE" captionStartId="55.[171,183,1583,1602]" captionTargetBox="[162,1176,216,1570]" captionTargetId="figure-14@55.[120,1176,216,1570]" captionTargetPageId="55" captionText="FIGURE 21. Trechus pseudoqiqiensis sp. nov.; a. Dorsal habitus (CASENT1014204). b. Map of locality record (red circle) for T. pseudoqiqiensis in the Gaoligong Shan region. Scale lines a = 0.5mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155324" httpUri="https://zenodo.org/record/13155324/files/figure.png" pageId="54" pageNumber="395">Figs. 21</figureCitation>
,
<figureCitation id="13442A0EFFA3FFBF3C8A610AFECD76C1" box="[221,259,379,401]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="54" pageNumber="395">42b</figureCitation>
,
<figureCitation id="13442A0EFFA3FFBF3D59610AFE8576C1" box="[270,331,379,401]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="54" pageNumber="395">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA3FFBF3CFC61D5FED775AC" pageId="54" pageNumber="395" type="materials_examined">
<paragraph id="8BC0368BFFA3FFBF3CFC61D5FD0C756C" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">
<emphasis id="B90BEA99FFA3FFBF3CFC61D5FE9676EA" bold="true" box="[171,344,420,444]" pageId="54" pageNumber="395">
<smallCapsWord id="8D26A057FFA3FFBF3CFC61D5FF2A76EB" baselines="438,438" box="[171,228,420,444]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="54" pageNumber="395">TYPE</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3CBC61D6FE9676EA" baselines="437" box="[235,344,423,442]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="54" pageNumber="395">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFA3FFBF3D2861D5FD0C756C" collectingDate="2004-05-08" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="3290" latitude="27.20676" location="Lishadi Township" longLatPrecision="1" longitude="98.71763" pageId="54" pageNumber="395" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFA3FFBF3D2861D5FE2B76EC" box="[383,485,420,444]" pageId="54" pageNumber="395" type="holotype">Holotype</typeStatus>
, a female, in IOZ, labeled: “CASENT 1014205”/ “
<collectingCountry id="F368761BFFA3FFBF3FA861D5FB9676EC" box="[1023,1112,420,444]" name="China" pageId="54" pageNumber="395">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA3FFBF383561D5FF68768C" country="China" name="Yunnan" pageId="54" pageNumber="395">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA3FFBF3CE861B4FEB2768C" box="[191,380,452,477]" pageId="54" pageNumber="395">Fugong County</collectingCounty>
,
<location id="8EA06050FFA3FFBF3DC261B5FDA0768C" LSID="urn:lsid:plazi:treatment:03D6879DFFA3FFB13C2C612AFEE174F1:8EA06050FFA3FFBF3DC261B5FDA0768C" box="[405,622,452,476]" country="China" county="Fugong County" latitude="27.20676" longLatPrecision="1" longitude="98.71763" name="Lishadi Township" pageId="54" pageNumber="395" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFA3FFBF3ED061B5FB5F768C" box="[647,1169,452,476]" location="Shibali on Yaping Road" pageId="54" pageNumber="395">
<quantity id="4C879B6EFFA3FFBF3ED061B5FD21768C" box="[647,751,452,476]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="54" pageNumber="395" unit="km" value="11.5">11.5 km</quantity>
above Shibali on Yaping Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFA3FFBF3C2C6195FF3176AC" box="[123,255,484,508]" degrees="27.20676" direction="north" orientation="latitude" pageId="54" pageNumber="395" precision="1" value="27.20676">N27.20676°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA3FFBF3D506195FE4976AC" box="[263,391,484,508]" degrees="98.71763" direction="east" orientation="longitude" pageId="54" pageNumber="395" precision="1" value="98.71763">E98.71763°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFA3FFBF3DE76195FDC676AC" box="[432,520,484,508]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="54" pageNumber="395" unit="m" value="3290.0">
<elevation id="0052D1B8FFA3FFBF3DE76195FDC676AC" box="[432,520,484,508]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="54" pageNumber="395" unit="m" value="3290.0">3290 m</elevation>
</quantity>
,
<date id="FFC1104BFFA3FFBF3E416195FD5276AC" box="[534,668,484,509]" pageId="54" pageNumber="395" value="2004-05-08">
<collectingDate id="EF85E9A3FFA3FFBF3E416195FD5276AC" box="[534,668,484,509]" pageId="54" pageNumber="395" value="2004-05-08">8 May 2004</collectingDate>
</date>
Stop #DHK-2004-040 D.H.
<taxonomicName id="4C7F4D08FFA3FFBF3F8B6195FB9676AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[988,1112,484,508]" pageId="54" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFA3FFBF38316195FF2F754C" pageId="54" pageNumber="395">C.E. Griswold</collectorName>
, Liang H.-B., Li X.-Y., &amp; Zhu B.-X. collectors”/ “
<typeStatus id="54C48829FFA3FFBF3EAD6275FC47754C" box="[762,905,516,540]" pageId="54" pageNumber="395" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA3FFBF3FC66275FEEB756C" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="395" phylum="Arthropoda" rank="species" species="pseudoqiqiensis" status="sp. nov.">
<emphasis id="B90BEA99FFA3FFBF3FC66275FB5B754C" box="[913,1173,516,540]" italics="true" pageId="54" pageNumber="395">Trechus pseudoqiqiensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFA3FFBF3D676254FE4D756C" box="[304,387,549,572]" pageId="54" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label].
</materialsCitation>
</paragraph>
<paragraph id="8BC0368BFFA3FFBF3CFC6235FD39752C" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">
<materialsCitation id="3B173CD6FFA3FFBF3CFC6235FD3D752C" country="China" county="Fugong County" elevation="3290" latitude="27.20676" location="Fugong County" longLatPrecision="1" longitude="98.71763" pageId="54" pageNumber="395" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFA3FFBF3CFC6235FE97750B" bold="true" box="[171,345,580,604]" pageId="54" pageNumber="395">
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<typeStatus id="54C48829FFA3FFBF3CFC6235FF2A750B" box="[171,228,580,604]" pageId="54" pageNumber="395">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3CB96239FE97750B" baselines="598" box="[238,345,584,603]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="54" pageNumber="395">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFA3FFBF3DD36235FE08750C" box="[388,454,580,604]" name="China" pageId="54" pageNumber="395">China</collectingCountry>
,
<collectingRegion id="49BBF869FFA3FFBF3D826234FDE7750C" box="[469,553,581,604]" country="China" name="Yunnan" pageId="54" pageNumber="395">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA3FFBF3E606234FD2C750C" box="[567,738,580,605]" pageId="54" pageNumber="395">Fugong County</collectingCounty>
,
<taxonomicName id="4C7F4D08FFA3FFBF3EA76235FC2B750C" authority="Township" authorityName="Township" box="[752,997,580,604]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="395" phylum="Arthropoda" rank="species" species="shiyueliang">Shiyueliang Township</taxonomicName>
,
<locationDeviation id="9A722FA9FFA3FFBF3FA56235FEB3752C" location="Shibali on Yaping Road" pageId="54" pageNumber="395">
<quantity id="4C879B6EFFA3FFBF3FA56235FB82750C" box="[1010,1100,580,604]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="54" pageNumber="395" unit="km" value="11.5">11.5 km</quantity>
above Shibali on Yaping Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFA3FFBF3DDF6215FDC5752C" box="[392,523,612,636]" degrees="27.20676" direction="north" orientation="latitude" pageId="54" pageNumber="395" precision="1" value="27.20676">N27.20676°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA3FFBF3E436215FD5D752C" box="[532,659,612,636]" degrees="98.71763" direction="east" orientation="longitude" pageId="54" pageNumber="395" precision="1" value="98.71763">E98.71763°</geoCoordinate>
,
<quantity id="4C879B6EFFA3FFBF3EC96215FD3D752C" box="[670,755,612,636]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="54" pageNumber="395" unit="m" value="3290.0">
<elevation id="0052D1B8FFA3FFBF3EC96215FD3D752C" box="[670,755,612,636]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="54" pageNumber="395" unit="m" value="3290.0">3290 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA3FFBF3CFC62F5FED775AC" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">
<emphasis id="B90BEA99FFA3FFBF3CFC62F5FE2275CB" bold="true" box="[171,492,644,668]" pageId="54" pageNumber="395">
<smallCapsWord id="8D26A057FFA3FFBF3CFC62F5FEE075CB" baselines="662,662" box="[171,302,644,668]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="54" pageNumber="395">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3D6362F9FE9E75CB" baselines="662" box="[308,336,648,667]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="54" pageNumber="395">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3D0262F9FE6675CB" baselines="662" box="[341,424,648,667]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="54" pageNumber="395">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3DF862F9FE2275CB" baselines="662" box="[431,492,648,667]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="54" pageNumber="395">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFA3FFBF3EBC62F5FC5B75CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[747,917,644,668]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="395" phylum="Arthropoda" rank="species" species="pseudoqiqiensis">
<emphasis id="B90BEA99FFA3FFBF3EBC62F5FC5B75CC" box="[747,917,644,668]" italics="true" pageId="54" pageNumber="395">pseudoqiqiensis</emphasis>
</taxonomicName>
, is a combination of the Greek prefix,
<emphasis id="B90BEA99FFA3FFBF3D4062D5FEAD75ED" box="[279,355,676,701]" italics="true" pageId="54" pageNumber="395">ψΕυδής</emphasis>
(translated into Latin as
<emphasis id="B90BEA99FFA3FFBF3ED762D5FD1A75EC" box="[640,724,676,700]" italics="true" pageId="54" pageNumber="395">pseudo-</emphasis>
) meaning false, and the species epithet,
<taxonomicName id="4C7F4D08FFA3FFBF3C2C62B4FF14758C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,218,709,732]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="395" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA3FFBF3C2C62B4FF14758C" box="[123,218,709,732]" italics="true" pageId="54" pageNumber="395">qiqiensis</emphasis>
</taxonomicName>
, in reference to the similarity of the unique
<typeStatus id="54C48829FFA3FFBF3EE962B5FCD5758C" box="[702,795,708,732]" pageId="54" pageNumber="395" type="holotype">holotype</typeStatus>
female of this species to members of
<taxonomicName id="4C7F4D08FFA3FFBF3CCF6294FEDC75AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[152,274,741,764]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="395" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA3FFBF3CCF6294FEDC75AC" box="[152,274,741,764]" italics="true" pageId="54" pageNumber="395">T. qiqiensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA3FFBF3CFC6375FE55748C" pageId="54" pageNumber="395" type="diagnosis">
<paragraph id="8BC0368BFFA3FFBF3CFC6375FE55748C" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">
<smallCapsWord id="8D26A057FFA3FFBF3CFC6375FEEE744B" baselines="790,790" box="[171,288,772,796]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="54" pageNumber="395">
<emphasis id="B90BEA99FFA3FFBF3CFC6375FEEE744B" bold="true" box="[171,288,772,796]" pageId="54" pageNumber="395">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFA3FFBF3E106375FD6D744D" box="[583,675,772,797]" captionStart="FIGURE" captionStartId="55.[171,183,1583,1602]" captionTargetBox="[162,1176,216,1570]" captionTargetId="figure-14@55.[120,1176,216,1570]" captionTargetPageId="55" captionText="FIGURE 21. Trechus pseudoqiqiensis sp. nov.; a. Dorsal habitus (CASENT1014204). b. Map of locality record (red circle) for T. pseudoqiqiensis in the Gaoligong Shan region. Scale lines a = 0.5mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155324" httpUri="https://zenodo.org/record/13155324/files/figure.png" pageId="54" pageNumber="395">Fig. 21a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: similar to
<taxonomicName id="4C7F4D08FFA3FFBF3F856354FB85746C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[978,1099,805,828]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="395" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA3FFBF3F856354FB85746C" box="[978,1099,805,828]" italics="true" pageId="54" pageNumber="395">T. qiqiensis</emphasis>
</taxonomicName>
in size (BL = 4.0 mm) and most features, except body color paler, more reddish; elytral striae more faintly punctate, almost impunctate toward elytral apex, more deeply impressed on center of disc and not attenuated near apex, recurrent stria less abruptly interrupted by a convexity and nearly continuous anteriorly with stria 7; preapical seta present and inserted on interval 3 next to stria 2 near apical one-fourth of elytra.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA3FFB13CFC6394FEE174F1" lastPageId="56" lastPageNumber="397" pageId="54" pageNumber="395" type="description">
<paragraph id="8BC0368BFFA3FFBF3CFC6394FD92734D" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">
<smallCapsWord id="8D26A057FFA3FFBF3CFC6394FEF474AB" baselines="1015,1014" box="[171,314,997,1021]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="54" pageNumber="395">
<emphasis id="B90BEA99FFA3FFBF3CFC6394FEF474AB" bold="true" box="[171,314,997,1021]" pageId="54" pageNumber="395">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size relatively large (BL = 4.0 mm). Body color bright brownish red, shiny, appendages yellowish-tan, palpi pale yellow.
</paragraph>
<paragraph id="8BC0368BFFA3FFBF3CFC6454FCA5726D" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">
Head. Moderate in size; eyes slightly convex, their diameter about 1.5 times as long as tempora, the latter short, convex and glabrous. Frons not flattened; frontal furrows deep, rounded, not or only slightly attenuated posterior to the eyes. Two pairs of supraorbital setae present, the anteri- or pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave.
<collectingRegion id="49BBF869FFA3FFBF3D6764D4FE4473EC" box="[304,394,1189,1212]" country="Myanmar" name="Chin" pageId="54" pageNumber="395">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFA3FFBF3EE964D4FCD673EC" box="[702,792,1189,1212]" country="Myanmar" name="Chin" pageId="54" pageNumber="395">Mentum</collectingRegion>
distinctly concave at middle, with medial tooth broad and apex slightly bifid, about one-half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae short, with about 2 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 4 about equal in length, antennomere 3 slightly longer.
</paragraph>
<paragraph id="8BC0368BFFA3FFBF3CFC6534FF34712D" blockId="54.[123,1174,420,1726]" pageId="54" pageNumber="395">Pronotum. Transverse (ratio PW/PL = 1.45), slightly narrowed posteriorly, greatest width near anterior one-third; lateral margins broadly rounded, not sinuate subbasally, straightened only immediately anterior to basal angles, the latter small and rectangular. Disc convex, smooth and glabrous, median longitudinal impression finely impressed, interrupted slightly posterior to anteri- or margin but extended to basal margin; basal foveae distinct, but shallower and slightly convex at center; median basal area mainly smooth, but with several small foveae anteriorly near the posterior transverse impression. Basal margin rounded and slightly projected medially. Lateral borders of pronotum slender, narrowly reflexed, lateral grooves distinctly impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFA3FFB13CFC66F4FE087688" blockId="54.[123,1174,420,1726]" lastBlockId="56.[123,1174,219,929]" lastPageId="56" lastPageNumber="397" pageId="54" pageNumber="395">
Elytra. Ovoid and only slightly elongate, humeri not very broad but rounded. Disc convex and smooth. Striae very finely punctate, nearly smooth in apical half, intervals moderately convex; striae 1 to 4 deeply impressed and not attenuated apically, stria 5 to 8 successively more faintly impressed by still evident. Parascutellar striole present, short. Recurrent stria less abruptly interrupted by a convexity and nearly continuous anteriorly with stria 7. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a discal position on interval 3 next to stria
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forward position at apical one-fourth of elytra. Umbilicate setal series with setae of humeral group equidistance from each other and setae of median group insert- ed slightly posterior to middle.
</paragraph>
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21.
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<emphasis id="B90BEA99FFA2FFBE3D5B665EFE127112" box="[268,476,1583,1602]" italics="true" pageId="55" pageNumber="396">Trechus pseudoqiqiensis</emphasis>
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; a. Dorsal habitus (CASENT1014204). b. Map of locality record (red circle) for
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<emphasis id="B90BEA99FFA2FFBE3CE86638FE93710B" box="[191,349,1608,1627]" italics="true" pageId="55" pageNumber="396">T. pseudoqiqiensis</emphasis>
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in the Gaoligong Shan region. Scale lines a = 0.5mm, b = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFADFFB13CFC6190FC4F76A8" blockId="56.[123,1174,219,929]" box="[171,897,480,505]" pageId="56" pageNumber="397">Legs. Slightly short, protibiae with only a faint longitudinal furrow.</paragraph>
<paragraph id="8BC0368BFFADFFB13CFC6270FC8C7569" blockId="56.[123,1174,219,929]" pageId="56" pageNumber="397">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of female apically with two pairs of paramedial setae.</paragraph>
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</emphasis>
.— The unique
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female of this species was found under a stone in an open area cleared of forest but surrounded by bamboo thickets and scattered
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<emphasis id="B90BEA99FFADFFB13FAA6212FBF6752B" box="[1021,1080,611,635]" italics="true" pageId="56" pageNumber="397">Abies</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFADFFB138106212FF1D75CC" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="56" pageNumber="397" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFADFFB138106212FF1D75CC" italics="true" pageId="56" pageNumber="397">Rhododendron</emphasis>
</taxonomicName>
and
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<emphasis id="B90BEA99FFADFFB13D5262F4FE9C75CC" box="[261,338,645,668]" italics="true" pageId="56" pageNumber="397">Prunus</emphasis>
species
</taxonomicName>
at an elevation of
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(
<figureCitation id="13442A0EFFADFFB13E9362F5FCEE75CC" box="[708,800,644,668]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="56" pageNumber="397">Fig. 42b</figureCitation>
).
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of
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<emphasis id="B90BEA99FFADFFB13FBC62F5FF5175EC" italics="true" pageId="56" pageNumber="397">Trechus shibalicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFADFFB13CF162D4FF3775EC" box="[166,249,677,700]" pageId="56" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
also was found at the same site.
</paragraph>
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<smallCapsWord id="8D26A057FFADFFB13DA862B8FD80758C" baselines="727" box="[511,590,713,732]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="56" pageNumber="397">WITHIN</smallCapsWord>
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</emphasis>
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<emphasis id="B90BEA99FFADFFB13EDA62B4FCDC758C" bold="true" box="[653,786,709,733]" pageId="56" pageNumber="397">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FFADFFB13F4A62B4FF22744E" county="Fugong County" location="This" pageId="56" pageNumber="397" specimenCount="1" stateProvince="Shan">
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<collectingRegion id="49BBF869FFADFFB13F4A62B4FC99758C" box="[797,855,709,733]" country="Myanmar" name="Shan" pageId="56" pageNumber="397">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFADFFB13FD362B4FC2A758D" box="[900,996,709,733]" captionStart="FIGURE" captionStartId="55.[171,183,1583,1602]" captionTargetBox="[162,1176,216,1570]" captionTargetId="figure-14@55.[120,1176,216,1570]" captionTargetPageId="55" captionText="FIGURE 21. Trechus pseudoqiqiensis sp. nov.; a. Dorsal habitus (CASENT1014204). b. Map of locality record (red circle) for T. pseudoqiqiensis in the Gaoligong Shan region. Scale lines a = 0.5mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155324" httpUri="https://zenodo.org/record/13155324/files/figure.png" pageId="56" pageNumber="397">Fig. 21b</figureCitation>
.
<location id="8EA06050FFADFFB13FA562B4FBEF758D" LSID="urn:lsid:plazi:treatment:03D6879DFFA3FFB13C2C612AFEE174F1:8EA06050FFADFFB13FA562B4FBEF758D" box="[1010,1057,709,733]" county="Fugong County" name="This" pageId="56" pageNumber="397" stateProvince="Shan">This</location>
species is known from a single locality in
<collectingCounty id="62A14E07FFADFFB13D876297FDB675AE" box="[464,632,742,766]" pageId="56" pageNumber="397">Fugong County</collectingCounty>
, on the eastern slope of the northcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFADFFB13CA06377FF08746F" location="Core Area" pageId="56" pageNumber="397" specimenCount="1" stateProvince="Shan">
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(see
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material above for exact collection data). This locality is in
<location id="8EA06050FFADFFB138356376FF61746F" LSID="urn:lsid:plazi:treatment:03D6879DFFA3FFB13C2C612AFEE174F1:8EA06050FFADFFB138356376FF61746F" name="Core Area" pageId="56" pageNumber="397" stateProvince="Shan">Core Area</location>
3
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFADFFB13CFC6339FEE174F1" blockId="56.[123,1174,219,929]" pageId="56" pageNumber="397">
<emphasis id="B90BEA99FFADFFB13CFC6339FD99740E" bold="true" box="[171,599,840,864]" pageId="56" pageNumber="397">
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<smallCapsWord id="8D26A057FFADFFB13D94633AFD99740E" baselines="857" box="[451,599,843,862]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="56" pageNumber="397">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFADFFB13E296339FEE474F1" country="China" location="Gaoligong Shan" pageId="56" pageNumber="397" specimenCount="1" stateProvince="Yunnan">
This species currently is known only from the
<typeStatus id="54C48829FFADFFB138306339FB5B7430" box="[1127,1173,840,864]" pageId="56" pageNumber="397">type</typeStatus>
locality on the eastern slope of the northcentral part of the
<location id="8EA06050FFADFFB13F5D6319FC7274D0" LSID="urn:lsid:plazi:treatment:03D6879DFFA3FFB13C2C612AFEE174F1:8EA06050FFADFFB13F5D6319FC7274D0" box="[778,956,872,896]" country="China" name="Gaoligong Shan" pageId="56" pageNumber="397" stateProvince="Yunnan">Gaoligong Shan</location>
, in western
<collectingRegion id="49BBF869FFADFFB138136318FF1374F1" country="China" name="Yunnan" pageId="56" pageNumber="397">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFADFFB13CB063F8FEE474F1" box="[231,298,905,929]" name="China" pageId="56" pageNumber="397">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,354 @@
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<mods:title id="000C92D4841AA46489FF71762EEF4F2C">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:affiliation id="C0358AF06BE665EDA08AF08F75FEBC4C">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="314A58D2E7899AA83AAA14E18E0294CC">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<taxonomicName id="4C7F4D08FFA5FFB93C2C62AAFD0175A3" ID-CoL="8M8YZ" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,719,731,756]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="389" phylum="Arthropoda" rank="species" species="mingguangensis" status="sp. nov." subGenus="Trechus">
<emphasis id="B90BEA99FFA5FFB93C2C62AAFF1A75A3" bold="true" box="[123,212,731,755]" italics="true" pageId="48" pageNumber="389">Trechus</emphasis>
(
<emphasis id="B90BEA99FFA5FFB93CB362AAFEF375A3" bold="true" box="[228,317,731,755]" italics="true" pageId="48" pageNumber="389">Trechus</emphasis>
)
<emphasis id="B90BEA99FFA5FFB93D1962AAFDCD75A3" bold="true" box="[334,515,731,755]" italics="true" pageId="48" pageNumber="389">mingguangensis</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFA5FFB93E8C62ADFCFC75A4" box="[731,818,732,756]" pageId="48" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA5FFB93C2C628DFE9C7441" box="[123,338,763,786]" pageId="48" pageNumber="389" type="description">
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(
<figureCitation id="13442A0EFFA5FFB93CD5628DFF1D7441" box="[130,211,764,786]" captionStart="FIGURE" captionStartId="49.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1571]" captionTargetId="figure-14@49.[120,1176,216,1571]" captionTargetPageId="49" captionText="FIGURE 19. Trechus mingguangensis sp. nov.; a. Dorsal habitus (CASENT1038692). b. Median lobe of aedeagus of male (CASENT1038692), left lateral aspect. c. Map of locality records (red circles) for T. mingguangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155320" httpUri="https://zenodo.org/record/13155320/files/figure.png" pageId="48" pageNumber="389">Figs. 19</figureCitation>
,
<figureCitation id="13442A0EFFA5FFB93C8A628DFECD7442" box="[221,259,764,786]" captionStart="FIGURE" captionStartId="103.[171,183,1626,1645]" captionTargetBox="[130,1167,217,1606]" captionTargetId="figure-14@103.[130,1167,217,1606]" captionTargetPageId="103" captionText="FIGURE 41. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Dulongjiang Township, Dulong Jiang at Elideng village, elevation 1640 m; habitat in which specimens of Trechus indicus Putzeys were collected. b. Tengchong County, Mingguang Township, slope SW of Eighth Boundary Post Pass, elevation 2887 m; habitat in which specimens of Trechus mingguangensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155368" httpUri="https://zenodo.org/record/13155368/files/figure.png" pageId="48" pageNumber="389">41b</figureCitation>
,
<figureCitation id="13442A0EFFA5FFB93D59628AFE857441" box="[270,331,763,785]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="48" pageNumber="389">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA5FFB93CFC6354FED1738D" pageId="48" pageNumber="389" type="materials_examined">
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</emphasis>
.—
<materialsCitation id="3B173CD6FFA5FFB93DD46355FE6274EC" collectingDate="2006-05-23" collectorName="R. L. Brett &amp; X. P. Wang &amp; D. Z. Dong" country="China" county="Tengchong County" elevation="2287" latitude="25.80984" location="Eighth Boundary Post Pass" longLatPrecision="1" longitude="98.62084" municipality="Mingguang Township" pageId="48" pageNumber="389" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFA5FFB93DD46355FE27746C" box="[387,489,804,828]" pageId="48" pageNumber="389" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1038692”/ “
<collectingCountry id="F368761BFFA5FFB93FAA6354FB98746D" box="[1021,1110,805,829]" name="China" pageId="48" pageNumber="389">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FFA5FFB93CEF6335FE5E740D" box="[184,400,836,861]" pageId="48" pageNumber="389">Tengchong County</collectingCounty>
,
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,
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/
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,
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,
<date id="FFC1104BFFA5FFB93D336314FE3B742C" box="[356,501,868,893]" pageId="48" pageNumber="389" value="2006-05-23">
<collectingDate id="EF85E9A3FFA5FFB93D336314FE3B742C" box="[356,501,868,893]" pageId="48" pageNumber="389" value="2006-05-23">23 May 2006</collectingDate>
</date>
,”/ “Stop # DHK-2006-037A, D.H.
<taxonomicName id="4C7F4D08FFA5FFB93F3C6315FC29742C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[875,999,868,892]" pageId="48" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFA5FFB93FA66314FB91742C" box="[1009,1119,869,892]" pageId="48" pageNumber="389">R.L. Brett</collectorName>
,
<collectorName id="268A535DFFA5FFB9383E6314FF7774CD" pageId="48" pageNumber="389">X.P. Wang</collectorName>
&amp;
<collectorName id="268A535DFFA5FFB93C8963F4FE8474CD" box="[222,330,901,925]" pageId="48" pageNumber="389">D.Z.Dong</collectorName>
collectors”/ “
<typeStatus id="54C48829FFA5FFB93D8863F4FDA074CD" box="[479,622,901,925]" pageId="48" pageNumber="389" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA5FFB93E2063F5FBFB74CC" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[631,1077,900,925]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="389" phylum="Arthropoda" rank="species" species="mingguangensis" status="sp. nov.">
<emphasis id="B90BEA99FFA5FFB93E2063F5FC4E74CC" box="[631,896,900,924]" italics="true" pageId="48" pageNumber="389">Trechus mingguangensis</emphasis>
Deuve &amp; Liang
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,
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designated 2016” [red label]
</materialsCitation>
.
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<typeStatus id="54C48829FFA5FFB93DE163D4FDD074ED" box="[438,542,933,957]" pageId="48" pageNumber="389" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
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) labeled: “CASENT 1038840”/ “
<collectingCountry id="F368761BFFA5FFB93CD363B4FF13748D" box="[132,221,965,989]" name="China" pageId="48" pageNumber="389">CHINA</collectingCountry>
,
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,
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,
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, small stream on SW-facing slope below 7.9 airkm
<locationDeviation id="9A722FA9FFA5FFB93D676394FE3F74AD" box="[304,497,997,1021]" location="Zizhi village" pageId="48" pageNumber="389">N of Zizhi village</locationDeviation>
,
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,”/ “
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,
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</quantity>
,
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<collectingDate id="EF85E9A3FFA5FFB93FBA6394FBB174AC" box="[1005,1151,996,1021]" pageId="48" pageNumber="389" value="2006-05-23">23 May 2006</collectingDate>
</date>
,”/ “Stop # DHK-2006-038A, D.H.
<taxonomicName id="4C7F4D08FFA5FFB93D836474FD83734D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[468,589,1029,1053]" pageId="48" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFA5FFB93E396474FD2E734D" box="[622,736,1029,1053]" pageId="48" pageNumber="389">D.Z. Dong</collectorName>
collectors”/ “
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<emphasis id="B90BEA99FFA5FFB93FA96474FF3D736C" italics="true" pageId="48" pageNumber="389">Trechus mingguangensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFA5FFB93DF86454FDCD736C" box="[431,515,1061,1084]" pageId="48" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA5FFB93CFC6434FD4B732C" blockId="48.[123,1174,804,1725]" pageId="48" pageNumber="389">
<materialsCitation id="3B173CD6FFA5FFB93CFC6434FD4F732C" country="China" county="Tengchong County" elevation="2287" latitude="25.80314" location="Eighth Boundary Post Pass" longLatPrecision="1" longitude="98.62117" municipality="Mingguang Township" pageId="48" pageNumber="389" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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.—
<collectingCountry id="F368761BFFA5FFB93D296434FE0E730D" box="[382,448,1093,1117]" name="China" pageId="48" pageNumber="389">China</collectingCountry>
,
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,
<collectingCounty id="62A14E07FFA5FFB93E7E6434FD38730D" box="[553,758,1093,1117]" pageId="48" pageNumber="389">Tengchong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA5FFB93F576434FC23730D" box="[768,1005,1093,1117]" pageId="48" pageNumber="389">Mingguang Township</collectingMunicipality>
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,
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,
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.
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<paragraph id="8BC0368BFFA5FFB93CFC64F4FED1738D" blockId="48.[123,1174,804,1725]" pageId="48" pageNumber="389">
<emphasis id="B90BEA99FFA5FFB93CFC64F4FE3873CB" bold="true" box="[171,502,1157,1181]" pageId="48" pageNumber="389">
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.— The species epithet,
<taxonomicName id="4C7F4D08FFA5FFB93F5364F7FC7F73CD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[772,945,1158,1181]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="389" phylum="Arthropoda" rank="species" species="mingguangensis">
<emphasis id="B90BEA99FFA5FFB93F5364F7FC7F73CD" box="[772,945,1158,1181]" italics="true" pageId="48" pageNumber="389">mingguangensis</emphasis>
</taxonomicName>
, is derived from the name of the township in which the
<typeStatus id="54C48829FFA5FFB93DB864D4FDD373ED" box="[495,541,1189,1213]" pageId="48" pageNumber="389">type</typeStatus>
locality is located, Mingguang, and the Latin suffix,
<emphasis id="B90BEA99FFA5FFB9380764D7FB4173ED" box="[1104,1167,1190,1213]" italics="true" pageId="48" pageNumber="389">-ensis</emphasis>
, denoting place.
</paragraph>
</subSubSection>
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<paragraph id="8BC0368BFFA5FFB93CFC6494FCC7714D" blockId="48.[123,1174,804,1725]" pageId="48" pageNumber="389">
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<emphasis id="B90BEA99FFA5FFB93CFC6494FEEE73AB" bold="true" box="[171,288,1253,1277]" pageId="48" pageNumber="389">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFFA5FFB93E106494FD6D73AD" box="[583,675,1253,1277]" captionStart="FIGURE" captionStartId="49.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1571]" captionTargetId="figure-14@49.[120,1176,216,1571]" captionTargetPageId="49" captionText="FIGURE 19. Trechus mingguangensis sp. nov.; a. Dorsal habitus (CASENT1038692). b. Median lobe of aedeagus of male (CASENT1038692), left lateral aspect. c. Map of locality records (red circles) for T. mingguangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155320" httpUri="https://zenodo.org/record/13155320/files/figure.png" pageId="48" pageNumber="389">Fig. 19a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL =
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), apterous; body color reddish-brown; tempora glabrous, short and convex; pronotum transverse (ratio PW/PL = 1.40), basal angles acute and sharp; elytra very slightly elongate, striae finely punctate, striae 1 to 4 deeply impressed, stria 5 more faintly impressed, striae 6 to 8 effaced or nearly so, recurrent stria terminated anteriorly at a slight convexity in line with the presumed location of stria 7 apically; two normal discal setae present, inserted next to stria 3, preapical seta present, inserted in a discal position next to stria 3 at apical one-fourth of elytra; median lobe of male aedeagus with a large sagittal aileron, apical lamella long and rectilinear, nearly straight, apex thin, endophallus with copulatory piece poorly defined (
<figureCitation id="13442A0EFFA5FFB93EC86674FD32714D" box="[671,764,1541,1565]" captionStart="FIGURE" captionStartId="49.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1571]" captionTargetId="figure-14@49.[120,1176,216,1571]" captionTargetPageId="49" captionText="FIGURE 19. Trechus mingguangensis sp. nov.; a. Dorsal habitus (CASENT1038692). b. Median lobe of aedeagus of male (CASENT1038692), left lateral aspect. c. Map of locality records (red circles) for T. mingguangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155320" httpUri="https://zenodo.org/record/13155320/files/figure.png" pageId="48" pageNumber="389">Fig. 19b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA5FFBB3CFC6654FBBC712D" lastPageId="50" lastPageNumber="391" pageId="48" pageNumber="389" type="description">
<paragraph id="8BC0368BFFA5FFB93CFC6654FE59712D" blockId="48.[123,1174,804,1725]" pageId="48" pageNumber="389">
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<emphasis id="B90BEA99FFA5FFB93CFC6654FEF4716C" bold="true" box="[171,314,1573,1597]" pageId="48" pageNumber="389">DESCRIPTION</emphasis>
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.— Size large, BL =
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. Body color dark reddish-brown, shiny, appendages paler, yellowish- or reddish-tan, palpi pale yellow. Body surface smooth, only head capsule faintly alutaceous.
</paragraph>
<paragraph id="8BC0368BFFA5FFBB3CFC66F4FC127684" blockId="48.[123,1174,804,1725]" lastBlockId="50.[123,1174,219,1661]" lastPageId="50" lastPageNumber="391" pageId="48" pageNumber="389">
Head. Moderate in size; eyes slightly reduced, moderately convex, but their diameter less than twice as long as tempora, the latter short, moderately convex and glabrous. Frons moderately flattened; frontal furrows deep, curved, prolonged and not attenuated posterior to the eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave.
<collectingRegion id="49BBF869FFA7FFBB3EF4616DFD337663" box="[675,765,284,307]" country="Myanmar" name="Chin" pageId="50" pageNumber="391">Mentum</collectingRegion>
and submentum not fused but suture between them only faintly impressed.
<collectingRegion id="49BBF869FFA7FFBB3E59614DFDA67603" box="[526,616,316,339]" country="Myanmar" name="Chin" pageId="50" pageNumber="391">Mentum</collectingRegion>
with medial tooth broad and apically truncate, about one-half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae short, not quite extended to the basal one-fourth of elytra, male with about 2.5 to 3 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter.
</paragraph>
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19.
<taxonomicName id="4C7F4D08FFA4FFB83D476642FE2A7116" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[272,484,1587,1606]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="390" phylum="Arthropoda" rank="species" species="mingguangensis" status="sp. nov.">
<emphasis id="B90BEA99FFA4FFB83D476642FE2A7116" box="[272,484,1587,1606]" italics="true" pageId="49" pageNumber="390">Trechus mingguangensis</emphasis>
</taxonomicName>
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; a. Dorsal habitus (CASENT1038692). b. Median lobe of aedeagus of male (CASENT1038692), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFA4FFB83F35663CFBCC710F" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[866,1026,1613,1631]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="390" phylum="Arthropoda" rank="species" species="mingguangensis">
<emphasis id="B90BEA99FFA4FFB83F35663CFBCC710F" box="[866,1026,1613,1631]" italics="true" pageId="49" pageNumber="390">T. mingguangensis</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFA7FFBB3CFC61ACFC8175A6" blockId="50.[123,1174,219,1661]" pageId="50" pageNumber="391">Pronotum. Moderately transverse (ratio PW/PL = 1.40), narrowed posteriorly, greatest width anterior to middle; lateral margins widely rounded, straightened only just anterior to basal angles, the latter small, but projected, acute and sharp. Disc convex, smooth and glabrous, median longitudinal impression sharply impressed, not quite extended to anterior margin; basal foveae distinct, subcircular; median basal area with several faint, longitudinal rugulae, delimited laterally by oblique furrows. Basal margin slightly bisinuate, slightly reflexed and oblique laterally, slightly projected medially. Lateral borders of pronotum moderately slender, regular, slightly reflexed, lateral grooves deeply impressed. Single midlateral setae on each side inserted slightly anterior to middle; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFA7FFBB3CFC628EFCAD7348" blockId="50.[123,1174,219,1661]" pageId="50" pageNumber="391">
Elytra. Ovoid, only very slightly elongate, more narrowed basally than apically, humeri distinct but rounded. Disc convex and smooth. Striae very finely punctate, striae 1 to 4 deeply impressed, stria 5 more faintly impressed, striae 6 to 8 effaced or nearly so. Parascutellar striole present. Recurrent stria terminated anteriorly by a slight convexity at the presumed location of stria 7 apically. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a discal position on interval 3, next to stria
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forward position at apical one-fourth of elytra. Umbilicate setal series with setae of humeral group equidistance from each other and setae of median group inserted slightly posterior to middle.
</paragraph>
<paragraph id="8BC0368BFFA7FFBB3CFC6450FE507309" blockId="50.[123,1174,219,1661]" pageId="50" pageNumber="391">Legs. Relatively short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFA7FFBB3CFC6410FB9E73C9" blockId="50.[123,1174,219,1661]" pageId="50" pageNumber="391">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with four pairs.</paragraph>
<paragraph id="8BC0368BFFA7FFBB3CFC64D0FF7E724A" blockId="50.[123,1174,219,1661]" pageId="50" pageNumber="391">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFA7FFBB3DBA64D0FD8673E9" box="[493,584,1185,1209]" captionStart="FIGURE" captionStartId="49.[171,183,1587,1606]" captionTargetBox="[120,1176,216,1571]" captionTargetId="figure-14@49.[120,1176,216,1571]" captionTargetPageId="49" captionText="FIGURE 19. Trechus mingguangensis sp. nov.; a. Dorsal habitus (CASENT1038692). b. Median lobe of aedeagus of male (CASENT1038692), left lateral aspect. c. Map of locality records (red circles) for T. mingguangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155320" httpUri="https://zenodo.org/record/13155320/files/figure.png" pageId="50" pageNumber="391">Fig. 19b</figureCitation>
) only slightly enlarged basally but with a large sagittal aileron present; shaft markedly narrowed at middle and broader subapically; apical lamella long, rectilinear, extremely thin in lateral view; endophallus voluminous but without well-defined sclerites.
</paragraph>
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</emphasis>
.—
<specimenCount id="9D79FD02FFA7FFBB3D876552FDA5726B" box="[464,619,1315,1339]" count="1" pageId="50" pageNumber="391" type="generic">One specimen</specimenCount>
of this species was found under stones and debris along a roadcut (
<figureCitation id="13442A0EFFA7FFBB3D796532FE47720B" box="[302,393,1347,1371]" captionStart="FIGURE" captionStartId="103.[171,183,1626,1645]" captionTargetBox="[130,1167,217,1606]" captionTargetId="figure-14@103.[130,1167,217,1606]" captionTargetPageId="103" captionText="FIGURE 41. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Dulongjiang Township, Dulong Jiang at Elideng village, elevation 1640 m; habitat in which specimens of Trechus indicus Putzeys were collected. b. Tengchong County, Mingguang Township, slope SW of Eighth Boundary Post Pass, elevation 2887 m; habitat in which specimens of Trechus mingguangensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155368" httpUri="https://zenodo.org/record/13155368/files/figure.png" pageId="50" pageNumber="391">Fig. 41b</figureCitation>
) through slightly disturbed primary forest of
<taxonomicName id="4C7F4D08FFA7FFBB3F3A6535FC62720B" box="[877,940,1348,1371]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="50" pageNumber="391" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA7FFBB3F3A6535FC62720B" box="[877,940,1348,1371]" italics="true" pageId="50" pageNumber="391">Tsuga</emphasis>
</taxonomicName>
, with small, scattered small bamboo thickets; and the other specimen was found under stones along a small stream crossing the road in the same general area. No other trechines were collected in either of these sites, which range in elevation from
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.
</paragraph>
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</emphasis>
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<emphasis id="B90BEA99FFA7FFBB3EDD65B5FCC1728A" bold="true" box="[650,783,1476,1500]" pageId="50" pageNumber="391">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FFA7FFBB3F4E65B5FDCF714C" county="Mingguang Township in Tengchong County" location="We" pageId="50" pageNumber="391" specimenCount="2" stateProvince="Shan">
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<collectingRegion id="49BBF869FFA7FFBB3F4E65B5FC9A728A" box="[793,852,1476,1500]" country="Myanmar" name="Shan" pageId="50" pageNumber="391">SHAN</collectingRegion>
</emphasis>
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.—
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.
<location id="8EA06050FFA7FFBB3FBC65B5FBC0728B" LSID="urn:lsid:plazi:treatment:03D6879DFFA5FFBB3C2C62AAFBBC712D:8EA06050FFA7FFBB3FBC65B5FBC0728B" box="[1003,1038,1476,1499]" county="Mingguang Township in Tengchong County" name="We" pageId="50" pageNumber="391" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFA7FFBB3C876595FE9972AC" box="[208,343,1508,1532]" count="2" pageId="50" pageNumber="391" type="generic">2 specimens</specimenCount>
, both from
<collectingCounty id="62A14E07FFA7FFBB3D826595FC7F72AC" box="[469,945,1508,1532]" pageId="50" pageNumber="391">Mingguang Township in Tengchong County</collectingCounty>
on the western slope of the southern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFA7FFBB3E516675FDC9716C" location="Core Area" pageId="50" pageNumber="391" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFA7FFBB3E516675FDF5714C" box="[518,571,1540,1564]" country="Myanmar" name="Shan" pageId="50" pageNumber="391">Shan</collectingRegion>
(see
<typeStatus id="54C48829FFA7FFBB3E266674FD68714D" box="[625,678,1541,1565]" pageId="50" pageNumber="391">Type</typeStatus>
material above for exact collection data). Both of these localities are in
<location id="8EA06050FFA7FFBB3DD56654FE3E716D" LSID="urn:lsid:plazi:treatment:03D6879DFFA5FFBB3C2C62AAFBBC712D:8EA06050FFA7FFBB3DD56654FE3E716D" box="[386,496,1573,1597]" name="Core Area" pageId="50" pageNumber="391" stateProvince="Shan">Core Area</location>
6
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA7FFBB3CFC6635FBBC712D" blockId="50.[123,1174,219,1661]" pageId="50" pageNumber="391">
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</emphasis>
.— This species currently is known only from the western slope of the southern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFA7FFBB3F326614FBEE712D" box="[869,1056,1637,1661]" country="China" name="Yunnan" pageId="50" pageNumber="391">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFA7FFBB387C6614FBA0712D" box="[1067,1134,1637,1661]" name="China" pageId="50" pageNumber="391">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="9B80E237A6837E7E07386072B1B0FBA1" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFA6FFBF3C2C60AAFE497663" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechus (Trechus) qiqiensis Deuve &amp; Kavanaugh &amp; Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="395" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="392" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="170748B133141F86C7EE8D7DD74EA8B1">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:role id="AD7585A94C2F26CFE93D608E46FEF3D5">
<mods:roleTerm id="E54FF644E1F47076436A2A604ECD5CA7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3AC7DFC4F5EE708CED1D910DC6548BE8">Deuve, Thierry</mods:namePart>
<mods:affiliation id="D992CF592FC7336306A1D33300BDA022">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
<mods:name id="76132B78E17CFB72FB3DA37D38F2E1C1" type="personal">
<mods:role id="03A098162EBFABE6F931F905606E0C0F">
<mods:roleTerm id="CC7A2DFB070162D1DF1F81F264CB813D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EDF6C93FB3F0DC2AF89AD525AA1F1C64">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="3BC64E7A589D6423A7C48EE776621578">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="555E893B5FA0B488262B3C07905314AE" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="48C931B680DB1620DB6855C662BA23D1" type="personal">
<mods:role id="7693BAD8B9CA8451641A98878AEBA567">
<mods:roleTerm id="8A7BC5204D765E754841AE25D7E2DACD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="452275F6A37567AA8C237C5D656CCB95">Liang, Hongbin</mods:namePart>
<mods:affiliation id="82E57E5EA78F7304BB3566B6AD2891C5">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:typeOfResource id="64964F311DFA1F73C8A97AC91D5D27CF">text</mods:typeOfResource>
<mods:relatedItem id="BECAD4523A3E73BC25A8CCEA03ECD668" type="host">
<mods:titleInfo id="FCE21377A92E0051B95B427261D44982">
<mods:title id="7514368728906A04FB3CA64B8A1E2518">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="C311DC98B967E5BD4C5E2B74451EAB9A">
<mods:date id="1F0A081C73E13AC0D0A0708A8F5AA916">2016</mods:date>
<mods:detail id="CF3DA3C40671563174FCAEADF1915F78" type="pubDate">
<mods:number id="C244631DF39C57DC8460FD39DDAAA180">2016-10-14</mods:number>
</mods:detail>
<mods:detail id="18C8B704B45F456728D27B307282A33C" type="volume">
<mods:number id="E565A957FF623E7E7AB6BE4786ED482A">63</mods:number>
</mods:detail>
<mods:detail id="B1D3B6A1973F21A64C8A1EFBE258E061" type="issue">
<mods:number id="689D27922C940EB8B565582317E05744">12</mods:number>
</mods:detail>
<mods:extent id="1B07FF8112B9192F791077D38ECECFB6" unit="page">
<mods:start id="BAE95053C03DC9DED08F4E9AE1CD8AD5">341</mods:start>
<mods:end id="4893A77C568DC4AFC255615DE44DBF9F">455</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="25175300B667046B2D9CE1674BDDBB13">journal article</mods:classification>
<mods:identifier id="3175709CF8F7A3D3E88A5568317739B4" type="DOI">10.5281/zenodo.13155283</mods:identifier>
<mods:identifier id="239614827FCB1871D5186D7A5D0C6C7F" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="020C58592FF118E79CECF36C93C784AD" type="Zenodo-Dep">13155283</mods:identifier>
<mods:identifier id="0803CE59D6EE25AED89DE7EAEFC8F2E3" type="ZooBank">4C790FE0-B735-4592-8827-EEF83C663CB1</mods:identifier>
</mods:mods>
<treatment id="03D6879DFFA6FFBF3C2C60AAFE497663" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663" httpUri="http://treatment.plazi.org/id/03D6879DFFA6FFBF3C2C60AAFE497663" lastPageId="54" lastPageNumber="395" pageId="51" pageNumber="392">
<subSubSection id="C3656500FFA6FFBA3C2C60AAFCD077A3" box="[123,798,219,243]" pageId="51" pageNumber="392" type="nomenclature">
<paragraph id="8BC0368BFFA6FFBA3C2C60AAFCD077A3" blockId="51.[123,798,219,273]" box="[123,798,219,243]" pageId="51" pageNumber="392">
<heading id="D08881E7FFA6FFBA3C2C60AAFCD077A3" box="[123,798,219,243]" fontSize="10" level="2" pageId="51" pageNumber="392" reason="4">
<emphasis id="B90BEA99FFA6FFBA3C2C60AAFCD077A3" bold="true" box="[123,798,219,243]" pageId="51" pageNumber="392">
<taxonomicName id="4C7F4D08FFA6FFBA3C2C60AAFE3377A3" ID-CoL="8M8J2" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,509,219,243]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis" subGenus="Trechus">
<emphasis id="B90BEA99FFA6FFBA3C2C60AAFF1A77A3" bold="true" box="[123,212,219,243]" italics="true" pageId="51" pageNumber="392">Trechus</emphasis>
(
<emphasis id="B90BEA99FFA6FFBA3CB360AAFEF377A3" bold="true" box="[228,317,219,243]" italics="true" pageId="51" pageNumber="392">Trechus</emphasis>
)
<emphasis id="B90BEA99FFA6FFBA3D1960AAFE6077A3" bold="true" box="[334,430,219,243]" italics="true" pageId="51" pageNumber="392">qiqiensis</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFA6FFBA3E9F60AAFCD077A3" box="[712,798,219,243]" pageId="51" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBA3C2C608AFE4F7641" box="[123,385,251,273]" pageId="51" pageNumber="392" type="description">
<paragraph id="8BC0368BFFA6FFBA3C2C608AFE4F7641" blockId="51.[123,798,219,273]" box="[123,385,251,273]" pageId="51" pageNumber="392">
(
<figureCitation id="13442A0EFFA6FFBA3CD5608AFF1D7640" box="[130,211,251,273]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="51" pageNumber="392">Figs. 20</figureCitation>
,
<figureCitation id="13442A0EFFA6FFBA3C8A608AFECD7641" box="[221,259,251,273]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="51" pageNumber="392">37b</figureCitation>
,
<figureCitation id="13442A0EFFA6FFBA3D59608AFEFD7640" box="[270,307,251,272]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="51" pageNumber="392">42a</figureCitation>
,
<figureCitation id="13442A0EFFA6FFBA3D6A608AFEB57641" box="[317,379,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="51" pageNumber="392">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBA3CFC6155FF74734D" pageId="51" pageNumber="392" type="materials_examined">
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</emphasis>
.—
<materialsCitation id="3B173CD6FFA6FFBA3DD56155FE5F76EC" collectingDate="2000-07-15" collectingDateMax="2000-07-19" collectingDateMin="2000-07-15" collectorName="C. E. Griswold" country="China" county="Gaoligong Shan" elevation="2775" latitude="27.71503" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" pageId="51" pageNumber="392" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFA6FFBA3DD56155FE26766C" box="[386,488,292,316]" pageId="51" pageNumber="392" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1007384”/ “
<collectingCountry id="F368761BFFA6FFBA3FAB6155FB9B766C" box="[1020,1109,292,316]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FFA6FFBA3D416135FE09760C" box="[278,455,324,348]" pageId="51" pageNumber="392">Gaoligong Shan</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA6FFBA3D856135FD6E760C" box="[466,672,324,348]" pageId="51" pageNumber="392">Nujiang Prefecture</collectingMunicipality>
,
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, No. 12
<location id="8EA06050FFA6FFBA381A6135FF3D762B" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA381A6135FF3D762B" country="China" county="Gaoligong Shan" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" name="Bridge Camp area" pageId="51" pageNumber="392" stateProvince="Yunnan">Bridge Camp area</location>
, 16.3 airkm
<locationDeviation id="9A722FA9FFA6FFBA3DD66115FDFD762C" box="[385,563,356,380]" location="Gongshan" pageId="51" pageNumber="392">W of Gongshan</locationDeviation>
,”/ “
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/
<geoCoordinate id="EE4B504CFFA6FFBA3EBF6115FCA7762C" box="[744,873,356,380]" degrees="98.50244" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.50244">E98.50244°</geoCoordinate>
,
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,
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, Stop#00-23A, D.H.
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,
<collectorName id="268A535DFFA6FFBA3D8E61F5FDBB76CC" box="[473,629,388,412]" pageId="51" pageNumber="392">C.E. Griswold</collectorName>
, Liang H.-B., D. Ubick, &amp; Dong D.-Z. collectors”/ “IMAGE” [green label]/ “
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<typeStatus id="54C48829FFA6FFBA3DC661D5FDEE76EC" box="[401,544,420,444]" pageId="51" pageNumber="392" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA6FFBA3E7F61D5FCE176EB" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[552,815,420,444]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA6FFBA3E7F61D5FD2C76EC" box="[552,738,420,444]" italics="true" pageId="51" pageNumber="392">Trechus qiqiensis</emphasis>
Deuve
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&amp; Kavanaugh,
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designat- ed 2016” [red label].
<materialsCitation id="3B173CD6FFA6FFBA3D3C61B5FF31756C" collectionCode="CAS, IOZ, MNHN" country="China" pageId="51" pageNumber="392" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFA6FFBA3D3C61B5FE1D768C" box="[363,467,452,476]" pageId="51" pageNumber="392" type="paratype">Paratypes</typeStatus>
(at total of 11):
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and
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(in
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,
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,
<collectionCode id="ED6EAE4EFFA6FFBA386C61B5FB40768C" box="[1083,1166,452,476]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="51" pageNumber="392">MNHN</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFA6FFBA3D6C6195FE5576AC" box="[315,411,484,508]" pageId="51" pageNumber="392" type="holotype">holotype</typeStatus>
, except first label “CASENT 1007380” to “CASENT 1007381” and “CASENT 1007382” to “CASENT 1007383” and “CASENT 1007385” to “CASENT 1007386”, respectively
</materialsCitation>
;
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and
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(in
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,
<collectionCode id="ED6EAE4EFFA6FFBA3E3D6255FD56756C" box="[618,664,548,572]" pageId="51" pageNumber="392">IOZ</collectionCode>
) labeled: “CASENT 1010345” and “CASENT 1010346” to “CASENT 1010347”/ “
<collectingCountry id="F368761BFFA6FFBA3DA96235FD99750C" box="[510,599,580,604]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3E376234FD7A750C" box="[608,692,581,604]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3EE96235FC4E750C" box="[702,896,580,604]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<location id="8EA06050FFA6FFBA3FDD6235FBFB750C" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3FDD6235FBFB750C" box="[906,1077,580,604]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44661" name="Cikai Township" pageId="51" pageNumber="392" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFA6FFBA38686235FDFF752C" location="Gongshan on Dulong Valley Road" pageId="51" pageNumber="392">
<quantity id="4C879B6EFFA6FFBA38686235FB5B750C" box="[1087,1173,580,604]" metricMagnitude="4" metricUnit="m" metricValue="5.26" pageId="51" pageNumber="392" unit="km" value="52.6">52.6 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFFA6FFBA3E6A6215FD1E752C" box="[573,720,612,636]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="51" pageNumber="392" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FFA6FFBA3E6A6215FD1E752C" box="[573,720,612,636]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="51" pageNumber="392" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFA6FFBA3EAD6215FCB1752C" box="[762,895,612,636]" degrees="27.77032" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3FD06215FBC9752C" box="[903,1031,612,636]" degrees="98.44661" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.44661">E98.44661°</geoCoordinate>
,
<date id="FFC1104BFFA6FFBA38446215FF7D75CC" pageId="51" pageNumber="392" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">
<collectingDate id="EF85E9A3FFA6FFBA38446215FF7D75CC" pageId="51" pageNumber="392" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">1-2 October 2002</collectingDate>
</date>
, Stop #DHK-2002-034, D.H.
<taxonomicName id="4C7F4D08FFA6FFBA3DA362F5FDA375CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[500,621,644,668]" pageId="51" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFA6FFBA3ED862F5FCC775CC" box="[655,777,644,668]" pageId="51" pageNumber="392">P.A. Marek</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA6FFBA3FD862F5FDA575AC" collectingDate="2004-11-13" collectionCode="CAS" country="China" county="Gongshan County" elevation="3330" latitude="27.78075" location="Cikai Township" longLatPrecision="1" longitude="98.47" pageId="51" pageNumber="392" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA6FFBA3FD862F5FC1C75CC" box="[911,978,644,668]" count="1" pageId="51" pageNumber="392" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA6FFBA3FAA62F5FBFC75CC" box="[1021,1074,644,668]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="51" pageNumber="392" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1015626”/ “
<collectingCountry id="F368761BFFA6FFBA3D2F62D5FE1F75EC" box="[376,465,676,700]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3D8A62D4FDFF75EC" box="[477,561,677,700]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3E6A62D5FCCF75EC" box="[573,769,676,700]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<location id="8EA06050FFA6FFBA3F5A62D5FC7475EC" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3F5A62D5FC7475EC" box="[781,954,676,700]" country="China" county="Gongshan County" latitude="27.78075" longLatPrecision="1" longitude="98.47" name="Cikai Township" pageId="51" pageNumber="392" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFA6FFBA3F9162D5FE73758C" location="Gongshan on Dulong Valley Road" pageId="51" pageNumber="392">
<quantity id="4C879B6EFFA6FFBA3F9162D5FBC475EC" box="[966,1034,676,700]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="51" pageNumber="392" unit="km" value="48.0">48 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFFA6FFBA3D9C62B5FDEF758C" box="[459,545,708,732]" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" pageId="51" pageNumber="392" unit="m" value="3330.0">
<elevation id="0052D1B8FFA6FFBA3D9C62B5FDEF758C" box="[459,545,708,732]" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" pageId="51" pageNumber="392" unit="m" value="3330.0">3330 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFA6FFBA3E1B62B5FD1E758C" box="[588,720,708,732]" degrees="27.78075" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.78075">N27.78075°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3E8F62B5FC94758C" box="[728,858,708,732]" degrees="98.47000" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.47">E98.47000°</geoCoordinate>
,
<date id="FFC1104BFFA6FFBA3F3F62B5FBF1758C" box="[872,1087,708,732]" pageId="51" pageNumber="392" value="2004-11-13">
<collectingDate id="EF85E9A3FFA6FFBA3F3F62B5FBF1758C" box="[872,1087,708,732]" pageId="51" pageNumber="392" value="2004-11-13">13 November 2004</collectingDate>
</date>
, Stop # DHK-2004-086, D.H.
<taxonomicName id="4C7F4D08FFA6FFBA3D2F6295FE3F75AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[376,497,740,764]" pageId="51" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA6FFBA3E2B6295FC2E742C" collectingDate="2006-08-19" collectionCode="IOZ" collectorName="Y. Liu &amp; All" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="51" pageNumber="392" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA6FFBA3E2B6295FD0475AC" box="[636,714,740,764]" count="1" pageId="51" pageNumber="392" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA6FFBA3F566295FCE175AC" box="[769,815,740,764]" pageId="51" pageNumber="392">IOZ</collectionCode>
) labeled: “CASENT 1025832”/ “
<collectingCountry id="F368761BFFA6FFBA3CD36375FF13744C" box="[132,221,772,796]" name="China" pageId="51" pageNumber="392">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3CB06374FEF5744C" box="[231,315,773,796]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3D116375FDC7744C" box="[326,521,772,796]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA6FFBA3E446375FCD2744C" box="[531,796,772,796]" pageId="51" pageNumber="392">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFA6FFBA3FE76375FBA7744C" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA3FE76375FBA7744C" box="[944,1129,772,796]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="51" pageNumber="392" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFA6FFBA38236375FE46746C" location="Chukuai Lake" pageId="51" pageNumber="392">
<quantity id="4C879B6EFFA6FFBA38236375FF53746C" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="51" pageNumber="392" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/ “
<geoCoordinate id="EE4B504CFFA6FFBA3E636355FD76746C" box="[564,696,804,828]" degrees="27.97686" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3E976355FC8F746C" box="[704,833,804,828]" degrees="98.47799" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.47799">E98.47799°</geoCoordinate>
,
<quantity id="4C879B6EFFA6FFBA3F1B6355FC6E746C" box="[844,928,804,828]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="51" pageNumber="392" unit="m" value="3750.0">
<elevation id="0052D1B8FFA6FFBA3F1B6355FC6E746C" box="[844,928,804,828]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="51" pageNumber="392" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFFA6FFBA3FFB6355FB97746C" box="[940,1113,804,828]" pageId="51" pageNumber="392" value="2006-08-19">
<collectingDate id="EF85E9A3FFA6FFBA3FFB6355FB97746C" box="[940,1113,804,828]" pageId="51" pageNumber="392" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-082,
<collectorName id="268A535DFFA6FFBA3D156334FE47740C" box="[322,393,836,860]" pageId="51" pageNumber="392">Y. Liu</collectorName>
collector”.
<collectorName id="268A535DFFA6FFBA3E586335FDFF740C" box="[527,561,836,860]" pageId="51" pageNumber="392">All</collectorName>
<typeStatus id="54C48829FFA6FFBA3E6A6334FD6D740D" box="[573,675,837,861]" pageId="51" pageNumber="392" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFA6FFBA38596335FB5B740C" box="[1038,1173,836,860]" pageId="51" pageNumber="392" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA6FFBA3C2C6315FE4C742C" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,386,868,892]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA6FFBA3C2C6315FEFA742C" box="[123,308,868,892]" italics="true" pageId="51" pageNumber="392">Trechus qiqiensis</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFA6FFBA3E7B6314FDB1742C" box="[556,639,869,892]" pageId="51" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA6FFBA3CFC63F5FE2474EC" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<materialsCitation id="3B173CD6FFA6FFBA3CFC63F5FE2874EC" country="China" county="Gongshan County" elevation="2775" latitude="27.71503" location="Camp" longLatPrecision="1" longitude="98.50244" municipality="Qiqi Trail" pageId="51" pageNumber="392" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFA6FFBA3CFC63F5FEAD74CB" bold="true" box="[171,355,900,924]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC63F5FF2874CB" baselines="918,918" box="[171,230,900,924]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="51" pageNumber="392">
<typeStatus id="54C48829FFA6FFBA3CFC63F5FF2874CB" box="[171,230,900,924]" pageId="51" pageNumber="392">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3CA163F9FEAD74CB" baselines="918" box="[246,355,904,923]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="51" pageNumber="392">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFA6FFBA3DC163F5FE1474CC" box="[406,474,900,924]" name="China" pageId="51" pageNumber="392">China</collectingCountry>
,
<collectingRegion id="49BBF869FFA6FFBA3DB863F4FD8B74CC" box="[495,581,901,924]" country="China" name="Yunnan" pageId="51" pageNumber="392">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA6FFBA3E0D63F5FCE574CD" box="[602,811,900,925]" pageId="51" pageNumber="392">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA6FFBA3F6863F5FC7A74CC" box="[831,948,900,924]" pageId="51" pageNumber="392">Qiqi Trail</collectingMunicipality>
at No. 12
<location id="8EA06050FFA6FFBA381963F4FB5C74CD" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA6FFBA381963F4FB5C74CD" box="[1102,1170,901,925]" country="China" county="Gongshan County" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Qiqi Trail" name="Camp" pageId="51" pageNumber="392" stateProvince="Yunnan">Camp</location>
,
<geoCoordinate id="EE4B504CFFA6FFBA3C2C63D5FF3374EC" box="[123,253,932,956]" degrees="27.71503" direction="north" orientation="latitude" pageId="51" pageNumber="392" precision="1" value="27.71503">N27.71503°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA6FFBA3D5263D5FE4874EC" box="[261,390,932,956]" degrees="98.50244" direction="east" orientation="longitude" pageId="51" pageNumber="392" precision="1" value="98.50244">E98.50244°</geoCoordinate>
,
<quantity id="4C879B6EFFA6FFBA3DC663D5FE2874EC" box="[401,486,932,956]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="51" pageNumber="392" unit="m" value="2775.0">
<elevation id="0052D1B8FFA6FFBA3DC663D5FE2874EC" box="[401,486,932,956]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="51" pageNumber="392" unit="m" value="2775.0">2775 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA6FFBA3CFC63B4FF74734D" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<emphasis id="B90BEA99FFA6FFBA3CFC63B4FE3F748B" bold="true" box="[171,497,965,989]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC63B4FEE0748B" baselines="983,982" box="[171,302,965,989]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="51" pageNumber="392">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3D6163B9FE9C748B" baselines="982" box="[310,338,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="51" pageNumber="392">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3D0E63B9FE62748B" baselines="982" box="[345,428,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="51" pageNumber="392">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFA6FFBA3DE363B9FE3F748B" baselines="982" box="[436,497,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="51" pageNumber="392">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFA6FFBA3EAE63B4FC96748C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[761,856,965,988]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="392" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA6FFBA3EAE63B4FC96748C" box="[761,856,965,988]" italics="true" pageId="51" pageNumber="392">qiqiensis</emphasis>
</taxonomicName>
, is derived from the name of the ancient trail, Qiqi Trail, passing through the
<typeStatus id="54C48829FFA6FFBA3E2A6394FD6574AD" box="[637,683,997,1021]" pageId="51" pageNumber="392">type</typeStatus>
locality, and the Latin suffix,
<emphasis id="B90BEA99FFA6FFBA3FBB6394FBE474AC" box="[1004,1066,997,1020]" italics="true" pageId="51" pageNumber="392">-ensis</emphasis>
, denoting place.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBA3CFC6454FD1C722D" pageId="51" pageNumber="392" type="diagnosis">
<paragraph id="8BC0368BFFA6FFBA3CFC6454FD1C722D" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC6454FEEE736B" baselines="1079,1078" box="[171,288,1061,1085]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="51" pageNumber="392">
<emphasis id="B90BEA99FFA6FFBA3CFC6454FEEE736B" bold="true" box="[171,288,1061,1085]" pageId="51" pageNumber="392">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFA6FFBA3E106454FD6D736D" box="[583,675,1061,1085]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="51" pageNumber="392">Fig. 20a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size relatively large (BL = 4.0 to
<quantity id="4C879B6EFFA6FFBA3CE96414FEDD732D" box="[190,275,1125,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="51" pageNumber="392" unit="mm" value="4.2">4.2 mm</quantity>
), apterous; body color reddish-brown; eyes slightly convex; tempora glabrous, short and convex; pronotum transverse (ratio PW/PL = 1.44), basal angles rectangular and sharp; elytra with striae 1 to 4 deeply impressed on disc but attenuated apically, remaining striae successively less distinct, the outermost effaced or nearly so; recurrent stria terminated anteriorly by a slight convexity in line with stria 7; two discal setae present, inserted on interval 3 next to stria 3; preapical seta present and inserted in a discal position on interval 3, next to either stria 2 or
<quantity id="4C879B6EFFA6FFBA38466574FBF4724D" box="[1041,1082,1285,1309]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="51" pageNumber="392" unit="in" value="3.0">3 in</quantity>
forward position near apical one-fourth of elytra; median lobe of male aedeagus large and robust, subapically bent ventrally, apical lamella sinuate and reflexed dorsally (
<figureCitation id="13442A0EFFA6FFBA3F186534FC61720D" box="[847,943,1349,1373]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="51" pageNumber="392">Fig. 20b</figureCitation>
), endophallus with a more heavily sclerotized voluminous lobe or scaly fold.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA6FFBF3CFC65F4FE497663" lastPageId="54" lastPageNumber="395" pageId="51" pageNumber="392" type="description">
<paragraph id="8BC0368BFFA6FFBA3CFC65F4FE8E728D" blockId="51.[123,1174,292,1725]" pageId="51" pageNumber="392">
<smallCapsWord id="8D26A057FFA6FFBA3CFC65F4FEF472CC" baselines="1431,1431" box="[171,314,1413,1437]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="51" pageNumber="392">
<emphasis id="B90BEA99FFA6FFBA3CFC65F4FEF472CC" bold="true" box="[171,314,1413,1437]" pageId="51" pageNumber="392">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size relatively large, BL = 4.0 to
<quantity id="4C879B6EFFA6FFBA3E9B65F4FCEC72CD" box="[716,802,1413,1437]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="51" pageNumber="392" unit="mm" value="4.2">4.2 mm</quantity>
. Body color reddish-brown, shiny, appendages concolorous but paler, palpi pale yellow. Body surface smooth, only head capsule faintly alutaceous.
</paragraph>
<paragraph id="8BC0368BFFA6FFBC3CFC6594FDD77624" blockId="51.[123,1174,292,1725]" lastBlockId="53.[123,1174,219,1726]" lastPageId="53" lastPageNumber="394" pageId="51" pageNumber="392">
Head. Moderate in size; eyes only slightly projected, slightly more convex, their diameter less than twice as long as tempora, the latter short, convex and glabrous. Frons slightly flattened; frontal furrows deep, curved, continuous but attenuated posterior to the eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Mandibles short; right mandible distinctly tridentate with middle tooth closer to basal tooth (premolar) than to distal tooth, left mandible with a small slightly bifid process.
<collectingRegion id="49BBF869FFA6FFBA3D4566D7FEA271ED" box="[274,364,1702,1725]" country="Myanmar" name="Chin" pageId="51" pageNumber="392">Mentum</collectingRegion>
and submentum not fused but suture between them only faintly impressed.
<collectingRegion id="49BBF869FFA0FFBC3C2C60AAFF1B77A2" box="[123,213,219,242]" country="Myanmar" name="Chin" pageId="53" pageNumber="394">Mentum</collectingRegion>
with medial tooth broad and apically truncate, about one-half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae short, extended posteriorly to basal one-sixth of elytra, with about 2 or 3 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter.
</paragraph>
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20.
<taxonomicName id="4C7F4D08FFA1FFBD3D46665EFE667111" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[273,424,1583,1602]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="393" phylum="Arthropoda" rank="species" species="qiqiensis" status="sp. nov.">
<emphasis id="B90BEA99FFA1FFBD3D46665EFE667111" box="[273,424,1583,1602]" italics="true" pageId="52" pageNumber="393">Trechus qiqiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA1FFBD3DF8665EFE397112" box="[431,503,1583,1602]" pageId="52" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFA1FFBD3F656638FC5C710B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[818,914,1609,1627]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="393" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA1FFBD3F656638FC5C710B" box="[818,914,1609,1627]" italics="true" pageId="52" pageNumber="393">T. qiqiensis</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFA0FFBC3CFC610CFDAA75C6" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">Pronotum. Transverse (ratio PW/PL = 1.44), narrowed posteriorly, greatest width near anteri- or one-fourth; lateral margins markedly rounded, with a short sinuation just anterior to basal angles, the latter small, rectangular and sharp. Disc convex, smooth and glabrous, median longitudinal impression finely impressed, but continuous between anterior and posterior margins; basal foveae distinct, circular; median basal area faintly rugulose, delimited laterally by short, oblique furrows. Basal margin nearly rectilinear, slightly projected medially. Lateral borders of pronotum moderately slender, regular, narrowly and regularly reflexed, lateral grooves narrow but distinctly impressed. Single midlateral setae on each side inserted slightly anterior to middle; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC62EFFDA674A8" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
Elytra. Ovoid, humeri distinct but rounded. Disc convex and smooth. Striae finely punctate, intervals faintly convex; striae 1 to 4 deeply impressed but attenuated apically, stria 5 more faintly impressed, striae 6 to 8 effaced or nearly so. Parascutellar striole present. Recurrent stria terminated anteriorly by a slight convexity at the presumed location of stria 7 apically. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a discal position on interval 3, next to either stria 2 or
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forward position at apical one-fourth of elytra in most specimens; in a few specimens two preapical setae present, one inserted farther forward next to stria 3, as a third discal seta, the second inserted less far forward and next to stria 2. Umbilicate setal series with setae of humeral group equidistance from each other and setae of medi- an group inserted slightly posterior to middle.
</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6470FD6A7368" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">Legs. Medium proportions, slightly short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6430FB857329" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC64F0FBE6738A" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFA0FFBC3E5364F0FDA873C9" box="[516,614,1153,1177]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="53" pageNumber="394">Fig. 20b</figureCitation>
) large and robust, with a moderate-sized sagittal aileron; shaft subapically bent ventrally; apical lamella sinuate and reflexed dorsally, bluntly point- ed apically; endophallus with a more heavily sclerotized voluminous lobe or scaly fold.
</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6493FD73716C" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
<emphasis id="B90BEA99FFA0FFBC3CFC6493FE6973A9" bold="true" box="[171,423,1250,1274]" pageId="53" pageNumber="394">
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<smallCapsWord id="8D26A057FFA0FFBC3D446497FE6973A9" baselines="1268" box="[275,423,1254,1273]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="53" pageNumber="394">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found at elevations ranging from
<quantity id="4C879B6EFFA0FFBC3CE06573FEAB724A" box="[183,357,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="3.2625" metricValueMax="3.75" metricValueMin="2.775" pageId="53" pageNumber="394" unit="m" value="3262.5" valueMax="3750.0" valueMin="2775.0">2775 to 3750 m</quantity>
in a variety of habitats. At the lowest elevation (
<quantity id="4C879B6EFFA0FFBC3F296573FC1B724B" box="[894,981,1282,1307]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="53" pageNumber="394" unit="m" value="2775.0">2775 m</quantity>
), specimens were collected by sifting meager leaf litter and mosses on the forest floor and on rotting logs (
<figureCitation id="13442A0EFFA0FFBC387A6552FB44726B" box="[1069,1162,1315,1339]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="53" pageNumber="394">Fig. 42a</figureCitation>
). This area had abundant conifers (
<taxonomicName id="4C7F4D08FFA0FFBC3DAF6532FDFD720B" box="[504,563,1347,1371]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="53" pageNumber="394" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA0FFBC3DAF6532FDFD720B" box="[504,563,1347,1371]" italics="true" pageId="53" pageNumber="394">Abies</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFA0FFBC3E116532FD4A720B" authorityName="C.Linnaeus" authorityYear="1753" box="[582,644,1347,1371]" class="Pinopsida" family="Cupressaceae" genus="Thuja" kingdom="Plantae" order="Pinales" pageId="53" pageNumber="394" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA0FFBC3E116532FD4A720B" box="[582,644,1347,1371]" italics="true" pageId="53" pageNumber="394">Thuja</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7F4D08FFA0FFBC3E9E6535FCAF720B" box="[713,865,1347,1371]" pageId="53" pageNumber="394">
<emphasis id="B90BEA99FFA0FFBC3E9E6535FCC8720B" box="[713,774,1348,1371]" italics="true" pageId="53" pageNumber="394">Picea</emphasis>
species
</taxonomicName>
) that formed a partly open canopy. Litter and mosses were moist and substrate underneath was composed of crumbling granitic sand. Specimens of
<taxonomicName id="4C7F4D08FFA0FFBC3DF065F2FD2572CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[423,747,1411,1435]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="monochaeta" status="sp. nov.">
<emphasis id="B90BEA99FFA0FFBC3DF065F2FD2572CB" box="[423,747,1411,1435]" italics="true" pageId="53" pageNumber="394">Trechepaphiopsis monochaeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA0FFBC3EA565F5FC8A72CB" box="[754,836,1412,1435]" pageId="53" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
were also collected in the same litter samples at this site. At higher elevations (above
<quantity id="4C879B6EFFA0FFBC3EE265D5FCC672EC" box="[693,776,1444,1468]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="53" pageNumber="394" unit="m" value="3300.0">3300 m</quantity>
), specimens were found under stones in meadows, talus slopes, in low
<taxonomicName id="4C7F4D08FFA0FFBC3D8B65B5FDB4728C" box="[476,634,1476,1500]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="53" pageNumber="394" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA0FFBC3D8B65B5FDB4728C" box="[476,634,1476,1500]" italics="true" pageId="53" pageNumber="394">Rhododendron</emphasis>
</taxonomicName>
thickets, and on heath-covered tundra slopes (
<figureCitation id="13442A0EFFA0FFBC383D65B5FF6972AC" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="53" pageNumber="394">Fig. 37b</figureCitation>
). In such areas,
<taxonomicName id="4C7F4D08FFA0FFBC3D096594FE1472AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[350,474,1509,1532]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFA0FFBC3D096594FE1472AC" box="[350,474,1509,1532]" italics="true" pageId="53" pageNumber="394">T. qiqiensis</emphasis>
</taxonomicName>
specimens were found syntopic with those of
<taxonomicName id="4C7F4D08FFA0FFBC3FB56595FE4D714C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="balli" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFA0FFBC3FB56595FB5872AC" box="[994,1174,1508,1532]" italics="true" pageId="53" pageNumber="394">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFA0FFBC3CD36675FE8A714C" box="[132,324,1540,1564]" italics="true" pageId="53" pageNumber="394">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FFA0FFBC3D056675FE4D714C" box="[338,387,1540,1564]" italics="true" pageId="53" pageNumber="394">balli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFA0FFBC3DE16675FCE9714C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[438,807,1540,1564]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FFA0FFBC3DE16675FDA4714C" box="[438,618,1540,1564]" italics="true" pageId="53" pageNumber="394">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFA0FFBC3E2E6674FD7F714C" box="[633,689,1541,1564]" italics="true" pageId="53" pageNumber="394">s. str.</emphasis>
)
<emphasis id="B90BEA99FFA0FFBC3E976675FCE9714C" box="[704,807,1540,1564]" italics="true" pageId="53" pageNumber="394">griswoldi</emphasis>
</taxonomicName>
<emphasis id="B90BEA99FFA0FFBC3F706675FCE5714C" box="[807,811,1540,1564]" italics="true" pageId="53" pageNumber="394">,</emphasis>
and at highest elevation (
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) also with members of
<taxonomicName id="4C7F4D08FFA0FFBC3D3D6655FDAC716C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[362,610,1572,1596]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="394" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FFA0FFBC3D3D6655FDAC716C" box="[362,610,1572,1596]" italics="true" pageId="53" pageNumber="394">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA0FFBC3E3D6654FD73716C" box="[618,701,1573,1596]" pageId="53" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFFA0FFBC3CFC6635FBDD71ED" blockId="53.[123,1174,219,1726]" pageId="53" pageNumber="394">
<emphasis id="B90BEA99FFA0FFBC3CFC6635FD4E710B" bold="true" box="[171,640,1604,1628]" pageId="53" pageNumber="394">
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<smallCapsWord id="8D26A057FFA0FFBC3D086639FE3D710B" baselines="1622" box="[351,499,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="53" pageNumber="394">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFA0FFBC3DAA6639FD82710B" baselines="1622" box="[509,588,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="53" pageNumber="394">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFA0FFBC3E016639FD4E710B" baselines="1622" box="[598,640,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="53" pageNumber="394">THE</smallCapsWord>
</emphasis>
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<emphasis id="B90BEA99FFA0FFBC3EDD6635FCC1710B" bold="true" box="[650,783,1604,1628]" pageId="53" pageNumber="394">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FFA0FFBC3F4E6635FBEB71CD" county="Cikai Townships in Gongshan County" location="Bingzhiongluo" municipality="We" pageId="53" pageNumber="394" specimenCount="24" specimenCount-female="7" specimenCount-male="5" stateProvince="Shan">
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<emphasis id="B90BEA99FFA0FFBC3F4E6635FC9A710B" bold="true" box="[793,852,1604,1628]" pageId="53" pageNumber="394">
<collectingRegion id="49BBF869FFA0FFBC3F4E6635FC9A710B" box="[793,852,1604,1628]" country="Myanmar" name="Shan" pageId="53" pageNumber="394">SHAN</collectingRegion>
</emphasis>
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.—
<figureCitation id="13442A0EFFA0FFBC3FD76634FC10710D" box="[896,990,1605,1629]" captionStart="FIGURE" captionStartId="52.[171,183,1583,1602]" captionTargetBox="[135,1176,216,1570]" captionTargetId="figure-12@52.[120,1176,216,1570]" captionTargetPageId="52" captionText="FIGURE 20. Trechus qiqiensis sp. nov.; a. Dorsal habitus (CASENT1007384). b. Median lobe of aedeagus of male (CASENT1007384), left lateral aspect. c. Map of locality records (red circles) for T. qiqiensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155322" httpUri="https://zenodo.org/record/13155322/files/figure.png" pageId="53" pageNumber="394">Fig. 20c</figureCitation>
.
<collectingMunicipality id="6BA4ACF1FFA0FFBC3FBC6634FBC0710C" box="[1003,1038,1605,1628]" pageId="53" pageNumber="394">We</collectingMunicipality>
examined a total of
<specimenCount id="9D79FD02FFA0FFBC3C846614FEA8712D" box="[211,358,1637,1661]" count="12" pageId="53" pageNumber="394" type="generic">12 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFA0FFBC3D2F6614FE03712D" box="[376,461,1637,1661]" count="5" pageId="53" pageNumber="394" type="male">5 males</specimenCount>
and
<specimenCount id="9D79FD02FFA0FFBC3E516614FDBA712D" box="[518,628,1637,1661]" count="7" pageId="53" pageNumber="394" type="female">7 females</specimenCount>
), all from
<location id="8EA06050FFA0FFBC3EBD6614FC47712D" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA0FFBC3EBD6614FC47712D" box="[746,905,1637,1661]" county="Cikai Townships in Gongshan County" municipality="We" name="Bingzhiongluo" pageId="53" pageNumber="394" stateProvince="Shan">Bingzhiongluo</location>
and
<collectingCounty id="62A14E07FFA0FFBC3F956614FEF071CD" pageId="53" pageNumber="394">Cikai Townships in Gongshan County</collectingCounty>
on the crest and eastern slope of the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFA0FFBC387A66F4FBC171ED" location="Core Area" pageId="53" pageNumber="394" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFA0FFBC387A66F4FBAC71CD" box="[1069,1122,1669,1693]" country="Myanmar" name="Shan" pageId="53" pageNumber="394">Shan</collectingRegion>
(see
<typeStatus id="54C48829FFA0FFBC3C2C66D7FF7E71EE" box="[123,176,1702,1726]" pageId="53" pageNumber="394">Type</typeStatus>
material above for exact collection data). These localities are all in
<location id="8EA06050FFA0FFBC3FDC66D4FC3671EE" LSID="urn:lsid:plazi:treatment:03D6879DFFA6FFBF3C2C60AAFE497663:8EA06050FFA0FFBC3FDC66D4FC3671EE" box="[907,1016,1701,1726]" name="Core Area" pageId="53" pageNumber="394" stateProvince="Shan">Core Area</location>
2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA3FFBF3CFC60AAFE497663" blockId="54.[123,1173,219,307]" pageId="54" pageNumber="395">
<emphasis id="B90BEA99FFA3FFBF3CFC60AAFD9477A1" bold="true" box="[171,602,219,243]" pageId="54" pageNumber="395">
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<smallCapsWord id="8D26A057FFA3FFBF3D4E60AFFE0E77A1" baselines="236" box="[281,448,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="54" pageNumber="395">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFA3FFBF3D9160AFFD9477A1" baselines="236" box="[454,602,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="54" pageNumber="395">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Gongshan County on the crest and eastern slope of the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFA3FFBF3C2C616DFEFB7663" box="[123,309,283,307]" country="China" name="Yunnan" pageId="54" pageNumber="395">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFA3FFBF3D17616AFE4D7663" box="[320,387,283,307]" name="China" pageId="54" pageNumber="395">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,801 @@
<document id="03F10A57E965C10B7F27E18FBACE0426" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFABFFCB3C2C648DFB9F7623" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechus (Trechus) shibalicus Deuve &amp; Kavanaugh &amp; Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="407" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="403" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C1B1442DDF557FC7E3E73730CDF5A86D">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:role id="B55923E452269D3B6780118FE2C3CD0A">
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<mods:namePart id="051EC4D9D1369595ACD4F5E97A575369">Deuve, Thierry</mods:namePart>
<mods:affiliation id="3D949FBA488B69492CE94BE516739A92">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
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<mods:role id="E3A2B694A4962510D9DBD86BEDBD2351">
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</mods:role>
<mods:namePart id="3569DD057D0BABA95ACFF4AA54C57D7E">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="CB09BAFB8CB02768BC51A0820FE276EA">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="6FC87478A7EDBCBE14E87AADC76350AC" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="9B2AAD25AEB339EADD30F39B3A699C6C" type="personal">
<mods:role id="7DA97238D527AFD3766B053875564531">
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</mods:role>
<mods:namePart id="8A92CEEC8FE3FEFA5232D9A91EA5B44D">Liang, Hongbin</mods:namePart>
<mods:affiliation id="E79616CDE1EC02B5910F1088435AD396">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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</mods:titleInfo>
<mods:part id="4402ED54A1AA28C460B51218F1436913">
<mods:date id="094689D04E49F6D23C98913934E3604F">2016</mods:date>
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<mods:number id="29BB0E8BA4D63E27C34C7548CFB25B96">2016-10-14</mods:number>
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<mods:number id="88AED6CEE61EBFAC927730D82FEF2D6A">63</mods:number>
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<mods:number id="75526713DFAE44D4C413F4E52BA77768">12</mods:number>
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<treatment id="03D6879DFFABFFCB3C2C648DFB9F7623" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623" httpUri="http://treatment.plazi.org/id/03D6879DFFABFFCB3C2C648DFB9F7623" lastPageId="66" lastPageNumber="407" pageId="62" pageNumber="403">
<subSubSection id="C3656500FFABFFB73C2C648DFCE37244" box="[123,813,1276,1300]" pageId="62" pageNumber="403" type="nomenclature">
<paragraph id="8BC0368BFFABFFB73C2C648DFCE37244" blockId="62.[123,813,1276,1330]" box="[123,813,1276,1300]" pageId="62" pageNumber="403">
<heading id="D08881E7FFABFFB73C2C648DFCE37244" box="[123,813,1276,1300]" fontSize="10" level="2" pageId="62" pageNumber="403" reason="4">
<emphasis id="B90BEA99FFABFFB73C2C648DFCE37244" bold="true" box="[123,813,1276,1300]" pageId="62" pageNumber="403">
<taxonomicName id="4C7F4D08FFABFFB73C2C648DFDC57244" ID-CoL="8MFHT" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,523,1276,1300]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="403" phylum="Arthropoda" rank="species" species="shibalicus" subGenus="Trechus">
<emphasis id="B90BEA99FFABFFB73C2C648DFF1A7244" bold="true" box="[123,212,1276,1300]" italics="true" pageId="62" pageNumber="403">Trechus</emphasis>
(
<emphasis id="B90BEA99FFABFFB73CB3648DFEF37244" bold="true" box="[228,317,1276,1300]" italics="true" pageId="62" pageNumber="403">Trechus</emphasis>
)
<emphasis id="B90BEA99FFABFFB73D19648DFE737244" bold="true" box="[334,445,1276,1300]" italics="true" pageId="62" pageNumber="403">shibalicus</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFABFFB73E80648DFCE37244" box="[727,813,1276,1300]" pageId="62" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFABFFB73C2C656DFE7D7262" box="[123,435,1308,1330]" pageId="62" pageNumber="403" type="description">
<paragraph id="8BC0368BFFABFFB73C2C656DFE7D7262" blockId="62.[123,813,1276,1330]" box="[123,435,1308,1330]" pageId="62" pageNumber="403">
(
<figureCitation id="13442A0EFFABFFB73CD5656DFF1D7261" box="[130,211,1308,1330]" captionStart="FIGURE" captionStartId="64.[171,183,1594,1613]" captionTargetBox="[120,1176,216,1574]" captionTargetId="figure-12@64.[120,1176,216,1575]" captionTargetPageId="64" captionText="FIGURE 24. Trechus shibalicus sp. nov.; a. Dorsal habitus (CASENT1017532). b. Median lobe of aedeagus of male (CASENT1017532), left lateral aspect. c. Map of locality records (red circles) for T. shibalicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155330" httpUri="https://zenodo.org/record/13155330/files/figure.png" pageId="62" pageNumber="403">Figs. 24</figureCitation>
,
<figureCitation id="13442A0EFFABFFB73C8A656DFECD7262" box="[221,259,1308,1330]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="62" pageNumber="403">16d</figureCitation>
,
<figureCitation id="13442A0EFFABFFB73D59656DFEFA7262" box="[270,308,1308,1330]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="62" pageNumber="403">40b</figureCitation>
,
<figureCitation id="13442A0EFFABFFB73D69656DFEAA7262" box="[318,356,1308,1330]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="62" pageNumber="403">42b</figureCitation>
,
<figureCitation id="13442A0EFFABFFB73D38656DFE627262" box="[367,428,1308,1330]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="62" pageNumber="403">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFABFFB63CFC6534FCDB72CB" lastPageId="63" lastPageNumber="404" pageId="62" pageNumber="403" type="materials_examined">
<paragraph id="8BC0368BFFABFFB63CFC6534FB4473AA" blockId="62.[123,1174,1349,1726]" lastBlockId="63.[123,1174,219,1725]" lastPageId="63" lastPageNumber="404" pageId="62" pageNumber="403">
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<smallCapsWord id="8D26A057FFABFFB73CBA6539FE94720B" baselines="1366" box="[237,346,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="62" pageNumber="403">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFABFFB73DD46534FCE5728D" collectingDate="2005-08-08" collectorName="H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Fugong County" elevation="3100" latitude="27.18416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" pageId="62" pageNumber="403" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFABFFB73DD46534FE27720D" box="[387,489,1349,1373]" pageId="62" pageNumber="403" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1017532”/ “
<collectingCountry id="F368761BFFABFFB73FAA6534FB98720D" box="[1021,1110,1349,1373]" name="China" pageId="62" pageNumber="403">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFABFFB738356537FF6A722D" country="China" name="Yunnan" pageId="62" pageNumber="403">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFABFFB73CF96517FE9B722D" box="[174,341,1381,1406]" pageId="62" pageNumber="403">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFABFFB73D096514FDD1722D" box="[350,543,1381,1405]" pageId="62" pageNumber="403">Lishadi Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FFABFFB73E7F6514FC74722D" box="[552,954,1381,1405]" location="Shibali on Shibali Road" pageId="62" pageNumber="403">
<quantity id="4C879B6EFFABFFB73E7F6514FDBF722D" box="[552,625,1381,1405]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="62" pageNumber="403" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
, north bank of
<location id="8EA06050FFABFFB738006514FEF372CD" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFABFFB738006514FEF372CD" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" name="North Fork of Yamu He" pageId="62" pageNumber="403" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFABFFB73D1D65F4FE0572CD" box="[330,459,1413,1437]" degrees="27.18416" direction="north" orientation="latitude" pageId="62" pageNumber="403" precision="1" value="27.18416">N27.18416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFABFFB73D8B65F4FD9072CD" box="[476,606,1413,1437]" degrees="98.72026" direction="east" orientation="longitude" pageId="62" pageNumber="403" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFABFFB73ED065F4FD1072CD" box="[647,734,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="62" pageNumber="403" unit="m" value="3100.0">
<elevation id="0052D1B8FFABFFB73ED065F4FD1072CD" box="[647,734,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="62" pageNumber="403" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFABFFB73EBC65F4FC4572CD" box="[747,907,1413,1437]" pageId="62" pageNumber="403" value="2005-08-08">
<collectingDate id="EF85E9A3FFABFFB73EBC65F4FC4572CD" box="[747,907,1413,1437]" pageId="62" pageNumber="403" value="2005-08-08">8 August 2005</collectingDate>
</date>
Stop #DHK-2005-067A D.H.
<taxonomicName id="4C7F4D08FFABFFB73CE365D4FEFE72ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[180,304,1445,1469]" pageId="62" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFABFFB73D6D65D7FE7A72ED" box="[314,436,1445,1469]" pageId="62" pageNumber="403">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFABFFB73DE965D7FDFC72EE" box="[446,562,1446,1470]" pageId="62" pageNumber="403">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFABFFB73E0065D7FD0B72ED" box="[599,709,1445,1469]" pageId="62" pageNumber="403">J.F. Zhang</collectorName>
collectors”/ “
<typeStatus id="54C48829FFABFFB73F0265D4FC2A72ED" box="[853,996,1445,1469]" pageId="62" pageNumber="403" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFABFFB73FBD65D4FF22728D" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="403" phylum="Arthropoda" rank="species" species="shibalicus">
<emphasis id="B90BEA99FFABFFB73FBD65D4FF51728D" italics="true" pageId="62" pageNumber="403">Trechus shibalicus</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFABFFB73DCE65B7FE23728D" box="[409,493,1478,1501]" pageId="62" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FFABFFB73F6065B7FD32714D" collectionCode="IOZ, CAS" country="China" pageId="62" pageNumber="403" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFABFFB73F6065B7FC51728E" box="[823,927,1478,1502]" pageId="62" pageNumber="403" type="paratype">Paratypes</typeStatus>
(a total of 43):
<specimenCount id="9D79FD02FFABFFB7381B65B4FB58728D" box="[1100,1174,1477,1501]" count="1" pageId="62" pageNumber="403" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFABFFB73CF06594FEC072AD" box="[167,270,1509,1533]" count="3" pageId="62" pageNumber="403" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFABFFB73D606594FEA872AD" box="[311,358,1509,1533]" pageId="62" pageNumber="403">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFABFFB73D276594FE6B72AD" box="[368,421,1509,1533]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="62" pageNumber="403" type="Museum">CAS</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFABFFB73E316594FD0872AD" box="[614,710,1509,1533]" pageId="62" pageNumber="403" type="holotype">holotype</typeStatus>
, except first label “CASENT 1017531” and “CASENT 1017533” to “CASENT 1017535”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FFABFFB73F5E6674FC1E712D" collectingDate="2004-05-08" collectionCode="CAS" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="3290" latitude="27.20676" location="Lishadi Township" longLatPrecision="1" longitude="98.71763" pageId="62" pageNumber="403" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFABFFB73F5E6674FC9C714D" box="[777,850,1541,1565]" count="1" pageId="62" pageNumber="403" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFABFFB73FD66674FC78714D" box="[897,950,1541,1565]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="62" pageNumber="403" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1014204”/ “
<collectingCountry id="F368761BFFABFFB73CAA6654FE98716D" box="[253,342,1573,1597]" name="China" pageId="62" pageNumber="403">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFABFFB73D366657FE7B716D" box="[353,437,1574,1597]" country="China" name="Yunnan" pageId="62" pageNumber="403">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFABFFB73D956657FDA5716D" box="[450,619,1573,1598]" pageId="62" pageNumber="403">Fugong County</collectingCounty>
,
<location id="8EA06050FFABFFB73E206654FCF4716D" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFABFFB73E206654FCF4716D" box="[631,826,1573,1597]" country="China" county="Fugong County" latitude="27.20676" longLatPrecision="1" longitude="98.71763" name="Lishadi Township" pageId="62" pageNumber="403" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFABFFB73F116654FF29710D" location="Shibali on Yaping Road" pageId="62" pageNumber="403">
<quantity id="4C879B6EFFABFFB73F116654FC53716D" box="[838,925,1573,1597]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="62" pageNumber="403" unit="km" value="11.5">11.5 km</quantity>
above Shibali on Yaping Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFABFFB73CAC6634FEB2710D" box="[251,380,1605,1629]" degrees="27.20676" direction="north" orientation="latitude" pageId="62" pageNumber="403" precision="1" value="27.20676">N27.20676°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFABFFB73DC36634FDD8710D" box="[404,534,1605,1629]" degrees="98.71763" direction="east" orientation="longitude" pageId="62" pageNumber="403" precision="1" value="98.71763">E98.71763°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFABFFB73E116634FD6B710D" box="[582,677,1605,1629]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="62" pageNumber="403" unit="m" value="3290.0">
<elevation id="0052D1B8FFABFFB73E116634FD6B710D" box="[582,677,1605,1629]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="62" pageNumber="403" unit="m" value="3290.0">3290 m</elevation>
</quantity>
,
<date id="FFC1104BFFABFFB73EEE6634FC85710D" box="[697,843,1605,1630]" pageId="62" pageNumber="403" value="2004-05-08">
<collectingDate id="EF85E9A3FFABFFB73EEE6634FC85710D" box="[697,843,1605,1630]" pageId="62" pageNumber="403" value="2004-05-08">8 May 2004</collectingDate>
</date>
Stop #DHK-2004-040 D.H.
<taxonomicName id="4C7F4D08FFABFFB73C2C6614FF39712D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,1637,1661]" pageId="62" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFABFFB73D516614FE69712D" box="[262,423,1637,1661]" pageId="62" pageNumber="403">C.E. Griswold</collectorName>
, Liang H.-B., Li X.-Y., &amp; Zhu B.-X. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFABFFB63F886614FCDE77A3" collectingDate="2005-08-07" collectionCode="IOZ" collectorName="H. B. Liang &amp; J. F. Zhang" country="China" county="Fugong County" elevation="3450" lastPageId="63" lastPageNumber="404" latitude="27.21034" location="North Fork Yamu He" longLatPrecision="1" longitude="98.70141" municipality="Lishadi Township" pageId="62" pageNumber="403" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFABFFB73F886614FBE2712D" box="[991,1068,1637,1661]" count="1" pageId="62" pageNumber="403" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFABFFB738376614FB40712D" box="[1120,1166,1637,1661]" pageId="62" pageNumber="403">IOZ</collectionCode>
) labeled: “CASENT 1018365”/ “
<collectingCountry id="F368761BFFABFFB73D8266F4FDE071CD" box="[469,558,1669,1693]" name="China" pageId="62" pageNumber="403">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFABFFB73E6E66F7FD4371CD" box="[569,653,1670,1693]" country="China" name="Yunnan" pageId="62" pageNumber="403">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFABFFB73ECE66F7FC8D71CD" box="[665,835,1669,1694]" pageId="62" pageNumber="403">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFABFFB73F1966F4FBDC71CD" box="[846,1042,1669,1693]" pageId="62" pageNumber="403">Lishadi Township</collectingMunicipality>
, headwaters of
<location id="8EA06050FFABFFB73CCE66D4FEB171ED" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFABFFB73CCE66D4FEB171ED" box="[153,383,1701,1725]" country="China" county="Fugong County" latitude="27.21034" longLatPrecision="1" longitude="98.70141" municipality="Lishadi Township" name="North Fork Yamu He" pageId="62" pageNumber="403" stateProvince="Yunnan">North Fork Yamu He</location>
just
<locationDeviation id="9A722FA9FFABFFB73DE066D7FD4671ED" box="[439,648,1701,1725]" location="Shibali Yakou" pageId="62" pageNumber="403">E of Shibali Yakou</locationDeviation>
,
<quantity id="4C879B6EFFABFFB73EC366D4FD2771ED" box="[660,745,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="62" pageNumber="403" unit="m" value="3450.0">
<elevation id="0052D1B8FFABFFB73EC366D4FD2771ED" box="[660,745,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="62" pageNumber="403" unit="m" value="3450.0">3450 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFABFFB73F4566D4FC5B71ED" box="[786,917,1701,1725]" degrees="27.21034" direction="north" orientation="latitude" pageId="62" pageNumber="403" precision="1" value="27.21034">N27.21034°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFABFFB73FF166D4FBE971ED" box="[934,1063,1701,1725]" degrees="98.70141" direction="east" orientation="longitude" pageId="62" pageNumber="403" precision="1" value="98.70141">E98.70141°</geoCoordinate>
,
<date id="FFC1104BFFABFFB6386366D7FF7D77A3" lastPageId="63" lastPageNumber="404" pageId="62" pageNumber="403" value="2005-08-07">
<collectingDate id="EF85E9A3FFABFFB6386366D7FF7D77A3" lastPageId="63" lastPageNumber="404" pageId="62" pageNumber="403" value="2005-08-07">7 August 2005</collectingDate>
</date>
, Stop# LHB-05-52,
<collectorName id="268A535DFFAAFFB63DDC60AAFDCC77A3" box="[395,514,219,243]" pageId="63" pageNumber="404">H.B. Liang</collectorName>
&amp;
<collectorName id="268A535DFFAAFFB63E7460AAFD5C77A3" box="[547,658,219,243]" pageId="63" pageNumber="404">J.F. Zhang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63F4D60AAFDE87604" collectingDate="2005-08-12" collectionCode="IOZ" collectorName="H. B. Liang &amp; J. F. Zhang" country="China" county="Fugong County" elevation="3450" latitude="27.21034" location="North Fork Yamu He" longLatPrecision="1" longitude="98.70141" municipality="Lishadi Township" pageId="63" pageNumber="404" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63F4D60AAFCAC77A3" box="[794,866,219,243]" count="1" pageId="63" pageNumber="404" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63FDB60AAFC7477A3" box="[908,954,219,243]" pageId="63" pageNumber="404">IOZ</collectionCode>
) labeled: “CASENT 1018605”/ “
<collectingCountry id="F368761BFFAAFFB63CAB608AFE9B7643" box="[252,341,251,275]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63D08608DFE7D7643" box="[351,435,252,275]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63DE9608DFDA87643" box="[446,614,251,276]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAAFFB63E26608AFCFD7643" box="[625,819,251,275]" pageId="63" pageNumber="404">Lishadi Township</collectingMunicipality>
, headwaters of
<location id="8EA06050FFAAFFB63F8E608AFF547663" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63F8E608AFF547663" country="China" county="Fugong County" latitude="27.21034" longLatPrecision="1" longitude="98.70141" municipality="Lishadi Township" name="North Fork Yamu He" pageId="63" pageNumber="404" stateProvince="Yunnan">North Fork Yamu He</location>
just
<locationDeviation id="9A722FA9FFAAFFB63C9A616DFE567663" box="[205,408,283,307]" location="Shibali Yakou" pageId="63" pageNumber="404">E of Shibali Yakou</locationDeviation>
,
<quantity id="4C879B6EFFAAFFB63DF5616AFE3B7664" box="[418,501,283,308]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="63" pageNumber="404" unit="m" value="3450.0">
<elevation id="0052D1B8FFAAFFB63DF5616AFE3B7664" box="[418,501,283,308]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="63" pageNumber="404" unit="m" value="3450.0">3450 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFAAFFB63E4B616DFD517664" box="[540,671,284,308]" degrees="27.21034" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.21034">N27.21034°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63EF9616DFCE17664" box="[686,815,284,308]" degrees="98.70141" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.70141">E98.70141°</geoCoordinate>
,
<date id="FFC1104BFFAAFFB63F6D616DFC2B7663" box="[826,997,283,308]" pageId="63" pageNumber="404" value="2005-08-12">
<collectingDate id="EF85E9A3FFAAFFB63F6D616DFC2B7663" box="[826,997,283,308]" pageId="63" pageNumber="404" value="2005-08-12">12 August 2005</collectingDate>
</date>
, Stop# LHB-05- 54,
<collectorName id="268A535DFFAAFFB63CF5614DFED77604" box="[162,281,316,340]" pageId="63" pageNumber="404">H.B. Liang</collectorName>
&amp;
<collectorName id="268A535DFFAAFFB63D6D614DFE667604" box="[314,424,316,340]" pageId="63" pageNumber="404">J.F. Zhang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63E67614DFF1F76E4" collectingDate="2005-08-04" collectingDateMax="2005-08-17" collectingDateMin="2005-08-04" collectionCode="CAS" collectorName="H. B. Liang &amp; P. Paquin &amp; D. Z. Dong" country="China" county="Fugong County" elevation="2535" latitude="27.16536" location="Shibali area" longLatPrecision="1" longitude="98.78003" municipality="Lishadi Township" pageId="63" pageNumber="404" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63E67614DFDB97604" box="[560,631,316,340]" count="1" pageId="63" pageNumber="404" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63EF6614DFD187604" box="[673,726,316,340]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1018844”/ “
<collectingCountry id="F368761BFFAAFFB6386F614DFB5F7604" box="[1080,1169,316,340]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63C2C612DFF017623" box="[123,207,348,371]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63C8D612DFE4D7624" box="[218,387,348,372]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAAFFB63DDA612DFD9E7624" box="[397,592,348,372]" pageId="63" pageNumber="404">Lishadi Township</collectingMunicipality>
,
<location id="8EA06050FFAAFFB63E0C612DFD137623" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63E0C612DFD137623" box="[603,733,348,372]" country="China" county="Fugong County" latitude="27.16536" longLatPrecision="1" longitude="98.78003" municipality="Lishadi Township" name="Shibali area" pageId="63" pageNumber="404" stateProvince="Yunnan">Shibali area</location>
,
<quantity id="4C879B6EFFAAFFB63EBF612DFCF27623" box="[744,828,348,372]" metricMagnitude="3" metricUnit="m" metricValue="2.535" pageId="63" pageNumber="404" unit="m" value="2535.0">
<elevation id="0052D1B8FFAAFFB63EBF612DFCF27623" box="[744,828,348,372]" metricMagnitude="3" metricUnit="m" metricValue="2.535" pageId="63" pageNumber="404" unit="m" value="2535.0">2535 m</elevation>
</quantity>
,
<geoCoordinate id="EE4B504CFFAAFFB63F10612DFC067624" box="[839,968,348,372]" degrees="27.16536" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.16536">N27.16536°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63F8F612DFB977624" box="[984,1113,348,372]" degrees="98.78003" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.78003">E98.78003°</geoCoordinate>
,
<date id="FFC1104BFFAAFFB63832612DFEC976C4" pageId="63" pageNumber="404" value="2005-08-04" valueMax="2005-08-17" valueMin="2005-08-04">
<collectingDate id="EF85E9A3FFAAFFB63832612DFEC976C4" pageId="63" pageNumber="404" value="2005-08-04" valueMax="2005-08-17" valueMin="2005-08-04">4-17 August 2005</collectingDate>
</date>
,”/ “Stop# DHK-2005-059 D.H.
<taxonomicName id="4C7F4D08FFAAFFB63E0A610DFD1776C4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[605,729,380,404]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAAFFB63EB3610CFC9076C4" box="[740,862,380,404]" pageId="63" pageNumber="404">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFAAFFB63F3F610CFC0076C4" box="[872,974,380,404]" pageId="63" pageNumber="404">P. Paquin</collectorName>
, &amp;
<collectorName id="268A535DFFAAFFB63FA4610CFBAB76C5" box="[1011,1125,381,405]" pageId="63" pageNumber="404">D.Z. Dong</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63CB761EDFCF876A5" collectingDate="2005-08-17" collectionCode="CAS" country="China" county="Fugong County" elevation="3675" latitude="27.20244" location="Lumadeng Township" longLatPrecision="1" longitude="98.69526" pageId="63" pageNumber="404" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63CB761EDFE8F76E4" box="[224,321,412,436]" count="1" pageId="63" pageNumber="404" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63D2461EDFE6676E4" box="[371,424,412,436]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1020015”/ “
<collectingCountry id="F368761BFFAAFFB63F4B61EDFCBB76E4" box="[796,885,412,436]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63FD461ECFC1976E4" box="[899,983,413,436]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63FB161ECFB5C76E4" box="[998,1170,412,437]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<location id="8EA06050FFAAFFB63C2C61CDFEAE7684" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63C2C61CDFEAE7684" box="[123,352,444,468]" country="China" county="Fugong County" latitude="27.20244" longLatPrecision="1" longitude="98.69526" name="Lumadeng Township" pageId="63" pageNumber="404" stateProvince="Yunnan">Lumadeng Township</location>
, second cirque
<locationDeviation id="9A722FA9FFAAFFB63E5161CCFD1D7684" box="[518,723,444,469]" location="Shibali Yakou" pageId="63" pageNumber="404">S of Shibali Yakou</locationDeviation>
,
<quantity id="4C879B6EFFAAFFB63E8961CDFCFC7685" box="[734,818,444,469]" metricMagnitude="3" metricUnit="m" metricValue="3.675" pageId="63" pageNumber="404" unit="m" value="3675.0">
<elevation id="0052D1B8FFAAFFB63E8961CDFCFC7685" box="[734,818,444,469]" metricMagnitude="3" metricUnit="m" metricValue="3.675" pageId="63" pageNumber="404" unit="m" value="3675.0">3675 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFAAFFB63F0C61CCFC107685" box="[859,990,445,469]" degrees="27.20244" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.20244">N27.20244°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63FB961CDFBA17684" box="[1006,1135,444,468]" degrees="98.69526" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.69526">E98.69526°</geoCoordinate>
,
<date id="FFC1104BFFAAFFB6382C61CCFEC976A5" pageId="63" pageNumber="404" value="2005-08-17">
<collectingDate id="EF85E9A3FFAAFFB6382C61CCFEC976A5" pageId="63" pageNumber="404" value="2005-08-17">17 August 2005</collectingDate>
</date>
, Stop# DHK-2005-093, D.H.
<taxonomicName id="4C7F4D08FFAAFFB63E1E61ACFD0C76A5" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[585,706,477,501]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63F1661ACFC8D7526" collectingDate="2005-08-07" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; P. Paquin &amp; D. Z. Dong" country="China" county="Fugong County" elevation="3612" latitude="27.21231" location="Shibali Yakou" longLatPrecision="1" longitude="98.69575" municipality="Lishadi Township" pageId="63" pageNumber="404" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63F1661ACFC5D76A5" box="[833,915,477,501]" count="2" pageId="63" pageNumber="404" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FFAAFFB63F9F61ACFBE176A5" box="[968,1071,477,501]" count="2" pageId="63" pageNumber="404" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB6380C61ACFB5F76A5" box="[1115,1169,477,501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAAFFB63C2C618CFF677545" box="[123,169,509,533]" pageId="63" pageNumber="404">IOZ</collectionCode>
) labeled: “CASENT 1020903” to “CASENT 1020904” and “CASENT 1020905” to “CASENT 1020906”, respectively/ “
<collectingCountry id="F368761BFFAAFFB63E52626CFD907565" box="[517,606,541,565]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63E3E626FFD737565" box="[617,701,542,565]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63E9E626FFCBF7565" box="[713,881,541,566]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAAFFB63F2B626CFBF17565" box="[892,1087,541,565]" pageId="63" pageNumber="404">Lishadi Township</collectingMunicipality>
,
<location id="8EA06050FFAAFFB6381D626CFF0C7505" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB6381D626CFF0C7505" country="China" county="Fugong County" latitude="27.21231" longLatPrecision="1" longitude="98.69575" municipality="Lishadi Township" name="Shibali Yakou" pageId="63" pageNumber="404" stateProvince="Yunnan">Shibali Yakou</location>
,
<quantity id="4C879B6EFFAAFFB63C82624CFEFF7505" box="[213,305,573,597]" metricMagnitude="3" metricUnit="m" metricValue="3.6119999999999997" pageId="63" pageNumber="404" unit="m" value="3612.0">
<elevation id="0052D1B8FFAAFFB63C82624CFEFF7505" box="[213,305,573,597]" metricMagnitude="3" metricUnit="m" metricValue="3.6119999999999997" pageId="63" pageNumber="404" unit="m" value="3612.0">3612 m</elevation>
</quantity>
,
<geoCoordinate id="EE4B504CFFAAFFB63D12624CFE087505" box="[325,454,573,597]" degrees="27.21231" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.21231">N27.21231°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63D89624CFD917505" box="[478,607,573,597]" degrees="98.69575" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.69575">E98.69575°</geoCoordinate>
,
<date id="FFC1104BFFAAFFB63E24624FFCED7505" box="[627,803,573,598]" pageId="63" pageNumber="404" value="2005-08-07">
<collectingDate id="EF85E9A3FFAAFFB63E24624FFCED7505" box="[627,803,573,598]" pageId="63" pageNumber="404" value="2005-08-07">7 August 2005</collectingDate>
</date>
,”/ “Stop# DHK-2005-066, D.H.
<taxonomicName id="4C7F4D08FFAAFFB63C2C622FFF397526" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,606,630]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAAFFB63D5D622FFE427526" box="[266,396,606,630]" pageId="63" pageNumber="404">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFAAFFB63DC9622FFDC37526" box="[414,525,606,630]" pageId="63" pageNumber="404">P. Paquin</collectorName>
, &amp;
<collectorName id="268A535DFFAAFFB63E14622FFD737526" box="[579,701,606,630]" pageId="63" pageNumber="404">D.Z. Dong</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63F01622FFD427586" collectingDate="2005-08-12" collectionCode="CAS" collectorName="P. Paquin &amp; D. Z. Dong" country="China" county="Fugong County" elevation="3460" latitude="27.21447" location="Lishadi Township" longLatPrecision="1" longitude="98.70064" pageId="63" pageNumber="404" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63F01622FFC757526" box="[854,955,606,630]" count="1" pageId="63" pageNumber="404" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63FA1622FFBE57526" box="[1014,1067,606,630]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1021290”/ “
<collectingCountry id="F368761BFFAAFFB63D22620FFE0075C6" box="[373,462,638,662]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63D80620FFDE575C5" box="[471,555,638,661]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63E61620FFD1075C6" box="[566,734,638,662]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<location id="8EA06050FFAAFFB63EB0620FFC6775C6" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63EB0620FFC6775C6" box="[743,937,638,662]" country="China" county="Fugong County" latitude="27.21447" longLatPrecision="1" longitude="98.70064" name="Lishadi Township" pageId="63" pageNumber="404" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFAAFFB63FE4620FFF0C75E6" location="Shibali Yakou" pageId="63" pageNumber="404">
<quantity id="4C879B6EFFAAFFB63FE4620FFC3375C6" box="[947,1021,638,662]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="404" unit="km" value="0.5">0.5 km</quantity>
NE of Shibali Yakou
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFAAFFB63C8362EFFE9B75E6" box="[212,341,670,694]" degrees="27.21447" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.21447">N27.21447°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63D3C62EFFE2075E6" box="[363,494,670,694]" degrees="98.70064" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.70064">E98.70064°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFAAFFB63E4B62EFFDB675E5" box="[540,632,670,694]" metricMagnitude="3" metricUnit="m" metricValue="3.46" pageId="63" pageNumber="404" unit="m" value="3460.0">
<elevation id="0052D1B8FFAAFFB63E4B62EFFDB675E5" box="[540,632,670,694]" metricMagnitude="3" metricUnit="m" metricValue="3.46" pageId="63" pageNumber="404" unit="m" value="3460.0">3460 m</elevation>
</quantity>
,
<date id="FFC1104BFFAAFFB63EDC62EFFC8B75E6" box="[651,837,670,694]" pageId="63" pageNumber="404" value="2005-08-12">
<collectingDate id="EF85E9A3FFAAFFB63EDC62EFFC8B75E6" box="[651,837,670,694]" pageId="63" pageNumber="404" value="2005-08-12">12 August 2005</collectingDate>
</date>
, Stop# DHK-2005-077, D.H.
<taxonomicName id="4C7F4D08FFAAFFB63C2C62CFFF397586" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,702,726]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAAFFB63D5362CEFEA27586" box="[260,364,702,726]" pageId="63" pageNumber="404">P. Paquin</collectorName>
, &amp;
<collectorName id="268A535DFFAAFFB63DC062CEFDC27587" box="[407,524,703,727]" pageId="63" pageNumber="404">D.Z. Dong</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63ECE62CFFC8A7407" collectingDate="2004-05-05" collectionCode="CAS, IOZ" country="China" county="Fugong County" elevation="3100" latitude="27.18416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" municipality="Lumadeng Township" pageId="63" pageNumber="404" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63ECE62CFFD237586" box="[665,749,702,726]" count="6" pageId="63" pageNumber="404" type="male">6 males</specimenCount>
and
<specimenCount id="9D79FD02FFAAFFB63F7162CFFC417586" box="[806,911,702,726]" count="5" pageId="63" pageNumber="404" type="female">5 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63FE862CFFC3B7586" box="[959,1013,702,726]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAAFFB6385462CFFBFF7586" box="[1027,1073,702,726]" pageId="63" pageNumber="404">IOZ</collectionCode>
) labeled: “CASENT 1022286” to “CASENT 1022291” and “CASENT 1022292” to “CASENT 1022296”, respectively/ “
<collectingCountry id="F368761BFFAAFFB63D41628EFEA17447" box="[278,367,767,791]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63D2C628EFE017446" box="[379,463,767,790]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63D8A628EFD497447" box="[477,647,767,791]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAAFFB63EC3628EFCB57447" box="[660,891,767,791]" pageId="63" pageNumber="404">Lumadeng Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FFAAFFB63FD0628FFEC67467" location="Shibali on Shibali Road" pageId="63" pageNumber="404">
<quantity id="4C879B6EFFAAFFB63FD0628FFC1A7447" box="[903,980,766,791]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="63" pageNumber="404" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<location id="8EA06050FFAAFFB63D45636EFDDD7467" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63D45636EFDDD7467" box="[274,531,799,823]" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" municipality="Lumadeng Township" name="North Fork of Yamu He" pageId="63" pageNumber="404" stateProvince="Yunnan">North Fork of Yamu He</location>
,”/ “
<geoCoordinate id="EE4B504CFFAAFFB63E6D636EFD707467" box="[570,702,799,823]" degrees="27.18416" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.18416">N27.18416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63E9A636EFC817467" box="[717,847,799,823]" degrees="98.72026" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFAAFFB63F21636EFC057466" box="[886,971,799,823]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">
<elevation id="0052D1B8FFAAFFB63F21636EFC057466" box="[886,971,799,823]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFAAFFB63F81636EFB947467" box="[982,1114,799,823]" pageId="63" pageNumber="404" value="2004-05-05">
<collectingDate id="EF85E9A3FFAAFFB63F81636EFB947467" box="[982,1114,799,823]" pageId="63" pageNumber="404" value="2004-05-05">5 May 2004</collectingDate>
</date>
, Stop #LHB-04-023 Lian H.-B., Li X.-Y., &amp; Zhu B.-Q. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63F01634EFC6E74E8" collectingDate="2004-05-06" collectionCode="CAS" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="3290" latitude="27.20676" location="Lishadi Township" longLatPrecision="1" longitude="98.77177" pageId="63" pageNumber="404" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63F01634EFC757407" box="[854,955,831,855]" count="1" pageId="63" pageNumber="404" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63FA1634EFBE57407" box="[1014,1067,831,855]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1023606”/ “
<collectingCountry id="F368761BFFAAFFB63DD4632EFE127427" box="[387,476,863,887]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63DBB6311FD8E7427" box="[492,576,864,887]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63E056311FCCF7427" box="[594,769,863,888]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<location id="8EA06050FFAAFFB63F45632EFC147427" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63F45632EFC147427" box="[786,986,863,887]" country="China" county="Fugong County" latitude="27.20676" longLatPrecision="1" longitude="98.77177" name="Lishadi Township" pageId="63" pageNumber="404" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFAAFFB63FBC632EFE4C74C7" location="Shibali on Shibali Road" pageId="63" pageNumber="404">
<quantity id="4C879B6EFFAAFFB63FBC632EFB867427" box="[1003,1096,863,887]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="63" pageNumber="404" unit="km" value="11.5">11.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFAAFFB63DD8630EFDDE74C7" box="[399,528,895,919]" degrees="27.20676" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.20676">N27.20676°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63E76630EFD7F74C7" box="[545,689,895,919]" degrees="98.771763" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.77177">E98.771763°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFAAFFB63E8D63F1FCFC74C7" box="[730,818,896,920]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="63" pageNumber="404" unit="m" value="3290.0">
<elevation id="0052D1B8FFAAFFB63E8D63F1FCFC74C7" box="[730,818,896,920]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="63" pageNumber="404" unit="m" value="3290.0">3290 m</elevation>
</quantity>
,
<date id="FFC1104BFFAAFFB63F68630EFC0A74C8" box="[831,964,895,920]" pageId="63" pageNumber="404" value="2004-05-06">
<collectingDate id="EF85E9A3FFAAFFB63F68630EFC0A74C8" box="[831,964,895,920]" pageId="63" pageNumber="404" value="2004-05-06">6 May 2004</collectingDate>
</date>
Stop #DHK-2004- 036 D.H.
<taxonomicName id="4C7F4D08FFAAFFB63CB463D1FE9174E8" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[227,351,928,952]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAAFFB63D3E63D1FDCB74E8" box="[361,517,928,952]" pageId="63" pageNumber="404">C.E. Griswold</collectorName>
, Liang H.-B., &amp; Zhu B.-X. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63FFD63D1FC637309" collectingDate="2004-05-07" collectionCode="CAS, IOZ, MNHN" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="3100" latitude="27.18416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" pageId="63" pageNumber="404" specimenCount="13" specimenCount-female="4" specimenCount-male="9" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63FFD63D1FC3274E8" box="[938,1020,928,952]" count="9" pageId="63" pageNumber="404" type="male">9 males</specimenCount>
and
<specimenCount id="9D79FD02FFAAFFB6387863D1FB5874E8" box="[1071,1174,928,952]" count="4" pageId="63" pageNumber="404" type="female">4 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63CF463B1FF177488" box="[163,217,960,984]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAAFFB63CBE63B1FED67488" box="[233,280,960,984]" pageId="63" pageNumber="404">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFAAFFB63D7063B1FEB47488" box="[295,378,960,984]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="63" pageNumber="404">MNHN</collectionCode>
) labeled: “CASENT 1023743” to “CASENT 1023745” and “CASENT 1023747” to “CASENT 1023752” and “CASENT 1023753” to “CASENT 1023756”, respectively/ “
<collectingCountry id="F368761BFFAAFFB63CFF6471FECF7348" box="[168,257,1024,1048]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63D5A6470FEAF7348" box="[269,353,1025,1048]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63D3A6470FDD97348" box="[365,535,1024,1049]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAAFFB63E756471FD287348" box="[546,742,1024,1048]" pageId="63" pageNumber="404">Lishadi Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FFAAFFB63EA66471FB5F7348" box="[753,1169,1024,1048]" location="Shibali on Shibali Road" pageId="63" pageNumber="404">
<quantity id="4C879B6EFFAAFFB63EA66471FCF37348" box="[753,829,1024,1048]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="63" pageNumber="404" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<location id="8EA06050FFAAFFB63C2C6451FE467368" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63C2C6451FE467368" box="[123,392,1056,1080]" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" name="North Fork of Yamu He" pageId="63" pageNumber="404" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFAAFFB63DC16451FDD97368" box="[406,535,1056,1080]" degrees="27.18416" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.18416">N27.18416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63E7E6451FD657368" box="[553,683,1056,1080]" degrees="98.72026" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFAAFFB63E826450FCE37368" box="[725,813,1057,1081]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">
<elevation id="0052D1B8FFAAFFB63E826450FCE37368" box="[725,813,1057,1081]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFAAFFB63F6B6450FC0C7369" box="[828,962,1057,1081]" pageId="63" pageNumber="404" value="2004-05-07">
<collectingDate id="EF85E9A3FFAAFFB63F6B6450FC0C7369" box="[828,962,1057,1081]" pageId="63" pageNumber="404" value="2004-05-07">7 May 2004</collectingDate>
</date>
Stop #DHK-2004- 038A D.H.
<taxonomicName id="4C7F4D08FFAAFFB63CA36430FEBE7309" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[244,368,1089,1113]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAAFFB63D2D6430FDDB7309" box="[378,533,1089,1113]" pageId="63" pageNumber="404">C.E. Griswold</collectorName>
, Liang H.-B., &amp; Zhu B.-X. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAAFFB63FE06430FB4B73AA" collectingDate="2004-05-07" collectionCode="CAS, IOZ" collectorName="All" country="China" county="Fugong County" elevation="3100" latitude="27.18326" location="Lumadeng Township" longLatPrecision="1" longitude="98.72002" pageId="63" pageNumber="404" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAAFFB63FE06430FC307309" box="[951,1022,1089,1113]" count="1" pageId="63" pageNumber="404" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFAAFFB638786430FB5B7309" box="[1071,1173,1089,1113]" count="3" pageId="63" pageNumber="404" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAAFFB63CF66410FF197329" box="[161,215,1121,1145]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="404" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAAFFB63CB26410FEDD7329" box="[229,275,1121,1145]" pageId="63" pageNumber="404">IOZ</collectionCode>
) labeled: “CASENT 1023775” and “CASENT 1023776” to “CASENT 1023778”, respectively/ “
<collectingCountry id="F368761BFFAAFFB63D4164F0FEA173C9" box="[278,367,1153,1177]" name="China" pageId="63" pageNumber="404">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63D2C64F3FE0173C9" box="[379,463,1154,1177]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63D8A64F3FD4973C9" box="[477,647,1153,1178]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<location id="8EA06050FFAAFFB63EC364F0FCB573C9" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63EC364F0FCB573C9" box="[660,891,1153,1177]" country="China" county="Fugong County" latitude="27.18326" longLatPrecision="1" longitude="98.72002" name="Lumadeng Township" pageId="63" pageNumber="404" stateProvince="Yunnan">Lumadeng Township</location>
,
<locationDeviation id="9A722FA9FFAAFFB63FD064F0FEC473E9" location="Shibali on Shibali Road" pageId="63" pageNumber="404">
<quantity id="4C879B6EFFAAFFB63FD064F0FC1A73C9" box="[903,980,1153,1177]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="63" pageNumber="404" unit="km" value="8.5">8.5 km</quantity>
above
<locationDeviation id="9A722FA9FFAAFFB6387164F0FEC473E9" pageId="63" pageNumber="404">Shibali on Shibali Road</locationDeviation>
</locationDeviation>
,
<locationDeviation id="9A722FA9FFAAFFB63D4064D0FDEE73EA" box="[279,544,1185,1210]" pageId="63" pageNumber="404">North Fork of Yamu He</locationDeviation>
,”/ “
<geoCoordinate id="EE4B504CFFAAFFB63E0364D0FD1B73E9" box="[596,725,1185,1209]" degrees="27.18326" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.18326">N27.18326°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63EB164D3FCA673EA" box="[742,872,1186,1210]" degrees="98.72002" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.72002">E98.72002°</geoCoordinate>
,
<quantity id="4C879B6EFFAAFFB63F2264D3FC0473E9" box="[885,970,1186,1210]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">
<elevation id="0052D1B8FFAAFFB63F2264D3FC0473E9" box="[885,970,1186,1210]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFAAFFB63F8064D3FB9573EA" box="[983,1115,1186,1210]" pageId="63" pageNumber="404" value="2004-05-07">
<collectingDate id="EF85E9A3FFAAFFB63F8064D3FB9573EA" box="[983,1115,1186,1210]" pageId="63" pageNumber="404" value="2004-05-07">7 May 2004</collectingDate>
</date>
Stop #DHK-2004-038B D.H.
<taxonomicName id="4C7F4D08FFAAFFB63DC364B3FDC3738A" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[404,525,1218,1242]" pageId="63" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”.
<collectorName id="268A535DFFAAFFB63ECE64B3FD75738A" box="[665,699,1218,1242]" pageId="63" pageNumber="404">All</collectorName>
<typeStatus id="54C48829FFAAFFB63E9D64B3FCFE738A" box="[714,816,1218,1242]" pageId="63" pageNumber="404" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFAAFFB63CD46493FEC473AA" box="[131,266,1250,1274]" pageId="63" pageNumber="404" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFAAFFB63D456493FDE973A9" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[274,551,1250,1274]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="404" phylum="Arthropoda" rank="species" species="shibalicus">
<emphasis id="B90BEA99FFAAFFB63D456493FE1773AA" box="[274,473,1250,1274]" italics="true" pageId="63" pageNumber="404">Trechus shibalicus</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFAAFFB63E866493FCEA73A9" box="[721,804,1250,1273]" pageId="63" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFAAFFB63CFC6573FBC8726A" blockId="63.[123,1174,219,1725]" pageId="63" pageNumber="404">
<materialsCitation id="3B173CD6FFAAFFB63CFC6573FBCC726A" country="China" county="Fugong County" elevation="3100" latitude="27.18416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" pageId="63" pageNumber="404" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFAAFFB63CFC6573FE957249" bold="true" box="[171,347,1282,1306]" pageId="63" pageNumber="404">
<smallCapsWord id="8D26A057FFAAFFB63CFC6573FF2A7249" baselines="1300,1300" box="[171,228,1282,1306]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="63" pageNumber="404">
<typeStatus id="54C48829FFAAFFB63CFC6573FF2A7249" box="[171,228,1282,1306]" pageId="63" pageNumber="404">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFAAFFB63CA76577FE957249" baselines="1300" box="[240,347,1286,1305]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="63" pageNumber="404">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFAAFFB63DD06573FE07724A" box="[391,457,1282,1306]" name="China" pageId="63" pageNumber="404">China</collectingCountry>
,
<collectingRegion id="49BBF869FFAAFFB63D8F6572FDE2724A" box="[472,556,1283,1306]" country="China" name="Yunnan" pageId="63" pageNumber="404">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAAFFB63E6B6572FD26724B" box="[572,744,1283,1307]" pageId="63" pageNumber="404">Fugong County</collectingCounty>
,
<taxonomicName id="4C7F4D08FFAAFFB63EA06573FC23724A" authority="Township" authorityName="Township" box="[759,1005,1282,1306]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="404" phylum="Arthropoda" rank="species" species="shiyueliang">Shiyueliang Township</taxonomicName>
, 8,
<locationDeviation id="9A722FA9FFAAFFB638586573FEB3726B" location="Shibali on Shibali Road" pageId="63" pageNumber="404">
<quantity id="4C879B6EFFAAFFB638586573FB84724A" box="[1039,1098,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="404" unit="km" value="5.0">5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<location id="8EA06050FFAAFFB63DDE6552FD42726A" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFAAFFB63DDE6552FD42726A" box="[393,652,1315,1339]" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" name="North Fork of Yamu He" pageId="63" pageNumber="404" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFAAFFB63ECF6552FCD4726B" box="[664,794,1315,1339]" degrees="27.18416" direction="north" orientation="latitude" pageId="63" pageNumber="404" precision="1" value="27.18416">N27.18416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAAFFB63F746552FC6D726B" box="[803,931,1315,1339]" degrees="98.72026" direction="east" orientation="longitude" pageId="63" pageNumber="404" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,
<quantity id="4C879B6EFFAAFFB63FF96552FBCC726A" box="[942,1026,1315,1339]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">
<elevation id="0052D1B8FFAAFFB63FF96552FBCC726A" box="[942,1026,1315,1339]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="404" unit="m" value="3100.0">3100 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFAAFFB63CFC6532FCDB72CB" blockId="63.[123,1174,219,1725]" pageId="63" pageNumber="404">
<emphasis id="B90BEA99FFAAFFB63CFC6532FE207209" bold="true" box="[171,494,1347,1371]" pageId="63" pageNumber="404">
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<smallCapsWord id="8D26A057FFAAFFB63D626537FE9F7209" baselines="1364" box="[309,337,1350,1369]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="63" pageNumber="404">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFAAFFB63D006537FE647209" baselines="1364" box="[343,426,1350,1369]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="63" pageNumber="404">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFAAFFB63DE66537FE207209" baselines="1364" box="[433,494,1350,1369]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="63" pageNumber="404">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFAAFFB63EA56532FC90720B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[754,862,1347,1371]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="404" phylum="Arthropoda" rank="species" species="shibalicus">
<emphasis id="B90BEA99FFAAFFB63EA56532FC90720B" box="[754,862,1347,1371]" italics="true" pageId="63" pageNumber="404">shibalicus</emphasis>
</taxonomicName>
, is derived from the name of settlement, Shibali, at and near which specimens of the
<typeStatus id="54C48829FFAAFFB63E8A6515FCC5722C" box="[733,779,1380,1404]" pageId="63" pageNumber="404">type</typeStatus>
series were collected, and the Latin adjectival suffix,
<emphasis id="B90BEA99FFAAFFB63D6265F5FEA472CB" box="[309,362,1412,1435]" italics="true" pageId="63" pageNumber="404">-icus</emphasis>
, meaning belonging to or pertaining to.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFAAFFB63CFC65D5FDDB71ED" pageId="63" pageNumber="404" type="diagnosis">
<paragraph id="8BC0368BFFAAFFB63CFC65D5FDDB71ED" blockId="63.[123,1174,219,1725]" pageId="63" pageNumber="404">
<smallCapsWord id="8D26A057FFAAFFB63CFC65D5FEEE72EA" baselines="1462,1461" box="[171,288,1444,1468]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="63" pageNumber="404">
<emphasis id="B90BEA99FFAAFFB63CFC65D5FEEE72EA" bold="true" box="[171,288,1444,1468]" pageId="63" pageNumber="404">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFAAFFB63E1065D5FD6D72EC" box="[583,675,1444,1468]" captionStart="FIGURE" captionStartId="64.[171,183,1594,1613]" captionTargetBox="[120,1176,216,1574]" captionTargetId="figure-12@64.[120,1176,216,1575]" captionTargetPageId="64" captionText="FIGURE 24. Trechus shibalicus sp. nov.; a. Dorsal habitus (CASENT1017532). b. Median lobe of aedeagus of male (CASENT1017532), left lateral aspect. c. Map of locality records (red circles) for T. shibalicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155330" httpUri="https://zenodo.org/record/13155330/files/figure.png" pageId="63" pageNumber="404">Fig. 24a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size small (BL =
<quantity id="4C879B6EFFAAFFB6387865B5FF6672AC" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="63" pageNumber="404" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
), apterous; body color reddish brown; tempora glabrous, short and distinctly convex; pronotum transverse (ratio PW/PL = 1.44), basal angles rectangular or slightly acute and sharp; elytra with striae finely punctate, lateral striae more or less effaced, recurrent stria abruptly interrupted at slight convexity at posterior end of stria 7, two typical discal setae present, preapical seta present and inserted in a discal, forward position near apical one-fourth of elytra on interval 3 next to stria 2; male aedeagus with sagittal aileron present but reduced, apical lamella slender with blunt apex, endophallus with a distinct scaly area.
</paragraph>
</subSubSection>
<caption id="DF006603FFD5FFC93CFC664BFDC471D0" ID-DOI="http://doi.org/10.5281/zenodo.13155330" ID-Zenodo-Dep="13155330" httpUri="https://zenodo.org/record/13155330/files/figure.png" pageId="64" pageNumber="405" startId="64.[171,183,1594,1613]" targetBox="[120,1176,216,1574]" targetPageId="64" targetType="figure">
<paragraph id="8BC0368BFFD5FFC93CFC664BFDC471D0" blockId="64.[123,1174,1594,1664]" pageId="64" pageNumber="405">
<smallCapsWord id="8D26A057FFD5FFC93CFC664BFF26711C" baselines="1608,1608" box="[171,232,1594,1613]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="64" pageNumber="405">FIGURE</smallCapsWord>
24.
<taxonomicName id="4C7F4D08FFD5FFC93D47664BFE7F711D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[272,433,1594,1613]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="405" phylum="Arthropoda" rank="species" species="shibalicus" status="sp. nov.">
<emphasis id="B90BEA99FFD5FFC93D47664BFE7F711D" box="[272,433,1594,1613]" italics="true" pageId="64" pageNumber="405">Trechus shibalicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD5FFC93DE0664BFE30711D" box="[439,510,1594,1613]" pageId="64" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1017532). b. Median lobe of aedeagus of male (CASENT1017532), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFD5FFC93F0F6625FC097136" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[856,967,1619,1638]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="405" phylum="Arthropoda" rank="species" species="shibalicus">
<emphasis id="B90BEA99FFD5FFC93F0F6625FC097136" box="[856,967,1619,1638]" italics="true" pageId="64" pageNumber="405">T. shibalicus</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<subSubSection id="C3656500FFD4FFCB3CFC60AAFB9F7623" lastPageId="66" lastPageNumber="407" pageId="65" pageNumber="406" type="description">
<paragraph id="8BC0368BFFD4FFC83CFC60AAFE547663" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">
<smallCapsWord id="8D26A057FFD4FFC83CFC60AAFEF477A1" baselines="237,236" box="[171,314,219,243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="65" pageNumber="406">
<emphasis id="B90BEA99FFD4FFC83CFC60AAFEF477A1" bold="true" box="[171,314,219,243]" pageId="65" pageNumber="406">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size small, BL =
<quantity id="4C879B6EFFD4FFC83E7160AAFD0677A2" box="[550,712,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="65" pageNumber="406" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
. Body color dark brown and shiny, elytra slightly reddish near sutural and lateral margins, appendages paler reddish yellow, palpi paler yellow. Body surface smooth.
</paragraph>
<paragraph id="8BC0368BFFD4FFC83CFC614DFE067505" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">
Head. Short and thick, eyes slightly convex, their diameter slightly greater than length of tempora, the latter glabrous and distinctly convex. Frons not flattened; frontal furrows deep, broadly rounded, not attenuated posteriorly. Two pairs of supraorbital setae present, the anterior pair insert- ed in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible as in
<figureCitation id="13442A0EFFD4FFC83D0C61CDFE787684" box="[347,438,444,468]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="65" pageNumber="406">Fig. 16d</figureCitation>
.
<collectingRegion id="49BBF869FFD4FFC83D9761CCFDD47684" box="[448,538,445,468]" country="Myanmar" name="Chin" pageId="65" pageNumber="406">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFD4FFC83F6A61CCFC597684" box="[829,919,445,468]" country="Myanmar" name="Chin" pageId="65" pageNumber="406">Mentum</collectingRegion>
with median tooth bifid apically, half as long as lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae rather short, with about 1.5 to 2 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter.
</paragraph>
<paragraph id="8BC0368BFFD4FFC83CFC622FFCF97407" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">Pronotum transverse (ratio PW/PL = 1.44), greatest width slightly anterior to middle, slightly narrowed posteriorly, lateral margins rounded, abruptly sinuate just anterior to basal angles, the latter rectangular or slightly acute and sharp. Disc convex; median longitudinal impression distinctly impressed, slightly widened and deepened in the median basal area, the latter also with several longitudinal rugulae and delimited laterally by short, oblique furrows. Basal foveae distinct, small and rounded. Lateral margination narrow and slightly widened basally, lateral border slightly reflexed, lateral grooves distinctly impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFD4FFC83CFC632EFB587368" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">
Elytra. Ovoid, humeri distinct but rounded. Disc convex; striae finely punctate, striae 1 to 4 distinctly impressed, the more lateral striae more or less effaced, stria 6 barely perceptible, stria 7 and 8 indistinct. Scutellar striole short but deeply impressed. Recurrent stria distinct, abruptly interrupted at slight convexity at posterior end of stria 7. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a prediscal position on interval 3 next to stria
<quantity id="4C879B6EFFD4FFC83D696450FEA67368" box="[318,360,1056,1081]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="65" pageNumber="406" unit="in" value="2.0">2 in</quantity>
forward position more than twice as far from apex as from sutural margin\.
</paragraph>
<paragraph id="8BC0368BFFD4FFC83CFC6430FE9E7329" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFD4FFC83CFC64F0FB8573E9" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFD4FFC83CFC64B3FC72724A" blockId="65.[123,1174,219,1726]" pageId="65" pageNumber="406">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFD4FFC83DBC64B3FD89738A" box="[491,583,1218,1242]" captionStart="FIGURE" captionStartId="64.[171,183,1594,1613]" captionTargetBox="[120,1176,216,1574]" captionTargetId="figure-12@64.[120,1176,216,1575]" captionTargetPageId="64" captionText="FIGURE 24. Trechus shibalicus sp. nov.; a. Dorsal habitus (CASENT1017532). b. Median lobe of aedeagus of male (CASENT1017532), left lateral aspect. c. Map of locality records (red circles) for T. shibalicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155330" httpUri="https://zenodo.org/record/13155330/files/figure.png" pageId="65" pageNumber="406">Fig. 24b</figureCitation>
) with base moderate in size, sagittal aileron present but reduced, shaft moderately thick subbasally, apical lamella narrowed, moderately elongate, with blunt apex; endophallus only faintly sclerotized but with a distinct scaly area.
</paragraph>
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</emphasis>
.— Members of this species have been found in a variety of microhabitats over a broad elevational range, from
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. They have been collected in daytime from under stones in thickets of bamboo (various elevations) (
<figureCitation id="13442A0EFFD4FFC83F7C6512FC45722B" box="[811,907,1379,1403]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="65" pageNumber="406">Fig. 42b</figureCitation>
), in subalpine forests of scattered
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</taxonomicName>
and
<emphasis id="B90BEA99FFD4FFC83D0765F2FE3572CB" box="[336,507,1411,1435]" italics="true" pageId="65" pageNumber="406">Rhodondendron</emphasis>
mixed with bamboo thickets (at
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), in rocky open areas cleared by snow avalanches but shaded by
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high herbaceous cover (
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) (
<figureCitation id="13442A0EFFD4FFC8385865D5FBA372EC" box="[1039,1133,1444,1468]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="65" pageNumber="406">Fig. 40b</figureCitation>
), in meadows adjacent to bamboo and
<emphasis id="B90BEA99FFD4FFC83DA465B5FD50728C" box="[499,670,1476,1500]" italics="true" pageId="65" pageNumber="406">Rhodondendron</emphasis>
thickets (
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) and along roadcuts through such thickets, and on talus and vegtetated slopes in a glacial cirque (
<quantity id="4C879B6EFFD4FFC83F196595FC6D72AC" box="[846,931,1508,1532]" metricMagnitude="3" metricUnit="m" metricValue="3.675" pageId="65" pageNumber="406" unit="m" value="3675.0">3675 m</quantity>
). At the lowest record- ed elevation (
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),
<specimenCount id="9D79FD02FFD4FFC83D206674FDC4714C" box="[375,522,1540,1564]" count="1" pageId="65" pageNumber="406" type="generic">one specimen</specimenCount>
was collected under a stone along a roadcut on moist, shad- ed ground. At both highest and lowest elevations, they were the only trechine encountered; but they were found syntopic with specimens of
<taxonomicName id="4C7F4D08FFD4FFC83E786634FCC8710D" box="[559,774,1605,1629]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="406" phylum="Arthropoda" rank="species" species="shiyueliang">
<emphasis id="B90BEA99FFD4FFC83E786634FCC8710D" box="[559,774,1605,1629]" italics="true" pageId="65" pageNumber="406">Trechus shiyueliang</emphasis>
</taxonomicName>
,
<emphasis id="B90BEA99FFD4FFC83F426634FE9B712D" italics="true" pageId="65" pageNumber="406">
<taxonomicName id="4C7F4D08FFD4FFC83F426634FBD0710D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[789,1054,1605,1629]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="406" phylum="Arthropoda" rank="species" species="pseudoqiqiensis">Trechus pseudoqiqiensis</taxonomicName>
,
<taxonomicName id="4C7F4D08FFD4FFC8387D6634FE9B712D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="406" phylum="Arthropoda" rank="species" species="unisetulosa" status="sp. nov.">Trechepaphiopsis unisetulosa</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23857E2FFD4FFC83D096614FE7A712C" box="[350,436,1637,1660]" pageId="65" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C7F4D08FFD4FFC83DB86614FCD8712D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[495,790,1637,1661]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="406" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFD4FFC83DB86614FCD8712D" box="[495,790,1637,1661]" italics="true" pageId="65" pageNumber="406">Trechepaphiopsis unipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD4FFC83F496614FCBC712C" box="[798,882,1637,1660]" pageId="65" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
at one or more mid-elevation (
<quantity id="4C879B6EFFD4FFC83CE466F4FEAE71CD" box="[179,352,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.195" metricValueMax="3.29" metricValueMin="3.1" pageId="65" pageNumber="406" unit="m" value="3195.0" valueMax="3290.0" valueMin="3100.0">3100 to 3290 m</quantity>
) sites.
</paragraph>
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<smallCapsWord id="8D26A057FFD4FFC83D0866D8FE3D71EC" baselines="1719" box="[351,499,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="65" pageNumber="406">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFD4FFC83DAA66D8FD8271EC" baselines="1719" box="[509,588,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="65" pageNumber="406">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFD4FFC83E0166D8FD4E71EC" baselines="1719" box="[598,640,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="65" pageNumber="406">THE</smallCapsWord>
</emphasis>
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<emphasis id="B90BEA99FFD4FFC83EDD66D4FCC171EC" bold="true" box="[650,783,1701,1725]" pageId="65" pageNumber="406">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFD4FFCB3F4E66D4FDA27643" county="Fugong County" lastPageId="66" lastPageNumber="407" location="We" pageId="65" pageNumber="406" specimenCount="88" specimenCount-female="21" specimenCount-male="23" stateProvince="Shan">
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<emphasis id="B90BEA99FFD4FFC83F4E66D4FC9A71EC" bold="true" box="[793,852,1701,1725]" pageId="65" pageNumber="406">
<collectingRegion id="49BBF869FFD4FFC83F4E66D4FC9A71EC" box="[793,852,1701,1725]" country="Myanmar" name="Shan" pageId="65" pageNumber="406">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
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.
<location id="8EA06050FFD4FFC83FBC66D7FBC071ED" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFD4FFC83FBC66D7FBC071ED" box="[1003,1038,1702,1725]" county="Fugong County" name="We" pageId="65" pageNumber="406" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFD7FFCB3C9960AAFE9077A3" box="[206,350,219,243]" count="44" pageId="66" pageNumber="407" type="generic">44 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFD7FFCB3D3A60AAFE0377A3" box="[365,461,219,243]" count="23" pageId="66" pageNumber="407" type="male">23 males</specimenCount>
and
<specimenCount id="9D79FD02FFD7FFCB3E5660AAFDB477A3" box="[513,634,219,243]" count="21" pageId="66" pageNumber="407" type="female">21 females</specimenCount>
), all from
<collectingCounty id="62A14E07FFD7FFCB3EBF60AAFC4177A3" box="[744,911,219,243]" pageId="66" pageNumber="407">Fugong County</collectingCounty>
on the eastern slope and crest of the northcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFD7FFCB3E24608AFD997663" location="Core Area" pageId="66" pageNumber="407" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFD7FFCB3E24608AFD667643" box="[627,680,251,275]" country="Myanmar" name="Shan" pageId="66" pageNumber="407">Shan</collectingRegion>
(see
<typeStatus id="54C48829FFD7FFCB3EB5608AFCD97643" box="[738,791,251,275]" pageId="66" pageNumber="407">Type</typeStatus>
material above for exact collection data). These localities are all in
<location id="8EA06050FFD7FFCB3D85616AFD8E7663" LSID="urn:lsid:plazi:treatment:03D6879DFFABFFCB3C2C648DFB9F7623:8EA06050FFD7FFCB3D85616AFD8E7663" box="[466,576,283,307]" name="Core Area" pageId="66" pageNumber="407" stateProvince="Shan">Core Area</location>
3
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFD7FFCB3CFC614AFB9F7623" blockId="66.[123,1174,219,371]" pageId="66" pageNumber="407">
<emphasis id="B90BEA99FFD7FFCB3CFC614AFD967601" bold="true" box="[171,600,315,339]" pageId="66" pageNumber="407">
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</emphasis>
.— This species currently is known only from Fugong County in the northcentral part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFD7FFCB3F12612DFC317623" box="[837,1023,347,371]" country="China" name="Yunnan" pageId="66" pageNumber="407">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFD7FFCB385D612AFB837623" box="[1034,1101,347,371]" name="China" pageId="66" pageNumber="407">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,347 @@
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<mods:title id="8CC350C5D842DD92EFF244CDD1DB045E">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:affiliation id="483CB1489D5705DDE3C6BA216BEF85BB">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="87F592A67F13462782CA3EA5ABE60BB3">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:classification id="EBF905F978D6FF000A327675A26849C9">journal article</mods:classification>
<mods:identifier id="2FD259DC744FE34F0F06C219D5CA1D6F" type="DOI">10.5281/zenodo.13155283</mods:identifier>
<mods:identifier id="42EECA0D25A1AE285E7FF16530D1E5CF" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="0591420724AF5CCE0BFC845F51B58916" type="Zenodo-Dep">13155283</mods:identifier>
<mods:identifier id="A7E03F97305E1DFDD780C9444B3935B5" type="ZooBank">4C790FE0-B735-4592-8827-EEF83C663CB1</mods:identifier>
</mods:mods>
<treatment id="03D6879DFFADFFB23C2C63BBFF0C76E7" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7" httpUri="http://treatment.plazi.org/id/03D6879DFFADFFB23C2C63BBFF0C76E7" lastPageId="59" lastPageNumber="400" pageId="56" pageNumber="397">
<subSubSection id="C3656500FFADFFB13C2C63BBFCC374B3" box="[123,781,970,995]" pageId="56" pageNumber="397" type="nomenclature">
<paragraph id="8BC0368BFFADFFB13C2C63BBFCC374B3" blockId="56.[123,781,970,1025]" box="[123,781,970,995]" pageId="56" pageNumber="397">
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<emphasis id="B90BEA99FFADFFB13C2C63BBFCC374B3" bold="true" box="[123,781,970,995]" pageId="56" pageNumber="397">
<taxonomicName id="4C7F4D08FFADFFB13C2C63BBFD6474B2" ID-CoL="8MDYW" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,682,970,995]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="397" phylum="Arthropoda" rank="species" species="luzhangensis" status="sp. nov." subGenus="Trechus">
<emphasis id="B90BEA99FFADFFB13C2C63BBFF1A74B2" bold="true" box="[123,212,970,994]" italics="true" pageId="56" pageNumber="397">Trechus</emphasis>
(
<emphasis id="B90BEA99FFADFFB13CB363BBFEF374B2" bold="true" box="[228,317,970,994]" italics="true" pageId="56" pageNumber="397">Trechus</emphasis>
)
<emphasis id="B90BEA99FFADFFB13D1963BBFE1074B2" bold="true" box="[334,478,970,994]" italics="true" pageId="56" pageNumber="397">luzhangensis</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFADFFB13EE163BAFCC374B3" box="[694,781,971,995]" pageId="56" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFADFFB13C2C639AFE9F7351" box="[123,337,1003,1025]" pageId="56" pageNumber="397" type="description">
<paragraph id="8BC0368BFFADFFB13C2C639AFE9F7351" blockId="56.[123,781,970,1025]" box="[123,337,1003,1025]" pageId="56" pageNumber="397">
(
<figureCitation id="13442A0EFFADFFB13CD5639AFF1D7350" box="[130,211,1003,1025]" captionStart="FIGURE" captionStartId="57.[171,183,1583,1602]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@57.[120,1176,216,1561]" captionTargetPageId="57" captionText="FIGURE 22. Trechus luzhangensis sp. nov.; a. Dorsal habitus (CASENT1017595). b. Median lobe of aedeagus of male (CASENT1017595), left lateral aspect. c. Map of locality records (red circles) for T. luzhangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155326" httpUri="https://zenodo.org/record/13155326/files/figure.png" pageId="56" pageNumber="397">Figs. 22</figureCitation>
,
<figureCitation id="13442A0EFFADFFB13C8A639AFECC7350" box="[221,258,1003,1024]" captionStart="FIGURE" captionStartId="105.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1606]" captionTargetId="figure-15@105.[120,1176,216,1607]" captionTargetPageId="105" captionText="FIGURE 43. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Luzhang Township, Piana Road 0.1 km E of Fengxue Yakou, elevation 3150 m; habitat in which specimens of Trechus luzhangensis sp. nov. were collected. b. Lushui County, Luzhang Township, 100 m S of Fengxue Yakou on east side of pass, elevation 3150 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve) were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155372" httpUri="https://zenodo.org/record/13155372/files/figure.png" pageId="56" pageNumber="397">43a</figureCitation>
,
<figureCitation id="13442A0EFFADFFB13D5B639AFE847351" box="[268,330,1003,1025]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="56" pageNumber="397">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFADFFB13CFC6469FD297265" pageId="56" pageNumber="397" type="materials_examined">
<paragraph id="8BC0368BFFADFFB13CFC6469FDFD73A4" blockId="56.[123,1174,1048,1725]" pageId="56" pageNumber="397">
<emphasis id="B90BEA99FFADFFB13CFC6469FE94737E" bold="true" box="[171,346,1048,1072]" pageId="56" pageNumber="397">
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<smallCapsWord id="8D26A057FFADFFB13CBA646AFE94737E" baselines="1065" box="[237,346,1051,1070]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="56" pageNumber="397">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFADFFB13DD46469FE4573E3" collectingDate="2005-05-11" collectorName="H. B. Liang &amp; C. E. Griswold &amp; D. Z. Dong &amp; K. J. Guo" country="China" county="Lushui County" elevation="3150" latitude="25.97228" location="Piana Road" longLatPrecision="1" longitude="98.68336" municipality="Luzhang Township" pageId="56" pageNumber="397" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFADFFB13DD46469FE277360" box="[387,489,1048,1072]" pageId="56" pageNumber="397" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1017595”/ “
<collectingCountry id="F368761BFFADFFB13FAA6469FB987360" box="[1021,1110,1048,1072]" name="China" pageId="56" pageNumber="397">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFADFFB138356469FF6A7300" country="China" name="Yunnan" pageId="56" pageNumber="397">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFADFFB13CF86448FE9E7301" box="[175,336,1081,1105]" pageId="56" pageNumber="397">Lushui County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFADFFB13D0C6448FDE57301" box="[347,555,1081,1105]" pageId="56" pageNumber="397">Luzhang Township</collectingMunicipality>
,
<location id="8EA06050FFADFFB13E616448FD7E7301" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7:8EA06050FFADFFB13E616448FD7E7301" box="[566,688,1081,1105]" country="China" county="Lushui County" latitude="25.97228" longLatPrecision="1" longitude="98.68336" municipality="Luzhang Township" name="Piana Road" pageId="56" pageNumber="397" stateProvince="Yunnan">Piana Road</location>
at
<location id="8EA06050FFADFFB13E856448FCB57301" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7:8EA06050FFADFFB13E856448FCB57301" box="[722,891,1081,1105]" country="China" county="Lushui County" latitude="25.97228" longLatPrecision="1" longitude="98.68336" municipality="Luzhang Township" name="Fengxue Yakou" pageId="56" pageNumber="397" stateProvince="Yunnan">Fengxue Yakou</location>
,
<geoCoordinate id="EE4B504CFFADFFB13FD16449FBC77300" box="[902,1033,1080,1104]" degrees="25.97228" direction="north" orientation="latitude" pageId="56" pageNumber="397" precision="1" value="25.97228">N25.97228°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFADFFB138466449FB5F7300" box="[1041,1169,1080,1104]" degrees="98.68336" direction="east" orientation="longitude" pageId="56" pageNumber="397" precision="1" value="98.68336">E98.68336°</geoCoordinate>
,
<quantity id="4C879B6EFFADFFB13C2C6428FF1F7321" box="[123,209,1113,1137]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="56" pageNumber="397" unit="m" value="3150.0">
<elevation id="0052D1B8FFADFFB13C2C6428FF1F7321" box="[123,209,1113,1137]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="56" pageNumber="397" unit="m" value="3150.0">3150 m</elevation>
</quantity>
,
<date id="FFC1104BFFADFFB13CB76428FEB67321" box="[224,376,1113,1138]" pageId="56" pageNumber="397" value="2005-05-11">
<collectingDate id="EF85E9A3FFADFFB13CB76428FEB67321" box="[224,376,1113,1138]" pageId="56" pageNumber="397" value="2005-05-11">11 May 2005</collectingDate>
</date>
,”/ “Stop# 2005-007, D.H.
<taxonomicName id="4C7F4D08FFADFFB13EF66428FCD37321" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[673,797,1113,1137]" pageId="56" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFADFFB13F7C642BFC677321" box="[811,937,1113,1137]" pageId="56" pageNumber="397">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFADFFB13FE06428FB987321" box="[951,1110,1113,1137]" pageId="56" pageNumber="397">C.E. Griswold</collectorName>
,
<collectorName id="268A535DFFADFFB13833642BFF7873C2" pageId="56" pageNumber="397">D.Z. Dong</collectorName>
&amp;
<collectorName id="268A535DFFADFFB13C8C640BFEF273C2" box="[219,316,1146,1170]" pageId="56" pageNumber="397">K.J. Guo</collectorName>
collectors”/ “
<typeStatus id="54C48829FFADFFB13D87640BFD9173C2" box="[464,607,1146,1170]" pageId="56" pageNumber="397" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFADFFB13E3F640BFBCF73C2" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[616,1025,1146,1170]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="397" phylum="Arthropoda" rank="species" species="luzhangensis" status="sp. nov.">
<emphasis id="B90BEA99FFADFFB13E3F640BFC8073C2" box="[616,846,1146,1170]" italics="true" pageId="56" pageNumber="397">Trechus luzhangensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFADFFB1385A640BFBAC73C1" box="[1037,1122,1146,1169]" pageId="56" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FFADFFB13DC164EAFDE173A4" collectionCode="CAS" country="China" pageId="56" pageNumber="397" specimenCount="3" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFADFFB13DC164EAFE3073E3" box="[406,510,1179,1203]" pageId="56" pageNumber="397" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="ED6EAE4EFFADFFB13EA664EAFCE873E3" box="[753,806,1179,1203]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="56" pageNumber="397" type="Museum">CAS</collectionCode>
) labeled same as
<typeStatus id="54C48829FFADFFB13FB264EAFB8B73E3" box="[997,1093,1179,1203]" pageId="56" pageNumber="397" type="holotype">holotype</typeStatus>
, except first label “CASENT 1017596” and last label “
<typeStatus id="54C48829FFADFFB13E3864CAFD387383" box="[623,758,1211,1235]" pageId="56" pageNumber="397" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFADFFB13EAA64CAFB5F7383" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[765,1169,1211,1235]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="397" phylum="Arthropoda" rank="species" species="luzhangensis" status="sp. nov.">
<emphasis id="B90BEA99FFADFFB13EAA64CAFC2F7383" box="[765,993,1211,1235]" italics="true" pageId="56" pageNumber="397">Trechus luzhangensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFADFFB13C2C64ADFF0073A3" box="[123,206,1244,1267]" pageId="56" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFADFFB13CFC648CFD297265" blockId="56.[123,1174,1048,1725]" pageId="56" pageNumber="397">
<materialsCitation id="3B173CD6FFADFFB13CFC648CFD2D7265" country="China" county="Lushui County" elevation="3150" latitude="25.97228" location="Pianma Road" longLatPrecision="1" longitude="98.68336" municipality="Luzhang Township" pageId="56" pageNumber="397" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54C48829FFADFFB13CFC648CFF2A7243" box="[171,228,1277,1301]" pageId="56" pageNumber="397">TYPE</typeStatus>
</smallCapsWord>
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</emphasis>
.—
<collectingCountry id="F368761BFFADFFB13DDD648CFE027245" box="[394,460,1277,1301]" name="China" pageId="56" pageNumber="397">China</collectingCountry>
,
<collectingRegion id="49BBF869FFADFFB13D89648CFDFC7244" box="[478,562,1277,1300]" country="China" name="Yunnan" pageId="56" pageNumber="397">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFADFFB13E14648CFD247245" box="[579,746,1277,1301]" pageId="56" pageNumber="397">Lushui County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFADFFB13EAD648CFC1E7245" box="[762,976,1277,1301]" pageId="56" pageNumber="397">Luzhang Township</collectingMunicipality>
,
<location id="8EA06050FFADFFB13FB7648CFBBB7245" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7:8EA06050FFADFFB13FB7648CFBBB7245" box="[992,1141,1277,1301]" country="China" county="Lushui County" latitude="25.97228" longLatPrecision="1" longitude="98.68336" municipality="Luzhang Township" name="Pianma Road" pageId="56" pageNumber="397" stateProvince="Yunnan">Pianma Road</location>
at
<location id="8EA06050FFADFFB13C2C656FFEEF7265" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7:8EA06050FFADFFB13C2C656FFEEF7265" box="[123,289,1309,1334]" country="China" county="Lushui County" latitude="25.97228" longLatPrecision="1" longitude="98.68336" municipality="Luzhang Township" name="Fengxue Yakou" pageId="56" pageNumber="397" stateProvince="Yunnan">Fengxue Yakou</location>
[Pass],
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/
<geoCoordinate id="EE4B504CFFADFFB13E54656CFD4D7265" box="[515,643,1309,1333]" degrees="98.68336" direction="east" orientation="longitude" pageId="56" pageNumber="397" precision="1" value="98.68336">E98.68336°</geoCoordinate>
,
<quantity id="4C879B6EFFADFFB13ED9656CFD2D7265" box="[654,739,1309,1333]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="56" pageNumber="397" unit="m" value="3150.0">
<elevation id="0052D1B8FFADFFB13ED9656CFD2D7265" box="[654,739,1309,1333]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="56" pageNumber="397" unit="m" value="3150.0">3150 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFADFFB13CFC654FFBF87227" pageId="56" pageNumber="397" type="etymology">
<paragraph id="8BC0368BFFADFFB13CFC654FFBF87227" blockId="56.[123,1174,1048,1725]" pageId="56" pageNumber="397">
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<emphasis id="B90BEA99FFADFFB13CFC654FFEFB7204" bold="true" box="[171,309,1342,1366]" pageId="56" pageNumber="397">ETYMOLOGY</emphasis>
</smallCapsWord>
.— The species epithet,
<taxonomicName id="4C7F4D08FFADFFB13E6D654FFD0A7206" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[570,708,1342,1366]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="397" phylum="Arthropoda" rank="species" species="luzhangensis">
<emphasis id="B90BEA99FFADFFB13E6D654FFD0A7206" box="[570,708,1342,1366]" italics="true" pageId="56" pageNumber="397">luzhangensis</emphasis>
</taxonomicName>
, is derived from the name of the township, Luzhang, in which the
<typeStatus id="54C48829FFADFFB13D25652EFE6E7227" box="[370,416,1375,1399]" pageId="56" pageNumber="397">type</typeStatus>
locality is found, and the Latin suffix,
<emphasis id="B90BEA99FFADFFB13F12652EFC4D7226" box="[837,899,1375,1398]" italics="true" pageId="56" pageNumber="397">-ensis</emphasis>
, denoting place.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFADFFB33CFC650EFEE07643" lastPageId="58" lastPageNumber="399" pageId="56" pageNumber="397" type="diagnosis">
<paragraph id="8BC0368BFFADFFB33CFC650EFEE07643" blockId="56.[123,1174,1048,1725]" lastBlockId="58.[123,1174,219,1725]" lastPageId="58" lastPageNumber="399" pageId="56" pageNumber="397">
<smallCapsWord id="8D26A057FFADFFB13CFC650EFEEE72C6" baselines="1425,1425" box="[171,288,1407,1431]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="56" pageNumber="397">
<emphasis id="B90BEA99FFADFFB13CFC650EFEEE72C6" bold="true" box="[171,288,1407,1431]" pageId="56" pageNumber="397">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFADFFB13E10650EFD6D72C7" box="[583,675,1407,1431]" captionStart="FIGURE" captionStartId="57.[171,183,1583,1602]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@57.[120,1176,216,1561]" captionTargetPageId="57" captionText="FIGURE 22. Trechus luzhangensis sp. nov.; a. Dorsal habitus (CASENT1017595). b. Median lobe of aedeagus of male (CASENT1017595), left lateral aspect. c. Map of locality records (red circles) for T. luzhangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155326" httpUri="https://zenodo.org/record/13155326/files/figure.png" pageId="56" pageNumber="397">Fig. 22a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
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), apterous; body color dark piceous, shiny, elytra with interval 1, lateral margins and apicomedial area reddish; tempora glabrous, convex, half as long as diameter of eyes; pronotum transverse (ratio PW/PL = 1.46), basal angles rectangular and sharp; elytra ovoid, convex, striae finely impressed, striae 1 to 3 deeply impressed on disc, not attenuated apically, remaining striae successively less distinct, the outermost very faint but still evident; recurrent stria terminated anteriorly at distinct convexity of interval 7; two discal setae present, inserted on interval 3 next to stria 3; preapical seta present and inserted in a discal position on interval 3, near stria
<quantity id="4C879B6EFFADFFB13F8666F4FC3271CD" box="[977,1020,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="56" pageNumber="397" unit="in" value="2.0">2 in</quantity>
forward position near apical one-fourth of elytra; median lobe of male aedeagus (
<figureCitation id="13442A0EFFADFFB13F0366D4FC6171ED" box="[852,943,1701,1725]" captionStart="FIGURE" captionStartId="57.[171,183,1583,1602]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@57.[120,1176,216,1561]" captionTargetPageId="57" captionText="FIGURE 22. Trechus luzhangensis sp. nov.; a. Dorsal habitus (CASENT1017595). b. Median lobe of aedeagus of male (CASENT1017595), left lateral aspect. c. Map of locality records (red circles) for T. luzhangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155326" httpUri="https://zenodo.org/record/13155326/files/figure.png" pageId="56" pageNumber="397">Fig. 22b</figureCitation>
) with moderate-sized sagittal aileron, apical lamella short and thick, with apex blunt, endophallus with a sclerotized line and a scaly area.
</paragraph>
</subSubSection>
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22.
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<emphasis id="B90BEA99FFACFFB03D5A665EFE0D7112" box="[269,451,1583,1602]" italics="true" pageId="57" pageNumber="398">Trechus luzhangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFACFFB03D9E665EFDC17112" box="[457,527,1583,1602]" pageId="57" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1017595). b. Median lobe of aedeagus of male (CASENT1017595), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFACFFB03F106638FC05710B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[839,971,1608,1627]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="398" phylum="Arthropoda" rank="species" species="luzhangensis">
<emphasis id="B90BEA99FFACFFB03F106638FC05710B" box="[839,971,1608,1627]" italics="true" pageId="57" pageNumber="398">T. luzhangensis</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<subSubSection id="C3656500FFAFFFB23CFC616DFF0C76E7" lastPageId="59" lastPageNumber="400" pageId="58" pageNumber="399" type="description">
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<emphasis id="B90BEA99FFAFFFB33CFC616DFEF47662" bold="true" box="[171,314,284,308]" pageId="58" pageNumber="399">DESCRIPTION</emphasis>
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.— Size medium, BL =
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. Body color dark piceous, shiny, elytra with interval 1, lateral margins and apicomedial area more or less reddish, appendages paler, yellowish-orange, palpi pale yellow. Body surface smooth, head capsule faintly alutaceous.
</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC610DFD767586" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">
Head. Moderate in size; eyes only slightly projected but convex, their diameter twice as long as tempora, the latter short, moderately convex and glabrous. Frons not flattened; frontal furrows deep, rounded, prolonged and not attenuated posterior to the eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Mandibles short; right mandible distinctly tridentate with middle tooth closer to basal tooth (premolar) than to distal tooth.
<collectingRegion id="49BBF869FFAFFFB33EA6626FFC857565" box="[753,843,542,565]" country="Myanmar" name="Chin" pageId="58" pageNumber="399">Mentum</collectingRegion>
and submentum not fused but nearly so, suture between them only faintly impressed.
<collectingRegion id="49BBF869FFAFFFB33E8F624FFCFC7505" box="[728,818,574,597]" country="Myanmar" name="Chin" pageId="58" pageNumber="399">Mentum</collectingRegion>
with medial tooth apically truncate, less than half the length of the lateral lobes. Submentum with six setae anteriorly, swollen anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae rather short, with about 2 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter.
</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC62AEFC5A7488" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">Pronotum. Transverse (ratio PW/PL = 1.46), moderately narrowed posteriorly, greatest width anterior to middle; lateral margins widely rounded, straightened only just anterior to basal angles, the latter very small, but projected, acute and sharp. Disc convex, smooth and glabrous, median longitudinal impression slender but sharply impressed, extended anteriorly to near anterior margin; basal foveae distinct, subcircular; median basal area faintly, longitudinally rugulose, delimited laterally by obliquely curved furrows. Lateral borders of pronotum moderately slender, narrowly reflexed, lateral grooves deeply impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC6391FC1873AA" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">
Elytra. Ovoid, only slightly more narrowed basally than apically, humeri distinct but rounded. Disc convex and smooth. Striae finely impressed, not or only faintly punctate; striae 1 to 3 deeply impressed on disc, not attenuated apically, remaining striae successively less distinct, the outermost very faint but still evident. Parascutellar striole present. Recurrent stria terminated anteriorly at distinct convexity of interval 7. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a prediscal position on interval 3, closer to stria 2 than
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forward position at apical one-fourth of elytra. Umbilicate setae of humeral group equidistance from each other and setae of median group inserted slightly posterior to middle.
</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC6572FE50726A" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">Legs. Relatively short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC6533FB85722B" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC65F2FCBF72AC" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">
Male aedeagus. Median lobe (
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) only moderately enlarged basally but with a mediumsized sagittal aileron present; shaft narrow subbasally, progressively thicker toward subapical portion; apical lamella short and thick, with apex blunt, endophallus with a sclerotized line and a scaly area, similar to that seen in males of
<taxonomicName id="4C7F4D08FFAFFFB33E5F6595FD5072AC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[520,670,1508,1532]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="399" phylum="Arthropoda" rank="species" species="shiyueliang" status="sp. nov.">
<emphasis id="B90BEA99FFAFFFB33E5F6595FD5072AC" box="[520,670,1508,1532]" italics="true" pageId="58" pageNumber="399">T. shiyueliang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFAFFFB33EF26595FD3672AB" box="[677,760,1508,1531]" pageId="58" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13442A0EFFAFFFB33F506595FCAB72AC" box="[775,869,1508,1532]" captionStart="FIGURE" captionStartId="47.[171,183,1569,1588]" captionTargetBox="[120,1132,216,1549]" captionTargetId="figure-14@47.[120,1132,216,1549]" captionTargetPageId="47" captionText="FIGURE 18. Trechus shiyueliang sp. nov.; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for T. shiyueliang in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155318" httpUri="https://zenodo.org/record/13155318/files/figure.png" pageId="58" pageNumber="399">Fig. 18b</figureCitation>
).
</paragraph>
<paragraph id="8BC0368BFFAFFFB33CFC6675FDF071ED" blockId="58.[123,1174,219,1725]" pageId="58" pageNumber="399">
<emphasis id="B90BEA99FFAFFFB33CFC6675FE68714B" bold="true" box="[171,422,1540,1564]" pageId="58" pageNumber="399">
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<smallCapsWord id="8D26A057FFAFFFB33D456679FE68714B" baselines="1558" box="[274,422,1544,1563]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="58" pageNumber="399">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found under stones on slopes above (
<figureCitation id="13442A0EFFAFFFB33C9E6655FEEC716D" box="[201,290,1572,1597]" captionStart="FIGURE" captionStartId="105.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1606]" captionTargetId="figure-15@105.[120,1176,216,1607]" captionTargetPageId="105" captionText="FIGURE 43. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Luzhang Township, Piana Road 0.1 km E of Fengxue Yakou, elevation 3150 m; habitat in which specimens of Trechus luzhangensis sp. nov. were collected. b. Lushui County, Luzhang Township, 100 m S of Fengxue Yakou on east side of pass, elevation 3150 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve) were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155372" httpUri="https://zenodo.org/record/13155372/files/figure.png" pageId="58" pageNumber="399">Fig. 43a</figureCitation>
) and below the road and both side of the pass at an elevation of
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. The habitat in this area includes broken scrub vegetation of two to four meter high bamboo and
<taxonomicName id="4C7F4D08FFAFFFB338776634FF62712D" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="58" pageNumber="399" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFAFFFB338776634FF62712D" italics="true" pageId="58" pageNumber="399">Rhododendron</emphasis>
</taxonomicName>
thickets on a thin layer of organic substrate, as well as open areas with stones on granitic sand substrate, and small seeps. Both specimens were found at the edges of thickets on organic substrate. No other trechines were found at this site.
</paragraph>
<paragraph id="8BC0368BFFAEFFB23CFC60AAFE9E7605" blockId="59.[123,1174,219,439]" pageId="59" pageNumber="400">
<emphasis id="B90BEA99FFAEFFB23CFC60AAFDB977A2" bold="true" box="[171,631,219,243]" pageId="59" pageNumber="400">
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<smallCapsWord id="8D26A057FFAEFFB23D0A60AFFE3F77A1" baselines="236" box="[349,497,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="59" pageNumber="400">DISTRIBUTION</smallCapsWord>
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</emphasis>
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<emphasis id="B90BEA99FFAEFFB23E2860AAFCCA77A1" bold="true" box="[639,772,219,243]" pageId="59" pageNumber="400">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFAEFFB23F5C60AAFB587644" location="We" pageId="59" pageNumber="400" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Shan">
<emphasis id="B90BEA99FFAEFFB23F5C60AAFC8077A3" bold="true" box="[779,846,219,243]" pageId="59" pageNumber="400">
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<collectingRegion id="49BBF869FFAEFFB23F5C60AAFC8877A1" box="[779,838,219,243]" country="Myanmar" name="Shan" pageId="59" pageNumber="400">SHAN</collectingRegion>
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.
</emphasis>
<figureCitation id="13442A0EFFAEFFB23F0360AAFC7E77A3" box="[852,944,219,243]" captionStart="FIGURE" captionStartId="57.[171,183,1583,1602]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@57.[120,1176,216,1561]" captionTargetPageId="57" captionText="FIGURE 22. Trechus luzhangensis sp. nov.; a. Dorsal habitus (CASENT1017595). b. Median lobe of aedeagus of male (CASENT1017595), left lateral aspect. c. Map of locality records (red circles) for T. luzhangensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155326" httpUri="https://zenodo.org/record/13155326/files/figure.png" pageId="59" pageNumber="400">Fig. 22c</figureCitation>
.
<location id="8EA06050FFAEFFB23FED60AAFC1377A2" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7:8EA06050FFAEFFB23FED60AAFC1377A2" box="[954,989,219,242]" name="We" pageId="59" pageNumber="400" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFAEFFB23CC0608DFED47644" box="[151,282,252,276]" count="2" pageId="59" pageNumber="400" type="generic">2 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFAEFFB23D7C608DFEBF7644" box="[299,369,252,276]" count="1" pageId="59" pageNumber="400" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFAEFFB23DF3608DFDCA7644" box="[420,516,252,276]" count="1" pageId="59" pageNumber="400" type="female">1 female</specimenCount>
), both from the crest of the southcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFAEFFB23C2C616DFE827605" county="Lushui County" location="Lushui County" pageId="59" pageNumber="400" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFAEFFB23C2C616DFF7E7664" box="[123,176,284,308]" country="Myanmar" name="Shan" pageId="59" pageNumber="400">Shan</collectingRegion>
in
<collectingCounty id="62A14E07FFAEFFB23C83616DFEBA7664" box="[212,372,284,308]" pageId="59" pageNumber="400">Lushui County</collectingCounty>
(see
<typeStatus id="54C48829FFAEFFB23DFA616CFE2C7665" box="[429,482,285,309]" pageId="59" pageNumber="400">Type</typeStatus>
material above for exact collection data). This locality straddles Core Areas 4 and 5
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFAEFFB23CFC612CFF0C76E7" blockId="59.[123,1174,219,439]" pageId="59" pageNumber="400">
<emphasis id="B90BEA99FFAEFFB23CFC612CFDAD7626" bold="true" box="[171,611,349,374]" pageId="59" pageNumber="400">
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.
</emphasis>
<materialsCitation id="3B173CD6FFAEFFB23E3D612FFF7076E7" country="China" location="Gaoligong Shan" pageId="59" pageNumber="400" specimenCount="1" stateProvince="Yunnan">
This species currently is known only from the
<typeStatus id="54C48829FFAEFFB23830612FFB5B7626" box="[1127,1173,350,374]" pageId="59" pageNumber="400">type</typeStatus>
locality on the crest of the southcentral part of the
<location id="8EA06050FFAEFFB23EF5610FFC9D76C6" LSID="urn:lsid:plazi:treatment:03D6879DFFADFFB23C2C63BBFF0C76E7:8EA06050FFAEFFB23EF5610FFC9D76C6" box="[674,851,382,406]" country="China" name="Gaoligong Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Gaoligong Shan</location>
, in western
<collectingRegion id="49BBF869FFAEFFB23F81610EFB5C76C6" box="[982,1170,382,406]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFAEFFB23C2C61EEFF7076E7" box="[123,190,415,439]" name="China" pageId="59" pageNumber="400">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,833 @@
<document id="FC15CD2775B62951CB38DFBDD2778FCE" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFAEFFB73C2C6191FBAD7384" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechus (Trechus) gongshanensis Deuve and Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="403" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="400" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="242DC1692C9C80D896A3BA256F813270">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="95E24BE55EB1CD9A053519DE6453EFFD">Deuve, Thierry</mods:namePart>
<mods:affiliation id="2F818A0B81929ECC57133CAF77CCA6D6">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="A091A6CD234824275A5E58B7FCE3DE08">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="E59BD6DCBC2B530818344E5A70C08DA3">Liang, Hongbin</mods:namePart>
<mods:affiliation id="333A0B1D0219F7635F372C088556C58B">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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Deuve and Liang
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(
<figureCitation id="13442A0EFFAEFFB23CD56270FF137547" box="[130,221,513,535]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="59" pageNumber="400">Figs. 16c</figureCitation>
,
<figureCitation id="13442A0EFFAEFFB23CBF6270FECC7546" box="[232,258,513,534]" captionStart="FIGURE" captionStartId="60.[171,183,1578,1597]" captionTargetBox="[132,1176,216,1563]" captionTargetId="figure-12@60.[120,1176,216,1563]" captionTargetPageId="60" captionText="FIGURE 23. Trechus gongshanensis sp. nov.; a. Dorsal habitus (CASENT1001929). b. Median lobe of aedeagus of male (CASENT1001929, left lateral aspect. c. Map of locality record s(red circles) for T. gongshanensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155328" httpUri="https://zenodo.org/record/13155328/files/figure.png" pageId="59" pageNumber="400">23</figureCitation>
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<figureCitation id="13442A0EFFAEFFB23D5B6270FEFF7546" box="[268,305,513,534]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="59" pageNumber="400">38a</figureCitation>
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<figureCitation id="13442A0EFFAEFFB23D6B6270FE5F7547" box="[316,401,513,535]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="59" pageNumber="400">39a, 39b</figureCitation>
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<figureCitation id="13442A0EFFAEFFB23DCB6270FE0F7546" box="[412,449,513,534]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="59" pageNumber="400">40a</figureCitation>
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<figureCitation id="13442A0EFFAEFFB23D9C6270FDC67547" box="[459,520,513,535]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="59" pageNumber="400">4648</figureCitation>
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<typeStatus id="54C48829FFAEFFB23C9062A3FEE175BA" box="[199,303,722,746]" pageId="59" pageNumber="400" type="paratype">Paratypes</typeStatus>
(a total of 51):
<specimenCount id="9D79FD02FFAEFFB23D8962A0FDE675B9" box="[478,552,721,745]" count="1" pageId="59" pageNumber="400" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFAEFFB23E0862A0FD0675B9" box="[607,712,721,745]" count="4" pageId="59" pageNumber="400" type="female">4 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23EA162A0FCE275B9" box="[758,812,721,745]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAEFFB23F6E62A0FCA975B9" box="[825,871,721,745]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFAEFFB2386562A0FB5C75B9" box="[1074,1170,721,745]" pageId="59" pageNumber="400" type="holotype">holotype</typeStatus>
, except first label “CASENT 1001928” and “CASENT 1001930” to “CASENT 1001932” and “CASENT 1008148”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB23E5F6362FDC374FD" collectingDate="2002-10-01" collectingDateMax="2002-10-02" collectingDateMin="2002-10-01" collectionCode="CAS, IOZ" collectorName="P. E. Marek" country="China" county="Gongshan County" elevation="3370" latitude="27.77032" location="Cikai Township" longLatPrecision="1" longitude="98.44661" pageId="59" pageNumber="400" specimenCount="12" specimenCount-female="4" specimenCount-male="8" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23E5F6362FD91747A" box="[520,607,786,811]" count="8" pageId="59" pageNumber="400" type="male">8 males</specimenCount>
and
<specimenCount id="9D79FD02FFAEFFB23ECB6362FCC6747A" box="[668,776,786,811]" count="4" pageId="59" pageNumber="400" type="female">4 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23F6B6362FCBC747B" box="[828,882,787,811]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAEFFB23FD56362FC7E747B" box="[898,944,787,811]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: “CASENT 1010348” to “CASENT 1010355” and “CASENT 1010356” to “CASENT 1010359”, respectively/ “
<collectingCountry id="F368761BFFAEFFB23CF06325FECE743C" box="[167,256,852,876]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23D5E6325FE93743B" box="[265,349,852,875]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23D306325FDE4743C" box="[359,554,852,876]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<location id="8EA06050FFAEFFB23E646325FD11743C" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23E646325FD11743C" box="[563,735,852,876]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44661" name="Cikai Township" pageId="59" pageNumber="400" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFAEFFB23EBF6325FF2C74DC" location="Gongshan on Dulong Valley Road" pageId="59" pageNumber="400">
<quantity id="4C879B6EFFAEFFB23EBF6325FCF1743C" box="[744,831,852,876]" metricMagnitude="4" metricUnit="m" metricValue="5.26" pageId="59" pageNumber="400" unit="km" value="52.6">52.6 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFFAEFFB23CA46305FE4774DD" box="[243,393,884,909]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="59" pageNumber="400" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FFAEFFB23CA46305FE4774DD" box="[243,393,884,909]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="59" pageNumber="400" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFAEFFB23DE16304FDF474DD" box="[438,570,885,909]" degrees="27.77032" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23E196305FD1E74DC" box="[590,720,884,908]" degrees="98.44661" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.44661">E98.44661°</geoCoordinate>
,
<quantity id="4C879B6EFFAEFFB23EB76305FCB974DC" box="[736,887,884,908]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="59" pageNumber="400" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FFAEFFB23EB76305FCB974DC" box="[736,887,884,908]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="59" pageNumber="400" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,
<date id="FFC1104BFFAEFFB23FD06304FB9F74DD" box="[903,1105,884,909]" pageId="59" pageNumber="400" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">
<collectingDate id="EF85E9A3FFAEFFB23FD06304FB9F74DD" box="[903,1105,884,909]" pageId="59" pageNumber="400" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">1-2 October 2002</collectingDate>
</date>
, D.H.
<taxonomicName id="4C7F4D08FFAEFFB23C2C63E4FF3A74FD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,244,917,941]" pageId="59" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFAEFFB23D4F63E7FE4174FD" box="[280,399,917,941]" pageId="59" pageNumber="400">P.E. Marek</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB23E4E63E4FDB9735F" collectingDate="2006-08-20" collectionCode="IOZ" collectorName="Y. Liu &amp; P. Hu &amp; D. Z. Dong &amp; J. Wang" country="China" county="Gongshan County" elevation="3950" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="59" pageNumber="400" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23E4E63E4FDB874FD" box="[537,630,917,941]" count="1" pageId="59" pageNumber="400" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23EF463E4FD1F74FD" box="[675,721,917,941]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: “CASENT 1024869”/ “
<collectingCountry id="F368761BFFAEFFB2386F63E4FB5F74FD" box="[1080,1169,917,941]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23C2C63C7FF01749D" box="[123,207,950,973]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23C8D63C7FE50749E" box="[218,414,950,974]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAEFFB23DFE63C7FD7D749E" box="[425,691,950,974]" pageId="59" pageNumber="400">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFAEFFB23F1D63C7FBCA749E" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23F1D63C7FBCA749E" box="[842,1028,950,974]" country="China" county="Gongshan County" latitude="27.98206" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Kawakarpu Shan</location>
, on slope
<locationDeviation id="9A722FA9FFAEFFB2382563C7FEFC74BE" location="Chukuai Lake" pageId="59" pageNumber="400">NE of Chukuai Lake</locationDeviation>
,
<quantity id="4C879B6EFFAEFFB23D6A63A7FE5F74BF" box="[317,401,982,1007]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="59" pageNumber="400" unit="m" value="3950.0">
<elevation id="0052D1B8FFAEFFB23D6A63A7FE5F74BF" box="[317,401,982,1007]" metricMagnitude="3" metricUnit="m" metricValue="3.95" pageId="59" pageNumber="400" unit="m" value="3950.0">3950 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFAEFFB23DEE63A7FDF274BE" box="[441,572,982,1006]" degrees="27.98206" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.98206">N27.98206°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23E1B63A6FD0374BF" box="[588,717,983,1007]" degrees="98.48027" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.48027">E98.48027°</geoCoordinate>
,
<date id="FFC1104BFFAEFFB23E8E63A6FC4D74BE" box="[729,899,982,1007]" pageId="59" pageNumber="400" value="2006-08-20">
<collectingDate id="EF85E9A3FFAEFFB23E8E63A6FC4D74BE" box="[729,899,982,1007]" pageId="59" pageNumber="400" value="2006-08-20">20 August 2006</collectingDate>
</date>
Stop #DHK-2006-086
<collectorName id="268A535DFFAEFFB2382863A6FF6D735F" pageId="59" pageNumber="400">Y. Liu</collectorName>
,
<collectorName id="268A535DFFAEFFB23CE76389FF3E735F" box="[176,240,1016,1039]" pageId="59" pageNumber="400">P. Hu</collectorName>
,
<collectorName id="268A535DFFAEFFB23CA96389FEBA7340" box="[254,372,1016,1040]" pageId="59" pageNumber="400">D.Z. Dong</collectorName>
&amp;
<collectorName id="268A535DFFAEFFB23DC96389FE387340" box="[414,502,1016,1040]" pageId="59" pageNumber="400">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB23ED26386FD4473A4" collectingDate="2006-08-21" collectionCode="CAS, IOZ" collectorName="D. Z. Dong &amp; Y. Liu &amp; P. Hu &amp; J. Wang" country="China" county="Gongshan County" elevation="3808" latitude="27.98231" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" pageId="59" pageNumber="400" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23ED26386FD1E735F" box="[645,720,1015,1039]" count="1" pageId="59" pageNumber="400" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23F556386FCF9735F" box="[770,823,1015,1039]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1025105”/ “
<collectingCountry id="F368761BFFAEFFB23CD36469FF137360" box="[132,221,1048,1072]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23CB06469FEF5737F" box="[231,315,1048,1071]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23D116469FDC77360" box="[326,521,1048,1072]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAEFFB23E446469FCD27360" box="[531,796,1048,1072]" pageId="59" pageNumber="400">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFAEFFB23FE76469FBA77360" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23FE76469FBA77360" box="[944,1129,1048,1072]" country="China" county="Gongshan County" latitude="27.98231" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFAEFFB238236469FE557300" location="Chukuai Lake" pageId="59" pageNumber="400">
<quantity id="4C879B6EFFAEFFB238236469FF537300" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="59" pageNumber="400" unit="km" value="0.4">0.4 km</quantity>
NW of Chukuai Lake
</locationDeviation>
,”/ “
<geoCoordinate id="EE4B504CFFAEFFB23D926448FD867301" box="[453,584,1081,1105]" degrees="27.98231" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.98231">N27.98231°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23E0C6449FD127300" box="[603,732,1080,1104]" degrees="98.47069" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.47069">E98.47069°</geoCoordinate>
,
<quantity id="4C879B6EFFAEFFB23EBC6448FC8F7300" box="[747,833,1081,1105]" metricMagnitude="3" metricUnit="m" metricValue="3.808" pageId="59" pageNumber="400" unit="m" value="3808.0">
<elevation id="0052D1B8FFAEFFB23EBC6448FC8F7300" box="[747,833,1081,1105]" metricMagnitude="3" metricUnit="m" metricValue="3.808" pageId="59" pageNumber="400" unit="m" value="3808.0">3808 m</elevation>
</quantity>
,
<date id="FFC1104BFFAEFFB23F076448FBCE7300" box="[848,1024,1080,1105]" pageId="59" pageNumber="400" value="2006-08-21">
<collectingDate id="EF85E9A3FFAEFFB23F076448FBCE7300" box="[848,1024,1080,1105]" pageId="59" pageNumber="400" value="2006-08-21">21 August 2006</collectingDate>
</date>
Stop #DHK-2006-094
<collectorName id="268A535DFFAEFFB23CB9642BFEA67322" box="[238,360,1114,1138]" pageId="59" pageNumber="400">D.Z. Dong</collectorName>
collector”/
<specimenCount id="9D79FD02FFAEFFB23DA36428FD837321" box="[500,589,1113,1137]" count="2" pageId="59" pageNumber="400" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FFAEFFB23ED86428FD337321" box="[655,765,1113,1137]" count="2" pageId="59" pageNumber="400" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23F626428FCA57321" box="[821,875,1113,1137]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAEFFB23F2A6428FC657321" box="[893,939,1113,1137]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: “CASENT 1025164” to “CASENT 1025165” and “CASENT 1025162” to “CASENT 1025163”, respectively/ “
<collectingCountry id="F368761BFFAEFFB23CF064EAFECE73E3" box="[167,256,1179,1203]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23D5C64EAFE9173E2" box="[267,351,1179,1202]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<location id="8EA06050FFAEFFB23D3D64EBFDE373E3" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23D3D64EBFDE373E3" box="[362,557,1178,1203]" country="China" county="Gongshan County" latitude="27.98231" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" name="Gongshan County" pageId="59" pageNumber="400" stateProvince="Yunnan">Gongshan County</location>
,
<location id="8EA06050FFAEFFB23E6F64EBFC8C73E2" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23E6F64EBFC8C73E2" box="[568,834,1178,1202]" country="China" county="Gongshan County" latitude="27.98231" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" name="Bingzhongluo Township" pageId="59" pageNumber="400" stateProvince="Yunnan">Bingzhongluo Township</location>
, SW slope of Kawakarpu Shan,
<locationDeviation id="9A722FA9FFAEFFB23C2C64CAFE0E7383" box="[123,448,1211,1235]" location="Chukuai Lake" pageId="59" pageNumber="400">
<quantity id="4C879B6EFFAEFFB23C2C64CAFF1F7383" box="[123,209,1211,1235]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="59" pageNumber="400" unit="km" value="0.75">0.75 km</quantity>
NW of Chukuai Lake
</locationDeviation>
,”/ “
<geoCoordinate id="EE4B504CFFAEFFB23DB164CAFDA77383" box="[486,617,1211,1235]" degrees="27.98631" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.98631">N27.98631°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23E2064CAFD377383" box="[631,761,1211,1235]" degrees="98.47069" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.47069">E98.47069°</geoCoordinate>
,
<quantity id="4C879B6EFFAEFFB23F5464CAFC987383" box="[771,854,1211,1235]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="59" pageNumber="400" unit="m" value="3820.0">
<elevation id="0052D1B8FFAEFFB23F5464CAFC987383" box="[771,854,1211,1235]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="59" pageNumber="400" unit="m" value="3820.0">3820 m</elevation>
</quantity>
,
<date id="FFC1104BFFAEFFB23F3764CAFBC67383" box="[864,1032,1211,1235]" pageId="59" pageNumber="400" value="2006-08-21">
<collectingDate id="EF85E9A3FFAEFFB23F3764CAFBC67383" box="[864,1032,1211,1235]" pageId="59" pageNumber="400" value="2006-08-21">21 August 2006</collectingDate>
</date>
Stop #DHK-2006-095
<collectorName id="268A535DFFAEFFB23CBD64ADFEFD73A4" box="[234,307,1244,1268]" pageId="59" pageNumber="400">Y. Liu</collectorName>
,
<collectorName id="268A535DFFAEFFB23D1664ADFE4F73A3" box="[321,385,1244,1267]" pageId="59" pageNumber="400">P. Hu</collectorName>
, &amp;
<collectorName id="268A535DFFAEFFB23DF864ADFDC673A4" box="[431,520,1244,1268]" pageId="59" pageNumber="400">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB23ECF64ADFE6D7226" collectingDate="2006-08-21" collectionCode="CAS, IOZ" collectorName="Y. Liu" country="China" county="Gongshan County" elevation="3820" latitude="27.98631" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" pageId="59" pageNumber="400" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23ECF64ADFD2A73A4" box="[664,740,1244,1268]" count="1" pageId="59" pageNumber="400" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFAEFFB23F4864ADFC4473A4" box="[799,906,1244,1268]" count="2" pageId="59" pageNumber="400" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23FEB64ADFC3C73A4" box="[956,1010,1244,1268]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAEFFB2385664ADFBE173A4" box="[1025,1071,1244,1268]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: “CASENT 1025833” and “CASENT 1025834” to “CASENT 1025835”, respectively/ “
<collectingCountry id="F368761BFFAEFFB2386F648CFB5F7245" box="[1080,1169,1277,1301]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23C2C656FFF017265" box="[123,207,1310,1333]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23C8C656CFE6E7265" box="[219,416,1309,1333]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAEFFB23DFB656CFD797265" box="[428,695,1309,1333]" pageId="59" pageNumber="400">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFAEFFB23F05656CFBC37265" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23F05656CFBC37265" box="[850,1037,1309,1333]" country="China" county="Gongshan County" latitude="27.98631" longLatPrecision="1" longitude="98.47069" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFAEFFB2384D656CFEE17206" location="Chukuai Lake" pageId="59" pageNumber="400">
<quantity id="4C879B6EFFAEFFB2384D656CFBAB7265" box="[1050,1125,1309,1333]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="59" pageNumber="400" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/ “
<geoCoordinate id="EE4B504CFFAEFFB23D8C654FFD907206" box="[475,606,1342,1366]" degrees="27.98631" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.98631">N27.98631°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23E39654FFD3E7206" box="[622,752,1342,1366]" degrees="98.47069" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.47069">E98.47069°</geoCoordinate>
,
<quantity id="4C879B6EFFAEFFB23EAC654FFC817205" box="[763,847,1342,1366]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="59" pageNumber="400" unit="m" value="3820.0">
<elevation id="0052D1B8FFAEFFB23EAC654FFC817205" box="[763,847,1342,1366]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="59" pageNumber="400" unit="m" value="3820.0">3820 m</elevation>
</quantity>
,
<date id="FFC1104BFFAEFFB23F0C654FFBCB7206" box="[859,1029,1342,1366]" pageId="59" pageNumber="400" value="2006-08-21">
<collectingDate id="EF85E9A3FFAEFFB23F0C654FFBCB7206" box="[859,1029,1342,1366]" pageId="59" pageNumber="400" value="2006-08-21">21 August 2006</collectingDate>
</date>
Stop #DHK-2006-082
<collectorName id="268A535DFFAEFFB23CBE652EFEE37226" box="[233,301,1374,1398]" pageId="59" pageNumber="400">Y. Liu</collectorName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB23DF8652EFD2A7288" collectingDate="2006-08-19" collectionCode="IOZ" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu &amp; P. Hu &amp; J. Wang" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="59" pageNumber="400" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23DF8652EFDC17226" box="[431,527,1374,1399]" count="1" pageId="59" pageNumber="400" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23E69652EFDA27227" box="[574,620,1375,1399]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: “CASENT 1025920”/ “
<collectingCountry id="F368761BFFAEFFB23F8F652EFBFF7227" box="[984,1073,1375,1399]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB2386A652EFB5F7226" box="[1085,1169,1375,1398]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23C2C650EFEF372C7" box="[123,317,1407,1431]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAEFFB23D1F650EFD9F72C7" box="[328,593,1407,1431]" pageId="59" pageNumber="400">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFAEFFB23EB2650EFC5572C7" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23EB2650EFC5572C7" box="[741,923,1407,1431]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Kawakarpu Shan</location>
at
<location id="8EA06050FFAEFFB23FEB650EFB9B72C7" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23FEB650EFB9B72C7" box="[956,1109,1407,1431]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="59" pageNumber="400" stateProvince="Yunnan">Chukuai Lake</location>
,
<quantity id="4C879B6EFFAEFFB23837650EFF5C72E8" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="59" pageNumber="400" unit="m" value="3720.0">
<elevation id="0052D1B8FFAEFFB23837650EFF5C72E8" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="59" pageNumber="400" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFAEFFB23CEB65D1FEF172E8" box="[188,319,1440,1464]" degrees="27.98121" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23D0665D1FE1D72E8" box="[337,467,1440,1464]" degrees="98.47580" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.4758">E98.47580°</geoCoordinate>
,
<date id="FFC1104BFFAEFFB23DB765D1FD4072E8" box="[480,654,1440,1464]" pageId="59" pageNumber="400" value="2006-08-19">
<collectingDate id="EF85E9A3FFAEFFB23DB765D1FD4072E8" box="[480,654,1440,1464]" pageId="59" pageNumber="400" value="2006-08-19">19 August 2006</collectingDate>
</date>
Stop #DHK-2006-095B D.H.
<taxonomicName id="4C7F4D08FFAEFFB23FB665D1FB9372E8" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[993,1117,1440,1464]" pageId="59" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAEFFB2383D65D1FF717288" pageId="59" pageNumber="400">J.A. Miller</collectorName>
,
<collectorName id="268A535DFFAEFFB23C9B65B0FE8A7289" box="[204,324,1473,1497]" pageId="59" pageNumber="400">D.Z. Dong</collectorName>
,
<collectorName id="268A535DFFAEFFB23D0765B0FE597288" box="[336,407,1472,1496]" pageId="59" pageNumber="400">Y. Liu</collectorName>
,
<collectorName id="268A535DFFAEFFB23DF365B0FE2C7288" box="[420,482,1473,1496]" pageId="59" pageNumber="400">P. Hu</collectorName>
, &amp;
<collectorName id="268A535DFFAEFFB23E5A65B0FDAA7289" box="[525,612,1473,1497]" pageId="59" pageNumber="400">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB23EA665B0FEBD710B" collectingDate="2006-08-19" collectionCode="CAS" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu &amp; P. Hu &amp; J. Wang" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="59" pageNumber="400" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23EA665B0FC9E7288" box="[753,848,1472,1497]" count="1" pageId="59" pageNumber="400" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23F2865B0FC7A7289" box="[895,948,1473,1497]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1025939”/ “
<collectingCountry id="F368761BFFAEFFB23D5E6590FEAB72A9" box="[265,357,1505,1529]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23D2E6593FE1E72A9" box="[377,464,1506,1529]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23DB26590FD7972A9" box="[485,695,1505,1529]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAEFFB23E9C6590FC2872A9" box="[715,998,1505,1529]" pageId="59" pageNumber="400">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFAEFFB23C2C6673FEFB714A" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23C2C6673FEFB714A" box="[123,309,1538,1562]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFAEFFB23D176673FDBD714A" box="[320,627,1538,1562]" location="Chukuai Lake" pageId="59" pageNumber="400">
<quantity id="4C879B6EFFAEFFB23D176673FE44714A" box="[320,394,1538,1562]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="59" pageNumber="400" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,
<geoCoordinate id="EE4B504CFFAEFFB23F556673FC4D714A" box="[770,899,1538,1562]" degrees="27.97686" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23FC56673FBDA714A" box="[914,1044,1538,1562]" degrees="98.47799" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.47799">E98.47799°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFAEFFB2386C6673FB5F7149" box="[1083,1169,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="59" pageNumber="400" unit="m" value="3750.0">
<elevation id="0052D1B8FFAEFFB2386C6673FB5F7149" box="[1083,1169,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="59" pageNumber="400" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFFAEFFB23C2C6652FEE8716A" box="[123,294,1570,1595]" pageId="59" pageNumber="400" value="2006-08-19">
<collectingDate id="EF85E9A3FFAEFFB23C2C6652FEE8716A" box="[123,294,1570,1595]" pageId="59" pageNumber="400" value="2006-08-19">19 August 2006</collectingDate>
</date>
Stop #DHK-2006-095C D.H.
<taxonomicName id="4C7F4D08FFAEFFB23E246653FD21716A" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[627,751,1570,1594]" pageId="59" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFAEFFB23EAB6652FCBD716A" box="[764,883,1570,1595]" pageId="59" pageNumber="400">J.A. Miller</collectorName>
,
<collectorName id="268A535DFFAEFFB23F286652FC38716B" box="[895,1014,1571,1595]" pageId="59" pageNumber="400">D.Z. Dong</collectorName>
,
<collectorName id="268A535DFFAEFFB238566652FB86716A" box="[1025,1096,1570,1594]" pageId="59" pageNumber="400">Y. Liu</collectorName>
,
<collectorName id="268A535DFFAEFFB238046652FB5F716A" box="[1107,1169,1571,1594]" pageId="59" pageNumber="400">P. Hu</collectorName>
, &amp;
<collectorName id="268A535DFFAEFFB23CCE6635FF3C710C" box="[153,242,1604,1628]" pageId="59" pageNumber="400">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFAEFFB43DD66632FD0877A3" collectingDate="2006-08-19" collectionCode="CAS, IOZ" collectorName="J. A. Miller &amp; D. Z. Dong" country="China" county="Gongshan County" elevation="3745" lastPageId="61" lastPageNumber="402" latitude="27.98393" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" pageId="59" pageNumber="400" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFAEFFB23DD66632FE18710B" box="[385,470,1603,1627]" count="4" pageId="59" pageNumber="400" type="male">4 males</specimenCount>
and
<specimenCount id="9D79FD02FFAEFFB23E466632FDB2710B" box="[529,636,1603,1627]" count="2" pageId="59" pageNumber="400" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFAEFFB23EFA6632FD2D710B" box="[685,739,1603,1627]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="400" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFAEFFB23EA56632FCEE710B" box="[754,800,1603,1627]" pageId="59" pageNumber="400">IOZ</collectionCode>
) labeled: “CASENT 1026196” to “CASENT 1026199” and “CASENT 1026200” to “CASENT 1026201”, respectively/ / “
<collectingCountry id="F368761BFFAEFFB2386F6615FB5F712C" box="[1080,1169,1636,1660]" name="China" pageId="59" pageNumber="400">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFAEFFB23C2C66F4FF0171CC" box="[123,207,1669,1692]" country="China" name="Yunnan" pageId="59" pageNumber="400">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFAEFFB23C8D66F5FE5371CD" box="[218,413,1668,1693]" pageId="59" pageNumber="400">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFAEFFB23DFF66F5FD7C71CC" box="[424,690,1668,1692]" pageId="59" pageNumber="400">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFAEFFB23F1F66F5FBCC71CC" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFAEFFB23F1F66F5FBCC71CC" box="[840,1026,1668,1692]" country="China" county="Gongshan County" latitude="27.98393" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="59" pageNumber="400" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFAEFFB2385A66F4FEF971ED" location="Chukuai Lake" pageId="59" pageNumber="400">
<quantity id="4C879B6EFFAEFFB2385A66F4FB9971CC" box="[1037,1111,1668,1693]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="59" pageNumber="400" unit="km" value="0.3">0.3 km</quantity>
NNE of Chukuai Lake
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFAEFFB23D1266D4FE0871ED" box="[325,454,1701,1725]" degrees="27.98393" direction="north" orientation="latitude" pageId="59" pageNumber="400" precision="1" value="27.98393">N27.98393°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFAEFFB23D8F66D4FD9471ED" box="[472,602,1701,1725]" degrees="98.47491" direction="east" orientation="longitude" pageId="59" pageNumber="400" precision="1" value="98.47491">E98.47491°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFAEFFB23ED266D4FD1371ED" box="[645,733,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="59" pageNumber="400" unit="m" value="3745.0">
<elevation id="0052D1B8FFAEFFB23ED266D4FD1371ED" box="[645,733,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="59" pageNumber="400" unit="m" value="3745.0">3745 m</elevation>
</quantity>
,
<date id="FFC1104BFFAEFFB23EBC66D4FC5471ED" box="[747,922,1701,1725]" pageId="59" pageNumber="400" value="2006-08-19">
<collectingDate id="EF85E9A3FFAEFFB23EBC66D4FC5471ED" box="[747,922,1701,1725]" pageId="59" pageNumber="400" value="2006-08-19">19 August 2006</collectingDate>
</date>
Stop #DHK-2006-081 D.H.
<taxonomicName id="4C7F4D08FFA8FFB43CE260AAFEFF77A3" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[181,305,219,243]" pageId="61" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFA8FFB43D6C60AAFE7E77A3" box="[315,432,219,243]" pageId="61" pageNumber="402">J.A. Miller</collectorName>
, &amp;
<collectorName id="268A535DFFA8FFB43D8160AAFD8677A3" box="[470,584,219,243]" pageId="61" pageNumber="402">D.Z. Dong</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA8FFB43E8760AAFBD27604" collectingDate="2006-08-13" collectionCode="CAS, IOZ" collectorName="J. A. Miller" country="China" county="Gongshan County" elevation="3720" latitude="27.76978" location="Cikai Township" longLatPrecision="1" longitude="98.44681" pageId="61" pageNumber="402" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA8FFB43E8760AAFCEF77A3" box="[720,801,219,243]" count="3" pageId="61" pageNumber="402" type="male">3 males</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA8FFB43F1C60AAFC4F77A3" box="[843,897,219,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="402" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFA8FFB43FDB60AAFC7477A3" box="[908,954,219,243]" pageId="61" pageNumber="402">IOZ</collectionCode>
) labeled: “CASENT 1026320” to “CASENT 1026322”, respectively/ “
<collectingCountry id="F368761BFFA8FFB43EE6608AFCC47643" box="[689,778,251,275]" name="China" pageId="61" pageNumber="402">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA8FFB43F4C608DFCA17643" box="[795,879,252,275]" country="China" name="Yunnan" pageId="61" pageNumber="402">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA8FFB43FD6608AFB857643" box="[897,1099,251,275]" pageId="61" pageNumber="402">Gongshan County</collectingCounty>
,
<location id="8EA06050FFA8FFB4380B608AFF297663" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFA8FFB4380B608AFF297663" country="China" county="Gongshan County" latitude="27.76978" longLatPrecision="1" longitude="98.44681" name="Cikai Township" pageId="61" pageNumber="402" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFA8FFB43CA5616DFDD67663" box="[242,536,283,308]" location="Heipu Yakou" pageId="61" pageNumber="402">
<quantity id="4C879B6EFFA8FFB43CA5616DFEF07663" box="[242,318,283,308]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="61" pageNumber="402" unit="km" value="0.1">0.1 km</quantity>
SE of Heipu Yakou
</locationDeviation>
in valley below tunnel,
<geoCoordinate id="EE4B504CFFA8FFB43F75616AFC6D7663" box="[802,931,283,307]" degrees="27.76978" direction="north" orientation="latitude" pageId="61" pageNumber="402" precision="1" value="27.76978">N27.76978°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA8FFB43FE4616AFBFB7663" box="[947,1077,283,307]" degrees="98.44681" direction="east" orientation="longitude" pageId="61" pageNumber="402" precision="1" value="98.44681">E98.44681°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFA8FFB43809616DFF5C7603" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="61" pageNumber="402" unit="m" value="3720.0">
<elevation id="0052D1B8FFA8FFB43809616DFF5C7603" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="61" pageNumber="402" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,
<date id="FFC1104BFFA8FFB43CC9614DFE827604" box="[158,332,316,340]" pageId="61" pageNumber="402" value="2006-08-13">
<collectingDate id="EF85E9A3FFA8FFB43CC9614DFE827604" box="[158,332,316,340]" pageId="61" pageNumber="402" value="2006-08-13">13 August 2006</collectingDate>
</date>
, Stop #DHK-2006-073 D.H.
<taxonomicName id="4C7F4D08FFA8FFB43EDD614DFCCD7604" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[650,771,316,340]" pageId="61" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFA8FFB43F70614DFC537604" box="[807,925,316,340]" pageId="61" pageNumber="402">J.A. Miller</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA8FFB43870614DFC2776E4" collectingDate="2006-08-19" collectionCode="CAS" collectorName="J. A. Miller" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="61" pageNumber="402" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA8FFB43870614DFBBE7604" box="[1063,1136,316,340]" count="1" pageId="61" pageNumber="402" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA8FFB43C2C612DFF7E7624" box="[123,176,348,372]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="402" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1026366”/ “
<collectingCountry id="F368761BFFA8FFB43E4C612DFDBA7624" box="[539,628,348,372]" name="China" pageId="61" pageNumber="402">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA8FFB43ED7612DFD1A7623" box="[640,724,348,371]" country="China" name="Yunnan" pageId="61" pageNumber="402">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA8FFB43EB7612DFC6B7624" box="[736,933,348,372]" pageId="61" pageNumber="402">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA8FFB43FE6612DFF6476C4" pageId="61" pageNumber="402">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFA8FFB43D1C610DFDCA76C4" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFA8FFB43D1C610DFDCA76C4" box="[331,516,380,404]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="61" pageNumber="402" stateProvince="Yunnan">Kawakarpu Shan</location>
at
<location id="8EA06050FFA8FFB43E7C610DFD0976C4" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFA8FFB43E7C610DFD0976C4" box="[555,711,380,404]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="61" pageNumber="402" stateProvince="Yunnan">Chukuai Lake</location>
,
<quantity id="4C879B6EFFA8FFB43E82610DFCE576C4" box="[725,811,380,404]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="61" pageNumber="402" unit="m" value="3720.0">
<elevation id="0052D1B8FFA8FFB43E82610DFCE576C4" box="[725,811,380,404]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="61" pageNumber="402" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFA8FFB43F01610DFC1776C4" box="[854,985,380,404]" degrees="27.98121" direction="north" orientation="latitude" pageId="61" pageNumber="402" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA8FFB43FBC610DFBA376C4" box="[1003,1133,380,404]" degrees="98.47580" direction="east" orientation="longitude" pageId="61" pageNumber="402" precision="1" value="98.4758">E98.47580°</geoCoordinate>
,
<date id="FFC1104BFFA8FFB4382C610DFEC976E4" pageId="61" pageNumber="402" value="2006-08-19">
<collectingDate id="EF85E9A3FFA8FFB4382C610DFEC976E4" pageId="61" pageNumber="402" value="2006-08-19">19 August 2006</collectingDate>
</date>
Stop #DHK-2006-080 D.H.
<taxonomicName id="4C7F4D08FFA8FFB43E1B61EDFD0B76E4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[588,709,412,436]" pageId="61" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFA8FFB43EB861EDFCA976E4" box="[751,871,412,436]" pageId="61" pageNumber="402">J.A. Miller</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFA8FFB43FA061EDFC467526" collectingDate="2006-08-18" collectionCode="CAS, IOZ, MNHN" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="61" pageNumber="402" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFA8FFB43FA061EDFB8276E4" box="[1015,1100,412,436]" count="8" pageId="61" pageNumber="402" type="male">8 males</specimenCount>
and
<specimenCount id="9D79FD02FFA8FFB438DF61EDFF007684" count="5" pageId="61" pageNumber="402" type="female">5 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFA8FFB43D5561CCFEF67685" box="[258,312,445,469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="402" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFA8FFB43D1F61CCFEB97685" box="[328,375,445,469]" pageId="61" pageNumber="402">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFA8FFB43DD061CCFE147685" box="[391,474,445,469]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="61" pageNumber="402">MNHN</collectionCode>
) labeled: “CASENT 10263824” to “CASENT 10263831” and “CASENT 10263819” to “CASENT 10263823”, respectively/ “
<collectingCountry id="F368761BFFA8FFB43F7761ACFCB776A5" box="[800,889,477,501]" name="China" pageId="61" pageNumber="402">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFA8FFB43FD561ACFC1876A4" box="[898,982,477,500]" country="China" name="Yunnan" pageId="61" pageNumber="402">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA8FFB43FB761ACFF5F7545" pageId="61" pageNumber="402">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFA8FFB43CF2618CFE767545" box="[165,440,509,533]" pageId="61" pageNumber="402">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFA8FFB43E23618CFCFA7545" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFA8FFB43E23618CFCFA7545" box="[628,820,509,533]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="61" pageNumber="402" stateProvince="Yunnan">Kawakarpu Shan</location>
at
<location id="8EA06050FFA8FFB43F3E618CFBC27545" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFA8FFB43F3E618CFBC27545" box="[873,1036,509,533]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="61" pageNumber="402" stateProvince="Yunnan">Chukuai Lake</location>
,
<quantity id="4C879B6EFFA8FFB43876618CFBB07545" box="[1057,1150,509,533]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="61" pageNumber="402" unit="m" value="3720.0">
<elevation id="0052D1B8FFA8FFB43876618CFBB07545" box="[1057,1150,509,533]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="61" pageNumber="402" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFA8FFB43CD3626CFEC97565" box="[132,263,541,565]" degrees="27.98121" direction="north" orientation="latitude" pageId="61" pageNumber="402" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA8FFB43D4F626CFE547565" box="[280,410,541,565]" degrees="98.47580" direction="east" orientation="longitude" pageId="61" pageNumber="402" precision="1" value="98.4758">E98.47580°</geoCoordinate>
,
<date id="FFC1104BFFA8FFB43DF0626CFD9D7565" box="[423,595,541,565]" pageId="61" pageNumber="402" value="2006-08-18">
<collectingDate id="EF85E9A3FFA8FFB43DF0626CFD9D7565" box="[423,595,541,565]" pageId="61" pageNumber="402" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-079 D.H.
<taxonomicName id="4C7F4D08FFA8FFB43FC6626CFBC37565" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[913,1037,541,565]" pageId="61" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFA8FFB4384D626CFB5F7565" box="[1050,1169,541,565]" pageId="61" pageNumber="402">J.A. Miller</collectorName>
,
<collectorName id="268A535DFFA8FFB43C2C624FFF3E7506" box="[123,240,574,598]" pageId="61" pageNumber="402">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFA8FFB43D43624FFE987505" box="[276,342,573,597]" pageId="61" pageNumber="402">Y. Liu</collectorName>
collectors”. All
<typeStatus id="54C48829FFA8FFB43E54624FFDA77506" box="[515,617,574,598]" pageId="61" pageNumber="402" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFA8FFB43FE3624CFBF57505" box="[948,1083,573,597]" pageId="61" pageNumber="402" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFA8FFB43815624CFE077526" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="402" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FFA8FFB43815624CFED67526" italics="true" pageId="61" pageNumber="402">Trechus gongshanensis</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFA8FFB43D83622FFDE67525" box="[468,552,606,629]" pageId="61" pageNumber="402" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="DF006603FFA9FFB53CFC665BFDC47120" ID-DOI="http://doi.org/10.5281/zenodo.13155328" ID-Zenodo-Dep="13155328" httpUri="https://zenodo.org/record/13155328/files/figure.png" pageId="60" pageNumber="401" startId="60.[171,183,1578,1597]" targetBox="[132,1176,216,1563]" targetPageId="60" targetType="figure">
<paragraph id="8BC0368BFFA9FFB53CFC665BFDC47120" blockId="60.[123,1174,1578,1648]" pageId="60" pageNumber="401">
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23.
<taxonomicName id="4C7F4D08FFA9FFB53D5C665BFE1F716D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[267,465,1578,1597]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="401" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FFA9FFB53D5C665BFE1F716D" box="[267,465,1578,1597]" italics="true" pageId="60" pageNumber="401">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA9FFB53D82665BFDD7716D" box="[469,537,1578,1597]" pageId="60" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1001929). b. Median lobe of aedeagus of male (CASENT1001929, left lateral aspect. c. Map of locality record s(red circles) for
<taxonomicName id="4C7F4D08FFA9FFB53F6D6635FC007106" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[826,974,1603,1622]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="401" phylum="Arthropoda" rank="species" species="gongshanensis">
<emphasis id="B90BEA99FFA9FFB53F6D6635FC007106" box="[826,974,1603,1622]" italics="true" pageId="60" pageNumber="401">T. gongshanensis</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFA8FFB43CFC620FFDE975E6" blockId="61.[123,1174,219,1725]" pageId="61" pageNumber="402">
<materialsCitation id="3B173CD6FFA8FFB43CFC620FFDED75E6" country="China" county="Gongshan County" elevation="3450" latitude="27.69655" location="Dongshaofang area" longLatPrecision="1" longitude="95.45389" pageId="61" pageNumber="402" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFA8FFB43CFC620FFEBA75C4" bold="true" box="[171,372,638,662]" pageId="61" pageNumber="402">
<smallCapsWord id="8D26A057FFA8FFB43CFC620FFF2875C5" baselines="656,656" box="[171,230,638,662]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="61" pageNumber="402">
<typeStatus id="54C48829FFA8FFB43CFC620FFF2875C5" box="[171,230,638,662]" pageId="61" pageNumber="402">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFA8FFB43D5162F0FEBA75C4" baselines="655" box="[262,372,641,660]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="61" pageNumber="402">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFA8FFB43DE1620FFE3575C6" box="[438,507,638,662]" name="China" pageId="61" pageNumber="402">China</collectingCountry>
,
<collectingRegion id="49BBF869FFA8FFB43E48620FFDB975C5" box="[543,631,638,661]" country="China" name="Yunnan" pageId="61" pageNumber="402">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFA8FFB43ECD620FFCB275C6" box="[666,892,638,662]" pageId="61" pageNumber="402">Gongshan County</collectingCounty>
,
<location id="8EA06050FFA8FFB43FC8620FFB5F75C5" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFA8FFB43FC8620FFB5F75C5" box="[927,1169,638,662]" country="China" county="Gongshan County" latitude="27.69655" longLatPrecision="1" longitude="95.45389" name="Dongshaofang area" pageId="61" pageNumber="402" stateProvince="Yunnan">Dongshaofang area</location>
,
<geoCoordinate id="EE4B504CFFA8FFB43C2C62EFFF3375E6" box="[123,253,670,694]" degrees="27.69655" direction="north" orientation="latitude" pageId="61" pageNumber="402" precision="1" value="27.69655">N27.69655°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFA8FFB43D5162EFFE4875E6" box="[262,390,670,694]" degrees="95.45389" direction="east" orientation="longitude" pageId="61" pageNumber="402" precision="1" value="95.45389">E95.45389°</geoCoordinate>
,
<quantity id="4C879B6EFFA8FFB43DC662EFFDED75E6" box="[401,547,670,694]" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="61" pageNumber="402" unit="m" value="3450.0" valueMax="3600.0" valueMin="3300.0">
<elevation id="0052D1B8FFA8FFB43DC662EFFDED75E6" box="[401,547,670,694]" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="61" pageNumber="402" unit="m" value="3450.0" valueMax="3600.0" valueMin="3300.0">3300-3600 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA8FFB43CFC62CEFE397447" blockId="61.[123,1174,219,1725]" pageId="61" pageNumber="402">
<emphasis id="B90BEA99FFA8FFB43CFC62CEFE357585" bold="true" box="[171,507,703,727]" pageId="61" pageNumber="402">
<smallCapsWord id="8D26A057FFA8FFB43CFC62CEFEE07585" baselines="721,720" box="[171,302,703,727]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="61" pageNumber="402">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFA8FFB43D6E62B3FE9B7585" baselines="720" box="[313,341,706,725]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="61" pageNumber="402">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFA8FFB43D0862B3FE7C7585" baselines="720" box="[351,434,706,725]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="61" pageNumber="402">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFA8FFB43DE962B3FE357585" baselines="720" box="[446,507,706,725]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="61" pageNumber="402">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFA8FFB43F5962CFFC657586" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[782,939,702,726]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="402" phylum="Arthropoda" rank="species" species="gongshanensis">
<emphasis id="B90BEA99FFA8FFB43F5962CFFC657586" box="[782,939,702,726]" italics="true" pageId="61" pageNumber="402">gongshanensis</emphasis>
</taxonomicName>
, is derived from the name of the county, Gongshan, in which all specimens of the
<typeStatus id="54C48829FFA8FFB43F4862AEFC8375A7" box="[799,845,735,759]" pageId="61" pageNumber="402">type</typeStatus>
series were collected, and the Latin suffix,
<emphasis id="B90BEA99FFA8FFB43D516371FE8A7447" box="[262,324,768,791]" italics="true" pageId="61" pageNumber="402">-ensis</emphasis>
, denoting place.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA8FFB43CFC636EFF237308" pageId="61" pageNumber="402" type="diagnosis">
<paragraph id="8BC0368BFFA8FFB43CFC636EFF237308" blockId="61.[123,1174,219,1725]" pageId="61" pageNumber="402">
<smallCapsWord id="8D26A057FFA8FFB43CFC636EFEEE7465" baselines="817,816" box="[171,288,799,823]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="61" pageNumber="402">
<emphasis id="B90BEA99FFA8FFB43CFC636EFEEE7465" bold="true" box="[171,288,799,823]" pageId="61" pageNumber="402">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFA8FFB43E10636EFD6D7467" box="[583,675,799,823]" captionStart="FIGURE" captionStartId="60.[171,183,1578,1597]" captionTargetBox="[132,1176,216,1563]" captionTargetId="figure-12@60.[120,1176,216,1563]" captionTargetPageId="60" captionText="FIGURE 23. Trechus gongshanensis sp. nov.; a. Dorsal habitus (CASENT1001929). b. Median lobe of aedeagus of male (CASENT1001929, left lateral aspect. c. Map of locality record s(red circles) for T. gongshanensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155328" httpUri="https://zenodo.org/record/13155328/files/figure.png" pageId="61" pageNumber="402">Fig. 23a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size small (BL =
<quantity id="4C879B6EFFA8FFB43878634EFF667428" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="61" pageNumber="402" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
), apterous; body color dark piceous, elytra slightly reddish near sutural and lateral margins; tempora glabrous and only slightly convex; pronotum transverse (ratio PW/PL = 1.45), basal angles acute and sharp; elytra convex, discal striae 1 to 3 or 4 finely impressed, striae 6 to 8 effaced, recurrent stria joined anteriorly with apex of stria 7, two discal setae present on interval 3 next to stria 3, preapical seta present and inserted in a discal, forward position near apical one-fourth of elytra on interval 3, near stria
<quantity id="4C879B6EFFA8FFB43D916470FE3F7349" box="[454,497,1025,1049]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="61" pageNumber="402" unit="in" value="2.0">2 in</quantity>
most individuals, nearer to stria
<quantity id="4C879B6EFFA8FFB43F0D6470FC4A7349" box="[858,900,1025,1049]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="61" pageNumber="402" unit="in" value="3.0">3 in</quantity>
a few; aedeagus of male with base robust, apical lamella elongate with apex blunt, endophallus with a slightly sclerotized scaly area.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFA8FFB73CFC6410FBAD7384" lastPageId="62" lastPageNumber="403" pageId="61" pageNumber="402" type="description">
<paragraph id="8BC0368BFFA8FFB43CFC6410FD3973EA" blockId="61.[123,1174,219,1725]" pageId="61" pageNumber="402">
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<emphasis id="B90BEA99FFA8FFB43CFC6410FEF47328" bold="true" box="[171,314,1121,1145]" pageId="61" pageNumber="402">DESCRIPTION</emphasis>
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.— Size small, BL =
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. Body color dark piceous and shiny, elytra slightly reddish near sutural and lateral margins, appendages paler reddish yellow, palpi paler yellow. Body surface smooth, head capsule faintly alutaceous.
</paragraph>
<paragraph id="8BC0368BFFA8FFB43CFC64B3FE4972AC" blockId="61.[123,1174,219,1725]" pageId="61" pageNumber="402">
Head. Moderate in size, eyes moderately convex, their diameter only slightly longer than length of tempora, the latter glabrous and only slightly convex. Frontal furrows deep, rounded, slightly attenuated posteriorly but prolonged posterior to the eyes. Clypeus with four setae. Labrum with six setae, apical margin distinctly concave. Right mandible as in
<figureCitation id="13442A0EFFA8FFB43F276552FC02726B" box="[880,972,1315,1339]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="61" pageNumber="402">Fig. 16c</figureCitation>
.
<collectingRegion id="49BBF869FFA8FFB43F806552FBFF726A" box="[983,1073,1315,1338]" country="Myanmar" name="Chin" pageId="61" pageNumber="402">Mentum</collectingRegion>
and submentum not fused but nearly so, suture between them very fine and only faintly impressed.
<collectingRegion id="49BBF869FFA8FFB4380B6532FF6A722B" country="Myanmar" name="Chin" pageId="61" pageNumber="402">Mentum</collectingRegion>
with medial area concave, median tooth short, apically bifid, less than half as long as lateral lobes. Submentum with six setae anteriorly, swollen anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae short, extended posteriorly to basal one-fifth of elytra, with about 2 antennomeres extended posteriorly beyond basal pronotal margin; antennomere 3 slightly longer than antennomere 2 or 4.
</paragraph>
<paragraph id="8BC0368BFFA8FFB73CFC6674FB8C7663" blockId="61.[123,1174,219,1725]" lastBlockId="62.[123,1174,219,1236]" lastPageId="62" lastPageNumber="403" pageId="61" pageNumber="402">Pronotum. Transverse (ratio PW/PL = 1.45), widest at anterior one-third, narrowed posteriorly, lateral margins rounded, then abruptly sinuate just anterior to basal angles, the latter acute, sharp and slightly projected laterally. Disc convex, smooth, median longitudinal impression finely impressed but prolonged to the basal margin, slightly widened and deepened in the median basal area, the latter also with several faint longitudinal rugulae and delimited anteriorly by an arcuate posterior transverse impression that is partially effaced medially. Basal foveae distinct but smooth. Basal margin nearly rectilinear. Lateral margination slender throughout and narrowly reflexed, lateral grooves distinctly impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted slightly anterior to apex of basal angle.</paragraph>
<paragraph id="8BC0368BFFABFFB73CFC614AFE557563" blockId="62.[123,1174,219,1236]" pageId="62" pageNumber="403">
Elytra. Ovoid, humeri distinct but rounded. Disc convex, with striae 1 to 3 or 4 finely impressed, the more lateral striae successively more faintly impressed, striae 7 and 8 effaced, striae slightly and irregularly punctate. Parascutellar striole rudimentary. Recurrent stria distinct, moderately deep, terminated at the posterior end of stria 7. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted in a prediscal position on interval 3, closer to stria 2 than
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most specimens (closer to stria
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a few) in forward position at apical one-fourth of elytra.
</paragraph>
<paragraph id="8BC0368BFFABFFB73CFC624DFE507523" blockId="62.[123,1174,219,1236]" pageId="62" pageNumber="403">Legs. Relatively short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFABFFB73CFC620AFB8575E3" blockId="62.[123,1174,219,1236]" pageId="62" pageNumber="403">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFABFFB73CFC62CAFDFB7443" blockId="62.[123,1174,219,1236]" pageId="62" pageNumber="403">
Male aedeagus. (
<figureCitation id="13442A0EFFABFFB73D3E62CAFE027583" box="[361,460,699,723]" captionStart="FIGURE" captionStartId="60.[171,183,1578,1597]" captionTargetBox="[132,1176,216,1563]" captionTargetId="figure-12@60.[120,1176,216,1563]" captionTargetPageId="60" captionText="FIGURE 23. Trechus gongshanensis sp. nov.; a. Dorsal habitus (CASENT1001929). b. Median lobe of aedeagus of male (CASENT1001929, left lateral aspect. c. Map of locality record s(red circles) for T. gongshanensis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155328" httpUri="https://zenodo.org/record/13155328/files/figure.png" pageId="62" pageNumber="403">Fig. 23b</figureCitation>
). Median lobe with basal bulb robust and with a thick sagittal aileron; apical lamella elongate, slightly tapered, with apex blunt. Endophallus nearly unarmed, with only a faintly sclerotized scaly area.
</paragraph>
<paragraph id="8BC0368BFFABFFB73CFC636DFDB87344" blockId="62.[123,1174,219,1236]" pageId="62" pageNumber="403">
<emphasis id="B90BEA99FFABFFB73CFC636DFE6A7462" bold="true" box="[171,420,796,820]" pageId="62" pageNumber="403">
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<smallCapsWord id="8D26A057FFABFFB73D47636EFE6A7462" baselines="813" box="[272,420,799,818]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="62" pageNumber="403">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in a variety of microhabitats in the alpine zone at elevations ranging from
<quantity id="4C879B6EFFABFFB73EC6634DFCF27403" box="[657,828,827,852]" metricMagnitude="3" metricUnit="m" metricValue="3.655" metricValueMax="3.95" metricValueMin="3.36" pageId="62" pageNumber="403" unit="m" value="3655.0" valueMax="3950.0" valueMin="3360.0">3360 to 3950 m</quantity>
. Specimens were collected from under stones in moist meadows, on tundra slopes and ridges with sparse to thick herbaceous vegetation (
<figureCitation id="13442A0EFFABFFB73C86630DFEE574C4" box="[209,299,892,916]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="62" pageNumber="403">Fig. 38a</figureCitation>
,
<figureCitation id="13442A0EFFABFFB73D62630DFE9174C4" box="[309,351,892,916]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="62" pageNumber="403">39b</figureCitation>
,
<figureCitation id="13442A0EFFABFFB73D3D630DFE5C74C4" box="[362,402,892,916]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="62" pageNumber="403">40a</figureCitation>
) and at the edges of small streams and seeps from talus slopes (
<figureCitation id="13442A0EFFABFFB73879630DFB4474C4" box="[1070,1162,892,916]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="62" pageNumber="403">Fig. 39a</figureCitation>
). They were also collected in pitfall traps placed at the edges of
<taxonomicName id="4C7F4D08FFABFFB73F7C63EDFC0774E4" box="[811,969,924,948]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="62" pageNumber="403" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFABFFB73F7C63EDFC0774E4" box="[811,969,924,948]" italics="true" pageId="62" pageNumber="403">Rhododendron</emphasis>
</taxonomicName>
thickets up to two meters tall. Members of this species have been found syntopic with specimens of
<taxonomicName id="4C7F4D08FFABFFB73FB563CDFE4A74A4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="403" phylum="Arthropoda" rank="species" species="balli" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFABFFB73FB563CDFB587484" box="[994,1174,956,980]" italics="true" pageId="62" pageNumber="403">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFABFFB73CD363ADFE8A74A4" box="[132,324,988,1012]" italics="true" pageId="62" pageNumber="403">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FFABFFB73D0463ADFE4A74A4" box="[339,388,988,1012]" italics="true" pageId="62" pageNumber="403">balli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFABFFB73DC563ADFCCD74A4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[402,771,988,1012]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="403" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FFABFFB73DC563ADFD8874A4" box="[402,582,988,1012]" italics="true" pageId="62" pageNumber="403">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFABFFB73E0163ACFD4174A4" box="[598,655,989,1012]" italics="true" pageId="62" pageNumber="403">s. str.</emphasis>
)
<emphasis id="B90BEA99FFABFFB73EC963ADFCCD74A4" box="[670,771,988,1012]" italics="true" pageId="62" pageNumber="403">griswoldi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFABFFB73F4663ADFF517344" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="403" phylum="Arthropoda" rank="species" species="gongshanicus">
<emphasis id="B90BEA99FFABFFB73F4663ADFC0B74A4" box="[785,965,988,1012]" italics="true" pageId="62" pageNumber="403">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFABFFB73F8263ACFBC074A4" box="[981,1038,989,1012]" italics="true" pageId="62" pageNumber="403">s. str.</emphasis>
)
<emphasis id="B90BEA99FFABFFB7384A63ADFF517344" italics="true" pageId="62" pageNumber="403">gongshanicus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7F4D08FFABFFB73C8D638DFE5A7344" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[218,404,1020,1044]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="403" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFABFFB73C8D638DFE5A7344" box="[218,404,1020,1044]" italics="true" pageId="62" pageNumber="403">Trechus qiqiensis</emphasis>
</taxonomicName>
at one or more sites.
</paragraph>
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<emphasis id="B90BEA99FFABFFB73CFC646DFD4E7363" bold="true" box="[171,640,1052,1076]" pageId="62" pageNumber="403">
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<materialsCitation id="3B173CD6FFABFFB73F4E646DFBEB7324" county="Cikai Townships in Gongshan County" location="Bingzhiongluo" municipality="We" pageId="62" pageNumber="403" specimenCount="104" specimenCount-female="30" specimenCount-male="22" stateProvince="Shan">
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<collectingRegion id="49BBF869FFABFFB73F4E646DFC9A7362" box="[793,852,1052,1076]" country="Myanmar" name="Shan" pageId="62" pageNumber="403">SHAN</collectingRegion>
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.—
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.
<collectingMunicipality id="6BA4ACF1FFABFFB73FBC646DFBC07363" box="[1003,1038,1052,1075]" pageId="62" pageNumber="403">We</collectingMunicipality>
examined a total of
<specimenCount id="9D79FD02FFABFFB73C98644DFEAF7304" box="[207,353,1084,1108]" count="52" pageId="62" pageNumber="403" type="generic">52 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFABFFB73D27644DFE1F7304" box="[368,465,1084,1108]" count="22" pageId="62" pageNumber="403" type="male">22 males</specimenCount>
and
<specimenCount id="9D79FD02FFABFFB73E50644DFD4E7304" box="[519,640,1084,1108]" count="30" pageId="62" pageNumber="403" type="female">30 females</specimenCount>
), all from
<location id="8EA06050FFABFFB73EA6644DFC5E7304" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFABFFB73EA6644DFC5E7304" box="[753,912,1084,1108]" county="Cikai Townships in Gongshan County" municipality="We" name="Bingzhiongluo" pageId="62" pageNumber="403" stateProvince="Shan">Bingzhiongluo</location>
and
<collectingCounty id="62A14E07FFABFFB73F92644DFEF07324" pageId="62" pageNumber="403">Cikai Townships in Gongshan County</collectingCounty>
on the crest and eastern slope of the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFABFFB7387A642DFBC173C4" location="Core Area" pageId="62" pageNumber="403" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFABFFB7387A642DFBAC7324" box="[1069,1122,1116,1140]" country="Myanmar" name="Shan" pageId="62" pageNumber="403">Shan</collectingRegion>
(see
<typeStatus id="54C48829FFABFFB73C2C640DFF7E73C4" box="[123,176,1148,1172]" pageId="62" pageNumber="403">Type</typeStatus>
material above for exact collection data). These localities are all in
<location id="8EA06050FFABFFB73FDC640DFC3673C4" LSID="urn:lsid:plazi:treatment:03D6879DFFAEFFB73C2C6191FBAD7384:8EA06050FFABFFB73FDC640DFC3673C4" box="[907,1016,1148,1172]" name="Core Area" pageId="62" pageNumber="403" stateProvince="Shan">Core Area</location>
2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFABFFB73CFC64EDFBAD7384" blockId="62.[123,1174,219,1236]" pageId="62" pageNumber="403">
<emphasis id="B90BEA99FFABFFB73CFC64EDFD9473E2" bold="true" box="[171,602,1180,1204]" pageId="62" pageNumber="403">
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</emphasis>
.— This species currently is known only from Gongshan County in the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFABFFB73F0164CCFBDF7384" box="[854,1041,1212,1236]" country="China" name="Yunnan" pageId="62" pageNumber="403">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFABFFB7384B64CDFB917384" box="[1052,1119,1212,1236]" name="China" pageId="62" pageNumber="403">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C2052D047CA74A3651BEF5F28EEB499E">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="4C01361790790A2E5E4433B3FBE3E039">Deuve, Thierry</mods:namePart>
<mods:affiliation id="78849EC9FDFCAE75B842F2047368BEDC">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="45D77BD176AED14CDCDBA8407E7E9F4A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="F5BEDCF6BC5CFE535A212008A2E8985C">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FFB1FFAD3C2C6292FEF875AB" bold="true" box="[123,310,739,763]" italics="true" pageId="36" pageNumber="377">Eocnides fragilis</emphasis>
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(
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,
<figureCitation id="13442A0EFFB1FFAD3C8A6375FED4744A" box="[221,282,772,794]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStartId-0="106.[171,183,1674,1693]" captionStartId-1="107.[171,183,1629,1648]" captionStartId-2="108.[171,183,1359,1378]" captionStartId-3="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,974,1660]" captionTargetBox-1="[120,1176,245,1580]" captionTargetBox-2="[120,1176,228,1340]" captionTargetBox-3="[197,1099,216,1598]" captionTargetId-0="figure-14@106.[120,1176,974,1660]" captionTargetId-1="figure-14@107.[120,1176,216,1608]" captionTargetId-2="figure-12@108.[120,1176,228,1340]" captionTargetId-3="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="106" captionTargetPageId-1="107" captionTargetPageId-2="108" captionTargetPageId-3="109" captionText-0="FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km." captionText-1="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-2="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-3="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155376" figureDoi-1="http://doi.org/10.5281/zenodo.13155378" figureDoi-2="http://doi.org/10.5281/zenodo.13155380" figureDoi-3="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155376/files/figure.png" httpUri-1="https://zenodo.org/record/13155378/files/figure.png" httpUri-2="https://zenodo.org/record/13155380/files/figure.png" httpUri-3="https://zenodo.org/record/13155382/files/figure.png" pageId="36" pageNumber="377">4548</figureCitation>
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<emphasis id="B90BEA99FFB1FFAD3C2C6341FEED7415" box="[123,291,816,837]" italics="true" pageId="36" pageNumber="377">Eocnides fragilis</emphasis>
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, a male, in NMST. Type locality:
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,
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, ca.
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<emphasis id="B90BEA99FFB1FFAD3C2C631FFE7774D3" box="[123,441,878,899]" italics="true" pageId="36" pageNumber="377">Agonotrechiotes longiantennatus</emphasis>
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<emphasis id="B90BEA99FFB1FFAD3CFC63E9FEEE74FE" bold="true" box="[171,288,920,944]" pageId="36" pageNumber="377">DIAGNOSIS</emphasis>
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.— Adults of this species (
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) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL =
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), fully-winged; body color pale, brownish-tan; head with eyes only moderate in size; frontal furrows attenuated; anterior pair of supraorbital setae not foveate; pronotum slightly transverse (ratio PW/PL= 1.48), median longitudinal impression distinctly impressed, especially basally, least so at middle; basal foveae prolonged anteriorly parallel to lateral margin in basal one-half; elytra with discal stria 2 partially effaced near base, striae 5 and 6 faintly but distinctly impressed; medi- an lobe of male aedeagus (
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) slender, apex slightly recurved ventrally.
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.— Members of this species were found by splashing gravel bars at the edges of a small, lowland stream adjacent to disturbed, agricultural areas, at elevations ranging from
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. This microhabitat is similar to that described by
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for specimens collected at Munigou He,
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, in Songpan County, northern
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. No other trechines were collected at these sites.
</paragraph>
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.
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examined a total of
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) from the following localities:
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:
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(
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, just below
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,
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,
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/
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,
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,
<collectorName id="268A535DFFB1FFAD3EC565D0FCC272E9" box="[658,780,1441,1465]" pageId="36" pageNumber="377">P.E. Marek</collectorName>
and
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collectors [
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(
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, just below
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,
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,
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,
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,
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,
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and
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collectors [
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;
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].
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of these localities are in
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2.
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</paragraph>
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.—
<figureCitation id="13442A0EFFB1FFAD3ED56652FD1C716B" box="[642,722,1571,1595]" captionStart="FIGURE" captionStartId="106.[171,183,1674,1693]" captionTargetBox="[120,1176,974,1660]" captionTargetId="figure-14@106.[120,1176,974,1660]" captionTargetPageId="106" captionText="FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.13155376" httpUri="https://zenodo.org/record/13155376/files/figure.png" pageId="36" pageNumber="377">Fig. 45</figureCitation>
. This species has been recorded previously from five localities in northern
<collectingRegion id="49BBF869FFB1FFAD3DA76635FD7F710C" box="[496,689,1604,1628]" country="China" name="Sichuan" pageId="36" pageNumber="377">Sichuan Province</collectingRegion>
, in Jiuzhaigou and Songpan Counties (
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,
<bibRefCitation id="EFEE4B7AFFB1FFAD3CE86615FECA712C" author="UENO, S. I." box="[191,260,1636,1660]" pageId="36" pageNumber="377" pagination="275 - 276" refId="ref61481" refString="UENO, S. I. 1999 b. Two new localities of Eocnides fragilis (Coleoptera, Trechinae), with brief notes on its habitats. Elytra 27: 275 - 276." type="journal article" year="1999">1999b</bibRefCitation>
) and in Luding County (
<bibRefCitation id="EFEE4B7AFFB1FFAD3E436615FD50712C" author="DEUVE, T." box="[532,670,1636,1660]" pageId="36" pageNumber="377" pagination="1 - 24" refId="ref59005" refString="DEUVE, T. 2010. Nouveaux Nebriidae, Broscidae et Trechidae de Chine et d'iran (Coleoptera, Caraboidea). Revue Francaise d'Entomologie, (N. S.) 32: 1 - 24." type="journal article" year="2010">Deuve 2010</bibRefCitation>
). We report here the following new record for
<collectingRegion id="49BBF869FFB1FFAD3C2C66F4FF1B71CD" box="[123,213,1669,1693]" country="China" name="Sichuan" pageId="36" pageNumber="377">Sichuan</collectingRegion>
:
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: Qionglai
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, Sanjiang Township (Sanjiang Nature Reserve, river at gate to reserve,
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,
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,
<date id="FFC1104BFFB1FFAD3E9666D7FC4971EE" box="[705,903,1701,1726]" pageId="36" pageNumber="377" value="2007-09-09">9 September 2007</date>
, D.H.
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collector [
<specimenCount id="9D79FD02FFB3FFAF3CFB60AAFEC177A3" box="[172,271,219,243]" count="1" pageId="38" pageNumber="379" type="female">1 female</specimenCount>
; CAS]. This record extends the range of
<taxonomicName id="4C7F4D08FFB3FFAF3E8A60ADFC8577A3" authorityName="Ueno. We" authorityYear="1989" box="[733,843,219,243]" class="Insecta" family="Carabidae" genus="Eocnides" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="379" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B90BEA99FFB3FFAF3E8A60ADFC8577A3" box="[733,843,219,243]" italics="true" pageId="38" pageNumber="379">E. fragilis</emphasis>
</taxonomicName>
in
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. The discovery of
<taxonomicName id="4C7F4D08FFB3FFAF3C2C608DFF287643" authorityName="Ueno. We" authorityYear="1989" box="[123,230,251,275]" class="Insecta" family="Carabidae" genus="Eocnides" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="379" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B90BEA99FFB3FFAF3C2C608DFF287643" box="[123,230,251,275]" italics="true" pageId="38" pageNumber="379">E. fragilis</emphasis>
</taxonomicName>
in the northern Gaoligong
<collectingRegion id="49BBF869FFB3FFAF3E5D608AFD407643" box="[522,654,251,275]" country="Myanmar" name="Shan" pageId="38" pageNumber="379">Shan region</collectingRegion>
, in western
<collectingRegion id="49BBF869FFB3FFAF3F58608DFC077643" box="[783,969,251,275]" country="China" name="Yunnan" pageId="38" pageNumber="379">Yunnan Province</collectingRegion>
, extends its known range an additional
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SW.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="1D695583EC7D53B7219C2EC4CF8F7224" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFB5FFAA3C2C626AFC5F7233" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Queinnectrechus (Gaoligongtrechus) balli Deuve &amp; Kavanaugh &amp; Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="376" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="373" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AF619C0F84356EABE7C3CC536F955DAD">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="08AEEA418F765B56A86B5A19297D872B">Deuve, Thierry</mods:namePart>
<mods:affiliation id="164BCC6C7383E42ADEF5DC7F6DCC1344">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="B839472ED5E27371FC9C4863F5018420">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="4FC0DE8A97921F4106BBD8E157C3132C">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:namePart id="88FA587EB98AEC3892DE5B867ECB7253">Liang, Hongbin</mods:namePart>
<mods:affiliation id="5213E1890772B9B8C1877A0BA69740A9">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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(
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Deuve
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and Kavanaugh,
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(
<figureCitation id="13442A0EFFB5FFA93CD5624AFF1D7500" box="[130,211,571,593]" captionStart="FIGURE" captionStartId="33.[171,183,1577,1596]" captionTargetBox="[144,1175,216,1563]" captionTargetId="figure-15@33.[120,1175,216,1563]" captionTargetPageId="33" captionText="FIGURE 14. Queinnectrechus (Gaoligongtrechus) balli sp. nov.; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for Q. balli in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155309" httpUri="https://zenodo.org/record/13155309/files/figure.png" pageId="32" pageNumber="373">Figs. 14</figureCitation>
,
<figureCitation id="13442A0EFFB5FFA93C8A624AFECD7501" box="[221,259,571,593]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="32" pageNumber="373">37b</figureCitation>
,
<figureCitation id="13442A0EFFB5FFA93D59624AFEFA7501" box="[270,308,571,593]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="32" pageNumber="373">39b</figureCitation>
,
<figureCitation id="13442A0EFFB5FFA93D69624AFEAD7500" box="[318,355,571,592]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="32" pageNumber="373">40a</figureCitation>
,
<figureCitation id="13442A0EFFB5FFA93D39624AFE657501" box="[366,427,571,593]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="32" pageNumber="373">4648</figureCitation>
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<emphasis id="B90BEA99FFB5FFA93ED16295FCF475AC" box="[646,826,740,764]" italics="true" pageId="32" pageNumber="373">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFB5FFA93F1E6295FBC775AC" box="[841,1033,740,764]" italics="true" pageId="32" pageNumber="373">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FFB5FFA9384F6295FB8775AC" box="[1048,1097,740,764]" italics="true" pageId="32" pageNumber="373">balli</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFB5FFA93D746374FEB7744C" box="[291,377,773,796]" pageId="32" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FFB5FFA93E9B6374FBE3740C" collectionCode="CAS, IOZ, MNHN" country="China" pageId="32" pageNumber="373" specimenCount="14" specimenCount-female="12" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFB5FFA93E9B6374FCFA744D" box="[716,820,773,797]" pageId="32" pageNumber="373" type="paratype">Paratypes</typeStatus>
(a total of 32):
<specimenCount id="9D79FD02FFB5FFA93FBC6375FB8E744C" box="[1003,1088,772,796]" count="2" pageId="32" pageNumber="373" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FFB5FFA9382C6375FF00746C" count="12" pageId="32" pageNumber="373" type="female">12 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA93CAB6355FEFC746C" box="[252,306,804,828]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="373" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFB5FFA93D696355FEA3746C" box="[318,365,804,828]" pageId="32" pageNumber="373">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFB5FFA93D2E6354FE02746D" box="[377,460,805,829]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="373">MNHN</collectionCode>
) labeled same as
<typeStatus id="54C48829FFB5FFA93ED86355FD22746C" box="[655,748,804,828]" pageId="32" pageNumber="373" type="holotype">holotype</typeStatus>
except first label “CASENT 1001933” to “CASENT 1001934” and “CASENT 1001936” to “CASENT 1001947”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA9386F6334FBCB74EC" collectingDate="2002-10-01" collectingDateMax="2002-10-02" collectingDateMin="2002-10-01" collectionCode="CAS" collectorName="P. E. Marek" country="China" county="Gongshan County" elevation="3370" latitude="27.77032" location="Cikai Township" longLatPrecision="1" longitude="98.44661" pageId="32" pageNumber="373" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA9386F6334FB58740C" box="[1080,1174,836,861]" count="1" pageId="32" pageNumber="373" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA93CF46314FF16742D" box="[163,216,869,893]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="373" type="Museum">CAS</collectionCode>
) labeled “CASENT 1010343”/ “
<collectingCountry id="F368761BFFB5FFA93E136314FD53742D" box="[580,669,869,893]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93EFC6314FD31742C" box="[683,767,869,892]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93F5A6315FC1A742D" box="[781,980,868,893]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<location id="8EA06050FFB5FFA93FB56315FB5C742C" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93FB56315FB5C742C" box="[994,1170,868,892]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44661" name="Cikai Township" pageId="32" pageNumber="373" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFB5FFA93C2C63F5FD4574CC" box="[123,651,900,924]" location="Gongshan on Dulong Valley Road" pageId="32" pageNumber="373">
<quantity id="4C879B6EFFB5FFA93C2C63F5FF1D74CC" box="[123,211,900,924]" metricMagnitude="4" metricUnit="m" metricValue="5.26" pageId="32" pageNumber="373" unit="km" value="52.6">52.6 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFFB5FFA93EC063F5FCE774CD" box="[663,809,900,925]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="32" pageNumber="373" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FFB5FFA93EC063F5FCE774CD" box="[663,809,900,925]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="32" pageNumber="373" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFB5FFA93F0663F4FC1B74CD" box="[849,981,901,925]" degrees="27.77032" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93FB263F5FBBD74CC" box="[997,1139,900,924]" degrees="098.44661" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.44661">E098.44661°</geoCoordinate>
,
<date id="FFC1104BFFB5FFA9382863F4FEE974ED" pageId="32" pageNumber="373" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">
<collectingDate id="EF85E9A3FFB5FFA9382863F4FEE974ED" pageId="32" pageNumber="373" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">1- 2 October 2002</collectingDate>
</date>
, Stop #DHK-2002-034, D.H.
<taxonomicName id="4C7F4D08FFB5FFA93E3963D5FD2974EC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[622,743,932,956]" pageId="32" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFB5FFA93F5963D4FC4B74EC" box="[782,901,932,956]" pageId="32" pageNumber="373">P.E. Marek</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA9384663D4FC50734D" collectingDate="2006-08-11" collectionCode="CAS" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3365" latitude="27.77032" location="Heipu Yakou" longLatPrecision="1" longitude="98.44674" municipality="Cikai Township" pageId="32" pageNumber="373" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA9384663D4FBBE74EC" box="[1041,1136,932,957]" count="1" pageId="32" pageNumber="373" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA93C2C63B4FF7E748D" box="[123,176,965,989]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="373" type="Museum">CAS</collectionCode>
) labeled “CASENT 1024375”/ “
<collectingCountry id="F368761BFFB5FFA93E5A63B4FDA8748D" box="[525,614,965,989]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93E2763B4FD0A748C" box="[624,708,965,988]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93E9863B5FC5C748D" box="[719,914,964,989]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB5FFA93FCB63B5FB86748C" box="[924,1096,964,988]" pageId="32" pageNumber="373">Cikai Township</collectingMunicipality>
, southeast slope of
<location id="8EA06050FFB5FFA93D5C6395FE5474AC" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93D5C6395FE5474AC" box="[267,410,996,1020]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44674" municipality="Cikai Township" name="Heipu Yakou" pageId="32" pageNumber="373" stateProvince="Yunnan">Heipu Yakou</location>
,
<quantity id="4C879B6EFFB5FFA93DF16395FE3574AC" box="[422,507,996,1020]" metricMagnitude="3" metricUnit="m" metricValue="3.3649999999999998" pageId="32" pageNumber="373" unit="m" value="3365.0">
<elevation id="0052D1B8FFB5FFA93DF16395FE3574AC" box="[422,507,996,1020]" metricMagnitude="3" metricUnit="m" metricValue="3.3649999999999998" pageId="32" pageNumber="373" unit="m" value="3365.0">3365 m</elevation>
</quantity>
,
<geoCoordinate id="EE4B504CFFB5FFA93E506394FD4674AD" box="[519,648,997,1021]" degrees="27.77032" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93ECE6395FCE774AC" box="[665,809,996,1020]" degrees="098.44674" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.44674">E098.44674°</geoCoordinate>
,”/ “
<date id="FFC1104BFFB5FFA93F056394FBCE74AC" box="[850,1024,996,1021]" pageId="32" pageNumber="373" value="2006-08-11">
<collectingDate id="EF85E9A3FFB5FFA93F056394FBCE74AC" box="[850,1024,996,1021]" pageId="32" pageNumber="373" value="2006-08-11">11 August 2006</collectingDate>
</date>
, Stop #DHK-2006-069A, D.H.
<taxonomicName id="4C7F4D08FFB5FFA93D6B6474FE76734D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[316,440,1029,1053]" pageId="32" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFB5FFA93D946474FDF6734D" box="[451,568,1029,1053]" pageId="32" pageNumber="373">J.A. Miller</collectorName>
,
<collectorName id="268A535DFFB5FFA93E146474FD77734D" box="[579,697,1029,1053]" pageId="32" pageNumber="373">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFB5FFA93E896474FCEE734D" box="[734,800,1029,1053]" pageId="32" pageNumber="373">Y. Liu</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA93FFF6474FC0473CD" collectingDate="2006-08-19" collectionCode="IOZ, MNHN" collectorName="Y. Liu" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.44779" municipality="Bingzhongluo Township" pageId="32" pageNumber="373" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA93FFF6474FC34734D" box="[936,1018,1029,1053]" count="3" pageId="32" pageNumber="373" type="male">3 males</specimenCount>
and
<specimenCount id="9D79FD02FFB5FFA938796474FB5B734D" box="[1070,1173,1029,1053]" count="2" pageId="32" pageNumber="373" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA93CF76454FF01736D" box="[160,207,1061,1085]" pageId="32" pageNumber="373">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFB5FFA93C8D6454FEE3736D" box="[218,301,1061,1085]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="373">MNHN</collectionCode>
) labeled “CASENT 1025840” to “CASENT 1025842” and “CASENT 1025838” to “CASENT 1025839”, respectively/ “
<collectingCountry id="F368761BFFB5FFA93E766434FDB4730D" box="[545,634,1093,1117]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93ED26434FD17730C" box="[645,729,1093,1116]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93EB36434FC66730D" box="[740,936,1093,1117]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB5FFA93FE56434FF64732D" pageId="32" pageNumber="373">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFB5FFA93D1C6414FDC6732D" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93D1C6414FDC6732D" box="[331,520,1125,1149]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.44779" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="32" pageNumber="373" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFB5FFA93E426414FC99732D" box="[533,855,1125,1149]" location="Chukuai Lake" pageId="32" pageNumber="373">
<quantity id="4C879B6EFFB5FFA93E426414FDAC732D" box="[533,610,1125,1149]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="373" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/ “
<geoCoordinate id="EE4B504CFFB5FFA9385D6414FB43732D" box="[1034,1165,1125,1149]" degrees="27.97686" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93C2C64F4FEC473CD" box="[123,266,1157,1181]" degrees="098.44779" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.44779">E098.44779°</geoCoordinate>
,
<quantity id="4C879B6EFFB5FFA93D4464F5FEA873CC" box="[275,358,1156,1180]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="32" pageNumber="373" unit="m" value="3750.0">
<elevation id="0052D1B8FFB5FFA93D4464F5FEA873CC" box="[275,358,1156,1180]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="32" pageNumber="373" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFFB5FFA93D2764F4FDD473CD" box="[368,538,1157,1181]" pageId="32" pageNumber="373" value="2006-08-19">
<collectingDate id="EF85E9A3FFB5FFA93D2764F4FDD473CD" box="[368,538,1157,1181]" pageId="32" pageNumber="373" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-082
<collectorName id="268A535DFFB5FFA93F4164F4FC9973CD" box="[790,855,1157,1181]" pageId="32" pageNumber="373">Y. Liu</collectorName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA93F8464F4FBB373AD" collectingDate="2006-08-19" collectionCode="CAS" collectorName="J. A. Miller &amp; D. Z. Dong" country="China" county="Gongshan County" elevation="3745" latitude="27.98393" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" pageId="32" pageNumber="373" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA93F8464F4FBFE73CD" box="[979,1072,1157,1181]" count="1" pageId="32" pageNumber="373" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA9380F64F4FB4373CD" box="[1112,1165,1157,1181]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="373" type="Museum">CAS</collectionCode>
) labeled “CASENT 1026202”/ “
<collectingCountry id="F368761BFFB5FFA93D9D64D4FDED73ED" box="[458,547,1189,1213]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93E7A64D4FD4F73EC" box="[557,641,1189,1212]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93EDB64D4FC8173ED" box="[652,847,1189,1213]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB5FFA93F0D64D4FBAD73ED" box="[858,1123,1189,1213]" pageId="32" pageNumber="373">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFB5FFA93C8F64B4FE5D738D" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93C8F64B4FE5D738D" box="[216,403,1221,1245]" country="China" county="Gongshan County" latitude="27.98393" longLatPrecision="1" longitude="98.47491" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="32" pageNumber="373" stateProvince="Yunnan">Kawakarpu Shan</location>
,
<locationDeviation id="9A722FA9FFB5FFA93DC964B4FD29738D" box="[414,743,1221,1245]" location="Chukuai Lake" pageId="32" pageNumber="373">
<quantity id="4C879B6EFFB5FFA93DC964B4FE27738D" box="[414,489,1221,1245]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="373" unit="km" value="0.3">0.3 km</quantity>
NNE of Chukuai Lake
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFB5FFA93EA464B4FCBA738D" box="[755,884,1221,1245]" degrees="27.98393" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.98393">N27.98393°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93FD464B4FBDD738D" box="[899,1043,1221,1245]" degrees="098.47491" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.47491">E098.47491°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFB5FFA9386C64B4FB5F738C" box="[1083,1169,1221,1245]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="32" pageNumber="373" unit="m" value="3745.0">
<elevation id="0052D1B8FFB5FFA9386C64B4FB5F738C" box="[1083,1169,1221,1245]" metricMagnitude="3" metricUnit="m" metricValue="3.745" pageId="32" pageNumber="373" unit="m" value="3745.0">3745 m</elevation>
</quantity>
,
<date id="FFC1104BFFB5FFA93C2C6494FEE673AD" box="[123,296,1253,1277]" pageId="32" pageNumber="373" value="2006-08-19">
<collectingDate id="EF85E9A3FFB5FFA93C2C6494FEE673AD" box="[123,296,1253,1277]" pageId="32" pageNumber="373" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-081 D.H.
<taxonomicName id="4C7F4D08FFB5FFA93E326494FD2F73AD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[613,737,1253,1277]" pageId="32" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFB5FFA93EBB6494FCAD73AD" box="[748,867,1253,1277]" pageId="32" pageNumber="373">J.A. Miller</collectorName>
, &amp;
<collectorName id="268A535DFFB5FFA93FDC6494FC3073AD" box="[907,1022,1253,1277]" pageId="32" pageNumber="373">D.Z. Dong</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA938DF6494FBB5722D" collectingDate="2006-08-13" collectionCode="CAS, IOZ" collectorName="J. A. Miller" country="China" county="Gongshan County" elevation="3720" latitude="27.76978" location="Cikai Township" longLatPrecision="1" longitude="98.44681" pageId="32" pageNumber="373" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA938DF6494FF77724D" count="5" pageId="32" pageNumber="373" type="male">5 males</specimenCount>
and
<specimenCount id="9D79FD02FFB5FFA93CA56574FE92724D" box="[242,348,1285,1309]" count="2" pageId="32" pageNumber="373" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA93DDA6574FE0D724D" box="[397,451,1285,1309]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="373" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFB5FFA93D866574FE31724D" box="[465,511,1285,1309]" pageId="32" pageNumber="373">IOZ</collectionCode>
) labeled ““CASENT 1026323” to “CASENT 1026327” and “CASENT 1026328” to “CASENT 1026329”, respectively/ “
<collectingCountry id="F368761BFFB5FFA93F5B6554FCAB726D" box="[780,869,1317,1341]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93F386554FC0D726C" box="[879,963,1317,1340]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93F996554FB5C726D" box="[974,1170,1317,1341]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<location id="8EA06050FFB5FFA93C2C6534FEE7720D" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93C2C6534FEE7720D" box="[123,297,1349,1373]" country="China" county="Gongshan County" latitude="27.76978" longLatPrecision="1" longitude="98.44681" name="Cikai Township" pageId="32" pageNumber="373" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FFB5FFA93D626534FD90720D" box="[309,606,1349,1373]" location="Heipu Yakou" pageId="32" pageNumber="373">
<quantity id="4C879B6EFFB5FFA93D626534FE4F720D" box="[309,385,1349,1373]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="373" unit="km" value="0.1">0.1 km</quantity>
SE of Heipu Yakou
</locationDeviation>
in valley below tunnel,
<geoCoordinate id="EE4B504CFFB5FFA93F3D6534FC22720D" box="[874,1004,1349,1373]" degrees="27.76978" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.76978">N27.76978°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93FAA6534FBB1720D" box="[1021,1151,1349,1373]" degrees="98.44681" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.44681">E98.44681°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFB5FFA93CD36514FF12722C" box="[132,220,1381,1405]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="32" pageNumber="373" unit="m" value="3720.0">
<elevation id="0052D1B8FFB5FFA93CD36514FF12722C" box="[132,220,1381,1405]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="32" pageNumber="373" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,
<date id="FFC1104BFFB5FFA93CBD6514FE52722D" box="[234,412,1381,1405]" pageId="32" pageNumber="373" value="2006-08-13">
<collectingDate id="EF85E9A3FFB5FFA93CBD6514FE52722D" box="[234,412,1381,1405]" pageId="32" pageNumber="373" value="2006-08-13">13 August 2006</collectingDate>
</date>
, Stop #DHK-2006-073 D.H.
<taxonomicName id="4C7F4D08FFB5FFA93EB66514FC94722D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[737,858,1381,1405]" pageId="32" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFB5FFA93FD46514FC34722D" box="[899,1018,1381,1405]" pageId="32" pageNumber="373">J.A. Miller</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA938DF6514FC97728D" collectingDate="2006-08-13" collectionCode="CAS" collectorName="J. A. Miller" country="China" county="Gongshan County" elevation="3350" latitude="27.77437" location="Heipu Yakou" longLatPrecision="1" longitude="98.44793" municipality="Dulongjiang Township" pageId="32" pageNumber="373" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA938DF6514FF6172CD" count="1" pageId="32" pageNumber="373" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA93C8965F4FEDD72CD" box="[222,275,1413,1437]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="373" type="Museum">CAS</collectionCode>
) labeled “CASENT 1026707”/ “
<collectingCountry id="F368761BFFB5FFA93E2F65F4FD1F72CD" box="[632,721,1413,1437]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93E8965F4FCFC72CC" box="[734,818,1413,1436]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93F6865F4FBCA72CD" box="[831,1028,1413,1437]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB5FFA9384765F4FF2972ED" pageId="32" pageNumber="373">Dulongjiang Township</collectingMunicipality>
, NW slope of
<location id="8EA06050FFB5FFA93DDB65D4FDD372ED" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93DDB65D4FDD372ED" box="[396,541,1445,1469]" country="China" county="Gongshan County" latitude="27.77437" longLatPrecision="1" longitude="98.44793" municipality="Dulongjiang Township" name="Heipu Yakou" pageId="32" pageNumber="373" stateProvince="Yunnan">Heipu Yakou</location>
,
<quantity id="4C879B6EFFB5FFA93E7C65D4FD4C72EC" box="[555,642,1445,1469]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="32" pageNumber="373" unit="m" value="3350.0">
<elevation id="0052D1B8FFB5FFA93E7C65D4FD4C72EC" box="[555,642,1445,1469]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="32" pageNumber="373" unit="m" value="3350.0">3350 m</elevation>
</quantity>
,
<geoCoordinate id="EE4B504CFFB5FFA93EC765D4FCDF72ED" box="[656,785,1445,1469]" degrees="27.77437" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.77437">N27.77437°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93F7465D4FC7D72ED" box="[803,947,1445,1469]" degrees="098.44793" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.44793">E098.44793°</geoCoordinate>
,”/ “
<date id="FFC1104BFFB5FFA93F8965D4FB5F72ED" box="[990,1169,1445,1469]" pageId="32" pageNumber="373" value="2006-08-13">
<collectingDate id="EF85E9A3FFB5FFA93F8965D4FB5F72ED" box="[990,1169,1445,1469]" pageId="32" pageNumber="373" value="2006-08-13">13 August 2006</collectingDate>
</date>
, Stop #DHK-2006-075 D.H.
<taxonomicName id="4C7F4D08FFB5FFA93DEF65B4FDFF728D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[440,561,1477,1501]" pageId="32" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFB5FFA93E0A65B4FD18728D" box="[605,726,1477,1501]" pageId="32" pageNumber="373">J.A. Miller</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB5FFA93F3F65B4FBFE712D" collectingDate="2006-08-18" collectionCode="IOZ" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="32" pageNumber="373" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB5FFA93F3F65B4FC1D728D" box="[872,979,1477,1501]" count="2" pageId="32" pageNumber="373" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB5FFA9385F65B4FBF8728D" box="[1032,1078,1477,1501]" pageId="32" pageNumber="373">IOZ</collectionCode>
) labeled “CASENT 1026817” and “CASENT 1026818”, respectively/ “
<collectingCountry id="F368761BFFB5FFA93F4A6594FCB872AD" box="[797,886,1509,1533]" name="China" pageId="32" pageNumber="373">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93FD76594FC1A72AC" box="[896,980,1509,1532]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB5FFA93F886594FF5F714D" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB5FFA93CF26674FE76714D" box="[165,440,1541,1565]" pageId="32" pageNumber="373">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFB5FFA93E236674FCFA714D" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93E236674FCFA714D" box="[628,820,1541,1565]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="32" pageNumber="373" stateProvince="Yunnan">Kawakarpu Shan</location>
at
<location id="8EA06050FFB5FFA93F3E6674FBC2714D" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93F3E6674FBC2714D" box="[873,1036,1541,1565]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="32" pageNumber="373" stateProvince="Yunnan">Chukuai Lake</location>
,
<quantity id="4C879B6EFFB5FFA938766674FBB0714D" box="[1057,1150,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="32" pageNumber="373" unit="m" value="3720.0">
<elevation id="0052D1B8FFB5FFA938766674FBB0714D" box="[1057,1150,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="32" pageNumber="373" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFB5FFA93CD36654FEC9716D" box="[132,263,1573,1597]" degrees="27.98121" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93D4F6654FE69716D" box="[280,423,1573,1597]" degrees="098.47580" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.4758">E098.47580°</geoCoordinate>
,
<date id="FFC1104BFFB5FFA93DE46654FD90716D" box="[435,606,1573,1597]" pageId="32" pageNumber="373" value="2006-08-18">
<collectingDate id="EF85E9A3FFB5FFA93DE46654FD90716D" box="[435,606,1573,1597]" pageId="32" pageNumber="373" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-079
<collectorName id="268A535DFFB5FFA93F0A6654FC1A716D" box="[861,980,1573,1597]" pageId="32" pageNumber="373">J.A. Miller</collectorName>
,
<collectorName id="268A535DFFB5FFA93FB76654FB99716D" box="[992,1111,1573,1597]" pageId="32" pageNumber="373">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFB5FFA938286654FF6E710D" pageId="32" pageNumber="373">Y. Liu</collectorName>
collectors”. All
<typeStatus id="54C48829FFB5FFA93D306637FE03710E" box="[359,461,1606,1630]" pageId="32" pageNumber="373" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFB5FFA93F1B6634FC1D710D" box="[844,979,1605,1629]" pageId="32" pageNumber="373" type="paratype">PARATYPE</typeStatus>
<emphasis id="B90BEA99FFB5FFA93FB56634FB45710D" box="[994,1163,1605,1629]" italics="true" pageId="32" pageNumber="373">Queinnectrechu</emphasis>
s
<taxonomicName id="4C7F4D08FFB5FFA93C2C6614FE1C712D" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,466,1637,1661]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="373" phylum="Arthropoda" rank="species" species="balli" subGenus="Gaoligongtrechus">
(
<emphasis id="B90BEA99FFB5FFA93CD36614FE8A712D" box="[132,324,1637,1661]" italics="true" pageId="32" pageNumber="373">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FFB5FFA93D046614FE4A712D" box="[339,388,1637,1661]" italics="true" pageId="32" pageNumber="373">balli</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFB5FFA93E2B6617FD01712D" box="[636,719,1638,1661]" pageId="32" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFB5FFA93CFC66F4FDAC71ED" blockId="32.[123,1174,612,1725]" pageId="32" pageNumber="373">
<materialsCitation id="3B173CD6FFB5FFA93CFC66F4FD9071ED" country="China" county="Gongshan County" elevation="3490" latitude="27.69655" location="Qiqi" longLatPrecision="1" longitude="98.45389" municipality="Gaoligong Shan" pageId="32" pageNumber="373" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFB5FFA93CFC66F4FE9771CC" bold="true" box="[171,345,1669,1693]" pageId="32" pageNumber="373">
<smallCapsWord id="8D26A057FFB5FFA93CFC66F4FF2A71CC" baselines="1687,1687" box="[171,228,1669,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="32" pageNumber="373">
<typeStatus id="54C48829FFB5FFA93CFC66F4FF2A71CC" box="[171,228,1669,1693]" pageId="32" pageNumber="373">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFB5FFA93CB966F8FE9771CC" baselines="1687" box="[238,345,1673,1692]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="32" pageNumber="373">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFB5FFA93DD466F4FE0B71CD" box="[387,453,1669,1693]" name="China" pageId="32" pageNumber="373">China</collectingCountry>
,
<collectingRegion id="49BBF869FFB5FFA93D8666F7FDEB71CD" box="[465,549,1670,1693]" country="China" name="Yunnan" pageId="32" pageNumber="373">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFB5FFA93E6566F4FD2A71CD" box="[562,740,1669,1693]" pageId="32" pageNumber="373">Gaoligong Shan</collectingMunicipality>
,
<collectingCounty id="62A14E07FFB5FFA93EA666F4FC7871CD" box="[753,950,1669,1693]" pageId="32" pageNumber="373">Gongshan County</collectingCounty>
,
<location id="8EA06050FFB5FFA93F9466F4FC3C71CD" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93F9466F4FC3C71CD" box="[963,1010,1669,1693]" country="China" county="Gongshan County" latitude="27.69655" longLatPrecision="1" longitude="98.45389" municipality="Gaoligong Shan" name="Qiqi" pageId="32" pageNumber="373" stateProvince="Yunnan">Qiqi</location>
/
<location id="8EA06050FFB5FFA93FAD66F4FB8771CD" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB5FFA93FAD66F4FB8771CD" box="[1018,1097,1669,1693]" country="China" county="Gongshan County" latitude="27.69655" longLatPrecision="1" longitude="98.45389" municipality="Gaoligong Shan" name="Dulong" pageId="32" pageNumber="373" stateProvince="Yunnan">Dulong</location>
divide area,
<geoCoordinate id="EE4B504CFFB5FFA93CE266D4FEF671ED" box="[181,312,1701,1725]" degrees="27.69655" direction="north" orientation="latitude" pageId="32" pageNumber="373" precision="1" value="27.69655">N27.69655°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB5FFA93D1766D4FE0E71ED" box="[320,448,1701,1725]" degrees="98.45389" direction="east" orientation="longitude" pageId="32" pageNumber="373" precision="1" value="98.45389">E98.45389°</geoCoordinate>
,
<quantity id="4C879B6EFFB5FFA93D9C66D4FD9071ED" box="[459,606,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.4899999999999998" metricValueMax="3.6799999999999997" metricValueMin="3.3" pageId="32" pageNumber="373" unit="m" value="3490.0" valueMax="3680.0" valueMin="3300.0">
<elevation id="0052D1B8FFB5FFA93D9C66D4FD9071ED" box="[459,606,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="3.4899999999999998" metricValueMax="3.6799999999999997" metricValueMin="3.3" pageId="32" pageNumber="373" unit="m" value="3490.0" valueMax="3680.0" valueMin="3300.0">3300-3680 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<caption id="DF006603FFB4FFA83CFC6658FD54713E" ID-DOI="http://doi.org/10.5281/zenodo.13155309" ID-Zenodo-Dep="13155309" httpUri="https://zenodo.org/record/13155309/files/figure.png" pageId="33" pageNumber="374" startId="33.[171,183,1577,1596]" targetBox="[144,1175,216,1563]" targetPageId="33" targetType="figure">
<paragraph id="8BC0368BFFB4FFA83CFC6658FD54713E" blockId="33.[123,1173,1576,1647]" pageId="33" pageNumber="374">
<smallCapsWord id="8D26A057FFB4FFA83CFC6658FF26716A" baselines="1591,1590" box="[171,232,1577,1596]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="33" pageNumber="374">FIGURE</smallCapsWord>
14.
<taxonomicName id="4C7F4D08FFB4FFA83D5A6659FDB8716B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[269,630,1576,1595]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="374" phylum="Arthropoda" rank="species" species="balli" status="sp. nov." subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFB4FFA83D5A6659FE53716B" box="[269,413,1576,1595]" italics="true" pageId="33" pageNumber="374">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FFB4FFA83DFE6659FD8C716B" box="[425,578,1576,1595]" italics="true" pageId="33" pageNumber="374">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FFB4FFA83E186659FDB8716B" box="[591,630,1576,1595]" italics="true" pageId="33" pageNumber="374">balli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFB4FFA83E2C6658FD0E716C" box="[635,704,1577,1596]" pageId="33" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFB4FFA838416633FB977105" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1046,1113,1602,1621]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="374" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B90BEA99FFB4FFA838416633FB977105" box="[1046,1113,1602,1621]" italics="true" pageId="33" pageNumber="374">Q. balli</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFB7FFAB3CFC60AAFBA47604" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">
<emphasis id="B90BEA99FFB7FFAB3CFC60AAFE2177A1" bold="true" box="[171,495,219,243]" pageId="34" pageNumber="375">
<smallCapsWord id="8D26A057FFB7FFAB3CFC60AAFEE077A1" baselines="237,236" box="[171,302,219,243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="34" pageNumber="375">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFB7FFAB3D6260AFFE9F77A1" baselines="236" box="[309,337,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="34" pageNumber="375">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFB7FFAB3D0060AFFE6477A1" baselines="236" box="[343,426,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="34" pageNumber="375">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFB7FFAB3DE560AFFE2177A1" baselines="236" box="[434,495,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="34" pageNumber="375">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFB7FFAB3EA560AAFCED77A3" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[754,803,219,243]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="375" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B90BEA99FFB7FFAB3EA560AAFCED77A3" box="[754,803,219,243]" italics="true" pageId="34" pageNumber="375">balli</emphasis>
</taxonomicName>
, is the Latinized form (in the genitive case) of the surname of George Eugene Ball, Professor Emeritus at the University of
<collectingRegion id="49BBF869FFB7FFAB3868608AFB5C7643" box="[1087,1170,251,275]" country="Canada" name="Alberta" pageId="34" pageNumber="375">Alberta</collectingRegion>
, Edmonton,
<collectingRegion id="49BBF869FFB7FFAB3CA1616AFE877663" box="[246,329,283,307]" country="Canada" name="Alberta" pageId="34" pageNumber="375">Alberta</collectingRegion>
,
<collectingCountry id="F368761BFFB7FFAB3D03616AFE6A7663" box="[340,420,283,307]" name="Canada" pageId="34" pageNumber="375">Canada</collectingCountry>
— our mentor, dear friend, and one of the worlds most accomplished and inspirational systematists. We are pleased to name this extraordinary species in his honor.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFB7FFAB3CFC612DFF0E76E5" pageId="34" pageNumber="375" type="diagnosis">
<paragraph id="8BC0368BFFB7FFAB3CFC612DFF0E76E5" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">
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<emphasis id="B90BEA99FFB7FFAB3CFC612DFEEE7622" bold="true" box="[171,288,348,372]" pageId="34" pageNumber="375">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFB7FFAB3E60612DFD5F7624" box="[567,657,348,372]" captionStart="FIGURE" captionStartId="33.[171,183,1577,1596]" captionTargetBox="[144,1175,216,1563]" captionTargetId="figure-15@33.[120,1175,216,1563]" captionTargetPageId="33" captionText="FIGURE 14. Queinnectrechus (Gaoligongtrechus) balli sp. nov.; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for Q. balli in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155309" httpUri="https://zenodo.org/record/13155309/files/figure.png" pageId="34" pageNumber="375">Fig. 14a</figureCitation>
) can be distinguished from those of all other trechine species in the region by the combination of character states noted in the diagnosis for this genus.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFB7FFAB3CFC61CCFDEB71ED" pageId="34" pageNumber="375" type="description">
<paragraph id="8BC0368BFFB7FFAB3CFC61CCFDAF7545" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">
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<emphasis id="B90BEA99FFB7FFAB3CFC61CCFEF47683" bold="true" box="[171,314,445,469]" pageId="34" pageNumber="375">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size moderate, BL =
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. Color of dorsum dark, forebody dark reddish brown, elytra piceous to black, antennae and legs reddish brown, palpi slightly paler, reddish tan; surface smooth and markedly shiny.
</paragraph>
<paragraph id="8BC0368BFFB7FFAB3CFC626CFCF574E8" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">
Head. Slightly elongate, with eyes small but convex, their diameter less than length of tempora. Tempora not or only slightly convex, glabrous, and joined to neck region at a markedly obtuse (ca. 150°) angle. Frons with deep frontal furrow that delimit three (two lateral and medial) very convex areas, however furrows abruptly interrupted or less impressed posteriorly; two pairs of supraorbital seta present, the anterior pair inserted near middle of eyes, the posterior pair inserted dorsally on the tempora near the postocular furrow. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible tridentate, with the middle tooth reduced, obtuse, left mandible with premolar tooth fused with retinaculum to form a small caniniform process with a sharp tip.
<collectingRegion id="49BBF869FFB7FFAB3DDB636EFE287466" box="[396,486,799,822]" country="Myanmar" name="Chin" pageId="34" pageNumber="375">Mentum</collectingRegion>
and submentum fused.
<collectingRegion id="49BBF869FFB7FFAB3EBC636EFC8B7466" box="[747,837,799,822]" country="Myanmar" name="Chin" pageId="34" pageNumber="375">Mentum</collectingRegion>
with medial tooth broad, truncate, about half as long as lateral lobes, the latter apically pointed. Submentum with six setae anteriorly. Genae with a single ventral seta on each side. Antennae pubescent from apical half of scape distally, extended posteriorly to basal one-third of elytra, with four antennomeres posterior to the pronotal base, antennomere 3 slightly longer than antennomere 4.
</paragraph>
<paragraph id="8BC0368BFFB7FFAB3CFC63B1FDCA73E9" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">Pronotum. Narrowly cordate, not transverse, about as long as wide (PW/PL = 1.0), markedly narrowed posteriorly, the lateral margins distinctly sinuate anterior to basal angles, the latter acute, projected, but without digitiform extensions. Disc markedly convex, globose, glabrous; median longitudinal impression faintly impressed; basal foveae formed as small, deep, circular pits, medi- an basal area smooth, faintly delimited, basal margin dilated medially as a broad, round projection. Lateral margination narrow, evident only in anterior one-third, effaced in posterior two-thirds. Two or three anteromedial setae and a single basal seta present on each side, the latter inserted slightly but distinctly anterior to basal angle.</paragraph>
<paragraph id="8BC0368BFFB7FFAB3CFC64B3FE77728B" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">Elytra. Elytral silhouette tear-shaped, narrower anteriorly than posteriorly, widest distinctly posterior to middle, humeri effaced, basal part of lateral explanation not visible from above. Disc markedly convex and smooth, with basal part abruptly and truncate, concave, fitted to convex base of pronotum, without distinct discal striae, except recurrent stria short but evidently impressed. Parascutellar setiferous pore present. Four to six discal setae present and aligned near presumed location of stria 3. Preapical seta present, inserted slightly more medially than the row of discal setae. Umbilicate setal series with setae of humeral group aggregated with distance between first and second setae less than between second and third and third and fourth, setae of median group distinctly posterior to middle.</paragraph>
<paragraph id="8BC0368BFFB7FFAB3CFC6595FBFB714C" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">Legs. Moderately long but slender; protibiae with longitudinal furrow, sparely pubescent apically on anterior surface. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFB7FFAB3CFC6655FD68710C" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side and ventrite VII of females with two setae on each side.</paragraph>
<paragraph id="8BC0368BFFB7FFAB3CFC6615FDEB71ED" blockId="34.[123,1174,219,1725]" pageId="34" pageNumber="375">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFB7FFAB3DBF6615FD8C712C" box="[488,578,1636,1660]" captionStart="FIGURE" captionStartId="33.[171,183,1577,1596]" captionTargetBox="[144,1175,216,1563]" captionTargetId="figure-15@33.[120,1175,216,1563]" captionTargetPageId="33" captionText="FIGURE 14. Queinnectrechus (Gaoligongtrechus) balli sp. nov.; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for Q. balli in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155309" httpUri="https://zenodo.org/record/13155309/files/figure.png" pageId="34" pageNumber="375">Fig. 14b</figureCitation>
) slender, bent basally about 90° to shaft, basal bulb with a large sagittal aileron, apex short and slightly narrowed, blunt; endophallus with a pair of slender sclerites tapered to points on both ends.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFB6FFAA3CFC60AAFC5F7233" pageId="35" pageNumber="376" type="discussion">
<paragraph id="8BC0368BFFB6FFAA3CFC60AAFEFB75AA" blockId="35.[123,1174,219,1379]" pageId="35" pageNumber="376">
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<emphasis id="B90BEA99FFB6FFAA3CFC60AAFEE677A1" bold="true" box="[171,296,219,243]" pageId="35" pageNumber="376">COMMENTS</emphasis>
</smallCapsWord>
.— Members of this new genus exhibit practically the same chaetotaxic pattern as those of genus
<taxonomicName id="4C7F4D08FFB6FFAA3D4A608AFDFA7643" authority="Jeannel (1935)" authorityName="Jeannel" authorityYear="1935" box="[285,564,251,275]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3D4A608AFE427643" box="[285,396,251,275]" italics="true" pageId="35" pageNumber="376">Kozlovites</emphasis>
<bibRefCitation id="EFEE4B7AFFB6FFAA3DC4608AFDFA7643" author="JEANNEL, R." box="[403,564,251,275]" pageId="35" pageNumber="376" pagination="273 - 282" refId="ref59618" refString="JEANNEL, R. 1935. Sur quelques Trechinae de l'Asie Centrale. Revue Francaise d'Entomologie 1: 273 - 282." type="journal article" year="1935">Jeannel (1935)</bibRefCitation>
</taxonomicName>
, with a row of four to six discal setae apparently aligned on interval
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or near stria 3 and a preapical seta apparently inserted on interval 3 near stria 2 but in a position forward of the level of the anterior tip of the recurrent stria. However,
<taxonomicName id="4C7F4D08FFB6FFAA3C2C612CFEF57625" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,315,349,373]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="subGenus" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFB6FFAA3C2C612CFEF57625" box="[123,315,349,373]" italics="true" pageId="35" pageNumber="376">Gaoligongtrechus</emphasis>
</taxonomicName>
members are distinguished from those of
<taxonomicName id="4C7F4D08FFB6FFAA3F57612CFCA17625" authorityName="Jeannel" authorityYear="1935" box="[768,879,349,373]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3F57612CFCA17625" box="[768,879,349,373]" italics="true" pageId="35" pageNumber="376">Kozlovites</emphasis>
</taxonomicName>
(and of
<taxonomicName id="4C7F4D08FFB6FFAA3F9D612FFEE376C5" authority="Belousov and Kabak 2016" authorityName="Belousov and Kabak" authorityYear="2016" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3F9D612FFBE77625" box="[970,1065,350,373]" italics="true" pageId="35" pageNumber="376">Uenoites</emphasis>
Belousov and Kabak 2016
</taxonomicName>
) by the extreme convexity of both the pronotum and elytra, which are of similar form to that seen in members of
<taxonomicName id="4C7F4D08FFB6FFAA3DB261EFFD5776E6" authorityName="Deuve" authorityYear="1992" box="[485,665,414,438]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3DB261EFFD5776E6" box="[485,665,414,438]" italics="true" pageId="35" pageNumber="376">Queinnectrechus</emphasis>
</taxonomicName>
(
<emphasis id="B90BEA99FFB6FFAA3EFC61EFFD2B76E5" box="[683,741,414,437]" italics="true" pageId="35" pageNumber="376">s. str.</emphasis>
), and by the posterior projection of the pronotal basal area and margin and its fit with the modified elytral base. Additional distinguishing features include the lateral marginations, each of which bears one or two anteromedial setae and is effaced in the posterior two-thirds, the effaced elytral discal striae, the medial tooth of the mentum truncate rather than bifid, the abdominal ventrites each with only a single pair of paramedial setae, except for the female ventrite VII which has two pairs of subapical paramedial setae, as is typical among
<taxonomicName id="4C7F4D08FFB6FFAA3C9C6211FEE67528" authorityName="Bonelli" authorityYear="1810" box="[203,296,608,632]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
. In addition, Belousov and Kabak (2016) considered the apex of the male median lobe formed as a large apical hook as a synapomorphy for
<taxonomicName id="4C7F4D08FFB6FFAA3EB762F0FC6775C9" box="[736,937,641,665]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90BEA99FFB6FFAA3EB762F0FC8175C9" box="[736,847,641,665]" italics="true" pageId="35" pageNumber="376">Kozlovites</emphasis>
species.
</taxonomicName>
The apex of the medi- an lobe of
<taxonomicName id="4C7F4D08FFB6FFAA3CB862D3FEBB75E9" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[239,373,673,698]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="balli" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFB6FFAA3CB862D3FEC775E9" box="[239,265,674,697]" italics="true" pageId="35" pageNumber="376">Q.</emphasis>
(
<emphasis id="B90BEA99FFB6FFAA3D4E62D3FEFD75E9" box="[281,307,674,697]" italics="true" pageId="35" pageNumber="376">G.</emphasis>
)
<emphasis id="B90BEA99FFB6FFAA3D1362D0FEBB75E9" box="[324,373,673,697]" italics="true" pageId="35" pageNumber="376">balli</emphasis>
</taxonomicName>
males (
<figureCitation id="13442A0EFFB6FFAA3D9D62D0FDEB75E9" box="[458,549,673,697]" captionStart="FIGURE" captionStartId="33.[171,183,1577,1596]" captionTargetBox="[144,1175,216,1563]" captionTargetId="figure-15@33.[120,1175,216,1563]" captionTargetPageId="33" captionText="FIGURE 14. Queinnectrechus (Gaoligongtrechus) balli sp. nov.; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for Q. balli in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155309" httpUri="https://zenodo.org/record/13155309/files/figure.png" pageId="35" pageNumber="376">Fig. 14b</figureCitation>
) has no trace of a hook; and the slender shaft and abruptly bent (at a 90° angle) basal region are unlike that seen in males of any described
<taxonomicName id="4C7F4D08FFB6FFAA385F62B3FBB9758A" authorityName="Jeannel" authorityYear="1935" box="[1032,1143,706,730]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA385F62B3FBB9758A" box="[1032,1143,706,730]" italics="true" pageId="35" pageNumber="376">Kozlovites</emphasis>
</taxonomicName>
or
<taxonomicName id="4C7F4D08FFB6FFAA3C2C6292FEFB75AA" box="[123,309,738,762]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90BEA99FFB6FFAA3C2C6292FF1475AA" box="[123,218,739,762]" italics="true" pageId="35" pageNumber="376">Uenoites</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8BC0368BFFB6FFAA3CFC6372FDD974ED" blockId="35.[123,1174,219,1379]" pageId="35" pageNumber="376">
Members of subgenus
<taxonomicName id="4C7F4D08FFB6FFAA3DE66372FDBF744B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[433,625,771,795]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="subGenus" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFB6FFAA3DE66372FDBF744B" box="[433,625,771,795]" italics="true" pageId="35" pageNumber="376">Gaoligongtrechus</emphasis>
</taxonomicName>
differ from those of subgenus
<taxonomicName id="4C7F4D08FFB6FFAA3FB56372FF0F746B" authority="Deuve" authorityName="Deuve" authorityYear="1992" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3FB56372FB58744B" box="[994,1174,771,795]" italics="true" pageId="35" pageNumber="376">Queinnectrechus</emphasis>
Deuve
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFB6FFAA3CA16352FD17746B" authority="Belousov and Kabak, (2003)" authorityName="Belousov and Kabak" authorityYear="2003" box="[246,729,803,827]" class="Insecta" family="Carabidae" genus="Dactylotrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3CA16352FE56746B" box="[246,408,803,827]" italics="true" pageId="35" pageNumber="376">Dactylotrechus</emphasis>
Belousov and Kabak, (2003)
</taxonomicName>
, in having the basal pronotal angles simple, without the digitiform extensions seen in members of these two genera. Like
<taxonomicName id="4C7F4D08FFB6FFAA3FA46335FB5B740C" authorityName="Belousov and Kabak" authorityYear="2003" box="[1011,1173,836,860]" class="Insecta" family="Carabidae" genus="Dactylotrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFB6FFAA3FA46335FB5B740C" box="[1011,1173,836,860]" italics="true" pageId="35" pageNumber="376">Dactylotrechus</emphasis>
</taxonomicName>
members, those of our new subgenus have anterolateral setae in the pronotal margins and a preapical seta apparently inserted near stia 2; however, this seta is inserted farther forward, in a subdiscal position, in
<taxonomicName id="4C7F4D08FFB6FFAA3D7663D4FE6874ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[289,422,933,957]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="balli" subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FFB6FFAA3D7663D4FEF574EC" box="[289,315,933,956]" italics="true" pageId="35" pageNumber="376">Q.</emphasis>
(
<emphasis id="B90BEA99FFB6FFAA3D1C63D4FEAB74EC" box="[331,357,933,956]" italics="true" pageId="35" pageNumber="376">G.</emphasis>
)
<emphasis id="B90BEA99FFB6FFAA3D2263D4FE6874ED" box="[373,422,933,957]" italics="true" pageId="35" pageNumber="376">balli</emphasis>
</taxonomicName>
members.
</paragraph>
<paragraph id="8BC0368BFFB6FFAA3CFC63B7FF0073F0" blockId="35.[123,1174,219,1379]" pageId="35" pageNumber="376">
<emphasis id="B90BEA99FFB6FFAA3CFC63B7FE68748C" bold="true" box="[171,422,966,990]" pageId="35" pageNumber="376">
<smallCapsWord id="8D26A057FFB6FFAA3CFC63B7FEC7748C" baselines="984,983" box="[171,265,966,990]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="35" pageNumber="376">HABITAT</smallCapsWord>
<smallCapsWord id="8D26A057FFB6FFAA3D4563B8FE68748C" baselines="983" box="[274,422,969,988]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="35" pageNumber="376">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found under stones on moist, organic substrate in alpine meadows, slopes and ridges with low, dense to sparse herbaceous vegetation, at elevations ranging from
<quantity id="4C879B6EFFB6FFAA3DAE6477FD69734E" box="[505,679,1030,1054]" metricMagnitude="3" metricUnit="m" metricValue="3.525" metricValueMax="3.75" metricValueMin="3.3" pageId="35" pageNumber="376" unit="m" value="3525.0" valueMax="3750.0" valueMin="3300.0">3300 to 3750 m</quantity>
(
<figureCitation id="13442A0EFFB6FFAA3EE06477FCDB734E" box="[695,789,1030,1054]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="35" pageNumber="376">Fig. 37b</figureCitation>
,
<figureCitation id="13442A0EFFB6FFAA3F756477FC82734E" box="[802,844,1030,1054]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="35" pageNumber="376">39b</figureCitation>
,
<figureCitation id="13442A0EFFB6FFAA3F0E6477FC4A734E" box="[857,900,1030,1054]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="35" pageNumber="376">40a</figureCitation>
).
<specimenCount id="9D79FD02FFB6FFAA3FCF6477FBFC734E" box="[920,1074,1030,1054]" count="1" pageId="35" pageNumber="376" type="generic">One specimen</specimenCount>
was collected at night, found walking on the barren slope of a roadcut at
<quantity id="4C879B6EFFB6FFAA3F106456FC57736E" box="[839,921,1063,1087]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="35" pageNumber="376" unit="m" value="3350.0">3350 m</quantity>
elevation. Members of this species have been found together (syntopic) repeatedly with specimens of
<taxonomicName id="4C7F4D08FFB6FFAA3FB56436FEC073D0" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="gongshanicus">
<emphasis id="B90BEA99FFB6FFAA3FB56436FEC073D0" italics="true" pageId="35" pageNumber="376">Queinnectrechus gongshanicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFB6FFAA3D4B6419FDF273D0" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[284,572,1128,1152]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FFB6FFAA3D4B6419FDF273D0" box="[284,572,1128,1152]" italics="true" pageId="35" pageNumber="376">Queinnectrechus griswoldi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFB6FFAA3E1C6419FC8D73D0" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[587,835,1128,1152]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FFB6FFAA3E1C6419FC8D73D0" box="[587,835,1128,1152]" italics="true" pageId="35" pageNumber="376">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFB6FFAA3F1C6419FC6F732F" box="[843,929,1128,1151]" pageId="35" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C7F4D08FFB6FFAA3F8C6419FB58732F" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[987,1174,1128,1152]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="376" phylum="Arthropoda" rank="species" species="qiqiensis" status="sp. nov.">
<emphasis id="B90BEA99FFB6FFAA3F8C6419FB58732F" box="[987,1174,1128,1152]" italics="true" pageId="35" pageNumber="376">Trechus qiqiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFB6FFAA3C2C64F8FF0073F0" box="[123,206,1161,1184]" pageId="35" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFFB6FFAA3CFC64D9FC927272" blockId="35.[123,1174,219,1379]" pageId="35" pageNumber="376">
<emphasis id="B90BEA99FFB6FFAA3CFC64D9FD4E73EF" bold="true" box="[171,640,1192,1216]" pageId="35" pageNumber="376">
<smallCapsWord id="8D26A057FFB6FFAA3CFC64D9FE9873EF" baselines="1210,1210" box="[171,342,1192,1216]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="35" pageNumber="376">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFB6FFAA3D0864DDFE3D73EF" baselines="1210" box="[351,499,1196,1215]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="35" pageNumber="376">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFB6FFAA3DAA64DDFD8273EF" baselines="1210" box="[509,588,1196,1215]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="35" pageNumber="376">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFB6FFAA3E0164DDFD4E73EF" baselines="1210" box="[598,640,1196,1215]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="35" pageNumber="376">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FFB6FFAA3EDD64D9FCC173EF" baselines="1210,1210" box="[650,783,1192,1216]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="35" pageNumber="376">
<emphasis id="B90BEA99FFB6FFAA3EDD64D9FCC173EF" bold="true" box="[650,783,1192,1216]" pageId="35" pageNumber="376">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFB6FFAA3F4E64D9FB9D73B1" location="We" pageId="35" pageNumber="376" specimenCount="66" specimenCount-female="21" specimenCount-male="12" stateProvince="Shan">
<smallCapsWord id="8D26A057FFB6FFAA3F4E64D9FC9A73EF" baselines="1210,1210" box="[793,852,1192,1216]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="35" pageNumber="376">
<emphasis id="B90BEA99FFB6FFAA3F4E64D9FC9A73EF" bold="true" box="[793,852,1192,1216]" pageId="35" pageNumber="376">
<collectingRegion id="49BBF869FFB6FFAA3F4E64D9FC9A73EF" box="[793,852,1192,1216]" country="Myanmar" name="Shan" pageId="35" pageNumber="376">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFB6FFAA3FD764D9FC107391" box="[896,990,1192,1217]" captionStart="FIGURE" captionStartId="33.[171,183,1577,1596]" captionTargetBox="[144,1175,216,1563]" captionTargetId="figure-15@33.[120,1175,216,1563]" captionTargetPageId="33" captionText="FIGURE 14. Queinnectrechus (Gaoligongtrechus) balli sp. nov.; a. Dorsal habitus (CASENT1001935). b. Median lobe of aedeagus of male (CASENT1001935), left lateral aspect. c. Map of locality records (red circles) for Q. balli in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155309" httpUri="https://zenodo.org/record/13155309/files/figure.png" pageId="35" pageNumber="376">Fig. 14c</figureCitation>
.
<location id="8EA06050FFB6FFAA3FBC64D8FBC07390" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB6FFAA3FBC64D8FBC07390" box="[1003,1038,1193,1216]" name="We" pageId="35" pageNumber="376" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFB6FFAA3C9A64B8FE9073B1" box="[205,350,1225,1249]" count="33" pageId="35" pageNumber="376" type="generic">33 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFB6FFAA3D3B64B8FE0273B1" box="[364,460,1225,1249]" count="12" pageId="35" pageNumber="376" type="male">12 males</specimenCount>
and
<specimenCount id="9D79FD02FFB6FFAA3E5764B8FDB673B1" box="[512,632,1225,1249]" count="21" pageId="35" pageNumber="376" type="female">21 females</specimenCount>
), all from the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFB6FFAA380E64B8FC967272" county="Dulongjiang Townships in Gongshan County" location="Cikai" municipality="Bingzhongluo" pageId="35" pageNumber="376" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFB6FFAA380E64B8FB5F73B1" box="[1113,1169,1225,1249]" country="Myanmar" name="Shan" pageId="35" pageNumber="376">Shan</collectingRegion>
, in
<collectingMunicipality id="6BA4ACF1FFB6FFAA3CC16498FEFE7251" box="[150,304,1257,1281]" pageId="35" pageNumber="376">Bingzhongluo</collectingMunicipality>
,
<location id="8EA06050FFB6FFAA3D6D6498FEBA7251" LSID="urn:lsid:plazi:treatment:03D6879DFFB5FFAA3C2C626AFC5F7233:8EA06050FFB6FFAA3D6D6498FEBA7251" box="[314,372,1257,1281]" county="Dulongjiang Townships in Gongshan County" municipality="Bingzhongluo" name="Cikai" pageId="35" pageNumber="376" stateProvince="Shan">Cikai</location>
and
<collectingCounty id="62A14E07FFB6FFAA3DF16498FC497252" box="[422,903,1257,1282]" pageId="35" pageNumber="376">Dulongjiang Townships in Gongshan County</collectingCounty>
(see
<typeStatus id="54C48829FFB6FFAA3FEA649BFC3C7252" box="[957,1010,1258,1282]" pageId="35" pageNumber="376">Type</typeStatus>
material above for exact collection data). These localities are in Core Areas 1 and 2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFB6FFAA3CFC655BFC5F7233" blockId="35.[123,1174,219,1379]" pageId="35" pageNumber="376">
<emphasis id="B90BEA99FFB6FFAA3CFC655BFDAF7211" bold="true" box="[171,609,1322,1346]" pageId="35" pageNumber="376">
<smallCapsWord id="8D26A057FFB6FFAA3CFC655BFEDC7211" baselines="1340,1340" box="[171,274,1322,1346]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="35" pageNumber="376">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFB6FFAA3D4A655FFE0A7211" baselines="1340" box="[285,452,1326,1345]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="35" pageNumber="376">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFB6FFAA3D9A655FFDAF7211" baselines="1340" box="[461,609,1326,1345]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="35" pageNumber="376">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFB6FFAA3ED3653AFCF17233" box="[644,831,1355,1379]" country="China" name="Yunnan" pageId="35" pageNumber="376">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFB6FFAA3F1D653AFC437233" box="[842,909,1355,1379]" name="China" pageId="35" pageNumber="376">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,512 @@
<document id="EBFDAD5D2A591822136276BB9DD8578D" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFB8FFB93C2C630AFC7975E3" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechus (Trechus) shiyueliang Deuve &amp; Kavanaugh &amp; Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="389" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="386" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="3A953E3BBC86C5E7B3B0AF64BB6092BE">
<mods:title id="A80CC62F627E14DFE0FC0CD0C6BB7A41">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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</mods:role>
<mods:namePart id="02CD0FE18B254AF249208FBF886A15AB">Deuve, Thierry</mods:namePart>
<mods:affiliation id="5726118C5E787AD6B99560B079CB1FCC">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="4B547ED049480B6F83627E25B49C66F0">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="81F2597E6387F3125A61EFAACAC3E6FA">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="D9CED34D3D35E27ABD06C0717CFE4932" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="78E23D9ED4A1EB40C196B3F2FF308669">Liang, Hongbin</mods:namePart>
<mods:affiliation id="78DE73288F3E342545CE6FD46E978859">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:date id="E9A2E87F36AF54113ECF4A67E525F29B">2016</mods:date>
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<subSubSection id="C3656500FFB8FFA43C2C630AFCF374C4" box="[123,829,891,916]" pageId="45" pageNumber="386" type="nomenclature">
<paragraph id="8BC0368BFFB8FFA43C2C630AFCF374C4" blockId="45.[123,829,891,946]" box="[123,829,891,916]" pageId="45" pageNumber="386">
<heading id="D08881E7FFB8FFA43C2C630AFCF374C4" box="[123,829,891,916]" fontSize="10" level="2" pageId="45" pageNumber="386" reason="4">
<emphasis id="B90BEA99FFB8FFA43C2C630AFCF374C4" bold="true" box="[123,829,891,916]" pageId="45" pageNumber="386">
<taxonomicName id="4C7F4D08FFB8FFA43C2C630AFDD274C4" ID-CoL="8MHS7" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,540,891,916]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="386" phylum="Arthropoda" rank="species" species="shiyueliang" subGenus="Trechus">
<emphasis id="B90BEA99FFB8FFA43C2C630AFF1A74C3" bold="true" box="[123,212,891,915]" italics="true" pageId="45" pageNumber="386">Trechus</emphasis>
(
<emphasis id="B90BEA99FFB8FFA43CB3630AFEF374C3" bold="true" box="[228,317,891,915]" italics="true" pageId="45" pageNumber="386">Trechus</emphasis>
)
<emphasis id="B90BEA99FFB8FFA43D19630AFE0374C3" bold="true" box="[334,461,891,915]" italics="true" pageId="45" pageNumber="386">shiyueliang</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFB8FFA43EB0630DFCF374C4" box="[743,829,892,916]" pageId="45" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFB8FFA43C2C63EDFE4D74E2" box="[123,387,924,946]" pageId="45" pageNumber="386" type="description">
<paragraph id="8BC0368BFFB8FFA43C2C63EDFE4D74E2" blockId="45.[123,829,891,946]" box="[123,387,924,946]" pageId="45" pageNumber="386">
(
<figureCitation id="13442A0EFFB8FFA43CD563EDFF1D74E1" box="[130,211,924,946]" captionStart="FIGURE" captionStartId="47.[171,183,1569,1588]" captionTargetBox="[120,1132,216,1549]" captionTargetId="figure-14@47.[120,1132,216,1549]" captionTargetPageId="47" captionText="FIGURE 18. Trechus shiyueliang sp. nov.; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for T. shiyueliang in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155318" httpUri="https://zenodo.org/record/13155318/files/figure.png" pageId="45" pageNumber="386">Figs. 18</figureCitation>
,
<figureCitation id="13442A0EFFB8FFA43C8A63EDFECD74E2" box="[221,259,924,946]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="45" pageNumber="386">40b</figureCitation>
,
<figureCitation id="13442A0EFFB8FFA43D5963EDFEFA74E2" box="[270,308,924,946]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="45" pageNumber="386">42b</figureCitation>
,
<figureCitation id="13442A0EFFB8FFA43D6963EDFEB274E2" box="[318,380,924,946]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="45" pageNumber="386">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFB8FFA73CFC63B4FD5B7643" lastPageId="46" lastPageNumber="387" pageId="45" pageNumber="386" type="materials_examined">
<paragraph id="8BC0368BFFB8FFA43CFC63B4FED9710D" blockId="45.[123,1174,965,1726]" pageId="45" pageNumber="386">
<emphasis id="B90BEA99FFB8FFA43CFC63B4FE97748B" bold="true" box="[171,345,965,989]" pageId="45" pageNumber="386">
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<smallCapsWord id="8D26A057FFB8FFA43CBB63B9FE97748B" baselines="982" box="[236,345,968,987]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="45" pageNumber="386">MATERIAL</smallCapsWord>
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.—
<materialsCitation id="3B173CD6FFB8FFA43DD563B4FF4A730D" collectingDate="2005-08-08" collectorName="H. B. Liang &amp; P. Paquin &amp; D. Z. Dong" country="China" county="Fugong County" elevation="3240" latitude="27.20654" location="Lishadi Township" longLatPrecision="1" longitude="98.71375" pageId="45" pageNumber="386" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFB8FFA43DD563B4FE26748D" box="[386,488,965,989]" pageId="45" pageNumber="386" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1021004”/ “
<collectingCountry id="F368761BFFB8FFA43FAB63B4FB9B748D" box="[1020,1109,965,989]" name="China" pageId="45" pageNumber="386">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FFB8FFA43CE66394FE9574AD" box="[177,347,997,1021]" pageId="45" pageNumber="386">Fugong County</collectingCounty>
,
<location id="8EA06050FFB8FFA43D306394FDE274AD" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFB8FFA43D306394FDE274AD" box="[359,556,997,1021]" country="China" county="Fugong County" latitude="27.20654" longLatPrecision="1" longitude="98.71375" name="Lishadi Township" pageId="45" pageNumber="386" stateProvince="Yunnan">Lishadi Township</location>
,
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<quantity id="4C879B6EFFB8FFA43E6F6394FD6E74AD" box="[568,672,997,1021]" metricMagnitude="4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="45" pageNumber="386" unit="km" value="10.5" valueMax="11.0" valueMin="10.0">10-11 km</quantity>
W of Shibali on Shibali Road
</locationDeviation>
,
<quantity id="4C879B6EFFB8FFA43FAA6394FB5F74AC" box="[1021,1169,997,1021]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" metricValueMax="3.28" metricValueMin="3.2" pageId="45" pageNumber="386" unit="m" value="3240.0" valueMax="3280.0" valueMin="3200.0">
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</quantity>
, NN27.19980°/
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to
<geoCoordinate id="EE4B504CFFB8FFA43D936475FD86734C" box="[452,584,1028,1052]" degrees="27.20654" direction="north" orientation="latitude" pageId="45" pageNumber="386" precision="1" value="27.20654">N27.20654°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB8FFA43E076475FD1F734C" box="[592,721,1028,1052]" degrees="98.71772" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.71772">E98.71772°</geoCoordinate>
,”/
<date id="FFC1104BFFB8FFA43EA46474FC50734C" box="[755,926,1028,1053]" pageId="45" pageNumber="386" value="2005-08-08">
<collectingDate id="EF85E9A3FFB8FFA43EA46474FC50734C" box="[755,926,1028,1053]" pageId="45" pageNumber="386" value="2005-08-08">8 August 2005</collectingDate>
</date>
, Stop#DHK-2005-068 D.H.
<taxonomicName id="4C7F4D08FFB8FFA43CEC6454FEF9736D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[187,311,1061,1085]" pageId="45" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFB8FFA43D1F6454FE06736D" box="[328,456,1061,1085]" pageId="45" pageNumber="386">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFB8FFA43D8E6454FD8B736D" box="[473,581,1061,1085]" pageId="45" pageNumber="386">P. Paquin</collectorName>
, &amp;
<collectorName id="268A535DFFB8FFA43E206454FD3E736D" box="[631,752,1061,1085]" pageId="45" pageNumber="386">D.Z. Dong</collectorName>
collectors”/ “IMAGE” [green label]/ “
</materialsCitation>
<typeStatus id="54C48829FFB8FFA43CD36434FEDD730D" box="[132,275,1093,1117]" pageId="45" pageNumber="386" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFB8FFA43D4A6434FD8B730C" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[285,581,1093,1117]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="386" phylum="Arthropoda" rank="species" species="shiyueliang">
<emphasis id="B90BEA99FFB8FFA43D4A6434FE3B730D" box="[285,501,1093,1117]" italics="true" pageId="45" pageNumber="386">Trechus shiyueliang</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFB8FFA43EA06434FC83730C" box="[759,845,1093,1116]" pageId="45" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label].
<materialsCitation id="3B173CD6FFB8FFA43C2C6414FF2973CD" collectionCode="IOZ" country="China" pageId="45" pageNumber="386" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFB8FFA43C2C6414FF2D732D" box="[123,227,1125,1149]" pageId="45" pageNumber="386" type="paratype">Paratypes</typeStatus>
(a total of 8):
<specimenCount id="9D79FD02FFB8FFA43D286414FE13732D" box="[383,477,1125,1149]" count="1" pageId="45" pageNumber="386" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB8FFA43E5E6414FDF9732D" box="[521,567,1125,1149]" pageId="45" pageNumber="386">IOZ</collectionCode>
) labeled same as
<typeStatus id="54C48829FFB8FFA43EA26414FC9B732D" box="[757,853,1125,1149]" pageId="45" pageNumber="386" type="holotype">holotype</typeStatus>
, except first label: “CASENT 1021005”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB8FFA43CA464F4FB5C738D" collectingDate="2005-08-16" collectionCode="IOZ" collectorName="P. Paquin" country="China" county="Fugong County" elevation="3221" latitude="27.20055" location="Lishadi Township" longLatPrecision="1" longitude="98.71399" pageId="45" pageNumber="386" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB8FFA43CA464F4FEF573CD" box="[243,315,1157,1181]" count="1" pageId="45" pageNumber="386" type="male">1 male</specimenCount>
and
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(in
<collectionCode id="ED6EAE4EFFB8FFA43DAD64F4FDE673CD" box="[506,552,1157,1181]" pageId="45" pageNumber="386">IOZ</collectionCode>
) labeled: “CASENT 1018374” and “CASENT 1018373”, respectively/ “
<collectingCountry id="F368761BFFB8FFA43D4464D4FEA273ED" box="[275,364,1189,1213]" name="China" pageId="45" pageNumber="386">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB8FFA43D2164D4FE0473EC" box="[374,458,1189,1212]" country="China" name="Yunnan" pageId="45" pageNumber="386">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB8FFA43D8364D4FDB273ED" box="[468,636,1189,1213]" pageId="45" pageNumber="386">Fugong County</collectingCounty>
,
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,
<locationDeviation id="9A722FA9FFB8FFA43F0564D4FF78738D" location="Shibali on Shibali Road" pageId="45" pageNumber="386">
<quantity id="4C879B6EFFB8FFA43F0564D4FC5B73ED" box="[850,917,1189,1213]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="386" unit="km" value="10.0">10 km</quantity>
W of Shibali on Shibali Road
</locationDeviation>
,
<quantity id="4C879B6EFFB8FFA43C9664B4FEDB738D" box="[193,277,1221,1245]" metricMagnitude="3" metricUnit="m" metricValue="3.221" pageId="45" pageNumber="386" unit="m" value="3221.0">
<elevation id="0052D1B8FFB8FFA43C9664B4FEDB738D" box="[193,277,1221,1245]" metricMagnitude="3" metricUnit="m" metricValue="3.221" pageId="45" pageNumber="386" unit="m" value="3221.0">3221 m</elevation>
</quantity>
,”/,
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/
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,
<date id="FFC1104BFFB8FFA43E0F64B4FCCB738D" box="[600,773,1221,1245]" pageId="45" pageNumber="386" value="2005-08-16">
<collectingDate id="EF85E9A3FFB8FFA43E0F64B4FCCB738D" box="[600,773,1221,1245]" pageId="45" pageNumber="386" value="2005-08-16">16 August 2005</collectingDate>
</date>
, Stop #PP-3805
<collectorName id="268A535DFFB8FFA43FEE64B4FBD3738D" box="[953,1053,1221,1245]" pageId="45" pageNumber="386">P. Paquin</collectorName>
collector”
</materialsCitation>
;
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<specimenCount id="9D79FD02FFB8FFA43C2C6494FF1373AD" box="[123,221,1253,1277]" count="1" pageId="45" pageNumber="386" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB8FFA43D466494FE8873AD" box="[273,326,1253,1277]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="45" pageNumber="386" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1023759”/ “
<collectingCountry id="F368761BFFB8FFA43EE96494FCD973AD" box="[702,791,1253,1277]" name="China" pageId="45" pageNumber="386">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB8FFA43F716494FCB473AC" box="[806,890,1253,1276]" country="China" name="Yunnan" pageId="45" pageNumber="386">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB8FFA43FDE6494FBF873AD" box="[905,1078,1253,1277]" pageId="45" pageNumber="386">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB8FFA438126494FF25724D" pageId="45" pageNumber="386">Lishadi Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FFB8FFA43D5F6574FCEE724D" box="[264,800,1285,1309]" location="Shibali on Shibali Road" pageId="45" pageNumber="386">
<quantity id="4C879B6EFFB8FFA43D5F6574FEA9724D" box="[264,359,1285,1309]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="45" pageNumber="386" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<location id="8EA06050FFB8FFA43F6A6574FB5C724C" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFB8FFA43F6A6574FB5C724C" box="[829,1170,1285,1309]" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" name="North Fork of Yamu He" pageId="45" pageNumber="386" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFB8FFA43C2C6554FF30726D" box="[123,254,1317,1341]" degrees="27.18416" direction="north" orientation="latitude" pageId="45" pageNumber="386" precision="1" value="27.18416">N27.18416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB8FFA43D506554FE49726D" box="[263,391,1317,1341]" degrees="98.72026" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFB8FFA43DF96554FDCA726C" box="[430,516,1317,1341]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="45" pageNumber="386" unit="m" value="3100.0">
<elevation id="0052D1B8FFB8FFA43DF96554FDCA726C" box="[430,516,1317,1341]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="45" pageNumber="386" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFB8FFA43E476554FD5D726D" box="[528,659,1317,1341]" pageId="45" pageNumber="386" value="2004-05-07">
<collectingDate id="EF85E9A3FFB8FFA43E476554FD5D726D" box="[528,659,1317,1341]" pageId="45" pageNumber="386" value="2004-05-07">7 May 2004</collectingDate>
</date>
Stop #DHK-2004-038A D.H.
<taxonomicName id="4C7F4D08FFB8FFA43F896554FB94726D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[990,1114,1317,1341]" pageId="45" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFB8FFA438316554FF2F720D" pageId="45" pageNumber="386">C.E. Griswold</collectorName>
, Liang H.-B., &amp; Zhu B.-X. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB8FFA43EC16534FF3C72ED" collectingDate="2005-08-12" collectionCode="IOZ" country="China" county="Fugong County" elevation="3740" latitude="27.20802" location="Lumadeng Township" longLatPrecision="1" longitude="98.69644" pageId="45" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB8FFA43EC16534FD2E720D" box="[662,736,1349,1373]" count="1" pageId="45" pageNumber="386" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB8FFA43F586534FCF3720D" box="[783,829,1349,1373]" pageId="45" pageNumber="386">IOZ</collectionCode>
) labeled: “CASENT 1020019”/ “
<collectingCountry id="F368761BFFB8FFA43CD26514FF2C722D" box="[133,226,1381,1405]" name="China" pageId="45" pageNumber="386">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB8FFA43CAF6514FE9E722C" box="[248,336,1381,1404]" country="China" name="Yunnan" pageId="45" pageNumber="386">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB8FFA43D306514FDEC722D" box="[359,546,1381,1405]" pageId="45" pageNumber="386">Fugong County</collectingCounty>
,
<location id="8EA06050FFB8FFA43E6D6514FCFD722D" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFB8FFA43E6D6514FCFD722D" box="[570,819,1381,1405]" country="China" county="Fugong County" latitude="27.20802" longLatPrecision="1" longitude="98.69644" name="Lumadeng Township" pageId="45" pageNumber="386" stateProvince="Yunnan">Lumadeng Township</location>
, ridge
<locationDeviation id="9A722FA9FFB8FFA43FCF6514FB5C722D" box="[920,1170,1381,1405]" location="Shibali Yakou" pageId="45" pageNumber="386">S of Shibali Yakou</locationDeviation>
,
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/
<geoCoordinate id="EE4B504CFFB8FFA43D5065F4FE4972CD" box="[263,391,1413,1437]" degrees="98.69644" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.69644">E98.69644°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFB8FFA43DE665F4FDC772CC" box="[433,521,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.7399999999999998" pageId="45" pageNumber="386" unit="m" value="3740.0">
<elevation id="0052D1B8FFB8FFA43DE665F4FDC772CC" box="[433,521,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.7399999999999998" pageId="45" pageNumber="386" unit="m" value="3740.0">3740 m</elevation>
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,
<date id="FFC1104BFFB8FFA43E4F65F4FD0472CD" box="[536,714,1413,1437]" pageId="45" pageNumber="386" value="2005-08-12">
<collectingDate id="EF85E9A3FFB8FFA43E4F65F4FD0472CD" box="[536,714,1413,1437]" pageId="45" pageNumber="386" value="2005-08-12">12 August 2005</collectingDate>
</date>
, Stop #DHK-2005-092 D. H.
<taxonomicName id="4C7F4D08FFB8FFA4384B65F4FB5B72CD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1052,1173,1413,1437]" pageId="45" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFB8FFA43CAA65D4FEDD710D" collectingDate="2005-08-13" collectionCode="CAS" collectorName="H. B. Liang &amp; D. Z. Dong &amp; G. Tang" country="China" county="Fugong County" elevation="3270" latitude="27.06429" location="Lao Shibali Yakou" longLatPrecision="1" longitude="98.75123" municipality="Lumadeng Township" pageId="45" pageNumber="386" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFB8FFA43CAA65D4FE8172ED" box="[253,335,1445,1469]" count="2" pageId="45" pageNumber="386" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FFB8FFA43DD365D4FE2F72ED" box="[388,481,1445,1469]" count="1" pageId="45" pageNumber="386" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFB8FFA43E5A65D4FD8C72ED" box="[525,578,1445,1469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="45" pageNumber="386" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1021323” to “CASENT 1021324” and “CASENT 1021325”, respectively/ “
<collectingCountry id="F368761BFFB8FFA43E6265B4FD40728D" box="[565,654,1477,1501]" name="China" pageId="45" pageNumber="386">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFB8FFA43ECE65B7FD23728D" box="[665,749,1478,1501]" country="China" name="Yunnan" pageId="45" pageNumber="386">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB8FFA43EAF65B7FC6F728D" box="[760,929,1477,1502]" pageId="45" pageNumber="386">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFB8FFA43FFA65B4FB5C728D" box="[941,1170,1477,1501]" pageId="45" pageNumber="386">Lumadeng Township</collectingMunicipality>
,
<location id="8EA06050FFB8FFA43C2C6597FE8B72AD" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFB8FFA43C2C6597FE8B72AD" box="[123,325,1509,1533]" country="China" county="Fugong County" latitude="27.06429" longLatPrecision="1" longitude="98.75123" municipality="Lumadeng Township" name="Lao Shibali Yakou" pageId="45" pageNumber="386" stateProvince="Yunnan">
Lao
<location id="8EA06050FFB8FFA43CFB6594FE8B72AD" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFB8FFA43CFB6594FE8B72AD" box="[172,325,1509,1533]" country="China" county="Fugong County" latitude="27.06429" longLatPrecision="1" longitude="98.75123" municipality="Lumadeng Township" name="Shibali Yakou" pageId="45" pageNumber="386" stateProvince="Yunnan">Shibali Yakou</location>
</location>
,
<quantity id="4C879B6EFFB8FFA43D066594FE6B72AD" box="[337,421,1509,1533]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="45" pageNumber="386" unit="m" value="3270.0">
<elevation id="0052D1B8FFB8FFA43D066594FE6B72AD" box="[337,421,1509,1533]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="45" pageNumber="386" unit="m" value="3270.0">3270 m</elevation>
</quantity>
,
<geoCoordinate id="EE4B504CFFB8FFA43DE66594FDFA72AD" box="[433,564,1509,1533]" degrees="27.06429" direction="north" orientation="latitude" pageId="45" pageNumber="386" precision="1" value="27.06429">N27.06429°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB8FFA43E6A6594FD7272AD" box="[573,700,1509,1533]" degrees="98.75123" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.75123">E98.75123°</geoCoordinate>
,
<date id="FFC1104BFFB8FFA43E9F6594FCB872AD" box="[712,886,1509,1533]" pageId="45" pageNumber="386" value="2005-08-13">
<collectingDate id="EF85E9A3FFB8FFA43E9F6594FCB872AD" box="[712,886,1509,1533]" pageId="45" pageNumber="386" value="2005-08-13">13 August 2005</collectingDate>
</date>
,”/ “Stop# DHK-2005-079, D.H.
<taxonomicName id="4C7F4D08FFB8FFA43CE06674FEFD714D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[183,307,1541,1565]" pageId="45" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFB8FFA43D686677FE74714D" box="[319,442,1541,1565]" pageId="45" pageNumber="386">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFB8FFA43D916677FDF3714E" box="[454,573,1542,1566]" pageId="45" pageNumber="386">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFB8FFA43E316674FD72714E" box="[614,700,1541,1566]" pageId="45" pageNumber="386">G. Tang</collectorName>
collectors”. All
<typeStatus id="54C48829FFB8FFA43F386677FC1B714E" box="[879,981,1542,1566]" pageId="45" pageNumber="386" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFB8FFA43D4D6654FE6F716D" box="[282,417,1573,1597]" pageId="45" pageNumber="386" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFB8FFA43DFD6654FD01716D" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[426,719,1573,1597]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="386" phylum="Arthropoda" rank="species" species="shiyueliang">
<emphasis id="B90BEA99FFB8FFA43DFD6654FD4F716D" box="[426,641,1573,1597]" italics="true" pageId="45" pageNumber="386">Trechus shiyueliang</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFB8FFA43F2A6657FC1C716D" box="[893,978,1574,1597]" pageId="45" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFB8FFA43CFC6614FDEE71CD" blockId="45.[123,1174,965,1726]" pageId="45" pageNumber="386">
<materialsCitation id="3B173CD6FFB8FFA43CFC6614FDD271CD" country="China" county="Fugong County" elevation="3240" location="Fugong County" pageId="45" pageNumber="386" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFB8FFA43CFC6614FE97712C" bold="true" box="[171,345,1637,1661]" pageId="45" pageNumber="386">
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<typeStatus id="54C48829FFB8FFA43CFC6614FF2A712C" box="[171,228,1637,1661]" pageId="45" pageNumber="386">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFB8FFA43CB96618FE97712C" baselines="1655" box="[238,345,1641,1660]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="45" pageNumber="386">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFB8FFA43DD46614FE0B712D" box="[387,453,1637,1661]" name="China" pageId="45" pageNumber="386">China</collectingCountry>
,
<collectingRegion id="49BBF869FFB8FFA43D856617FDE8712D" box="[466,550,1638,1661]" country="China" name="Yunnan" pageId="45" pageNumber="386">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFB8FFA43E646617FD10712D" box="[563,734,1637,1662]" pageId="45" pageNumber="386">Fugong County</collectingCounty>
,
<taxonomicName id="4C7F4D08FFB8FFA43EBC6614FC11712D" authority="Township" authorityName="Township" box="[747,991,1637,1661]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="386" phylum="Arthropoda" rank="species" species="shiyueliang">Shiyueliang Township</taxonomicName>
,
<locationDeviation id="9A722FA9FFB8FFA43FBB6614FEB371CD" location="Shibali on Shibali Road" pageId="45" pageNumber="386">
<quantity id="4C879B6EFFB8FFA43FBB6614FB9A712D" box="[1004,1108,1637,1661]" metricMagnitude="4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="45" pageNumber="386" unit="km" value="10.5" valueMax="11.0" valueMin="10.0">10-11 km</quantity>
W of Shibali on Shibali Road
</locationDeviation>
,
<quantity id="4C879B6EFFB8FFA43DDE66F4FDD271CD" box="[393,540,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" metricValueMax="3.28" metricValueMin="3.2" pageId="45" pageNumber="386" unit="m" value="3240.0" valueMax="3280.0" valueMin="3200.0">
<elevation id="0052D1B8FFB8FFA43DDE66F4FDD271CD" box="[393,540,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" metricValueMax="3.28" metricValueMin="3.2" pageId="45" pageNumber="386" unit="m" value="3240.0" valueMax="3280.0" valueMin="3200.0">3200-3280 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFB8FFA73CFC66D7FD5B7643" blockId="45.[123,1174,965,1726]" lastBlockId="46.[123,1174,219,1725]" lastPageId="46" lastPageNumber="387" pageId="45" pageNumber="386">
<emphasis id="B90BEA99FFB8FFA43CFC66D7FE3971EC" bold="true" box="[171,503,1702,1726]" pageId="45" pageNumber="386">
<smallCapsWord id="8D26A057FFB8FFA43CFC66D7FEE071EC" baselines="1720,1719" box="[171,302,1702,1726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="45" pageNumber="386">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFB8FFA43D6066D8FE9D71EC" baselines="1719" box="[311,339,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="45" pageNumber="386">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFB8FFA43D0B66D8FE6171EC" baselines="1719" box="[348,431,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="45" pageNumber="386">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFB8FFA43DED66D8FE3971EC" baselines="1719" box="[442,503,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="45" pageNumber="386">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFB8FFA43F5266D4FC4E71ED" box="[773,896,1701,1725]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="386" phylum="Arthropoda" rank="species" species="shiyueliang">
<emphasis id="B90BEA99FFB8FFA43F5266D4FC4E71ED" box="[773,896,1701,1725]" italics="true" pageId="45" pageNumber="386">shiyueliang</emphasis>
</taxonomicName>
, a noun in apposition, is derived from the name of the township in which the
<typeStatus id="54C48829FFBBFFA73EED60AAFD2677A3" box="[698,744,219,243]" pageId="46" pageNumber="387">type</typeStatus>
locality is located, Shiyueliang. This is the current name for the former Lishadi township.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBBFFA73CFC616DFCBD7545" pageId="46" pageNumber="387" type="diagnosis">
<paragraph id="8BC0368BFFBBFFA73CFC616DFCBD7545" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">
<smallCapsWord id="8D26A057FFBBFFA73CFC616DFEEE7662" baselines="302,301" box="[171,288,284,308]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="46" pageNumber="387">
<emphasis id="B90BEA99FFBBFFA73CFC616DFEEE7662" bold="true" box="[171,288,284,308]" pageId="46" pageNumber="387">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFBBFFA73E10616AFD6D7664" box="[583,675,283,308]" captionStart="FIGURE" captionStartId="47.[171,183,1569,1588]" captionTargetBox="[120,1132,216,1549]" captionTargetId="figure-14@47.[120,1132,216,1549]" captionTargetPageId="47" captionText="FIGURE 18. Trechus shiyueliang sp. nov.; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for T. shiyueliang in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155318" httpUri="https://zenodo.org/record/13155318/files/figure.png" pageId="46" pageNumber="387">Fig. 18a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size relatively large (BL = 4.0 to
<quantity id="4C879B6EFFBBFFA73C92612DFED07624" box="[197,286,348,372]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="46" pageNumber="387" unit="mm" value="4.2">4.2 mm</quantity>
), apterous; body color reddish-brown; head slightly elongate; tempora glabrous; pronotum moderately transverse (ratio PW/PL = 1.42); elytra with striae not or faintly punctate, recurrent stria continuous anteriorly with stria 5, two discal setae present and inserted next to stria 3, preapical seta present and inserted next to stria
<quantity id="4C879B6EFFBBFFA73ECB61CCFD087684" box="[668,710,444,469]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="46" pageNumber="387" unit="in" value="2.0">2 in</quantity>
forward position near apical one-fourth of elytra; median lobe of male aedeagus with apex thin and faintly reflexed dorsally in lateral view (
<figureCitation id="13442A0EFFBBFFA73CD4618CFF117545" box="[131,223,509,533]" captionStart="FIGURE" captionStartId="47.[171,183,1569,1588]" captionTargetBox="[120,1132,216,1549]" captionTargetId="figure-14@47.[120,1132,216,1549]" captionTargetPageId="47" captionText="FIGURE 18. Trechus shiyueliang sp. nov.; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for T. shiyueliang in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155318" httpUri="https://zenodo.org/record/13155318/files/figure.png" pageId="46" pageNumber="387">Fig. 18b</figureCitation>
), endophallus with only a single, small thin elongate sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBBFFB93CFC626CFC7975E3" lastPageId="48" lastPageNumber="389" pageId="46" pageNumber="387" type="description">
<paragraph id="8BC0368BFFBBFFA73CFC626CFC167505" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">
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<emphasis id="B90BEA99FFBBFFA73CFC626CFEF47564" bold="true" box="[171,314,541,565]" pageId="46" pageNumber="387">DESCRIPTION</emphasis>
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.— Size relatively large, BL = 4.0 to
<quantity id="4C879B6EFFBBFFA73E9B626CFCEC7565" box="[716,802,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="46" pageNumber="387" unit="mm" value="4.2">4.2 mm</quantity>
. Body color reddish-brown, shiny, appendages slightly paler. Body surface smooth, head capsule finely alutaceous.
</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC622FFCD574E8" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">
Head. Moderate in size, slightly elongate and thick; eyes small and moderately convex, their diameter about 1.5 times as long as tempora, the latter short, moderately convex and glabrous. Frons slightly flattened; frontal furrows deep, rounded, slightly attenuated posterior to the eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Mandibles short; right mandible tridentate with middle tooth closer to basal tooth (premolar) than to distal tooth.
<collectingRegion id="49BBF869FFBBFFA73FFE628EFBCD7446" box="[937,1027,767,790]" country="Myanmar" name="Chin" pageId="46" pageNumber="387">Mentum</collectingRegion>
and submentum not fully fused but nearly so, suture between them only faintly impressed.
<collectingRegion id="49BBF869FFBBFFA73F96636EFBD57466" box="[961,1051,799,822]" country="Myanmar" name="Chin" pageId="46" pageNumber="387">Mentum</collectingRegion>
with medial tooth apically truncate, less than one-half the length of the lateral lobes. Submentum with six setae anteriorly, swollen anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae short, with less than two antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 3 and 4 slightly longer than antennomere 2.
</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC63B0FB4073E9" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">Pronotum. Moderately transverse (ratio PW/PL = 1.42), narrowed posteriorly, greatest width near anterior one-third; lateral margins markedly rounded, straightened only just anterior to small, rectangular and sharp basal angles. Disc convex, smooth and glabrous, median longitudinal impression deeply impressed, interrupted anteriorly a short distance from anterior margin; basal foveae distinct; median basal area with several faint, irregular, punctiform depressions, delimited laterally by oblique furrows. Lateral borders of pronotum moderately slender, regular, slightly reflexed, lateral grooves deeply impressed. Single midlateral setae on each side inserted at anteri- or one-third; single basolateral seta on each side, inserted slightly anterior to apex of basal angle.</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC64B3FC60722B" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">
Elytra. Ovoid, humeri distinct but rounded. Disc convex and smooth. Striae not or only faintly punctate, striae 1 to 5 deeply impressed, striae 6 to 8 very faintly impressed but evident. Parascutellar striole present. Recurrent stria continuous anteriorly with stria 5. Basal setiferous pore present at common origin of striae 1 and 2. Two discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of elytra. Preapical seta present and inserted on interval 3 next to stria
<quantity id="4C879B6EFFBBFFA73D386512FE56722B" box="[367,408,1379,1403]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="46" pageNumber="387" unit="in" value="2.0">2 in</quantity>
forward position near apical one-fourth of elytra.
</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC65F2FD5372EB" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">Umbilicate setal series with setae of humeral group equidistance from each other and setae of median group inserted slightly posterior to middle.</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC65B5FE9E72AC" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC6675FB85716C" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFBBFFA73CFC6635FEFF71ED" blockId="46.[123,1174,219,1725]" pageId="46" pageNumber="387">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFBBFFA73DBA6635FD87710C" box="[493,585,1604,1628]" captionStart="FIGURE" captionStartId="47.[171,183,1569,1588]" captionTargetBox="[120,1132,216,1549]" captionTargetId="figure-14@47.[120,1132,216,1549]" captionTargetPageId="47" captionText="FIGURE 18. Trechus shiyueliang sp. nov.; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for T. shiyueliang in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155318" httpUri="https://zenodo.org/record/13155318/files/figure.png" pageId="46" pageNumber="387">Fig. 18b</figureCitation>
) moderately broad basally with sagittal aileron present and moderate in size; shaft gradually narrowed toward apex; apical lamella short, rectilinear, extremely thin and slightly recurved dorsally in lateral view; endophallus with a single thin and elongate sclerite.
</paragraph>
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18.
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<emphasis id="B90BEA99FFBAFFA63D586651FE747163" box="[271,442,1568,1587]" italics="true" pageId="47" pageNumber="388">Trechus shiyueliang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBAFFA63D976650FDC97164" box="[448,519,1569,1588]" pageId="47" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for
<taxonomicName id="4C7F4D08FFBAFFA63F07664AFC07711D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[848,969,1594,1613]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="388" phylum="Arthropoda" rank="species" species="shiyueliang">
<emphasis id="B90BEA99FFBAFFA63F07664AFC07711D" box="[848,969,1594,1613]" italics="true" pageId="47" pageNumber="388">T. shiyueliang</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFA5FFB93CFC60AAFBBD76A3" blockId="48.[123,1174,219,691]" pageId="48" pageNumber="389">
<emphasis id="B90BEA99FFA5FFB93CFC60AAFE6A77A1" bold="true" box="[171,420,219,243]" pageId="48" pageNumber="389">
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<smallCapsWord id="8D26A057FFA5FFB93D4760AFFE6A77A1" baselines="236" box="[272,420,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="48" pageNumber="389">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in a variety of microhabitats at elevations ranging from
<quantity id="4C879B6EFFA5FFB93D86608AFDB17643" box="[465,639,251,275]" metricMagnitude="3" metricUnit="m" metricValue="3.42" metricValueMax="3.7399999999999998" metricValueMin="3.1" pageId="48" pageNumber="389" unit="m" value="3420.0" valueMax="3740.0" valueMin="3100.0">3100 to 3740 m</quantity>
. They have been collected in daytime from under stones in thickets of bamboo and
<taxonomicName id="4C7F4D08FFA5FFB93DBA616DFDF47663" box="[493,570,284,307]" class="Magnoliopsida" family="Rosaceae" genus="Prunus" kingdom="Plantae" order="Rosales" pageId="48" pageNumber="389" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA5FFB93DBA616DFDF47663" box="[493,570,284,307]" italics="true" pageId="48" pageNumber="389">Prunus</emphasis>
</taxonomicName>
, in rocky open areas cleared by snow avalanches (
<figureCitation id="13442A0EFFA5FFB9383D616AFF687603" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="48" pageNumber="389">Fig. 40b</figureCitation>
), in meadows adjacent to bamboo and
<taxonomicName id="4C7F4D08FFA5FFB93E06614AFD217603" box="[593,751,315,339]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="48" pageNumber="389" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA5FFB93E06614AFD217603" box="[593,751,315,339]" italics="true" pageId="48" pageNumber="389">Rhododendron</emphasis>
</taxonomicName>
thickets (
<figureCitation id="13442A0EFFA5FFB93F0C614AFC777603" box="[859,953,315,339]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="48" pageNumber="389">Fig. 42b</figureCitation>
) and along roadcuts through such thickets. They have also been collected at night, found walking on the substrate along roadcuts through areas of bamboo with an overstory of scattered
<taxonomicName id="4C7F4D08FFA5FFB93F63610AFCA176C3" box="[820,879,379,403]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="48" pageNumber="389" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFA5FFB93F63610AFCA176C3" box="[820,879,379,403]" italics="true" pageId="48" pageNumber="389">Abies</emphasis>
</taxonomicName>
trees. At higher elevations, they were the only trechine encountered in this area; but they were found syntopic with specimens of
<taxonomicName id="4C7F4D08FFA5FFB93CCF61CAFE917683" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[152,351,443,467]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="389" phylum="Arthropoda" rank="species" species="shibalicus" status="sp. nov.">
<emphasis id="B90BEA99FFA5FFB93CCF61CAFE917683" box="[152,351,443,467]" italics="true" pageId="48" pageNumber="389">Trechus shibalicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA5FFB93D3061CDFE727683" box="[359,444,444,467]" pageId="48" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FFA5FFB93D9061CAFD237683" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[455,749,443,467]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="389" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFA5FFB93D9061CAFD237683" box="[455,749,443,467]" italics="true" pageId="48" pageNumber="389">Trechepaphiopsis unipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA5FFB93EA261CDFC867683" box="[757,840,444,467]" pageId="48" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C7F4D08FFA5FFB93F2A61CAFF6976A3" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="389" phylum="Arthropoda" rank="species" species="unisetulosa" status="sp. nov.">
<emphasis id="B90BEA99FFA5FFB93F2A61CAFF6976A3" italics="true" pageId="48" pageNumber="389">Trechepaphiopsis unisetulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFA5FFB93CF961ADFECC76A3" box="[174,258,476,499]" pageId="48" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
at the upper limit of the altitudinal range of this last mentioned species (
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).
</paragraph>
<paragraph id="8BC0368BFFA5FFB93CFC618AFDD67524" blockId="48.[123,1174,219,691]" pageId="48" pageNumber="389">
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<smallCapsWord id="8D26A057FFA5FFB93D08618EFE3D7542" baselines="525" box="[351,499,511,530]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="48" pageNumber="389">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFA5FFB93DAA618EFD827542" baselines="525" box="[509,588,511,530]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="48" pageNumber="389">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFA5FFB93E01618EFD4E7542" baselines="525" box="[598,640,511,530]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="48" pageNumber="389">THE</smallCapsWord>
</emphasis>
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<emphasis id="B90BEA99FFA5FFB93EDD618AFCC17542" bold="true" box="[650,783,507,531]" pageId="48" pageNumber="389">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FFA5FFB93F4E618AFDE07503" county="Fugong County" location="We" pageId="48" pageNumber="389" specimenCount="18" specimenCount-female="4" specimenCount-male="5" stateProvince="Shan">
<smallCapsWord id="8D26A057FFA5FFB93F4E618AFC9A7542" baselines="525,525" box="[793,852,507,531]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="48" pageNumber="389">
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<collectingRegion id="49BBF869FFA5FFB93F4E618AFC9A7542" box="[793,852,507,531]" country="Myanmar" name="Shan" pageId="48" pageNumber="389">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFA5FFB93FD7618AFC137543" box="[896,989,507,531]" captionStart="FIGURE" captionStartId="47.[171,183,1569,1588]" captionTargetBox="[120,1132,216,1549]" captionTargetId="figure-14@47.[120,1132,216,1549]" captionTargetPageId="47" captionText="FIGURE 18. Trechus shiyueliang sp. nov.; a. Dorsal habitus (CASENT1021004). b. Median lobe of aedeagus of male (CASENT1021004), left lateral aspect. c. Map of locality record s(red circles) for T. shiyueliang in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155318" httpUri="https://zenodo.org/record/13155318/files/figure.png" pageId="48" pageNumber="389">Fig. 18c</figureCitation>
.
<location id="8EA06050FFA5FFB93FBC618DFBC07543" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFA5FFB93FBC618DFBC07543" box="[1003,1038,508,531]" county="Fugong County" name="We" pageId="48" pageNumber="389" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFA5FFB93C9B626AFE807563" box="[204,334,539,563]" count="9" pageId="48" pageNumber="389" type="generic">9 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFA5FFB93D0B626AFE607563" box="[348,430,539,563]" count="5" pageId="48" pageNumber="389" type="male">5 males</specimenCount>
and
<specimenCount id="9D79FD02FFA5FFB93DB7626AFD847563" box="[480,586,539,563]" count="4" pageId="48" pageNumber="389" type="female">4 females</specimenCount>
), all from
<collectingCounty id="62A14E07FFA5FFB93EE2626DFC927563" box="[693,860,539,564]" pageId="48" pageNumber="389">Fugong County</collectingCounty>
on the eastern slope and crest of the northcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFA5FFB93E62624AFDDA7524" location="Core Area" pageId="48" pageNumber="389" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFA5FFB93E62624AFDA47503" box="[565,618,571,595]" country="Myanmar" name="Shan" pageId="48" pageNumber="389">Shan</collectingRegion>
(see
<typeStatus id="54C48829FFA5FFB93EF4624DFD167504" box="[675,728,572,596]" pageId="48" pageNumber="389">Type</typeStatus>
material above for exact collection data). These localities are all in
<location id="8EA06050FFA5FFB93DD8622DFE337524" LSID="urn:lsid:plazi:treatment:03D6879DFFB8FFB93C2C630AFC7975E3:8EA06050FFA5FFB93DD8622DFE337524" box="[399,509,604,628]" name="Core Area" pageId="48" pageNumber="389" stateProvince="Shan">Core Area</location>
3
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFA5FFB93CFC620AFC7975E3" blockId="48.[123,1174,219,691]" pageId="48" pageNumber="389">
<emphasis id="B90BEA99FFA5FFB93CFC620AFDAF75C2" bold="true" box="[171,609,635,659]" pageId="48" pageNumber="389">
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<smallCapsWord id="8D26A057FFA5FFB93D4A620EFE0A75C2" baselines="653" box="[285,452,639,658]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="48" pageNumber="389">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFA5FFB93D9A620EFDAF75C2" baselines="653" box="[461,609,639,658]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="48" pageNumber="389">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from the northcentral part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FFA5FFB93EFC62EDFCA875E3" box="[683,870,667,691]" country="China" name="Yunnan" pageId="48" pageNumber="389">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFA5FFB93F2762EAFC7D75E3" box="[880,947,667,691]" name="China" pageId="48" pageNumber="389">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,663 @@
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<mods:title id="437BC5DEBA30096634BD388F069101C6">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="601A33BEDD56B0A0D25676FA96C80A2C">Deuve, Thierry</mods:namePart>
<mods:affiliation id="0670AFC32EA6F560351DC1E134CA337E">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="A5FDDE8CB86555FB92EBC2B0BB6DD82A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="820FD1CE53681F0D0A8D265B08F85871">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<subSubSection id="C3656500FFBCFFA03C2C6357FD8E746E" box="[123,576,806,831]" pageId="41" pageNumber="382" type="nomenclature">
<paragraph id="8BC0368BFFBCFFA03C2C6357FD8E746E" blockId="41.[123,576,806,861]" box="[123,576,806,831]" pageId="41" pageNumber="382">
<heading id="D08881E7FFBCFFA03C2C6357FD8E746E" box="[123,576,806,831]" fontSize="10" level="2" pageId="41" pageNumber="382" reason="4">
<taxonomicName id="4C7F4D08FFBCFFA03C2C6357FD8E746E" ID-CoL="57ZNR" authority="Putzeys, 1870" authorityName="Putzeys Size" authorityYear="1870" box="[123,576,806,831]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="indicus" subGenus="Trechus">
<emphasis id="B90BEA99FFBCFFA03C2C6357FD8E746E" bold="true" box="[123,576,806,831]" pageId="41" pageNumber="382">
<emphasis id="B90BEA99FFBCFFA03C2C6357FF1A746E" bold="true" box="[123,212,806,830]" italics="true" pageId="41" pageNumber="382">Trechus</emphasis>
(
<emphasis id="B90BEA99FFBCFFA03CB36357FEF3746E" bold="true" box="[228,317,806,830]" italics="true" pageId="41" pageNumber="382">Trechus</emphasis>
)
<emphasis id="B90BEA99FFBCFFA03D196357FE50746E" bold="true" box="[334,414,806,830]" italics="true" pageId="41" pageNumber="382">indicus</emphasis>
<bibRefCitation id="EFEE4B7AFFBCFFA03DF26356FD8E746E" author="PUTZEYS, J." box="[421,576,806,831]" pageId="41" pageNumber="382" pagination="7 - 48" refId="ref60457" refString="PUTZEYS, J. 1870. Trechorum oculatorum monographia. Stettiner Entomologische Zeitung 31: 7 - 48, 145 - 201." type="journal article" year="1870">Putzeys, 1870</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBCFFA03C2C6336FE68740D" box="[123,422,839,861]" pageId="41" pageNumber="382" type="description">
<paragraph id="8BC0368BFFBCFFA03C2C6336FE68740D" blockId="41.[123,576,806,861]" box="[123,422,839,861]" pageId="41" pageNumber="382">
(
<figureCitation id="13442A0EFFBCFFA03CD56336FF10740D" box="[130,222,839,861]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="41" pageNumber="382">Figs. 16b</figureCitation>
,
<figureCitation id="13442A0EFFBCFFA03CBE6336FECD740C" box="[233,259,839,860]" captionStart="FIGURE" captionStartId="44.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1559]" captionTargetId="figure-12@44.[120,1176,216,1559]" captionTargetPageId="44" captionText="FIGURE 17. Trechus indicus Putzeys; a. Dorsal habitus (CASENT1016115). b. Median lobe of aedeagus of male (CASENT1015874), left lateral aspect. c. Map of locality records (red circles) for T. indicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155315" httpUri="https://zenodo.org/record/13155315/files/figure.png" pageId="41" pageNumber="382">17</figureCitation>
,
<figureCitation id="13442A0EFFBCFFA03D596336FEFD740C" box="[270,307,839,860]" captionStart="FIGURE" captionStartId="103.[171,183,1626,1645]" captionTargetBox="[130,1167,217,1606]" captionTargetId="figure-14@103.[130,1167,217,1606]" captionTargetPageId="103" captionText="FIGURE 41. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Dulongjiang Township, Dulong Jiang at Elideng village, elevation 1640 m; habitat in which specimens of Trechus indicus Putzeys were collected. b. Tengchong County, Mingguang Township, slope SW of Eighth Boundary Post Pass, elevation 2887 m; habitat in which specimens of Trechus mingguangensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155368" httpUri="https://zenodo.org/record/13155368/files/figure.png" pageId="41" pageNumber="382">41a</figureCitation>
,
<figureCitation id="13442A0EFFBCFFA03D6A6336FE99740C" box="[317,343,839,860]" captionStart="FIGURE" captionStartId="106.[171,183,895,914]" captionTargetBox="[120,1176,216,881]" captionTargetId="figure-12@106.[120,1176,215,882]" captionTargetPageId="106" captionText="FIGURE 44. Photograph of habitat for trechine species in the Gaoligong Shan region. Longling County, Longjiang Township, Xiaoheishan Forest Reserve, elevation 2020 m; habitat in which specimens of Agonotrechus xiaoheishan sp. nov. and Trechus indicus Putzeys were collected. Photo by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155374" httpUri="https://zenodo.org/record/13155374/files/figure.png" pageId="41" pageNumber="382">44</figureCitation>
,
<figureCitation id="13442A0EFFBCFFA03D356336FE51740D" box="[354,415,839,861]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStartId-0="106.[171,183,1674,1693]" captionStartId-1="107.[171,183,1629,1648]" captionStartId-2="108.[171,183,1359,1378]" captionStartId-3="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,974,1660]" captionTargetBox-1="[120,1176,245,1580]" captionTargetBox-2="[120,1176,228,1340]" captionTargetBox-3="[197,1099,216,1598]" captionTargetId-0="figure-14@106.[120,1176,974,1660]" captionTargetId-1="figure-14@107.[120,1176,216,1608]" captionTargetId-2="figure-12@108.[120,1176,228,1340]" captionTargetId-3="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="106" captionTargetPageId-1="107" captionTargetPageId-2="108" captionTargetPageId-3="109" captionText-0="FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km." captionText-1="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-2="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-3="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155376" figureDoi-1="http://doi.org/10.5281/zenodo.13155378" figureDoi-2="http://doi.org/10.5281/zenodo.13155380" figureDoi-3="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155376/files/figure.png" httpUri-1="https://zenodo.org/record/13155378/files/figure.png" httpUri-2="https://zenodo.org/record/13155380/files/figure.png" httpUri-3="https://zenodo.org/record/13155382/files/figure.png" pageId="41" pageNumber="382">4548</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBCFFA03C2C6302FB5B74BA" pageId="41" pageNumber="382" type="reference_group">
<paragraph id="8BC0368BFFBCFFA03C2C6302FB5B74D8" blockId="41.[123,1174,882,1225]" box="[123,1173,882,905]" pageId="41" pageNumber="382">
<treatmentCitationGroup id="AB6F11A5FFBCFFA03C2C6302FB5B74D8" box="[123,1173,882,905]" pageId="41" pageNumber="382">
<treatmentCitation id="0ADE109AFFBCFFA03C2C6302FE0074D8" author="PUTZEYS, J." box="[123,462,882,904]" page="175" pageId="41" pageNumber="382" year="1870">
<taxonomicName id="4C7F4D08FFBCFFA03C2C6302FE0074D8" ID-CoL="57ZNR" authority="Putzeys, 1870: 175" authorityName="Putzeys Size" authorityPageNumber="175" authorityYear="1870" box="[123,462,882,904]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFBCFFA03C2C6302FEDF74D8" box="[123,273,883,904]" italics="true" pageId="41" pageNumber="382">Trechus indicus</emphasis>
<bibRefCitation id="EFEE4B7AFFBCFFA03D4F6302FE0074D8" author="PUTZEYS, J." box="[280,462,882,904]" pageId="41" pageNumber="382" pagination="7 - 48" refId="ref60457" refString="PUTZEYS, J. 1870. Trechorum oculatorum monographia. Stettiner Entomologische Zeitung 31: 7 - 48, 145 - 201." type="journal article" year="1870">Putzeys, 1870:175</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54C48829FFBCFFA03D806302FDFD74D9" box="[471,563,883,905]" pageId="41" pageNumber="382" type="holotype">Holotype</typeStatus>
, lost (see
<bibRefCitation id="EFEE4B7AFFBCFFA03EC46302FCDA74D8" author="JEANNEL, R." box="[659,788,883,905]" pageId="41" pageNumber="382" pagination="393 - 435" refId="ref59489" refString="JEANNEL, R. 1923. Les Trechinae (Coleoptera, Carabidae) de la Region Orientale. Annals and Magazine of Natural History (9) 12: 393 - 435." type="journal article" year="1923">Jeannel 1923</bibRefCitation>
, footnote p. 416). Type locality: eastern
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC0368BFFBCFFA03CFC63E2FB5B74BA" blockId="41.[123,1174,882,1225]" pageId="41" pageNumber="382">
<collectingCountry id="F368761BFFBCFFA03CFC63E2FF1174F9" box="[171,223,915,937]" name="India" pageId="41" pageNumber="382">India</collectingCountry>
.
<taxonomicName id="4C7F4D08FFBCFFA03C2C63C5FE7E7499" ID-CoL="57ZNR" authority="Putzeys" authorityName="Putzeys Size" authorityYear="1870" box="[123,432,948,969]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFBCFFA03C2C63C5FF087499" box="[123,198,948,969]" italics="true" pageId="41" pageNumber="382">Trechus</emphasis>
(
<emphasis id="B90BEA99FFBCFFA03C8363C5FEC67499" box="[212,264,948,969]" italics="true" pageId="41" pageNumber="382">s. str.</emphasis>
)
<emphasis id="B90BEA99FFBCFFA03D4163C5FE957499" box="[278,347,948,969]" italics="true" pageId="41" pageNumber="382">indicus</emphasis>
Putzeys
</taxonomicName>
:
<bibRefCitation id="EFEE4B7AFFBCFFA03DEE63C5FDA5749A" author="JEANNEL, R." box="[441,619,948,970]" pageId="41" pageNumber="382" pagination="1 - 592" refId="ref59554" refString="JEANNEL, R. 1927. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres. (Deuxieme livraison). L'Abeille 33: 1 - 592." type="journal article" year="1927">Jeannel, 1927:157</bibRefCitation>
, 158 (in part)
<taxonomicName id="4C7F4D08FFBCFFA03C2C63A5FDE474BA" ID-CoL="8LP5X" authority="Jeannel, 1927: 157" authorityName="Jeannel" authorityPageNumber="157" authorityYear="1927" box="[123,554,980,1002]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="B90BEA99FFBCFFA03C2C63A5FF0874B9" box="[123,198,980,1001]" italics="true" pageId="41" pageNumber="382">Trechus</emphasis>
(
<emphasis id="B90BEA99FFBCFFA03C8163A4FEC274BA" box="[214,268,981,1002]" italics="true" pageId="41" pageNumber="382">s. str.</emphasis>
)
<emphasis id="B90BEA99FFBCFFA03D4B63A4FEA074BA" box="[284,366,981,1002]" italics="true" pageId="41" pageNumber="382">macrops</emphasis>
<bibRefCitation id="EFEE4B7AFFBCFFA03D2163A5FDE474BA" author="JEANNEL, R." box="[374,554,980,1002]" pageId="41" pageNumber="382" pagination="1 - 592" refId="ref59554" refString="JEANNEL, R. 1927. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres. (Deuxieme livraison). L'Abeille 33: 1 - 592." type="journal article" year="1927">Jeannel, 1927:157</bibRefCitation>
</taxonomicName>
, 160 (in part).
<typeStatus id="54C48829FFBCFFA03E9663A4FD3F74BA" box="[705,753,981,1002]" pageId="41" pageNumber="382">Type</typeStatus>
locality:
<collectingCountry id="F368761BFFBCFFA03F0663A5FC4374BA" box="[849,909,980,1002]" name="China" pageId="41" pageNumber="382">China</collectingCountry>
,
<collectingRegion id="49BBF869FFBCFFA03FCE63A4FC2874BA" box="[921,998,981,1002]" country="China" name="Yunnan" pageId="41" pageNumber="382">Yunnan</collectingRegion>
. Synonymized by
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBCFFA03CFC6382FF397399" pageId="41" pageNumber="382" type="description">
<paragraph id="8BC0368BFFBCFFA03CFC6382FF397399" blockId="41.[123,1174,882,1225]" pageId="41" pageNumber="382">
<bibRefCitation id="EFEE4B7AFFBCFFA03CFC6382FEA37359" author="JEANNEL, R." box="[171,365,1011,1033]" pageId="41" pageNumber="382" pagination="273 - 282" refId="ref59618" refString="JEANNEL, R. 1935. Sur quelques Trechinae de l'Asie Centrale. Revue Francaise d'Entomologie 1: 273 - 282." type="journal article" year="1935">Jeannel (1935: 275)</bibRefCitation>
.
<taxonomicName id="4C7F4D08FFBCFFA03C2C6463FEBF7378" authority="Jeannel" authorityName="Jeannel" box="[123,369,1042,1064]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="B90BEA99FFBCFFA03C2C6463FED07378" box="[123,286,1042,1064]" italics="true" pageId="41" pageNumber="382">Trechus macrops</emphasis>
Jeannel
</taxonomicName>
:
<bibRefCitation id="EFEE4B7AFFBCFFA03D2D6463FDD67378" author="ANDREWES, H. E." box="[378,536,1042,1064]" pageId="41" pageNumber="382" refId="ref57956" refString="ANDREWES, H. E. 1935. The fauna of British India, including Ceylon and Burma. Coleoptera. Carabidae. Vol. II. Carabinae - 1. Taylor &amp; Francis, London, xvi + 323 pp. + 5 pl. + 1 map." type="book" year="1935">Andrewes, 1935</bibRefCitation>
63, 67.
<taxonomicName id="4C7F4D08FFBCFFA03C2C6442FEA87318" authority="Putzeys" authorityName="Putzeys Size" authorityYear="1870" box="[123,358,1075,1096]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFBCFFA03C2C6442FEDF7318" box="[123,273,1075,1096]" italics="true" pageId="41" pageNumber="382">Trechus indicus</emphasis>
Putzeys
</taxonomicName>
:
<bibRefCitation id="EFEE4B7AFFBCFFA03D276442FDEC7319" author="JEANNEL, R." box="[368,546,1075,1097]" pageId="41" pageNumber="382" pagination="273 - 282" refId="ref59618" refString="JEANNEL, R. 1935. Sur quelques Trechinae de l'Asie Centrale. Revue Francaise d'Entomologie 1: 273 - 282." type="journal article" year="1935">Jeannel, 1935:275</bibRefCitation>
.
<taxonomicName id="4C7F4D08FFBCFFA03C2C6422FEA87339" authority="Putzeys" authorityName="Putzeys Size" authorityYear="1870" box="[123,358,1107,1129]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFBCFFA03C2C6422FEDF7338" box="[123,273,1107,1128]" italics="true" pageId="41" pageNumber="382">Trechus indicus</emphasis>
Putzeys
</taxonomicName>
:
<bibRefCitation id="EFEE4B7AFFBCFFA03D276425FDC37339" author="DEUVE, T." box="[368,525,1108,1129]" pageId="41" pageNumber="382" pagination="79 - 88" refId="ref58750" refString="DEUVE, T. 1988. Nouveaux Trechinae de la Region Himalayenne (Col. Trechidae). Bulletin de la Societe Entomologique de France 93: 79 - 88." type="journal article" year="1988">Deuve, 1988:80</bibRefCitation>
.
<taxonomicName id="4C7F4D08FFBCFFA03C2C6405FE7E73D9" authority="Putzeys" authorityName="Putzeys Size" authorityYear="1870" box="[123,432,1140,1162]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFBCFFA03C2C6405FF0873D9" box="[123,198,1140,1161]" italics="true" pageId="41" pageNumber="382">Trechus</emphasis>
(
<emphasis id="B90BEA99FFBCFFA03C836404FEC673DA" box="[212,264,1141,1162]" italics="true" pageId="41" pageNumber="382">s. str.</emphasis>
)
<emphasis id="B90BEA99FFBCFFA03D416405FE9573D9" box="[278,347,1140,1161]" italics="true" pageId="41" pageNumber="382">indicus</emphasis>
Putzeys
</taxonomicName>
:
<bibRefCitation id="EFEE4B7AFFBCFFA03DEE6405FD9F73D9" author="UENO, S. I." box="[441,593,1140,1161]" pageId="41" pageNumber="382" pagination="175 - 196" refId="ref61071" refString="UENO, S. I. 1977. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Carabidae Subfam. Trechinae. Entomologica Basiliensia 2: 175 - 196." type="journal article" year="1977">Uéno 1977:181</bibRefCitation>
.
<taxonomicName id="4C7F4D08FFBCFFA03C2C64E4FE7273FB" authority="Jeannel" authorityName="Jeannel" box="[123,444,1173,1195]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="B90BEA99FFBCFFA03C2C64E4FF0873FA" box="[123,198,1173,1194]" italics="true" pageId="41" pageNumber="382">Trechus</emphasis>
(
<emphasis id="B90BEA99FFBCFFA03C8264E4FEC773FA" box="[213,265,1173,1194]" italics="true" pageId="41" pageNumber="382">s. str.</emphasis>
)
<emphasis id="B90BEA99FFBCFFA03D4064E4FEA773FA" box="[279,361,1173,1194]" italics="true" pageId="41" pageNumber="382">macrops</emphasis>
Jeannel
</taxonomicName>
:
<bibRefCitation id="EFEE4B7AFFBCFFA03D9164E4FD6773FA" author="UENO, S. I. &amp; W. Y. YIN" box="[454,681,1172,1195]" pageId="41" pageNumber="382" pagination="353 - 361" refId="ref61516" refString="UENO, S. I., AND W. Y. YIN. 1993. Notes on the trechine fauna (Coleoptera, Trechinae) of the Diancang Shan Mountains in western Yunnan, Southwest China. Elytra 21: 353 - 361." type="journal article" year="1993">Uéno &amp; Yin, 1993:354</bibRefCitation>
. These authors considered
<taxonomicName id="4C7F4D08FFBCFFA03FE764E4FBD573FA" box="[944,1051,1173,1194]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="382" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="B90BEA99FFBCFFA03FE764E4FBD573FA" box="[944,1051,1173,1194]" italics="true" pageId="41" pageNumber="382">T. macrops</emphasis>
</taxonomicName>
as a distinct species.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBCFFA03CFC64ACFCB0712C" pageId="41" pageNumber="382" type="diagnosis">
<paragraph id="8BC0368BFFBCFFA03CFC64ACFCB0712C" blockId="41.[123,1174,1245,1726]" pageId="41" pageNumber="382">
<smallCapsWord id="8D26A057FFBCFFA03CFC64ACFEEE73A4" baselines="1263,1263" box="[171,288,1245,1269]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="41" pageNumber="382">
<emphasis id="B90BEA99FFBCFFA03CFC64ACFEEE73A4" bold="true" box="[171,288,1245,1269]" pageId="41" pageNumber="382">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFBCFFA03E1064ACFD6D73A5" box="[583,675,1245,1269]" captionStart="FIGURE" captionStartId="44.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1559]" captionTargetId="figure-12@44.[120,1176,216,1559]" captionTargetPageId="44" captionText="FIGURE 17. Trechus indicus Putzeys; a. Dorsal habitus (CASENT1016115). b. Median lobe of aedeagus of male (CASENT1015874), left lateral aspect. c. Map of locality records (red circles) for T. indicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155315" httpUri="https://zenodo.org/record/13155315/files/figure.png" pageId="41" pageNumber="382">Fig. 17a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size large for genus (BL = 4.2 to 5.0 mm); fully-winged; eyes markedly large, tempora very short, joined to neck region at nearly a right angle; frontal furrows angulate at midlength; right mandible as in
<figureCitation id="13442A0EFFBCFFA03F81654EFBFC7207" box="[982,1074,1343,1367]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="41" pageNumber="382">Fig. 16b</figureCitation>
; mentum and submentum not fused; mentum with medial tooth simple, triangular; antennae with antennomeres 3 and 4 subequal in length, antennomere 2 slightly shorter; pronotum transverse (ratio PW/PL = 1.43), with lateral margins not sinuate posteriorly, straight just anterior to basal angles, the latter small and rectangular, lateral explanation very wide basally, basal margin slightly convex medally; elytra oblong, lateral discal striae effaced, only the medial four or five striae distinctly impressed, recurrent stria markedly impressed; preapical seta absent or vestigial, two discal setae present, inserted in stria 3 at anterior one-fifth and at middle, respectively; median lobe of male aedeagus (
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) not arcuate, with apex slender, ventrally hooked, endophallus with two elongate, projecting sclerites, one apically tapered, the other apically lobate.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFBCFFA43CFC66F4FC1E7404" lastPageId="45" lastPageNumber="386" pageId="41" pageNumber="382" type="discussion">
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<emphasis id="B90BEA99FFBCFFA03CFC66F4FEE671CB" bold="true" box="[171,296,1669,1693]" pageId="41" pageNumber="382">COMMENTS</emphasis>
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.—
<bibRefCitation id="EFEE4B7AFFBCFFA03D1866F4FDCB71CD" author="UENO, S. I." box="[335,517,1669,1693]" pageId="41" pageNumber="382" pagination="175 - 196" refId="ref61071" refString="UENO, S. I. 1977. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Carabidae Subfam. Trechinae. Entomologica Basiliensia 2: 175 - 196." type="journal article" year="1977">Uéno (1977:182)</bibRefCitation>
correctly pointed out that this widely distributed species, with members fully-winged, is morphologically varied across its range. In particular, the preapical seta tends to be absent from members of eastern populations. The many specimens we examined from localities in the study area confirm his observation. Among them, the prepical seta is absent from most specimens. If present, the seta is very small or vestigial, and only present unilaterally in most such cases. This led
<bibRefCitation id="EFEE4B7AFFBFFFA33D0C614DFDEF7604" author="UENO, S. I." box="[347,545,316,340]" pageId="42" pageNumber="383" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno (1999a:215)</bibRefCitation>
, while reporting the presence of this species in the Gaoligong
<collectingRegion id="49BBF869FFBFFFA33CE0612DFF227624" box="[183,236,348,372]" country="Myanmar" name="Shan" pageId="42" pageNumber="383">Shan</collectingRegion>
for the first time, to consider the eastern populations, including those in the study area, as a separate species,
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<emphasis id="B90BEA99FFBFFFA33D32610CFE1576C4" box="[357,475,381,404]" italics="true" pageId="42" pageNumber="383">T. macrops</emphasis>
Jeannel.
</taxonomicName>
</paragraph>
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<emphasis id="B90BEA99FFBFFFA33CFC61ECFE6A76E3" bold="true" box="[171,420,413,437]" pageId="42" pageNumber="383">
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</emphasis>
.— Members of this species have been found in a variety of microhabitats at elevations ranging from
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. They have been collected in daytime from under stones on the shaded and open banks of small to large streams, along roadcuts, and in closed canopy forest. At night, they have been found active on open sandy beaches and floodplain flats of larger streams (
<figureCitation id="13442A0EFFBFFFA33D73626CFEB17565" box="[292,383,541,565]" captionStart="FIGURE" captionStartId="103.[171,183,1626,1645]" captionTargetBox="[130,1167,217,1606]" captionTargetId="figure-14@103.[130,1167,217,1606]" captionTargetPageId="103" captionText="FIGURE 41. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Dulongjiang Township, Dulong Jiang at Elideng village, elevation 1640 m; habitat in which specimens of Trechus indicus Putzeys were collected. b. Tengchong County, Mingguang Township, slope SW of Eighth Boundary Post Pass, elevation 2887 m; habitat in which specimens of Trechus mingguangensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155368" httpUri="https://zenodo.org/record/13155368/files/figure.png" pageId="42" pageNumber="383">Fig. 41a</figureCitation>
) and on the ground along roadcuts. In almost all of the localities where specimens of
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<emphasis id="B90BEA99FFBFFFA33D58624FFEB67505" box="[271,376,573,597]" italics="true" pageId="42" pageNumber="383">T. indicus</emphasis>
</taxonomicName>
were founded, they were the only trechine collected. However, they were found syntopic with specimens of
<taxonomicName id="4C7F4D08FFBFFFA33DBC622FFCC87526" authorityName="Forest Reserve, N" authorityYear="2067" box="[491,774,606,630]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="383" phylum="Arthropoda" rank="species" species="xiaoheishan">
<emphasis id="B90BEA99FFBFFFA33DBC622FFCC87526" box="[491,774,606,630]" italics="true" pageId="42" pageNumber="383">Agonotrechus xiaoheishan</emphasis>
</taxonomicName>
at
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in the southern part of the study area.
<bibRefCitation id="EFEE4B7AFFBFFFA33D8D620FFDBE75C6" author="UENO, S. I." box="[474,624,638,662]" pageId="42" pageNumber="383" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno (1999a)</bibRefCitation>
reported that specimens of this species (which he recorded as
<taxonomicName id="4C7F4D08FFBFFFA33CAA62EFFDC275E6" authority="Jeannel" authorityName="Jeannel" box="[253,524,670,694]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="383" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="B90BEA99FFBFFFA33CAA62EFFE7D75E6" box="[253,435,670,694]" italics="true" pageId="42" pageNumber="383">Trechus macrops</emphasis>
Jeannel
</taxonomicName>
) were found in the same area as specimens of
<taxonomicName id="4C7F4D08FFBFFFA3385562EFFF2F7586" authorityName="Ueno Size" authorityYear="1999" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="383" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B90BEA99FFBFFFA3385562EFFF2F7586" italics="true" pageId="42" pageNumber="383">Agonotrechus yunnanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFBFFFA33D4162CFFDA77587" authority="(Ueno)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2016" box="[278,617,702,727]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="383" phylum="Arthropoda" rank="species" species="fortipes" status="comb. nov.">
<emphasis id="B90BEA99FFBFFFA33D4162CFFDD87587" box="[278,534,702,727]" italics="true" pageId="42" pageNumber="383">Epaphiotrechus fortipes</emphasis>
(Uéno)
</taxonomicName>
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</paragraph>
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</emphasis>
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<emphasis id="B90BEA99FFBFFFA33EDD62AFFCC175A5" bold="true" box="[650,783,734,758]" pageId="42" pageNumber="383">GAOLIGONG</emphasis>
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</emphasis>
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.—
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. We examined a total of
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(
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and
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) from the following localities:
<emphasis id="B90BEA99FFBFFFA33F8B628EFB5B7447" bold="true" box="[988,1173,767,791]" pageId="42" pageNumber="383">
<emphasis id="B90BEA99FFBFFFA33F8B628EFB427447" bold="true" box="[988,1164,767,791]" italics="true" pageId="42" pageNumber="383">Fugong County</emphasis>
:
</emphasis>
Lumadeng Township (Lumadeng,
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,
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/
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,
<date id="FFC1104BFFBFFFA33FF7636EFB827467" box="[928,1100,799,823]" pageId="42" pageNumber="383" value="2004-04-23">23 April 2004</date>
, D.H.
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collector [
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; CAS]).
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<emphasis id="B90BEA99FFBFFFA33E76634EFD277407" bold="true" box="[545,745,831,855]" italics="true" pageId="42" pageNumber="383">Gongshan County</emphasis>
:
</emphasis>
Dulongjiang Township (Bapo,
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,
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/
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,
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, H.B. Liang collector [
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; IOZ]; Bapo area, Dulong Jiang at Mulangdang,
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,
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/
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,
<date id="FFC1104BFFBFFFA33F1263F1FBC074C8" box="[837,1038,896,920]" pageId="42" pageNumber="383" value="2004-11-04">4 November 2004</date>
, H.B. Liang collector [
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; IOZ];
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N of Dizhengdang village on Dulong Jiang,
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,
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/
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,
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, D.H.
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, D.Z. Dong &amp; G. Tang collectors [
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and
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; CAS, IOZ]; Dulong Jiang at Elideng village,
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,
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/
<geoCoordinate id="EE4B504CFFBFFFA338456391FB5F74A8" box="[1042,1169,992,1016]" degrees="98.32145" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.32145">E98.32145°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33C2C6470FE8A7349" box="[123,324,1025,1049]" pageId="42" pageNumber="383" value="2004-11-03">3 November 2004</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA33DD86470FDC57349" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[399,523,1025,1049]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
, D.Z. Dong &amp; G. Tang collectors [
<specimenCount id="9D79FD02FFBFFFA33FC66470FC247349" box="[913,1002,1025,1049]" count="13" pageId="42" pageNumber="383" type="male">13 male</specimenCount>
and
<specimenCount id="9D79FD02FFBFFFA338736470FB5F7349" box="[1060,1169,1025,1049]" count="8" pageId="42" pageNumber="383" type="female">8 females</specimenCount>
; CAS, IOZ]; Dulong Jiang at Xianjiudang village,
<quantity id="4C879B6EFFBFFFA33ECF6450FD227368" box="[664,748,1057,1081]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="42" pageNumber="383" unit="m" value="1580.0">1580 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33EAE6450FCB57369" box="[761,891,1057,1081]" degrees="27.94092" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="27.94092">N27.94092°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33FD36450FBCA7369" box="[900,1028,1057,1081]" degrees="98.33340" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.3334">E98.33340°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA338586450FF7D7309" pageId="42" pageNumber="383" value="2004-11-04">4 November 2004</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA33CA16430FEBC7309" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[246,370,1089,1113]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
, M.A. Dixon, D.Z. Dong &amp; G. Tang collectors [
<specimenCount id="9D79FD02FFBFFFA33F396430FC7B7309" box="[878,949,1089,1113]" count="1" pageId="42" pageNumber="383" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFBFFFA33FB06433FB9F7309" box="[999,1105,1089,1113]" count="7" pageId="42" pageNumber="383" type="female">7 females</specimenCount>
; CAS, IOZ];
<quantity id="4C879B6EFFBFFFA33CE96413FEC67329" box="[190,264,1121,1146]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="383" unit="km" value="0.2">0.2 km</quantity>
S of confluence of Dulong Jiang and Muke Wang,
<quantity id="4C879B6EFFBFFFA33F7E6410FCB27329" box="[809,892,1121,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="42" pageNumber="383" unit="m" value="1450.0">1450 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33FD16410FBC77329" box="[902,1033,1121,1145]" degrees="27.84125" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="27.84125">N27.84125°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA338466410FB5F7329" box="[1041,1169,1121,1145]" degrees="98.33979" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.33979">E98.33979°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33C2C64F3FEF173CA" box="[123,319,1154,1178]" pageId="42" pageNumber="383" value="2004-11-07">7 November 2004</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA33DD264F3FDCF73CA" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[389,513,1154,1178]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
, V.F. Lee &amp; D.Z. Dong collectors [
<specimenCount id="9D79FD02FFBFFFA33F2D64F3FC1473CA" box="[890,986,1154,1178]" count="1" pageId="42" pageNumber="383" type="female">1 female</specimenCount>
; CAS];
<quantity id="4C879B6EFFBFFFA3386764F3FBB573CA" box="[1072,1147,1154,1178]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="383" unit="km" value="0.5">0.5 km</quantity>
N of Kongdang,
<quantity id="4C879B6EFFBFFFA33D4064D3FEA573E9" box="[279,363,1186,1210]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="383" unit="m" value="1500.0">1500 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33D2F64D3FE3773EA" box="[376,505,1186,1210]" degrees="27.88111" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="27.88111">N27.88111°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33E5564D3FD4F73EA" box="[514,641,1186,1210]" degrees="98.34063" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.34063">E98.34063°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33EDA64D3FC8873EA" box="[653,838,1186,1210]" pageId="42" pageNumber="383" value="2004-10-25">25 October 2004</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA33FD964D3FBC473EA" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[910,1034,1186,1210]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
, H.B. Liang, D.Z. Dong &amp; G. Tang collectors [
<specimenCount id="9D79FD02FFBFFFA33DB764B3FDF1738A" box="[480,575,1218,1242]" count="1" pageId="42" pageNumber="383" type="female">1 female</specimenCount>
; CAS]; Moqie Wang at Gongshan-Dulong Road Km 91,
<quantity id="4C879B6EFFBFFFA33C2C6493FF0073AA" box="[123,206,1250,1274]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="42" pageNumber="383" unit="m" value="1550.0">1550 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33C8E6492FE9273AB" box="[217,348,1251,1275]" degrees="27.89934" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="27.89934">N27.89934°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33D336492FE2A73AB" box="[356,484,1251,1275]" degrees="98.34999" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.34999">E98.34999°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33DB86493FD7C73AB" box="[495,690,1250,1275]" pageId="42" pageNumber="383" value="2004-11-06">6 November 2004</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA33EA06493FCBE73AA" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[759,880,1250,1274]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp; H.B. Liang collectors [
<specimenCount id="9D79FD02FFBFFFA338D16492FF77724B" count="2" pageId="42" pageNumber="383" type="male">2 males</specimenCount>
and
<specimenCount id="9D79FD02FFBFFFA33CA56572FE9B724B" box="[242,341,1283,1307]" count="4" pageId="42" pageNumber="383" type="female">4 female</specimenCount>
; CAS, IOZ]).
<emphasis id="B90BEA99FFBFFFA33DA36573FD75724B" bold="true" box="[500,699,1282,1307]" pageId="42" pageNumber="383">
<emphasis id="B90BEA99FFBFFFA33DA36573FD7C724B" bold="true" box="[500,690,1282,1307]" italics="true" pageId="42" pageNumber="383">Longling County</emphasis>
:
</emphasis>
Longjiang Township (small stream
<quantity id="4C879B6EFFBFFFA3381E6572FB5B724B" box="[1097,1173,1283,1307]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="42" pageNumber="383" unit="km" value="1.2">1.2 km</quantity>
SSE of Km 23.5 on Route 23.5,
<quantity id="4C879B6EFFBFFFA33D816552FDE4726A" box="[470,554,1315,1339]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="42" pageNumber="383" unit="m" value="2020.0">2020 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33E626552FD65726B" box="[565,683,1315,1339]" degrees="24.2888" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="5" value="24.2888">N24.2888°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33EE46552FCFD726B" box="[691,819,1315,1339]" degrees="98.76001" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.76001">E98.76001°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33F696552FC01726B" box="[830,975,1315,1339]" pageId="42" pageNumber="383" value="2005-05-25">25 May 2005</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA338426552FB5F726B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1045,1169,1315,1339]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
, H.B. Liang &amp; D.Z. Dong collectors [
<specimenCount id="9D79FD02FFBFFFA33E656532FD4D720B" box="[562,643,1347,1371]" count="1" pageId="42" pageNumber="383" type="male">1 male</specimenCount>
; CAS];
<taxonomicName id="4C7F4D08FFBFFFA33EBF6532FBA2720B" authority="Forest Reserve, 2067" authorityName="Forest Reserve, N" authorityYear="2067" box="[744,1132,1347,1371]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="383" phylum="Arthropoda" rank="species" species="xiaoheishan">
Xiaoheishan Forest Reserve,
<quantity id="4C879B6EFFBFFFA338606532FBA2720B" box="[1079,1132,1347,1371]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="42" pageNumber="383" unit="m" value="2067.0">2067</quantity>
</taxonomicName>
m,
<geoCoordinate id="EE4B504CFFBFFFA33C2C6512FF33722B" box="[123,253,1379,1403]" degrees="24.83671" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.83671">N24.83671°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33D526512FE48722B" box="[261,390,1379,1403]" degrees="98.76185" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.76185">E98.76185°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33DC46512FDE4722B" box="[403,554,1379,1404]" pageId="42" pageNumber="383" value="2005-05-28">28 May 2005</date>
, H.B. Liang, K.J. Guo &amp; H.M. Yan collectors [
<specimenCount id="9D79FD02FFBFFFA3386F6512FB5C722B" box="[1080,1170,1379,1403]" count="3" pageId="42" pageNumber="383" type="male">3 males</specimenCount>
; CAS, IOZ]); Zhenan Township (Bangbie village,
<quantity id="4C879B6EFFBFFFA33EF565F2FD3972CB" box="[674,759,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="42" pageNumber="383" unit="m" value="1540.0">1540 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33F5365F2FC4972CB" box="[772,903,1411,1435]" degrees="24.81306" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.81306">N24.81306°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33FD865F2FBC172CB" box="[911,1039,1411,1435]" degrees="98.83306" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.83306">E98.83306°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA3384B65F5FF7D72EC" pageId="42" pageNumber="383" value="2003-10-30">30 October 2003</date>
, H.B. Liang &amp; X.C. Shi collectors [
<specimenCount id="9D79FD02FFBFFFA33E1965D5FD5272EC" box="[590,668,1444,1468]" count="1" pageId="42" pageNumber="383" type="male">1 male</specimenCount>
; IOZ]).
<emphasis id="B90BEA99FFBFFFA33EAC65D5FC0072EC" bold="true" box="[763,974,1444,1468]" italics="true" pageId="42" pageNumber="383">Longyang County:</emphasis>
Bawan Township (Baoshan-Tenchong Road Km 24 at Nankang Yakou,
<quantity id="4C879B6EFFBFFFA33E9765B5FCDA728B" box="[704,788,1476,1500]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="42" pageNumber="383" unit="m" value="2130.0">2130 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33F7765B5FC6C728C" box="[800,930,1476,1500]" degrees="24.82583" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.82583">N24.82583°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33FFD65B5FBE5728C" box="[938,1067,1476,1500]" degrees="98.77222" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.77222">E98.77222°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA3386165B5FF1372AC" pageId="42" pageNumber="383" value="2003-10-26">26 October 2003</date>
, H.B. Liang &amp; X.C. Shi collectors [
<specimenCount id="9D79FD02FFBFFFA33E326595FD0B72AC" box="[613,709,1508,1532]" count="1" pageId="42" pageNumber="383" type="female">1 female</specimenCount>
; IOZ]; Bawan-Tengchong Road at Km 40- 41,
<quantity id="4C879B6EFFBFFFA33CF56675FF3B714C" box="[162,245,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="2.404" pageId="42" pageNumber="383" unit="m" value="2404.0">2404 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33D576675FE4D714C" box="[256,387,1540,1564]" degrees="24.93750" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.9375">N24.93750°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33DDB6675FDC5714C" box="[396,523,1540,1564]" degrees="98.75083" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.75083">E98.75083°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33E426675FD04714C" box="[533,714,1540,1564]" pageId="42" pageNumber="383" value="2003-10-12">12 October 2003</date>
, H.B. Liang &amp; X.C. Shi collectors [
<specimenCount id="9D79FD02FFBFFFA338106675FB5F714C" box="[1095,1169,1540,1564]" count="1" pageId="42" pageNumber="383" type="male">1 male</specimenCount>
; IOZ]; Bawan-Tengchong Road at Km 41,
<quantity id="4C879B6EFFBFFFA33E026655FD62716C" box="[597,684,1572,1596]" metricMagnitude="3" metricUnit="m" metricValue="2.4859999999999998" pageId="42" pageNumber="383" unit="m" value="2486.0">2486 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33EEC6655FCF0716C" box="[699,830,1572,1596]" degrees="24.93750" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.9375">N24.93750°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33F106655FC08716C" box="[839,966,1572,1596]" degrees="98.75083" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.75083">E98.75083°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33F826654FB5F716D" box="[981,1169,1573,1597]" pageId="42" pageNumber="383" value="2003-10-11">11 October 2003</date>
, H.B. Liang collector [1; female; IOZ]; Bawan-Tengchong Road at Km 42-46,
<quantity id="4C879B6EFFBFFFA338646634FB5F710C" box="[1075,1169,1605,1629]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="42" pageNumber="383" unit="m" value="2290.0">2290 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33C2C6614FF32712D" box="[123,252,1637,1661]" degrees="24.95361" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.95361">N24.95361°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33D526614FE48712D" box="[261,390,1637,1661]" degrees="98.74222" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.74222">E98.74222°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33DC36614FD9F712D" box="[404,593,1637,1661]" pageId="42" pageNumber="383" value="2003-10-14">14 October 2003</date>
, H.B. Liang &amp; X.C. Shi collectors [
<specimenCount id="9D79FD02FFBFFFA33FBF6614FB85712D" box="[1000,1099,1637,1661]" count="1" pageId="42" pageNumber="383" type="female">1 female</specimenCount>
; IOZ]; Luoshuidong area at Sancha He,
<quantity id="4C879B6EFFBFFFA33DA766F4FD8671CD" box="[496,584,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="42" pageNumber="383" unit="m" value="2300.0">2300 m</quantity>
,
<geoCoordinate id="EE4B504CFFBFFFA33E0066F4FD1771CD" box="[599,729,1669,1693]" degrees="24.94833" direction="north" orientation="latitude" pageId="42" pageNumber="383" precision="1" value="24.94833">N24.94833°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFBFFFA33EB566F4FCAC71CD" box="[738,866,1669,1693]" degrees="98.75667" direction="east" orientation="longitude" pageId="42" pageNumber="383" precision="1" value="98.75667">E98.75667°</geoCoordinate>
,
<date id="FFC1104BFFBFFFA33F2666F4FB9D71CD" box="[881,1107,1669,1693]" pageId="42" pageNumber="383" value="1998-10-26" valueMax="1998-10-31" valueMin="1998-10-26">26-31 October 1998</date>
, D.H.
<taxonomicName id="4C7F4D08FFBFFFA33C2C66D4FF3A71ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,244,1701,1725]" pageId="42" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp; C.E. Griswold collectors [
<specimenCount id="9D79FD02FFBFFFA33E6D66D7FD5271ED" box="[570,668,1701,1726]" count="1" pageId="42" pageNumber="383" type="female">1 female</specimenCount>
; CAS]).
<emphasis id="B90BEA99FFBFFFA33EA866D4FC1571ED" bold="true" box="[767,987,1701,1725]" italics="true" pageId="42" pageNumber="383">Tengchong County:</emphasis>
Jietou Township (Zhoujia-po village,
<quantity id="4C879B6EFFB8FFA43D3660AAFE7777A2" box="[353,441,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="45" pageNumber="386" unit="m" value="1740.0">1740 m</quantity>
,
<geoCoordinate id="EE4B504CFFB8FFA43D9D60AAFD8377A3" box="[458,589,219,243]" degrees="25.33222" direction="north" orientation="latitude" pageId="45" pageNumber="386" precision="1" value="25.33222">N25.33222°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB8FFA43E0160AAFD1A77A3" box="[598,724,219,243]" degrees="98.67611" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.67611">E98.67611°</geoCoordinate>
,
<date id="FFC1104BFFB8FFA43EB360AAFC6A77A3" box="[740,932,219,243]" pageId="45" pageNumber="386" value="2003-10-24">24 October 2003</date>
, D.Z. Dong collector [
<specimenCount id="9D79FD02FFB8FFA43CD3608AFF1A7643" box="[132,212,251,275]" count="1" pageId="45" pageNumber="386" type="male">1 male</specimenCount>
; CAS]); Qushi Township (Longchuan Jiang at Xiaojiangqiao,
<quantity id="4C879B6EFFB8FFA43FC1608AFC327643" box="[918,1020,251,275]" metricMagnitude="-6" metricUnit="m" metricValue="1.4449999999999998" pageId="45" pageNumber="386" unit="nm" value="1445.0">1445 nm</quantity>
,
<geoCoordinate id="EE4B504CFFB8FFA4385B608AFB437643" box="[1036,1165,251,275]" degrees="25.23944" direction="north" orientation="latitude" pageId="45" pageNumber="386" precision="1" value="25.23944">N25.23944°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFB8FFA43C2C616AFF327663" box="[123,252,283,307]" degrees="98.63722" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.63722">E98.63722°</geoCoordinate>
,
<date id="FFC1104BFFB8FFA43D50616AFE727663" box="[263,444,283,307]" pageId="45" pageNumber="386" value="2003-10-21">21 October 2003</date>
, H.B. Liang &amp; X.C. Shi collectors [
<specimenCount id="9D79FD02FFB8FFA43F6F616AFC4C7663" box="[824,898,283,307]" count="1" pageId="45" pageNumber="386" type="male">1 male</specimenCount>
; IOZ]); Shangying Township (Bawan-Tengchong Road Km 46-51,
<quantity id="4C879B6EFFB8FFA43E03614AFD657603" box="[596,683,315,339]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="45" pageNumber="386" unit="m" value="2220.0">2220 m</quantity>
,
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,
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, H.B. Liang &amp; X.C. Shi collectors [
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; IOZ]); Bawan-Tengchong Road Km 48-51 at Dahaoping Forest Station,
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,
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,
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, H.B. Liang collector [
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and
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; CAS, IOZ, MNHN]; Bawan-Tengchong Road Km 65, beside Longchuanjiang,
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,
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,
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, H.B. Liang &amp; X.C. Shi collectors [
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and
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; CAS IOZ]; Bawan-Tengchong Road Km 65 at Longwenqiao,
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,
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,
<date id="FFC1104BFFB8FFA43DC2618AFD9D7543" box="[405,595,507,531]" pageId="45" pageNumber="386" value="2003-10-20">20 October 2003</date>
, H.B. Liang &amp; X.C. Shi collectors [
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; IOZ]); Wuhe Township (
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Station,
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,
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<geoCoordinate id="EE4B504CFFB8FFA43F07626AFC1E7563" box="[848,976,539,563]" degrees="98.76000" direction="east" orientation="longitude" pageId="45" pageNumber="386" precision="1" value="98.76">E98.76000°</geoCoordinate>
,
<date id="FFC1104BFFB8FFA43F8C626AFB5F7563" box="[987,1169,539,563]" pageId="45" pageNumber="386" value="2003-10-29">29 October 2003</date>
, H.B. Liang collector [
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and
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; CAS, IOZ]).
</paragraph>
<caption id="DF006603FFBEFFA23CFC663AFDEE7195" ID-DOI="http://doi.org/10.5281/zenodo.13155313" ID-Zenodo-Dep="13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="43" pageNumber="384" startId="43.[171,183,1611,1630]" targetBox="[153,1126,230,1591]" targetPageId="43" targetType="figure">
<paragraph id="8BC0368BFFBEFFA23CFC663AFDEE7195" blockId="43.[123,1174,1611,1733]" pageId="43" pageNumber="384">
<smallCapsWord id="8D26A057FFBEFFA23CFC663AFF26710D" baselines="1625,1625" box="[171,232,1611,1630]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="43" pageNumber="384">FIGURE</smallCapsWord>
16. Trechine right mandibles, ventral aspect. a.
<taxonomicName id="4C7F4D08FFBEFFA23ED4663AFCAB710E" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[643,869,1611,1630]" class="Insecta" family="Carabidae" genus="Agonotrechus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="xiaoheishan" status="sp. nov.">
<emphasis id="B90BEA99FFBEFFA23ED4663AFCAB710E" box="[643,869,1611,1630]" italics="true" pageId="43" pageNumber="384">Agonotrechus xiaoheishan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBEFFA23F3C663AFC60710E" box="[875,942,1611,1630]" pageId="43" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
b.
<taxonomicName id="4C7F4D08FFBEFFA23F9F663AFB5B710E" authority="Putzeys" authorityName="Putzeys Size" authorityYear="1870" box="[968,1173,1611,1630]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFBEFFA23F9F663AFB80710E" box="[968,1102,1611,1630]" italics="true" pageId="43" pageNumber="384">Trechus indicus</emphasis>
Putzeys
</taxonomicName>
with dentition enlarged. c.
<taxonomicName id="4C7F4D08FFBEFFA23D3E6615FDFC7127" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[361,562,1636,1655]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FFBEFFA23D3E6615FDFC7127" box="[361,562,1636,1655]" italics="true" pageId="43" pageNumber="384">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBEFFA23E6D6614FDB17128" box="[570,639,1637,1656]" pageId="43" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
d.
<taxonomicName id="4C7F4D08FFBEFFA23EC96615FCF17127" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[670,831,1636,1655]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="shibalicus" status="sp. nov.">
<emphasis id="B90BEA99FFBEFFA23EC96615FCF17127" box="[670,831,1636,1655]" italics="true" pageId="43" pageNumber="384">Trechus shibalicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBEFFA23F106614FC437128" box="[839,909,1637,1656]" pageId="43" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
e.
<taxonomicName id="4C7F4D08FFBEFFA23FFD6615FF7D71C2" authority="Deuve" authorityName="Deuve" class="Insecta" family="Carabidae" genus="Pseudepaphius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="gonggaicus">
<emphasis id="B90BEA99FFBEFFA23FFD6615FB5B7127" box="[938,1173,1636,1655]" italics="true" pageId="43" pageNumber="384">Pseudepaphius gonggaicus</emphasis>
Deuve
</taxonomicName>
with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f.
<taxonomicName id="4C7F4D08FFBEFFA23857660FFF1F71FA" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFBEFFA23857660FFF1F71FA" italics="true" pageId="43" pageNumber="384">Trechepaphiopsis uniporosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBEFFA23C8066E9FED571FB" box="[215,283,1688,1707]" pageId="43" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
with dentition enlarged. g.
<taxonomicName id="4C7F4D08FFBEFFA23E5D66E9FCCB71FB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[522,773,1688,1707]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="fortipesoides" status="sp. nov.">
<emphasis id="B90BEA99FFBEFFA23E5D66E9FCCB71FB" box="[522,773,1688,1707]" italics="true" pageId="43" pageNumber="384">Epaphiotrechus fortipesoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBEFFA23F5B66E9FC9E71FB" box="[780,848,1688,1707]" pageId="43" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
with dentition enlarged. h.
<taxonomicName id="4C7F4D08FFBEFFA2381766E9FED97194" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="384" phylum="Arthropoda" rank="species" species="gaoligong" status="sp. nov.">
<emphasis id="B90BEA99FFBEFFA2381766E9FED97194" italics="true" pageId="43" pageNumber="384">Trechepaphiama gaoligong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFBEFFA23D4A66C3FEAE7195" box="[285,352,1714,1733]" pageId="43" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
Scale lines = 0.5 mm.
</paragraph>
</caption>
<caption id="DF006603FFB9FFA53CFC6656FE06713D" ID-DOI="http://doi.org/10.5281/zenodo.13155315" ID-Zenodo-Dep="13155315" httpUri="https://zenodo.org/record/13155315/files/figure.png" pageId="44" pageNumber="385" startId="44.[171,183,1575,1594]" targetBox="[120,1176,216,1559]" targetPageId="44" targetType="figure">
<paragraph id="8BC0368BFFB9FFA53CFC6656FE06713D" blockId="44.[123,1175,1575,1646]" pageId="44" pageNumber="385">
<smallCapsWord id="8D26A057FFB9FFA53CFC6656FF267169" baselines="1589,1589" box="[171,232,1575,1594]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="44" pageNumber="385">FIGURE</smallCapsWord>
17.
<taxonomicName id="4C7F4D08FFB9FFA53D436656FE25716A" authority="Putzeys" authorityName="Putzeys Size" authorityYear="1870" box="[276,491,1575,1594]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="385" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFB9FFA53D436656FE53716A" box="[276,413,1575,1594]" italics="true" pageId="44" pageNumber="385">Trechus indicus</emphasis>
Putzeys
</taxonomicName>
; a. Dorsal habitus (CASENT1016115). b. Median lobe of aedeagus of male (CASENT1015874), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFB9FFA53F6C6630FC417103" authorityName="Putzeys Size" authorityYear="1870" box="[827,911,1600,1619]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="385" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B90BEA99FFB9FFA53F6C6630FC417103" box="[827,911,1600,1619]" italics="true" pageId="44" pageNumber="385">T. indicus</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFB8FFA43CFC622AFE717583" blockId="45.[123,1174,219,852]" pageId="45" pageNumber="386">Members of this species have been collected in both northern and southern parts of the study area and on both eastern and western slopes of the Gaoligongshan. We have recorded this species from Core Areas 1, 3, 6, and 7. The absence of records from Core Areas 2, 4 and 5 is likely the result of inadequate sampling.</paragraph>
<paragraph id="8BC0368BFFB8FFA43CFC62AAFC1E7404" blockId="45.[123,1174,219,852]" pageId="45" pageNumber="386">
<emphasis id="B90BEA99FFB8FFA43CFC62AAFD9375A2" bold="true" box="[171,605,731,755]" pageId="45" pageNumber="386">
<smallCapsWord id="8D26A057FFB8FFA43CFC62AAFEDC75A2" baselines="749,749" box="[171,274,731,755]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="45" pageNumber="386">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFB8FFA43D4C62AEFE0C75A2" baselines="749" box="[283,450,735,754]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="45" pageNumber="386">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFB8FFA43D9E62AEFD9375A2" baselines="749" box="[457,605,735,754]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="45" pageNumber="386">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<figureCitation id="13442A0EFFB8FFA43ED062ADFD1775A3" box="[647,729,731,756]" captionStart="FIGURE" captionStartId="106.[171,183,1674,1693]" captionTargetBox="[120,1176,974,1660]" captionTargetId="figure-14@106.[120,1176,974,1660]" captionTargetPageId="106" captionText="FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.13155376" httpUri="https://zenodo.org/record/13155376/files/figure.png" pageId="45" pageNumber="386">Fig. 45</figureCitation>
. This species is widely distributed along the southern edge of the Himalayan Mountains and Qinghai-Xizang (Tibetan) Plateau from eastern
<collectingCountry id="F368761BFFB8FFA43C2C636DFF337464" box="[123,253,796,820]" name="Afghanistan" pageId="45" pageNumber="386">Afghanistan</collectingCountry>
to
<collectingRegion id="49BBF869FFB8FFA43D75636DFE2C7464" box="[290,482,796,820]" country="China" name="Sichuan" pageId="45" pageNumber="386">Sichuan Province</collectingRegion>
,
<collectingCountry id="F368761BFFB8FFA43DB9636DFDFF7464" box="[494,561,796,820]" name="China" pageId="45" pageNumber="386">China</collectingCountry>
. Attainment and maintenance of this broad geographical range is no doubt facilitated by the flight capability of members of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,347 @@
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<mods:title id="8F788DF581095E07F295434E8B9B889C">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="95F20964DADA5C114CFE48F455C9843A">Deuve, Thierry</mods:namePart>
<mods:affiliation id="E7474E1C02845253D0E35E45E2021930">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:affiliation id="5F1747966B626DC3402FD2A40C6EBE1E">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:affiliation id="1923587C18EBD27FDF68BE3F3D8AC148">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:title id="FC2AF84E7034531D03015550899A8604">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="F8F42EB5ABDCF04D5138433551660C3B">2016</mods:date>
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<mods:number id="5CE2C2C4634A284DA56B2DFC270F7FF9">63</mods:number>
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<subSubSection id="C3656500FFC1FFDD3C2C6307FDD574DE" box="[123,539,885,910]" pageId="84" pageNumber="425" type="nomenclature">
<paragraph id="8BC0368BFFC1FFDD3C2C6307FDD574DE" blockId="84.[123,539,885,940]" box="[123,539,885,910]" pageId="84" pageNumber="425">
<heading id="D08881E7FFC1FFDD3C2C6307FDD574DE" box="[123,539,885,910]" fontSize="10" level="2" pageId="84" pageNumber="425" reason="4">
<taxonomicName id="4C7F4D08FFC1FFDD3C2C6307FDD574DE" ID-CoL="8MMR7" authority="(Ueno), 1999" authorityName="" authorityYear="1999" baseAuthorityName="Ueno" baseAuthorityYear="1999" box="[123,539,885,910]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC1FFDD3C2C6307FDD574DE" bold="true" box="[123,539,885,910]" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3C2C6307FE4A74DE" bold="true" box="[123,388,886,910]" italics="true" pageId="84" pageNumber="425">Epaphiotrechus fortipes</emphasis>
(Uéno), 1999
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC1FFDD3C2C63E7FEDA74FC" box="[123,276,918,940]" pageId="84" pageNumber="425" type="description">
<paragraph id="8BC0368BFFC1FFDD3C2C63E7FEDA74FC" blockId="84.[123,539,885,940]" box="[123,276,918,940]" pageId="84" pageNumber="425">
(
<figureCitation id="13442A0EFFC1FFDD3CD563E7FF0774FB" box="[130,201,918,940]" captionStart="FIGURE" captionStartId="85.[699,711,1580,1599]" captionTargetBox="[123,1176,216,1723]" captionTargetId="figure-308@85.[648,1176,216,1567]" captionTargetPageId="85" captionText="FIGURE 32. Epaphiotrechus fortipes (Uéno); Map of locality record (red circle) in the Gaoligong Shan region. Scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155344" httpUri="https://zenodo.org/record/13155344/files/figure.png" pageId="84" pageNumber="425">Fig. 32</figureCitation>
,
<figureCitation id="13442A0EFFC1FFDD3C8363E7FEC374FC" box="[212,269,918,940]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="84" pageNumber="425">46-48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC1FFDD3C2C63CFFDB774A3" pageId="84" pageNumber="425" type="reference_group">
<paragraph id="8BC0368BFFC1FFDD3C2C63CFFDD77484" blockId="84.[123,633,958,1012]" box="[123,537,958,980]" pageId="84" pageNumber="425">
<treatmentCitationGroup id="AB6F11A5FFC1FFDD3C2C63CFFDD77484" box="[123,537,958,980]" pageId="84" pageNumber="425">
<treatmentCitation id="0ADE109AFFC1FFDD3C2C63CFFDDB7484" author="UENO, S. I." box="[123,533,958,980]" page="219" pageId="84" pageNumber="425" year="1999">
<taxonomicName id="4C7F4D08FFC1FFDD3C2C63CFFDDB7484" ID-CoL="8LP5V" authority="Ueno, 1999 a: 219" authorityName="Ueno" authorityPageNumber="219" authorityYear="1999" box="[123,533,958,980]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC1FFDD3C2C63CFFF087483" box="[123,198,958,979]" italics="true" pageId="84" pageNumber="425">Trechus</emphasis>
(
<emphasis id="B90BEA99FFC1FFDD3C8363CEFEC67484" box="[212,264,959,980]" italics="true" pageId="84" pageNumber="425">s. str.</emphasis>
)
<emphasis id="B90BEA99FFC1FFDD3D4163CFFE917483" box="[278,351,958,979]" italics="true" pageId="84" pageNumber="425">fortipes</emphasis>
<bibRefCitation id="EFEE4B7AFFC1FFDD3D3163CFFDDB7484" author="UENO, S. I." box="[358,533,958,980]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno, 1999a: 219</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BC0368BFFC1FFDD3C2C63AFFDB774A3" blockId="84.[123,633,958,1012]" box="[123,633,990,1012]" pageId="84" pageNumber="425">
<taxonomicName id="4C7F4D08FFC1FFDD3C2C63AFFE6174A4" ID-CoL="8MMR7" authority="(Ueno)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2016" box="[123,431,990,1012]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes" status="comb. nov.">
<emphasis id="B90BEA99FFC1FFDD3C2C63AFFEAF74A3" box="[123,353,990,1011]" italics="true" pageId="84" pageNumber="425">Epaphiotrechus fortipes</emphasis>
(Uéno)
</taxonomicName>
,
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<emphasis id="B90BEA99FFC1FFDD3DEE63AEFE2674A2" bold="true" box="[441,488,991,1012]" pageId="84" pageNumber="425">NEW</emphasis>
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<emphasis id="B90BEA99FFC1FFDD3DB963AFFDB774A3" bold="true" box="[494,633,990,1012]" pageId="84" pageNumber="425">COMBINATION</emphasis>
</smallCapsWord>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC1FFDD3CFC6474FE6273CD" pageId="84" pageNumber="425" type="materials_examined">
<paragraph id="8BC0368BFFC1FFDD3CFC6474FD16736C" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3CFC6474FE92734B" bold="true" box="[171,348,1029,1053]" pageId="84" pageNumber="425">
<smallCapsWord id="8D26A057FFC1FFDD3CFC6474FF2A734B" baselines="1047,1046" box="[171,228,1029,1053]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="84" pageNumber="425">TYPE</smallCapsWord>
<smallCapsWord id="8D26A057FFC1FFDD3CB86479FE92734B" baselines="1046" box="[239,348,1032,1051]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="84" pageNumber="425">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFC1FFDD3DD06474FD16736C" country="China" county="Baoshan County" elevation="2350" latitude="24.95" location="Gaoligong Shan" longLatPrecision="1249" longitude="98.75" pageId="84" pageNumber="425" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFC1FFDD3DD06474FE23734D" box="[391,493,1029,1053]" pageId="84" pageNumber="425" type="holotype">Holotype</typeStatus>
, a male, in NSMT. Type locality:
<collectingCountry id="F368761BFFC1FFDD3F266474FC7D734D" box="[881,947,1029,1053]" name="China" pageId="84" pageNumber="425">China</collectingCountry>
,
<collectingRegion id="49BBF869FFC1FFDD3F956474FBD8734C" box="[962,1046,1029,1052]" country="China" name="Yunnan" pageId="84" pageNumber="425">Yunnan</collectingRegion>
,
<location id="8EA06050FFC1FFDD38726474FF7D736D" LSID="urn:lsid:plazi:treatment:03D6879DFFC1FFD13C2C6307FB5875CC:8EA06050FFC1FFDD38726474FF7D736D" country="China" county="Baoshan County" latitude="24.95" longLatPrecision="1249" longitude="98.75" name="Gaoligong Shan" pageId="84" pageNumber="425" stateProvince="Yunnan">Gaoligong Shan</location>
,
<collectingCounty id="62A14E07FFC1FFDD3CE96454FEBF736D" box="[190,369,1061,1085]" pageId="84" pageNumber="425">Baoshan County</collectingCounty>
,
<geoCoordinate id="EE4B504CFFC1FFDD3D2B6452FE1A736C" box="[380,468,1059,1084]" degrees="24" direction="north" minutes="57" orientation="latitude" pageId="84" pageNumber="425" precision="925" value="24.95">N24°57</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFC1FFDD3DB36452FDF2736C" box="[484,572,1059,1084]" degrees="98" direction="east" minutes="45" orientation="longitude" pageId="84" pageNumber="425" precision="925" value="98.75">E98°45</geoCoordinate>
,
<quantity id="4C879B6EFFC1FFDD3E1F6455FD16736C" box="[584,728,1060,1084]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="84" pageNumber="425" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">
<elevation id="0052D1B8FFC1FFDD3E1F6455FD16736C" box="[584,728,1060,1084]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="84" pageNumber="425" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">2200-2500 m</elevation>
</quantity>
</materialsCitation>
</paragraph>
<paragraph id="8BC0368BFFC1FFDD3CFC6434FE6273CD" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3CFC6434FE00730B" bold="true" box="[171,462,1093,1117]" pageId="84" pageNumber="425">
<smallCapsWord id="8D26A057FFC1FFDD3CFC6434FF3D730B" baselines="1111,1110" box="[171,243,1093,1117]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Notes" pageId="84" pageNumber="425">NOTES</smallCapsWord>
<smallCapsWord id="8D26A057FFC1FFDD3CAB6439FED4730B" baselines="1110" box="[252,282,1096,1115]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="on" pageId="84" pageNumber="425">ON</smallCapsWord>
<smallCapsWord id="8D26A057FFC1FFDD3D746439FE96730B" baselines="1110" box="[291,344,1096,1115]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="type" pageId="84" pageNumber="425">TYPE</smallCapsWord>
<smallCapsWord id="8D26A057FFC1FFDD3D366439FE00730B" baselines="1110" box="[353,462,1096,1115]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="84" pageNumber="425">MATERIAL</smallCapsWord>
</emphasis>
.— We have not had an opportunity to study the
<typeStatus id="54C48829FFC1FFDD3FBD6434FB89730D" box="[1002,1095,1093,1117]" pageId="84" pageNumber="425" type="holotype">holotype</typeStatus>
or any
<typeStatus id="54C48829FFC1FFDD3C2C6414FEE3732D" box="[123,301,1125,1149]" isOtherTypeStatus="true" pageId="84" pageNumber="425">other specimens</typeStatus>
of this species. Features noted below are based on
<bibRefCitation id="EFEE4B7AFFC1FFDD3FDC6414FBF8732D" author="UENO, S. I." box="[907,1078,1125,1149]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uénos (1999a)</bibRefCitation>
original description and illustrations.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC1FFDF3CFC64D4FD957565" lastPageId="86" lastPageNumber="427" pageId="84" pageNumber="425" type="diagnosis">
<paragraph id="8BC0368BFFC1FFDD3CFC64D4FC22738C" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
<smallCapsWord id="8D26A057FFC1FFDD3CFC64D4FEEE73EB" baselines="1207,1206" box="[171,288,1189,1213]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3CFC64D4FEEE73EB" bold="true" box="[171,288,1189,1213]" pageId="84" pageNumber="425">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (see Uéno 199a,
<figureCitation id="13442A0EFFC1FFDD3EB964D4FCE073ED" box="[750,814,1189,1213]" captionStart="FIGURES" captionStartId="4.[171,183,1157,1176]" captionTargetBox="[120,1176,216,1144]" captionTargetId="figure-163@4.[120,1176,216,1144]" captionTargetPageId="4" captionText="FIGURES 24. Fig. 2. Map showing natural extent of study area, colored in green (however, sampling was permitted only in those portions in Yunnan Province. Fig. 3. Map showing location of core sampling areas. Fig. 4. Map showing locations of all entomological sampling sites." figureDoi="http://doi.org/10.5281/zenodo.13155287" httpUri="https://zenodo.org/record/13155287/files/figure.png" pageId="84" pageNumber="425">Fig. 4</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states:
</paragraph>
<paragraph id="8BC0368BFFC1FFDD3CFC6494FEC3710D" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
Size large (BL =
<quantity id="4C879B6EFFC1FFDD3D3D6494FE0A73AD" box="[362,452,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="84" pageNumber="425" unit="mm" value="4.5">4.5 mm</quantity>
), apterous; female
<typeStatus id="54C48829FFC1FFDD3ECD6494FD3973AD" box="[666,759,1253,1277]" pageId="84" pageNumber="425" type="holotype">holotype</typeStatus>
with unusually broad protarsomeres; body color dark piceous, slightly paler reddish on elytral interval 1; right mandible tridentate; mentum with median tooth short and simple; pronotum slightly narrowed (ratio PW/PL = 1.26), with basal angles small and rectangular; elytra elongate, slightly flattened medially, striae finely impressed, medial three or four striae distinctly crenulate (due to strial punctures), more lateral striae slightly effaced, but striae 6 and 7 evident, stria 5 deepened at its base; parascutellar striole present, slightly elongate; recurrent stria short, terminated near apex of stria 5; two discal setiferous pores present, inserted at anterior one-sixth and slightly posterior to middle, respectively, on interval 3 near stria 3; preapical seta present, inserted on interval 3 next to stria 2 and about equidistant from apical and sutural elytral margins; median lobe of male aedeagus (see
<bibRefCitation id="EFEE4B7AFFC1FFDD3FEF6674FBF0714D" author="UENO, S. I." box="[952,1086,1541,1565]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno 1999a</bibRefCitation>
,
<figureCitation id="13442A0EFFC1FFDD381C6674FB5B714D" box="[1099,1173,1541,1565]" captionStart="FIGURE" captionStartId="8.[171,183,1613,1632]" captionTargetBox="[120,1176,224,1597]" captionTargetId="figure-11@8.[120,1176,224,1597]" captionTargetPageId="8" captionText="FIGURE 5. Perileptus imaicus Jeannel; a. Dorsal habitus (CASENT1036287). b. Median lobe of aedeagus of male (CASENT10363020), left lateral aspect. c. Map of locality records (red circles) for P. imaicus in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155291" httpUri="https://zenodo.org/record/13155291/files/figure.png" pageId="84" pageNumber="425">Figs. 5</figureCitation>
and
<figureCitation id="13442A0EFFC1FFDD3CFE6654FF79716D" box="[169,183,1573,1597]" captionStart="FIGURE" captionStartId="10.[171,183,1609,1628]" captionTargetBox="[144,1176,216,1593]" captionTargetId="figure-11@10.[120,1176,216,1593]" captionTargetPageId="10" captionText="FIGURE 6. Perileptus pusilloides sp. nov.; a. Dorsal habitus (CASENT1039260). b. Median lobe of aedeagus of male (CASENT1039259), left lateral aspect. c. Map of locality records (red circles) for P. pusilloides in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155293" httpUri="https://zenodo.org/record/13155293/files/figure.png" pageId="84" pageNumber="425">6</figureCitation>
) with apex, expanded, securiform in lateral view; endophallus with distinct, elongate-triangular sclerite.
</paragraph>
<paragraph id="8BC0368BFFC1FFDD3CFC6614FB5871ED" blockId="84.[123,1174,1029,1726]" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3CFC6614FE6A712C" bold="true" box="[171,420,1637,1661]" pageId="84" pageNumber="425">
<smallCapsWord id="8D26A057FFC1FFDD3CFC6614FEC7712C" baselines="1655,1655" box="[171,265,1637,1661]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="84" pageNumber="425">HABITAT</smallCapsWord>
<smallCapsWord id="8D26A057FFC1FFDD3D476618FE6A712C" baselines="1655" box="[272,420,1641,1660]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="84" pageNumber="425">DISTRIBUTION</smallCapsWord>
</emphasis>
.— No members of this species were found during field collecting for this project, and no habitat information accompanied the
<typeStatus id="54C48829FFC1FFDD3EB866F7FCD371CE" box="[751,797,1670,1694]" pageId="84" pageNumber="425">type</typeStatus>
material.
<bibRefCitation id="EFEE4B7AFFC1FFDD3FDA66F4FBEA71CD" author="UENO, S. I." box="[909,1060,1669,1693]" pageId="84" pageNumber="425" pagination="215 - 223" refId="ref61432" refString="UENO, S. I. 1999 a. Two new trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in Yunnan, Southwest China. Bulletin of the National Science Museum, Ser. A (Zool.) 25: 215 - 223." type="journal article" year="1999">Uéno (1999a)</bibRefCitation>
suggested that the
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locality was located “near the lower edge of the
<taxonomicName id="4C7F4D08FFC1FFDD3F4F66D4FC7871ED" box="[792,950,1701,1725]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="84" pageNumber="425" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFC1FFDD3F4F66D4FC7871ED" box="[792,950,1701,1725]" italics="true" pageId="84" pageNumber="425">Rhododendron</emphasis>
</taxonomicName>
zone. He visited the
</paragraph>
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32.
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<emphasis id="B90BEA99FFC0FFDC3F7F665DFC34716F" box="[808,1018,1580,1599]" italics="true" pageId="85" pageNumber="426">Epaphiotrechus fortipes</emphasis>
(Uéno)
</taxonomicName>
; Map of locality record (red circle) in the Gaoligong Shan region.
</paragraph>
<paragraph id="8BC0368BFFC0FFDC3EDC662EFCF37122" blockId="85.[651,1175,1580,1650]" box="[651,829,1631,1650]" pageId="85" pageNumber="426">Scale line = 100 km.</paragraph>
</caption>
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<emphasis id="B90BEA99FFC3FFDF3CFC60AAFEEE77A1" bold="true" box="[171,288,219,243]" pageId="86" pageNumber="427">DIAGNOSIS</emphasis>
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.— Adults of this species (
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) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL =
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), apterous; body color piceous, sutural and lateral elytral margins slightly paler, reddish; tempora glabrous; median tooth of mentum apically truncate, almost bifurcate; antennae long; pronotum slightly narrowed (ratio PW/PL = 1.28), basal angles small, sharp, rectangular; elytra oblong, slightly flattened along median suture, striae finely impressed, finely punctate, striae 1 to 2 or 3 more deeply impressed, more lateral striae more or less effaced but still evident, stria 5 not more deeply impressed at its base; recurrent stria long, deeply impressed, abruptly terminated on interval 7; two discal setiferous pores present, inserted at anterior one-sixth and slightly posterior to middle, respectively, on interval 3 near stria 3; preapical seta present, inserted 1.5 times as far from apical elytral margin as from sutural margin.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC3FFDF3CFC624CFC8C71CC" pageId="86" pageNumber="427" type="description">
<paragraph id="8BC0368BFFC3FFDF3CFC624CFBB475E6" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">
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<emphasis id="B90BEA99FFC3FFDF3CFC624CFEF47504" bold="true" box="[171,314,573,597]" pageId="86" pageNumber="427">DESCRIPTION</emphasis>
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.— Size large, BL =
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. Body color dark piceous, head, basal and lateral pronotal margins, and lateral and sutural margins of elytra more or less reddish, appendages paler reddish brown, palpi yellowish brown, dorsum smooth, shiny with pronotum and elytra moderately iridescent, head faintly alutaceous due to moderately impressed isodiametric microsculpture.
</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC62CFFEEC7348" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">
Head. Broad, slightly elongate. Eyes slightly convex, their diameter slightly less than twice length of tempora, the latter short but distinctly convex. Frons convex, with frontal furrows sharply impressed, not attenuated posterior to eyes. Two pairs of supraorbital setae present, the anterior pair inserted in slight foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible (
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Fig.
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) with premolar not fused with retinaculum but closely associated with the latter, anterior tip of retinaculum enlarged as a distinct tooth and displaced anteriorly.
<collectingRegion id="49BBF869FFC3FFDF3CE563F1FEC274C7" box="[178,268,896,919]" country="Myanmar" name="Chin" pageId="86" pageNumber="427">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFC3FFDF3E6D63F1FD5A74C7" box="[570,660,896,919]" country="Myanmar" name="Chin" pageId="86" pageNumber="427">Mentum</collectingRegion>
with medial tooth apically truncate, about one-half the length of the lateral lobes. Submentum with six setae anteriorly. Genae with a single ventral seta one each side. Antennae moderate in length, with about 2.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 3 and 4 about equal in length, antennomere 2 slightly shorter.
</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6450FE2F726A" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">Pronotum. Slightly narrowed (ratio PW/PL = 1.28), with greatest width anterior to middle, only slightly narrowed posteriorly; lateral margins rounded, straightened for a short distance posteriorly, then slightly sinuate just anterior to basal angles, the latter small, sharp and rectangular; basal margin with median region slightly and convexly projected posteriorly. Disc convex, with median longitudinal impression distinct, markedly deepened in median basal area, not extended anteriorly to apical margin; basal foveae moderate in size and depth; median basal area delimited laterally by a pair of short, oblique furrows, smooth except for a pair of paramedial longitudinal foveae. Single midlateral setae on each side inserted anterior to middle; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6532FB8D716C" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">Elytra. Elongate ovoid, slightly narrowed anteriorly, humeri rounded and only slightly evident. Disc moderately convex, slightly flattened in sutural area; striae finely impressed, finely punctate, striae 1 to 2 or 3 more deeply impressed, more lateral striae more or less effaced but still evident, stria 5 not more deeply impressed at its base. Parascutellar striole present, deeply impressed and long. Recurrent stria long, deeply impressed, abruptly terminated on interval 7 slightly anterior to the insertion point of preapical seta. Two discal setiferous pores present, inserted at anterior one-sixth and near middle, respectively, on interval 3 near stria 3. Preapical seta present, inserted 1.5 times as far from apical elytral margin as from sutural margin on interval 3 next to stria 2.</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6634FD67710C" blockId="86.[123,1174,219,1725]" box="[171,681,1604,1629]" pageId="86" pageNumber="427">Legs. Short; protibiae with longitudinal furrow.</paragraph>
<paragraph id="8BC0368BFFC3FFDF3CFC6615FC8C71CC" blockId="86.[123,1174,219,1725]" pageId="86" pageNumber="427">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of female apically with two pairs of paramedial setae.</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC3FFD13CFC66D4FB5875CC" lastPageId="88" lastPageNumber="429" pageId="86" pageNumber="427" type="discussion">
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<emphasis id="B90BEA99FFC3FFDF3CFC66D4FEE671EC" bold="true" box="[171,296,1701,1725]" pageId="86" pageNumber="427">COMMENTS</emphasis>
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.— This species appears to be closely related to
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<emphasis id="B90BEA99FFC3FFDF3F7266D7FC5A71ED" box="[805,916,1701,1725]" italics="true" pageId="86" pageNumber="427">E. fortipes</emphasis>
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, but its members can be distinguished from those of the latter by the truncate, almost bifurcate, median tooth of the mentum (simple in
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<emphasis id="B90BEA99FFCDFFD13D76608DFE5F7644" box="[289,401,252,276]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members), elytral stria 5 not more deeply impressed basally (distinctly more deeply impressed in
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<emphasis id="B90BEA99FFCDFFD13DC2616CFDCA7664" box="[405,516,284,308]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members), recurrent stria longer (shorter in
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<emphasis id="B90BEA99FFCDFFD13FB6616CFB9E7664" box="[993,1104,284,308]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members) and terminated on interval 7 (terminated on interval
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members), and the preapical seta inserted 1.5 times as far from apical elytral margin as from sutural margin (inserted about equidistant from apical and sutural margins in
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<emphasis id="B90BEA99FFCDFFD13E27610EFD1176C7" box="[624,735,383,407]" italics="true" pageId="88" pageNumber="429">E. fortipes</emphasis>
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members).
</paragraph>
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33.
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<emphasis id="B90BEA99FFC2FFDE3D5C6646FDCA711A" box="[267,516,1591,1610]" italics="true" pageId="87" pageNumber="428">Epaphiotrechus fortipesoides</emphasis>
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; a. Dorsal habitus (CASENT1039089). b. Map of locality record (red circle) for
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<emphasis id="B90BEA99FFC2FFDE3C816620FE927133" box="[214,348,1616,1635]" italics="true" pageId="87" pageNumber="428">E. fortipesoides</emphasis>
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in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km.
</paragraph>
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</emphasis>
.— The unique
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female of this species was found on the ground under a stone or log in a managed conifer forest along a roadcut at an elevation of
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. No other trechines were found in this area.
</paragraph>
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</emphasis>
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<materialsCitation id="3B173CD6FFCDFFD13F4A6270FE23750B" county="western Tengchong County" location="This" pageId="88" pageNumber="429" specimenCount="1" stateProvince="Shan">
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.—
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.
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species is known only from a single female specimen from the
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locality in
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in the southern part of the Gaoligong
</materialsCitation>
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<collectingRegion id="49BBF869FFCDFFD13DA36232FDB7750B" box="[500,633,579,603]" country="Myanmar" name="Shan" pageId="88" pageNumber="429">Shan region</collectingRegion>
. This locality is in
<location id="8EA06050FFCDFFD13F116232FC7D750B" LSID="urn:lsid:plazi:treatment:03D6879DFFC1FFD13C2C6307FB5875CC:8EA06050FFCDFFD13F116232FC7D750B" box="[838,947,579,603]" name="Core Area" pageId="88" pageNumber="429" stateProvince="Shan">Core Area</location>
6
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFCDFFD13CFC6212FB5875CC" blockId="88.[123,1174,219,668]" pageId="88" pageNumber="429">
<emphasis id="B90BEA99FFCDFFD13CFC6212FD92752A" bold="true" box="[171,604,611,635]" pageId="88" pageNumber="429">
<smallCapsWord id="8D26A057FFCDFFD13CFC6212FEDC752A" baselines="629,629" box="[171,274,611,635]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="88" pageNumber="429">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFCDFFD13D4D6216FE0F752A" baselines="629" box="[282,449,615,634]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="88" pageNumber="429">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFCDFFD13D9F6216FD92752A" baselines="629" box="[456,604,615,634]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="88" pageNumber="429">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Tengchong County in the southern part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFCDFFD13FDC62F5FB8B75CC" box="[907,1093,644,668]" country="China" name="Yunnan" pageId="88" pageNumber="429">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFCDFFD1381862F5FB5C75CC" box="[1103,1170,644,668]" name="China" pageId="88" pageNumber="429">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,537 @@
<document id="F6DC32937A46B32C35A1F785C562BFBB" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFC5FFDA3C2C64EDFB4F73A6" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechepaphiopsis unipilosa Deuve and Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="424" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="421" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F51E816054171FF00CFF9BFB95A2E918">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:role id="EAC53E1F9BF939B926DDE8565319BC6D">
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<mods:namePart id="841F9B6855EACCF415DD78D05644F24E">Deuve, Thierry</mods:namePart>
<mods:affiliation id="A73907B10B1B1FDF4FAA317B293FF77F">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="5ED39F8DE80D8C4367E4EBCA3FAC670D">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="2A8BB445ED5BAD0B076A0537DB338DC5">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="5630876A11E696FC812BA0F5EA9BD4BA" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="E98373F35B6CBBE21A5B44A3DA9C8298" type="personal">
<mods:role id="5BD037A7B9CEC5862F0BDEE4388B663A">
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</mods:role>
<mods:namePart id="21EE327130BFB01A679F2A0148BCF557">Liang, Hongbin</mods:namePart>
<mods:affiliation id="09FBCF602D1D467A3828E0735AA3D278">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:date id="C82D732A489A0AC7E95A28E5EC3FCB04">2016</mods:date>
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<treatment id="03D6879DFFC5FFDA3C2C64EDFB4F73A6" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6" httpUri="http://treatment.plazi.org/id/03D6879DFFC5FFDA3C2C64EDFB4F73A6" lastPageId="83" lastPageNumber="424" pageId="80" pageNumber="421">
<subSubSection id="C3656500FFC5FFD93C2C64EDFD1673E4" box="[123,728,1180,1204]" pageId="80" pageNumber="421" type="nomenclature">
<paragraph id="8BC0368BFFC5FFD93C2C64EDFD1673E4" blockId="80.[123,728,1180,1234]" box="[123,728,1180,1204]" pageId="80" pageNumber="421">
<heading id="D08881E7FFC5FFD93C2C64EDFD1673E4" box="[123,728,1180,1204]" fontSize="10" level="2" pageId="80" pageNumber="421" reason="4">
<emphasis id="B90BEA99FFC5FFD93C2C64EDFD1673E4" bold="true" box="[123,728,1180,1204]" pageId="80" pageNumber="421">
<taxonomicName id="4C7F4D08FFC5FFD93C2C64EDFDBB73E4" ID-CoL="8MBQ3" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,629,1180,1204]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFC5FFD93C2C64EDFE6773E4" bold="true" box="[123,425,1180,1204]" italics="true" pageId="80" pageNumber="421">Trechepaphiopsis unipilosa</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFC5FFD93ED564EDFD1673E4" box="[642,728,1180,1204]" pageId="80" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC5FFD93C2C64CDFE9C7382" box="[123,338,1212,1234]" pageId="80" pageNumber="421" type="description">
<paragraph id="8BC0368BFFC5FFD93C2C64CDFE9C7382" blockId="80.[123,728,1180,1234]" box="[123,338,1212,1234]" pageId="80" pageNumber="421">
(
<figureCitation id="13442A0EFFC5FFD93CD564CDFF1D7381" box="[130,211,1212,1234]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[137,1176,216,1573]" captionTargetId="figure-11@82.[120,1176,216,1573]" captionTargetPageId="82" captionText="FIGURE 31. Trechepaphiopsis unipilosa sp. nov.; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for T. unipilosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="80" pageNumber="421">Figs. 31</figureCitation>
,
<figureCitation id="13442A0EFFC5FFD93C8A64CDFECD7382" box="[221,259,1212,1234]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="80" pageNumber="421">40b</figureCitation>
,
<figureCitation id="13442A0EFFC5FFD93D5964CDFE857382" box="[270,331,1212,1234]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="80" pageNumber="421">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC5FFD83CFC6494FF017526" lastPageId="81" lastPageNumber="422" pageId="80" pageNumber="421" type="materials_examined">
<paragraph id="8BC0368BFFC5FFD83CFC6494FD2776E4" blockId="80.[123,1174,1253,1726]" lastBlockId="81.[123,1174,219,1725]" lastPageId="81" lastPageNumber="422" pageId="80" pageNumber="421">
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<smallCapsWord id="8D26A057FFC5FFD93CBA6499FE9473AB" baselines="1270" box="[237,346,1256,1275]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="80" pageNumber="421">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFC5FFD93DD46494FC95722D" collectingDate="2004-05-07" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="3100" latitude="27.18416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" pageId="80" pageNumber="421" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFC5FFD93DD46494FE2773AD" box="[387,489,1253,1277]" pageId="80" pageNumber="421" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1023757”/ “
<collectingCountry id="F368761BFFC5FFD93FAA6494FB9873AD" box="[1021,1110,1253,1277]" name="China" pageId="80" pageNumber="421">CHINA</collectingCountry>
,
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,
<collectingCounty id="62A14E07FFC5FFD93CE56574FE90724D" box="[178,350,1285,1309]" pageId="80" pageNumber="421">Fugong County</collectingCounty>
,
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above Shibali on Shibali Road
</locationDeviation>
,
<location id="8EA06050FFC5FFD93FAD6574FF24726C" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC5FFD93FAD6574FF24726C" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" name="North Fork of Yamu He" pageId="80" pageNumber="421" stateProvince="Yunnan">North Fork of Yamu He</location>
,
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/
<geoCoordinate id="EE4B504CFFC5FFD93DDE6554FDC4726D" box="[393,522,1317,1341]" degrees="98.72026" direction="east" orientation="longitude" pageId="80" pageNumber="421" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFC5FFD93E6F6554FD58726C" box="[568,662,1317,1341]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="80" pageNumber="421" unit="m" value="3100.0">
<elevation id="0052D1B8FFC5FFD93E6F6554FD58726C" box="[568,662,1317,1341]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="80" pageNumber="421" unit="m" value="3100.0">3100 m</elevation>
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,
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<collectingDate id="EF85E9A3FFC5FFD93EFE6554FCF5726D" box="[681,827,1317,1341]" pageId="80" pageNumber="421" value="2004-05-07">7 May 2004</collectingDate>
</date>
Stop #DHK-2004-038A D.H.
<taxonomicName id="4C7F4D08FFC5FFD93C2C6534FF39720D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,1349,1373]" pageId="80" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFC5FFD93D556534FE51720D" box="[258,415,1349,1373]" pageId="80" pageNumber="421">C.E. Griswold</collectorName>
, Liang H.-B., &amp; Zhu B.-X. collectors”/ “
<typeStatus id="54C48829FFC5FFD93F016534FC2B720D" box="[854,997,1349,1373]" pageId="80" pageNumber="421" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFC5FFD93FBA6534FE77722D" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFC5FFD93FBA6534FECD722D" italics="true" pageId="80" pageNumber="421">Trechepaphiopsis unipilosa</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFC5FFD93D916517FDD5722D" box="[454,539,1382,1405]" pageId="80" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FFC5FFD93F3F6517FE3472ED" collectionCode="CAS, IOZ" country="China" pageId="80" pageNumber="421" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFC5FFD93F3F6517FC1E722E" box="[872,976,1382,1406]" pageId="80" pageNumber="421" type="paratype">Paratypes</typeStatus>
(at total of 17):
<specimenCount id="9D79FD02FFC5FFD938DF6514FF0072CD" count="2" pageId="80" pageNumber="421" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFC5FFD93D5665F4FEF972CD" box="[257,311,1413,1437]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="80" pageNumber="421" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFC5FFD93D1165F4FEBA72CD" box="[326,372,1413,1437]" pageId="80" pageNumber="421">IOZ</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFC5FFD93E1E65F4FD6772CD" box="[585,681,1413,1437]" pageId="80" pageNumber="421" type="holotype">holotype</typeStatus>
, except first label “CASENT 1023746” and “CASENT 1023758”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FFC5FFD93E5D65D4FC88716D" collectingDate="2005-08-08" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; D. Z. Dong &amp; J. F. Zhang" country="China" county="Fugong County" elevation="3100" latitude="27.18326" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" pageId="80" pageNumber="421" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFC5FFD93E5D65D4FD9672ED" box="[522,600,1445,1469]" count="1" pageId="80" pageNumber="421" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFC5FFD93EC065D4FCCD72ED" box="[663,771,1445,1469]" count="2" pageId="80" pageNumber="421" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFC5FFD93F6E65D4FCA172ED" box="[825,879,1445,1469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="80" pageNumber="421" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFC5FFD93FD765D4FC6072ED" box="[896,942,1445,1469]" pageId="80" pageNumber="421">IOZ</collectionCode>
) labeled: “CASENT 1017488” and “CASENT 10174896” to “CASENT 1017490”, respectively/ “
<collectingCountry id="F368761BFFC5FFD93F8165B4FBE1728D" box="[982,1071,1477,1501]" name="China" pageId="80" pageNumber="421">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFC5FFD9386A65B7FB5F728D" box="[1085,1169,1478,1501]" country="China" name="Yunnan" pageId="80" pageNumber="421">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFC5FFD93C2C6597FEED72AD" box="[123,291,1509,1534]" pageId="80" pageNumber="421">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFC5FFD93D7A6594FDDC72AD" box="[301,530,1509,1533]" pageId="80" pageNumber="421">Lumadeng Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FFC5FFD93E4B6594FC7A72AD" box="[540,948,1509,1533]" location="Shibali on Shibali Road" pageId="80" pageNumber="421">
<quantity id="4C879B6EFFC5FFD93E4B6594FDA872AD" box="[540,614,1509,1533]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="80" pageNumber="421" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
, south bank of
<location id="8EA06050FFC5FFD938006594FE8C714D" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC5FFD938006594FE8C714D" country="China" county="Fugong County" latitude="27.18326" longLatPrecision="1" longitude="98.72002" municipality="Lumadeng Township" name="North Fork of Yamu He" pageId="80" pageNumber="421" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFC5FFD93D076674FE1F714D" box="[336,465,1541,1565]" degrees="27.18326" direction="north" orientation="latitude" pageId="80" pageNumber="421" precision="1" value="27.18326">N27.18326°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFC5FFD93DB46674FDAA714D" box="[483,612,1541,1565]" degrees="98.72002" direction="east" orientation="longitude" pageId="80" pageNumber="421" precision="1" value="98.72002">E98.72002°</geoCoordinate>
”/
<quantity id="4C879B6EFFC5FFD93ED76674FD18714D" box="[640,726,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="80" pageNumber="421" unit="m" value="3100.0">
<elevation id="0052D1B8FFC5FFD93ED76674FD18714D" box="[640,726,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="80" pageNumber="421" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFC5FFD93EB26674FC46714D" box="[741,904,1541,1565]" pageId="80" pageNumber="421" value="2005-08-08">
<collectingDate id="EF85E9A3FFC5FFD93EB26674FC46714D" box="[741,904,1541,1565]" pageId="80" pageNumber="421" value="2005-08-08">8 August 2005</collectingDate>
</date>
Stop #DHK-2005-067B D.H.
<taxonomicName id="4C7F4D08FFC5FFD93CE26654FEFF716D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[181,305,1573,1597]" pageId="80" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFC5FFD93D6C6657FE7B716D" box="[315,437,1573,1597]" pageId="80" pageNumber="421">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFC5FFD93DE86657FDFA716E" box="[447,564,1574,1598]" pageId="80" pageNumber="421">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFC5FFD93E0E6657FD06716D" box="[601,712,1573,1597]" pageId="80" pageNumber="421">J.F. Zhang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFC5FFD93F076657FB5C71CD" collectingDate="2005-08-16" collectionCode="CAS, IOZ, MNHN" collectorName="P. Paquin" country="China" county="Fugong County" elevation="3250" latitude="27.20055" location="Lishadi Township" longLatPrecision="1" longitude="98.71399" pageId="80" pageNumber="421" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFC5FFD93F076657FC6F716D" box="[848,929,1573,1597]" count="7" pageId="80" pageNumber="421" type="male">7 males</specimenCount>
and
<specimenCount id="9D79FD02FFC5FFD93F836654FBFF716D" box="[980,1073,1573,1597]" count="1" pageId="80" pageNumber="421" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFC5FFD9380C6654FB5F716D" box="[1115,1169,1573,1597]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="80" pageNumber="421" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFC5FFD93C2C6634FF64710D" box="[123,170,1605,1629]" pageId="80" pageNumber="421">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFC5FFD93CEF6634FEC5710D" box="[184,267,1605,1629]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="80" pageNumber="421">MNHN</collectionCode>
) labeled: “CASENT 1018375” to “CASENT 1018381” and “CASENT 1018382”, respectively/ “
<collectingCountry id="F368761BFFC5FFD93D446614FEA2712D" box="[275,364,1637,1661]" name="China" pageId="80" pageNumber="421">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFC5FFD93D216617FE04712D" box="[374,458,1638,1661]" country="China" name="Yunnan" pageId="80" pageNumber="421">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFC5FFD93D836617FDB2712D" box="[468,636,1637,1662]" pageId="80" pageNumber="421">Fugong County</collectingCounty>
,
<location id="8EA06050FFC5FFD93ED16614FC86712D" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC5FFD93ED16614FC86712D" box="[646,840,1637,1661]" country="China" county="Fugong County" latitude="27.20055" longLatPrecision="1" longitude="98.71399" name="Lishadi Township" pageId="80" pageNumber="421" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFC5FFD93F056614FF7871CD" location="Shibali on Shibali Road" pageId="80" pageNumber="421">
<quantity id="4C879B6EFFC5FFD93F056614FC5B712D" box="[850,917,1637,1661]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="80" pageNumber="421" unit="km" value="10.0">10 km</quantity>
W of Shibali on Shibali Road
</locationDeviation>
,
<quantity id="4C879B6EFFC5FFD93C9566F4FED871CD" box="[194,278,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.25" pageId="80" pageNumber="421" unit="m" value="3250.0">
<elevation id="0052D1B8FFC5FFD93C9566F4FED871CD" box="[194,278,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.25" pageId="80" pageNumber="421" unit="m" value="3250.0">3250 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFC5FFD93D1766F4FE0D71CD" box="[320,451,1669,1693]" degrees="27.20055" direction="north" orientation="latitude" pageId="80" pageNumber="421" precision="1" value="27.20055">N27.20055°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFC5FFD93D9B66F4FD8371CD" box="[460,589,1669,1693]" degrees="98.71399" direction="east" orientation="longitude" pageId="80" pageNumber="421" precision="1" value="98.71399">E98.71399°</geoCoordinate>
,
<date id="FFC1104BFFC5FFD93E0E66F4FCCB71CD" box="[601,773,1669,1693]" pageId="80" pageNumber="421" value="2005-08-16">
<collectingDate id="EF85E9A3FFC5FFD93E0E66F4FCCB71CD" box="[601,773,1669,1693]" pageId="80" pageNumber="421" value="2005-08-16">16 August 2005</collectingDate>
</date>
Stop #PP-3805
<collectorName id="268A535DFFC5FFD93FEF66F7FBD371CD" box="[952,1053,1669,1693]" pageId="80" pageNumber="421">P. Paquin</collectorName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFC5FFD83C2C66D4FDDB7643" collectingDate="2005-08-17" collectionCode="CAS" collectorName="P. Paquin" country="China" county="Fugong County" elevation="3221" lastPageId="81" lastPageNumber="422" latitude="27.20192" location="Lishadi Township" longLatPrecision="1" longitude="98.71329" pageId="80" pageNumber="421" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFC5FFD93C2C66D4FF0D71ED" box="[123,195,1701,1725]" count="1" pageId="80" pageNumber="421" type="male">1 male</specimenCount>
(in
<collectionCode id="ED6EAE4EFFC5FFD93CBA66D4FEEC71ED" box="[237,290,1701,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="80" pageNumber="421" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1018392”/ “
<collectingCountry id="F368761BFFC5FFD93ED166D4FD1171ED" box="[646,735,1701,1725]" name="China" pageId="80" pageNumber="421">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFC5FFD93EBE66D7FCF371ED" box="[745,829,1702,1725]" country="China" name="Yunnan" pageId="80" pageNumber="421">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFC5FFD93F1F66D7FC3E71ED" box="[840,1008,1701,1726]" pageId="80" pageNumber="421">Fugong County</collectingCounty>
,
<location id="8EA06050FFC5FFD83FAD66D4FF6477A3" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC5FFD83FAD66D4FF6477A3" country="China" county="Fugong County" lastPageId="81" lastPageNumber="422" latitude="27.20192" longLatPrecision="1" longitude="98.71329" name="Lishadi Township" pageId="80" pageNumber="421" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFC4FFD83CE360AAFD8177A3" box="[180,591,219,243]" location="Shibali on Shibali Road" pageId="81" pageNumber="422">
<quantity id="4C879B6EFFC4FFD83CE360AAFEC277A3" box="[180,268,219,243]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="81" pageNumber="422" unit="km" value="10.5">10.5 km</quantity>
W of Shibali on Shibali Road
</locationDeviation>
,
<quantity id="4C879B6EFFC4FFD83E0D60AAFD6377A3" box="[602,685,219,243]" metricMagnitude="3" metricUnit="m" metricValue="3.221" pageId="81" pageNumber="422" unit="m" value="3221.0">
<elevation id="0052D1B8FFC4FFD83E0D60AAFD6377A3" box="[602,685,219,243]" metricMagnitude="3" metricUnit="m" metricValue="3.221" pageId="81" pageNumber="422" unit="m" value="3221.0">3221 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFFC4FFD83E8360AAFC9777A3" box="[724,857,219,243]" degrees="27.20192" direction="north" orientation="latitude" pageId="81" pageNumber="422" precision="1" value="27.20192">N27.20192°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFC4FFD83F3660AAFC2C77A3" box="[865,994,219,243]" degrees="98.71329" direction="east" orientation="longitude" pageId="81" pageNumber="422" precision="1" value="98.71329">E98.71329°</geoCoordinate>
,
<date id="FFC1104BFFC4FFD83FBA60AAFB5B77A3" box="[1005,1173,219,243]" pageId="81" pageNumber="422" value="2005-08-17">
<collectingDate id="EF85E9A3FFC4FFD83FBA60AAFB5B77A3" box="[1005,1173,219,243]" pageId="81" pageNumber="422" value="2005-08-17">17 August 2005</collectingDate>
</date>
Stop #PP-4105
<collectorName id="268A535DFFC4FFD83D67608DFE547643" box="[304,410,251,275]" pageId="81" pageNumber="422">P. Paquin</collectorName>
collector”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFC4FFD83E71608AFD2A76E4" collectingDate="2004-05-06" collectionCode="CAS, IOZ" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="3290" latitude="27.20676" location="Lishadi Township" longLatPrecision="1" longitude="98.71763" pageId="81" pageNumber="422" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFC4FFD83E71608AFDB17643" box="[550,639,251,275]" count="3" pageId="81" pageNumber="422" type="male">3 males</specimenCount>
(in
<collectionCode id="ED6EAE4EFFC4FFD83EEF608AFD207643" box="[696,750,251,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="81" pageNumber="422" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFC4FFD83F57608AFCE07643" box="[768,814,251,275]" pageId="81" pageNumber="422">IOZ</collectionCode>
) labeled: “CASENT 1023607”, “CASENT 1023608” and “CASENT 1023820”, respectively/ “
<collectingCountry id="F368761BFFC4FFD83F74616DFCB27664" box="[803,892,284,308]" name="China" pageId="81" pageNumber="422">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFC4FFD83FDF616DFC127663" box="[904,988,284,307]" country="China" name="Yunnan" pageId="81" pageNumber="422">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFC4FFD83FBF616DFB5C7664" box="[1000,1170,284,308]" pageId="81" pageNumber="422">Fugong County</collectingCounty>
,
<location id="8EA06050FFC4FFD83C2C614DFEF37604" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC4FFD83C2C614DFEF37604" box="[123,317,316,340]" country="China" county="Fugong County" latitude="27.20676" longLatPrecision="1" longitude="98.71763" name="Lishadi Township" pageId="81" pageNumber="422" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFC4FFD83D10614AFD257604" box="[327,747,315,340]" location="Shibali on Shibali Road" pageId="81" pageNumber="422">
<quantity id="4C879B6EFFC4FFD83D10614AFE537604" box="[327,413,315,340]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="81" pageNumber="422" unit="km" value="11.5">11.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFC4FFD83EA2614DFCB77604" box="[757,889,316,340]" degrees="27.20676" direction="north" orientation="latitude" pageId="81" pageNumber="422" precision="1" value="27.20676">N27.20676°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFC4FFD83FD5614DFBCC7604" box="[898,1026,316,340]" degrees="98.71763" direction="east" orientation="longitude" pageId="81" pageNumber="422" precision="1" value="98.71763">E98.71763°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFC4FFD8387F614DFBB37603" box="[1064,1149,316,340]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="81" pageNumber="422" unit="m" value="3290.0">
<elevation id="0052D1B8FFC4FFD8387F614DFBB37603" box="[1064,1149,316,340]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="81" pageNumber="422" unit="m" value="3290.0">3290 m</elevation>
</quantity>
,
<date id="FFC1104BFFC4FFD838DF614DFF267624" pageId="81" pageNumber="422" value="2004-05-06">
<collectingDate id="EF85E9A3FFC4FFD838DF614DFF267624" pageId="81" pageNumber="422" value="2004-05-06">6 May 2004</collectingDate>
</date>
Stop #DHK-2004-036 D.H.
<taxonomicName id="4C7F4D08FFC4FFD83E48612DFD557624" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[543,667,348,372]" pageId="81" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFC4FFD83EF1612DFC8D7624" box="[678,835,348,372]" pageId="81" pageNumber="422">C.E. Griswold</collectorName>
, Liang H.-B., &amp; Zhu B.-X. collectors”. All
<typeStatus id="54C48829FFC4FFD83D5C610CFEBF76C5" box="[267,369,381,405]" pageId="81" pageNumber="422" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFC4FFD83E8E610DFCAE76C4" box="[729,864,380,404]" pageId="81" pageNumber="422" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFC4FFD83F3B610DFEEB76E4" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="422" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFC4FFD83F3B610DFB5B76C4" box="[876,1173,380,404]" italics="true" pageId="81" pageNumber="422">Trechepaphiopsis unipilosa</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFC4FFD83D6761ECFE4D76E4" box="[304,387,413,436]" pageId="81" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFC4FFD83CFC61CCFBC876A4" blockId="81.[123,1174,219,1725]" pageId="81" pageNumber="422">
<materialsCitation id="3B173CD6FFC4FFD83CFC61CCFBCC76A4" country="China" county="Fugong County" elevation="3100" latitude="27.18416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" pageId="81" pageNumber="422" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFC4FFD83CFC61CCFE957683" bold="true" box="[171,347,445,469]" pageId="81" pageNumber="422">
<smallCapsWord id="8D26A057FFC4FFD83CFC61CCFF2A7683" baselines="463,462" box="[171,228,445,469]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="81" pageNumber="422">
<typeStatus id="54C48829FFC4FFD83CFC61CCFF2A7683" box="[171,228,445,469]" pageId="81" pageNumber="422">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFC4FFD83CA761B1FE957683" baselines="462" box="[240,347,448,467]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="81" pageNumber="422">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFC4FFD83DD061CCFE077685" box="[391,457,445,469]" name="China" pageId="81" pageNumber="422">China</collectingCountry>
,
<collectingRegion id="49BBF869FFC4FFD83D8F61CCFDE27684" box="[472,556,445,468]" country="China" name="Yunnan" pageId="81" pageNumber="422">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFC4FFD83E6B61CCFD267685" box="[572,744,445,469]" pageId="81" pageNumber="422">Fugong County</collectingCounty>
,
<taxonomicName id="4C7F4D08FFC4FFD83EA061CCFC237685" authority="Township" authorityName="Township" box="[759,1005,445,469]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="422" phylum="Arthropoda" rank="species" species="shiyueliang">Shiyueliang Township</taxonomicName>
,
<locationDeviation id="9A722FA9FFC4FFD83FAB61CDFEB376A5" location="Shibali on Shibali Road" pageId="81" pageNumber="422">
<quantity id="4C879B6EFFC4FFD83FAB61CDFB847685" box="[1020,1098,444,469]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="81" pageNumber="422" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<location id="8EA06050FFC4FFD83DDE61ACFD4276A4" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC4FFD83DDE61ACFD4276A4" box="[393,652,477,501]" country="China" county="Fugong County" latitude="27.18416" longLatPrecision="1" longitude="98.72026" name="North Fork of Yamu He" pageId="81" pageNumber="422" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFC4FFD83ECF61ACFCD476A5" box="[664,794,477,501]" degrees="27.18416" direction="north" orientation="latitude" pageId="81" pageNumber="422" precision="1" value="27.18416">N27.18416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFC4FFD83F7461ACFC6D76A5" box="[803,931,477,501]" degrees="98.72026" direction="east" orientation="longitude" pageId="81" pageNumber="422" precision="1" value="98.72026">E98.72026°</geoCoordinate>
,
<quantity id="4C879B6EFFC4FFD83FF961ACFBCC76A4" box="[942,1026,477,501]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="81" pageNumber="422" unit="m" value="3100.0">
<elevation id="0052D1B8FFC4FFD83FF961ACFBCC76A4" box="[942,1026,477,501]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="81" pageNumber="422" unit="m" value="3100.0">3100 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFC4FFD83CFC618CFF017526" blockId="81.[123,1174,219,1725]" pageId="81" pageNumber="422">
<emphasis id="B90BEA99FFC4FFD83CFC618CFE3E7544" bold="true" box="[171,496,509,533]" pageId="81" pageNumber="422">
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<smallCapsWord id="8D26A057FFC4FFD83D626271FE9F7543" baselines="526" box="[309,337,512,531]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="81" pageNumber="422">OF</smallCapsWord>
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</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFC4FFD83EA1618CFC977545" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[758,857,509,533]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="422" phylum="Arthropoda" rank="species" species="unipilosa">
<emphasis id="B90BEA99FFC4FFD83EA1618CFC977545" box="[758,857,509,533]" italics="true" pageId="81" pageNumber="422">unipilosa</emphasis>
</taxonomicName>
, is an adjective derived from the Latin word,
<emphasis id="B90BEA99FFC4FFD83D7F626FFE947565" box="[296,346,542,565]" italics="true" pageId="81" pageNumber="422">unus</emphasis>
, meaning one, and the Greek word,
<emphasis id="B90BEA99FFC4FFD83EB0626CFCD17565" box="[743,799,541,565]" italics="true" pageId="81" pageNumber="422">πίΛΟς</emphasis>
(transliterated into Latin as
<emphasis id="B90BEA99FFC4FFD83805626CFB487565" box="[1106,1158,541,565]" italics="true" pageId="81" pageNumber="422">pilus</emphasis>
), meaning hair. The name refers to the single discal seta found on the elytra of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC4FFD83CFC620FFB5874C8" pageId="81" pageNumber="422" type="diagnosis">
<paragraph id="8BC0368BFFC4FFD83CFC620FFB5874C8" blockId="81.[123,1174,219,1725]" pageId="81" pageNumber="422">
<smallCapsWord id="8D26A057FFC4FFD83CFC620FFEEE75C4" baselines="656,655" box="[171,288,638,662]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="81" pageNumber="422">
<emphasis id="B90BEA99FFC4FFD83CFC620FFEEE75C4" bold="true" box="[171,288,638,662]" pageId="81" pageNumber="422">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFC4FFD83E10620FFD6D75C6" box="[583,675,638,662]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[137,1176,216,1573]" captionTargetId="figure-11@82.[120,1176,216,1573]" captionTargetPageId="82" captionText="FIGURE 31. Trechepaphiopsis unipilosa sp. nov.; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for T. unipilosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="81" pageNumber="422">Fig. 31a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size small (BL =
<quantity id="4C879B6EFFC4FFD8387862EFFF667587" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="81" pageNumber="422" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7 to 3.1 mm</quantity>
), apterous; body color reddish brown, dorsum shiny, pronotum and elytra slightly iridescent; eyes small but convex; tempora convex and sparsely pubescent; pronotum transverse (ratio PW/PL = 1.42), with basal angles obtuse and rounded; elytra convex, medial striae (1 to 5 or 6) distinct, finely punctate, more lateral striae more or less effaced; recurrent stria terminated anteriorly on interval 6; only one discal setiferous pore present, inserted at anterior one-third next to stria 3; preapical seta present, inserted next to stria 2; median lobe of male aedeagus (
<figureCitation id="13442A0EFFC4FFD83F95632EFBD37427" box="[962,1053,863,887]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[137,1176,216,1573]" captionTargetId="figure-11@82.[120,1176,216,1573]" captionTargetPageId="82" captionText="FIGURE 31. Trechepaphiopsis unipilosa sp. nov.; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for T. unipilosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="81" pageNumber="422">Fig. 31b</figureCitation>
) with apex slender and rounded, endophallus with a faintly sclerotized scaly area distinctly narrowed distally.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC4FFDA3CFC63D1FCA175C4" lastPageId="83" lastPageNumber="424" pageId="81" pageNumber="422" type="description">
<paragraph id="8BC0368BFFC4FFD83CFC63D1FEA07349" blockId="81.[123,1174,219,1725]" pageId="81" pageNumber="422">
<smallCapsWord id="8D26A057FFC4FFD83CFC63D1FEF474E6" baselines="946,945" box="[171,314,928,952]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="81" pageNumber="422">
<emphasis id="B90BEA99FFC4FFD83CFC63D1FEF474E6" bold="true" box="[171,314,928,952]" pageId="81" pageNumber="422">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size small, BL =
<quantity id="4C879B6EFFC4FFD83E4863D1FD7774E7" box="[543,697,928,952]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="81" pageNumber="422" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7 to 3.1 mm</quantity>
. Body color reddish brown, antennmae concolorous, legs slightly paler, yellowish brown, palpi paler yellow; dorsum shiny, pronotum and elytra slightly iridescent, head slightly alutaceous from more deeply impressed, irregularly isodiametric microsculpture.
</paragraph>
<paragraph id="8BC0368BFFC4FFD83CFC6450FED2726A" blockId="81.[123,1174,219,1725]" pageId="81" pageNumber="422">
Head. Broad; eyes small but convex, their diameter about equal to length of tempora, the latter short, distinctly convex and sparsely pubescent Frons convex, frontal furrows sharply impressed and rounded, not attenuated posterior to eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly emarginate.
<collectingRegion id="49BBF869FFC4FFD83D5764D3FE9473E9" box="[256,346,1186,1209]" country="Myanmar" name="Chin" pageId="81" pageNumber="422">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFC4FFD83EDA64D3FD2973E9" box="[653,743,1186,1209]" country="Myanmar" name="Chin" pageId="81" pageNumber="422">Mentum</collectingRegion>
with medial tooth apically bifid, about half the length of the lateral lobes. Submentum with six setae anteriorly. Gula broad. Genae with a single ventral seta one each side. Antennae short, with only about two antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 4 about equal in length, antennomere 3 slightly longer.
</paragraph>
<paragraph id="8BC0368BFFC4FFD83CFC6532FC38716C" blockId="81.[123,1174,219,1725]" pageId="81" pageNumber="422">Pronotum. Transverse (ratio PW/PL = 1.42), with greatest width at anterior one-third; lateral margins rounded, slightly more so anteriorly, not or only slightly straighten posteriorly just anteri- or to basal angles, the latter obtuse and blunted. Disc convex and smooth, median longitudinal impression finely impressed, continuous from middle of apical median swelling to posterior margin; basal foveae small and shallow, faintly impressed; median basal area well defined but short and transverse, smooth or with a few small longitudinal foveae. Lateral borders slender and finely relexed dorsally, lateral grooves deeply impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFC4FFDA3CFC6635FF2676E4" blockId="81.[123,1174,219,1725]" lastBlockId="83.[123,1174,219,1271]" lastPageId="83" lastPageNumber="424" pageId="81" pageNumber="422">Elytra. Ovoid only slightly elongate, not or only very slightly more narrowed anteriorly than posteriorly, humeri distinct but rounded. Disc convex, striae finely punctate, only medial striae 1 to 3 distinctly impressed, more lateral striae more or less effaced, but still evident in most specimens. Parascutellar striole present, distinct but short. Recurrent stria deeply impressed, its anterior end abruptly terminated in the posterior end of stria 5. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Only one discal setiferous pore present, inserted at anterior two-fifth of elytral length next to stria 3. Only one discal setiferous pore present, inserted slightly anterior to middle next to stria 3. Preapical seta present, inserted on interval 3 next to stria 2 and about equidistant from apical and sutural elytral margin. Umbilicate setal series with setae of humeral group equidistance from each other and setae of median group inserted slightly posterior to middle.</paragraph>
<caption id="DF006603FFC7FFDB3CFC6640FDC47126" ID-DOI="http://doi.org/10.5281/zenodo.13155342" ID-Zenodo-Dep="13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="82" pageNumber="423" startId="82.[171,183,1585,1604]" targetBox="[137,1176,216,1573]" targetPageId="82" targetType="figure">
<paragraph id="8BC0368BFFC7FFDB3CFC6640FDC47126" blockId="82.[123,1174,1584,1655]" pageId="82" pageNumber="423">
<smallCapsWord id="8D26A057FFC7FFDB3CFC6640FF267112" baselines="1599,1598" box="[171,232,1585,1604]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="82" pageNumber="423">FIGURE</smallCapsWord>
31.
<taxonomicName id="4C7F4D08FFC7FFDB3D5B6641FE387113" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[268,502,1584,1603]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="423" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFC7FFDB3D5B6641FE387113" box="[268,502,1584,1603]" italics="true" pageId="82" pageNumber="423">Trechepaphiopsis unipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFC7FFDB3DAB6640FD8F7114" box="[508,577,1585,1604]" pageId="82" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFC7FFDB3F3C663AFC01710D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[875,975,1610,1629]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="423" phylum="Arthropoda" rank="species" species="unipilosa">
<emphasis id="B90BEA99FFC7FFDB3F3C663AFC01710D" box="[875,975,1610,1629]" italics="true" pageId="82" pageNumber="423">T. unipilosa</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFC6FFDA3CFC61CDFE9E76A4" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFC6FFDA3CFC618DFB857564" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFC6FFDA3CFC624DFCA175C4" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFC6FFDA3DA9624DFD937504" box="[510,605,572,596]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[137,1176,216,1573]" captionTargetId="figure-11@82.[120,1176,216,1573]" captionTargetPageId="82" captionText="FIGURE 31. Trechepaphiopsis unipilosa sp. nov.; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for T. unipilosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="83" pageNumber="424">Fig. 31b</figureCitation>
) with sagittal aileron very small, shaft moderately broad basally, then gradually narrowed apically to a narrowly rounded apex; endophallus with a faintly sclerotized scaly area distinctly elongate and narrowed distally.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC6FFDA3CFC62EDFB4F73A6" pageId="83" pageNumber="424" type="discussion">
<paragraph id="8BC0368BFFC6FFDA3CFC62EDFD457465" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">
<smallCapsWord id="8D26A057FFC6FFDA3CFC62EDFEE675E3" baselines="686,686" box="[171,296,668,692]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC62EDFEE675E3" bold="true" box="[171,296,668,692]" pageId="83" pageNumber="424">COMMENTS</emphasis>
</smallCapsWord>
.— This species appears to be very closely related (perhaps sister species) to
<taxonomicName id="4C7F4D08FFC6FFDA3C2C62CCFEED7585" authorityName="Deuve &amp; Kavanaugh" authorityYear="2016" box="[123,291,701,725]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="monochaetus">
<emphasis id="B90BEA99FFC6FFDA3C2C62CCFEED7585" box="[123,291,701,725]" italics="true" pageId="83" pageNumber="424">T. monochaetus</emphasis>
</taxonomicName>
, but its male members can be distinguished from those of the latter by having the base of the median lobe of the male aedeagus narrower (broader in
<taxonomicName id="4C7F4D08FFC6FFDA3F0B62AFFC3475A5" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[860,1018,733,757]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFC6FFDA3F0B62AFFC3475A5" box="[860,1018,733,757]" italics="true" pageId="83" pageNumber="424">T. monochaeta</emphasis>
</taxonomicName>
members, see
<figureCitation id="13442A0EFFC6FFDA3C2C628CFF197445" box="[123,215,765,789]" captionStart="FIGURE" captionStartId="79.[171,183,1585,1604]" captionTargetBox="[126,1176,216,1572]" captionTargetId="figure-15@79.[120,1176,216,1573]" captionTargetPageId="79" captionText="FIGURE 30. Trechepaphiopsis monochaeta sp. nov.; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for T. monochaeta in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="83" pageNumber="424">Fig. 30b</figureCitation>
) and the scaly sclerotized area of the endophallus distinctly narrowed distally (of more equal width throughout in
<taxonomicName id="4C7F4D08FFC6FFDA3DCF636FFDFB7465" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[408,565,797,821]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFC6FFDA3DCF636FFDFB7465" box="[408,565,797,821]" italics="true" pageId="83" pageNumber="424">T. monochaeta</emphasis>
</taxonomicName>
males).
</paragraph>
<paragraph id="8BC0368BFFC6FFDA3CFC634CFDCF7366" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC634CFE687403" bold="true" box="[171,422,829,853]" pageId="83" pageNumber="424">
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<smallCapsWord id="8D26A057FFC6FFDA3D456331FE687403" baselines="846" box="[274,422,832,851]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="83" pageNumber="424">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been collected by sifting leaf litter from forests of scattered, large
<taxonomicName id="4C7F4D08FFC6FFDA3D86632CFDFA7425" box="[465,564,861,885]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="83" pageNumber="424" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEA99FFC6FFDA3D86632CFDC27425" box="[465,524,861,885]" italics="true" pageId="83" pageNumber="424">Abies</emphasis>
sp.
</taxonomicName>
trees with a dense understory of bamboo or
<taxonomicName id="4C7F4D08FFC6FFDA3877632CFF2C74C5" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="83" pageNumber="424" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEA99FFC6FFDA3877632CFF6274C5" italics="true" pageId="83" pageNumber="424">Rhododendron</emphasis>
spp.
</taxonomicName>
at elevations ranging from
<quantity id="4C879B6EFFC6FFDA3E43630CFD0974C5" box="[532,711,893,917]" metricMagnitude="3" metricUnit="m" metricValue="3.2555" metricValueMax="3.29" metricValueMin="3.221" pageId="83" pageNumber="424" unit="m" value="3255.5" valueMax="3290.0" valueMin="3221.0">3221 to 3290 m</quantity>
. Specimens of
<emphasis id="B90BEA99FFC6FFDA3F27630CFF2974E5" italics="true" pageId="83" pageNumber="424">
<taxonomicName id="4C7F4D08FFC6FFDA3F27630CFB8974C5" box="[880,1095,893,917]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="shiyueliang">Trechus shiyueliang</taxonomicName>
and
<taxonomicName id="4C7F4D08FFC6FFDA38D5630FFF2974E5" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="shibalicus">T. shibalicus</taxonomicName>
</emphasis>
were collected in one or more of the same litter samples. Specimens of
<taxonomicName id="4C7F4D08FFC6FFDA3F8863EFFB9574E5" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[991,1115,925,949]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="unipilosa">
<emphasis id="B90BEA99FFC6FFDA3F8863EFFB9574E5" box="[991,1115,925,949]" italics="true" pageId="83" pageNumber="424">T. unipilosa</emphasis>
</taxonomicName>
were also collected at
<quantity id="4C879B6EFFC6FFDA3D6163CFFE447485" box="[310,394,958,982]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="83" pageNumber="424" unit="m" value="3100.0">3100 m</quantity>
elevation under stones at the open edges of a small stream draining a north-facing glacial cirque with a large snowfield in its basin (
<figureCitation id="13442A0EFFC6FFDA3F4F63AFFCB874A6" box="[792,886,990,1014]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="83" pageNumber="424">Fig. 40b</figureCitation>
). A specimen of
<taxonomicName id="4C7F4D08FFC6FFDA387D63AFFE987346" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="unisetulosa" status="sp. nov.">
<emphasis id="B90BEA99FFC6FFDA387D63AFFE987346" italics="true" pageId="83" pageNumber="424">Trechepaphiopsis unisetulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFC6FFDA3D37638FFE7B7345" box="[352,437,1022,1045]" pageId="83" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
was also collected in this area, at the upper limit of the altitudinal range of this last mentioned species.
</paragraph>
<paragraph id="8BC0368BFFC6FFDA3CFC644FFDCD73E6" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC644FFD4E7305" bold="true" box="[171,640,1086,1110]" pageId="83" pageNumber="424">
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<smallCapsWord id="8D26A057FFC6FFDA3D086430FE3D7304" baselines="1103" box="[351,499,1089,1108]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="83" pageNumber="424">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFC6FFDA3DAA6433FD827305" baselines="1104" box="[509,588,1090,1109]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="83" pageNumber="424">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFC6FFDA3E016433FD4E7305" baselines="1104" box="[598,640,1090,1109]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="83" pageNumber="424">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FFC6FFDA3EDD644FFCC17304" baselines="1104,1103" box="[650,783,1086,1110]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3EDD644FFCC17304" bold="true" box="[650,783,1086,1110]" pageId="83" pageNumber="424">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFC6FFDA3F4E644FFEAC73C6" location="We" pageId="83" pageNumber="424" specimenCount="36" specimenCount-female="5" specimenCount-male="13" stateProvince="Shan">
<smallCapsWord id="8D26A057FFC6FFDA3F4E644FFC9A7304" baselines="1104,1103" box="[793,852,1086,1110]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3F4E644FFC9A7304" bold="true" box="[793,852,1086,1110]" pageId="83" pageNumber="424">
<collectingRegion id="49BBF869FFC6FFDA3F4E644FFC9A7304" box="[793,852,1086,1110]" country="Myanmar" name="Shan" pageId="83" pageNumber="424">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFC6FFDA3FD7644FFC107306" box="[896,990,1086,1110]" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[137,1176,216,1573]" captionTargetId="figure-11@82.[120,1176,216,1573]" captionTargetPageId="82" captionText="FIGURE 31. Trechepaphiopsis unipilosa sp. nov.; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for T. unipilosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="83" pageNumber="424">Fig. 31c</figureCitation>
.
<location id="8EA06050FFC6FFDA3FBC644FFBC07305" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC6FFDA3FBC644FFBC07305" box="[1003,1038,1086,1109]" name="We" pageId="83" pageNumber="424" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFC6FFDA3C98642FFEAF7326" box="[207,353,1118,1142]" count="18" pageId="83" pageNumber="424" type="generic">18 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFC6FFDA3D26642FFE1F7326" box="[369,465,1118,1142]" count="13" pageId="83" pageNumber="424" type="male">13 males</specimenCount>
and
<specimenCount id="9D79FD02FFC6FFDA3E50642FFDBF7326" box="[519,625,1118,1142]" count="5" pageId="83" pageNumber="424" type="female">5 females</specimenCount>
) from sites on the eastern slope of the northcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFC6FFDA3D3E640FFE3173E6" county="Fugong County" location="Core Area" pageId="83" pageNumber="424" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFC6FFDA3D3E640FFE5073C6" box="[361,414,1150,1174]" country="Myanmar" name="Shan" pageId="83" pageNumber="424">Shan</collectingRegion>
in
<collectingCounty id="62A14E07FFC6FFDA3D96640EFDA773C6" box="[449,617,1150,1175]" pageId="83" pageNumber="424">Fugong County</collectingCounty>
(see
<typeStatus id="54C48829FFC6FFDA3EF6640EFD1873C7" box="[673,726,1151,1175]" pageId="83" pageNumber="424">Type</typeStatus>
material above for exact collection data). All of these sites are in
<location id="8EA06050FFC6FFDA3D2D64EFFE2673E6" LSID="urn:lsid:plazi:treatment:03D6879DFFC5FFDA3C2C64EDFB4F73A6:8EA06050FFC6FFDA3D2D64EFFE2673E6" box="[378,488,1182,1206]" county="Fugong County" name="Core Area" pageId="83" pageNumber="424" stateProvince="Shan">Core Area</location>
3
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFC6FFDA3CFC64CFFB4F73A6" blockId="83.[123,1174,219,1271]" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC64CFFD967385" bold="true" box="[171,600,1214,1238]" pageId="83" pageNumber="424">
<smallCapsWord id="8D26A057FFC6FFDA3CFC64CFFEDC7385" baselines="1232,1232" box="[171,274,1214,1238]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="83" pageNumber="424">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFC6FFDA3D4F64B3FE717385" baselines="1232" box="[280,447,1218,1237]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="83" pageNumber="424">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFC6FFDA3D9364B3FD967385" baselines="1232" box="[452,600,1218,1237]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="83" pageNumber="424">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Fugong County in the northcentral part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFC6FFDA3F2364AEFBE173A6" box="[884,1071,1246,1270]" country="China" name="Yunnan" pageId="83" pageNumber="424">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFC6FFDA386D64AFFBB373A6" box="[1082,1149,1246,1270]" name="China" pageId="83" pageNumber="424">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,164 @@
<document id="C549B0670340F2FCF52D1C8765575021" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFC6FFDD3D1E6568FB587563" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Epaphiotrechus Deuve &amp; Kavanaugh &amp; Liang 2016, gen. nov." docType="treatment" docVersion="2" lastPageNumber="425" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="424" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="68297D116BD5C20C2CB17353283636D4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="76B8FB2C7A1ABA47CBE09428C83FE0CB">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:name id="F8396336ACA249EBDF1B41F0D2C1DFD4" type="personal">
<mods:role id="0195162869DCE63BB24B84EEDF5F9A49">
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</mods:role>
<mods:namePart id="2F56285F18106C2C7FCF2266242B541C">Deuve, Thierry</mods:namePart>
<mods:affiliation id="E8494E3F4CBDCE03ADC55402E3A80799">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
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<mods:role id="FEBA491E520DE26FF2D4D5549B2C1A4E">
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</mods:role>
<mods:namePart id="01C92B58D389867381ED1E690166DEC1">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="0DB60B06B24B58C70644C2AEAC4C493A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="94043828CB513BC440EBB2276A2207D8" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="4AA335CD2BF51EBB86198A9481FEF8D3" type="personal">
<mods:role id="1E695A1C101D804464243695A90AD0ED">
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</mods:role>
<mods:namePart id="A98B3100D2E0F31A2B41AE97301EC8AA">Liang, Hongbin</mods:namePart>
<mods:affiliation id="D8DCBE82E97C62EE2F75FCD0B92ED6A2">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:part id="D63DBFB8FCA199A90B52DB5CFE1ED6C7">
<mods:date id="2BE853C691D5D40E5F59E79349D27CFF">2016</mods:date>
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<mods:number id="460E74D0FC8669DBCCA8B7F5B19CFC38">2016-10-14</mods:number>
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<treatment id="03D6879DFFC6FFDD3D1E6568FB587563" LSID="urn:lsid:plazi:treatment:03D6879DFFC6FFDD3D1E6568FB587563" httpUri="http://treatment.plazi.org/id/03D6879DFFC6FFDD3D1E6568FB587563" lastPageId="84" lastPageNumber="425" pageId="83" pageNumber="424">
<subSubSection id="C3656500FFC6FFDA3D1E6568FC097261" box="[329,967,1305,1329]" pageId="83" pageNumber="424" type="nomenclature">
<paragraph id="8BC0368BFFC6FFDA3D1E6568FC097261" blockId="83.[329,967,1305,1329]" box="[329,967,1305,1329]" pageId="83" pageNumber="424">
<heading id="D08881E7FFC6FFDA3D1E6568FC097261" box="[329,967,1305,1329]" centered="true" fontSize="11" level="2" pageId="83" pageNumber="424" reason="2">
<emphasis id="B90BEA99FFC6FFDA3D1E6568FC097261" bold="true" box="[329,967,1305,1329]" pageId="83" pageNumber="424">
Genus
<taxonomicName id="4C7F4D08FFC6FFDA3DCE6568FD587261" ID-CoL="8M7XY" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[409,662,1305,1329]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFC6FFDA3DCE6568FD867261" bold="true" box="[409,584,1305,1329]" italics="true" pageId="83" pageNumber="424">Epaphiotrechus</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFC6FFDA3F356568FC097261" box="[866,967,1305,1329]" pageId="83" pageNumber="424" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC6FFDA3CFC6535FD2C72CC" pageId="83" pageNumber="424" type="type_taxon">
<paragraph id="8BC0368BFFC6FFDA3CFC6535FD32720C" blockId="83.[123,1174,1348,1725]" box="[171,764,1348,1372]" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC6535FEF1720B" bold="true" box="[171,319,1348,1372]" pageId="83" pageNumber="424">
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<typeStatus id="54C48829FFC6FFDA3CFC6535FF2A720B" box="[171,228,1348,1372]" pageId="83" pageNumber="424">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFC6FFDA3CBB6539FEF1720B" baselines="1366" box="[236,319,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="83" pageNumber="424">SPECIES</smallCapsWord>
</emphasis>
.—
<taxonomicName id="4C7F4D08FFC6FFDA3D3F6535FD6F720C" ID-CoL="8M9KW" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[360,673,1348,1372]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="fortipesoides" status="sp. nov.">
<emphasis id="B90BEA99FFC6FFDA3D3F6535FD6F720C" box="[360,673,1348,1372]" italics="true" pageId="83" pageNumber="424">Epaphiotrechus fortipesoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFC6FFDA3EFF6534FD32720C" box="[680,764,1349,1372]" pageId="83" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFFC6FFDA3CFC6515FD2C72CC" blockId="83.[123,1174,1348,1725]" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC6515FDD5722C" bold="true" box="[171,539,1380,1404]" pageId="83" pageNumber="424">Derivation of genus group name</emphasis>
.— The genus group name (masculine) is a combination of two other trechine generic names,
<taxonomicName id="4C7F4D08FFC6FFDA3DBA65F5FD9C72CC" ID-CoL="8KWTX" authorityName="Samouelle" authorityYear="1819" box="[493,594,1412,1436]" class="Insecta" family="Carabidae" genus="Epaphius" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFC6FFDA3DBA65F5FD9C72CC" box="[493,594,1412,1436]" italics="true" pageId="83" pageNumber="424">Epaphius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFC6FFDA3ED065F5FD1472CC" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[647,730,1412,1436]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFC6FFDA3ED065F5FD1472CC" box="[647,730,1412,1436]" italics="true" pageId="83" pageNumber="424">Trechus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC6FFDD3CFC65D4FDA87603" lastPageId="84" lastPageNumber="425" pageId="83" pageNumber="424" type="diagnosis">
<paragraph id="8BC0368BFFC6FFDD3CFC65D4FDA87603" blockId="83.[123,1174,1348,1725]" lastBlockId="84.[123,1174,219,563]" lastPageId="84" lastPageNumber="425" pageId="83" pageNumber="424">
<smallCapsWord id="8D26A057FFC6FFDA3CFC65D4FEEE72EB" baselines="1463,1462" box="[171,288,1445,1469]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA3CFC65D4FEEE72EB" bold="true" box="[171,288,1445,1469]" pageId="83" pageNumber="424">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this genus (
<figureCitation id="13442A0EFFC6FFDA3E7C65D5FD4D72ED" box="[555,643,1444,1469]" captionStart="FIGURE" captionStartId="87.[171,183,1591,1610]" captionTargetBox="[120,1176,216,1574]" captionTargetId="figure-15@87.[120,1176,216,1575]" captionTargetPageId="87" captionText="FIGURE 33. Epaphiotrechus fortipesoides sp. nov.; a. Dorsal habitus (CASENT1039089). b. Map of locality record (red circle) for E. fortipesoides in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155346" httpUri="https://zenodo.org/record/13155346/files/figure.png" pageId="83" pageNumber="424">Fig. 33a</figureCitation>
) can be recognized by the following combination of character states: size large (BL =
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), apterous; body color dark piceous with elytral interval 1 and lateral areas of pronotum and elytra more or less reddish; dorsum shiny, distinctly iridescent, dorsal surface glabrous except for isolated fixed setae typical for trechines, eyes also glabrous; right mandible (
<figureCitation id="13442A0EFFC6FFDA3DC76654FE25716D" box="[400,491,1573,1597]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="83" pageNumber="424">
Fig.
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</figureCitation>
) with premolar not fused with retinaculum but closely associat- ed with the latter [possibly representative of an intermediate state in the evolution of the “bidentate” mandibular
<typeStatus id="54C48829FFC6FFDA3D176617FEA0712E" box="[320,366,1638,1662]" pageId="83" pageNumber="424">type</typeStatus>
(see discussion above for genus
<taxonomicName id="4C7F4D08FFC6FFDA3EA56614FC8B712D" authorityName="Clairville" authorityYear="1806" box="[754,837,1637,1661]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFC6FFDA3EA56614FC8B712D" box="[754,837,1637,1661]" italics="true" pageId="83" pageNumber="424">Trechus</emphasis>
</taxonomicName>
)], anterior tip of retinaculum enlarged as a distinct tooth and displaced anteriorly (but not quite as far as in members of
<taxonomicName id="4C7F4D08FFC6FFDA386066F4FEF071ED" pageId="83" pageNumber="424">
<emphasis id="B90BEA99FFC6FFDA386066F4FF2F71ED" italics="true" pageId="83" pageNumber="424">Trechepaphiopsis</emphasis>
species
</taxonomicName>
); pronotum with basal angles small and rectangular; elytra elongate, oblong, slightly flattened along the median suture area, striae finely impressed, crenulate or finely punctate, lateral striae partly effaced but striae 6 and 7 still evident; two discal setae present on interval 3 next to stria 3, inserted near the anterior one-sixth and near mid-elytral length, respectively; preapical seta present, inserted next to stria 2.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC1FFDD3CFC612AFB587563" pageId="84" pageNumber="425" type="discussion">
<paragraph id="8BC0368BFFC1FFDD3CFC612AFD9176A3" blockId="84.[123,1174,219,563]" pageId="84" pageNumber="425">
<smallCapsWord id="8D26A057FFC1FFDD3CFC612AFEE67622" baselines="365,365" box="[171,296,347,371]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3CFC612AFEE67622" bold="true" box="[171,296,347,371]" pageId="84" pageNumber="425">COMMENTS</emphasis>
</smallCapsWord>
.— When Uéno (1999) described
<taxonomicName id="4C7F4D08FFC1FFDD3ECD612AFC547623" box="[666,922,347,371]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B90BEA99FFC1FFDD3ECD612AFD237623" box="[666,749,347,371]" italics="true" pageId="84" pageNumber="425">Trechus</emphasis>
(
<emphasis id="B90BEA99FFC1FFDD3EA8612DFCF77623" box="[767,825,348,371]" italics="true" pageId="84" pageNumber="425">s. str.</emphasis>
)
<emphasis id="B90BEA99FFC1FFDD3F1E612AFC547623" box="[841,922,347,371]" italics="true" pageId="84" pageNumber="425">fortipes</emphasis>
</taxonomicName>
, he noted that it was a “strange species similar to certain
<taxonomicName id="4C7F4D08FFC1FFDD3DBA610AFDBF76C3" authorityName="Ueno" authorityYear="1953" box="[493,625,379,403]" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFC1FFDD3DBA610AFDBF76C3" box="[493,625,379,403]" italics="true" pageId="84" pageNumber="425">Epaphiopsis</emphasis>
</taxonomicName>
”, with the distinct premolar on the right mandible. However, characters of the male aedeagus, particularly the presence of a distinct endophallic sclerite, led him to consider this species as a basal member of genus
<taxonomicName id="4C7F4D08FFC1FFDD3F6E61CAFC427683" authorityName="Clairville" authorityYear="1806" box="[825,908,443,467]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFC1FFDD3F6E61CAFC427683" box="[825,908,443,467]" italics="true" pageId="84" pageNumber="425">Trechus</emphasis>
</taxonomicName>
in which the plesiomorphic right mandibular dentition was retained.
</paragraph>
<paragraph id="8BC0368BFFC1FFDD3CFC618AFB587563" blockId="84.[123,1174,219,563]" pageId="84" pageNumber="425">
<emphasis id="B90BEA99FFC1FFDD3CFC618AFE347543" bold="true" box="[171,506,507,531]" pageId="84" pageNumber="425">
<smallCapsWord id="8D26A057FFC1FFDD3CFC618AFE987542" baselines="525,525" box="[171,342,507,531]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="84" pageNumber="425">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFC1FFDD3D37618EFE3A7542" baselines="525" box="[352,500,511,530]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="84" pageNumber="425">DISTRIBUTION</smallCapsWord>
.
</emphasis>
This genus currently is known from only two species, both found only in the southern part of the Gaoligong Shan region of western
<collectingRegion id="49BBF869FFC1FFDD3FDD626DFB8A7563" box="[906,1092,539,563]" country="China" name="Yunnan" pageId="84" pageNumber="425">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFC1FFDD3818626AFB5C7563" box="[1103,1170,539,563]" name="China" pageId="84" pageNumber="425">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,300 @@
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<mods:title id="78FC786C141A2E2682D6245C9E853590">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="8CC70432538B7E8255D9C91EBF6434BD">Deuve, Thierry</mods:namePart>
<mods:affiliation id="4A7E361709C11BBD77C007E328E87E0C">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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</mods:role>
<mods:namePart id="F30DA4ED66BA795B086789B483011CF3">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="E4DDB657EF405E498ADE07C93DD9D485">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="17C7C1EEB1002720B0028476B8838DDC" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
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<mods:role id="FAFC5F1ACA919CA437B9A298F4819A42">
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<mods:namePart id="C5B1C3B82AB31F401D58448B27453F5A">Liang, Hongbin</mods:namePart>
<mods:affiliation id="93D30D5B547B91FCE6D48424E672C645">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:date id="2A50E990B24A11600D1F9DD51604E655">2016</mods:date>
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<treatment id="03D6879DFFCCFFD23C2C60AAFB4C7492" LSID="urn:lsid:plazi:treatment:03D6879DFFCCFFD23C2C60AAFB4C7492" httpUri="http://treatment.plazi.org/id/03D6879DFFCCFFD23C2C60AAFB4C7492" lastPageId="91" lastPageNumber="432" pageId="89" pageNumber="430">
<subSubSection id="C3656500FFCCFFD03C2C60AAFCD777A3" box="[123,793,219,243]" pageId="89" pageNumber="430" type="nomenclature">
<paragraph id="8BC0368BFFCCFFD03C2C60AAFCD777A3" blockId="89.[123,793,219,274]" box="[123,793,219,243]" pageId="89" pageNumber="430">
<heading id="D08881E7FFCCFFD03C2C60AAFCD777A3" box="[123,793,219,243]" fontSize="10" level="2" pageId="89" pageNumber="430" reason="4">
<emphasis id="B90BEA99FFCCFFD03C2C60AAFCD777A3" bold="true" box="[123,793,219,243]" pageId="89" pageNumber="430">
<taxonomicName id="4C7F4D08FFCCFFD03C2C60AAFE3677A3" ID-CoL="8MJTV" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,504,219,243]" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="430" phylum="Arthropoda" rank="species" species="gaoligong">
<emphasis id="B90BEA99FFCCFFD03C2C60AAFE6477A3" bold="true" box="[123,426,219,243]" italics="true" pageId="89" pageNumber="430">Trechepaphiama gaoligong</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFCCFFD03E9460AAFCD777A3" box="[707,793,219,243]" pageId="89" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFCCFFD03C2C608DFE4F7642" box="[123,385,252,274]" pageId="89" pageNumber="430" type="description">
<paragraph id="8BC0368BFFCCFFD03C2C608DFE4F7642" blockId="89.[123,793,219,274]" box="[123,385,252,274]" pageId="89" pageNumber="430">
(
<figureCitation id="13442A0EFFCCFFD03CD5608DFF117642" box="[130,223,252,274]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="89" pageNumber="430">Figs. 16h</figureCitation>
,
<figureCitation id="13442A0EFFCCFFD03CBE608DFECD7641" box="[233,259,252,273]" captionStart="FIGURE" captionStartId="90.[171,183,1578,1597]" captionTargetBox="[138,1176,216,1565]" captionTargetId="figure-11@90.[120,1176,216,1565]" captionTargetPageId="90" captionText="FIGURE 34. Trechepaphiama gaoligong sp. nov.; a. Dorsal habitus (CASENT1001926). b. Map of locality record (red circle) for T. gaoligong in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155348" httpUri="https://zenodo.org/record/13155348/files/figure.png" pageId="89" pageNumber="430">34</figureCitation>
,
<figureCitation id="13442A0EFFCCFFD03D59608DFEFC7641" box="[270,306,252,273]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="89" pageNumber="430">37a</figureCitation>
,
<figureCitation id="13442A0EFFCCFFD03D6A608DFEB57642" box="[317,379,252,274]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="89" pageNumber="430">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFCCFFD03CFC6158FF0C75D7" pageId="89" pageNumber="430" type="materials_examined">
<paragraph id="8BC0368BFFCCFFD03CFC6158FF7176B4" blockId="89.[123,1174,296,1725]" pageId="89" pageNumber="430">
<emphasis id="B90BEA99FFCCFFD03CFC6158FE96766F" bold="true" box="[171,344,297,321]" pageId="89" pageNumber="430">
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</emphasis>
.—
<materialsCitation id="3B173CD6FFCCFFD03D286159FF4A7693" collectingDate="1998-10-15" collectingDateMax="1998-10-18" collectingDateMin="1998-10-15" collectorName="C. E. Griswold &amp; C. Ferraris &amp; C. - L. Long" country="China" location="Nujiang Prefecture" municipality="Gaoligongshan Mountains" pageId="89" pageNumber="430" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFCCFFD03D286159FE2B7610" box="[383,485,296,320]" pageId="89" pageNumber="430" type="holotype">Holotype</typeStatus>
, a female, in IOZ, labeled: “CASENT 1001926”/ “
<collectingCountry id="F368761BFFCCFFD03FA96158FB997611" box="[1022,1111,297,321]" name="China" pageId="89" pageNumber="430">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFCCFFD038356158FEC47631" country="China" name="Yunnan" pageId="89" pageNumber="430">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFCCFFD03D426138FDFD7631" box="[277,563,329,353]" pageId="89" pageNumber="430">Gaoligongshan Mountains</collectingMunicipality>
,
<location id="8EA06050FFCCFFD03E696138FCC57631" LSID="urn:lsid:plazi:treatment:03D6879DFFCCFFD23C2C60AAFB4C7492:8EA06050FFCCFFD03E696138FCC57631" box="[574,779,329,353]" country="China" municipality="Gaoligongshan Mountains" name="Nujiang Prefecture" pageId="89" pageNumber="430" stateProvince="Yunnan">Nujiang Prefecture</location>
, 9 [actually 9.3] km
<locationDeviation id="9A722FA9FFCCFFD03FBD6138FB5C7631" box="[1002,1170,329,353]" location="Pianma" pageId="89" pageNumber="430">ESE of Pianma</locationDeviation>
, 25°59.6N/ 98°37.6E.”/ “2450 [actually 2460-2470] m,
<date id="FFC1104BFFCCFFD03E81611BFC6176D2" box="[726,943,362,386]" pageId="89" pageNumber="430" value="1998-10-15" valueMax="1998-10-18" valueMin="1998-10-15">
<collectingDate id="EF85E9A3FFCCFFD03E81611BFC6176D2" box="[726,943,362,386]" pageId="89" pageNumber="430" value="1998-10-15" valueMax="1998-10-18" valueMin="1998-10-15">15-18 October 1998</collectingDate>
</date>
, Stop #98-118D D.H.
<taxonomicName id="4C7F4D08FFCCFFD03C2C61FBFF3976F2" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,394,418]" pageId="89" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFCCFFD03D5E61FAFE6376F2" box="[265,429,394,419]" pageId="89" pageNumber="430">C.E. Griswold</collectorName>
,
<collectorName id="268A535DFFCCFFD03DE861FAFDF776F2" box="[447,569,394,419]" pageId="89" pageNumber="430">C. Ferraris</collectorName>
&amp;
<collectorName id="268A535DFFCCFFD03E3C61FAFD2776F3" box="[619,745,395,419]" pageId="89" pageNumber="430">C.-L. Long</collectorName>
collectors”/ “IMAGE” [green label]/ “
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<taxonomicName id="4C7F4D08FFCCFFD03D4A61DAFD5A7693" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[285,660,427,451]" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="430" phylum="Arthropoda" rank="species" species="gaoligong">
<emphasis id="B90BEA99FFCCFFD03D4A61DAFD8B7693" box="[285,581,427,451]" italics="true" pageId="89" pageNumber="430">Trechepaphiama gaoligong</emphasis>
Deuve
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&amp; Kavanaugh,
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designated 2016” [red label].
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.—
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,
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,
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,
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ESE of Pianma
</locationDeviation>
,
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/
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,
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.
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.—
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species epithet,
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<emphasis id="B90BEA99FFCCFFD03F5A625FFCB77516" box="[781,889,558,582]" italics="true" pageId="89" pageNumber="430">gaoligong</emphasis>
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, is a noun in apposition, derived from the name of the mountain range, the
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<collectingRegion id="49BBF869FFCCFFD03F5D623FFC8C7536" box="[778,834,590,614]" country="Myanmar" name="Shan" pageId="89" pageNumber="430">Shan</collectingRegion>
, in which the
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was collected
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.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFCCFFD03CFC62E1FD9274FD" pageId="89" pageNumber="430" type="diagnosis">
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<emphasis id="B90BEA99FFCCFFD03CFC62E1FEEE75F6" bold="true" box="[171,288,656,680]" pageId="89" pageNumber="430">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFFCCFFD03E1062E1FD6D75F8" box="[583,675,656,680]" captionStart="FIGURE" captionStartId="90.[171,183,1578,1597]" captionTargetBox="[138,1176,216,1565]" captionTargetId="figure-11@90.[120,1176,216,1565]" captionTargetPageId="90" captionText="FIGURE 34. Trechepaphiama gaoligong sp. nov.; a. Dorsal habitus (CASENT1001926). b. Map of locality record (red circle) for T. gaoligong in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155348" httpUri="https://zenodo.org/record/13155348/files/figure.png" pageId="89" pageNumber="430">Fig. 34a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL =
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), apterous; body color reddish brown, dorsum shiny; eyes reduced, their diameter about equal to length of tempora, the latter distinctly convex and glabrous; pronotum moderately transverse (ratio PW/PL = 1.26), basal angles subrectangular; elytra broad, markedly convex, slightly inflated, humeri broadly rounded, striae 1 to 5 deeply impressed and slightly punctate, more lateral striae more or less effaced, intervals 1 to 4 slightly convex, more lateral intervals flat; recurrent stria anteriorly nearly continuous with posterior end of stria 5; discal setiferous pores absent; preapical seta present, inserted on interval 3 next to stria 2.
</paragraph>
</subSubSection>
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<paragraph id="8BC0368BFFCCFFD03CFC63C7FC2F735F" blockId="89.[123,1174,296,1725]" pageId="89" pageNumber="430">
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.— Size large, BL
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. Body color reddish brown, appendages concolorous, except palpi yellowish brown, dorsum smooth, shiny, pronotum and elytra very faintly iridescent, head faintly alutaceous due to moderately impressed isodiametric microsculpture
</paragraph>
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Head. Moderate in size, slightly broad; eyes small, only slightly projected laterally, their diameter about equal to length of tempora, the latter short, convex and glabrous. Frons only slightly convex, with frontal furrows deeply impressed but less so posterior to hind margins of eyes. Two pairs of supraorbital setae present, the anterior pair inserted in distinct foveae. Clypeus with four setae. Labrum with six setae, anterior margin moderately concave. Right mandible (
<figureCitation id="13442A0EFFCCFFD03F9164EAFBED73E2" box="[966,1059,1178,1203]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="89" pageNumber="430">Fig. 16h</figureCitation>
) with premolar tooth fused with retinaculum, anterior tip of retinaculum enlarged to form a distinct tooth and displaced distally, posterior tip of retinaculum reduced, nearly flattened. Left mandible with only a relatively small, feebly trifid tooth.
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and submentum not fused.
<collectingRegion id="49BBF869FFCCFFD03F97648CFBD47244" box="[960,1050,1277,1300]" country="Myanmar" name="Chin" pageId="89" pageNumber="430">Mentum</collectingRegion>
with medial tooth broad, apically rounded, about one-half the length of the lateral lobes. Submentum with four setae anteriorly. Gula broad. Genae with a single ventral seta one each side. Antennae rather short, only about two antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 4 about equal in length, antennomere 3 slightly longer.
</paragraph>
<paragraph id="8BC0368BFFCCFFD23CFC65D1FCA57644" blockId="89.[123,1174,296,1725]" lastBlockId="91.[123,1174,219,962]" lastPageId="91" lastPageNumber="432" pageId="89" pageNumber="430">Pronotum. Slightly narrow (ratio PW/PL = 1.26), with greatest width anterior to middle, only slightly narrowed posteriorly; lateral margins rounded, straightened for a short distance posteriorly, then very slightly sinuate just anterior to basal angles, the latter blunt and subrectangular, very slightly and bluntly projected laterally; basal margin with median region slightly and convexly projected posteriorly. Disc distinctly convex, with median longitudinal impression very finely impressed, superficial, effaced in the median anterior area but extended to basal margin; basal foveae distinct but shallow; median basal area small, delimited laterally by a pair of oblique furrows, smooth except for several short longitudinal depressions; basal margin slightly bisinuate, slightly convex medially; lateral borders slender, gradually widened posteriorly, faintly reflexed dorsally, lateral grooves widened near base. Single midlateral setae on each side inserted anterior to middle; single basolateral seta on each side, inserted at basal angle.</paragraph>
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34.
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<emphasis id="B90BEA99FFCFFFD33D5A665BFE36716D" box="[269,504,1578,1597]" italics="true" pageId="90" pageNumber="431">Trechepaphiama gaoligong</emphasis>
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; a. Dorsal habitus (CASENT1001926). b. Map of locality record (red circle) for
<taxonomicName id="4C7F4D08FFCFFFD33C816635FE8C7106" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[214,322,1603,1622]" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="431" phylum="Arthropoda" rank="species" species="gaoligong">
<emphasis id="B90BEA99FFCFFFD33C816635FE8C7106" box="[214,322,1603,1622]" italics="true" pageId="90" pageNumber="431">T. gaoligong</emphasis>
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in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFCEFFD23CFC616DFDA67549" blockId="91.[123,1174,219,962]" pageId="91" pageNumber="432">Elytra. Broad, ovoid, slightly more narrowed posteriorly than anteriorly, markedly convex, slightly inflated; humeri broadly rounded; striae 1 to 5 deeply impressed and slightly punctate, more lateral striae more or less effaced, barely evident; intervals 1 to 4 slightly convex, more lateral intervals flat; basal margination terminated medially at origin of stria 5. Parascutellar striole present, short but distinct. Recurrent stria anteriorly nearly continuous with posterior end of stria 5. Discal setiferous pores absent. Preapical seta present, inserted on interval 3 next to stria 2. Umbilicate setal series with setae of humeral group equidistance from each other and setae of medi- an group inserted slightly posterior to middle.</paragraph>
<paragraph id="8BC0368BFFCEFFD23CFC6253FD67756A" blockId="91.[123,1174,219,962]" box="[171,681,546,570]" pageId="91" pageNumber="432">Legs. Short; protibiae with longitudinal furrow.</paragraph>
<paragraph id="8BC0368BFFCEFFD23CFC6233FC8C752A" blockId="91.[123,1174,219,962]" pageId="91" pageNumber="432">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of female apically with two pairs of paramedial setae.</paragraph>
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.— The unique
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female of this species was collected by sifting leaf litter taken on sandy substrate on a secondary floodplain covered by a broadleaf forest canopy at an elevation of
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(
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). Specimens of
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<emphasis id="B90BEA99FFCEFFD23EF862B4FC10758D" box="[687,990,709,733]" italics="true" pageId="91" pageNumber="432">Trechepaphiopsis uniporosa</emphasis>
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and
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<emphasis id="B90BEA99FFCEFFD2384262B7FB58758D" box="[1045,1174,710,733]" italics="true" pageId="91" pageNumber="432">T. unisetosa</emphasis>
</taxonomicName>
were collected in the same sifted litter samples at this site.
</paragraph>
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<materialsCitation id="3B173CD6FFCEFFD23F4A6377FD227430" county="western Lushui County" location="This" pageId="91" pageNumber="432" specimenCount="1" stateProvince="Shan">
<smallCapsWord id="8D26A057FFCEFFD23F4A6377FC99744D" baselines="792,792" box="[797,855,774,798]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="91" pageNumber="432">
<emphasis id="B90BEA99FFCEFFD23F4A6377FC99744D" bold="true" box="[797,855,774,798]" pageId="91" pageNumber="432">
<collectingRegion id="49BBF869FFCEFFD23F4A6377FC99744D" box="[797,855,774,798]" country="Myanmar" name="Shan" pageId="91" pageNumber="432">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFCEFFD23FD36377FC2A744E" box="[900,996,774,798]" captionStart="FIGURE" captionStartId="90.[171,183,1578,1597]" captionTargetBox="[138,1176,216,1565]" captionTargetId="figure-11@90.[120,1176,216,1565]" captionTargetPageId="90" captionText="FIGURE 34. Trechepaphiama gaoligong sp. nov.; a. Dorsal habitus (CASENT1001926). b. Map of locality record (red circle) for T. gaoligong in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155348" httpUri="https://zenodo.org/record/13155348/files/figure.png" pageId="91" pageNumber="432">Fig. 34b</figureCitation>
.
<location id="8EA06050FFCEFFD23FA56377FBEF744E" LSID="urn:lsid:plazi:treatment:03D6879DFFCCFFD23C2C60AAFB4C7492:8EA06050FFCEFFD23FA56377FBEF744E" box="[1010,1057,774,798]" county="western Lushui County" name="This" pageId="91" pageNumber="432" stateProvince="Shan">This</location>
species is known only from a single female specimen from the
<typeStatus id="54C48829FFCEFFD23EF96359FD127410" box="[686,732,808,832]" pageId="91" pageNumber="432">type</typeStatus>
locality in
<collectingCounty id="62A14E07FFCEFFD23F036359FB80746F" box="[852,1102,807,831]" pageId="91" pageNumber="432">western Lushui County</collectingCounty>
on the western slope of the southcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFCEFFD23EA06339FF0874D0" location="Core Area" pageId="91" pageNumber="432" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFCEFFD23EA06339FC4E7430" box="[759,896,840,864]" country="Myanmar" name="Shan" pageId="91" pageNumber="432">Shan region</collectingRegion>
. This locality is in
<location id="8EA06050FFCEFFD238366339FF6174D1" LSID="urn:lsid:plazi:treatment:03D6879DFFCCFFD23C2C60AAFB4C7492:8EA06050FFCEFFD238366339FF6174D1" name="Core Area" pageId="91" pageNumber="432" stateProvince="Shan">Core Area</location>
4
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFCEFFD23CFC63F8FB4C7492" blockId="91.[123,1174,219,962]" pageId="91" pageNumber="432">
<emphasis id="B90BEA99FFCEFFD23CFC63F8FD9774CF" bold="true" box="[171,601,905,929]" pageId="91" pageNumber="432">
<smallCapsWord id="8D26A057FFCEFFD23CFC63F8FEDC74F0" baselines="923,923" box="[171,274,905,929]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="91" pageNumber="432">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFCEFFD23D4E63FDFE0E74CF" baselines="922" box="[281,448,908,927]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="91" pageNumber="432">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFCEFFD23D9263FDFD9774CF" baselines="922" box="[453,601,908,927]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="91" pageNumber="432">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Lushui County in the southcentral part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFCEFFD23F2263DBFBFE7492" box="[885,1072,938,962]" country="China" name="Yunnan" pageId="91" pageNumber="432">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFCEFFD2386C63DBFBB07492" box="[1083,1150,938,962]" name="China" pageId="91" pageNumber="432">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,169 @@
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<mods:title id="9B86E746B0DF0A716A8D9044D27E4C40">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="31C9F5CE31EA08ECDD97EEC8C37D1339">Deuve, Thierry</mods:namePart>
<mods:affiliation id="5CBEFADA4AB625D79B3A2F1C05476B92">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="0425FC3F57E2E8738C1B9718C0551CC0">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="857E935F68A069D647D08CE9C0DF2D1A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="030462D22F8D17C0E2CCCEDF6D3BA27D" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:name id="22F0DFAE1E65E9D88FCDD61B2CC1F913" type="personal">
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<mods:namePart id="4BD3D36AA1C1B54506BCCAFACEDDEA3B">Liang, Hongbin</mods:namePart>
<mods:affiliation id="955260A270B1712758FB793809C973E3">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:date id="D6225FF5A32B4272C72725AFDE1745C1">2016</mods:date>
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<treatment id="03D6879DFFCDFFD13D1362B4FEE171CD" LSID="urn:lsid:plazi:treatment:03D6879DFFCDFFD13D1362B4FEE171CD" httpUri="http://treatment.plazi.org/id/03D6879DFFCDFFD13D1362B4FEE171CD" lastPageNumber="429" pageId="88" pageNumber="429">
<subSubSection id="C3656500FFCDFFD13D1362B4FC02758D" box="[324,972,709,733]" pageId="88" pageNumber="429" type="nomenclature">
<paragraph id="8BC0368BFFCDFFD13D1362B4FC02758D" blockId="88.[324,972,709,733]" box="[324,972,709,733]" pageId="88" pageNumber="429">
<heading id="D08881E7FFCDFFD13D1362B4FC02758D" box="[324,972,709,733]" centered="true" fontSize="11" level="2" pageId="88" pageNumber="429" reason="2">
<emphasis id="B90BEA99FFCDFFD13D1362B4FC02758D" bold="true" box="[324,972,709,733]" pageId="88" pageNumber="429">
Genus
<taxonomicName id="4C7F4D08FFCDFFD13DC362B4FD52758D" ID-CoL="8MDGC" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[404,668,709,733]" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13DC362B4FD80758D" bold="true" box="[404,590,709,733]" italics="true" pageId="88" pageNumber="429">Trechepaphiama</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFCDFFD13F3062B4FC02758D" box="[871,972,709,733]" pageId="88" pageNumber="429" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFCDFFD13CFC6280FC85741A" pageId="88" pageNumber="429" type="type_taxon">
<paragraph id="8BC0368BFFCDFFD13CFC6280FD267459" blockId="88.[123,1174,753,1693]" box="[171,744,753,777]" pageId="88" pageNumber="429">
<emphasis id="B90BEA99FFCDFFD13CFC6280FEF17458" bold="true" box="[171,319,753,777]" pageId="88" pageNumber="429">
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<typeStatus id="54C48829FFCDFFD13CFC6280FF2A7458" box="[171,228,753,777]" pageId="88" pageNumber="429">TYPE</typeStatus>
</smallCapsWord>
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.—
<taxonomicName id="4C7F4D08FFCDFFD13D3F6280FD437459" ID-CoL="8MJTV" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[360,653,753,777]" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="species" species="gaoligong" status="sp. nov.">
<emphasis id="B90BEA99FFCDFFD13D3F6280FD437459" box="[360,653,753,777]" italics="true" pageId="88" pageNumber="429">Trechepaphiama gaoligong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFCDFFD13EC26283FD267459" box="[661,744,754,777]" pageId="88" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFFCDFFD13CFC6363FC85741A" blockId="88.[123,1174,753,1693]" pageId="88" pageNumber="429">
<emphasis id="B90BEA99FFCDFFD13CFC6363FDFC7478" bold="true" box="[171,562,786,810]" pageId="88" pageNumber="429">
<smallCapsWord id="8D26A057FFCDFFD13CFC6363FEE07478" baselines="804,803" box="[171,302,786,810]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="88" pageNumber="429">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFCDFFD13D616364FE9C7478" baselines="803" box="[310,338,789,808]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="88" pageNumber="429">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFCDFFD13D0E6364FE507478" baselines="803" box="[345,414,789,808]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="genus" pageId="88" pageNumber="429">GENUS</smallCapsWord>
<smallCapsWord id="8D26A057FFCDFFD13DF16364FE207478" baselines="803" box="[422,494,789,808]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="group" pageId="88" pageNumber="429">GROUP</smallCapsWord>
<smallCapsWord id="8D26A057FFCDFFD13DA26364FDFC7478" baselines="803" box="[501,562,789,808]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="88" pageNumber="429">NAME</smallCapsWord>
</emphasis>
.— The genus group name (masculine) is a combination of two other trechine genus group names,
<taxonomicName id="4C7F4D08FFCDFFD13E176342FD5D741B" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[576,659,819,843]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13E176342FD5D741B" box="[576,659,819,843]" italics="true" pageId="88" pageNumber="429">Trechus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFCDFFD13E9E6342FC8A741B" authorityName="Jeannel" authorityYear="1962" box="[713,836,819,843]" class="Insecta" family="Carabidae" genus="Epaphiama" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13E9E6342FC8A741B" box="[713,836,819,843]" italics="true" pageId="88" pageNumber="429">Epaphiama</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFCDFFD13CFC6325FDA87245" pageId="88" pageNumber="429" type="diagnosis">
<paragraph id="8BC0368BFFCDFFD13CFC6325FDA87245" blockId="88.[123,1174,753,1693]" pageId="88" pageNumber="429">
<smallCapsWord id="8D26A057FFCDFFD13CFC6325FEEE743A" baselines="870,869" box="[171,288,852,876]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="88" pageNumber="429">
<emphasis id="B90BEA99FFCDFFD13CFC6325FEEE743A" bold="true" box="[171,288,852,876]" pageId="88" pageNumber="429">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this genus (
<figureCitation id="13442A0EFFCDFFD13E7C6322FD4A743C" box="[555,644,851,876]" captionStart="FIGURE" captionStartId="90.[171,183,1578,1597]" captionTargetBox="[138,1176,216,1565]" captionTargetId="figure-11@90.[120,1176,216,1565]" captionTargetPageId="90" captionText="FIGURE 34. Trechepaphiama gaoligong sp. nov.; a. Dorsal habitus (CASENT1001926). b. Map of locality record (red circle) for T. gaoligong in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155348" httpUri="https://zenodo.org/record/13155348/files/figure.png" pageId="88" pageNumber="429">Fig. 34a</figureCitation>
) can be recognized by the following combination of character states: size large (BL =
<quantity id="4C879B6EFFCDFFD13E536305FD9774DC" box="[516,601,884,908]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="88" pageNumber="429" unit="mm" value="4.2">4.2 mm</quantity>
), apterous; body color reddish brown, dorsum shiny, slightly iridescent, dorsal surface glabrous except for isolated fixed setae typical for trechines, eyes also glabrous, small, only slightly convex; tempora glabrous; right mandible (
<figureCitation id="13442A0EFFCDFFD13F9063C4FBED749D" box="[967,1059,949,973]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="88" pageNumber="429">Fig. 16h</figureCitation>
) with premolar tooth fused with retinaculum, anterior tip of retinaculum enlarged to form a distinct tooth and displaced distally, posterior tip of retinaculum reduced, nearly flattened; mentum and submentum not fused; mentum with median tooth simple with apex rounded; submentum [probably] with six setae [only five setae visible in unique female
<typeStatus id="54C48829FFCDFFD13EDC6449FD267300" box="[651,744,1080,1104]" pageId="88" pageNumber="429" type="holotype">holotype</typeStatus>
of type species]; pronotum slightly convex, disc glabrous, basal foveae only slightly distinct, median basal area delimited laterally by oblique furrows, basal margin slightly curved medially, slightly sinuate and oblique laterally; elytra broadly rounded, more narrowed apically than basally, humeri broad, disc convex with striae finely impressed, slightly punctate, lateral striae attenuated, recurrent stria deeply impressed, its anterior end straight, almost continuous anteriorly with the apex of stria 5; no discal setae present, preapical seta present, inserted next to stria 2.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFCDFFD13CFC656CFEE171CD" pageId="88" pageNumber="429" type="discussion">
<paragraph id="8BC0368BFFCDFFD13CFC656CFDA9716B" blockId="88.[123,1174,753,1693]" pageId="88" pageNumber="429">
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<emphasis id="B90BEA99FFCDFFD13CFC656CFEE67264" bold="true" box="[171,296,1309,1333]" pageId="88" pageNumber="429">COMMENTS</emphasis>
</smallCapsWord>
.— This genus, known from only a single female specimen, belongs to the “
<taxonomicName id="4C7F4D08FFCDFFD13836656CFF1A7206" authorityName="Ueno" authorityYear="1953" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="455" phylum="Arthropoda" rank="genus">Epaphiopsis</taxonomicName>
Complex” of genera as evidenced by the dentition of the mandibles. Its members can be distinguished from those of other genera of this lineage by the absence of elytral discal setae, which is exceptional among trechines, and by their markedly convex, dorsally inflated elytra, imparting an appearance similar to that of
<taxonomicName id="4C7F4D08FFCDFFD13D8765D1FD3E72E8" authority="Jeannel (1962)" authorityName="Jeannel" authorityYear="1962" box="[464,752,1440,1464]" class="Insecta" family="Carabidae" genus="Epaphiama" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13D8765D1FD8572E8" box="[464,587,1440,1464]" italics="true" pageId="88" pageNumber="429">Epaphiama</emphasis>
<bibRefCitation id="EFEE4B7AFFCDFFD13E0565D1FD3E72E8" author="JEANNEL, R." box="[594,752,1440,1464]" pageId="88" pageNumber="429" pagination="171 - 207" refId="ref59689" refString="JEANNEL, R. 1962. Les Trechini de l'Extreme-Orient. Revue Francaise d'Entomologie 29: 171 - 207." type="journal article" year="1962">Jeannel (1962)</bibRefCitation>
</taxonomicName>
members — short and convex. In contrast,
<taxonomicName id="4C7F4D08FFCDFFD13CE165B0FEBF7289" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[182,369,1473,1497]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13CE165B0FEBF7289" box="[182,369,1473,1497]" italics="true" pageId="88" pageNumber="429">Trechepaphiopsis</emphasis>
</taxonomicName>
members are similar in habitus to small
<taxonomicName id="4C7F4D08FFCDFFD13F7065B0FC657289" authorityName="Ueno" authorityYear="1953" box="[807,939,1473,1497]" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13F7065B0FC657289" box="[807,939,1473,1497]" italics="true" pageId="88" pageNumber="429">Epaphiopsis</emphasis>
</taxonomicName>
members while those of
<taxonomicName id="4C7F4D08FFCDFFD13CC06590FEF172A9" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[151,319,1505,1529]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFCDFFD13CC06590FEF172A9" box="[151,319,1505,1529]" italics="true" pageId="88" pageNumber="429">Epaphiotrechus</emphasis>
</taxonomicName>
are more similar in habitus to large
<emphasis id="B90BEA99FFCDFFD13EE86590FCC672A9" box="[703,776,1505,1529]" italics="true" pageId="88" pageNumber="429">Trechu</emphasis>
s members. A better understanding of phylogenetic affinities of this new genus must await discovery of a male and study of its genital structure and/or comparative molecular study.
</paragraph>
<paragraph id="8BC0368BFFCDFFD13CFC6632FEE171CD" blockId="88.[123,1174,753,1693]" pageId="88" pageNumber="429">
<emphasis id="B90BEA99FFCDFFD13CFC6632FE3C710A" bold="true" box="[171,498,1603,1627]" pageId="88" pageNumber="429">
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<smallCapsWord id="8D26A057FFCDFFD13D096636FE3C710A" baselines="1621" box="[350,498,1607,1626]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="88" pageNumber="429">DISTRIBUTION</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFCDFFD13E4C6632FEE471CD" country="China" location="Gaoligong Shan region" pageId="88" pageNumber="429" specimenCount="1" stateProvince="Yunnan">
This genus currently is known only from the
<typeStatus id="54C48829FFCDFFD1385C6635FBF7710C" box="[1035,1081,1604,1628]" pageId="88" pageNumber="429">type</typeStatus>
species, which is known only from the southcentral part of the
<location id="8EA06050FFCDFFD13E9D6615FC0D712C" LSID="urn:lsid:plazi:treatment:03D6879DFFCDFFD13D1362B4FEE171CD:8EA06050FFCDFFD13E9D6615FC0D712C" box="[714,963,1636,1660]" country="China" name="Gaoligong Shan region" pageId="88" pageNumber="429" stateProvince="Yunnan">Gaoligong Shan region</location>
of western
<collectingRegion id="49BBF869FFCDFFD138136614FF1371CD" country="China" name="Yunnan" pageId="88" pageNumber="429">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFCDFFD13CB066F4FEE471CD" box="[231,298,1669,1693]" name="China" pageId="88" pageNumber="429">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,361 @@
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<mods:title id="2DB3C6F0F99AD4B998094FCF9A123A85">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:affiliation id="B2D3D847EF7AE2200A33267954C9AA7B">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FFD3FFCF3C2C60AAFE6577A3" bold="true" box="[123,427,219,243]" italics="true" pageId="70" pageNumber="411">Trechepaphiopsis unisetosa</emphasis>
(Deuve), 2004
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(sp. 2)
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</paragraph>
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<subSubSection id="C3656500FFD3FFCF3C2C608AFE4F7641" box="[123,385,251,273]" pageId="70" pageNumber="411" type="description">
<paragraph id="8BC0368BFFD3FFCF3C2C608AFE4F7641" blockId="70.[123,668,219,273]" box="[123,385,251,273]" pageId="70" pageNumber="411">
(
<figureCitation id="13442A0EFFD3FFCF3CD5608AFF1D7641" box="[130,211,251,273]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Figs. 27</figureCitation>
,
<figureCitation id="13442A0EFFD3FFCF3C8A608DFECC7641" box="[221,258,252,273]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="70" pageNumber="411">37a</figureCitation>
,
<figureCitation id="13442A0EFFD3FFCF3D5B608AFEFD7641" box="[268,307,251,273]" captionStart="FIGURE" captionStartId="105.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1606]" captionTargetId="figure-15@105.[120,1176,216,1607]" captionTargetPageId="105" captionText="FIGURE 43. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Luzhang Township, Piana Road 0.1 km E of Fengxue Yakou, elevation 3150 m; habitat in which specimens of Trechus luzhangensis sp. nov. were collected. b. Lushui County, Luzhang Township, 100 m S of Fengxue Yakou on east side of pass, elevation 3150 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve) were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155372" httpUri="https://zenodo.org/record/13155372/files/figure.png" pageId="70" pageNumber="411">43b</figureCitation>
,
<figureCitation id="13442A0EFFD3FFCF3D6A608AFEB57641" box="[317,379,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="70" pageNumber="411">4648</figureCitation>
)
</paragraph>
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<emphasis id="B90BEA99FFD3FFCF3C2C6156FEE1766D" box="[123,303,295,317]" italics="true" pageId="70" pageNumber="411">Trechus unisetosus</emphasis>
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.
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, a female, in SCAU. Type locality:
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,
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,
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,
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,
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[Pass],
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<taxonomicName id="4C7F4D08FFD3FFCF3C2C6114FE12762A" ID-CoL="8MH87" authority="(Deuve)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2004" box="[123,476,357,379]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa" status="comb. nov.">
<emphasis id="B90BEA99FFD3FFCF3C2C6114FE4B762B" box="[123,389,357,379]" italics="true" pageId="70" pageNumber="411">Trechepaphiopsis unisetosa</emphasis>
(Deuve)
</taxonomicName>
,
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<emphasis id="B90BEA99FFD3FFCF3E4B6114FD68762A" bold="true" box="[540,678,357,379]" pageId="70" pageNumber="411">COMBINATION</emphasis>
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</subSubSection>
<subSubSection id="C3656500FFD3FFCF3CFC61FEFE9875BD" pageId="70" pageNumber="411" type="diagnosis">
<paragraph id="8BC0368BFFD3FFCF3CFC61FEFE9875BD" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
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<emphasis id="B90BEA99FFD3FFCF3CFC61FEFEEE76F5" bold="true" box="[171,288,399,423]" pageId="70" pageNumber="411">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFFD3FFCF3E1061FEFD6D76F7" box="[583,675,399,423]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Fig. 27a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
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), apterous; body color light brown, dorsum slightly iridescent; eyes small but convex; tempora very convex, sparsely pubescent; pronotum transverse (ratio PW/PL = 1.35), with basal angles obtuse and rounded; elytra convex, with median 2 or 3 striae deeply impressed, lateral striae faintly impressed to effaced; recurrent stria terminated anteriorly in presumed location of interval 6; only one discal setiferous pore present, inserted at anterior one-third next to stria 3; preapical seta present, inserted next to stria 2; median lobe of male aedeagus (
<figureCitation id="13442A0EFFD3FFCF3FDB6202FC2775DB" box="[908,1001,627,651]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Fig. 27b</figureCitation>
) highly distinctive, with ventral margin of shaft sinuous and apex short and thin in lateral view, with broad apical projects in ventral view; endophallus without sclerites, but with a long tubular sleeve covered with long, slender spines.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD3FFCF3CFC6284FB4C710C" pageId="70" pageNumber="411" type="discussion">
<paragraph id="8BC0368BFFD3FFCF3CFC6284FF2B74C0" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
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<emphasis id="B90BEA99FFD3FFCF3CFC6284FEE6745C" bold="true" box="[171,296,757,781]" pageId="70" pageNumber="411">COMMENTS</emphasis>
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.— Among males of all the species of
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<emphasis id="B90BEA99FFD3FFCF3EEB6284FCB9745D" box="[700,887,757,781]" italics="true" pageId="70" pageNumber="411">Trechepaphiopsis</emphasis>
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, those of
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<emphasis id="B90BEA99FFD3FFCF3F896287FB93745D" box="[990,1117,758,781]" italics="true" pageId="70" pageNumber="411">T. unisetosa</emphasis>
</taxonomicName>
have an endophlallus most like that typical for members of the
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<emphasis id="B90BEA99FFD3FFCF3EA06367FCB5747E" box="[759,891,790,814]" italics="true" pageId="70" pageNumber="411">Epaphiopsis</emphasis>
</taxonomicName>
complex of genera, especially of certain species of
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<emphasis id="B90BEA99FFD3FFCF3DF36347FDC7741E" box="[420,521,822,846]" italics="true" pageId="70" pageNumber="411">Epaphius</emphasis>
</taxonomicName>
described from
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(
<bibRefCitation id="EFEE4B7AFFD3FFCF3F5A6347FC6E741E" author="JEANNEL, R." box="[781,928,822,846]" pageId="70" pageNumber="411" pagination="171 - 207" refId="ref59689" refString="JEANNEL, R. 1962. Les Trechini de l'Extreme-Orient. Revue Francaise d'Entomologie 29: 171 - 207." type="journal article" year="1962">Jeannel 1962</bibRefCitation>
,
<bibRefCitation id="EFEE4B7AFFD3FFCF3FFA6346FB8C741E" author="DEUVE, T." box="[941,1090,822,846]" pageId="70" pageNumber="411" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve 1992b</bibRefCitation>
). These also have a long tubular sleeve covered with long spines, symmetrically arranged in ventral view (
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)
</paragraph>
<paragraph id="8BC0368BFFD3FFCF3CFC63E9FE6473C4" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
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</emphasis>
.— Members of this species have been found in mixed broadleaf evergreen/ deciduous forest at elevations ranging from
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and collected using both pitfall traps and sifting of leaf litter from the forest floor (
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). Specimens of
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<emphasis id="B90BEA99FFD3FFCF3FE363A8FF0A7341" italics="true" pageId="70" pageNumber="411">Trechepaphiosis uniporosa</emphasis>
</taxonomicName>
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and
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<emphasis id="B90BEA99FFD3FFCF3D1A638BFDBC7342" box="[333,626,1018,1042]" italics="true" pageId="70" pageNumber="411">Trechepaphiama goaligong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD3FFCF3E2D638BFD097341" box="[634,711,1018,1041]" pageId="70" pageNumber="411" rank="species">sp. nov</taxonomicNameLabel>
were collected in the same sifted leaf litter samples and are therefore syntopic with
<taxonomicName id="4C7F4D08FFD3FFCF3E7A646AFD627362" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2004" box="[557,684,1051,1074]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFD3FFCF3E7A646AFD627362" box="[557,684,1051,1074]" italics="true" pageId="70" pageNumber="411">T. unisetosa</emphasis>
</taxonomicName>
at this site. Additional specimens of
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<emphasis id="B90BEA99FFD3FFCF386E646AFF697303" italics="true" pageId="70" pageNumber="411">T. unisetosa</emphasis>
</taxonomicName>
were collected in pitfall traps set in
<taxonomicName id="4C7F4D08FFD3FFCF3E61644AFD1A7303" box="[566,724,1083,1107]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="70" pageNumber="411" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFD3FFCF3E61644AFD1A7303" box="[566,724,1083,1107]" italics="true" pageId="70" pageNumber="411">Rhododendron</emphasis>
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and bamboo thickets on the east flank of the summit ridge just south of the Fengxue Yakou at
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(
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), a site slightly above the
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locality for the species.
</paragraph>
<paragraph id="8BC0368BFFD3FFCF3CFC64EDFF0E72E9" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
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</emphasis>
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<emphasis id="B90BEA99FFD3FFCF3EDD64EDFCC073E3" bold="true" box="[650,782,1180,1204]" pageId="70" pageNumber="411">GAOLIGONG</emphasis>
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.—
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.
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examined a total of
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(
<specimenCount id="9D79FD02FFD3FFCF3D2064CCFE147385" box="[375,474,1213,1237]" count="10" pageId="70" pageNumber="411" type="male">10 males</specimenCount>
and
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)
</materialsCitation>
<materialsCitation id="3B173CD6FFD3FFCF3ECB64CCFF7272E9" collectingDate="1998-10-15" collectingDateMax="2005-05-21" collectingDateMin="1998-10-15" collectionCode="CAS, IOZ, MNHN" collectorName="J.P. DUMBACHER &amp; C. E. Griswold &amp; K. J. Guo &amp; Lushui &amp; Changyan He &amp; C. Ferraris &amp; C. L. Long" county="Lushui County" elevation="100" latitude="25.99363" location="Fengxue Yakou" longLatPrecision="1" longitude="97.66651" municipality="Luzhang Township" pageId="70" pageNumber="411" specimenCount="25" specimenCount-female="15" specimenCount-male="10" stateProvince="SHAN">
from the following localities:
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<collectingCounty id="62A14E07FFD3FFCF3FB264CCFB407385" box="[997,1166,1213,1237]" pageId="70" pageNumber="411">Lushui County</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BA4ACF1FFD3FFCF3C2C64AFFE9373A6" box="[123,349,1246,1270]" pageId="70" pageNumber="411">Luzhang Township</collectingMunicipality>
(
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S of
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on east side of pass,
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</quantity>
,
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<paragraph id="8BC0368BFFD3FFCF3CFC65B0FF73714B" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">Members of this species have been collected only in the southcentral part of the study area, on both western and eastern slopes of the Gaoligongshan, which are in Core Areas 4 and 5, respectively.</paragraph>
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.— This species currently is known only from Lushui County in the southcentral part of the Gaoligong Shan region, western
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,
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.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

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<mods:title id="D08FCEB0507387C896EBC48753BB9FDD">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:affiliation id="D84E60AE772E2BEA8FCC54898600D1D8">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<emphasis id="B90BEA99FFD6FFCA3C2C653EFE5D7237" bold="true" box="[123,403,1359,1383]" italics="true" pageId="67" pageNumber="408">Trechepaphiopsis asetosa</emphasis>
(Uéno), 1997
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<paragraph id="8BC0368BFFD6FFCA3C2C6501FEEC72D6" blockId="67.[123,554,1359,1414]" box="[123,290,1392,1414]" pageId="67" pageNumber="408">(Figs. 25, 4648)</paragraph>
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<emphasis id="B90BEA99FFD6FFCA3C2C65EAFEBD72E1" box="[123,371,1435,1457]" italics="true" pageId="67" pageNumber="408">Trechus (s. str.) asetosus</emphasis>
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, a male, in NMST. Type locality:
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,
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,
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,
<collectingCounty id="62A14E07FFD6FFCA3D0465CAFDC27280" box="[339,524,1467,1489]" pageId="67" pageNumber="408">Tengchong County</collectingCounty>
,
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,
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.
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</paragraph>
<paragraph id="8BC0368BFFD6FFCA3C2C65AAFD4772A0" blockId="67.[123,1174,1435,1521]" box="[123,649,1499,1521]" pageId="67" pageNumber="408">
<taxonomicName id="4C7F4D08FFD6FFCA3C2C65AAFE7172A0" ID-CoL="8MKZ6" authority="(Ueno)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="1997" box="[123,447,1499,1521]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="408" phylum="Arthropoda" rank="species" species="asetosa" status="comb. nov.">
<emphasis id="B90BEA99FFD6FFCA3C2C65AAFEBC72A1" box="[123,370,1499,1521]" italics="true" pageId="67" pageNumber="408">Trechepaphiopsis asetosa</emphasis>
(Uéno)
</taxonomicName>
,
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<emphasis id="B90BEA99FFD6FFCA3D9E65AAFE3672BF" bold="true" box="[457,504,1499,1520]" pageId="67" pageNumber="408">NEW</emphasis>
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<emphasis id="B90BEA99FFD6FFCA3DA965AAFD4772A0" bold="true" box="[510,649,1499,1521]" pageId="67" pageNumber="408">COMBINATION</emphasis>
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</subSubSection>
<subSubSection id="C3656500FFD6FFCA3CFC6674FE91710D" pageId="67" pageNumber="408" type="description">
<paragraph id="8BC0368BFFD6FFCA3CFC6674FE91710D" blockId="67.[123,1174,1540,1726]" pageId="67" pageNumber="408">
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<smallCapsWord id="8D26A057FFD6FFCA3D366679FE00714B" baselines="1558" box="[353,462,1544,1563]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="67" pageNumber="408">MATERIAL</smallCapsWord>
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.— We have not had an opportunity to study the
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or any
<typeStatus id="54C48829FFD6FFCA3C2C6654FEE9716D" box="[123,295,1573,1597]" isOtherTypeStatus="true" pageId="67" pageNumber="408">other specimens</typeStatus>
of this species. Features noted below are based on
<bibRefCitation id="EFEE4B7AFFD6FFCA3F106654FC2E716D" author="UENO, S. I." box="[839,992,1573,1597]" pageId="67" pageNumber="408" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uénos (1997)</bibRefCitation>
original description and illustrations.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD6FFCD3CFC6614FD8F7283" lastPageId="68" lastPageNumber="409" pageId="67" pageNumber="408" type="diagnosis">
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<emphasis id="B90BEA99FFD6FFCA3CFC6614FEEE712B" bold="true" box="[171,288,1637,1661]" pageId="67" pageNumber="408">DIAGNOSIS</emphasis>
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.— Adults of this species (see
<bibRefCitation id="EFEE4B7AFFD6FFCA3E3E6614FD2D712D" author="UENO, S. I." box="[617,739,1637,1661]" pageId="67" pageNumber="408" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno 1997</bibRefCitation>
,
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) can be distinguished from those of all other species in the region by the following combination of character states: size small to medium (BL= 3.0 to
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), apterous; body color dark brown to piceous, dorsum slightly iridescent; eyes very small, flattened, not projected laterally beyond tempora; tempora convex, slightly inflated; pronotum large and transverse (ratio PW/PL = 1.32 to 1.49), widest near anterior two-fifths, lateral margins slightly more curved in apical half than in basal half, basal angles obtuse and rounded; elytra ovoid, relative long and narrow, convex, striae 1 and 2 distinct, nearly complete and finely punctate, striae 3 and 4 faintly evident, stria 5 to 7 effaced, striae 8 evident but interrupted in apical half; recurrent stria terminated anteriorly in presumed location of interval 6; discal setae absent; preapical seta present, inserted next stria 2, inserted closer to sutural than to apical margin; median lobe of male aedeagus (see
<bibRefCitation id="EFEE4B7AFFD1FFCD3C2C62AAFF3A75A3" author="UENO, S. I." box="[123,244,731,755]" pageId="68" pageNumber="409" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno 1997</bibRefCitation>
,
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) with apex short and narrow, faintly recurved dorsally, endophallus without sclerites, but covered with scales.
</paragraph>
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</emphasis>
.— According to
<bibRefCitation id="EFEE4B7AFFD1FFCD3C2C632AFEC47423" author="UENO, S. I." box="[123,266,859,883]" pageId="68" pageNumber="409" pagination="181 - 192" refId="ref61319" refString="UENO, S. I. 1997. New trechine beetles (Coleoptera, Trechinae) from the Gaoligong Shan Mountains in western Yunnan. Elytra 25: 181 - 192." type="journal article" year="1997">Uéno (1997)</bibRefCitation>
, members of this species were collected by sifting moist leaf litter accumulations on the ground in the “
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<emphasis id="B90BEA99FFD1FFCD3D8763EAFDA074E3" box="[464,622,923,947]" italics="true" pageId="68" pageNumber="409">Rhododendron</emphasis>
</taxonomicName>
zone” at elevations ranging from
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. At one of three sites where specimens of
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<emphasis id="B90BEA99FFD1FFCD3C2C638DFF287343" box="[123,230,1020,1043]" italics="true" pageId="68" pageNumber="409">T. asetosa</emphasis>
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were found (at
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), specimens of
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<emphasis id="B90BEA99FFD1FFCD3CCD646AFE5C7363" box="[154,402,1051,1075]" italics="true" pageId="68" pageNumber="409">Minutotrechus minutus</emphasis>
</taxonomicName>
were also collected in the same litter samples.
</paragraph>
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<emphasis id="B90BEA99FFD1FFCD3CFC642BFF6B73C2" bold="true" pageId="68" pageNumber="409">
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<smallCapsWord id="8D26A057FFD1FFCD3D38642FFDC77321" baselines="1132" box="[367,521,1118,1137]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="68" pageNumber="409">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFD1FFCD3E4B642FFDA07321" baselines="1132" box="[540,622,1118,1137]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="68" pageNumber="409">WITHIN</smallCapsWord>
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</emphasis>
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<emphasis id="B90BEA99FFD1FFCD3CF9640AFEFD73C1" bold="true" box="[174,307,1147,1171]" pageId="68" pageNumber="409">GAOLIGONG</emphasis>
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<smallCapsWord id="8D26A057FFD1FFCD3D6B640AFEB973C1" baselines="1165,1164" box="[316,375,1147,1171]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="68" pageNumber="409">
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<collectingRegion id="49BBF869FFD1FFCD3D6B640AFEB973C1" box="[316,375,1147,1171]" country="Myanmar" name="Shan" pageId="68" pageNumber="409">SHAN</collectingRegion>
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.— Fig. 25.
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examined a single
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specimen from the type locality, the only site from which the species has been recorded, in
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in the southern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFD1FFCD3C2C656AFDE87263" box="[123,550,1307,1331]" location="Core Area" pageId="68" pageNumber="409" specimenCount="1" stateProvince="Shan" typeStatus="paratype">
<collectingRegion id="49BBF869FFD1FFCD3C2C656AFF317263" box="[123,255,1307,1331]" country="Myanmar" name="Shan" pageId="68" pageNumber="409">Shan region</collectingRegion>
. This site is in
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6
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.
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<emphasis id="B90BEA99FFD1FFCD3CFC654BFF607221" bold="true" pageId="68" pageNumber="409">
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</emphasis>
.—
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This species currently is known only from the
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locality, in
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in the southern part of the
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, in western
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,
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</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,220 @@
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<mods:title id="C07258FC50C92E8C9CF65F82EE2E2E18">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="B8B214A118F0F979E9D8EC70F09DD583">Deuve, Thierry</mods:namePart>
<mods:affiliation id="0693AC081DF4DADA026F731B67A73306">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
<mods:name id="36F425AA6E3EB04568ABE0C684C5E040" type="personal">
<mods:role id="D414F1657E83530E5C4C314ED0174FFD">
<mods:roleTerm id="BF16828A988F696E429BB05D8B62318E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0FDE787494A2FFD50D9D68B61FEADBC">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="303E24D105D5D67A3E018BFDCAED7D91">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="CB856D38E783CF1AA3A3232CD92DB3AC" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="2A7EAD10B8EF5CD139BC2DF0F4829251" type="personal">
<mods:role id="4DAC60D2F081025EE5C3C61F3B39318D">
<mods:roleTerm id="A6A85B2D9F003105A95B02341BAF09F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A56F1A4CB20BB3CD96215DD0A8BB1F9">Liang, Hongbin</mods:namePart>
<mods:affiliation id="5EE002272D291CB9AA450D6E18FE3CC9">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:typeOfResource id="6F966757C00F9EAB0ACE2A8A948BB338">text</mods:typeOfResource>
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<mods:title id="635D6507B3813FA85CFB5EFE52547F2C">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="0F05E7B550DB7A74DFCFF8FE606BBD18">
<mods:date id="365A1CC8131800592B9999F9F79EFA33">2016</mods:date>
<mods:detail id="301115378C8075C0D155CBA683D4A0DC" type="pubDate">
<mods:number id="E9525CD7AFCF71432E96C9F70504FA45">2016-10-14</mods:number>
</mods:detail>
<mods:detail id="00EF05B3ECDF7922553F41B80F5AC9B8" type="volume">
<mods:number id="3E5D5F40508C3D9CF3B64746EBA8653D">63</mods:number>
</mods:detail>
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<mods:number id="4468E0599EDC9F0F36EF2927A80B9FD1">12</mods:number>
</mods:detail>
<mods:extent id="3F42F31F4CC535F447AB4B8545A1F377" unit="page">
<mods:start id="B1207EB75B65399D22B0AFA5EB076A6B">341</mods:start>
<mods:end id="FDDB76FD634FB3D7F23D1F257938B7E3">455</mods:end>
</mods:extent>
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<mods:classification id="D3A83E91E2C90A2871311420B56CE5D4">journal article</mods:classification>
<mods:identifier id="A23683C0CE91D2F73C2B3C519A167915" type="DOI">10.5281/zenodo.13155283</mods:identifier>
<mods:identifier id="D911F5A1E3E2347BB7BED59357425CB8" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="03E99993B1908A8D3960C48B3460AB69" type="Zenodo-Dep">13155283</mods:identifier>
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<heading id="D08881E7FFD7FFCB3D1761EBFC1E76E3" box="[320,976,410,435]" centered="true" fontSize="11" level="2" pageId="66" pageNumber="407" reason="2">
<emphasis id="B90BEA99FFD7FFCB3D1761EBFC1E76E3" bold="true" box="[320,976,410,435]" pageId="66" pageNumber="407">
Genus
<taxonomicName id="4C7F4D08FFD7FFCB3DC761EBFD6E76E3" ID-CoL="8M67V" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[400,672,410,435]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3DC761EBFD9F76E2" bold="true" box="[400,593,410,434]" italics="true" pageId="66" pageNumber="407">Trechepaphiopsis</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFD7FFCB3F3C61EAFC1E76E3" box="[875,976,411,435]" pageId="66" pageNumber="407" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD7FFCB3CFC61B4FF2A750D" pageId="66" pageNumber="407" type="type_taxon">
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<typeStatus id="54C48829FFD7FFCB3CFC61B4FF2A768C" box="[171,228,453,477]" pageId="66" pageNumber="407">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFD7FFCB3CBB61B8FEF1768C" baselines="471" box="[236,319,457,476]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="66" pageNumber="407">SPECIES</smallCapsWord>
</emphasis>
.—
<taxonomicName id="4C7F4D08FFD7FFCB3D3F61B4FD5B768D" ID-CoL="8MLL2" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[360,661,453,477]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFD7FFCB3D3F61B4FD5B768D" box="[360,661,453,477]" italics="true" pageId="66" pageNumber="407">Trechepaphiopsis uniporosa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFD7FFCB3EF361B7FD39768D" box="[676,759,454,477]" pageId="66" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFFD7FFCB3CFC6194FF2A750D" blockId="66.[123,1174,453,1725]" pageId="66" pageNumber="407">
<emphasis id="B90BEA99FFD7FFCB3CFC6194FDE376AC" bold="true" box="[171,557,485,509]" pageId="66" pageNumber="407">
<smallCapsWord id="8D26A057FFD7FFCB3CFC6194FEE076AC" baselines="503,503" box="[171,302,485,509]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="66" pageNumber="407">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFD7FFCB3D636198FE9E76AC" baselines="503" box="[308,336,489,508]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="66" pageNumber="407">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFD7FFCB3D016198FE5576AC" baselines="503" box="[342,411,489,508]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="genus" pageId="66" pageNumber="407">GENUS</smallCapsWord>
<smallCapsWord id="8D26A057FFD7FFCB3DF56198FE2476AC" baselines="503" box="[418,490,489,508]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="group" pageId="66" pageNumber="407">GROUP</smallCapsWord>
<smallCapsWord id="8D26A057FFD7FFCB3DA76198FDE376AC" baselines="503" box="[496,557,489,508]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="66" pageNumber="407">NAME</smallCapsWord>
</emphasis>
.— The genus group name (feminine) is a combination of two other trechine generic names,
<taxonomicName id="4C7F4D08FFD7FFCB3DA26274FD86754D" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[501,584,517,541]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3DA26274FD86754D" box="[501,584,517,541]" italics="true" pageId="66" pageNumber="407">Trechus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFD7FFCB3ED66274FD28754D" ID-CoL="8KWTX" authorityName="Samouelle" authorityYear="1819" box="[641,742,517,541]" class="Insecta" family="Carabidae" genus="Epaphius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3ED66274FD28754D" box="[641,742,517,541]" italics="true" pageId="66" pageNumber="407">Epaphius</emphasis>
</taxonomicName>
, plus the Greek suffix, οψις (translated into Latin as -
<emphasis id="B90BEA99FFD7FFCB3D4F6257FE81756D" box="[280,335,550,573]" italics="true" pageId="66" pageNumber="407">opsis</emphasis>
) meaning having the aspect of, here referring to a similarity with
<taxonomicName id="4C7F4D08FFD7FFCB38666254FB58756D" ID-CoL="8KWTX" authorityName="Samouelle" authorityYear="1819" box="[1073,1174,549,573]" class="Insecta" family="Carabidae" genus="Epaphius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB38666254FB58756D" box="[1073,1174,549,573]" italics="true" pageId="66" pageNumber="407">Epaphius</emphasis>
</taxonomicName>
members.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD7FFCB3CFC6214FCA9730D" pageId="66" pageNumber="407" type="diagnosis">
<paragraph id="8BC0368BFFD7FFCB3CFC6214FCA9730D" blockId="66.[123,1174,453,1725]" pageId="66" pageNumber="407">
<smallCapsWord id="8D26A057FFD7FFCB3CFC6214FEEE752C" baselines="631,631" box="[171,288,613,637]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="66" pageNumber="407">
<emphasis id="B90BEA99FFD7FFCB3CFC6214FEEE752C" bold="true" box="[171,288,613,637]" pageId="66" pageNumber="407">DIAGNOSIS</emphasis>
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.— Adults of this genus (Figs. 2531) can be recognized by the following combination of character states: size small to moderate (BL =
<quantity id="4C879B6EFFD7FFCB3E8362F4FCA075CD" box="[724,878,645,669]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.7" metricValueMin="2.8" pageId="66" pageNumber="407" unit="mm" value="3.25" valueMax="3.7" valueMin="2.8">2.8 to 3.7 mm</quantity>
), apterous; body color light to dark brown, reddish-brown, or piceous-brown, most members with dorsum slightly iridescent, dorsal surface glabrous except for isolated fixed setae typical for trechines, eyes also glabrous; head short, with eyes small; tempora convex, swollen in some members, sparsely pubescent; right mandible (
<figureCitation id="13442A0EFFD7FFCB3CBA6374FE89744D" box="[237,327,773,797]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="66" pageNumber="407">Fig. 16e</figureCitation>
) with premolar tooth not fused with retinaculum and anterior point of the retinaculum free and displaced distally to form a separate tooth; mentum and submentum not fused; mentum with median tooth apical truncate or bifid; submentum with six setae; pronotum transverse, disc glabrous, basal foveae only slightly distinct, median basal area short and very transverse, delimited laterally by short, obliquely curved furrows, basal margin slightly projected posteriorly in most members, basal angles small, obtuse, rounded; elytra distinctly convex, with striae finely impressed, more or less punctate, lateral striae attenuated or effaced, recurrent stria terminated anteriorly with a bend or hook on interval 5 or 6, with a single discal seta on interval 3 next to stria three or without discal setae; preapical seta present, inserted next to stria 2; median lobe of male aedeagus of varied form, but endophallus membraneous, with spiny or scaly areas in some members, more or less sclerotized but without distinct sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD7FFCA3CFC6414FB587604" lastPageId="67" lastPageNumber="408" pageId="66" pageNumber="407" type="discussion">
<paragraph id="8BC0368BFFD7FFCB3CFC6414FCE572ED" blockId="66.[123,1174,453,1725]" pageId="66" pageNumber="407">
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<emphasis id="B90BEA99FFD7FFCB3CFC6414FEE6732C" bold="true" box="[171,296,1125,1149]" pageId="66" pageNumber="407">COMMENTS</emphasis>
</smallCapsWord>
.— In subtropical
<collectingCountry id="F368761BFFD7FFCB3DAF6414FDF4732D" box="[504,570,1125,1149]" name="China" pageId="66" pageNumber="407">China</collectingCountry>
, the “
<taxonomicName id="4C7F4D08FFD7FFCB3E286414FCCA732D" authorityName="Ueno" authorityYear="1953" box="[639,772,1125,1149]" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="455" phylum="Arthropoda" rank="genus">Epaphiopsis</taxonomicName>
Complex” of genera is represented mainly by
<taxonomicName id="4C7F4D08FFD7FFCB3CB964F4FDEF73CD" authority="Ueno (1962)" authorityName="Ueno" authorityYear="1962" box="[238,545,1157,1181]" class="Insecta" family="Carabidae" genus="Pseudepaphius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3CB964F4FE4173CD" box="[238,399,1157,1181]" italics="true" pageId="66" pageNumber="407">Pseudepaphius</emphasis>
<bibRefCitation id="EFEE4B7AFFD7FFCB3DC064F4FDEF73CD" author="UENO, S. I." box="[407,545,1157,1181]" pageId="66" pageNumber="407" pagination="41 - 74" refId="ref60936" refString="UENO, S. I. 1962. Primitive trechines of the subgenus Epaphiopsis. Memoirs of the College of Science, Kyoto Imperial University, Series B, 29: 41 - 74." type="journal article" year="1962">Uéno (1962)</bibRefCitation>
</taxonomicName>
, members of which are distinguished from true
<taxonomicName id="4C7F4D08FFD7FFCB387264F4FF5973ED" authorityName="Ueno" authorityYear="1953" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB387264F4FF5973ED" italics="true" pageId="66" pageNumber="407">Epaphiopsis</emphasis>
</taxonomicName>
members by their smooth pronotum and the presence of a single discal seta on interval 5 next to stria 5 (
<bibRefCitation id="EFEE4B7AFFD7FFCB3CB164B4FEA3738D" author="DEUVE, T." box="[230,365,1221,1245]" pageId="66" pageNumber="407" pagination="5 - 18" refId="ref58882" refString="DEUVE, T. 1995. Contribution a l'inventaire des Trechidae Trechinae de Chine et de Thailande (Coleoptera). Revue Francaise d'Entomologie (N. S.) 17: 5 - 18." type="journal article" year="1995">Deuve 1995</bibRefCitation>
). Members of our new genus,
<taxonomicName id="4C7F4D08FFD7FFCB3EE664B4FCA2738D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[689,876,1221,1245]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3EE664B4FCA2738D" box="[689,876,1221,1245]" italics="true" pageId="66" pageNumber="407">Trechepaphiopsis</emphasis>
</taxonomicName>
, are easily distinguished by their elytra chaetotaxy. Genus
<taxonomicName id="4C7F4D08FFD7FFCB3D9F6494FCAC73AD" authority="Ueno and Yin (1993)" authorityName="Ueno and Yin" authorityYear="1993" box="[456,866,1253,1277]" class="Insecta" family="Carabidae" genus="Junnanotrechus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3D9F6494FDBF73AD" box="[456,625,1253,1277]" italics="true" pageId="66" pageNumber="407">Junnanotrechus</emphasis>
Uéno and Yin (1993)
</taxonomicName>
also belongs in this generic complex and is probably closely related to
<taxonomicName id="4C7F4D08FFD7FFCB3E056574FD3D724D" authorityName="Ueno" authorityYear="1962" box="[594,755,1285,1309]" class="Insecta" family="Carabidae" genus="Pseudepaphius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3E056574FD3D724D" box="[594,755,1285,1309]" italics="true" pageId="66" pageNumber="407">Pseudepaphius</emphasis>
</taxonomicName>
, based on both morphological (
<bibRefCitation id="EFEE4B7AFFD7FFCB38196574FF70726D" author="DEUVE, T." pageId="66" pageNumber="407" pagination="67 - 72" refId="ref59043" refString="DEUVE, T. 2013 a. Nouveaux Trechini de la faune des litieres du Shaanxi et du Yunnan (Coleoptera, Caraboidea, Trechidae). Revue Francaise d'Entomologie, (N. S.) 33 [2011]: 67 - 72." type="journal article" year="2013">Deuve 2013a</bibRefCitation>
) and molecular (A. Faille, unpublished) data. The illustration of the mandibles of
<taxonomicName id="4C7F4D08FFD7FFCB38666557FDF8720D" authority="Belousov and Kabak" authorityName="Belousov and Kabak" class="Insecta" family="Carabidae" genus="Junnanotrechus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B90BEA99FFD7FFCB38666557FE85720D" italics="true" pageId="66" pageNumber="407">Junnanotrechus elegantulus</emphasis>
Belousov and Kabak
</taxonomicName>
(2014b,
<figureCitation id="13442A0EFFD7FFCB3ECF6534FD19720D" box="[664,727,1349,1373]" captionStart="FIGURE" captionStartId="2.[171,183,961,980]" captionTargetBox="[120,1176,216,952]" captionTargetId="figure-315@2.[120,1176,216,952]" captionTargetPageId="2" captionText="FIGURE 1. Map of Asia with study region outlined; scale line = m 500 km. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_ oceans_12000.jpg&gt; [last accessed 28 September 2016]." figureDoi="http://doi.org/10.5281/zenodo.13155285" httpUri="https://zenodo.org/record/13155285/files/figure.png" pageId="66" pageNumber="407">Fig. 1</figureCitation>
) confirms this phylogenetic affinity. The right mandible presents the same morphological features shared with
<taxonomicName id="4C7F4D08FFD7FFCB3F0E6514FC34722D" authorityName="Ueno" authorityYear="1962" box="[857,1018,1381,1405]" class="Insecta" family="Carabidae" genus="Pseudepaphius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3F0E6514FC34722D" box="[857,1018,1381,1405]" italics="true" pageId="66" pageNumber="407">Pseudepaphius</emphasis>
</taxonomicName>
(
<figureCitation id="13442A0EFFD7FFCB385F6514FBAE722D" box="[1032,1120,1381,1405]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="66" pageNumber="407">Fig. 16e</figureCitation>
) and
<taxonomicName id="4C7F4D08FFD7FFCB3C2C65F4FEF872CD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,310,1413,1437]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3C2C65F4FEF872CD" box="[123,310,1413,1437]" italics="true" pageId="66" pageNumber="407">Trechepaphiopsis</emphasis>
</taxonomicName>
members (
<figureCitation id="13442A0EFFD7FFCB3DFB65F4FDCB72CD" box="[428,517,1413,1437]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="66" pageNumber="407">Fig. 16f</figureCitation>
), namely the unfused premolar and the anterior tip of the retinaculum distinctly displaced anteriorly to form a separate tooth.
</paragraph>
<paragraph id="8BC0368BFFD7FFCB3CFC65B4FC39710D" blockId="66.[123,1174,453,1725]" pageId="66" pageNumber="407">
<emphasis id="B90BEA99FFD7FFCB3CFC65B4FE3A728C" bold="true" box="[171,500,1477,1501]" pageId="66" pageNumber="407">
<smallCapsWord id="8D26A057FFD7FFCB3CFC65B4FE98728C" baselines="1495,1495" box="[171,342,1477,1501]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="66" pageNumber="407">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFD7FFCB3D3765B8FE3A728C" baselines="1495" box="[352,500,1481,1500]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="66" pageNumber="407">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This genus currently is known only from the Gaoligong Shan region of western
<collectingRegion id="49BBF869FFD7FFCB3D206597FDFF72AD" box="[375,561,1509,1533]" country="China" name="Yunnan" pageId="66" pageNumber="407">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFD7FFCB3E6C6594FDB372AD" box="[571,637,1509,1533]" name="China" pageId="66" pageNumber="407">China</collectingCountry>
, where it is represented by the seven species treat- ed here. Three were previously described, all in genus
<taxonomicName id="4C7F4D08FFD7FFCB3EE86674FCDC714D" authorityName="Clairville" authorityYear="1806" box="[703,786,1541,1565]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD7FFCB3EE86674FCDC714D" box="[703,786,1541,1565]" italics="true" pageId="66" pageNumber="407">Trechus</emphasis>
</taxonomicName>
, and four are described here as new. Each species apparently occupies only a narrow geographical range within the Gaoligong Shan, but their combined known ranges cover all but the northernmost part of the study area.
</paragraph>
<paragraph id="8BC0368BFFD7FFCA3CFC6617FB587604" blockId="66.[123,1174,453,1725]" lastBlockId="67.[123,1174,219,340]" lastPageId="67" lastPageNumber="408" pageId="66" pageNumber="407">
As is reflected in the key to species presented below, only male members of most of these species can be reliably distinguished, and that only by extraction and examination of their genitalic structure. To date, only two of these species have been recorded as sympatric and syntopic, namely
<taxonomicName id="4C7F4D08FFD6FFCA3C8660ADFE7E77A3" authority="(Deuve)" baseAuthorityName="Deuve" box="[209,432,219,243]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="408" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFD6FFCA3C8660ADFE9C77A3" box="[209,338,220,243]" italics="true" pageId="67" pageNumber="408">T. unisetosa</emphasis>
(Deuve)
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFD6FFCA3DB060ADFDA077A3" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[487,622,220,243]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="408" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFD6FFCA3DB060ADFDA077A3" box="[487,622,220,243]" italics="true" pageId="67" pageNumber="408">T. uniporosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD6FFCA3E2160ADFD0477A3" box="[630,714,220,243]" pageId="67" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
(in Core Area 4). Two species,
<taxonomicName id="4C7F4D08FFD6FFCA387360ADFF697643" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="408" phylum="Arthropoda" rank="species" species="unisetulosa" status="sp. nov.">
<emphasis id="B90BEA99FFD6FFCA387360ADFF697643" italics="true" pageId="67" pageNumber="408">T. unisetulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD6FFCA3CF9608DFECF7643" box="[174,257,252,275]" pageId="67" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C7F4D08FFD6FFCA3D63608DFE7F7643" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[308,433,251,275]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="408" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFD6FFCA3D63608DFE7F7643" box="[308,433,251,275]" italics="true" pageId="67" pageNumber="408">T. unipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD6FFCA3DEF608DFDC47643" box="[440,522,252,275]" pageId="67" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
occur in the same general area but apparently have different, non-overlapping altitudinal ranges. So, at least for the present, locality data may aid in tentative identifications of females and undissected males, except for the first two species mentioned above.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,463 @@
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<mods:title id="28D63F629B449CF23895B245ED2B8AE9">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="A19A6D6C829DFF993DA988F7443ACE25">Deuve, Thierry</mods:namePart>
<mods:affiliation id="128D79540A05C02BF390B8A3B625C21A">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="6F92BF29C83BDC7431F4A3DD57521611">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="0C324FBD12CA6EB9453DC028BBF57BD2">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="DF3BF74F6FC3009AA7F7484AF9A57E2A" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:namePart id="73344CC942DCCB8EB736C8C7AF0AC493">Liang, Hongbin</mods:namePart>
<mods:affiliation id="976DD8B86F86A7DEACEBA0D03FDC1665">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<paragraph id="8BC0368BFFD8FFC43C2C64ADFCF473A4" blockId="77.[123,826,1244,1298]" box="[123,826,1244,1268]" pageId="77" pageNumber="418">
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<emphasis id="B90BEA99FFD8FFC43C2C64ADFCF473A4" bold="true" box="[123,826,1244,1268]" pageId="77" pageNumber="418">
<taxonomicName id="4C7F4D08FFD8FFC43C2C64ADFDD673A4" ID-CoL="8MC9H" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,536,1244,1268]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="418" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFD8FFC43C2C64ADFE0473A4" bold="true" box="[123,458,1244,1268]" italics="true" pageId="77" pageNumber="418">Trechepaphiopsis monochaeta</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFD8FFC43EB364ADFCF473A4" box="[740,826,1244,1268]" pageId="77" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD8FFC43C2C648DFE4F7242" box="[123,385,1276,1298]" pageId="77" pageNumber="418" type="description">
<paragraph id="8BC0368BFFD8FFC43C2C648DFE4F7242" blockId="77.[123,826,1244,1298]" box="[123,385,1276,1298]" pageId="77" pageNumber="418">
(
<figureCitation id="13442A0EFFD8FFC43CD5648DFF1D7241" box="[130,211,1276,1298]" captionStart="FIGURE" captionStartId="79.[171,183,1585,1604]" captionTargetBox="[126,1176,216,1572]" captionTargetId="figure-15@79.[120,1176,216,1573]" captionTargetPageId="79" captionText="FIGURE 30. Trechepaphiopsis monochaeta sp. nov.; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for T. monochaeta in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="77" pageNumber="418">Figs. 30</figureCitation>
,
<figureCitation id="13442A0EFFD8FFC43C8A648DFECD7242" box="[221,259,1276,1298]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="77" pageNumber="418">38b</figureCitation>
,
<figureCitation id="13442A0EFFD8FFC43D59648DFEFD7241" box="[270,307,1276,1297]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="77" pageNumber="418">42a</figureCitation>
,
<figureCitation id="13442A0EFFD8FFC43D6A648DFEB57242" box="[317,379,1276,1298]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="77" pageNumber="418">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD8FFC73CFC6554FD687505" lastPageId="78" lastPageNumber="419" pageId="77" pageNumber="418" type="materials_examined">
<paragraph id="8BC0368BFFD8FFC43CFC6554FB5B71ED" blockId="77.[123,1174,1317,1725]" pageId="77" pageNumber="418">
<emphasis id="B90BEA99FFD8FFC43CFC6554FEAF726B" bold="true" box="[171,353,1317,1341]" pageId="77" pageNumber="418">
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</emphasis>
.—
<materialsCitation id="3B173CD6FFD8FFC43DC56554FDFC72ED" collectingDate="2000-07-15" collectingDateMax="2000-07-19" collectingDateMin="2000-07-15" collectorName="C. E. Griswold" country="China" county="Gaoligong Shan" elevation="2775" latitude="27.71503" location="Nujiang State Nature Reserve" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" pageId="77" pageNumber="418" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFD8FFC43DC56554FE36726D" box="[402,504,1317,1341]" pageId="77" pageNumber="418" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: labeled: “CASENT 1007387”/ “
<collectingCountry id="F368761BFFD8FFC43CD36534FF13720D" box="[132,221,1349,1373]" name="China" pageId="77" pageNumber="418">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFD8FFC43CBD6537FE68720D" box="[234,422,1349,1373]" country="China" name="Yunnan" pageId="77" pageNumber="418">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFD8FFC43DE56534FDAD720D" box="[434,611,1349,1373]" pageId="77" pageNumber="418">Gaoligong Shan</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFD8FFC43E276534FCF0720D" box="[624,830,1349,1373]" pageId="77" pageNumber="418">Nujiang Prefecture</collectingMunicipality>
,
<location id="8EA06050FFD8FFC43F1C6534FB5C720D" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFD8FFC43F1C6534FB5C720D" box="[843,1170,1349,1373]" country="China" county="Gaoligong Shan" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" name="Nujiang State Nature Reserve" pageId="77" pageNumber="418" stateProvince="Yunnan">Nujiang State Nature Reserve</location>
, No. 12
<location id="8EA06050FFD8FFC43C9E6514FE43722D" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFD8FFC43C9E6514FE43722D" box="[201,397,1381,1405]" country="China" county="Gaoligong Shan" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Nujiang Prefecture" name="Bridge Camp area" pageId="77" pageNumber="418" stateProvince="Yunnan">Bridge Camp area</location>
, 16.3 airkm
<locationDeviation id="9A722FA9FFD8FFC43E466517FD72722D" box="[529,700,1381,1405]" location="Gongshan" pageId="77" pageNumber="418">W of Gongshan</locationDeviation>
,”/ “
<geoCoordinate id="EE4B504CFFD8FFC43EB56514FCAB722D" box="[738,869,1381,1405]" degrees="27.71503" direction="north" orientation="latitude" pageId="77" pageNumber="418" precision="1" value="27.71503">N27.71503°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFD8FFC43F396514FC21722D" box="[878,1007,1381,1405]" degrees="98.50244" direction="east" orientation="longitude" pageId="77" pageNumber="418" precision="1" value="98.50244">E98.50244°</geoCoordinate>
,
<quantity id="4C879B6EFFD8FFC43FAD6514FB82722D" box="[1018,1100,1381,1405]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="77" pageNumber="418" unit="m" value="2775.0">
<elevation id="0052D1B8FFD8FFC43FAD6514FB82722D" box="[1018,1100,1381,1405]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="77" pageNumber="418" unit="m" value="2775.0">2775 m</elevation>
</quantity>
,
<date id="FFC1104BFFD8FFC438006514FF2972CD" pageId="77" pageNumber="418" value="2000-07-15" valueMax="2000-07-19" valueMin="2000-07-15">
<collectingDate id="EF85E9A3FFD8FFC438006514FF2972CD" pageId="77" pageNumber="418" value="2000-07-15" valueMax="2000-07-19" valueMin="2000-07-15">15-19 July 2000</collectingDate>
</date>
, Stop#00-23A, D.H.
<taxonomicName id="4C7F4D08FFD8FFC43D9B65F4FD8672CD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[460,584,1413,1437]" pageId="77" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFD8FFC43E0465F4FD3E72CD" box="[595,752,1413,1437]" pageId="77" pageNumber="418">C.E. Griswold</collectorName>
, Liang H.-B., D. Ubick, &amp; Dong D.-Z. collectors”/ “IMAGE” [green label]/ “
</materialsCitation>
<typeStatus id="54C48829FFD8FFC43E6565D4FD0F72ED" box="[562,705,1445,1469]" pageId="77" pageNumber="418" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFD8FFC43E8765D4FBBC72ED" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[720,1138,1445,1469]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="418" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFD8FFC43E8765D4FBD372ED" box="[720,1053,1445,1469]" italics="true" pageId="77" pageNumber="418">Trechepaphiopsis monochaeta</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFD8FFC43D5265B7FE92728D" box="[261,348,1478,1501]" pageId="77" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label].
<materialsCitation id="3B173CD6FFD8FFC43EE565B7FD44716D" collectionCode="CAS, IOZ, MNHN" country="China" pageId="77" pageNumber="418" specimenCount="31" specimenCount-female="17" specimenCount-male="14" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFD8FFC43EE565B7FCD4728E" box="[690,794,1478,1502]" pageId="77" pageNumber="418" type="paratype">Paratypes</typeStatus>
(at total of 56):
<specimenCount id="9D79FD02FFD8FFC43F8C65B4FBF1728D" box="[987,1087,1477,1501]" count="14" pageId="77" pageNumber="418" type="male">14 males</specimenCount>
and
<specimenCount id="9D79FD02FFD8FFC4382C65B4FF0072AD" count="17" pageId="77" pageNumber="418" type="female">17 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFD8FFC43CAE6594FEE172AD" box="[249,303,1509,1533]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="77" pageNumber="418" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFD8FFC43D6C6594FEA472AD" box="[315,362,1509,1533]" pageId="77" pageNumber="418">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFD8FFC43D226594FE0672AD" box="[373,456,1509,1533]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="77" pageNumber="418">MNHN</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFD8FFC43EDA6594FD2372AD" box="[653,749,1509,1533]" pageId="77" pageNumber="418" type="holotype">holotype</typeStatus>
, except first label “CASENT 1007388” to “CASENT 1007400” and “CASENT 1021923” and “CASENT 1007401” to “CASENT 1007416” and “CASENT 1021924”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FFD8FFC43EC26654FB5F71ED" collectingDate="2000-06-30" collectingDateMax="2000-07-05" collectingDateMin="2000-06-30" collectionCode="CAS, IOZ" collectorName="Stop &amp; J.P. DUMBACHER &amp; C. E. Griswold" country="China" county="Gongshan County" elevation="2830" latitude="27.62947" location="Nujiang Prefecture" longLatPrecision="1" longitude="98.6201" municipality="Gaoligong Shan" pageId="77" pageNumber="418" specimenCount="24" specimenCount-female="14" specimenCount-male="10" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFD8FFC43EC26654FD3A716D" box="[661,756,1573,1597]" count="10" pageId="77" pageNumber="418" type="male">10 males</specimenCount>
and
<specimenCount id="9D79FD02FFD8FFC43F706654FC55716D" box="[807,923,1573,1597]" count="14" pageId="77" pageNumber="418" type="female">14 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFD8FFC43F926654FC35716D" box="[965,1019,1573,1597]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="77" pageNumber="418" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFD8FFC438516654FBFA716D" box="[1030,1076,1573,1597]" pageId="77" pageNumber="418">IOZ</collectionCode>
) labeled: “CASENT 1006508” to “CASENT 1006517” and “CASENT 1006518” to “CASENT 1006530” and “CASENT 1021922”, respectively/ “
<collectingCountry id="F368761BFFD8FFC43E796614FD49712D" box="[558,647,1637,1661]" name="China" pageId="77" pageNumber="418">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFD8FFC43EC66617FC85712D" box="[657,843,1637,1661]" country="China" name="Yunnan" pageId="77" pageNumber="418">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFD8FFC43F036614FBCD712D" box="[852,1027,1637,1661]" pageId="77" pageNumber="418">Gaoligong Shan</collectingMunicipality>
,
<location id="8EA06050FFD8FFC4385A6614FF0471CD" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFD8FFC4385A6614FF0471CD" country="China" county="Gongshan County" latitude="27.62947" longLatPrecision="1" longitude="98.6201" municipality="Gaoligong Shan" name="Nujiang Prefecture" pageId="77" pageNumber="418" stateProvince="Yunnan">Nujiang Prefecture</location>
,
<collectingCounty id="62A14E07FFD8FFC43C8966F4FE6071CD" box="[222,430,1669,1693]" pageId="77" pageNumber="418">Gongshan County</collectingCounty>
,
<location id="8EA06050FFD8FFC43D9466F4FDF471CD" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFD8FFC43D9466F4FDF471CD" box="[451,570,1669,1693]" country="China" county="Gongshan County" latitude="27.62947" longLatPrecision="1" longitude="98.6201" municipality="Gaoligong Shan" name="Dazhu He" pageId="77" pageNumber="418" stateProvince="Yunnan">Dazhu He</location>
drainage, 13.5 airkm
<locationDeviation id="9A722FA9FFD8FFC43F0666F4FBEC71CD" box="[849,1058,1669,1693]" location="Gongshan" pageId="77" pageNumber="418">SW of Gongshan</locationDeviation>
,
<quantity id="4C879B6EFFD8FFC4386066F4FB4A71CD" box="[1079,1156,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="2.83" pageId="77" pageNumber="418" unit="m" value="2830.0">
<elevation id="0052D1B8FFD8FFC4386066F4FB4A71CD" box="[1079,1156,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="2.83" pageId="77" pageNumber="418" unit="m" value="2830.0">2830m</elevation>
</quantity>
”/ “
<geoCoordinate id="EE4B504CFFD8FFC43CD366D4FEC671ED" box="[132,264,1701,1725]" degrees="27.62947" direction="north" orientation="latitude" pageId="77" pageNumber="418" precision="1" value="27.62947">N27.62947°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFD8FFC43D4766D4FE5F71ED" box="[272,401,1701,1725]" degrees="98.62010" direction="east" orientation="longitude" pageId="77" pageNumber="418" precision="1" value="98.6201">E98.62010°</geoCoordinate>
,
<date id="FFC1104BFFD8FFC43DC866D4FD4471ED" box="[415,650,1701,1725]" pageId="77" pageNumber="418" value="2000-06-30" valueMax="2000-07-05" valueMin="2000-06-30">
<collectingDate id="EF85E9A3FFD8FFC43DC866D4FD4471ED" box="[415,650,1701,1725]" pageId="77" pageNumber="418" value="2000-06-30" valueMax="2000-07-05" valueMin="2000-06-30">30 June- 5 July 2000</collectingDate>
</date>
,
<collectorName id="268A535DFFD8FFC43EC066D4FD0571ED" box="[663,715,1701,1725]" pageId="77" pageNumber="418">Stop</collectorName>
#00-17I,
<collectorName id="268A535DFFD8FFC43F7B66D7FCAE71ED" box="[812,864,1702,1725]" name="J.P. DUMBACHER" pageId="77" pageNumber="418">D.H.</collectorName>
<taxonomicName id="4C7F4D08FFD8FFC43F3E66D4FC2B71ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[873,997,1701,1725]" pageId="77" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFD8FFC43FA466D4FB5F71ED" box="[1011,1169,1701,1725]" pageId="77" pageNumber="418">C.E. Griswold</collectorName>
</materialsCitation>
,
</paragraph>
<paragraph id="8BC0368BFFDBFFC73C2C60AAFE6F76C4" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">
<materialsCitation id="3B173CD6FFDBFFC73C2C60AAFE5376C4" collectingDate="2006-08-18" collectionCode="CAS" collectorName="Liang H. - B. &amp; Dong D. - Z." country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="78" pageNumber="419" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="268A535DFFDBFFC73C2C60AAFECA77A2" box="[123,260,219,243]" pageId="78" pageNumber="419">Liang H.-B.</collectorName>
, D. Ubick, &amp;
<collectorName id="268A535DFFDBFFC73DED60AAFDF377A2" box="[442,573,219,243]" pageId="78" pageNumber="419">Dong D.-Z.</collectorName>
collectors”;
<specimenCount id="9D79FD02FFDBFFC73E8260AAFCF477A3" box="[725,826,219,243]" count="1" pageId="78" pageNumber="419" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFDBFFC73F2360AAFC6777A3" box="[884,937,219,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="419" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1025812”/ “
<collectingCountry id="F368761BFFDBFFC73D5E608AFEAB7643" box="[265,357,251,275]" name="China" pageId="78" pageNumber="419">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDBFFC73D2E608DFE1E7643" box="[377,464,252,275]" country="China" name="Yunnan" pageId="78" pageNumber="419">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFDBFFC73DB3608AFD787643" box="[484,694,251,275]" pageId="78" pageNumber="419">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFDBFFC73E9C608AFC287643" box="[715,998,251,275]" pageId="78" pageNumber="419">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FFDBFFC73C2C616AFEFF7663" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFDBFFC73C2C616AFEFF7663" box="[123,305,283,307]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="78" pageNumber="419" stateProvince="Yunnan">Kawakarpu Shan</location>
<locationDeviation id="9A722FA9FFDBFFC73D60616DFDAB7663" box="[311,613,283,308]" location="Chukuai Lake" pageId="78" pageNumber="419">
<quantity id="4C879B6EFFDBFFC73D60616DFE4E7663" box="[311,384,283,308]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="78" pageNumber="419" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/, “
<geoCoordinate id="EE4B504CFFDBFFC73F43616AFC597663" box="[788,919,283,307]" degrees="27.97686" direction="north" orientation="latitude" pageId="78" pageNumber="419" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFDBFFC73FF1616DFBFA7664" box="[934,1076,284,308]" degrees="098.47799" direction="east" orientation="longitude" pageId="78" pageNumber="419" precision="1" value="98.47799">E098.47799°</geoCoordinate>
,
<quantity id="4C879B6EFFDBFFC73868616AFB5F7663" box="[1087,1169,283,307]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="78" pageNumber="419" unit="m" value="3750.0">
<elevation id="0052D1B8FFDBFFC73868616AFB5F7663" box="[1087,1169,283,307]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="78" pageNumber="419" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFFDBFFC73C2C614DFEED7604" box="[123,291,316,340]" pageId="78" pageNumber="419" value="2006-08-18">
<collectingDate id="EF85E9A3FFDBFFC73C2C614DFEED7604" box="[123,291,316,340]" pageId="78" pageNumber="419" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-078 D.H.
<taxonomicName id="4C7F4D08FFDBFFC73E0F614DFD1F7604" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[600,721,316,340]" pageId="78" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”. All
<typeStatus id="54C48829FFDBFFC73F22614DFC157604" box="[885,987,316,340]" pageId="78" pageNumber="419" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFDBFFC73D4B612DFE6D7624" box="[284,419,348,372]" pageId="78" pageNumber="419" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDBFFC73DFA612DFC8A7623" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[429,836,348,372]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="419" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFDBFFC73DFA612DFD3A7624" box="[429,756,348,372]" italics="true" pageId="78" pageNumber="419">Trechepaphiopsis monochaeta</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFDBFFC73FA1612DFB857623" box="[1014,1099,348,371]" pageId="78" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDBFFC73CFC61EDFC237684" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">
<materialsCitation id="3B173CD6FFDBFFC73CFC61EDFC277684" country="China" county="Gongshan County" elevation="2775" latitude="27.71503" location="Qiqi Trail" longLatPrecision="1" longitude="98.50244" municipality="Gaoligong Shan" pageId="78" pageNumber="419" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFDBFFC73CFC61EDFE9676E3" bold="true" box="[171,344,412,436]" pageId="78" pageNumber="419">
<smallCapsWord id="8D26A057FFDBFFC73CFC61EDFF2A76E3" baselines="430,430" box="[171,228,412,436]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="78" pageNumber="419">
<typeStatus id="54C48829FFDBFFC73CFC61EDFF2A76E3" box="[171,228,412,436]" pageId="78" pageNumber="419">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFDBFFC73CBA61D1FE9676E3" baselines="430" box="[237,344,416,435]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="78" pageNumber="419">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFDBFFC73DD661EDFE0D76E4" box="[385,451,412,436]" name="China" pageId="78" pageNumber="419">China</collectingCountry>
,
<collectingRegion id="49BBF869FFDBFFC73D8761ECFDEA76E4" box="[464,548,413,436]" country="China" name="Yunnan" pageId="78" pageNumber="419">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFDBFFC73E6761EDFD2F76E4" box="[560,737,412,436]" pageId="78" pageNumber="419">Gaoligong Shan</collectingMunicipality>
,
<collectingCounty id="62A14E07FFDBFFC73EBA61EDFC7C76E4" box="[749,946,412,436]" pageId="78" pageNumber="419">Gongshan County</collectingCounty>
,
<location id="8EA06050FFDBFFC73FEA61EDFBE676E4" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFDBFFC73FEA61EDFBE676E4" box="[957,1064,412,436]" country="China" county="Gongshan County" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Gaoligong Shan" name="Qiqi Trail" pageId="78" pageNumber="419" stateProvince="Yunnan">Qiqi Trail</location>
at No. 12
<location id="8EA06050FFDBFFC73C2C61CCFE8F7684" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFDBFFC73C2C61CCFE8F7684" box="[123,321,445,469]" country="China" county="Gongshan County" latitude="27.71503" longLatPrecision="1" longitude="98.50244" municipality="Gaoligong Shan" name="Bridge Camp area" pageId="78" pageNumber="419" stateProvince="Yunnan">Bridge Camp area</location>
, 16.3 airkm
<locationDeviation id="9A722FA9FFDBFFC73D9061CCFDBA7685" box="[455,628,445,469]" location="Gongshan" pageId="78" pageNumber="419">W of Gongshan</locationDeviation>
,
<geoCoordinate id="EE4B504CFFDBFFC73E2861CDFCCF7684" box="[639,769,444,468]" degrees="27.71503" direction="north" orientation="latitude" pageId="78" pageNumber="419" precision="1" value="27.71503">N27.71503°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFDBFFC73F5E61CDFC447684" box="[777,906,444,468]" degrees="98.50244" direction="east" orientation="longitude" pageId="78" pageNumber="419" precision="1" value="98.50244">E98.50244°</geoCoordinate>
,
<quantity id="4C879B6EFFDBFFC73FC261CDFC277684" box="[917,1001,444,468]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="78" pageNumber="419" unit="m" value="2775.0">
<elevation id="0052D1B8FFDBFFC73FC261CDFC277684" box="[917,1001,444,468]" metricMagnitude="3" metricUnit="m" metricValue="2.775" pageId="78" pageNumber="419" unit="m" value="2775.0">2775 m</elevation>
</quantity>
</materialsCitation>
,
</paragraph>
<paragraph id="8BC0368BFFDBFFC73CFC61ACFD687505" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">
<emphasis id="B90BEA99FFDBFFC73CFC61ACFE3676A3" bold="true" box="[171,504,477,501]" pageId="78" pageNumber="419">
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<smallCapsWord id="8D26A057FFDBFFC73D6F6191FE9A76A3" baselines="494" box="[312,340,480,499]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="78" pageNumber="419">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFDBFFC73D0A6191FE7E76A3" baselines="494" box="[349,432,480,499]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="78" pageNumber="419">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFDBFFC73DEC6191FE3676A3" baselines="494" box="[443,504,480,499]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="78" pageNumber="419">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFDBFFC73F5F61ACFC4476A5" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[776,906,477,501]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="419" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFDBFFC73F5F61ACFC4476A5" box="[776,906,477,501]" italics="true" pageId="78" pageNumber="419">monochaeta</emphasis>
</taxonomicName>
, is an adjective derived from the Greek words,
<emphasis id="B90BEA99FFDBFFC73D23618CFE7D7545" box="[372,435,509,533]" italics="true" pageId="78" pageNumber="419">μόΝΟς</emphasis>
(transliterated into Latin as
<emphasis id="B90BEA99FFDBFFC73EB4618FFCD07545" box="[739,798,510,533]" italics="true" pageId="78" pageNumber="419">mono</emphasis>
), meaning one or single, and
<emphasis id="B90BEA99FFDBFFC7380A618CFB5B7545" box="[1117,1173,509,533]" italics="true" pageId="78" pageNumber="419">χαίτα</emphasis>
(transliterated into Latin as
<emphasis id="B90BEA99FFDBFFC73DFE626CFE3E7565" box="[425,496,541,565]" italics="true" pageId="78" pageNumber="419">chaeta</emphasis>
), meaning hair or bristle. The name refers to the single discal seta found on the elytra of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDBFFC73CFC622FFDD774C8" pageId="78" pageNumber="419" type="diagnosis">
<paragraph id="8BC0368BFFDBFFC73CFC622FFDD774C8" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">
<smallCapsWord id="8D26A057FFDBFFC73CFC622FFEEE7524" baselines="624,623" box="[171,288,606,630]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="78" pageNumber="419">
<emphasis id="B90BEA99FFDBFFC73CFC622FFEEE7524" bold="true" box="[171,288,606,630]" pageId="78" pageNumber="419">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFDBFFC73E10622FFD6D7526" box="[583,675,606,630]" captionStart="FIGURE" captionStartId="79.[171,183,1585,1604]" captionTargetBox="[126,1176,216,1572]" captionTargetId="figure-15@79.[120,1176,216,1573]" captionTargetPageId="79" captionText="FIGURE 30. Trechepaphiopsis monochaeta sp. nov.; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for T. monochaeta in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="78" pageNumber="419">Fig. 30a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size small (BL = 2.8 to 3.0 mm), apterous; body color reddish brown, dorsum shiny, very slightly iridescent; eyes small but convex; tempora convex and sparsely pubescent; pronotum transverse (PW/PL = 1.38), with basal angles obtuse and rounded; elytra convex, medial four or five striae distinctly impressed, more lateral striae more or less effaced; recurrent stria terminated anteriorly in presumed location of interval 6; only one discal setiferous pore present, inserted anterior to and next to stria 3; preapical seta present, inserted next to stria 2; median lobe of male aedeagus (
<figureCitation id="13442A0EFFDBFFC73F6B634EFC577407" box="[828,921,831,855]" captionStart="FIGURE" captionStartId="79.[171,183,1585,1604]" captionTargetBox="[126,1176,216,1572]" captionTargetId="figure-15@79.[120,1176,216,1573]" captionTargetPageId="79" captionText="FIGURE 30. Trechepaphiopsis monochaeta sp. nov.; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for T. monochaeta in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="78" pageNumber="419">Fig. 30b</figureCitation>
) with apex moderately broad and apically rounded, slightly deflected ventrally; endophallus with an elongate and scaly sclerotized area not narrowed apically.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDBFFD93CFC63D1FF7A7563" lastPageId="80" lastPageNumber="421" pageId="78" pageNumber="419" type="description">
<paragraph id="8BC0368BFFDBFFC73CFC63D1FBC47488" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">
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<emphasis id="B90BEA99FFDBFFC73CFC63D1FEF474E6" bold="true" box="[171,314,928,952]" pageId="78" pageNumber="419">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size small, BL = 2.8 to 3.0 mm. Body color reddish brown, appendages slightly paler, yellowish brown, palpi paler yellow; dorsum shiny, slightly iridescent.
</paragraph>
<paragraph id="8BC0368BFFDBFFC73CFC6391FD76724A" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">
Head. Medium in size; eyes small but moderately convex, their diameter about equal to length of tempora, the latter distinctly convex and sparsely pubescent. Frons convex, with frontal furrows slightly angulate and deeply impressed, continuous to posterior margins of tempora, not or slightly attenuated posterior to margins of eyes. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly emarginate or concave.
<collectingRegion id="49BBF869FFDBFFC73D1F64F3FE6C73C9" box="[328,418,1154,1177]" country="Myanmar" name="Chin" pageId="78" pageNumber="419">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFDBFFC73E8764F3FCE473C9" box="[720,810,1154,1177]" country="Myanmar" name="Chin" pageId="78" pageNumber="419">Mentum</collectingRegion>
with medial tooth broad and apically truncate or faintly bifid, about half the length of the lateral lobes. Submentum with six setae anteriorly. Gula broad. Genae with a single ventral seta one each side. Antennae short, with only about 1.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter.
</paragraph>
<paragraph id="8BC0368BFFDBFFC73CFC6552FF72716C" blockId="78.[123,1174,219,1725]" pageId="78" pageNumber="419">Pronotum. Transverse (ratio PW/PL = 1.38), with greatest width slightly anterior to middle; lateral margins rounded, slightly more so anteriorly, not or only slightly straighten posteriorly just anterior to basal angles, the latter obtuse and bluntly or rounded. Disc convex and smooth, median longitudinal impression finely impressed, continuous from middle of apical median swelling to posterior margin; basal foveae very small and shallow, faintly impressed; median basal area well defined but short and transverse, smooth but with several small longitudinal foveae. Lateral borders slender and finely relexed dorsally, lateral grooves deeply impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
<paragraph id="8BC0368BFFDBFFD93CFC6635FDA97663" blockId="78.[123,1174,219,1725]" lastBlockId="80.[123,1174,219,1140]" lastPageId="80" lastPageNumber="421" pageId="78" pageNumber="419">Elytra. Ovoid, only slightly narrowed anteriorly and posteriorly, humeri distinct but rounded. Disc convex, striae finely impressed and not or only faintly punctate, the medial three or four striae clearly impressed, the more lateral striae more or less effaced, but still evident. Parascutellar striole present and distinct. Recurrent stria deeply impressed, its anterior recurved end terminated in the posterior end of stria 5. Only one discal setiferous pore present, inserted slightly anterior to middle next to stria 3. Preapical seta present, inserted on interval 3 next to stria 2 and about equidistant from apical and sutural elytral margin.</paragraph>
<caption id="DF006603FFDAFFC63CFC6640FDF47126" ID-DOI="http://doi.org/10.5281/zenodo.13155340" ID-Zenodo-Dep="13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="79" pageNumber="420" startId="79.[171,183,1585,1604]" targetBox="[126,1176,216,1572]" targetPageId="79" targetType="figure">
<paragraph id="8BC0368BFFDAFFC63CFC6640FDF47126" blockId="79.[123,1173,1584,1655]" pageId="79" pageNumber="420">
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30.
<taxonomicName id="4C7F4D08FFDAFFC63D5B6641FDC17113" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[268,527,1584,1603]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="420" phylum="Arthropoda" rank="species" species="monochaeta" status="sp. nov.">
<emphasis id="B90BEA99FFDAFFC63D5B6641FDC17113" box="[268,527,1584,1603]" italics="true" pageId="79" pageNumber="420">Trechepaphiopsis monochaeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFDAFFC63E426640FD947114" box="[533,602,1585,1604]" pageId="79" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFDAFFC63FD5663AFC31710D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[898,1023,1610,1629]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="420" phylum="Arthropoda" rank="species" species="monochaeta">
<emphasis id="B90BEA99FFDAFFC63FD5663AFC31710D" box="[898,1023,1610,1629]" italics="true" pageId="79" pageNumber="420">T. monochaeta</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFC5FFD93CFC614DFE9E7623" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFC5FFD93CFC610AFB8576E3" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFC5FFD93CFC61CAFF7A7563" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFC5FFD93DA961CAFD907683" box="[510,606,443,467]" captionStart="FIGURE" captionStartId="79.[171,183,1585,1604]" captionTargetBox="[126,1176,216,1572]" captionTargetId="figure-15@79.[120,1176,216,1573]" captionTargetPageId="79" captionText="FIGURE 30. Trechepaphiopsis monochaeta sp. nov.; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for T. monochaeta in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="80" pageNumber="421">Fig. 30b</figureCitation>
) with sagittal aileron very small, shaft moderately broad basally, then gradually narrowed, with apex moderately broad and apically rounded, slightly deflected ventrally; endophallus with an elongate and scaly sclerotized area not narrowed apically.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFC5FFD93CFC624AFB5C7324" pageId="80" pageNumber="421" type="discussion">
<paragraph id="8BC0368BFFC5FFD93CFC624AFDAE7583" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">
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<emphasis id="B90BEA99FFC5FFD93CFC624AFEE67502" bold="true" box="[171,296,571,595]" pageId="80" pageNumber="421">COMMENTS</emphasis>
</smallCapsWord>
.— This species appears to be very closely related (perhaps sister species) to
<taxonomicName id="4C7F4D08FFC5FFD938D5624DFF107523" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFC5FFD938D5624DFF107523" italics="true" pageId="80" pageNumber="421">T. unipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFC5FFD93CB3622DFEF77523" box="[228,313,604,627]" pageId="80" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
, but its male members can be distinguished from those of the latter by having the base of the median lobe of the male aedeagus broader (narrower in
<taxonomicName id="4C7F4D08FFC5FFD93F19620DFC0575C3" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[846,971,635,659]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="unipilosa">
<emphasis id="B90BEA99FFC5FFD93F19620DFC0575C3" box="[846,971,635,659]" italics="true" pageId="80" pageNumber="421">T. unipilosa</emphasis>
</taxonomicName>
members, see
<figureCitation id="13442A0EFFC5FFD9383C620AFF6D75E3" captionStart="FIGURE" captionStartId="82.[171,183,1585,1604]" captionTargetBox="[137,1176,216,1573]" captionTargetId="figure-11@82.[120,1176,216,1573]" captionTargetPageId="82" captionText="FIGURE 31. Trechepaphiopsis unipilosa sp. nov.; a. Dorsal habitus (CASENT1023757). b. Median lobe of aedeagus of male (CASENT1023757), left lateral aspect. c. Map of locality records (red circles) for T. unipilosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155342" httpUri="https://zenodo.org/record/13155342/files/figure.png" pageId="80" pageNumber="421">Fig. 31b</figureCitation>
) and the scaly sclerotized area of the endophallus of more equal width throughout and not narrowed distally (as it is in
<taxonomicName id="4C7F4D08FFC5FFD93DDB62CDFDC47583" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[396,522,699,723]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="unipilosa">
<emphasis id="B90BEA99FFC5FFD93DDB62CDFDC47583" box="[396,522,699,723]" italics="true" pageId="80" pageNumber="421">T. unipilosa</emphasis>
</taxonomicName>
males).
</paragraph>
<paragraph id="8BC0368BFFC5FFD93CFC62ADFD3A74E3" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">
<emphasis id="B90BEA99FFC5FFD93CFC62ADFE6875A2" bold="true" box="[171,422,732,756]" pageId="80" pageNumber="421">
<smallCapsWord id="8D26A057FFC5FFD93CFC62ADFEC775A2" baselines="750,749" box="[171,265,732,756]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="80" pageNumber="421">HABITAT</smallCapsWord>
<smallCapsWord id="8D26A057FFC5FFD93D4562AEFE6875A2" baselines="749" box="[274,422,735,754]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="80" pageNumber="421">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been collected by sifting leaf litter and mosses from the floor of closed canopy forest with an understory of ferns and large-leafed (ca.
<quantity id="4C879B6EFFC5FFD93C2C636DFF737463" box="[123,189,796,820]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="80" pageNumber="421" unit="cm" value="60.0">60 cm</quantity>
long leaves)
<taxonomicName id="4C7F4D08FFC5FFD93D1D636DFDC37463" box="[330,525,796,820]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="80" pageNumber="421" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEA99FFC5FFD93D1D636DFE267464" box="[330,488,796,820]" italics="true" pageId="80" pageNumber="421">Rhododendron</emphasis>
sp.
</taxonomicName>
at elevations ranging from
<quantity id="4C879B6EFFC5FFD93F66636AFC177463" box="[817,985,795,820]" metricMagnitude="3" metricUnit="m" metricValue="2.8024999999999998" metricValueMax="2.83" metricValueMin="2.775" pageId="80" pageNumber="421" unit="m" value="2802.5" valueMax="2830.0" valueMin="2775.0">2775 to 2830 m</quantity>
(
<figureCitation id="13442A0EFFC5FFD93FB0636DFB8A7464" box="[999,1092,796,820]" captionStart="FIGURE" captionStartId="104.[171,183,1626,1645]" captionTargetBox="[134,1163,217,1607]" captionTargetId="figure-12@104.[134,1163,217,1607]" captionTargetPageId="104" captionText="FIGURE 42. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Cikai Township, Qiqi Trail at No. 12 Bridge Camp area, elevation 2775 m; habitat in which specimens of Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. b. Fugong County, Shiyueliang Township, 11.5 km above Shibali on Shibali Road, elevation 3290 m; habitat in which the holotype of Trechus pseudoqiqiensis sp. nov. was collect- ed. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155370" httpUri="https://zenodo.org/record/13155370/files/figure.png" pageId="80" pageNumber="421">Fig. 42a</figureCitation>
). At the lowest elevation, specimens of
<taxonomicName id="4C7F4D08FFC5FFD93D84634DFD417403" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[467,655,828,852]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="qiqiensis">
<emphasis id="B90BEA99FFC5FFD93D84634DFD417403" box="[467,655,828,852]" italics="true" pageId="80" pageNumber="421">Trechus qiqiensis</emphasis>
</taxonomicName>
were collected in the same litter samples.
<specimenCount id="9D79FD02FFC5FFD9383E634DFF2E7424" count="1" pageId="80" pageNumber="421" type="generic">One specimen</specimenCount>
was also found under stones or wood chips in a disturbed open meadow area surround- ed by one meter high
<taxonomicName id="4C7F4D08FFC5FFD93D32630DFDCD74C4" box="[357,515,892,916]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="80" pageNumber="421" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFC5FFD93D32630DFDCD74C4" box="[357,515,892,916]" italics="true" pageId="80" pageNumber="421">Rhododendron</emphasis>
</taxonomicName>
thickets at an elevation of
<quantity id="4C879B6EFFC5FFD93F71630DFCB674C3" box="[806,888,892,916]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="80" pageNumber="421" unit="m" value="3750.0">3750 m</quantity>
(
<figureCitation id="13442A0EFFC5FFD93FD0630DFC2B74C4" box="[903,997,892,916]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="80" pageNumber="421">Fig. 38b</figureCitation>
); and specimens of
<taxonomicName id="4C7F4D08FFC5FFD93CCF63EDFE7674E4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[152,440,924,948]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="421" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FFC5FFD93CCF63EDFE7674E4" box="[152,440,924,948]" italics="true" pageId="80" pageNumber="421">Queinnectrechus griswoldi</emphasis>
</taxonomicName>
were found in this same area
</paragraph>
<paragraph id="8BC0368BFFC5FFD93CFC63CAFD5B7364" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">
<emphasis id="B90BEA99FFC5FFD93CFC63CAFD4E7483" bold="true" box="[171,640,955,979]" pageId="80" pageNumber="421">
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<emphasis id="B90BEA99FFC5FFD93EDD63CDFCC17482" bold="true" box="[650,783,956,980]" pageId="80" pageNumber="421">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FFC5FFD93F4E63CDFE0B7344" location="We" pageId="80" pageNumber="421" specimenCount="114" specimenCount-female="34" specimenCount-male="23" stateProvince="Shan">
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<collectingRegion id="49BBF869FFC5FFD93F4E63CDFC9A7482" box="[793,852,956,980]" country="Myanmar" name="Shan" pageId="80" pageNumber="421">SHAN</collectingRegion>
</emphasis>
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.—
<figureCitation id="13442A0EFFC5FFD93FD763CDFC107484" box="[896,990,956,980]" captionStart="FIGURE" captionStartId="79.[171,183,1585,1604]" captionTargetBox="[126,1176,216,1572]" captionTargetId="figure-15@79.[120,1176,216,1573]" captionTargetPageId="79" captionText="FIGURE 30. Trechepaphiopsis monochaeta sp. nov.; a. Dorsal habitus (CASENT1007411). b. Median lobe of aedeagus of male (CASENT1007411), left lateral aspect. c. Map of locality records (red circles) for T. monochaeta in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155340" httpUri="https://zenodo.org/record/13155340/files/figure.png" pageId="80" pageNumber="421">Fig. 30c</figureCitation>
.
<location id="8EA06050FFC5FFD93FBC63CDFBC07483" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFC5FFD93FBC63CDFBC07483" box="[1003,1038,956,979]" name="We" pageId="80" pageNumber="421" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFC5FFD93C8463ADFEA874A4" box="[211,358,988,1012]" count="57" pageId="80" pageNumber="421" type="generic">57 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFC5FFD93D2F63ADFE1574A4" box="[376,475,988,1012]" count="23" pageId="80" pageNumber="421" type="male">23 males</specimenCount>
and
<specimenCount id="9D79FD02FFC5FFD93E4263ADFD4074A4" box="[533,654,988,1012]" count="34" pageId="80" pageNumber="421" type="female">34 females</specimenCount>
) from sites on the crest or eastern slope of the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFC5FFD93D9B638DFD5F7364" county="Gongshan County" location="Core Area" pageId="80" pageNumber="421" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFC5FFD93D9B638DFDCF7344" box="[460,513,1020,1044]" country="Myanmar" name="Shan" pageId="80" pageNumber="421">Shan</collectingRegion>
in
<collectingCounty id="62A14E07FFC5FFD93E71638DFD277344" box="[550,745,1020,1044]" pageId="80" pageNumber="421">Gongshan County</collectingCounty>
(see
<typeStatus id="54C48829FFC5FFD93F75638DFC997344" box="[802,855,1020,1044]" pageId="80" pageNumber="421">Type</typeStatus>
material above for exact collection data). All of these sites are in
<location id="8EA06050FFC5FFD93E5B646DFDB47364" LSID="urn:lsid:plazi:treatment:03D6879DFFD8FFD93C2C64ADFB5C7324:8EA06050FFC5FFD93E5B646DFDB47364" box="[524,634,1052,1076]" county="Gongshan County" name="Core Area" pageId="80" pageNumber="421" stateProvince="Shan">Core Area</location>
2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFC5FFD93CFC644DFB5C7324" blockId="80.[123,1174,219,1140]" pageId="80" pageNumber="421">
<emphasis id="B90BEA99FFC5FFD93CFC644DFD947302" bold="true" box="[171,602,1084,1108]" pageId="80" pageNumber="421">
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<smallCapsWord id="8D26A057FFC5FFD93D91644EFD947302" baselines="1101" box="[454,602,1087,1106]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="80" pageNumber="421">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Gongshan County in the northern part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFC5FFD93FD1642DFB8E7324" box="[902,1088,1116,1140]" country="China" name="Yunnan" pageId="80" pageNumber="421">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFC5FFD9381C642DFB407324" box="[1099,1166,1116,1140]" name="China" pageId="80" pageNumber="421">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,349 @@
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<mods:title id="EB068AB7FAC1FAE43D86B5F3F62C2A54">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="30BB27F8ECFE981BC6B4B37617BFDFA2">Deuve, Thierry</mods:namePart>
<mods:affiliation id="9549DB5A661B98C9F36573AEF0A6DEBB">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="CDC0DEAFB53EFB30411F07B75E1ADEEB">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="10DAF552503634D27B92C62E68803E82">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="4F33AFBF786C1FE2E9E2B07505A9D665" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="D5F15433D8A30E2E02B02B1651D35DA4" type="personal">
<mods:role id="14AF1B8B8D6E0FF5BAD7E8E8BA5E492C">
<mods:roleTerm id="434219FF2EF87FA676EE4EE391DF28E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B69A2D87620FFF59598D7142AC747D6D">Liang, Hongbin</mods:namePart>
<mods:affiliation id="D92BA4F25A266DED95D061692640B1BB">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:part id="ED8BB980F5FFBAF1577BB7B19EB78C48">
<mods:date id="763E6AE3A9D6EACF157AABB734EFF749">2016</mods:date>
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<emphasis id="B90BEA99FFDDFFC13C2C60AAFD2F77A3" bold="true" box="[123,737,219,243]" pageId="72" pageNumber="413">
<taxonomicName id="4C7F4D08FFDDFFC13C2C60AAFDB077A3" ID-CoL="8MLL2" authority="Deuve and Liang" authorityName="Deuve and Liang" authorityYear="2016" box="[123,638,219,243]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDDFFC13C2C60AAFE7C77A3" bold="true" box="[123,434,219,243]" italics="true" pageId="72" pageNumber="413">Trechepaphiopsis uniporosa</emphasis>
Deuve and Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFDDFFC13EDC60AAFD2F77A3" box="[651,737,219,243]" pageId="72" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC13C2C608AFEB37641" box="[123,381,251,273]" pageId="72" pageNumber="413" type="description">
<paragraph id="8BC0368BFFDDFFC13C2C608AFEB37641" blockId="72.[123,737,219,273]" box="[123,381,251,273]" pageId="72" pageNumber="413">
(
<figureCitation id="13442A0EFFDDFFC13CD5608AFF157641" box="[130,219,251,273]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="72" pageNumber="413">Figs. 16f</figureCitation>
,
<figureCitation id="13442A0EFFDDFFC13CB2608AFF317640" box="[229,255,251,272]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="72" pageNumber="413">28</figureCitation>
,
<figureCitation id="13442A0EFFDDFFC13D5D608AFEE07640" box="[266,302,251,272]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="72" pageNumber="413">37a</figureCitation>
,
<figureCitation id="13442A0EFFDDFFC13D6E608AFEB97641" box="[313,375,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="72" pageNumber="413">4648</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC13CFC6155FC46758C" pageId="72" pageNumber="413" type="materials_examined">
<paragraph id="8BC0368BFFDDFFC13CFC6155FBD0756C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<emphasis id="B90BEA99FFDDFFC13CFC6155FE97766A" bold="true" box="[171,345,292,316]" pageId="72" pageNumber="413">
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<smallCapsWord id="8D26A057FFDDFFC13CBB6156FE97766A" baselines="309" box="[236,345,295,314]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="72" pageNumber="413">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FFDDFFC13DD56155FF4A76EC" collectingDate="1998-10-15" collectingDateMax="1998-10-18" collectingDateMin="1998-10-15" collectorName="C. E. Griswold &amp; C. Ferraris &amp; C. - L. Long" country="China" location="Nujiang Prefecture" municipality="Gaoligongshan Mountains" pageId="72" pageNumber="413" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFDDFFC13DD56155FE26766C" box="[386,488,292,316]" pageId="72" pageNumber="413" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1001923”/ “
<collectingCountry id="F368761BFFDDFFC13FAB6155FB9B766C" box="[1020,1109,292,316]" name="China" pageId="72" pageNumber="413">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDDFFC138366155FEC4760C" country="China" name="Yunnan" pageId="72" pageNumber="413">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFDDFFC13D426135FDFD760C" box="[277,563,324,348]" pageId="72" pageNumber="413">Gaoligongshan Mountains</collectingMunicipality>
,
<location id="8EA06050FFDDFFC13E696135FCC5760C" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDDFFC13E696135FCC5760C" box="[574,779,324,348]" country="China" municipality="Gaoligongshan Mountains" name="Nujiang Prefecture" pageId="72" pageNumber="413" stateProvince="Yunnan">Nujiang Prefecture</location>
, 9 [actually 9.3] km
<locationDeviation id="9A722FA9FFDDFFC13FBD6135FB5C760C" box="[1002,1170,324,348]" location="Pianma" pageId="72" pageNumber="413">ESE of Pianma</locationDeviation>
, 25°59.6N/ 98°37.6E.”/ “2450 [actually 2460-2470] m,
<date id="FFC1104BFFDDFFC13E816115FC61762C" box="[726,943,356,380]" pageId="72" pageNumber="413" value="1998-10-15" valueMax="1998-10-18" valueMin="1998-10-15">
<collectingDate id="EF85E9A3FFDDFFC13E816115FC61762C" box="[726,943,356,380]" pageId="72" pageNumber="413" value="1998-10-15" valueMax="1998-10-18" valueMin="1998-10-15">15-18 October 1998</collectingDate>
</date>
, Stop #98-118D D.H.
<taxonomicName id="4C7F4D08FFDDFFC13C2C61F5FF3976CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,247,388,412]" pageId="72" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFDDFFC13D5E61F5FE6376CC" box="[265,429,388,412]" pageId="72" pageNumber="413">C.E. Griswold</collectorName>
,
<collectorName id="268A535DFFDDFFC13DE861F5FDF776CC" box="[447,569,388,412]" pageId="72" pageNumber="413">C. Ferraris</collectorName>
&amp;
<collectorName id="268A535DFFDDFFC13E3C61F5FD2776CC" box="[619,745,388,412]" pageId="72" pageNumber="413">C.-L. Long</collectorName>
collectors”/ “IMAGE” [green label]/ “
</materialsCitation>
<typeStatus id="54C48829FFDDFFC13CD361D5FEDD76EC" box="[132,275,420,444]" pageId="72" pageNumber="413" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDDFFC13D4D61D5FD3776EC" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[282,761,420,444]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDDFFC13D4D61D5FD8676EC" box="[282,584,420,444]" italics="true" pageId="72" pageNumber="413">Trechepaphiopsis uniporosa</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFDDFFC13F5361D5FC9676EB" box="[772,856,420,443]" pageId="72" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label].
<materialsCitation id="3B173CD6FFDDFFC13C2C61B5FBD7756C" collectionCode="CAS, IOZ, MNHN" country="China" pageId="72" pageNumber="413" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFDDFFC13C2C61B5FF2D768C" box="[123,227,452,476]" pageId="72" pageNumber="413" type="paratype">Paratypes</typeStatus>
(a total of 5):
<specimenCount id="9D79FD02FFDDFFC13D2861B5FE1C768C" box="[383,466,452,476]" count="3" pageId="72" pageNumber="413" type="male">3 males</specimenCount>
and
<specimenCount id="9D79FD02FFDDFFC13E5061B5FDA1768C" box="[519,623,452,476]" count="2" pageId="72" pageNumber="413" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFDDFFC13ECB61B5FD1C768C" box="[668,722,452,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="72" pageNumber="413" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFDDFFC13E8A61B5FCC2768C" box="[733,780,452,476]" pageId="72" pageNumber="413">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFDDFFC13F4F61B5FCA5768C" box="[792,875,452,476]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="72" pageNumber="413">MNHN</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFDDFFC1386661B5FB5F768C" box="[1073,1169,452,476]" pageId="72" pageNumber="413" type="holotype">holotype</typeStatus>
, except first label “CASENT 1001920” to “CASENT 1001922” and “CASENT 1001924” to “CASENT 1001925”, respectively. All
<typeStatus id="54C48829FFDDFFC13E606274FD53754D" box="[567,669,517,541]" pageId="72" pageNumber="413" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFDDFFC138596275FB5B754C" box="[1038,1173,516,540]" pageId="72" pageNumber="413" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDDFFC13C2C6255FD94756C" authority="Deuve &amp; Liang" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,602,548,572]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDDFFC13C2C6255FE66756C" box="[123,424,548,572]" italics="true" pageId="72" pageNumber="413">Trechepaphiopsis uniporosa</emphasis>
Deuve &amp; Liang
</taxonomicName>
,
<taxonomicNameLabel id="A23857E2FFDDFFC13E326254FD76756C" box="[613,696,549,572]" pageId="72" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC6235FD29752C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<materialsCitation id="3B173CD6FFDDFFC13CFC6235FD2D752C" country="China" county="Lushui County" elevation="2465" location="Pianma Township" municipality="Gaoligong Shan" pageId="72" pageNumber="413" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFDDFFC13CFC6235FE97750B" bold="true" box="[171,345,580,604]" pageId="72" pageNumber="413">
<smallCapsWord id="8D26A057FFDDFFC13CFC6235FF2A750B" baselines="598,598" box="[171,228,580,604]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="72" pageNumber="413">
<typeStatus id="54C48829FFDDFFC13CFC6235FF2A750B" box="[171,228,580,604]" pageId="72" pageNumber="413">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFDDFFC13CB96239FE97750B" baselines="598" box="[238,345,584,603]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="72" pageNumber="413">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFDDFFC13DD46235FE0B750C" box="[387,453,580,604]" name="China" pageId="72" pageNumber="413">China</collectingCountry>
,
<collectingRegion id="49BBF869FFDDFFC13D856234FDE8750C" box="[466,550,581,604]" country="China" name="Yunnan" pageId="72" pageNumber="413">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFDDFFC13E636235FD28750C" box="[564,742,580,604]" pageId="72" pageNumber="413">Gaoligong Shan</collectingMunicipality>
,
<collectingCounty id="62A14E07FFDDFFC13EA46235FC58750C" box="[755,918,580,604]" pageId="72" pageNumber="413">Lushui County</collectingCounty>
,
<location id="8EA06050FFDDFFC13FF46235FBA9750C" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDDFFC13FF46235FBA9750C" box="[931,1127,580,604]" country="China" county="Lushui County" municipality="Gaoligong Shan" name="Pianma Township" pageId="72" pageNumber="413" stateProvince="Yunnan">Pianma Township</location>
,
<locationDeviation id="9A722FA9FFDDFFC138236235FE85752C" location="Pianma" pageId="72" pageNumber="413">
<quantity id="4C879B6EFFDDFFC138236235FF53752C" metricMagnitude="3" metricUnit="m" metricValue="9.3" pageId="72" pageNumber="413" unit="km" value="9.3">9.3 km</quantity>
ESE of Pianma
</locationDeviation>
, 25.99363°/ 98.66651°,
<quantity id="4C879B6EFFDDFFC13E076215FD2D752C" box="[592,739,612,636]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="72" pageNumber="413" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">
<elevation id="0052D1B8FFDDFFC13E076215FD2D752C" box="[592,739,612,636]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="72" pageNumber="413" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">2460-2470 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC62F5FC46758C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<emphasis id="B90BEA99FFDDFFC13CFC62F5FE2075CB" bold="true" box="[171,494,644,668]" pageId="72" pageNumber="413">
<smallCapsWord id="8D26A057FFDDFFC13CFC62F5FEE075CB" baselines="662,662" box="[171,302,644,668]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="72" pageNumber="413">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFDDFFC13D6262F9FE9F75CB" baselines="662" box="[309,337,648,667]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="72" pageNumber="413">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFDDFFC13D0062F9FE6475CB" baselines="662" box="[343,426,648,667]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="72" pageNumber="413">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFDDFFC13DE662F9FE2075CB" baselines="662" box="[433,494,648,667]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="72" pageNumber="413">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFDDFFC13EA662F4FC9275CC" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[753,860,645,668]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="413" phylum="Arthropoda" rank="species" species="uniporosa">
<emphasis id="B90BEA99FFDDFFC13EA662F4FC9275CC" box="[753,860,645,668]" italics="true" pageId="72" pageNumber="413">uniporosa</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B90BEA99FFDDFFC13D6662D4FEAD75EC" box="[305,355,677,700]" italics="true" pageId="72" pageNumber="413">unus</emphasis>
, meaning one, and
<emphasis id="B90BEA99FFDDFFC13E6062D4FDBA75EC" box="[567,628,677,700]" italics="true" pageId="72" pageNumber="413">porus</emphasis>
, meaning pore or hole. The name refers to the single discal setiferous pore found on the elytra of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC13CFC6295FF23734C" pageId="72" pageNumber="413" type="diagnosis">
<paragraph id="8BC0368BFFDDFFC13CFC6295FF23734C" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<smallCapsWord id="8D26A057FFDDFFC13CFC6295FEEE75AB" baselines="758,758" box="[171,288,740,764]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="72" pageNumber="413">
<emphasis id="B90BEA99FFDDFFC13CFC6295FEEE75AB" bold="true" box="[171,288,740,764]" pageId="72" pageNumber="413">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFDDFFC13E106295FD6D75AC" box="[583,675,740,764]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="72" pageNumber="413">Fig. 28a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
<quantity id="4C879B6EFFDDFFC138006375FF1E746D" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="72" pageNumber="413" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
), apterous; body color light brown, slightly iridescent; eyes small; tempora distinctly convex, sparsely pubescent; pronotum transverse (ratio PW/PL = 1.34), with basal angles obtuse and rounded; elytra convex, with stria 1 deeply impressed and finely punctate, other striae more faintly impressed and lateral striae effaced; recurrent stria terminated anteriorly in presumed location of interval 6; only one discal setiferous pore present, inserted at anterior one-third next to stria 3; preapical seta present, inserted next to stria 3; median lobe of male aedeagus (
<figureCitation id="13442A0EFFDDFFC13F9563B5FBD0748C" box="[962,1054,964,988]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="72" pageNumber="413">Fig. 28b</figureCitation>
) with apex slender but bluntly rounded and slightly recurved dorsally, endophallus with a faintly sclerotized scaly area.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDDFFC33CFC6454FEE17304" lastPageId="74" lastPageNumber="415" pageId="72" pageNumber="413" type="description">
<paragraph id="8BC0368BFFDDFFC13CFC6454FCDA730D" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<smallCapsWord id="8D26A057FFDDFFC13CFC6454FEF4736B" baselines="1079,1078" box="[171,314,1061,1085]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="72" pageNumber="413">
<emphasis id="B90BEA99FFDDFFC13CFC6454FEF4736B" bold="true" box="[171,314,1061,1085]" pageId="72" pageNumber="413">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size medium BL =
<quantity id="4C879B6EFFDDFFC13E626454FD01736C" box="[565,719,1061,1085]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="72" pageNumber="413" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
. Body color light brown, appendages concolorous, except palpi paler, dorsum shiny, slightly iridescent.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC6414FBE972CD" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">
<materialsCitation id="3B173CD6FFDDFFC13CFC6414FBED72CD" location="Antennae" municipality="Submentum" pageId="72" pageNumber="413" specimenCount="16" stateProvince="Chin">
Head. Moderate in size, short; eyes small, not projected, their diameter about equal to length of tempora but their convexity less than that of tempora, the latter markedly convex, inflated and sparsely pubescent. Frontal furrows thin and linear, deeply impressed, arcuate, continuous posteriorly to hind margins of tempora. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave or emarginate. Mandibles short and slender, right mandible as in
<figureCitation id="13442A0EFFDDFFC13F756574FCB2724D" box="[802,892,1285,1309]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="72" pageNumber="413">
Fig.
<specimenCount id="9D79FD02FFDDFFC13F026574FCB2724D" box="[853,892,1285,1309]" count="16" pageId="72" pageNumber="413" type="generic">16f</specimenCount>
</figureCitation>
.
<collectingRegion id="49BBF869FFDDFFC13FDF6574FC2C724C" box="[904,994,1285,1308]" country="Myanmar" name="Chin" pageId="72" pageNumber="413">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFDDFFC13CA66554FE85726C" box="[241,331,1317,1340]" country="Myanmar" name="Chin" pageId="72" pageNumber="413">Mentum</collectingRegion>
with medial tooth apically truncate, about one-half the length of the lateral lobes.
<collectingMunicipality id="6BA4ACF1FFDDFFC13C956534FE8D720D" box="[194,323,1349,1373]" pageId="72" pageNumber="413">Submentum</collectingMunicipality>
with six setae anteriorly.
<location id="8EA06050FFDDFFC13E0D6534FD0E720D" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDDFFC13E0D6534FD0E720D" box="[602,704,1349,1373]" municipality="Submentum" name="Antennae" pageId="72" pageNumber="413" stateProvince="Chin">Antennae</location>
rather short, not quite extended posteriorly to basal one-fourth of the elytra, only 2.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3 about equal in length, antennomere 4 slightly shorter
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDDFFC13CFC65D7FDFA71ED" blockId="72.[123,1174,292,1725]" pageId="72" pageNumber="413">Pronotum. Distinctly transverse (ratio PW/PL = 1.34), with greatest width near anterior one-third, only slightly narrowed posteriorly; lateral margins rounded, slightly more so anteriorly, slightly straighten just anterior to basal angles, the latter obtuse and rounded. Disc smooth and convex, median longitudinal impression finely impressed, but continuous between anterior and posterior margins; basal foveae shallow, faintly impressed; median basal area reduced, faintly delimit- ed, slightly punctate in some specimens. Basal margin nearly rectilinear. Lateral pronotal borders moderately slender, regular, narrowly and regularly reflexed, lateral grooves narrow but distinctly impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
<caption id="DF006603FFDCFFC03CFC6648FD02712F" ID-DOI="http://doi.org/10.5281/zenodo.13155336" ID-Zenodo-Dep="13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="73" pageNumber="414" startId="73.[171,183,1593,1612]" targetBox="[120,1176,216,1570]" targetPageId="73" targetType="figure">
<paragraph id="8BC0368BFFDCFFC03CFC6648FD02712F" blockId="73.[123,1174,1593,1664]" pageId="73" pageNumber="414">
<smallCapsWord id="8D26A057FFDCFFC03CFC6648FF26711B" baselines="1607,1607" box="[171,232,1593,1612]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="73" pageNumber="414">FIGURE</smallCapsWord>
28.
<taxonomicName id="4C7F4D08FFDCFFC03D5B6648FE33711C" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[268,509,1593,1612]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="414" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFDCFFC03D5B6648FE33711C" box="[268,509,1593,1612]" italics="true" pageId="73" pageNumber="414">Trechepaphiopsis uniporosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFDCFFC03E556648FD89711C" box="[514,583,1593,1612]" pageId="73" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFDCFFC0387C6622FB587135" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1067,1174,1619,1637]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="414" phylum="Arthropoda" rank="species" species="uniporosa">
<emphasis id="B90BEA99FFDCFFC0387C6622FB587135" box="[1067,1174,1619,1637]" italics="true" pageId="73" pageNumber="414">T. uniporosa</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFDFFFC33CFC60AAFC287683" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">Elytra. Ovoid, only slightly more narrowed anteriorly than posteriorly, humeri distinct but rounded. Disc convex, striae faintly impressed and finely punctate, the medial three or four striae distinctly impressed, the more lateral striae more or less effaced, but perceptible if only by the presence of fine punctures seen as rows of brown dots visible through the integument. Parascutellar striole present. Recurrent stria abruptly terminated anteriorly in presumed location of interval 6. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Only one discal setiferous pore present, inserted slightly anterior to middle next to stria 3. Preapical seta present on interval 3 near stria 3, closer to elytra apical margin that to sutural margin.</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC61ADFE9E7543" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC626AFB857503" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC622AFF7F7584" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFDFFFC33DA9622AFD907523" box="[510,606,603,627]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="74" pageNumber="415">Fig. 28b</figureCitation>
) with sagittal aileron very small, shaft moderately broad basally, expanded near mid-length, then gradually narrowed apically to a thin, bluntly round- ed apex, the latter slightly recurved dorsally; endophallus with a small scaly sclerotized area (
<figureCitation id="13442A0EFFDFFFC3383D62EAFF6B7584" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="74" pageNumber="415">Fig. 28c</figureCitation>
).
</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC62ADFE7A7424" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">
<emphasis id="B90BEA99FFDFFFC33CFC62ADFE6A75A2" bold="true" box="[171,420,732,756]" pageId="74" pageNumber="415">
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<smallCapsWord id="8D26A057FFDFFFC33D4762AEFE6A75A2" baselines="749" box="[272,420,735,754]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="74" pageNumber="415">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in mixed broadleaf evergreen/ deciduous forest at elevations ranging from
<quantity id="4C879B6EFFDFFFC33ECE628DFC8F7443" box="[665,833,764,788]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="74" pageNumber="415" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">2460 to 2470 m</quantity>
and collected by sifting leaf litter on the forest floor (
<figureCitation id="13442A0EFFDFFFC33D2C636DFE147464" box="[379,474,796,820]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="74" pageNumber="415">Fig. 37a</figureCitation>
). Specimens of
<taxonomicName id="4C7F4D08FFDFFFC33EDA636DFC677463" baseAuthorityName="Deuve" baseAuthorityYear="2004" box="[653,937,796,820]" class="Insecta" family="Carabidae" genus="Trechepaphiosis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="415" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFDFFFC33EDA636DFC677463" box="[653,937,796,820]" italics="true" pageId="74" pageNumber="415">Trechepaphiosis unisetosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFDFFFC33FB4636DFF297404" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="415" phylum="Arthropoda" rank="species" species="gaoligong" status="sp. nov.">
<emphasis id="B90BEA99FFDFFFC33FB4636DFF297404" italics="true" pageId="74" pageNumber="415">Trechepaphiama gaoligong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFDFFFC33CB8634DFEF07403" box="[239,318,828,851]" pageId="74" pageNumber="415" rank="species">sp. nov</taxonomicNameLabel>
were collected in the same sifted leaf litter samples and are therefore syntopic with
<taxonomicName id="4C7F4D08FFDFFFC33CE6632DFEF97423" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[177,311,860,883]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="415" phylum="Arthropoda" rank="species" species="uniporosa">
<emphasis id="B90BEA99FFDFFFC33CE6632DFEF97423" box="[177,311,860,883]" italics="true" pageId="74" pageNumber="415">T. uniporosa</emphasis>
</taxonomicName>
at this site.
</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC630AFD9474A4" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">
<emphasis id="B90BEA99FFDFFFC33CFC630AFDB174C2" bold="true" box="[171,639,891,915]" pageId="74" pageNumber="415">
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<smallCapsWord id="8D26A057FFDFFFC33D08630EFE3D74C2" baselines="909" box="[351,499,895,914]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="74" pageNumber="415">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFDFFFC33DAA630EFD8274C2" baselines="909" box="[509,588,895,914]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="74" pageNumber="415">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFDFFFC33E02630EFDB174C2" baselines="909" box="[597,639,895,914]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="74" pageNumber="415">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FFDFFFC33EDD630AFCC074C2" baselines="909,909" box="[650,782,891,915]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="74" pageNumber="415">
<emphasis id="B90BEA99FFDFFFC33EDD630AFCC074C2" bold="true" box="[650,782,891,915]" pageId="74" pageNumber="415">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFDFFFC33F4F630AFE257484" location="We" pageId="74" pageNumber="415" specimenCount="12" specimenCount-female="2" specimenCount-male="4" stateProvince="Shan">
<smallCapsWord id="8D26A057FFDFFFC33F4F630AFC9D74C2" baselines="909,909" box="[792,851,891,915]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="74" pageNumber="415">
<emphasis id="B90BEA99FFDFFFC33F4F630AFC9D74C2" bold="true" box="[792,851,891,915]" pageId="74" pageNumber="415">
<collectingRegion id="49BBF869FFDFFFC33F4F630AFC9D74C2" box="[792,851,891,915]" country="Myanmar" name="Shan" pageId="74" pageNumber="415">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFDFFFC33F28630DFC1174C4" box="[895,991,892,916]" captionStart="FIGURE" captionStartId="73.[171,183,1593,1612]" captionTargetBox="[120,1176,216,1570]" captionTargetId="figure-15@73.[120,1176,216,1570]" captionTargetPageId="73" captionText="FIGURE 28. Trechepaphiopsis uniporosa sp. nov.; a. Dorsal habitus (CASENT1001923). b-c. Median lobe of aedeagus of male (CASENT1001923); b. left lateral aspect; c. ventral aspect. d..Map of locality records (red circles) for T. uniporosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155336" httpUri="https://zenodo.org/record/13155336/files/figure.png" pageId="74" pageNumber="415">Fig. 28d</figureCitation>
.
<location id="8EA06050FFDFFFC33FBC630DFBC074C3" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDFFFC33FBC630DFBC074C3" box="[1003,1038,892,915]" name="We" pageId="74" pageNumber="415" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFDFFFC33C8463EDFE9674E4" box="[211,344,924,948]" count="6" pageId="74" pageNumber="415" type="generic">6 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFDFFFC33D3E63EDFE7174E4" box="[361,447,924,948]" count="4" pageId="74" pageNumber="415" type="male">4 males</specimenCount>
and
<specimenCount id="9D79FD02FFDFFFC33DAF63EDFDAD74E4" box="[504,611,924,948]" count="2" pageId="74" pageNumber="415" type="female">2 females</specimenCount>
) from the
<typeStatus id="54C48829FFDFFFC33E8D63EDFCC674E4" box="[730,776,924,948]" pageId="74" pageNumber="415">type</typeStatus>
locality on the western slope of the southcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFDFFFC33DA563CDFD9874A4" county="Lushui County" location="Core Area" pageId="74" pageNumber="415" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFDFFFC33DA563CDFDE97484" box="[498,551,956,980]" country="Myanmar" name="Shan" pageId="74" pageNumber="415">Shan</collectingRegion>
in
<collectingCounty id="62A14E07FFDFFFC33E1C63CDFD257484" box="[587,747,956,980]" pageId="74" pageNumber="415">Lushui County</collectingCounty>
(see
<typeStatus id="54C48829FFDFFFC33F7363CDFC977484" box="[804,857,956,980]" pageId="74" pageNumber="415">Type</typeStatus>
material above for exact collection data). This locality is in
<location id="8EA06050FFDFFFC33D8663ADFDF074A4" LSID="urn:lsid:plazi:treatment:03D6879DFFDDFFC33C2C60AAFEE17304:8EA06050FFDFFFC33D8663ADFDF074A4" box="[465,574,988,1012]" county="Lushui County" name="Core Area" pageId="74" pageNumber="415" stateProvince="Shan">Core Area</location>
4
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDFFFC33CFC638DFEE17304" blockId="74.[123,1174,219,1108]" pageId="74" pageNumber="415">
<emphasis id="B90BEA99FFDFFFC33CFC638DFD927342" bold="true" box="[171,604,1020,1044]" pageId="74" pageNumber="415">
<smallCapsWord id="8D26A057FFDFFFC33CFC638DFEDC7342" baselines="1038,1037" box="[171,274,1020,1044]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Overall" pageId="74" pageNumber="415">OVERALL</smallCapsWord>
<smallCapsWord id="8D26A057FFDFFFC33D4D638EFE0F7342" baselines="1037" box="[282,449,1023,1042]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="74" pageNumber="415">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFDFFFC33D9F638EFD927342" baselines="1037" box="[456,604,1023,1042]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="74" pageNumber="415">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from western Lushui County in the southcentral part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFDFFFC33813646DFF137304" country="China" name="Yunnan" pageId="74" pageNumber="415">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFDFFFC33CB0644DFEE47304" box="[231,298,1084,1108]" name="China" pageId="74" pageNumber="415">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,542 @@
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<mods:title id="813E1787905C658C72937DFC07006FCA">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:namePart id="D9DEB5018866F1191D836B46F7D9BE53">Deuve, Thierry</mods:namePart>
<mods:affiliation id="7DAE4CEC12CFC43F06F4AFDE87FA47B1">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="B79E1B069E12DBECF4DD1201FFC6D0C5">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="2977186469B8D507B8CF1ED77414C557">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="4B4F14CFC03CECFEBC8B0EB519235169" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:affiliation id="0FA761171127DFAFCBA75AB16BA98994">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<subSubSection id="C3656500FFDFFFC33C2C640DFCFF73C4" box="[123,817,1148,1172]" pageId="74" pageNumber="415" type="nomenclature">
<paragraph id="8BC0368BFFDFFFC33C2C640DFCFF73C4" blockId="74.[123,817,1148,1202]" box="[123,817,1148,1172]" pageId="74" pageNumber="415">
<heading id="D08881E7FFDFFFC33C2C640DFCFF73C4" box="[123,817,1148,1172]" fontSize="10" level="2" pageId="74" pageNumber="415" reason="4">
<emphasis id="B90BEA99FFDFFFC33C2C640DFCFF73C4" bold="true" box="[123,817,1148,1172]" pageId="74" pageNumber="415">
<taxonomicName id="4C7F4D08FFDFFFC33C2C640DFDC173C4" ID-CoL="8MG35" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,527,1148,1172]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="415" phylum="Arthropoda" rank="species" species="unisetulosa">
<emphasis id="B90BEA99FFDFFFC33C2C640DFE0F73C4" bold="true" box="[123,449,1148,1172]" italics="true" pageId="74" pageNumber="415">Trechepaphiopsis unisetulosa</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FFDFFFC33E8C640DFCFF73C4" box="[731,817,1148,1172]" pageId="74" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDFFFC33C2C64EDFE9C73E2" box="[123,338,1180,1202]" pageId="74" pageNumber="415" type="description">
<paragraph id="8BC0368BFFDFFFC33C2C64EDFE9C73E2" blockId="74.[123,817,1148,1202]" box="[123,338,1180,1202]" pageId="74" pageNumber="415">
(
<figureCitation id="13442A0EFFDFFFC33CD564EDFF1D73E1" box="[130,211,1180,1202]" captionStart="FIGURE" captionStartId="76.[171,183,1581,1600]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-11@76.[120,1176,216,1564]" captionTargetPageId="76" captionText="FIGURE 29. Trechepaphiopsis unisetulosa sp. nov.; a. Dorsal habitus (CASENT1018407). b. Median lobe of aedeagus of male (CASENT1001903), left lateral aspect. c. Map of locality records (red circles) for T. unisetulosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155338" httpUri="https://zenodo.org/record/13155338/files/figure.png" pageId="74" pageNumber="415">Figs. 29</figureCitation>
,
<figureCitation id="13442A0EFFDFFFC33C8A64EDFECD73E2" box="[221,259,1180,1202]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="74" pageNumber="415">40b</figureCitation>
,
<figureCitation id="13442A0EFFDFFFC33D5964EDFE8573E2" box="[270,331,1180,1202]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="74" pageNumber="415">4648</figureCitation>
)
</paragraph>
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.—
<materialsCitation id="3B173CD6FFDFFFC33DD564B4FED1726D" collectingDate="2005-08-18" collectorName="P. Paquin" country="China" county="Fugong County" elevation="2357" latitude="27.1665" location="Lishadi Township" longLatPrecision="1" longitude="98.77936" pageId="74" pageNumber="415" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FFDFFFC33DD564B4FE26738D" box="[386,488,1221,1245]" pageId="74" pageNumber="415" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1018407”/ “
<collectingCountry id="F368761BFFDFFFC33FAB64B4FB9B738D" box="[1020,1109,1221,1245]" name="China" pageId="74" pageNumber="415">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDFFFC3383664B4FED373AD" country="China" name="Yunnan" pageId="74" pageNumber="415">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFDFFFC33D616494FE3C73AD" box="[310,498,1253,1277]" pageId="74" pageNumber="415">Fugong County</collectingCounty>
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<quantity id="4C879B6EFFDFFFC33EAC6494FC9B73AD" box="[763,853,1253,1277]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="74" pageNumber="415" unit="km" value="0.2">0.2 km</quantity>
W of Shibali
</locationDeviation>
,
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,
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/
<geoCoordinate id="EE4B504CFFDFFFC33D526574FE5C724D" box="[261,402,1285,1309]" degrees="098.77936" direction="east" orientation="longitude" pageId="74" pageNumber="415" precision="1" value="98.77936">E098.77936°</geoCoordinate>
”/ “
<date id="FFC1104BFFDFFFC33DE16574FDAD724D" box="[438,611,1285,1309]" pageId="74" pageNumber="415" value="2005-08-18">
<collectingDate id="EF85E9A3FFDFFFC33DE16574FDAD724D" box="[438,611,1285,1309]" pageId="74" pageNumber="415" value="2005-08-18">18 August 2005</collectingDate>
</date>
, in leaf litter, Stop# PP-4405,
<collectorName id="268A535DFFDFFFC33FFF6574FBC0724D" box="[936,1038,1285,1309]" pageId="74" pageNumber="415">P. Paquin</collectorName>
”/ “IMAGE” [green label]/ “
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<typeStatus id="54C48829FFDFFFC33D486554FE60726D" box="[287,430,1317,1341]" pageId="74" pageNumber="415" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDFFFC33DEF6554FC8B726C" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[440,837,1317,1341]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="415" phylum="Arthropoda" rank="species" species="unisetulosa">
<emphasis id="B90BEA99FFDFFFC33DEF6554FD3B726D" box="[440,757,1317,1341]" italics="true" pageId="74" pageNumber="415">Trechepaphiopsis unisetulosa</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
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designated 2016” [red label].
<materialsCitation id="3B173CD6FFDFFFC33DDE6534FDF3722D" collectionCode="IOZ" country="China" pageId="74" pageNumber="415" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FFDFFFC33DDE6534FE3F720D" box="[393,497,1349,1373]" pageId="74" pageNumber="415" type="paratype">Paratypes</typeStatus>
(a total of 26):
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(in
<collectionCode id="ED6EAE4EFFDFFFC33F626534FCAD720D" box="[821,867,1349,1373]" pageId="74" pageNumber="415">IOZ</collectionCode>
) labeled: same as
<typeStatus id="54C48829FFDFFFC338666534FB5F720D" box="[1073,1169,1349,1373]" pageId="74" pageNumber="415" type="holotype">holotype</typeStatus>
, except first label “CASENT 1018408”
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;
<materialsCitation id="3B173CD6FFDFFFC33E076514FF1F72AD" collectingDate="2004-05-08" collectingDateMax="2004-05-11" collectingDateMin="2004-05-08" collectionCode="CAS" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="2475" latitude="27.16337" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.78208" municipality="Lishadi Township" pageId="74" pageNumber="415" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
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(in
<collectionCode id="ED6EAE4EFFDFFFC33EA56514FCE9722D" box="[754,807,1381,1405]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="74" pageNumber="415" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1014246”/ “
<collectingCountry id="F368761BFFDFFFC33CD365F4FF1372CD" box="[132,221,1413,1437]" name="China" pageId="74" pageNumber="415">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDFFFC33CBE65F7FE6D72CD" box="[233,419,1413,1437]" country="China" name="Yunnan" pageId="74" pageNumber="415">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFDFFFC33DF965F7FD9872CD" box="[430,598,1413,1438]" pageId="74" pageNumber="415">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFDFFFC33E3665F4FCED72CD" box="[609,803,1413,1437]" pageId="74" pageNumber="415">Lishadi Township</collectingMunicipality>
,
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<quantity id="4C879B6EFFDFFFC33F7965F4FCB672CD" box="[814,888,1413,1437]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="74" pageNumber="415" unit="km" value="0.4">0.4 km</quantity>
SE of Shibali
</locationDeviation>
along
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,
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/
<geoCoordinate id="EE4B504CFFDFFFC33DB465D4FDBF72ED" box="[483,625,1445,1469]" degrees="098.78208" direction="east" orientation="longitude" pageId="74" pageNumber="415" precision="1" value="98.78208">E098.78208°</geoCoordinate>
,
<quantity id="4C879B6EFFDFFFC33ED665D4FD1472ED" box="[641,730,1445,1469]" metricMagnitude="3" metricUnit="m" metricValue="2.475" pageId="74" pageNumber="415" unit="m" value="2475.0">
<elevation id="0052D1B8FFDFFFC33ED665D4FD1472ED" box="[641,730,1445,1469]" metricMagnitude="3" metricUnit="m" metricValue="2.475" pageId="74" pageNumber="415" unit="m" value="2475.0">2475 m</elevation>
</quantity>
,
<date id="FFC1104BFFDFFFC33EBC65D4FC5572ED" box="[747,923,1445,1470]" pageId="74" pageNumber="415" value="2004-05-08" valueMax="2004-05-11" valueMin="2004-05-08">
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</date>
”/ “from mini-Winkler extraction of leaf litter siftate, Stop #CGY37,
<collectorName id="268A535DFFDFFFC33E3365B4FCC9728D" box="[612,775,1477,1501]" pageId="74" pageNumber="415">C. E. Griswold</collectorName>
, D. H.
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, &amp; Yan H.-M. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFDFFFC33CB76594FC0A716D" collectingDate="2004-05-03" collectionCode="CAS" collectorName="C. E. Griswold." country="China" county="Fugong County" elevation="2560" latitude="27.16786" location="Shibali Road" longLatPrecision="1" longitude="98.77741" municipality="Lishadi Township" pageId="74" pageNumber="415" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFDFFFC33CB76594FEE372AD" box="[224,301,1509,1533]" count="1" pageId="74" pageNumber="415" type="male">1 male</specimenCount>
(in
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) labeled: “CASENT 1019222”/ “
<collectingCountry id="F368761BFFDFFFC33F5A6594FCA872AD" box="[781,870,1509,1533]" name="China" pageId="74" pageNumber="415">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDFFFC33F216597FBFB72AD" box="[886,1077,1509,1533]" country="China" name="Yunnan" pageId="74" pageNumber="415">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFDFFFC338136597FF05714D" pageId="74" pageNumber="415">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFDFFFC33C826674FE56714D" box="[213,408,1541,1565]" pageId="74" pageNumber="415">Lishadi Township</collectingMunicipality>
,
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<location id="8EA06050FFDFFFC33DF56674FE20714D" LSID="urn:lsid:plazi:treatment:03D6879DFFDFFFC43C2C640DFB4F73E4:8EA06050FFDFFFC33DF56674FE20714D" box="[418,494,1541,1565]" country="China" county="Fugong County" latitude="27.16786" longLatPrecision="1" longitude="98.77741" municipality="Lishadi Township" name="Shibali" pageId="74" pageNumber="415" stateProvince="Yunnan">Shibali</location>
Road
</location>
at
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,
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/
<geoCoordinate id="EE4B504CFFDFFFC33F636674FC0F714D" box="[820,961,1541,1565]" degrees="098.77741" direction="east" orientation="longitude" pageId="74" pageNumber="415" precision="1" value="98.77741">E098.77741°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFFDFFFC33FA66674FB8B714D" box="[1009,1093,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="2.56" pageId="74" pageNumber="415" unit="m" value="2560.0">
<elevation id="0052D1B8FFDFFFC33FA66674FB8B714D" box="[1009,1093,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="2.56" pageId="74" pageNumber="415" unit="m" value="2560.0">2560 m</elevation>
</quantity>
,
<date id="FFC1104BFFDFFFC338076674FF7E716D" pageId="74" pageNumber="415" value="2004-05-03">
<collectingDate id="EF85E9A3FFDFFFC338076674FF7E716D" pageId="74" pageNumber="415" value="2004-05-03">3 May 2004</collectingDate>
</date>
Stop #DHK-2004-024, D.H.
<taxonomicName id="4C7F4D08FFDFFFC33DA06654FDBE716D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[503,624,1573,1597]" pageId="74" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFDFFFC33EC06654FC8A716D" box="[663,836,1573,1597]" pageId="74" pageNumber="415">C. E. Griswold.</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFDFFFC33F866654FDB471ED" collectingDate="2005-08-09" collectionCode="IOZ" collectorName="H. B. Liang &amp; D. Z. Dong &amp; P. Paquin" country="China" county="Fugong County" elevation="3100" latitude="27.118416" location="North Fork of Yamu He" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" pageId="74" pageNumber="415" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFDFFFC33F866654FBFE716D" box="[977,1072,1573,1597]" count="1" pageId="74" pageNumber="415" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFDFFFC338376654FB40716D" box="[1120,1166,1573,1597]" pageId="74" pageNumber="415">IOZ</collectionCode>
) labeled: “CASENT 1020952”/ “
<collectingCountry id="F368761BFFDFFFC33D986634FDE6710D" box="[463,552,1605,1629]" name="China" pageId="74" pageNumber="415">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDFFFC33E646637FD23710D" box="[563,749,1605,1629]" country="China" name="Yunnan" pageId="74" pageNumber="415">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFDFFFC33EA06637FC51710D" box="[759,927,1605,1630]" pageId="74" pageNumber="415">Fugong County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FFDFFFC33FFF6634FBA4710D" box="[936,1130,1605,1629]" pageId="74" pageNumber="415">Lishadi Township</collectingMunicipality>
,
<locationDeviation id="9A722FA9FFDFFFC338236634FD9C712D" location="Shibali on Shibali Road" pageId="74" pageNumber="415">
<quantity id="4C879B6EFFDFFFC338236634FF50712D" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="74" pageNumber="415" unit="km" value="8.5">8.5 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
, north bank of
<location id="8EA06050FFDFFFC33F176614FB5C712D" LSID="urn:lsid:plazi:treatment:03D6879DFFDFFFC43C2C640DFB4F73E4:8EA06050FFDFFFC33F176614FB5C712D" box="[832,1170,1637,1661]" country="China" county="Fugong County" latitude="27.118416" longLatPrecision="1" longitude="98.72026" municipality="Lishadi Township" name="North Fork of Yamu He" pageId="74" pageNumber="415" stateProvince="Yunnan">North Fork of Yamu He</location>
,
<geoCoordinate id="EE4B504CFFDFFFC33C2C66F4FEC471CD" box="[123,266,1669,1693]" degrees="27.118416" direction="north" orientation="latitude" pageId="74" pageNumber="415" precision="1" value="27.118416">N27.118416°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFDFFFC33D4566F4FE5071CD" box="[274,414,1669,1693]" degrees="098.72026" direction="east" orientation="longitude" pageId="74" pageNumber="415" precision="1" value="98.72026">E098.72026°</geoCoordinate>
”/ “
<quantity id="4C879B6EFFDFFFC33D9466F4FDD571CD" box="[451,539,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="74" pageNumber="415" unit="m" value="3100.0">
<elevation id="0052D1B8FFDFFFC33D9466F4FDD571CD" box="[451,539,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="74" pageNumber="415" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<date id="FFC1104BFFDFFFC33E7E66F4FD0271CD" box="[553,716,1669,1693]" pageId="74" pageNumber="415" value="2005-08-09">
<collectingDate id="EF85E9A3FFDFFFC33E7E66F4FD0271CD" box="[553,716,1669,1693]" pageId="74" pageNumber="415" value="2005-08-09">9 August 2005</collectingDate>
</date>
, Stop DHK-2005-067A D.H.
<taxonomicName id="4C7F4D08FFDFFFC3384266F4FB5F71CD" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1045,1169,1669,1693]" pageId="74" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFDFFFC33C2C66D7FF3A71ED" box="[123,244,1701,1725]" pageId="74" pageNumber="415">H.B. Liang</collectorName>
,
<collectorName id="268A535DFFDFFFC33CA866D7FEBA71EE" box="[255,372,1702,1726]" pageId="74" pageNumber="415">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFFDFFFC33DCE66D7FE3271ED" box="[409,508,1701,1725]" pageId="74" pageNumber="415">P. Paquin</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFDFFFC23ED366D4FF3C7605" collectingDate="2004-05-03" collectingDateMax="2004-05-11" collectingDateMin="2004-05-03" collectionCode="CAS" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="2563" lastPageId="75" lastPageNumber="416" latitude="27.11679" location="Lishadi Township" longLatPrecision="1" longitude="98.77655" pageId="74" pageNumber="415" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFDFFFC33ED366D4FD0571ED" box="[644,715,1701,1725]" count="1" pageId="74" pageNumber="415" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FFDFFFC33EA966D4FC9571ED" box="[766,859,1701,1725]" count="1" pageId="74" pageNumber="415" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFFDFFFC33FD266D4FC7471ED" box="[901,954,1701,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="74" pageNumber="415" type="Museum">CAS</collectionCode>
) labeled: “CASENT 1021953” and “CASENT 1021952”, res/ “
<collectingCountry id="F368761BFFDEFFC23E0260AAFD6077A3" box="[597,686,219,243]" name="China" pageId="75" pageNumber="416">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDEFFC23EEA60AAFCB277A3" box="[701,892,219,243]" country="China" name="Yunnan" pageId="75" pageNumber="416">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFDEFFC23FDD60AAFBF977A3" box="[906,1079,219,243]" pageId="75" pageNumber="416">Fugong County</collectingCounty>
,
<location id="8EA06050FFDEFFC2381260AAFF297644" LSID="urn:lsid:plazi:treatment:03D6879DFFDFFFC43C2C640DFB4F73E4:8EA06050FFDEFFC2381260AAFF297644" country="China" county="Fugong County" latitude="27.11679" longLatPrecision="1" longitude="98.77655" name="Lishadi Township" pageId="75" pageNumber="416" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFDEFFC23CA6608DFD477644" box="[241,649,252,276]" location="Shibali on Shibali Road" pageId="75" pageNumber="416">
<quantity id="4C879B6EFFDEFFC23CA6608DFEF57644" box="[241,315,252,276]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="416" unit="km" value="0.3">0.3 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFDEFFC23EC3608DFCED7644" box="[660,803,252,276]" degrees="27.116791" direction="north" orientation="latitude" pageId="75" pageNumber="416" precision="1" value="27.11679">N27.116791°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFDEFFC23F7C608DFC767644" box="[811,952,252,276]" degrees="098.77655" direction="east" orientation="longitude" pageId="75" pageNumber="416" precision="1" value="98.77655">E098.77655°</geoCoordinate>
”/ “
<quantity id="4C879B6EFFDEFFC23F8D608DFBE17643" box="[986,1071,252,276]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="75" pageNumber="416" unit="m" value="2563.0">
<elevation id="0052D1B8FFDEFFC23F8D608DFBE17643" box="[986,1071,252,276]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="75" pageNumber="416" unit="m" value="2563.0">2563 m</elevation>
</quantity>
,
<collectingMethod id="523E4E9CFFDEFFC2386D608DFF697664" pageId="75" pageNumber="416">in pitfall trap</collectingMethod>
,
<date id="FFC1104BFFDEFFC23CE6616CFE997665" box="[177,343,285,309]" pageId="75" pageNumber="416" value="2004-05-03" valueMax="2004-05-11" valueMin="2004-05-03">
<collectingDate id="EF85E9A3FFDEFFC23CE6616CFE997665" box="[177,343,285,309]" pageId="75" pageNumber="416" value="2004-05-03" valueMax="2004-05-11" valueMin="2004-05-03">3-11 May 2004</collectingDate>
</date>
, Stop #CGY21,
<collectorName id="268A535DFFDEFFC23E50616CFD647664" box="[519,682,284,309]" pageId="75" pageNumber="416">C. E. Griswold</collectorName>
, D. H.
<taxonomicName id="4C7F4D08FFDEFFC23EA2616DFCBF7664" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[757,881,284,308]" pageId="75" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
, Liang H.-B., &amp; Yan H.-M. collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FFDEFFC23D57614CFBAD76A9" collectingDate="2004-05-04" collectionCode="CAS, IOZ, MNHN" collectorName="C. E. Griswold" country="China" county="Fugong County" elevation="2563" latitude="27.116636" location="Lishadi Township" longLatPrecision="1" longitude="98.77667" pageId="75" pageNumber="416" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FFDEFFC23D57614CFEAD7605" box="[256,355,317,341]" count="10" pageId="75" pageNumber="416" type="male">10 males</specimenCount>
and
<specimenCount id="9D79FD02FFDEFFC23DC8614CFDD97605" box="[415,535,317,341]" count="10" pageId="75" pageNumber="416" type="female">10 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFFDEFFC23E1E614CFDB17605" box="[585,639,317,341]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="75" pageNumber="416" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFDEFFC23ED9614CFD737605" box="[654,701,317,341]" pageId="75" pageNumber="416">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFFDEFFC23E9B614CFCD17605" box="[716,799,317,341]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="75" pageNumber="416">MNHN</collectionCode>
) labeled: “CASENT 1023285” to “CASENT 1023294” and “CASENT 1023295” to “CASENT 1023304”, respectively/ “
<collectingCountry id="F368761BFFDEFFC2386F612FFB5F7626" box="[1080,1169,350,374]" name="China" pageId="75" pageNumber="416">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FFDEFFC23C2C610EFEF476C6" box="[123,314,382,406]" country="China" name="Yunnan" pageId="75" pageNumber="416">Yunnan Province</collectingRegion>
,
<collectingCounty id="62A14E07FFDEFFC23D1E610EFE3876C7" box="[329,502,383,407]" pageId="75" pageNumber="416">Fugong County</collectingCounty>
,
<location id="8EA06050FFDEFFC23E52610FFD0276C6" LSID="urn:lsid:plazi:treatment:03D6879DFFDFFFC43C2C640DFB4F73E4:8EA06050FFDEFFC23E52610FFD0276C6" box="[517,716,382,406]" country="China" county="Fugong County" latitude="27.116636" longLatPrecision="1" longitude="98.77667" name="Lishadi Township" pageId="75" pageNumber="416" stateProvince="Yunnan">Lishadi Township</location>
,
<locationDeviation id="9A722FA9FFDEFFC23E8C610EFB5F76C6" box="[731,1169,382,407]" location="Shibali on Shibali Road" pageId="75" pageNumber="416">
<quantity id="4C879B6EFFDEFFC23E8C610EFCE476C6" box="[731,810,382,407]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="416" unit="km" value="0.3">0.3 km</quantity>
above Shibali on Shibali Road
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFDEFFC23C2C61EEFED976E7" box="[123,279,415,439]" degrees="27.1166361" direction="north" orientation="latitude" pageId="75" pageNumber="416" precision="1" value="27.116636">N27.1166361°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFDEFFC23D4861EEFE6276E7" box="[287,428,415,439]" degrees="098.77667" direction="east" orientation="longitude" pageId="75" pageNumber="416" precision="1" value="98.77667">E098.77667°</geoCoordinate>
”/ “
<quantity id="4C879B6EFFDEFFC23D9861EEFDEB76E7" box="[463,549,415,439]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="75" pageNumber="416" unit="m" value="2563.0">
<elevation id="0052D1B8FFDEFFC23D9861EEFDEB76E7" box="[463,549,415,439]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="75" pageNumber="416" unit="m" value="2563.0">2563 m</elevation>
</quantity>
,
<collectingMethod id="523E4E9CFFDEFFC23E6661EEFD0E76E7" box="[561,704,415,439]" pageId="75" pageNumber="416">in pitfall trap</collectingMethod>
,
<date id="FFC1104BFFDEFFC23E9B61EEFC9F76E7" box="[716,849,415,440]" pageId="75" pageNumber="416" value="2004-05-04">
<collectingDate id="EF85E9A3FFDEFFC23E9B61EEFC9F76E7" box="[716,849,415,440]" pageId="75" pageNumber="416" value="2004-05-04">4 May 2004</collectingDate>
</date>
, Stop #DHK-2004-026, D. H.
<taxonomicName id="4C7F4D08FFDEFFC23C2C61B1FF3A7688" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,244,448,472]" pageId="75" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFFDEFFC23D4261B1FE7D7688" box="[277,435,448,472]" pageId="75" pageNumber="416">C. E. Griswold</collectorName>
collectors”. All
<typeStatus id="54C48829FFDEFFC23E0861B1FD0B7688" box="[607,709,448,472]" pageId="75" pageNumber="416" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FFDEFFC2385961B1FB5B7688" box="[1038,1173,448,472]" pageId="75" pageNumber="416" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FFDEFFC23C2C6190FDCA76A8" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,516,481,505]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="416" phylum="Arthropoda" rank="species" species="unisetulosa">
<emphasis id="B90BEA99FFDEFFC23C2C6190FE7876A9" box="[123,438,481,505]" italics="true" pageId="75" pageNumber="416">Trechepaphiopsis unisetulosa</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FFDEFFC23EF96190FCCC76A8" box="[686,770,481,504]" pageId="75" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFDEFFC23CFC627CFD9E7515" blockId="75.[123,1174,524,1693]" pageId="75" pageNumber="416">
<materialsCitation id="3B173CD6FFDEFFC23CFC627CFD827515" country="China" county="Fugong County" elevation="2537" latitude="27.1665" location="Fugong County" longLatPrecision="1" longitude="98.77936" pageId="75" pageNumber="416" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90BEA99FFDEFFC23CFC627CFE957573" bold="true" box="[171,347,525,549]" pageId="75" pageNumber="416">
<smallCapsWord id="8D26A057FFDEFFC23CFC627CFF2A7573" baselines="543,542" box="[171,228,525,549]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="75" pageNumber="416">
<typeStatus id="54C48829FFDEFFC23CFC627CFF2A7573" box="[171,228,525,549]" pageId="75" pageNumber="416">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D26A057FFDEFFC23CA76261FE957573" baselines="542" box="[240,347,528,547]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="75" pageNumber="416">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="F368761BFFDEFFC23DDF627DFE047574" box="[392,458,524,548]" name="China" pageId="75" pageNumber="416">China</collectingCountry>
,
<collectingRegion id="49BBF869FFDEFFC23D8D627CFDE07574" box="[474,558,525,548]" country="China" name="Yunnan" pageId="75" pageNumber="416">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FFDEFFC23E69627CFD257575" box="[574,747,525,549]" pageId="75" pageNumber="416">Fugong County</collectingCounty>
,
<taxonomicName id="4C7F4D08FFDEFFC23EAD627DFC3F7574" authority="Township" authorityName="Township" box="[762,1009,524,548]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="416" phylum="Arthropoda" rank="species" species="shiyueliang">Shiyueliang Township</taxonomicName>
,
<locationDeviation id="9A722FA9FFDEFFC23857627CFF077515" location="Shibali" pageId="75" pageNumber="416">
<quantity id="4C879B6EFFDEFFC23857627CFB817574" box="[1024,1103,524,549]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="75" pageNumber="416" unit="km" value="0.2">0.2 km</quantity>
W of Shibali
</locationDeviation>
,
<geoCoordinate id="EE4B504CFFDEFFC23C83625CFE997515" box="[212,343,557,581]" degrees="27.16650" direction="north" orientation="latitude" pageId="75" pageNumber="416" precision="1" value="27.1665">N27.16650°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFFDEFFC23D08625CFE237515" box="[351,493,557,581]" degrees="098.77936" direction="east" orientation="longitude" pageId="75" pageNumber="416" precision="1" value="98.77936">E098.77936°</geoCoordinate>
,
<quantity id="4C879B6EFFDEFFC23DAF625CFD827515" box="[504,588,557,581]" metricMagnitude="3" metricUnit="m" metricValue="2.537" pageId="75" pageNumber="416" unit="m" value="2537.0">
<elevation id="0052D1B8FFDEFFC23DAF625CFD827515" box="[504,588,557,581]" metricMagnitude="3" metricUnit="m" metricValue="2.537" pageId="75" pageNumber="416" unit="m" value="2537.0">2537 m</elevation>
</quantity>
</materialsCitation>
,
</paragraph>
<paragraph id="8BC0368BFFDEFFC23CFC623FFC7475F7" blockId="75.[123,1174,524,1693]" pageId="75" pageNumber="416">
<emphasis id="B90BEA99FFDEFFC23CFC623FFE257534" bold="true" box="[171,491,590,614]" pageId="75" pageNumber="416">
<smallCapsWord id="8D26A057FFDEFFC23CFC623FFEE07534" baselines="608,607" box="[171,302,590,614]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="75" pageNumber="416">DERIVATION</smallCapsWord>
<smallCapsWord id="8D26A057FFDEFFC23D646220FE817534" baselines="607" box="[307,335,593,612]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="75" pageNumber="416">OF</smallCapsWord>
<smallCapsWord id="8D26A057FFDEFFC23D036220FE697534" baselines="607" box="[340,423,593,612]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="75" pageNumber="416">SPECIES</smallCapsWord>
<smallCapsWord id="8D26A057FFDEFFC23DF96220FE257534" baselines="607" box="[430,491,593,612]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="75" pageNumber="416">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FFDEFFC23EBE623FFCAC7536" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[745,866,590,614]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="416" phylum="Arthropoda" rank="species" species="unisetulosa">
<emphasis id="B90BEA99FFDEFFC23EBE623FFCAC7536" box="[745,866,590,614]" italics="true" pageId="75" pageNumber="416">unisetulosa</emphasis>
</taxonomicName>
, is an adjective derived from the Latin words,
<emphasis id="B90BEA99FFDEFFC23D61621EFEA675D6" box="[310,360,623,646]" italics="true" pageId="75" pageNumber="416">unus</emphasis>
, meaning one, and
<emphasis id="B90BEA99FFDEFFC23E15621EFD6A75D7" box="[578,676,623,647]" italics="true" pageId="75" pageNumber="416">setulosus</emphasis>
, meaning bearing bristles or setae. The name refers to the single discal seta found on the elytra of members of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDEFFC23CFC62C1FE6274BE" pageId="75" pageNumber="416" type="diagnosis">
<paragraph id="8BC0368BFFDEFFC23CFC62C1FE6274BE" blockId="75.[123,1174,524,1693]" pageId="75" pageNumber="416">
<smallCapsWord id="8D26A057FFDEFFC23CFC62C1FEEE7596" baselines="706,705" box="[171,288,688,712]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="75" pageNumber="416">
<emphasis id="B90BEA99FFDEFFC23CFC62C1FEEE7596" bold="true" box="[171,288,688,712]" pageId="75" pageNumber="416">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFFDEFFC23E1062C1FD6D7598" box="[583,675,688,712]" captionStart="FIGURE" captionStartId="76.[171,183,1581,1600]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-11@76.[120,1176,216,1564]" captionTargetPageId="76" captionText="FIGURE 29. Trechepaphiopsis unisetulosa sp. nov.; a. Dorsal habitus (CASENT1018407). b. Median lobe of aedeagus of male (CASENT1001903), left lateral aspect. c. Map of locality records (red circles) for T. unisetulosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155338" httpUri="https://zenodo.org/record/13155338/files/figure.png" pageId="75" pageNumber="416">Fig. 29a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size small to medium (BL =
<quantity id="4C879B6EFFDEFFC23CC76280FEE4745A" box="[144,298,753,778]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="75" pageNumber="416" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
), apterous; body color dark reddish brown, dorsum shiny, very slightly iridescent; eyes small, only slightly projected laterally; tempora convex and sparsely pubescent; pronotum transverse (ratio PW/PL = 1.40), with basal angles obtuse and rounded; elytra convex, with medial three or four stria distinct, finely punctate, more lateral striae more or less effaced; recurrent stria terminated anteriorly in presumed location of interval 6; only one discal setiferous pore present, inserted near anterior one-third next to stria 3; preapical seta present, inserted next to stria 2; medi- an lobe of male aedeagus (
<figureCitation id="13442A0EFFDEFFC23DF763C4FE30749D" box="[416,510,949,973]" captionStart="FIGURE" captionStartId="76.[171,183,1581,1600]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-11@76.[120,1176,216,1564]" captionTargetPageId="76" captionText="FIGURE 29. Trechepaphiopsis unisetulosa sp. nov.; a. Dorsal habitus (CASENT1018407). b. Median lobe of aedeagus of male (CASENT1001903), left lateral aspect. c. Map of locality records (red circles) for T. unisetulosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155338" httpUri="https://zenodo.org/record/13155338/files/figure.png" pageId="75" pageNumber="416">Fig. 29b</figureCitation>
) with apex lobated and broadly rounded; endophallus with a large, scaly sclerorized area.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFDEFFC43CFC6386FB4F73E4" lastPageId="77" lastPageNumber="418" pageId="75" pageNumber="416" type="description">
<paragraph id="8BC0368BFFDEFFC23CFC6386FDA17321" blockId="75.[123,1174,524,1693]" pageId="75" pageNumber="416">
<smallCapsWord id="8D26A057FFDEFFC23CFC6386FEF4735D" baselines="1033,1032" box="[171,314,1015,1039]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="75" pageNumber="416">
<emphasis id="B90BEA99FFDEFFC23CFC6386FEF4735D" bold="true" box="[171,314,1015,1039]" pageId="75" pageNumber="416">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size small to medium, BL =
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. Body color dark reddish brown, interval 1 of elytra slightly paler, reddish, appendages yellowish brown, palpi paler yellow; dorsum smooth, pronotum and elytra shiny, very slightly iridescent, head slightly alutaceous from more deeply impressed isodiametric microsculpture.
</paragraph>
<paragraph id="8BC0368BFFDEFFC23CFC640BFCA572C7" blockId="75.[123,1174,524,1693]" pageId="75" pageNumber="416">
Head. Rather broad; eyes small, only slightly projected laterally, their diameter about equal to length of tempora, the latter convex and sparsely pubescent. Frons convex, with frontal furrows evenly rounded and deeply impressed, continuous to posterior margins of tempora. Two pairs of supraorbital setae present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly emarginate.
<collectingRegion id="49BBF869FFDEFFC23EE1648CFCDE7244" box="[694,784,1277,1300]" country="Myanmar" name="Chin" pageId="75" pageNumber="416">Mentum</collectingRegion>
and submentum not fused.
<collectingRegion id="49BBF869FFDEFFC2386C648CFB5B7244" box="[1083,1173,1277,1300]" country="Myanmar" name="Chin" pageId="75" pageNumber="416">Mentum</collectingRegion>
with medial tooth broad and apically bifid, less than half the length of the lateral lobes. Submentum with six setae anteriorly. Gula broad. Genae with a single ventral seta one each side. Antennae short, with only about 1.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 4 about equal in length, antennomere 3 slightly longer.
</paragraph>
<paragraph id="8BC0368BFFDEFFC23CFC65D1FDB071CD" blockId="75.[123,1174,524,1693]" pageId="75" pageNumber="416">Pronotum. Transverse (ratio PW/PL = 1.40), with greatest width near anterior one-third, moderately narrowed posteriorly; lateral margins rounded, slightly more so anteriorly, slightly straighten, but not at all sinuate, just anterior to basal angles, the latter bluntly obtuse. Disc convex, medi- an longitudinal impression finely impressed, continuous from posterior side of apical median swelling to posterior margin; basal foveae shallow, faintly impressed; median basal area well defined but short and transverse. Lateral borders slender and finely relexed dorsally, lateral grooves deeply impressed. Single midlateral setae on each side inserted near anterior one-third; single basolateral seta on each side, inserted at basal angle.</paragraph>
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29.
<taxonomicName id="4C7F4D08FFD9FFC53D5A665CFDC47110" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[269,522,1581,1600]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="417" phylum="Arthropoda" rank="species" species="unisetulosa" status="sp. nov.">
<emphasis id="B90BEA99FFD9FFC53D5A665CFDC47110" box="[269,522,1581,1600]" italics="true" pageId="76" pageNumber="417">Trechepaphiopsis unisetulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD9FFC53E58665CFD9B7110" box="[527,597,1581,1600]" pageId="76" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1018407). b. Median lobe of aedeagus of male (CASENT1001903), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FFD9FFC53FDF6636FC307109" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[904,1022,1606,1625]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="417" phylum="Arthropoda" rank="species" species="unisetulosa">
<emphasis id="B90BEA99FFD9FFC53FDF6636FC307109" box="[904,1022,1606,1625]" italics="true" pageId="76" pageNumber="417">T. unisetulosa</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFFD8FFC43CFC60AAFE687543" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">Elytra. Ovoid, not or only slightly elongate, about equally narrowed anteriorly and posteriorly, humeri distinct but rounded. Disc convex, striae faintly impressed and finely punctate, the medial three or four striae distinctly impressed, the more lateral striae more or less effaced, bearly perceptible. Parascutellar striole present, longer than average for genus. Recurrent stria abruptly terminated anteriorly in presumed location of posterior end of stria 5. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Only one discal setiferous pore present, inserted at anterior two-fifth of elytral length next to stria 3. Preapical seta present on interval 3 next to stria 2 and about equidistant from apical and sutural elytral margins. Umbilicate setal series with setae of humeral group equidistance from each other and setae of median group inserted slightly posterior to middle.</paragraph>
<paragraph id="8BC0368BFFD8FFC43CFC626DFE9E7503" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFFD8FFC43CFC622AFB8575C3" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.</paragraph>
<paragraph id="8BC0368BFFD8FFC43CFC62EAFDAA75A3" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFFD8FFC43DAC62EAFD9575E3" box="[507,603,667,691]" captionStart="FIGURE" captionStartId="76.[171,183,1581,1600]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-11@76.[120,1176,216,1564]" captionTargetPageId="76" captionText="FIGURE 29. Trechepaphiopsis unisetulosa sp. nov.; a. Dorsal habitus (CASENT1018407). b. Median lobe of aedeagus of male (CASENT1001903), left lateral aspect. c. Map of locality records (red circles) for T. unisetulosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155338" httpUri="https://zenodo.org/record/13155338/files/figure.png" pageId="77" pageNumber="418">Fig. 29b</figureCitation>
) with sagittal aileron moderate in size, shaft broad basally, then gradually narrowed apically to a broadly rounded apex; endophallus with a large, scaly sclerorized area with two parts or folds.
</paragraph>
<paragraph id="8BC0368BFFD8FFC43CFC628DFD1274A4" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">
<emphasis id="B90BEA99FFD8FFC43CFC628DFE6B7442" bold="true" box="[171,421,764,788]" pageId="77" pageNumber="418">
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<smallCapsWord id="8D26A057FFD8FFC43D46628EFE6B7442" baselines="781" box="[273,421,767,786]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="77" pageNumber="418">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been collected in pitfall traps set in slightly disturbed broadleaf forest with large trees, dense understory of ferns and other herbs, and a deep leaf litter layer, and also by sifting leaf litter in this same habitat.
<specimenCount id="9D79FD02FFD8FFC43FD9634DFBE67404" box="[910,1064,828,852]" count="1" pageId="77" pageNumber="418" type="generic">One specimen</specimenCount>
was beaten from roadside vegetation with suspended leaf and twig debris in that vegetation in the same area.
<specimenCount id="9D79FD02FFD8FFC43C2C630DFEDC74C4" box="[123,274,892,916]" count="1" pageId="77" pageNumber="418" type="generic">One specimen</specimenCount>
was collected at a higher elevation (
<quantity id="4C879B6EFFD8FFC43EC6630DFD2A74C4" box="[657,740,892,916]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="77" pageNumber="418" unit="m" value="3100.0">3100 m</quantity>
) from under stones in a rocky open areas cleared by snow avalanches but shaded by three meter high herbaceous cover (
<figureCitation id="13442A0EFFD8FFC43F8A63EDFBF374E4" box="[989,1085,924,948]" captionStart="FIGURE" captionStartId="102.[175,187,1623,1642]" captionTargetBox="[128,1169,216,1606]" captionTargetId="figure-12@102.[128,1169,216,1606]" captionTargetPageId="102" captionText="FIGURE 40. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, 21 air km W of Cikai on east slope of Qiqi/Dulong divide, elevation 3600 m; habitat in which specimens of Queinnectrechus balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, elevation 3100 m; habitat in which specimens of Trechus shiyueliang sp. nov., T. shibalicus sp. nov., Trechepaphiopsis unisetulosa sp. nov. and T. unipilosa sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155364" httpUri="https://zenodo.org/record/13155364/files/figure.png" pageId="77" pageNumber="418">Fig. 40b</figureCitation>
). Specimens of
<taxonomicName id="4C7F4D08FFD8FFC43C8263CDFE677484" box="[213,425,956,980]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="418" phylum="Arthropoda" rank="species" species="shiyueliang">
<emphasis id="B90BEA99FFD8FFC43C8263CDFE677484" box="[213,425,956,980]" italics="true" pageId="77" pageNumber="418">Trechus shiyueliang</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D08FFD8FFC43DE163CCFDF57484" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[438,571,956,980]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="418" phylum="Arthropoda" rank="species" species="shibalicus">
<emphasis id="B90BEA99FFD8FFC43DE163CCFDF57484" box="[438,571,956,980]" italics="true" pageId="77" pageNumber="418">T. shibalicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D08FFD8FFC43E3963CDFC5C7484" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[622,914,956,980]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="418" phylum="Arthropoda" rank="species" species="unipilosa" status="sp. nov.">
<emphasis id="B90BEA99FFD8FFC43E3963CDFC5C7484" box="[622,914,956,980]" italics="true" pageId="77" pageNumber="418">Trechepaphiopsis unipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD8FFC43FCF63CDFC277483" box="[920,1001,956,979]" pageId="77" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
were found syntopic with the specimen of
<taxonomicName id="4C7F4D08FFD8FFC43DF763ACFDFD74A4" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[416,563,988,1012]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="418" phylum="Arthropoda" rank="species" species="unisetulosa">
<emphasis id="B90BEA99FFD8FFC43DF763ACFDFD74A4" box="[416,563,988,1012]" italics="true" pageId="77" pageNumber="418">T. unisetulosa</emphasis>
</taxonomicName>
at this last site.
</paragraph>
<paragraph id="8BC0368BFFD8FFC43CFC638DFDCD7324" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">
<emphasis id="B90BEA99FFD8FFC43CFC638DFD4E7343" bold="true" box="[171,640,1020,1044]" pageId="77" pageNumber="418">
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<smallCapsWord id="8D26A057FFD8FFC43D08638EFE3D7342" baselines="1037" box="[351,499,1023,1042]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="77" pageNumber="418">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFD8FFC43DAA6471FD827343" baselines="1038" box="[509,588,1024,1043]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="77" pageNumber="418">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFD8FFC43E016471FD4E7343" baselines="1038" box="[598,640,1024,1043]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="77" pageNumber="418">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FFD8FFC43EDD638DFCC17342" baselines="1038,1037" box="[650,783,1020,1044]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="77" pageNumber="418">
<emphasis id="B90BEA99FFD8FFC43EDD638DFCC17342" bold="true" box="[650,783,1020,1044]" pageId="77" pageNumber="418">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFD8FFC43F4E638DFEAC7304" location="We" pageId="77" pageNumber="418" specimenCount="54" specimenCount-female="14" specimenCount-male="13" stateProvince="Shan">
<smallCapsWord id="8D26A057FFD8FFC43F4E638DFC9A7342" baselines="1038,1037" box="[793,852,1020,1044]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Shan" pageId="77" pageNumber="418">
<emphasis id="B90BEA99FFD8FFC43F4E638DFC9A7342" bold="true" box="[793,852,1020,1044]" pageId="77" pageNumber="418">
<collectingRegion id="49BBF869FFD8FFC43F4E638DFC9A7342" box="[793,852,1020,1044]" country="Myanmar" name="Shan" pageId="77" pageNumber="418">SHAN</collectingRegion>
</emphasis>
</smallCapsWord>
.—
<figureCitation id="13442A0EFFD8FFC43FD7638DFC107344" box="[896,990,1020,1044]" captionStart="FIGURE" captionStartId="76.[171,183,1581,1600]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-11@76.[120,1176,216,1564]" captionTargetPageId="76" captionText="FIGURE 29. Trechepaphiopsis unisetulosa sp. nov.; a. Dorsal habitus (CASENT1018407). b. Median lobe of aedeagus of male (CASENT1001903), left lateral aspect. c. Map of locality records (red circles) for T. unisetulosa in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155338" httpUri="https://zenodo.org/record/13155338/files/figure.png" pageId="77" pageNumber="418">Fig. 29c</figureCitation>
.
<location id="8EA06050FFD8FFC43FBC638DFBC07343" LSID="urn:lsid:plazi:treatment:03D6879DFFDFFFC43C2C640DFB4F73E4:8EA06050FFD8FFC43FBC638DFBC07343" box="[1003,1038,1020,1043]" name="We" pageId="77" pageNumber="418" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFD8FFC43C99646DFE907364" box="[206,350,1052,1076]" count="27" pageId="77" pageNumber="418" type="generic">27 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFD8FFC43D3A646DFE037364" box="[365,461,1052,1076]" count="13" pageId="77" pageNumber="418" type="male">13 males</specimenCount>
and
<specimenCount id="9D79FD02FFD8FFC43E55646DFDB67364" box="[514,632,1052,1076]" count="14" pageId="77" pageNumber="418" type="female">14 females</specimenCount>
) from sites on the eastern slope of the northcentral part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FFD8FFC43D3E644DFE317324" county="Fugong County" location="Core Area" pageId="77" pageNumber="418" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FFD8FFC43D3E644DFE507304" box="[361,414,1084,1108]" country="Myanmar" name="Shan" pageId="77" pageNumber="418">Shan</collectingRegion>
in
<collectingCounty id="62A14E07FFD8FFC43D96644DFDA77304" box="[449,617,1084,1108]" pageId="77" pageNumber="418">Fugong County</collectingCounty>
(see
<typeStatus id="54C48829FFD8FFC43EF6644DFD187304" box="[673,726,1084,1108]" pageId="77" pageNumber="418">Type</typeStatus>
material above for exact collection data). All of these sites are in
<location id="8EA06050FFD8FFC43D2D642DFE267324" LSID="urn:lsid:plazi:treatment:03D6879DFFDFFFC43C2C640DFB4F73E4:8EA06050FFD8FFC43D2D642DFE267324" box="[378,488,1116,1140]" county="Fugong County" name="Core Area" pageId="77" pageNumber="418" stateProvince="Shan">Core Area</location>
3
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFD8FFC43CFC640DFB4F73E4" blockId="77.[123,1174,219,1204]" pageId="77" pageNumber="418">
<emphasis id="B90BEA99FFD8FFC43CFC640DFD9673C2" bold="true" box="[171,600,1148,1172]" pageId="77" pageNumber="418">
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<smallCapsWord id="8D26A057FFD8FFC43D4F640EFE7173C2" baselines="1165" box="[280,447,1151,1170]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="77" pageNumber="418">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FFD8FFC43D93640EFD9673C2" baselines="1165" box="[452,600,1151,1170]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="77" pageNumber="418">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species currently is known only from Fugong County in the northcentral part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFD8FFC43F2364ECFBE173E4" box="[884,1071,1180,1204]" country="China" name="Yunnan" pageId="77" pageNumber="418">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFD8FFC4386D64EDFBB373E4" box="[1082,1149,1180,1204]" name="China" pageId="77" pageNumber="418">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,455 @@
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<mods:title id="296E4FB2F4CE07810F6CD9BDCE03A01C">A Synopsis of the Eastern and Central Atlantic Combers of the Genus Serranus (Teleostei: Scorpaeniformes: Serranidae)</mods:title>
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<mods:affiliation id="98EDD9BF11B62151A1F655489DCEDEA1">Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118 U. S. A. e-mail: tiwamoto @ calacademy. org.</mods:affiliation>
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<mods:date id="CA37C043EC6F37B78E3C60892092972B">2018</mods:date>
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<mods:number id="9E21BE3654DC41711B71FEE15DCFA126">2018-09-28</mods:number>
</mods:detail>
<mods:detail id="757211924673AEA792FDCD2DB87C638A" type="volume">
<mods:number id="4440850D529D9C3FDBAB5BF9CB890613">65</mods:number>
</mods:detail>
<mods:detail id="C85ECEACE418F5081FDD6ADEF810776D" type="issue">
<mods:number id="838EF25CD5B300D09FDAC992F76C9DFA">1</mods:number>
</mods:detail>
<mods:extent id="8F67E9BE0AC705D0E07FF7AC82A54FF0" unit="page">
<mods:start id="86092397925C3F4D147A15408ABF1630">1</mods:start>
<mods:end id="C62B132FFAFCD6AC9DC97525E3E6C0A0">39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D96822DD973B3985C966B0E4C3D01431">journal article</mods:classification>
<mods:identifier id="0BBBA09695633C49735C55AF79C103DC" type="DOI">10.5281/zenodo.13156001</mods:identifier>
<mods:identifier id="53A0A640F6E4F6A815B020CF54FCF744" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="82589318A66D2BBAE0D0496AC4A79AC0" type="Zenodo-Dep">13156001</mods:identifier>
<mods:identifier id="5422667725C55D96E07E849F5A5B95FA" type="ZooBank">C84C1C06-23EC-4BDC-B868-8BA658E7E9D4</mods:identifier>
</mods:mods>
<treatment id="3E1987F70403FF91FFEEFBF4A5E7FDA8" LSID="urn:lsid:plazi:treatment:3E1987F70403FF91FFEEFBF4A5E7FDA8" httpUri="http://treatment.plazi.org/id/3E1987F70403FF91FFEEFBF4A5E7FDA8" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<subSubSection id="FEAA656A0403FF9CFFEEFBF4A54DFBF1" box="[123,639,1096,1120]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="B60F36E10403FF9CFFEEFBF4A54DFBF1" blockId="7.[123,639,1096,1151]" box="[123,639,1096,1120]" pageId="7" pageNumber="8">
<heading id="ED47818D0403FF9CFFEEFBF4A54DFBF1" box="[123,639,1096,1120]" fontSize="10" level="2" pageId="7" pageNumber="8" reason="4">
<taxonomicName id="71B04D620403FF9CFFEEFBF4A54DFBF1" ID-CoL="4WWXZ" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[123,639,1096,1120]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cabrilla" subGenus="Serranus">
<emphasis id="84C4EAF30403FF9CFFEEFBF4A54DFBF1" bold="true" box="[123,639,1096,1120]" pageId="7" pageNumber="8">
<emphasis id="84C4EAF30403FF9CFFEEFBF4A7D2FBF1" bold="true" box="[123,224,1096,1120]" italics="true" pageId="7" pageNumber="8">Serranus</emphasis>
(
<emphasis id="84C4EAF30403FF9CFF65FBF4A667FBF1" bold="true" box="[240,341,1096,1120]" italics="true" pageId="7" pageNumber="8">Serranus</emphasis>
)
<emphasis id="84C4EAF30403FF9CFEF0FBF4A68BFBF1" bold="true" box="[357,441,1096,1120]" italics="true" pageId="7" pageNumber="8">cabrilla</emphasis>
(Linnaeus, 1758)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0403FF9CFFEEFBD5A552FB3D" pageId="7" pageNumber="8" type="description">
<paragraph id="B60F36E10403FF9CFFEEFBD5A7DFFBEF" blockId="7.[123,639,1096,1151]" box="[123,237,1128,1151]" pageId="7" pageNumber="8">
<figureCitation id="2E8B2A640403FF9CFFEEFBD5A7DFFBEF" box="[123,237,1128,1151]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="8.[155,167,732,751]" captionStartId-1="8.[155,167,1217,1236]" captionStartId-2="8.[155,167,1674,1693]" captionTargetBox-0="[120,1176,216,720]" captionTargetBox-1="[120,1176,773,1205]" captionTargetBox-2="[162,1152,1272,1661]" captionTargetId-0="figure-49@8.[120,1176,215,721]" captionTargetId-1="figure-51@8.[120,1176,772,1206]" captionTargetId-2="figure-53@8.[120,1176,1257,1663]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionTargetPageId-2="8" captionText-0="FIGURE 5. Serranus cabrilla. Adult from near Hyères, Var, Provence, France. Photograph by Lucas Berenger." captionText-1="FIGURE 6. Serranus cabrilla. Adult from the Canary Islands. Photograph by Rogelio Herrera." captionText-2="FIGURE 7. Serranus cabrilla. Yellow morph from Ivory Coast (4°34ʹN, 6°37ʹW), collected by R/V Dr. Fridtjof Nansen by bottom trawl in 81 m. Photograph by Oddgeir Alvheim." figureDoi-0="http://doi.org/10.5281/zenodo.13156015" figureDoi-1="http://doi.org/10.5281/zenodo.13156019" figureDoi-2="http://doi.org/10.5281/zenodo.13156021" httpUri-0="https://zenodo.org/record/13156015/files/figure.png" httpUri-1="https://zenodo.org/record/13156019/files/figure.png" httpUri-2="https://zenodo.org/record/13156021/files/figure.png" pageId="7" pageNumber="8">Figures 57</figureCitation>
</paragraph>
<paragraph id="B60F36E10403FF9CFFEEFB24A552FB3D" blockId="7.[123,1173,1175,1286]" box="[123,608,1175,1197]" pageId="7" pageNumber="8">
<taxonomicName id="71B04D620403FF9CFFEEFB24A693FB3C" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[123,417,1175,1197]" family="Percidae" genus="Perca" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="84C4EAF30403FF9CFFEEFB24A634FB3D" box="[123,262,1175,1197]" italics="true" pageId="7" pageNumber="8">Perca cabrilla</emphasis>
Linnaeus, 1758
</taxonomicName>
(no locality given).
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0403FF9CFFEEFB09A350FA97" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="B60F36E10403FF9CFFEEFB09A372FB5B" blockId="7.[123,1173,1175,1286]" box="[123,1088,1205,1227]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="96A011CF0403FF9CFFEEFB09A372FB5B" box="[123,1088,1205,1227]" pageId="7" pageNumber="8">
<taxonomicName id="71B04D620403FF9CFFEEFB09A644FB5B" ID-CoL="4CP2P" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[123,374,1205,1226]" family="Serranidae" genus="Paracentropristis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="84C4EAF30403FF9CFFEEFB09A644FB5B" box="[123,374,1205,1226]" italics="true" pageId="7" pageNumber="8">Paracentropristis cabrilla</emphasis>
</taxonomicName>
:
<materialsCitation id="06D83CBC0403FF9CFE16FB09A308FB5B" box="[387,1082,1205,1227]" location="Madeira, Canaries, Azores" pageId="7" pageNumber="8" specimenCount="14" stateProvince="Madeira, Canaries, Azores">
<treatmentCitation id="371110F00403FF9CFE16FB09A50BFB5B" author="FOWLER, H. W." box="[387,569,1205,1227]" page="1291" pageId="7" pageNumber="8" year="1936">
<bibRefCitation id="D2214B100403FF9CFE16FB09A50BFB5B" author="FOWLER, H. W." box="[387,569,1205,1227]" pageId="7" pageNumber="8" pagination="607 - 1493" refId="ref20427" refString="FOWLER, H. W. 1936. The marine fishes of West Africa, based on collections of the American Museum Congo Expedition, 1908 - 15. Bulletin of the American Museum of Natural History 70 (1): 1 - 106 (2): 607 - 1493." type="journal article" year="1936">Fowler, 1936:1291</bibRefCitation>
</treatmentCitation>
(
<specimenCount id="A0B6FD680403FF9CFDDDFB09A5A5FB5B" box="[584,663,1205,1226]" count="14" pageId="7" pageNumber="8" type="generic">14 spec.</specimenCount>
,
<quantity id="71489B040403FF9CFD31FB09A411FB5B" box="[676,803,1205,1227]" metricMagnitude="-1" metricUnit="m" metricValue="2.31" metricValueMax="3.25" metricValueMin="1.3699999999999999" pageId="7" pageNumber="8" unit="mm" value="231.0" valueMax="325.0" valueMin="137.0">137325 mm</quantity>
;
<collectingRegion id="7474F8030403FF9CFCA4FB09A300FB5B" box="[817,1074,1205,1227]" isEnumeration="true" pageId="7" pageNumber="8">Madeira, Canaries, Azores</collectingRegion>
)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B60F36E10403FF9CFFEEFB6FA350FA97" blockId="7.[123,1173,1175,1286]" pageId="7" pageNumber="8">
<emphasis id="84C4EAF30403FF9CFFEEFB6FA61FFB79" box="[123,301,1235,1256]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="71B04D620403FF9CFFEEFB6FA617FB79" ID-CoL="4WWXZ" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[123,293,1235,1256]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cabrilla">Serranus cabrilla</taxonomicName>
:
</emphasis>
<bibRefCitation id="D2214B100403FF9CFEA1FB6FA6E6FB79" author="POLL, M." box="[308,468,1234,1257]" pageId="7" pageNumber="8" pagination="1 - 390" refId="ref21292" refString="POLL, M. 1954. Poissons. IV. Teleosteens acanthopterygiens. (Premiere partie). Resultats Scientifiques. Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de l'Atlantique Sud (1948 - 1949), 4 (3 A): 1 - 390, pls. I - IX." type="journal article" year="1954">Poll 1954:6972</bibRefCitation>
, fig. 19 (
<specimenCount id="A0B6FD680403FF9CFDA5FB6FA5B2FB79" box="[560,640,1235,1256]" count="17" pageId="7" pageNumber="8" type="generic">17 spec.</specimenCount>
,
<quantity id="71489B040403FF9CFD18FB6FA430FB79" box="[653,770,1235,1256]" metricMagnitude="-1" metricUnit="m" metricValue="1.595" metricValueMax="2.4" metricValueMin="0.79" pageId="7" pageNumber="8" unit="mm" value="159.5" valueMax="240.0" valueMin="79.0">79240 mm</quantity>
;
<collectingCountry id="CEA776710403FF9CFC85FB6FA462FB78" box="[784,848,1235,1257]" name="Gabon" pageId="7" pageNumber="8">Gabon</collectingCountry>
to s.
<collectingCountry id="CEA776710403FF9CFC12FB6FA4FCFB78" box="[903,974,1235,1257]" name="Angola" pageId="7" pageNumber="8">Angola</collectingCountry>
). Robins and Starck 1961:261262, figs. 1a, 7c, 8a, 8c (
<specimenCount id="A0B6FD680403FF9CFE67FB4DA507FA97" box="[498,565,1265,1286]" count="4" pageId="7" pageNumber="8" type="generic">4 spec.</specimenCount>
,
<collectingCountry id="CEA776710403FF9CFDD7FB4CA54BFA97" box="[578,633,1264,1286]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
and
<collectingCountry id="CEA776710403FF9CFD3CFB4CA5E7FA97" box="[681,725,1264,1286]" name="Italy" pageId="7" pageNumber="8">Italy</collectingCountry>
).
<bibRefCitation id="D2214B100403FF9CFD7CFB4DA4DEFA97" author="QUERO, J. C. &amp; J. C. HUREAU &amp; C. KARRER &amp; A. POST AND &amp; SALDANHA, EDS" box="[745,1004,1264,1286]" pageId="7" pageNumber="8" pagination="520 - 1080" refId="ref21346" refString="QUERO, J. C., J. C. HUREAU, C. KARRER, A. POST AND L. SALDANHA, EDS. 1990. Clofeta II. Check-list of the fishes of the eastern tropical Atlantic. Catalogue des poisons de l'Atlantique oriental tropical. Junta Nacional de Investigacao Cientifica e Tecnologica, Lisbon, Portugal. Pages [3 unnumbered] + 520 - 1080." type="journal article" year="1990">Quero et al. 1990:704705</bibRefCitation>
(compiled).
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0403FF9CFF3EFA9CA5E8FA28" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="B60F36E10403FF9CFF3EFA9CA5E8FA28" blockId="7.[123,1174,1311,1725]" pageId="7" pageNumber="8">
<smallCapsWord id="B0E9A03D0403FF9CFF3EFA9CA612FAA7" baselines="1330,1329" box="[171,288,1312,1336]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="7" pageNumber="8">
<emphasis id="84C4EAF30403FF9CFF3EFA9CA612FAA7" bold="true" box="[171,288,1312,1336]" pageId="7" pageNumber="8">DIAGNOSIS</emphasis>
</smallCapsWord>
.— D X,1314; P 1416; V III,7; gill rakers 68+1216 (1824 total); pored lateral-line scales 7077; circumpeduncular scales 3640. Caudal fin shallowly forked, upper lobe slightly longer than lower lobe, a very short streamer on upper lobe in juveniles, but none developed in adults. Interradial membranes of dorsal and anal fins scaled on basal one-half to one-third. Nape, opercle, cheek, pectoral-fin base and chest scaled.
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0403FF91FF3EFA7EA5E7FDA8" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="description">
<paragraph id="B60F36E10403FF9CFF3EFA7EA33EF92C" blockId="7.[123,1174,1311,1725]" pageId="7" pageNumber="8">
<smallCapsWord id="B0E9A03D0403FF9CFF3EFA7EA608FA49" baselines="1492,1491" box="[171,314,1474,1498]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="7" pageNumber="8">
<emphasis id="84C4EAF30403FF9CFF3EFA7EA608FA49" bold="true" box="[171,314,1474,1498]" pageId="7" pageNumber="8">DESCRIPTION</emphasis>
</smallCapsWord>
.— Body compressed, width across pectoral fin base about
<quantity id="71489B040403FF9CFC23FA7EA4EDFA4B" box="[950,991,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="8" unit="in" value="2.0">2 in</quantity>
HL; depth about
<quantity id="71489B040403FF9CFFEEFA5FA791FA6B" box="[123,163,1506,1531]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="8" unit="in" value="3.0">3 in</quantity>
SL; head about 2.7
<quantity id="71489B040403FF9CFE15FA5FA68EFA6B" box="[384,444,1506,1531]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" pageId="7" pageNumber="8" unit="in" value="2.8">2.8 in</quantity>
SL. Dorsal and ventral profiles gently curved; the snout pointed and longer than orbit diameter; interorbital width (fleshy) slightly greater than orbit diameter; lower jaw projecting beyond upper jaw; maxilla extending to below midorbit. Anterior nostrils with low rims lacking fingerlike fringes or cirri. Opercular spines three, the upper two fairly well developed, the lowermost obscure in adults, but more visible in juveniles. Preopercle margin coarsely serrat- ed, largest spines near angle. Interopercle smooth, (a few imbedded scales in large specimens); snout, interorbit, occiput, suborbit, jaws, gular and branchiostegal membranes naked.
</paragraph>
<caption id="E2CF6669040CFF93FF0EFD60A371FD7E" ID-DOI="http://doi.org/10.5281/zenodo.13156015" ID-Zenodo-Dep="13156015" box="[155,1091,731,751]" httpUri="https://zenodo.org/record/13156015/files/figure.png" pageId="8" pageNumber="9" startId="8.[155,167,732,751]" targetBox="[120,1176,216,720]" targetPageId="8" targetType="figure">
<paragraph id="B60F36E1040CFF93FF0EFD60A371FD7E" blockId="8.[155,1091,731,751]" box="[155,1091,731,751]" pageId="8" pageNumber="9">
<smallCapsWord id="B0E9A03D040CFF93FF0EFD60A7EAFD7C" baselines="746,745" box="[155,216,732,751]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="8" pageNumber="9">FIGURE</smallCapsWord>
5.
<emphasis id="84C4EAF3040CFF93FF61FD60A6BDFD7F" box="[244,399,731,751]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="71B04D62040CFF93FF61FD60A6B9FD7F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[244,395,731,751]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cabrilla">Serranus cabrilla</taxonomicName>
.
</emphasis>
Adult from near Hyères, Var, Provence, France. Photograph by Lucas Berenger.
</paragraph>
</caption>
<caption id="E2CF6669040CFF93FF0EFB7DA48BFB45" ID-DOI="http://doi.org/10.5281/zenodo.13156019" ID-Zenodo-Dep="13156019" box="[155,953,1216,1236]" httpUri="https://zenodo.org/record/13156019/files/figure.png" pageId="8" pageNumber="9" startId="8.[155,167,1217,1236]" targetBox="[120,1176,773,1205]" targetPageId="8" targetType="figure">
<paragraph id="B60F36E1040CFF93FF0EFB7DA48BFB45" blockId="8.[155,953,1216,1236]" box="[155,953,1216,1236]" pageId="8" pageNumber="9">
<smallCapsWord id="B0E9A03D040CFF93FF0EFB7DA7EAFB43" baselines="1231,1230" box="[155,216,1217,1236]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="8" pageNumber="9">FIGURE</smallCapsWord>
6.
<emphasis id="84C4EAF3040CFF93FF61FB7DA6BDFB42" box="[244,399,1216,1236]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="71B04D62040CFF93FF61FB7DA6B9FB42" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[244,395,1216,1236]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cabrilla">Serranus cabrilla</taxonomicName>
.
</emphasis>
Adult from the Canary Islands. Photograph by Rogelio Herrera.
</paragraph>
</caption>
<caption id="E2CF6669040CFF93FF0EF936A589F929" ID-DOI="http://doi.org/10.5281/zenodo.13156021" ID-Zenodo-Dep="13156021" httpUri="https://zenodo.org/record/13156021/files/figure.png" pageId="8" pageNumber="9" startId="8.[155,167,1674,1693]" targetBox="[162,1152,1272,1661]" targetPageId="8" targetType="figure">
<paragraph id="B60F36E1040CFF93FF0EF936A362F90C" blockId="8.[123,1104,1673,1720]" box="[155,1104,1673,1693]" pageId="8" pageNumber="9">
<smallCapsWord id="B0E9A03D040CFF93FF0EF936A7EAF90A" baselines="1688,1687" box="[155,216,1674,1693]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="8" pageNumber="9">FIGURE</smallCapsWord>
7.
<emphasis id="84C4EAF3040CFF93FF61F936A6BDF90D" box="[244,399,1674,1693]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="71B04D62040CFF93FF61F936A6B9F90C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[244,395,1674,1693]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cabrilla">Serranus cabrilla</taxonomicName>
.
</emphasis>
Yellow morph from Ivory Coast (4°34ʹN, 6°37ʹW), collected by R/V Dr. Fridtjof
</paragraph>
<paragraph id="B60F36E1040CFF93FFEEF919A589F929" blockId="8.[123,1104,1673,1720]" box="[123,699,1700,1720]" pageId="8" pageNumber="9">Nansen by bottom trawl in 81 m. Photograph by Oddgeir Alvheim.</paragraph>
</caption>
<paragraph id="B60F36E1040DFF92FF3EFF67A340FEC2" blockId="9.[123,1174,219,1726]" pageId="9" pageNumber="10">Premaxillary teeth in several irregular rows, the outer enlarged with one or more large canines at anterior end. A single file of enlarged, spaced teeth on dentary, largest teeth midlength on jaw; much smaller teeth interspersed on sides of larger teeth and a cluster of small teeth at anterior end. Vomer broadly V-shaped with small teeth. Palatine teeth all small, in a short to elongate patch.</paragraph>
<paragraph id="B60F36E1040DFF92FF3EFEE0A419FE45" blockId="9.[123,1174,219,1726]" pageId="9" pageNumber="10">Scales small, ctenoid, present on sides of head and nape to margin of frontals, but absent on interorbital, snout, suborbital region, jaws, gular and branchiostegal membranes, interopercle bone, outer margin of preopercle, and lower margin of subopercle. Small scales at bases of dorsal, anal, pectoral and caudal fins, and minimally (if at all) on pelvic fins.</paragraph>
<paragraph id="B60F36E1040DFF92FF3EFE61A6EDFD47" blockId="9.[123,1174,219,1726]" pageId="9" pageNumber="10">
First 46 spines of dorsal fin graduated, the 6
<superScript id="41C59BA9040DFF92FD3DFE60A585FE7C" attach="left" box="[680,695,476,493]" fontSize="7" pageId="9" pageNumber="10">th</superScript>
spine usually longest, the spines following subequal; the soft rays slightly higher, forming a slight rise (but no distinct notch) in profile of dorsal fin. Anal-fin spines much shorter than soft rays, the first spine somewhat more than half length of second and third spines, the last three or four soft rays longest. Pectoral fin broad-based, its origin slightly behind that of pelvic fins and about on same vertical as origin of dorsal fin, its distal tip over anus, slightly behind origin of anal fin. Pelvic fin falling well short of anus. Caudal fin shallowly lunate, almost truncate, upper and lower lobes about equal, but upper lobe with a very short streamer in small juveniles.
</paragraph>
<paragraph id="B60F36E1040DFF92FF3EFD62A64FFBE8" blockId="9.[123,1174,219,1726]" pageId="9" pageNumber="10">
<emphasis id="84C4EAF3040DFF92FF3EFD62A7C0FD66" bold="true" box="[171,242,734,759]" pageId="9" pageNumber="10">
<emphasis id="84C4EAF3040DFF92FF3EFD62A7DBFD67" bold="true" box="[171,233,734,758]" italics="true" pageId="9" pageNumber="10">Color</emphasis>
:
</emphasis>
(
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) A series of about nine dark vertical bands along the light brown to reddish body, interrupted in some specimens by two or three white stripes; underside of head and belly mostly white; bands on body reduced to dark lateral blotches in some. Caudal and soft dorsal fins punctated with small bluish dots; in some individuals the basal one-third to one-half of soft dorsal fin dark but distally pale with small bluish spots; tips of caudal fin occasionally reddish (
<figureCitation id="2E8B2A64040DFF92FBD6FCE3A3B7FCE6" box="[1091,1157,863,887]" captionStart="FIGURE" captionStartId="8.[155,167,1217,1236]" captionTargetBox="[120,1176,773,1205]" captionTargetId="figure-51@8.[120,1176,772,1206]" captionTargetPageId="8" captionText="FIGURE 6. Serranus cabrilla. Adult from the Canary Islands. Photograph by Rogelio Herrera." figureDoi="http://doi.org/10.5281/zenodo.13156019" httpUri="https://zenodo.org/record/13156019/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
). Posterior margin of caudal fin occasionally blackish but never with black lobes as in
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<emphasis id="84C4EAF3040DFF92FB99FC3CA3BDFC09" box="[1036,1167,896,920]" italics="true" pageId="9" pageNumber="10">S. atricauda</emphasis>
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. Juveniles may be quite different: a white midlateral stripe bordered by two thick black stripes, the upper stripe running from tip of snout through middle of eye to upper margin of operculum onto trunk above midlateral line to upper half of caudal peduncle, the lower stripe from base of pectoral fin to lower half of caudal peduncle; the dark stripes often partially broken into dark blotches. The juvenile color pattern is often very similar to that of
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<emphasis id="84C4EAF3040DFF92FD26FB9DA405FBA8" box="[691,823,1057,1081]" italics="true" pageId="9" pageNumber="10">S. atricauda</emphasis>
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(compare with
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), although
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(CAS 234559, 44.450.
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SL) trawled in 53
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off
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lacked the prominent dark lateral stripes.
</paragraph>
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As in many other fish species,
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<emphasis id="84C4EAF3040DFF92FE62FB3EA555FB0B" box="[503,615,1154,1178]" italics="true" pageId="9" pageNumber="10">S. cabrilla</emphasis>
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specimens from deep water tend to be more yellow (
<bibRefCitation id="D2214B10040DFF92FF11FB1EA668FB2B" author="MEDIONI, E. &amp; R. LECOMTE FINIGER &amp; N. LOUVEIRO &amp; S. PLANES" box="[132,346,1186,1210]" pageId="9" pageNumber="10" pagination="113 - 1124" refId="ref20985" refString="MEDIONI, E., R. LECOMTE FINIGER, N. LOUVEIRO, AND S. PLANES. 2001. Genetic and demographic variation among color morphs of cabrilla seabass. Journal of Fish Biology 58: 113 - 1124." type="journal article" year="2001">Medioni et al. 2001</bibRefCitation>
). In one individual from deeper waters at the
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(
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), ground color golden-yellow on flanks, yellowish-orange on head, and brownish to tan on dorsum; four or five metallic-blue longitudinal lines or stripes on body and three diagonal stripes on head, the lowest stripe on head originating on snout and running to preopercle angle onto subopercle; the upper two stripes originating below orbit and terminating at margin of opercle. A fourth diagonal stripe present in some, originating on posterior portion of maxilla and extending a short distance onto preopercle. Fins yellowish; the soft dorsal with dark basal half, the dark bands on body extending onto dorsal-fin rays.
</paragraph>
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<emphasis id="84C4EAF3040DFF92FF3EFA18A7E5FA2D" bold="true" box="[171,215,1444,1468]" italics="true" pageId="9" pageNumber="10">Size</emphasis>
:
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TL.
</paragraph>
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.— Over hard bottoms from the shore to
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; from the British Isles to
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, including the Azores, Madeira, the Canary Islands,
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(based on photo by the second author) and
<collectingCountry id="CEA77671040DFF92FE71F9B8A5B4F98D" box="[484,646,1540,1564]" name="Principe Island" pageId="9" pageNumber="10">Príncipe Island</collectingCountry>
(first record based on CAS 234287), apparently here and only in deep water at the
<collectingCountry id="CEA77671040DFF92FE78F999A55BF9AD" box="[493,617,1572,1597]" name="Cape Verde" pageId="9" pageNumber="10">Cape Verde</collectingCountry>
Islands (Freitas et al., in prep.); also throughout the Mediterranean and into the Black Sea. Its presence in the Red Sea is attributed to invasion from the Mediterranean Sea after the Suez Canal was opened in 1869 (
<bibRefCitation id="D2214B10040DFF92FCAFF9D9A4E0F9EC" author="NORMAN, J. R." box="[826,978,1637,1661]" pageId="9" pageNumber="10" pagination="375 - 390" refId="ref21165" refString="NORMAN, J. R. 1927. Cambridge Expedition to the Suez Canal, 1924. Part 3. Report on the fishes. Transactions of the Zoological Society of London 22 (12): 375 - 390." type="journal article" year="1927">Norman 1927</bibRefCitation>
;
<bibRefCitation id="D2214B10040DFF92FC71F9D9A3BCF9EC" author="TORTONESE, E." box="[996,1166,1637,1661]" pageId="9" pageNumber="10" pagination="93 - 103" refId="ref21806" refString="TORTONESE, E. 1954. Zoogeography of the Mediterranean sea perches (Pisces Serranidae). Rapport Proc. Commission Internationale pour l'Exploration Scientific de la mer Mediterraneanee 12: 93 - 103." type="journal article" year="1954">Tortonese 1954</bibRefCitation>
; Meisler 1988:156). Records of the species from
<collectingCountry id="CEA77671040DFF92FD17F939A43EF90C" box="[642,780,1669,1693]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
are apparently of the closely similar
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<emphasis id="84C4EAF3040DFF92FFEEF91AA627F92C" box="[123,277,1701,1725]" italics="true" pageId="9" pageNumber="10">S. knysnaensis</emphasis>
Gilchrist, 1904
</taxonomicName>
(Heemstra and Anderson, 2016).
</paragraph>
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(
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).—
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<emphasis id="84C4EAF3040EFF91FCBBFF67A405FF62" bold="true" box="[814,823,219,243]" pageId="10" pageNumber="11">:</emphasis>
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(
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SL);
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, purchased dockside in
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from fisherman selling catch taken off
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;
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.
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(3, 44.450.
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SL); 15°19.1ʹN, 16°55.5ʹW, 53
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;
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2012, sta. 98,
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.
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(
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SL); 5°54.77ʹN, 1°16.15ʹE;
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Survey
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(
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SL); collector
<collectorName id="1B455337040EFF91FD7AFEC2A468FE07" box="[751,858,382,406]" pageId="10" pageNumber="11">A.I. Good</collectorName>
; 1940.
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<collectingCountry id="CEA77671040EFF91FC3FFEC2A4EDFE07" box="[938,991,382,406]" name="Italy" pageId="10" pageNumber="11">Italy</collectingCountry>
</emphasis>
</materialsCitation>
:
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<specimenCode id="E6169E9A040EFF91FC78FEC2A3A4FE07" box="[1005,1174,382,406]" collectionCode="CAS-SU" country="0" httpUri="http://grbio.org/cool/fxd1-0prj" name="California Academy of Sciences, Stanford University Collection" pageId="10" pageNumber="11">CAS-SU 20933</specimenCode>
(3, 142-
<quantity id="71489B04040EFF91FF47FE23A61FFE27" box="[210,301,415,439]" metricMagnitude="-1" metricUnit="m" metricValue="2.16" pageId="10" pageNumber="11" unit="mm" value="216.0">216 mm</quantity>
SL); and
</materialsCitation>
<materialsCitation id="06D83CBC040EFF91FE34FE23A7EAFE46" collectorName="E. C. Starks" country="Italy" location="Naples" pageId="10" pageNumber="11" specimenCode="CAS-SU 20933" specimenCount="1">
<specimenCode id="E6169E9A040EFF91FE34FE23A57EFE26" box="[417,588,415,439]" collectionCode="CAS-SU" country="0" httpUri="http://grbio.org/cool/fxd1-0prj" name="California Academy of Sciences, Stanford University Collection" pageId="10" pageNumber="11">CAS-SU 20933</specimenCode>
(3, 142
<quantity id="71489B04040EFF91FD27FE23A43EFE27" box="[690,780,415,439]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" pageId="10" pageNumber="11" unit="mm" value="175.0">175 mm</quantity>
SL);
<location id="B36F603A040EFF91FCC5FE23A4A8FE26" LSID="urn:lsid:plazi:treatment:3E1987F70403FF91FFEEFBF4A5E7FDA8:B36F603A040EFF91FCC5FE23A4A8FE26" box="[848,922,415,439]" country="Italy" name="Naples" pageId="10" pageNumber="11">Naples</location>
; collector
<collectorName id="1B455337040EFF91FB87FE23A3BCFE26" box="[1042,1166,415,439]" pageId="10" pageNumber="11">E.C. Starks</collectorName>
; ca. 1907
</materialsCitation>
.
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<specimenCode id="E6169E9A040EFF91FF7DFE03A6B2FE46" box="[232,384,447,471]" collectionCode="CAS-IU" country="0" httpUri="http://grbio.org/cool/tb6w-1bpn" name="California Academy of Science, Indiana University Collection" pageId="10" pageNumber="11">CAS-IU 7545</specimenCode>
(
<quantity id="71489B04040EFF91FE06FE03A539FE46" box="[403,523,447,471]" metricMagnitude="0" metricUnit="m" metricValue="1.162" pageId="10" pageNumber="11" unit="mm" value="1162.0">1, 162 mm</quantity>
SL);
<collectingRegion id="7474F803040EFF91FDDBFE03A5BEFE46" box="[590,652,447,471]" country="Italy" name="Sicilia" pageId="10" pageNumber="11">Sicily</collectingRegion>
,
<location id="B36F603A040EFF91FD08FE03A5C4FE46" LSID="urn:lsid:plazi:treatment:3E1987F70403FF91FFEEFBF4A5E7FDA8:B36F603A040EFF91FD08FE03A5C4FE46" box="[669,758,447,471]" country="Spain" name="Palermo" pageId="10" pageNumber="11" stateProvince="Sicilia">Palermo</location>
; collector
<collectorName id="1B455337040EFF91FCFBFE7CA4C7FE46" box="[878,1013,447,471]" pageId="10" pageNumber="11">P. Doderlein</collectorName>
; 1886.
<collectingCountry id="CEA77671040EFF91FBDEFE03A3BEFE46" box="[1099,1164,447,471]" name="Spain" pageId="10" pageNumber="11">
<emphasis id="84C4EAF3040EFF91FBDEFE03A3BEFE46" bold="true" box="[1099,1164,447,471]" pageId="10" pageNumber="11">Spain</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="84C4EAF3040EFF91FB19FE03A3A7FE46" bold="true" box="[1164,1173,447,471]" pageId="10" pageNumber="11">:</emphasis>
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<specimenCode id="E6169E9A040EFF91FFEEFE5CA614FE69" box="[123,294,480,504]" collectionCode="CAS-SU" country="0" httpUri="http://grbio.org/cool/fxd1-0prj" name="California Academy of Sciences, Stanford University Collection" pageId="10" pageNumber="11">CAS-SU 69064</specimenCode>
(
<quantity id="71489B04040EFF91FEA2FE5CA69CFE66" box="[311,430,480,504]" metricMagnitude="0" metricUnit="m" metricValue="1.1709999999999998" pageId="10" pageNumber="11" unit="mm" value="1171.0">1, 171 mm</quantity>
SL);
<location id="B36F603A040EFF91FE64FE5CA58EFE69" LSID="urn:lsid:plazi:treatment:3E1987F70403FF91FFEEFBF4A5E7FDA8:B36F603A040EFF91FE64FE5CA58EFE69" box="[497,700,480,504]" country="Spain" name="Mediterranean Sea" pageId="10" pageNumber="11" stateProvince="Sicilia">Mediterranean Sea</location>
;
<collectorName id="1B455337040EFF91FD59FE5CA419FE69" box="[716,811,480,504]" pageId="10" pageNumber="11">Gibralter</collectorName>
,
<collectorName id="1B455337040EFF91FCAEFE5CA4C2FE69" box="[827,1008,480,504]" pageId="10" pageNumber="11">Gibralter Harbor</collectorName>
; collector
</materialsCitation>
<materialsCitation id="06D83CBC040EFF91FBF3FE5CA6F4FD89" collectingDate="1947-08-12" collectionCode="T" collectorName="H. Work &amp; Red Sea" country="Spain" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Sicilia">
<collectionCode id="D0A1AE24040EFF91FBF3FE5CA346FE66" box="[1126,1140,480,503]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="11">T</collectionCode>
.
<collectorName id="1B455337040EFF91FBEEFE5CA784FD89" pageId="10" pageNumber="11">H. Work</collectorName>
;
<date id="C20E1021040EFF91FF50FDBDA66BFD88" box="[197,345,513,537]" pageId="10" pageNumber="11" value="1947-08-12">
<collectingDate id="D24AE9C9040EFF91FF50FDBDA66BFD88" box="[197,345,513,537]" pageId="10" pageNumber="11" value="1947-08-12">12 Aug. 1947</collectingDate>
</date>
.
<collectorName id="1B455337040EFF91FEFDFDBDA6F4FD89" box="[360,454,512,537]" pageId="10" pageNumber="11">
<emphasis id="84C4EAF3040EFF91FEFDFDBDA6F4FD89" bold="true" box="[360,454,512,537]" pageId="10" pageNumber="11">Red Sea</emphasis>
</collectorName>
</materialsCitation>
<emphasis id="84C4EAF3040EFF91FE53FDBDA6FDFD88" bold="true" box="[454,463,513,537]" pageId="10" pageNumber="11">:</emphasis>
<materialsCitation id="06D83CBC040EFF91FE42FDBDA5FCFDA8" collectingDate="1967-09-22" collectionCode="USNM" collectorName="Egypt &amp; H. Steinitz" country="Spain" location="Abu Zenima" pageId="10" pageNumber="11" specimenCode="USNM 203668" specimenCount="1" stateProvince="Sicilia">
<specimenCode id="E6169E9A040EFF91FE42FDBDA54FFD89" box="[471,637,512,537]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM 203668</specimenCode>
(4,
<quantity id="71489B04040EFF91FD3FFDBCA42CFD89" box="[682,798,512,537]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" metricValueMax="9.8" metricValueMin="6.0" pageId="10" pageNumber="11" unit="mm" value="79.0" valueMax="98.0" valueMin="60.0">6098 mm</quantity>
SL); “Abu Zueima” [
<emphasis id="84C4EAF3040EFF91FB85FDBDA31CFD89" box="[1040,1070,513,536]" italics="true" pageId="10" pageNumber="11">sic</emphasis>
, possibly
<location id="B36F603A040EFF91FFEEFD9DA633FDA8" LSID="urn:lsid:plazi:treatment:3E1987F70403FF91FFEEFBF4A5E7FDA8:B36F603A040EFF91FFEEFD9DA633FDA8" box="[123,257,545,569]" country="Spain" name="Abu Zenima" pageId="10" pageNumber="11" stateProvince="Sicilia">Abu Zenima</location>
, in
<collectorName id="1B455337040EFF91FEBEFD9EA659FDAB" box="[299,363,546,570]" pageId="10" pageNumber="11">Egypt</collectorName>
]; coll.
<collectorName id="1B455337040EFF91FE2CFD9EA519FDA8" box="[441,555,545,569]" pageId="10" pageNumber="11">H. Steinitz</collectorName>
,
<date id="C20E1021040EFF91FDACFD9DA5FCFDA8" box="[569,718,545,569]" pageId="10" pageNumber="11" value="1967-09-22">
<collectingDate id="D24AE9C9040EFF91FDACFD9DA5FCFDA8" box="[569,718,545,569]" pageId="10" pageNumber="11" value="1967-09-22">22 Sept. 1967</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,181 @@
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<mods:title id="5BF68AA846C7245F09E1E5546B932637">A Synopsis of the Eastern and Central Atlantic Combers of the Genus Serranus (Teleostei: Scorpaeniformes: Serranidae)</mods:title>
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<subSubSection id="FEAA656A0406FF99FE77FDE2A41FFDE7" box="[482,813,606,630]" pageId="2" pageNumber="3" type="nomenclature">
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<emphasis id="84C4EAF30406FF99FE77FDE2A41FFDE7" bold="true" box="[482,813,606,630]" pageId="2" pageNumber="3">
Genus
<taxonomicName id="71B04D620406FF99FDA7FDE2A41FFDE7" ID-CoL="7FWT" authority="Cuvier, 1817" authorityName="Cuvier" authorityYear="1817" box="[562,813,606,630]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30406FF99FDA7FDE2A5A5FDE7" bold="true" box="[562,663,606,630]" italics="true" pageId="2" pageNumber="3">Serranus</emphasis>
Cuvier, 1817
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="B60F36E10406FF99FF2DFD36A36AFD0E" blockId="2.[184,1112,650,672]" box="[184,1112,650,672]" pageId="2" pageNumber="3">
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<typeStatus id="690B88430406FF99FF2DFD36A7DAFD0E" box="[184,232,650,671]" pageId="2" pageNumber="3">Type</typeStatus>
species
<taxonomicName id="71B04D620406FF99FEAFFD36A552FD31" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[314,608,650,672]" family="Percidae" genus="Perca" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cabrilla">
<emphasis id="84C4EAF30406FF99FEAFFD36A6F7FD0E" box="[314,453,650,671]" italics="true" pageId="2" pageNumber="3">Perca cabrilla</emphasis>
Linnaeus, 1758
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(as designated by
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, Official List, Opinion 93)
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.
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<subSubSection id="FEAA656A0406FF99FF3EFD05A33CFC65" pageId="2" pageNumber="3" type="diagnosis">
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.— D X,1215; A III,68, usually 7; P 1218; V I,5; BR 7; vertebrae 10+14. Dorsal fin undivided, spinous and soft rays broadly united; caudal fin truncate, emarginate, or moderately forked, with 17 principal rays (15 branched); supramaxilla absent; teeth on vomer and palatines, none on tongue. Opercle with two or (usually) three flattened spines, the lowermost sometimes obscure or undeveloped; central spine largest and directed horizontally. A spinousedged suprascapular scale present. Scales mostly ctenoid, but some species have cycloid scales on head or trunk and
<taxonomicName id="71B04D620406FF99FED5FCC0A69CFC02" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[320,430,891,915]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="drewesi" status="n. sp.">
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</taxonomicName>
<taxonomicNameLabel id="9FF757880406FF99FE20FCC0A6DDFC02" box="[437,495,892,915]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
has all scales cycloid; interorbit, occiput, upper part of postorbital region and interopercle variously scaled or naked; lateral line complete, with pored scales; maxilla naked. Species probably all synchronous hermaphrodites. Most species small, less than
<quantity id="71489B040406FF99FBEEFC00A7AEFC65" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="cm" value="20.0">20 cm</quantity>
TL, but a few attain about
<quantity id="71489B040406FF99FE57FC61A537FC65" box="[450,517,989,1013]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="3" unit="cm" value="40.0">40 cm</quantity>
TL (mostly from Robins and Starck, 1961:261).
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0406FF98FF3EFC41A3A4FEA2" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
<paragraph id="B60F36E10406FF99FF3EFC41A47CFAA9" blockId="2.[123,1174,697,1726]" pageId="2" pageNumber="3">
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<emphasis id="84C4EAF30406FF99FF3EFC41A624FB82" bold="true" box="[171,278,1021,1045]" pageId="2" pageNumber="3">REMARKS</emphasis>
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.— Members of the genus are found on both sides of the Atlantic (including the Mediterranean and Black Seas, and in the Red Sea as an invasive), in the tropical eastern Pacific, and in the southwestern Indian Ocean off
<collectingCountry id="CEA776710406FF99FDA9FB82A5F1FBC7" box="[572,707,1086,1110]" name="South Africa" pageId="2" pageNumber="3">South Africa</collectingCountry>
. The highest diversity is found in the western central Atlantic (14 spp.), followed by the eastern Atlantic (10 spp.), the eastern Pacific (6 spp.), and the Indian Ocean (2 spp.). The species are apparently endemic to each region, as no species is found in more than one region.
<bibRefCitation id="D2214B100406FF99FE41FB23A5BBFB26" author="KUITER, R." box="[468,649,1183,1207]" pageId="2" pageNumber="3" refId="ref20896" refString="KUITER, R. 2004. Serranidae and Plesiopidae. A Comprehensive Guide to Basslets, Hamlets, Longfins and Relatives. Aquatic Photographics, Seaford, Australia. 216 pp." type="book" year="2004">Kuiter (2004:80)</bibRefCitation>
listed 30 described species of
<taxonomicName id="71B04D620406FF99FC42FB23A305FB27" authorityName="Cuvier" authorityYear="1817" box="[983,1079,1183,1206]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30406FF99FC42FB23A305FB27" box="[983,1079,1183,1206]" italics="true" pageId="2" pageNumber="3">Serranus</emphasis>
</taxonomicName>
and two undescribed species, but one of the described species he listed is
<taxonomicName id="71B04D620406FF99FCA8FB03A782FB69" authority="Cadenat, 1960" authorityName="Cadenat" authorityYear="1960" family="Serranidae" genus="Chelidoperca" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="africana">
<emphasis id="84C4EAF30406FF99FCA8FB03A31DFB46" box="[829,1071,1215,1239]" italics="true" pageId="2" pageNumber="3">Chelidoperca africana</emphasis>
Cadenat, 1960
</taxonomicName>
, which we consider as the only member of
<taxonomicName id="71B04D620406FF99FD15FB5CA422FB69" authorityName="Boulenger" authorityYear="1895" box="[640,784,1248,1272]" family="Serranidae" genus="Chelidoperca" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30406FF99FD15FB5CA422FB69" box="[640,784,1248,1272]" italics="true" pageId="2" pageNumber="3">Chelidoperca</emphasis>
</taxonomicName>
in the Atlantic.
<taxonomicName id="71B04D620406FF99FC2EFB5CA379FB69" authorityName="Boulenger" authorityYear="1895" box="[955,1099,1248,1272]" family="Serranidae" genus="Chelidoperca" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30406FF99FC2EFB5CA379FB69" box="[955,1099,1248,1272]" italics="true" pageId="2" pageNumber="3">Chelidoperca</emphasis>
</taxonomicName>
is otherwise known from 10 other species of the Indo-West Pacific (
<bibRefCitation id="D2214B100406FF99FC9AFABDA333FA89" author="MATSUNUMA, M. &amp; T. YAMAKAWA &amp; J. T. WILLIAMS" box="[783,1025,1280,1304]" pageId="2" pageNumber="3" pagination="210 - 30" refId="ref20928" refString="MATSUNUMA, M., T. YAMAKAWA, AND J. T. WILLIAMS. 2018. Chelidoperca tosaensis, a new species of perchlet (Serranidae) from Japan and the Philippines, with geographic range extension of C. stella to the northwestern Pacific Ocean. Ichthyological Research 65: 210 - 30." type="journal article" year="2018">Matsunuma et al. 2018</bibRefCitation>
). Table 1 lists 32 species of
<taxonomicName id="71B04D620406FF99FE9AFA9DA65DFAA9" authorityName="Cuvier" authorityYear="1817" box="[271,367,1313,1336]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30406FF99FE9AFA9DA65DFAA9" box="[271,367,1313,1336]" italics="true" pageId="2" pageNumber="3">Serranus</emphasis>
</taxonomicName>
, based on Kuiter (loc. cit.) and our findings.
</paragraph>
<paragraph id="B60F36E10406FF99FF3EFAFDA622F9ED" blockId="2.[123,1174,697,1726]" pageId="2" pageNumber="3">
Robins and Starck (1961) comprehensively treated the western Atlantic species of
<taxonomicName id="71B04D620406FF99FBA0FAFDA3A7FAC9" authorityName="Cuvier" authorityYear="1817" box="[1077,1173,1345,1368]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30406FF99FBA0FAFDA3A7FAC9" box="[1077,1173,1345,1368]" italics="true" pageId="2" pageNumber="3">Serranus</emphasis>
</taxonomicName>
and provided valuable additional information on the genus, its included species, and relationships to other serranines; they also gave excellent descriptions of three eastern Atlantic species for which they had specimens. They considered that the anal fin-ray count of III,7 was the common number, but we found that our
<specimenCount id="A0B6FD680406FF99FEE3FA7FA524FA4A" box="[374,534,1475,1499]" count="2" pageId="2" pageNumber="3" type="generic">two specimens</specimenCount>
of
<taxonomicName id="71B04D620406FF99FDD5FA7FA5F7FA4A" baseAuthorityName="Gunther" baseAuthorityYear="1874" box="[576,709,1475,1499]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="atricauda">
<emphasis id="84C4EAF30406FF99FDD5FA7FA5F7FA4A" box="[576,709,1475,1499]" italics="true" pageId="2" pageNumber="3">S. atricauda</emphasis>
</taxonomicName>
had III,8, as did one of our specimens of
<taxonomicName id="71B04D620406FF99FFEEFA5FA7EEFA6A" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[123,220,1507,1531]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="scriba">
<emphasis id="84C4EAF30406FF99FFEEFA5FA7EEFA6A" box="[123,220,1507,1531]" italics="true" pageId="2" pageNumber="3">S. scriba</emphasis>
</taxonomicName>
(CAS-SU 20897).
<bibRefCitation id="D2214B100406FF99FE23FA5FA5BBFA6A" author="MEISLER, M. R." box="[438,649,1507,1531]" pageId="2" pageNumber="3" refId="ref21027" refString="MEISLER, M. R. 1987. Limits and relationships of serranine seabasses, with revisions of Serranus and Mentiperca (Pisces: Serranidae). Unpublished Ph. D. thesis. University of Southern California, Los Angeles, California, USA. 250 pages." type="book" year="1987">Meisler (1987:152)</bibRefCitation>
stated that,
<taxonomicName id="71B04D620406FF99FC83FA5FA4AFFA6A" baseAuthorityName="Gunther" baseAuthorityYear="1874" box="[790,925,1507,1531]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="atricauda">
<emphasis id="84C4EAF30406FF99FC83FA5FA4AFFA6A" box="[790,925,1507,1531]" italics="true" pageId="2" pageNumber="3">S. atricauda</emphasis>
</taxonomicName>
“is the only
<taxonomicName id="71B04D620406FF99FBA0FA5FA7FAF98A" pageId="2" pageNumber="3">
<emphasis id="84C4EAF30406FF99FBA0FA5FA3A7FA6B" box="[1077,1173,1507,1530]" italics="true" pageId="2" pageNumber="3">Serranus</emphasis>
species
</taxonomicName>
to have exclusively eight anal rays.“ Heemstra and Anderson (2016:2410) gave the anal fin-ray count for
<taxonomicName id="71B04D620406FF99FEA7F998A692F9AD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[306,416,1572,1596]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30406FF99FEA7F998A692F9AD" box="[306,416,1572,1596]" italics="true" pageId="2" pageNumber="3">S. hepatus</emphasis>
</taxonomicName>
as III,6 or 7, but seven of our
<specimenCount id="A0B6FD680406FF99FD71F998A446F9AD" box="[740,884,1572,1596]" count="10" pageId="2" pageNumber="3" type="generic">10 specimens</specimenCount>
of that species had a count of III,7; the other had III,6. The
<typeStatus id="690B88430406FF99FE46F9F8A502F9CD" box="[467,560,1604,1628]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
and only specimen of
<taxonomicName id="71B04D620406FF99FCB6F9F9A4BDF9CD" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[803,911,1604,1628]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="drewesi" status="n.sp.">
<emphasis id="84C4EAF30406FF99FCB6F9F9A4BDF9CD" box="[803,911,1604,1628]" italics="true" pageId="2" pageNumber="3">S. drewesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF757880406FF99FC03F9F9A4FBF9CD" box="[918,969,1605,1628]" pageId="2" pageNumber="3" rank="species">n.sp.</taxonomicNameLabel>
has an anal fin-ray count of III,6.
</paragraph>
<paragraph id="B60F36E10406FF98FF3EF93AA3A4FEA2" blockId="2.[123,1174,697,1726]" lastBlockId="3.[123,1174,219,307]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
It is regrettable that Martin R. Meislers Ph.D. dissertation (1987) was never published inasmuch as it is a comprehensive revisionary work on the group, based on study of most of the species of
<taxonomicName id="71B04D620407FF98FF7FFF67A678FF63" authorityName="Cuvier" authorityYear="1817" box="[234,330,219,242]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30407FF98FF7FFF67A678FF63" box="[234,330,219,242]" italics="true" pageId="3" pageNumber="4">Serranus</emphasis>
</taxonomicName>
and
<emphasis id="84C4EAF30407FF98FEE9FF60A6C4FF62" box="[380,502,220,243]" italics="true" pageId="3" pageNumber="4">Mentiperca</emphasis>
(as well as numerous other serranid genera) and a phylogenetic analysis using a variety of osteological, myological, and external characters. It contains a wealth of original information not elsewhere available and the bibliography is comprehensive to that time.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,287 @@
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<mods:title id="5BFE77981FAC7B133BBFBE18267F6B88">A Synopsis of the Eastern and Central Atlantic Combers of the Genus Serranus (Teleostei: Scorpaeniformes: Serranidae)</mods:title>
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<paragraph id="B60F36E1040EFF91FFEEFDDEA5D3FDEB" blockId="10.[123,737,610,665]" box="[123,737,610,634]" pageId="10" pageNumber="11">
<heading id="ED47818D040EFF91FFEEFDDEA5D3FDEB" box="[123,737,610,634]" fontSize="10" level="2" pageId="10" pageNumber="11" reason="4">
<emphasis id="84C4EAF3040EFF91FFEEFDDEA5D3FDEB" bold="true" box="[123,737,610,634]" pageId="10" pageNumber="11">
<taxonomicName id="71B04D62040EFF91FFEEFDDEA54FFDEB" authority="Iwamoto" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[123,637,610,634]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="drewesi" status="sp. nov." subGenus="Paracentropristis">
<emphasis id="84C4EAF3040EFF91FFEEFDDEA7D2FDEB" bold="true" box="[123,224,610,634]" italics="true" pageId="10" pageNumber="11">Serranus</emphasis>
(
<emphasis id="84C4EAF3040EFF91FF65FDDEA69CFDEB" bold="true" box="[240,430,610,634]" italics="true" pageId="10" pageNumber="11">Paracentropristis</emphasis>
)
<emphasis id="84C4EAF3040EFF91FE2BFDDEA523FDEB" bold="true" box="[446,529,610,634]" italics="true" pageId="10" pageNumber="11">drewesi</emphasis>
Iwamoto
</taxonomicName>
,
<taxonomicNameLabel id="9FF75788040EFF91FD1EFDDEA5D3FDEB" box="[651,737,610,634]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A040EFF91FFEEFD3FA7FEFD09" box="[123,204,643,665]" pageId="10" pageNumber="11" type="description">
<paragraph id="B60F36E1040EFF91FFEEFD3FA7FEFD09" blockId="10.[123,737,610,665]" box="[123,204,643,665]" pageId="10" pageNumber="11">
<figureCitation id="2E8B2A64040EFF91FFEEFD3FA7FEFD09" box="[123,204,643,665]" captionStart="FIGURE" captionStartId="11.[155,167,1550,1569]" captionTargetBox="[120,1176,993,1535]" captionTargetId="figure-2176@11.[120,1176,991,1536]" captionTargetPageId="11" captionText="FIGURE 8. Serranus drewesi Iwamoto, new species. Holotype (CAS 234050 (53.5 mm SL) from submarine cave on Santana Islet, São Tomé Island. Photograph by David Catania." figureDoi="http://doi.org/10.5281/zenodo.13156025" httpUri="https://zenodo.org/record/13156025/files/figure.png" pageId="10" pageNumber="11">Figure 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A040EFF91FF3EFD0EA6C9FC5E" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="B60F36E1040EFF91FF3EFD0EA6C9FC5E" blockId="10.[123,1174,690,1725]" pageId="10" pageNumber="11">
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<emphasis id="84C4EAF3040EFF91FF3EFD0EA612FD58" bold="true" box="[171,288,690,714]" pageId="10" pageNumber="11">DIAGNOSIS</emphasis>
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.— D X,11; P 14; A III,6; gill rakers 3+11; scales below 1D origin about 5, below 2D 4+1+8 or 9, pored lateral-line scales 44; circumpeduncular scales 1+9+1+10 (21 total). Anteri- or nostril tubular, rim greatly elevated posteriorly and bearing about 7 or 8 long cirri at tip. Scales all cycloid (except for a hard, sharply spinulated suprascapular scale bone above the upper opening of the gill cover). Caudal fin weakly emarginate, upper and lower lobes about equal. Interradial membranes of soft dorsal and anal fins scaly on basal one-half to one-third. Nape, opercle, preopercle, interopercle, mandibular ramus, pectoral-fin base, and chest covered with cycloid scales. A bold black spot between last three spines and first soft dorsal ray formed above broad black saddle on dorsal half of body.
</paragraph>
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<emphasis id="84C4EAF3040EFF91FF3EFC64A608FC7F" bold="true" box="[171,314,984,1008]" pageId="10" pageNumber="11">DESCRIPTION</emphasis>
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.— Head and body laterally compressed, greatest width of head 6.3 times in SL, greatest body width 5.9 times in SL. Snout pointed, longer than orbit diameter; interorbital space narrow, bony width about 60% of greatest orbit diameter. Orbit about
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HL; pupil sharply eggshaped with pronounced anterior aphakic (lensless) space. Jaws equal; posterior margin of maxilla below middle of orbit. Two broad, flat, weak opercular spines developed, the dorsal spine partially obscured by skin and scales, lower spine more prominent and forming posterodorsal corner of gill cover. Anterior nostril tubular, rim low anteriorly but elevated into a long flap posteriorly, the flap fringed with 7 or 8 long cirri, a few filaments longer than height of flap. Gill opening ventrally extending forward to about vertical passing through hind edge of orbit. Gill rakers relatively short, stout and recurved, the uppermost on lower limb (ceratobranchial) about
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in length, the succeeding rakers gradually shorter with last three or so tubercular. About 11 pseudobranchs.
</paragraph>
<paragraph id="B60F36E1040EFF91FF3EFAE3A57DF9AB" blockId="10.[123,1174,690,1725]" pageId="10" pageNumber="11">Spinous dorsal fin with middle spines longest, a prominent dip between spinous and soft-rayed portions of dorsal fin; the longest dorsal spine shorter than longest dorsal soft rays. Size and shape of pectoral fin could not be determined owing to left fin torn off in capture and right fin cut off for tissue sample. Pelvic fin below fourth spine of dorsal fin and behind origin of pectoral fin, its tip when depressed not extending to origin of anal fin. Anal fin with three strong spines, the first about half length of second, which is slightly longer than third spine; six soft rays forming narrow round- ed tip to fin. Caudal fin weakly emarginate.</paragraph>
<paragraph id="B60F36E1040EFF90FF3EF9F8A6D2FE03" blockId="10.[123,1174,690,1725]" lastBlockId="11.[122,1174,219,979]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">Scales all cycloid (excepting a hard, sharply spinulated suprascapular scale bone above the upper opening of the gill cover). Snout and dorsal surfaces of head naked anterior to a vertical passing through hind margin of preopercle. Interopercle finely scaled anteriorly and dorsally immediately below preopercular margin. Mandibular ramus with tiny thin scales covering most of posterior surfaces and extending forward onto lateral surfaces. Cheeks (viz, preopercle) finely scaled; hind (vertical) margin of preopercle finely serrated, but ventral margin smooth. Opercle and subopercle with large scales in two major rows bordered posteriorly with small scales. Chest and nape fully scaled. Eight scale rows along dorsal crest of nape before origin of first dorsal fin. Small scales at base of second dorsal fin (and sparsely on posterior base of spinous dorsal fin), and on bases of pectoral and caudal fins.</paragraph>
<paragraph id="B60F36E1040FFF90FF3FFE27A4D1FDE3" blockId="11.[122,1174,219,979]" pageId="11" pageNumber="12">Anterior tip of premaxillary with cluster of small teeth and one enlarged canine more laterally placed; the cluster of teeth followed by a tapered band of small teeth. Dentary teeth consist of a cluster of small, short, conical teeth anteriorly that are rigid along outer margin but depressible inward of outer row. Posterior to these, a series of four larger canine-like teeth, with an outer small- er tooth row, these in turn followed posteriorly by a row of smaller teeth. Vomerine teeth small, in a broad short band. Presence of palatine teeth could not be adequately assessed for fear of damaging specimen; however, there appeared to be a row of small teeth along the bone.</paragraph>
<paragraph id="B60F36E1040FFF90FF3FFDC6A573FC43" blockId="11.[122,1174,219,979]" pageId="11" pageNumber="12">
<emphasis id="84C4EAF3040FFF90FF3FFDC6A7DAFD03" bold="true" box="[170,232,634,658]" italics="true" pageId="11" pageNumber="12">Color</emphasis>
(fresh): (
<figureCitation id="2E8B2A64040FFF90FEC5FDC7A6BDFD02" box="[336,399,635,659]" captionStart="FIGURE" captionStartId="11.[155,167,1550,1569]" captionTargetBox="[120,1176,993,1535]" captionTargetId="figure-2176@11.[120,1176,991,1536]" captionTargetPageId="11" captionText="FIGURE 8. Serranus drewesi Iwamoto, new species. Holotype (CAS 234050 (53.5 mm SL) from submarine cave on Santana Islet, São Tomé Island. Photograph by David Catania." figureDoi="http://doi.org/10.5281/zenodo.13156025" httpUri="https://zenodo.org/record/13156025/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
) Head with five or six small black spots behind eye, two in interorbital space, one on snout on median line, and diffuse ill-defined spots on sides of snout and head. Dorsal half of body and head darker than ventral half; nape and dorsum posteriorly to below third dorsal spine dark brown; a broad dark saddle below last three spines and first two soft rays of dorsal fin; a second dark saddle below 5
<superScript id="41C59BA9040FFF90FD82FD46A514FC9A" attach="left" box="[535,550,762,779]" fontSize="7" pageId="11" pageNumber="12">th</superScript>
through 9
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soft rays, followed by third saddle (partially merged with second) below last two soft rays and anterior one-third or so of caudal peduncle, a fourth saddle on posterior third of caudal peduncle ending sharply at base of caudal fin. Spinous dorsal fin irregularly brown with faint pale blotches, a bold black spot between posterior part of spinous dorsal and anteriormost end of soft dorsal fin; pectoral fin missing on both sides; pelvic fins pale to light yellow; anal fin dark brown overall; caudal fin mostly orangish to pink with darker upper and lower edges. Iris orange.
</paragraph>
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<paragraph id="B60F36E1040FFF90FF0EF9B2A34AF9B0" blockId="11.[123,1144,1550,1596]" box="[155,1144,1550,1569]" pageId="11" pageNumber="12">
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8.
<taxonomicName id="71B04D62040FFF90FF61F9B2A6E8F9B0" authority="Iwamoto" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[244,474,1550,1569]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="drewesi" status="sp. nov.">
<emphasis id="84C4EAF3040FFF90FF61F9B2A6BBF9B0" box="[244,393,1550,1569]" italics="true" pageId="11" pageNumber="12">Serranus drewesi</emphasis>
Iwamoto
</taxonomicName>
,
<taxonomicNameLabel id="9FF75788040FFF90FE70F9B2A57FF9B0" box="[485,589,1550,1569]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
. Holotype (CAS 234050 (53.5 mm SL) from submarine cave on
</paragraph>
<paragraph id="B60F36E1040FFF90FFEEF995A5A0F9AD" blockId="11.[123,1144,1550,1596]" box="[123,658,1576,1596]" pageId="11" pageNumber="12">Santana Islet, São Tomé Island. Photograph by David Catania.</paragraph>
</caption>
<paragraph id="B60F36E1040FFF97FF3FF9E7A3A4FDA8" blockId="11.[122,1173,1627,1715]" lastBlockId="12.[123,1174,219,1321]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="84C4EAF3040FFF90FF3FF9E7A677F9E2" bold="true" box="[170,325,1627,1651]" pageId="11" pageNumber="12">
<emphasis id="84C4EAF3040FFF90FF3FF9E7A609F9E2" bold="true" box="[170,315,1627,1651]" italics="true" pageId="11" pageNumber="12">Comparisons</emphasis>
:
</emphasis>
<taxonomicName id="71B04D62040FFF90FED9F9E7A537F9E2" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[332,517,1627,1651]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="drewesi">
<emphasis id="84C4EAF3040FFF90FED9F9E7A537F9E2" box="[332,517,1627,1651]" italics="true" pageId="11" pageNumber="12">Serranus drewesi</emphasis>
</taxonomicName>
is most similar in overall characters to
<taxonomicName id="71B04D62040FFF90FC38F9E7A329F9E2" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[941,1051,1627,1651]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF3040FFF90FC38F9E7A329F9E2" box="[941,1051,1627,1651]" italics="true" pageId="11" pageNumber="12">S. hepatus</emphasis>
</taxonomicName>
, notably in having six anal fin rays and closely similar fin ray counts, somewhat similar body shape, and general color pattern, including a prominent black spot on the dorsal fin. That spot, however, is not ocellated as in
<taxonomicName id="71B04D620408FF97FE8BFF67A6BFFF62" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[286,397,219,243]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30408FF97FE8BFF67A6BFFF62" box="[286,397,219,243]" italics="true" pageId="12" pageNumber="13">S. hepatus</emphasis>
</taxonomicName>
and it merges with the broad saddle on the dorsum immediately below. The saddles on the body do not extend to the ventral body margin and do not sharply contrast with a generally pale body as in
<taxonomicName id="71B04D620408FF97FE0EFEA1A53BFEA5" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[411,521,284,308]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30408FF97FE0EFEA1A53BFEA5" box="[411,521,284,308]" italics="true" pageId="12" pageNumber="13">S. hepatus</emphasis>
</taxonomicName>
. The pelvic fin is overall yellowish (dusky in preserved specimen) in contrast to black in
<taxonomicName id="71B04D620408FF97FE50FE81A505FEC4" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[453,567,317,341]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30408FF97FE50FE81A505FEC4" box="[453,567,317,341]" italics="true" pageId="12" pageNumber="13">S. hepatus</emphasis>
</taxonomicName>
and the anal fin is uniformly dark brown without the pronounced black anterior and clear distal parts so notable in
<taxonomicName id="71B04D620408FF97FCA6FEE2A497FEE7" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[819,933,350,374]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30408FF97FCA6FEE2A497FEE7" box="[819,933,350,374]" italics="true" pageId="12" pageNumber="13">S. hepatus</emphasis>
</taxonomicName>
. Other differentiating features include: the absence in
<taxonomicName id="71B04D620408FF97FE40FEC3A571FE07" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[469,579,382,406]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="drewesi">
<emphasis id="84C4EAF30408FF97FE40FEC3A571FE07" box="[469,579,382,406]" italics="true" pageId="12" pageNumber="13">S. drewesi</emphasis>
</taxonomicName>
of narrow gold to orange stripes on the head (vs. present in
<taxonomicName id="71B04D620408FF97FF29FE23A61BFE26" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[188,297,415,439]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30408FF97FF29FE23A61BFE26" box="[188,297,415,439]" italics="true" pageId="12" pageNumber="13">S. hepatus</emphasis>
</taxonomicName>
); black spots on the head (lacking in
<taxonomicName id="71B04D620408FF97FD21FE23A413FE26" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[692,801,415,439]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hepatus">
<emphasis id="84C4EAF30408FF97FD21FE23A413FE26" box="[692,801,415,439]" italics="true" pageId="12" pageNumber="13">S. hepatus</emphasis>
</taxonomicName>
), two opercular spines in
<taxonomicName id="71B04D620408FF97FBA7FE23A7BFFE46" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="drewesi">
<emphasis id="84C4EAF30408FF97FBA7FE23A7BFFE46" italics="true" pageId="12" pageNumber="13">S. drewesi</emphasis>
</taxonomicName>
(vs. three); snout longer than orbit (vs. about equal to); gill rakers on first arch 14 total (3+1+10) (vs. 19-23); long cirri on flap of anterior nostril 7 or 8 (vs. 1-3 stubby cirri); cycloid scales on body (vs. ctenoid scales); circumpeduncular scale rows 21 (vs. 23-26); scale rows below origin of dorsal fin 5 (vs. 6-8); interorbital space naked (vs. scaled); mandibular ramus finely scaled (vs. naked).
</paragraph>
<paragraph id="B60F36E10408FF97FF3EFDFFA401FD4D" blockId="12.[123,1174,219,1321]" pageId="12" pageNumber="13">
<taxonomicName id="71B04D620408FF97FF3EFDFFA657FDCB" authorityName="Iwamoto &amp; Wirtz" authorityYear="2018" box="[171,357,578,602]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="drewesi">
<emphasis id="84C4EAF30408FF97FF3EFDFFA657FDCB" box="[171,357,578,602]" italics="true" pageId="12" pageNumber="13">Serranus drewesi</emphasis>
</taxonomicName>
is readily differentiated from
<emphasis id="84C4EAF30408FF97FD3AFDFFA4A7FDCB" box="[687,917,578,602]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="71B04D620408FF97FD3AFDFFA43CFDCB" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[687,782,578,602]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="scriba">S. scriba</taxonomicName>
,
<taxonomicName id="71B04D620408FF97FC88FDFFA4BCFDCB" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[797,910,578,602]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="cabrilla">S. cabrilla</taxonomicName>
,
</emphasis>
and
<taxonomicName id="71B04D620408FF97FC59FDFFA363FDCB" authorityName="Gunther" authorityYear="1874" box="[972,1105,578,602]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="atricauda">
<emphasis id="84C4EAF30408FF97FC59FDFFA363FDCB" box="[972,1105,578,602]" italics="true" pageId="12" pageNumber="13">S. atricauda</emphasis>
</taxonomicName>
by its fewer lateral-line scales and its distinctive color pattern. The combination of relatively low fin-ray counts of soft dorsal, pectoral, and anal fins, and the fewer gill rakers, the absence of ctenoid scales, as well as the color pattern, which includes a large black spot on the dorsal fin, distinguish the new species from the remaining eastern Atlantic species of the genus.
</paragraph>
<paragraph id="B60F36E10408FF97FF3EFD59A6FCFD6C" blockId="12.[123,1174,219,1321]" box="[171,462,741,766]" pageId="12" pageNumber="13">
<emphasis id="84C4EAF30408FF97FF3EFD59A7D2FD6F" bold="true" box="[171,224,741,766]" pageId="12" pageNumber="13">
<emphasis id="84C4EAF30408FF97FF3EFD59A7E5FD6C" bold="true" box="[171,215,741,765]" italics="true" pageId="12" pageNumber="13">Size</emphasis>
:
</emphasis>
To at least
<quantity id="71489B040408FF97FEC8FD59A694FD6C" box="[349,422,741,765]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="13" unit="cm" value="6.5">6.5 cm</quantity>
TL
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0408FF97FF3EFCBAA54FFCAF" pageId="12" pageNumber="13" type="distribution">
<paragraph id="B60F36E10408FF97FF3EFCBAA54FFCAF" blockId="12.[123,1174,219,1321]" pageId="12" pageNumber="13">
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<emphasis id="84C4EAF30408FF97FF3EFCBAA671FC8C" bold="true" box="[171,323,774,798]" pageId="12" pageNumber="13">DISTRIBUTION</emphasis>
</smallCapsWord>
.—
<materialsCitation id="06D83CBC0408FF97FEFEFCBBA544FCAE" country="Sao Tome and Principe" location="Santana Islet" municipality="Republic" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
Known only from the
<typeStatus id="690B88430408FF97FDC8FCBAA588FC8F" box="[605,698,774,798]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
taken by mini-spear in a submarine cave in
<location id="B36F603A0408FF97FFEEFC9BA636FCAE" LSID="urn:lsid:plazi:treatment:3E1987F7040EFF97FFEEFDDEA444FAB9:B36F603A0408FF97FFEEFC9BA636FCAE" box="[123,260,807,831]" country="Sao Tome and Principe" municipality="Republic" name="Santana Islet" pageId="12" pageNumber="13">Santana Islet</location>
,
<collectingMunicipality id="566BAC9B0408FF97FE87FC9BA640FCAE" box="[274,370,807,831]" pageId="12" pageNumber="13">Republic</collectingMunicipality>
of
<collectingCountry id="CEA776710408FF97FE02FC9BA544FCAE" box="[407,630,807,831]" name="Sao Tome and Principe" pageId="12" pageNumber="13">São Tomé e Príncipe</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0408FF97FF3EFCF4A3B6FC50" pageId="12" pageNumber="13" type="etymology">
<paragraph id="B60F36E10408FF97FF3EFCF4A3B6FC50" blockId="12.[123,1174,219,1321]" pageId="12" pageNumber="13">
<smallCapsWord id="B0E9A03D0408FF97FF3EFCF4A607FCCF" baselines="858,857" box="[171,309,840,864]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Etymology" pageId="12" pageNumber="13">
<emphasis id="84C4EAF30408FF97FF3EFCF4A607FCCF" bold="true" box="[171,309,840,864]" pageId="12" pageNumber="13">ETYMOLOGY</emphasis>
</smallCapsWord>
.— Named in honor of Dr. Robert C. Drewes of the California Academy of Sciences, for his dedicated efforts in leading 12 separate scientific and educational expeditions to
<collectingCountry id="CEA776710408FF97FBF8FCD4A61BFC30" name="Sao Tome and Principe" pageId="12" pageNumber="13">São Tomé e Príncipe</collectingCountry>
to explore and document the diverse fauna and flora of that country and to inspire and educate the countrys citizens as to the biological wealth and uniqueness of where they live.
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0408FF97FF3EFC76A369FB57" pageId="12" pageNumber="13" type="discussion">
<paragraph id="B60F36E10408FF97FF3EFC76A369FB57" blockId="12.[123,1174,219,1321]" pageId="12" pageNumber="13">
<smallCapsWord id="B0E9A03D0408FF97FF3EFC76A624FC70" baselines="988,988" box="[171,278,970,994]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Remarks" pageId="12" pageNumber="13">
<emphasis id="84C4EAF30408FF97FF3EFC76A624FC70" bold="true" box="[171,278,970,994]" pageId="12" pageNumber="13">REMARKS</emphasis>
</smallCapsWord>
.— We describe this species with some reluctance owing to the absence of additional specimens and the small size of the only representative. We did not examine the internal organs, notably the gonads, to determine whether it is a juvenile or an adult. If the former, it is likely that the adult pigmentation pattern of this species differs substantially from the
<typeStatus id="690B88430408FF97FC3FFB90A335FBD5" box="[938,1031,1068,1092]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
. We also did not examine the myological, arthrological, and osteological characters used by
<bibRefCitation id="D2214B100408FF97FC46FBF1A3A7FBF4" author="MEISLER, M. R." box="[979,1173,1101,1125]" pageId="12" pageNumber="13" refId="ref21027" refString="MEISLER, M. R. 1987. Limits and relationships of serranine seabasses, with revisions of Serranus and Mentiperca (Pisces: Serranidae). Unpublished Ph. D. thesis. University of Southern California, Los Angeles, California, USA. 250 pages." type="book" year="1987">Meisler (1987:80)</bibRefCitation>
defining the subgenera
<taxonomicName id="71B04D620408FF97FEECFBD2A506FB14" authorityName="Klunzinger" authorityYear="1884" box="[377,564,1134,1157]" family="Serranidae" genus="Paracentropristis" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30408FF97FEECFBD2A506FB14" box="[377,564,1134,1157]" italics="true" pageId="12" pageNumber="13">Paracentropristis</emphasis>
</taxonomicName>
and
<taxonomicName id="71B04D620408FF97FDFEFBD2A5F9FB14" authorityName="Cuvier" authorityYear="1817" box="[619,715,1134,1157]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="84C4EAF30408FF97FDFEFBD2A5F9FB14" box="[619,715,1134,1157]" italics="true" pageId="12" pageNumber="13">Serranus</emphasis>
</taxonomicName>
(e.g., levator arcus palatini muscle inserts on preopercle and dorsal origin of ethmomaxillary ligament medial on mesethmoid). We await the collection of additional specimens to expand on the description of this obscure little species.
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0408FF97FF3EFB6CA444FAB9" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="B60F36E10408FF97FF3EFB6CA444FAB9" blockId="12.[123,1174,219,1321]" pageId="12" pageNumber="13">
<materialsCitation id="06D83CBC0408FF97FF3EFB6CA444FAB9" collectingDate="2009-01-28" collectionCode="TYPE, SPECIMEN" collectorName="Is. &amp; Santana Islet &amp; J. E. McCosker &amp; J. - L. Testori" country="Sao Tome e Principe, Sao Tome" elevation="19" latitude="0.24252778" location="Sao Tome e Principe, Sao Tome" longLatPrecision="1" longitude="-6.760028" pageId="12" pageNumber="13" specimenCode="CAS 234050" specimenCount="1" typeStatus="holotype">
<emphasis id="84C4EAF30408FF97FF3EFB6CA666FB77" bold="true" box="[171,340,1232,1256]" pageId="12" pageNumber="13">
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<collectionCode id="D0A1AE240408FF97FF3EFB6CA7D6FB77" box="[171,228,1232,1256]" pageId="12" pageNumber="13">TYPE</collectionCode>
</smallCapsWord>
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<collectionCode id="D0A1AE240408FF97FF7FFB6FA666FB77" box="[234,340,1235,1254]" pageId="12" pageNumber="13">SPECIMEN</collectionCode>
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</emphasis>
.—
<typeStatus id="690B88430408FF97FEE9FB73A6EDFB76" box="[380,479,1231,1255]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
,
<specimenCode id="E6169E9A0408FF97FE79FB6CA547FB76" box="[492,629,1231,1256]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="12" pageNumber="13" type="Museum">CAS 234050</specimenCode>
(
<quantity id="71489B040408FF97FD11FB73A5D0FB76" box="[644,738,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" pageId="12" pageNumber="13" unit="mm" value="53.5">53.5 mm</quantity>
SL);
<collectingCountry id="CEA776710408FF97FC8BFB6CA35CFB79" box="[798,1134,1231,1256]" isEnumeration="true" pageId="12" pageNumber="13">São Tomé e Príncipe, São Tomé</collectingCountry>
<collectorName id="1B4553370408FF97FBE1FB6CA3BCFB76" box="[1140,1166,1232,1255]" pageId="12" pageNumber="13">Is.</collectorName>
;
<collectorName id="1B4553370408FF97FFEEFB4CA63EFA99" box="[123,268,1264,1288]" pageId="12" pageNumber="13">Santana Islet</collectorName>
(
<geoCoordinate id="D38450260408FF97FEB6FB4CA69AFA99" box="[291,424,1263,1288]" degrees="0" direction="north" minutes="14" orientation="latitude" pageId="12" pageNumber="13" precision="1" seconds="33. 1" value="0.24252778">0°14ʹ33.1ʺN</geoCoordinate>
,
<geoCoordinate id="D38450260408FF97FE28FB4CA575FA99" box="[445,583,1263,1288]" degrees="6" direction="west" minutes="45" orientation="longitude" pageId="12" pageNumber="13" precision="1" seconds="36. 1" value="-6.760028">6°45ʹ36.1ʺW</geoCoordinate>
); collected by
<collectorName id="1B4553370408FF97FD6AFB4DA497FA99" box="[767,933,1264,1288]" pageId="12" pageNumber="13">J.E. McCosker</collectorName>
and
<collectorName id="1B4553370408FF97FC7CFB4DA340FA99" box="[1001,1138,1264,1288]" pageId="12" pageNumber="13">J.-L. Testori</collectorName>
in submarine cave at
<quantity id="71489B040408FF97FED1FAADA644FAB8" box="[324,374,1297,1321]" metricMagnitude="1" metricUnit="m" metricValue="1.88976" pageId="12" pageNumber="13" unit="ft" value="62.0">
<elevation id="3D9DD1D20408FF97FED1FAADA644FAB8" box="[324,374,1297,1321]" metricMagnitude="1" metricUnit="m" metricValue="1.88976" pageId="12" pageNumber="13" unit="ft" value="62.0">62 ft</elevation>
</quantity>
[
<quantity id="71489B040408FF97FE13FAADA6E3FAB9" box="[390,465,1297,1321]" metricMagnitude="1" metricUnit="m" metricValue="1.89" pageId="12" pageNumber="13" unit="m" value="18.9">
<elevation id="3D9DD1D20408FF97FE13FAADA6E3FAB9" box="[390,465,1297,1321]" metricMagnitude="1" metricUnit="m" metricValue="1.89" pageId="12" pageNumber="13" unit="m" value="18.9">18.9 m</elevation>
</quantity>
] depth;
<date id="C20E10210408FF97FDBEFAADA586FAB8" box="[555,692,1297,1321]" pageId="12" pageNumber="13" value="2009-01-28">
<collectingDate id="D24AE9C90408FF97FDBEFAADA586FAB8" box="[555,692,1297,1321]" pageId="12" pageNumber="13" value="2009-01-28">28 Jan. 2009</collectingDate>
</date>
; mini-hand spear.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,388 @@
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<mods:title id="DCFA6D21D13007D0D9335CBAD7768850">A Synopsis of the Eastern and Central Atlantic Combers of the Genus Serranus (Teleostei: Scorpaeniformes: Serranidae)</mods:title>
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<mods:affiliation id="7901B1E5E044D8809410C0B6DBDB5118">Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118 U. S. A. e-mail: tiwamoto @ calacademy. org.</mods:affiliation>
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<treatment id="3E1987F70411FF82FFEEFA9AA5B2FE46" LSID="urn:lsid:plazi:treatment:3E1987F70411FF82FFEEFA9AA5B2FE46" httpUri="http://treatment.plazi.org/id/3E1987F70411FF82FFEEFA9AA5B2FE46" lastPageId="25" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="FEAA656A0411FF8EFFEEFA9AA476FAAF" box="[123,836,1318,1343]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="B60F36E10411FF8EFFEEFA9AA476FAAF" blockId="21.[123,836,1318,1375]" box="[123,836,1318,1343]" pageId="21" pageNumber="22">
<heading id="ED47818D0411FF8EFFEEFA9AA476FAAF" box="[123,836,1318,1343]" fontSize="10" level="2" pageId="21" pageNumber="22" reason="4">
<taxonomicName id="71B04D620411FF8EFFEEFA9AA476FAAF" ID-CoL="4WX6K" authority="Wirtz and Iwamoto, 2016" authorityName="Wirtz and Iwamoto" authorityYear="2016" box="[123,836,1318,1343]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="pulcher" subGenus="Paracentropristis">
<emphasis id="84C4EAF30411FF8EFFEEFA9AA476FAAF" bold="true" box="[123,836,1318,1343]" pageId="21" pageNumber="22">
<emphasis id="84C4EAF30411FF8EFFEEFA9AA7D2FAAF" bold="true" box="[123,224,1318,1342]" italics="true" pageId="21" pageNumber="22">Serranus</emphasis>
(
<emphasis id="84C4EAF30411FF8EFF65FA9AA69CFAAF" bold="true" box="[240,430,1318,1342]" italics="true" pageId="21" pageNumber="22">Paracentropristis</emphasis>
)
<emphasis id="84C4EAF30411FF8EFE50FA9AA52BFAAF" bold="true" box="[453,537,1318,1342]" italics="true" pageId="21" pageNumber="22">pulcher</emphasis>
Wirtz and Iwamoto, 2016
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0411FF8EFFEEFAFBA311FA3B" pageId="21" pageNumber="22" type="description">
<paragraph id="B60F36E10411FF8EFFEEFAFBA627FACE" blockId="21.[123,836,1318,1375]" box="[123,277,1351,1375]" pageId="21" pageNumber="22">
<figureCitation id="2E8B2A640411FF8EFFEEFAFBA627FACE" box="[123,277,1351,1375]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStartId-0="23.[155,167,737,756]" captionStartId-1="23.[155,167,1387,1406]" captionStartId-2="24.[155,167,613,632]" captionStartId-3="24.[155,167,1269,1288]" captionTargetBox-0="[373,923,216,720]" captionTargetBox-1="[120,1176,781,1371]" captionTargetBox-2="[120,1176,216,600]" captionTargetBox-3="[120,1176,648,1255]" captionTargetId-0="figure-703@23.[373,923,216,720]" captionTargetId-1="figure-705@23.[120,1176,781,1372]" captionTargetId-2="figure-951@24.[120,1176,215,602]" captionTargetId-3="figure-860@24.[120,1176,647,1256]" captionTargetPageId-0="23" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="24" captionText-0="FIGURE 18. Serranus pulcher from São Tomé Island showing color pattern on chin and throat. Photo by Peter Wirtz." captionText-1="FIGURE19. Serranus pulcher A freshly collected specimen (paratype, CAS 227758, 36.3 mm SL) from Santana Islet, São Tomé. Photograph by David Catania." captionText-2="FIGURE 20. Serranus pulcher from Príncipe Island. Photograph by by Luiz Rocha." captionText-3="FIGURE 21. Serranus pulcher. A different-colored morph from Príncipe Island. Photograph by Luiz Rocha." figureDoi-0="http://doi.org/10.5281/zenodo.13156055" figureDoi-1="http://doi.org/10.5281/zenodo.13156059" figureDoi-2="http://doi.org/10.5281/zenodo.13156063" figureDoi-3="http://doi.org/10.5281/zenodo.13156065" httpUri-0="https://zenodo.org/record/13156055/files/figure.png" httpUri-1="https://zenodo.org/record/13156059/files/figure.png" httpUri-2="https://zenodo.org/record/13156063/files/figure.png" httpUri-3="https://zenodo.org/record/13156065/files/figure.png" pageId="21" pageNumber="22">Figures 1821</figureCitation>
</paragraph>
<paragraph id="B60F36E10411FF8EFFEEFACBA311FA3B" blockId="21.[123,1173,1398,1450]" pageId="21" pageNumber="22">
<taxonomicName id="71B04D620411FF8EFFEEFACBA7C1FA1D" box="[123,243,1399,1420]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="84C4EAF30411FF8EFFEEFACBA7E0FA1D" box="[123,210,1399,1420]" italics="true" pageId="21" pageNumber="22">Serranus</emphasis>
sp.
</taxonomicName>
<bibRefCitation id="D2214B100411FF8EFF6CFACBA6AEFA1D" author="KUITER, R." box="[249,412,1398,1421]" pageId="21" pageNumber="22" refId="ref20896" refString="KUITER, R. 2004. Serranidae and Plesiopidae. A Comprehensive Guide to Basslets, Hamlets, Longfins and Relatives. Aquatic Photographics, Seaford, Australia. 216 pp." type="book" year="2004">Kuiter, 2004:162</bibRefCitation>
figs. AD (
<collectingCountry id="CEA776710411FF8EFD98FACBA55FFA1D" box="[525,621,1399,1420]" name="Sao Tome and Principe" pageId="21" pageNumber="22">São Tomé</collectingCountry>
Comber; photographs taken by Peter Wirtz in
<collectingCountry id="CEA776710411FF8EFBB8FACBA3BFFA1D" box="[1069,1165,1399,1420]" name="Sao Tome and Principe" pageId="21" pageNumber="22">São Tomé</collectingCountry>
)
<taxonomicName id="71B04D620411FF8EFFEEFA29A548FA38" authority="Wirtz and Iwamoto, 2016: 192 - 199" authorityName="Wirtz and Iwamoto" authorityPageNumber="192 - 199" authorityYear="2016" box="[123,634,1428,1450]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84C4EAF30411FF8EFFEEFA29A613FA38" box="[123,289,1428,1450]" italics="true" pageId="21" pageNumber="22">Serranus pulcher</emphasis>
Wirtz and Iwamoto, 2016:192199
</taxonomicName>
, figs. 115 (
<collectingCountry id="CEA776710411FF8EFD63FA28A48CFA3B" box="[758,958,1428,1450]" name="Sao Tome and Principe" pageId="21" pageNumber="22">São Tomé e Príncipe</collectingCountry>
;
<quantity id="71489B040411FF8EFC5EFA28A327FA3B" box="[971,1045,1428,1450]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="3.0" metricValueMin="0.1" pageId="21" pageNumber="22" unit="m" value="15.5" valueMax="30.0" valueMin="1.0">130 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0411FF8DFF3EFA7FA7D4FEA2" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="B60F36E10411FF8DFF3EFA7FA7D4FEA2" blockId="21.[123,1174,1475,1725]" lastBlockId="22.[123,1174,219,1726]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<smallCapsWord id="B0E9A03D0411FF8EFF3EFA7FA612FA48" baselines="1493,1492" box="[171,288,1475,1499]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="21" pageNumber="22">
<emphasis id="84C4EAF30411FF8EFF3EFA7FA612FA48" bold="true" box="[171,288,1475,1499]" pageId="21" pageNumber="22">DIAGNOSIS</emphasis>
</smallCapsWord>
.— D X,12; P 15, rarely 14 or16; A III,7; gill rakers 69+1214 (1923 total); pored lateral-line scales 4249; circumpeduncular scales 2224; scales below origin of first dorsal fin 56. Caudal fin truncate, the upper lobe slightly produced, lower lobe rounded. Dorsal, anal, and pectoral fins scaly near base. Anterior nostril tubelike, rim low anteriorly but rising to a high narrow flap posteriorly, with 46 long cirri at distal tip; posterior nostril a simple opening lacking a raised rim. Seven or eight transverse bands on body; lips red with dark bands; a short moustache-like red streak behind end of maxillary, running across hind margin of dentary and almost meeting opposite streak at midventral line and enclosing ivory-white of mandibular rami; another red diagonal streak running from upper edge of maxillary, across lower edge of preopercle, across interopercle and branchiostegal rays, to base of pelvic fin, and enclosing white of preopercle, interopercle and chest.
</paragraph>
</subSubSection>
<subSubSection id="FEAA656A0412FF82FF3EFE80A5B2FE46" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23" type="description">
<paragraph id="B60F36E10412FF8DFF3EFE80A63BFDC4" blockId="22.[123,1174,219,1726]" pageId="22" pageNumber="23">
<smallCapsWord id="B0E9A03D0412FF8DFF3EFE80A608FEC3" baselines="334,333" box="[171,314,316,340]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="22" pageNumber="23">
<emphasis id="84C4EAF30412FF8DFF3EFE80A608FEC3" bold="true" box="[171,314,316,340]" pageId="22" pageNumber="23">DESCRIPTION</emphasis>
</smallCapsWord>
.— Body relatively slender to moderately deep and compressed, width over pectoral bases about half of HL, greatest body depth about 2.7
<quantity id="71489B040412FF8DFCB3FEE0A456FEE5" box="[806,868,348,372]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="22" pageNumber="23" unit="in" value="3.2">3.2 in</quantity>
SL and less than HL, which is about 2.4
<quantity id="71489B040412FF8DFE92FEC0A674FE05" box="[263,326,380,404]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="22" pageNumber="23" unit="in" value="2.6">2.6 in</quantity>
SL. Dorsal and ventral profiles gently curved from tip of snout to caudal fin. Snout shorter in length than orbit diameter; both substantially more than interorbital width. Lower jaw projecting slightly beyond upper jaw; maxilla extending to below posterior half of pupil. Anterior nostril tubular, the rim posteriorly elevated into a high narrow flap with 46 long slender cirri at distal tip; posterior nostril lacking raised rim. Opercular spines three, the uppermost small and often obscure, middle spine longest. Preopercle margin serrated with flattened spines, those at angle largest.
</paragraph>
<paragraph id="B60F36E10412FF8DFF3EFDE2A367FC26" blockId="22.[123,1174,219,1726]" pageId="22" pageNumber="23">
Premaxillary teeth in narrow band, the outer series spaced and slightly enlarged; one or more large canines at anterior end. Dentary with a band of small teeth flanked by a series of slightly enlarged outer teeth that become larger posteriorly. Vomerine tooth band broadly V-shaped, each arm followed by narrow band of small palatine teeth. First four to five spines of dorsal fin graduated, the 3
<superScript id="41C59BA90412FF8DFF7EFD62A7C9FD7E" attach="left" box="[235,251,734,751]" fontSize="7" pageId="22" pageNumber="23">rd</superScript>
to 5
<superScript id="41C59BA90412FF8DFEB9FD62A609FD7E" attach="left" box="[300,315,734,751]" fontSize="7" pageId="22" pageNumber="23">th</superScript>
spines longest, the spines following subequal; the soft rays slightly higher than longest spines, the 3
<superScript id="41C59BA90412FF8DFECCFD42A65BFC9E" attach="left" box="[345,361,766,783]" fontSize="7" pageId="22" pageNumber="23">rd</superScript>
to 5
<superScript id="41C59BA90412FF8DFE0EFD42A698FC9E" attach="left" box="[411,426,766,783]" fontSize="7" pageId="22" pageNumber="23">th</superScript>
ray longest; a slight notch in fin profile. Anal fin relatively high, its posterior tip somewhat pointed; spines shorter than soft rays, the first spine more than half length of second and third spines, the second spine longer and stouter than the others. Pectoral fin broad-based, its origin about on same vertical as that of dorsal and pelvic fins; the tip of pectoral fin extends to, or almost to, anus, that of pelvic fin falls short of anus, both fins falling well short of anal-fin origin. Caudal fin truncate; dorsal lobe slightly pointed, ventral lobe rounded at tip.
</paragraph>
<paragraph id="B60F36E10412FF8DFF3EFC7CA417FBC8" blockId="22.[123,1174,219,1726]" pageId="22" pageNumber="23">Scales ctenoid, body fully covered; head naked over snout, interorbital, suborbital region, outer margin of preopercle and over subopercle. Small scales on interradial membranes of dorsal fin, mostly confined to basal one-fourth to one-half of soft rays, but a few on membrane between spines. Anal and caudal fins also with small scales at base, but on caudal fin, scales extending more distally. Scales on pectoral fin limited to immediate base of fin.</paragraph>
<paragraph id="B60F36E10412FF8DFF3EFBDDA404F90C" blockId="22.[123,1174,219,1726]" pageId="22" pageNumber="23">
<emphasis id="84C4EAF30412FF8DFF3EFBDDA6AFFBE8" bold="true" box="[171,413,1121,1145]" pageId="22" pageNumber="23">
<emphasis id="84C4EAF30412FF8DFF3EFBDDA6A6FBE8" bold="true" box="[171,404,1121,1145]" italics="true" pageId="22" pageNumber="23">Color highly variable</emphasis>
:
</emphasis>
in a fresh specimen from
<collectingCountry id="CEA776710412FF8DFD24FBDDA450FBE8" box="[689,866,1121,1145]" name="Sao Tome Island" pageId="22" pageNumber="23">São Tomé Island</collectingCountry>
(CAS 227751,
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SL) dorsal aspects of head and body grayish brown to olive, this color extending down sides of head onto entire surface of opercle and most of preopercle but not interopercle and jaws; on body the ground color extends to mid-lateral line, below which the color becomes whitish to reddish and is dissected by seven or eight broad, red or brownish vertical bands that originate along dorsal outline of body and extend dorsally onto base of dorsal fin and ventrally to, or near, ventral outline. A faint brownish longitudinal stripe runs from upper lobe of opercle just above mid-lateral line to caudal fin base. Upper and lower lips deep red, but marked with dark bluish-gray bands and spots; lower lip with few faint bands or spots. A prominent red diagonal stripe from upper edge of maxillary, across interopercle, branchiostegal rays, and chest to base of pelvic fin. A short tapered moustache-like red streak behind end of maxilla, running across hind margin of dentary and almost meeting opposite streak at median-ventral line (see
<figureCitation id="2E8B2A640412FF8DFCB2FA78A446FA4D" box="[807,884,1476,1500]" captionStart="FIGURE" captionStartId="23.[155,167,737,756]" captionTargetBox="[373,923,216,720]" captionTargetId="figure-703@23.[373,923,216,720]" captionTargetPageId="23" captionText="FIGURE 18. Serranus pulcher from São Tomé Island showing color pattern on chin and throat. Photo by Peter Wirtz." figureDoi="http://doi.org/10.5281/zenodo.13156055" httpUri="https://zenodo.org/record/13156055/files/figure.png" pageId="22" pageNumber="23">Fig. 18</figureCitation>
). Two other diffuse diagonal red streaks behind lower half of opercle extend onto pectoral-fin base. Branchiostegal rays, gular membrane, ramus of lower jaw, chest, and antero-ventral parts of belly white except where marked by red streaks. Soft dorsal and anal fins dark grey-brown with small orange spots that form narrow, broken, diagonal to horizontal lines on fins. Pectoral fin clear with orange tint. Pelvic fins dusky with small orange spots; caudal fin yellow on upper and lower lobes, membrane between rays clear with small orange spots that form vertical lines on tail.
</paragraph>
<paragraph id="B60F36E10412FF8CFF3EF91AA5CAF9E3" blockId="22.[123,1174,219,1726]" lastBlockId="23.[122,1174,1467,1715]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
In a photograph of a
<quantity id="71489B040412FF8DFE34F919A536F92C" box="[417,516,1701,1726]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="22" pageNumber="23" unit="mm" value="35.3">35.3 mm</quantity>
SL specimen (CAS 227758) (
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) from
<collectingCountry id="CEA776710412FF8DFB97F919A369F92C" box="[1026,1115,1701,1725]" name="Sao Tome and Principe" pageId="22" pageNumber="23">Príncipe</collectingCountry>
(Isla Santana), brownish color of sides of head interrupted by four or five thin, horizontal to slightly diagonal, reddish to orange lines; orange spots scattered on top of head and on snout. Vertical bands on body with dark squarish blocks mid-laterally. Anal and soft dorsal fins dark; spinous dorsal paler with red spots at base where body bands meet fin; tips of spines red. Ivory-white of gular and branchiostegal membranes, interopercle, and chest bordered by prominent dark-red streaks. Pelvic fins overall clear to light dusky with brownish-yellow rays.
</paragraph>
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<paragraph id="B60F36E10413FF8CFF0EFD5DA34DFD65" blockId="23.[155,1151,736,756]" box="[155,1151,736,756]" pageId="23" pageNumber="24">
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18.
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<emphasis id="84C4EAF30413FF8CFF6BFD5DA6A0FD62" box="[254,402,736,756]" italics="true" pageId="23" pageNumber="24">Serranus pulcher</emphasis>
</taxonomicName>
from São Tomé Island showing color pattern on chin and throat. Photo by Peter Wirtz.
</paragraph>
</caption>
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<paragraph id="B60F36E10413FF8CFF0EFAD7A3B5FAEF" blockId="23.[123,1159,1387,1433]" box="[155,1159,1387,1406]" pageId="23" pageNumber="24">
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19.
<taxonomicName id="71B04D620413FF8CFF6CFAD7A6BEFAEF" authorityName="Wirtz and Iwamoto" authorityYear="2016" box="[249,396,1387,1406]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84C4EAF30413FF8CFF6CFAD7A6BEFAEF" box="[249,396,1387,1406]" italics="true" pageId="23" pageNumber="24">Serranus pulcher</emphasis>
</taxonomicName>
A freshly collected specimen (paratype, CAS 227758, 36.3 mm SL) from Santana Islet,
</paragraph>
<paragraph id="B60F36E10413FF8CFFEEFA3AA6EDFA08" blockId="23.[123,1159,1387,1433]" box="[123,479,1414,1433]" pageId="23" pageNumber="24">São Tomé. Photograph by David Catania.</paragraph>
</caption>
<paragraph id="B60F36E10413FF83FF3FF9C7A6CDFAC6" blockId="23.[122,1174,1467,1715]" lastBlockId="24.[122,1174,1312,1720]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
A fish from
<collectingCountry id="CEA776710413FF8CFEA3F9C7A6BDF902" box="[310,399,1659,1683]" name="Sao Tome and Principe" pageId="23" pageNumber="24">Príncipe</collectingCountry>
(
<figureCitation id="2E8B2A640413FF8CFE37F9C7A6C3F902" box="[418,497,1659,1683]" captionStart="FIGURE" captionStartId="24.[155,167,613,632]" captionTargetBox="[120,1176,216,600]" captionTargetId="figure-951@24.[120,1176,215,602]" captionTargetPageId="24" captionText="FIGURE 20. Serranus pulcher from Príncipe Island. Photograph by by Luiz Rocha." figureDoi="http://doi.org/10.5281/zenodo.13156063" httpUri="https://zenodo.org/record/13156063/files/figure.png" pageId="23" pageNumber="24">Fig. 20</figureCitation>
) shows a prominent, broad, brownishred, broken midlateral stripe extending from midorbit to middle of caudal fin, bordered above and below by irregular white stripes, the dorsum with broad brownish patches between white patches, and the ventral aspects of the body a solid dark red.
</paragraph>
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<paragraph id="B60F36E1041CFF83FF0EFDD9A469FDE9" blockId="24.[155,859,613,632]" box="[155,859,613,632]" pageId="24" pageNumber="25">
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20.
<taxonomicName id="71B04D62041CFF83FF6BFDD9A6A0FDE9" authorityName="Wirtz and Iwamoto" authorityYear="2016" box="[254,402,613,632]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84C4EAF3041CFF83FF6BFDD9A6A0FDE9" box="[254,402,613,632]" italics="true" pageId="24" pageNumber="25">Serranus pulcher</emphasis>
</taxonomicName>
from Príncipe Island. Photograph by by Luiz Rocha.
</paragraph>
</caption>
<caption id="E2CF6669041CFF83FF0EFB49A300FA99" ID-DOI="http://doi.org/10.5281/zenodo.13156065" ID-Zenodo-Dep="13156065" box="[155,1074,1269,1288]" httpUri="https://zenodo.org/record/13156065/files/figure.png" pageId="24" pageNumber="25" startId="24.[155,167,1269,1288]" targetBox="[120,1176,648,1255]" targetPageId="24" targetType="figure">
<paragraph id="B60F36E1041CFF83FF0EFB49A300FA99" blockId="24.[155,1074,1269,1288]" box="[155,1074,1269,1288]" pageId="24" pageNumber="25">
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21.
<taxonomicName id="71B04D62041CFF83FF6BFB49A6A0FA99" authorityName="Wirtz and Iwamoto" authorityYear="2016" box="[254,402,1269,1288]" family="Serranidae" genus="Serranus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84C4EAF3041CFF83FF6BFB49A6A0FA99" box="[254,402,1269,1288]" italics="true" pageId="24" pageNumber="25">Serranus pulcher</emphasis>
</taxonomicName>
. A different-colored morph from Príncipe Island. Photograph by Luiz Rocha.
</paragraph>
</caption>
<paragraph id="B60F36E1041CFF83FF3EFADCA6ECF986" blockId="24.[122,1174,1312,1720]" pageId="24" pageNumber="25">
In a photograph (
<figureCitation id="2E8B2A64041CFF83FEF6FADCA69DFAE9" box="[355,431,1376,1400]" captionStart="FIGURE" captionStartId="24.[155,167,1269,1288]" captionTargetBox="[120,1176,648,1255]" captionTargetId="figure-860@24.[120,1176,647,1256]" captionTargetPageId="24" captionText="FIGURE 21. Serranus pulcher. A different-colored morph from Príncipe Island. Photograph by Luiz Rocha." figureDoi="http://doi.org/10.5281/zenodo.13156065" httpUri="https://zenodo.org/record/13156065/files/figure.png" pageId="24" pageNumber="25">Fig. 21</figureCitation>
) of an individual from
<collectingCountry id="CEA77671041CFF83FD3CFADCA430FAE9" box="[681,770,1376,1400]" name="Sao Tome and Principe" pageId="24" pageNumber="25">Príncipe</collectingCountry>
, ground color orange with bold white stripes bordering a dark midlateral stripe that originates behind eye and extends to base of caudal fin; top of head, suborbital region, and snout dark gray with orange spots and vermiculations. In preserved specimens &lt;
<quantity id="71489B04041CFF83FEE9FA7CA6F8FA46" box="[380,458,1472,1496]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="25" unit="mm" value="40.0">40 mm</quantity>
SL, midlateral stripe most prominent and other markings rather obscure. This color morph appears to predominate in
<collectingCountry id="CEA77671041CFF83FD2DFA5CA423FA69" box="[696,785,1504,1528]" name="Sao Tome and Principe" pageId="24" pageNumber="25">Príncipe</collectingCountry>
, where the species is quite common (Luiz Rocha, pers. comm. 2015).
</paragraph>
<paragraph id="B60F36E1041CFF83FF3FF99CA65FF9A6" blockId="24.[122,1174,1312,1720]" box="[170,365,1568,1592]" pageId="24" pageNumber="25">
<emphasis id="84C4EAF3041CFF83FF3FF99CA7D2F9A9" bold="true" box="[170,224,1568,1592]" pageId="24" pageNumber="25">
<emphasis id="84C4EAF3041CFF83FF3FF99CA7E4F9A9" bold="true" box="[170,214,1568,1592]" italics="true" pageId="24" pageNumber="25">Size</emphasis>
:
</emphasis>
To
<quantity id="71489B04041CFF83FE9FF99CA60DF9A6" box="[266,319,1568,1592]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="cm" value="9.0">9 cm</quantity>
TL.
</paragraph>
<paragraph id="B60F36E1041CFF83FF3FF9FCA552F906" blockId="24.[122,1174,1312,1720]" pageId="24" pageNumber="25">
<emphasis id="84C4EAF3041CFF83FF3FF9FCA6E4F9C7" bold="true" box="[170,470,1600,1624]" pageId="24" pageNumber="25">
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</emphasis>
.— Hard bottoms (rock, gravel, coral rubble, or marl) from about
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depth (juveniles) to at least
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.
</specimenCount>
São Tomé e Príncipe, off
<collectingCountry id="CEA77671041CFF83FCADF9DCA4C8F9E9" box="[824,1018,1632,1656]" isEnumeration="true" pageId="24" pageNumber="25">Ghana and Gabon</collectingCountry>
, and probably more widely along Gulf of Guinea mainland.
</paragraph>
<paragraph id="B60F36E1041CFF82FF3EF923A5B2FE46" blockId="24.[122,1174,1312,1720]" lastBlockId="25.[123,1174,218,471]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<emphasis id="84C4EAF3041CFF83FF3EF923A6AAF927" bold="true" box="[171,408,1695,1719]" pageId="24" pageNumber="25">
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</emphasis>
(
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).—
<collectingCountry id="CEA77671041CFF83FDB4F923A437F926" box="[545,773,1695,1720]" name="Sao Tome and Principe" pageId="24" pageNumber="25">
<emphasis id="84C4EAF3041CFF83FDB4F923A437F926" bold="true" box="[545,773,1695,1720]" pageId="24" pageNumber="25">São Tomé e Príncipe</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="84C4EAF3041CFF83FC93F91CA43DF929" bold="true" box="[774,783,1696,1720]" pageId="24" pageNumber="25">:</emphasis>
<materialsCitation id="06D83CBC041CFF82FC81F91CA6CFFE85" collectingDate="2009-01-20" collectionCode="CAS" collectorName="Bom Bom Is. &amp; J. E. McCosker &amp; D. Catania" country="Sao Tome and Principe" elevation="15" lastPageId="25" lastPageNumber="26" latitude="1.6955556" location="Is." longLatPrecision="15" longitude="7.4000835" pageId="24" pageNumber="25" specimenCode="CAS 227754" specimenCount="1" stateProvince="Principe">
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(4, 37.260.
<quantity id="71489B04041CFF83FBB1F91CA352F926" box="[1060,1120,1696,1720]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="mm" value="2.0">2 mm</quantity>
SL);
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, nw side of
<collectorName id="1B455337041DFF82FE05FF67A51BFF63" box="[400,553,219,242]" pageId="25" pageNumber="26">Bom Bom Is.</collectorName>
;
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,
<geoCoordinate id="D3845026041DFF82FD2BFF67A403FF62" box="[702,817,218,243]" degrees="7" direction="east" minutes="24" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="0. 3" value="7.4000835">7°24ʹ0.3ʺE</geoCoordinate>
;
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</quantity>
[
<quantity id="71489B04041DFF82FC1AFF67A4EDFF63" box="[911,991,219,243]" metricMagnitude="1" metricUnit="m" metricValue="1.46" pageId="25" pageNumber="26" unit="m" value="14.6">
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</quantity>
]; collectors
<collectorName id="1B455337041DFF82FBF8FF67A7DBFE85" pageId="25" pageNumber="26">J.E. McCosker</collectorName>
,
<collectorName id="1B455337041DFF82FF63FF40A655FE85" box="[246,359,252,276]" pageId="25" pageNumber="26">D. Catania</collectorName>
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View file

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<collectingRegion id="1C784080FFC55947FE81FA2207EB84B7" box="[341,418,1428,1452]" country="Mexico" name="Jalisco" pageId="44" pageNumber="85">Jalisco</collectingRegion>
</emphasis>
:
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<quantity id="19442387FFC55947FE64FA2307A884B6" box="[432,481,1429,1453]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="44" pageNumber="85" unit="mi" value="8.0">8 mi</quantity>
SW of Pihuamo
</location>
[ca.
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,
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],
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</quantity>
,
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<collectingDate id="BA46514AFFC55947FF49FA03075E84D6" box="[157,279,1461,1485]" pageId="44" pageNumber="85" value="1959-12-06">6-XII-1959</collectingDate>
</date>
(flr),
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<emphasis id="ECC85270FFC55947FE83FA00072E84D6" box="[343,359,1462,1485]" italics="true" pageId="44" pageNumber="85">R</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFC55947FEB3FA00045184D6" box="[359,536,1461,1485]" italics="true" pageId="44" pageNumber="85">
.
<materialsCitation id="6ED4843FFFC55947FEA1FA03045B84D6" box="[373,530,1461,1485]" collectionCode="W" country="Mexico" location="McVaugh" pageId="44" pageNumber="85" specimenCount="1" stateProvince="Jalisco">
<location id="DB63D8B9FFC55947FEA1FA03079384D6" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC55947FEA1FA03079384D6" box="[373,474,1461,1485]" country="Mexico" name="McVaugh" pageId="44" pageNumber="85" stateProvince="Jalisco">McVaugh</location>
&amp;
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</materialsCitation>
.
</emphasis>
<materialsCitation id="6ED4843FFFC55947FDCBFA03026784D6" box="[543,1070,1461,1485]" collectionCode="MICH, MEXU" country="Mexico" location="Koelz" pageId="44" pageNumber="85" specimenCount="2" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="ECC85270FFC55947FDCBFA0304D184D6" box="[543,664,1461,1485]" italics="true" pageId="44" pageNumber="85">
<location id="DB63D8B9FFC55947FDCBFA03041584D6" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC55947FDCBFA03041584D6" box="[543,604,1461,1485]" country="Mexico" name="Koelz" pageId="44" pageNumber="85" stateProvince="Jalisco">Koelz</location>
1507
</emphasis>
(
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:
<collectionCode id="B8AD16A7FFC55947FCC1FA03051284D6" box="[789,859,1461,1485]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="44" pageNumber="85" type="Herbarium">MICH</collectionCode>
!;
<typeStatus id="010730C0FFC55947FCA6FA0305F684D6" box="[882,959,1461,1485]" pageId="44" pageNumber="85" type="isotype">isotype</typeStatus>
:
<collectionCode id="B8AD16A7FFC55947FC1AFA00025484D6" box="[974,1053,1462,1485]" country="Mexico" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15658" lsid="urn:lsid:biocol.org:col:15658" name="Universidad Nacional Autónoma de México" pageId="44" pageNumber="85" type="Herbarium">MEXU</collectionCode>
!)
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.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFC55946FF7FFA5405CD82D6" lastPageId="45" lastPageNumber="86" pageId="44" pageNumber="85" type="description">
<paragraph id="DE038E62FFC55946FF7FFA54045082BA" blockId="44.[123,1174,1505,1726]" lastBlockId="45.[123,1174,219,929]" lastPageId="45" lastPageNumber="86" pageId="44" pageNumber="85">
Trees to
<quantity id="19442387FFC55947FEC4FA57070284E2" box="[272,331,1505,1529]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="85" unit="m" value="10.0">10 m</quantity>
tall. Older (woody) stems subquadrate, irregularly fissured-striate; younger (herbaceous) stems stubby, quadrate, glabrous. Leaves deciduous, clustered at apex of old growth, petiolate, petioles to
<quantity id="19442387FFC55947FEB2F99507FC8721" box="[358,437,1571,1595]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="44" pageNumber="85" unit="mm" value="73.0">73 mm</quantity>
long, blades membranaceous, ovate to elliptic,
<quantity id="19442387FFC55947FC04F995021D8721" box="[976,1108,1571,1595]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="2.2" metricValueMin="0.9" pageId="44" pageNumber="85" unit="mm" value="155.0" valueMax="220.0" valueMin="90.0">90220 mm</quantity>
long,
<quantity id="19442387FFC55947FFAFF9F506B28740" box="[123,251,1603,1627]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="10.0" metricValueMin="3.5" pageId="44" pageNumber="85" unit="mm" value="67.5" valueMax="100.0" valueMin="35.0">35100 mm</quantity>
wide, 22.7 × longer than wide, acute to acuminate at apex, rounded to attenuate and asymmetric at base, surfaces glabrous, margin entire. Inflorescence of axillary and/or terminal subsessile to pedunculate thyrses (main axis with lateral pedunculate dichasia appearing as branches) to
<quantity id="19442387FFC55947FF1CF913076F87A6" box="[200,294,1701,1725]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="44" pageNumber="85" unit="mm" value="250.0">250 mm</quantity>
long, peduncle to
<quantity id="19442387FFC55947FDD2F913041F87A6" box="[518,598,1701,1725]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="44" pageNumber="85" unit="mm" value="50.0">50 mm</quantity>
long, pubescent with erect to flexuose eglandular trichomes 0.050.2 (1) mm long, rachis pubescent with erect to flexuose eglandular trichomes 0.10.4 (1) mm long; dichasia opposite, pedunculate, 13 (or more)-flowered, to
<quantity id="19442387FFC45946FBDAFF4A02158008" box="[1038,1116,252,276]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="45" pageNumber="86" unit="mm" value="63.0">63 mm</quantity>
long (excluding corollas), dichasial peduncles
<quantity id="19442387FFC45946FDE9FEAA04ED802F" box="[573,676,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="2.0" metricValueMin="0.3" pageId="45" pageNumber="86" unit="mm" value="11.5" valueMax="20.0" valueMin="3.0">320 mm</quantity>
long, pubescent like rachis. Bracts caducous, lance-elliptic to elliptic,
<quantity id="19442387FFC45946FE55FE8B07BA804E" box="[385,499,317,341]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="45" pageNumber="86" unit="mm" value="14.5" valueMax="17.0" valueMin="12.0">1217 mm</quantity>
long, 3.5
<quantity id="19442387FFC45946FDB3FE8B04ED804E" box="[615,676,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="45" pageNumber="86" unit="mm" value="5.0">5 mm</quantity>
wide, abaxially pubescent with erect to flexuose eglandular trichomes 0.05
<quantity id="19442387FFC45946FE18FEE80457806E" box="[460,542,350,374]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="86" unit="mm" value="0.2">0.2 mm</quantity>
long. Bracteoles and secondary bracteoles usually present on young inflorescences but becoming deciduous as inflorescence matures, lance-elliptic to elliptic,
<quantity id="19442387FFC45946FF70FE16074080AC" box="[164,265,415,439]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.4" metricValueMin="0.7" pageId="45" pageNumber="86" unit="mm" value="10.5" valueMax="14.0" valueMin="7.0">714 mm</quantity>
long, 23.
<quantity id="19442387FFC45946FEAAFE2907F580AC" box="[382,444,415,439]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="45" pageNumber="86" unit="mm" value="5.0">5 mm</quantity>
wide, abaxially pubescent like bracts. Flowers pedicellate, pedicels
<quantity id="19442387FFC45946FFAFFE7606B880CC" box="[123,241,448,472]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="45" pageNumber="86" unit="mm" value="30.0" valueMax="50.0" valueMin="10.0">1050 mm</quantity>
long, pubescent like rachis. Calyx
<quantity id="19442387FFC45946FD54FE7604BF80CC" box="[640,758,448,472]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.8" metricValueMin="1.7" pageId="45" pageNumber="86" unit="mm" value="22.5" valueMax="28.0" valueMin="17.0">1728 mm</quantity>
long, lobes subheteromorphic, membranaceous, abaxially pubescent with erect (to ± antrorse) eglandular trichomes 12 (2.3) mm long, posterior lobe planar to ± conduplicate, ovate-elliptic,
<quantity id="19442387FFC45946FD2DFDB705228302" box="[761,875,513,537]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="45" pageNumber="86" unit="mm" value="21.0" valueMax="25.0" valueMin="17.0">1725 mm</quantity>
long,
<quantity id="19442387FFC45946FC7AFDB702698302" box="[942,1056,513,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="45" pageNumber="86" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">1115 mm</quantity>
wide, usually slightly larger than lateral lobes, acuminate at apex, lateral lobes ovate-elliptic to elliptic to obovate, 15.
<quantity id="19442387FFC45946FED4FDF407218341" box="[256,360,578,603]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.2" metricValueMin="0.5" pageId="45" pageNumber="86" unit="mm" value="13.5" valueMax="22.0" valueMin="5.0">522 mm</quantity>
long,
<quantity id="19442387FFC45946FE65FDF504508341" box="[433,537,578,603]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="45" pageNumber="86" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">915 mm</quantity>
wide, rounded to acute at apex, veins of lobes prominent (ca. 9 major veins). Corolla “pale green” to greenish yellow,
<quantity id="19442387FFC45946FCDDFDD505358360" box="[777,892,611,635]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="45" pageNumber="86" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">5055 mm</quantity>
long, externally glabrous, tube
<quantity id="19442387FFC45946FF7AFD3207698380" box="[174,288,644,668]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="45" pageNumber="86" unit="mm" value="31.0" valueMax="33.0" valueMin="29.0">2933 mm</quantity>
long, narrow proximal portion
<quantity id="19442387FFC45946FDB8FD3204998380" box="[620,720,644,668]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="45" pageNumber="86" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">913 mm</quantity>
long,
<quantity id="19442387FFC45946FCC7FD32053E8380" box="[787,887,644,668]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="45" pageNumber="86" unit="mm" value="12.5" valueMax="16.0" valueMin="9.0">916 mm</quantity>
in diameter near midpoint, throat
<quantity id="19442387FFC45946FF16FD13077F83A7" box="[194,310,677,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="45" pageNumber="86" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">1520 mm</quantity>
long,
<quantity id="19442387FFC45946FEAAFD1307BB83A7" box="[382,498,677,701]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="45" pageNumber="86" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">3035 mm</quantity>
in diameter at mouth, lobes spreading to recurved, “narrowly ovate to subcircular” (fide Miranda and McVaugh 1962),
<quantity id="19442387FFC45946FD0BFD73051883C6" box="[735,849,709,733]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="45" pageNumber="86" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">1824 mm</quantity>
long,
<quantity id="19442387FFC45946FC40FD73024F83C6" box="[916,1030,709,733]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.7" metricValueMin="1.7" pageId="45" pageNumber="86" unit="mm" value="22.0" valueMax="27.0" valueMin="17.0">1727 mm</quantity>
wide, round- ed to acute at apex. Stamens 4,
<quantity id="19442387FFC45946FE1AFD50040883E5" box="[462,577,742,766]" metricMagnitude="-2" metricUnit="m" metricValue="8.1" metricValueMax="9.0" metricValueMin="7.2" pageId="45" pageNumber="86" unit="mm" value="81.0" valueMax="90.0" valueMin="72.0">7290 mm</quantity>
long, filaments glabrous distally, pubescent proximally with eglandular trichomes, thecae
<quantity id="19442387FFC45946FE27FCB1042E8205" box="[499,615,775,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="45" pageNumber="86" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
long; staminode (if present) not seen. Style
<quantity id="19442387FFC45946FB87FCB106ED8224" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="45" pageNumber="86" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">5055 mm</quantity>
long, glabrous, stigma lobes
<quantity id="19442387FFC45946FE0DFC9E04608224" box="[473,553,807,832]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="86" unit="mm" value="1.5">1.5 mm</quantity>
long, shape and width not determined. Capsule
<quantity id="19442387FFC45946FBF7FC9102DC8224" box="[1059,1173,807,832]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="45" pageNumber="86" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">2530 mm</quantity>
long,
<quantity id="19442387FFC45946FF6FFCFF076A827B" box="[187,291,840,864]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="45" pageNumber="86" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">710 mm</quantity>
in diameter, pubescent throughout with erect to flexuose to antrorse eglandular trichomes 0.05
<quantity id="19442387FFC45946FEF3FCDF0731829B" box="[295,376,873,897]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="86" unit="mm" value="0.2">0.2 mm</quantity>
long, stipe 1.5
<quantity id="19442387FFC45946FDF1FCDF042B829B" box="[549,610,873,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="45" pageNumber="86" unit="mm" value="2.0">2 mm</quantity>
long. Seeds up to 16 per capsule,
<quantity id="19442387FFC45946FC05FCDF0260829B" box="[977,1065,873,897]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="45" pageNumber="86" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long,
<quantity id="19442387FFC45946FBB9FCDF06ED82BA" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="45" pageNumber="86" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
wide, surfaces minutely papillose.
</paragraph>
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<emphasis id="ECC85270FFC45946FF7FFC00077882D7" bold="true" box="[171,305,950,974]" pageId="45" pageNumber="86">PHENOLOGY</emphasis>
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.— Flowering: DecemberFebruary; fruiting: February.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFC45946FF7FFC6006AD84C3" pageId="45" pageNumber="86" type="distribution">
<paragraph id="DE038E62FFC45946FF7FFC6005438534" blockId="45.[123,1174,949,1726]" pageId="45" pageNumber="86">
<emphasis id="ECC85270FFC45946FF7FFC60079A82F6" bold="true" box="[171,467,982,1006]" pageId="45" pageNumber="86">
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<smallCapsWord id="D8E518BEFFC45946FEAEFC6C079A82F6" baselines="1000" box="[378,467,986,1005]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="habitat" pageId="45" pageNumber="86">HABITAT</smallCapsWord>
</emphasis>
.— West-central
<collectingRegion id="1C784080FFC45946FD52FC60049E82F5" box="[646,727,982,1006]" country="Mexico" name="Mexico" pageId="45" pageNumber="86">Mexico</collectingRegion>
(
<collectingRegion id="1C784080FFC45946FD33FC6005FB82F5" box="[743,946,982,1006]" isEnumeration="true" pageId="45" pageNumber="86">Jalisco, Michoacán</collectingRegion>
;
<figureCitation id="468792E7FFC45946FC14FC6005B782F5" box="[960,1022,982,1006]" captionStart="FIGURE" captionStartId="19.[155,167,950,969]" captionTargetBox="[127,1170,219,938]" captionTargetId="figure-78@19.[120,1176,216,945]" captionTargetPageId="19" captionText="FIGURE 8. Map of Mexico (showing states) and northern Central America with distributions of species of Louteridium endemic to Mexico: L. brevicalyx, L. dendropilosum, L. koelzii, L. parayi, L. rzedowskianum, and L. tamaulipense. Sym- bols may pertain to multiple collections." figureDoi="http://doi.org/10.5281/zenodo.13155723" httpUri="https://zenodo.org/record/13155723/files/figure.png" pageId="45" pageNumber="86">Fig. 8</figureCitation>
); plants occur on limestone slopes in “high forest dominated by
<taxonomicName id="19BCF5E1FFC45946FD4CFC4E054C8514" box="[664,773,1016,1039]" class="Magnoliopsida" family="Moraceae" genus="Brosimum" kingdom="Plantae" order="Rosales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="genus">
<emphasis id="ECC85270FFC45946FD4CFC4E054C8514" box="[664,773,1016,1039]" italics="true" pageId="45" pageNumber="86">Brosimum</emphasis>
</taxonomicName>
” in regions of tropical subdeciduous and deciduous forests at elevations from 500 to 1400 meters.
</paragraph>
<paragraph id="DE038E62FFC45946FF7FFB8E05FB854B" blockId="45.[123,1174,949,1726]" box="[171,946,1080,1104]" pageId="45" pageNumber="86">
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<emphasis id="ECC85270FFC45946FF7FFB8E07198554" bold="true" box="[171,336,1080,1104]" pageId="45" pageNumber="86">ILLUSTRATIONS</emphasis>
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.—
<figureCitation id="468792E7FFC45946FEADFB8E07A8854B" box="[377,481,1080,1104]" captionStart="FIGURE" captionStartId="46.[155,167,1620,1639]" captionTargetBox="[120,1174,222,1611]" captionTargetId="figure-59@46.[120,1176,216,1617]" captionTargetPageId="46" captionText="FIGURE 14. Louteridium koelzii. A. Apex of shoot with flush of new leaves (McVaugh &amp; Koelz 1797). B. Branch with inflorescence in flower (McVaugh &amp; Koelz 1507). C. Leaf (McVaugh &amp; Koelz 1797). D. Corolla bud opened (McVaugh &amp; Koelz 1797). E. Dehisced corolla with epipetalous stamens (McVaugh 26180). F. Distal portion of style and stigma (McVaugh 26180). G. Partially dehisced capsule with calyx and pedicel (McVaugh 26180). H. Seed (McVaugh 26180). Drawn by Karin Douthit, copyright reserved to University of Michigan Herbarium, used with permission." figureDoi="http://doi.org/10.5281/zenodo.13155739" httpUri="https://zenodo.org/record/13155739/files/figure.png" pageId="45" pageNumber="86">Figure 14</figureCitation>
; Miranda and McVaugh (1962:183, fig. 2).
</paragraph>
<paragraph id="DE038E62FFC45946FF7FFBEF06AD84C3" blockId="45.[123,1174,949,1726]" pageId="45" pageNumber="86">
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<emphasis id="ECC85270FFC45946FF7FFBEF07078574" bold="true" box="[171,334,1113,1137]" pageId="45" pageNumber="86">CONSERVATION</emphasis>
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.—
<taxonomicName id="19BCF5E1FFC45946FEA2FBEF040E856A" box="[374,583,1113,1137]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="ECC85270FFC45946FEA2FBEF040E856A" box="[374,583,1113,1137]" italics="true" pageId="45" pageNumber="86">Louteridium koelzii</emphasis>
</taxonomicName>
<materialsCitation id="6ED4843FFFC45946FD9AFBEF069484C2" collectionCode="EOO, EN" country="Mexico" county="The" location="Based" municipality="McVaugh" pageId="45" pageNumber="86" specimenCode="B1" specimenCount="1" stateProvince="Jalisco">
is currently known from a very limited region of western
<collectingCountry id="A6ABCEF2FFC45946FF71FBCC06BF8589" box="[165,246,1146,1170]" name="Mexico" pageId="45" pageNumber="86">Mexico</collectingCountry>
(
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=
<quantity id="19442387FFC45946FE8BFBCF07E28589" box="[351,427,1145,1170]" metricMagnitude="3" metricUnit="m" metricValue="5.3" pageId="45" pageNumber="86" unit="km" value="5.3">5.3 km</quantity>
<superScript id="29C9232AFFC45946FE7FFBCF07FD8591" attach="left" box="[427,436,1145,1162]" fontSize="7" pageId="45" pageNumber="86">2</superScript>
; AOO =
<quantity id="19442387FFC45946FDC8FBCC04288589" box="[540,609,1146,1170]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="45" pageNumber="86" unit="km" value="12.0">12 km</quantity>
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).
<collectingCounty id="3762F6EEFFC45946FD56FBCC04E28589" box="[642,683,1146,1170]" pageId="45" pageNumber="86">The</collectingCounty>
species is not known to occur in a protected area, and it has not been recollected since 1975 (i.e.,
<emphasis id="ECC85270FFC45946FD04FB2C05CB85A9" box="[720,898,1178,1202]" italics="true" pageId="45" pageNumber="86">
<collectingMunicipality id="3E671418FFC45946FD04FB2C057C85A9" box="[720,821,1178,1202]" pageId="45" pageNumber="86">McVaugh</collectingMunicipality>
26180
</emphasis>
; although see discussion below about an undetermined plant from a nearby region of
<collectingRegion id="1C784080FFC45946FD2BFB0D050185C8" box="[767,840,1211,1235]" country="Mexico" name="Jalisco" pageId="45" pageNumber="86">Jalisco</collectingRegion>
).
<location id="DB63D8B9FFC45946FC8AFB0D05D685C8" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC45946FC8AFB0D05D685C8" box="[862,927,1211,1235]" country="Mexico" county="The" municipality="McVaugh" name="Based" pageId="45" pageNumber="86" stateProvince="Jalisco">Based</location>
on current knowledge, there appear to be two subpopulations about
<quantity id="19442387FFC45946FDBDFB6A04E785EF" box="[617,686,1244,1268]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="45" pageNumber="86" unit="km" value="80.0">80 km</quantity>
apart, one in
<collectingRegion id="1C784080FFC45946FC9EFB6A05DA85EF" box="[842,915,1244,1268]" country="Mexico" name="Jalisco" pageId="45" pageNumber="86">Jalisco</collectingRegion>
and one in
<collectingRegion id="1C784080FFC45946FBCCFB6A02C785EF" box="[1048,1166,1244,1268]" country="Mexico" name="Michoacan de Ocampo" pageId="45" pageNumber="86">Michoacán</collectingRegion>
.
<location id="DB63D8B9FFC45946FFAFFB4A06BA840F" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC45946FFAFFB4A06BA840F" box="[123,243,1276,1300]" country="Mexico" county="The" municipality="McVaugh" name="Comparing" pageId="45" pageNumber="86" stateProvince="Jalisco">Comparing</location>
historic landsat images (2006 to 2017) via
<bibRefCitation id="BA2DF393FFC45946FD6AFB4A058F840E" author="GOOGLE EARTH PRO" box="[702,966,1276,1301]" pageId="45" pageNumber="86" refId="ref48440" refString="GOOGLE EARTH PRO. 2018. Version 7. 3. 1. 4507 (64 - bit). &lt;https: // earth. google. com / download-earth. html&gt; [accessed 24 May 2018]" type="url" year="2018">Google Earth Pro (2018)</bibRefCitation>
in the region of the known collections from
<collectingRegion id="1C784080FFC45946FE5FFAAB079D842E" box="[395,468,1309,1333]" country="Mexico" name="Jalisco" pageId="45" pageNumber="86">Jalisco</collectingRegion>
, it is evident that deforestation for agriculture has reduced the natural vegetation there substantially (ca. ≥ 30%), thus creating a threat to this species, at least in that region.
<location id="DB63D8B9FFC45946FF2FFAE90755846D" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC45946FF2FFAE90755846D" box="[251,284,1375,1398]" country="Mexico" county="The" municipality="McVaugh" name="No" pageId="45" pageNumber="86" stateProvince="Jalisco">No</location>
specific threat has been identified for the subpopulation in
<collectingRegion id="1C784080FFC45946FC4DFAE80246846D" box="[921,1039,1374,1398]" country="Mexico" name="Michoacan de Ocampo" pageId="45" pageNumber="86">Michoacán</collectingRegion>
.
<location id="DB63D8B9FFC45946FBC9FAE8021B846D" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC45946FBC9FAE8021B846D" box="[1053,1106,1374,1398]" country="Mexico" county="The" municipality="McVaugh" name="Thus" pageId="45" pageNumber="86" stateProvince="Jalisco">Thus</location>
, there are two locations for the species, and a threat is evident for one of them.
<location id="DB63D8B9FFC45946FC44FAC90598848C" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC45946FC44FAC90598848C" box="[912,977,1407,1431]" country="Mexico" county="The" municipality="McVaugh" name="Based" pageId="45" pageNumber="86" stateProvince="Jalisco">Based</location>
on these observations, a preliminary conservation assessment of
<location id="DB63D8B9FFC45946FD77FA16056B84A3" LSID="urn:lsid:plazi:treatment:56153F74FFC55944FF7FFAC204328308:DB63D8B9FFC45946FD77FA16056B84A3" box="[675,802,1440,1464]" country="Mexico" county="The" municipality="McVaugh" name="Endangered" pageId="45" pageNumber="86" stateProvince="Jalisco">Endangered</location>
(
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;
<specimenCode id="8E1A2619FFC45946FCA5FA1605D984A3" box="[881,912,1440,1464]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="45" pageNumber="86" type="Herbarium">B1</specimenCode>
, a, b) is proposed for
<taxonomicName id="19BCF5E1FFC45946FFAFFA77069484C2" box="[123,221,1473,1497]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="ECC85270FFC45946FFAFFA77069484C2" box="[123,221,1473,1497]" italics="true" pageId="45" pageNumber="86">L. koelzii</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFC45944FF7FFA5704328308" lastPageId="47" lastPageNumber="88" pageId="45" pageNumber="86" type="discussion">
<paragraph id="DE038E62FFC45944FF7FFA5704238088" blockId="45.[123,1174,949,1726]" lastBlockId="47.[123,1174,219,532]" lastPageId="47" lastPageNumber="88" pageId="45" pageNumber="86">
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<emphasis id="ECC85270FFC45946FF7FFA57076384E3" bold="true" box="[171,298,1505,1529]" pageId="45" pageNumber="86">DISCUSSION</emphasis>
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.— An undetermined specimen of
<taxonomicName id="19BCF5E1FFC45946FD66FA57057F84E2" box="[690,822,1505,1529]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="genus">
<emphasis id="ECC85270FFC45946FD66FA57057F84E2" box="[690,822,1505,1529]" italics="true" pageId="45" pageNumber="86">Louteridium</emphasis>
</taxonomicName>
from western
<collectingCountry id="A6ABCEF2FFC45946FC0BFA57027984E2" box="[991,1072,1505,1529]" name="Mexico" pageId="45" pageNumber="86">Mexico</collectingCountry>
(
<collectingRegion id="1C784080FFC45946FB91FA5702C784E2" box="[1093,1166,1505,1529]" country="Mexico" name="Jalisco" pageId="45" pageNumber="86">Jalisco</collectingRegion>
: Mpio. Jilotlán de los Dolores, Cerro de San Antonio,
<quantity id="19442387FFC45946FD6FF9B404B68701" box="[699,767,1538,1562]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="86" unit="km" value="15.0">15 km</quantity>
N de Tazumbos, matorral subtropical en suelos con pedregocidad de toba volcánica,
<quantity id="19442387FFC45946FDB9F99504F98721" box="[621,688,1571,1595]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="45" pageNumber="86" unit="m" value="900.0">900 m</quantity>
,
<date id="AA02A8A2FFC45946FD69F995057D8720" box="[701,820,1571,1595]" pageId="45" pageNumber="86" value="1982-02-06">6 Feb 1982</date>
,
<emphasis id="ECC85270FFC45946FC95F99502778720" box="[833,1086,1571,1595]" italics="true" pageId="45" pageNumber="86">F. Santana Michel 1081</emphasis>
, IBUG) does not seem to pertain to either species known from nearby regions (i.e.,
<taxonomicName id="19BCF5E1FFC45946FC37F9F2023A8740" box="[995,1139,1603,1627]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="species" species="brevicalyx">
<emphasis id="ECC85270FFC45946FC37F9F2023A8740" box="[995,1139,1603,1627]" italics="true" pageId="45" pageNumber="86">L. brevicalyx</emphasis>
</taxonomicName>
or
<taxonomicName id="19BCF5E1FFC45946FFAFF9D306978767" box="[123,222,1636,1660]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="ECC85270FFC45946FFAFF9D306978767" box="[123,222,1636,1660]" italics="true" pageId="45" pageNumber="86">L. koelzii</emphasis>
</taxonomicName>
) or other species of the genus. The plant, which lacks mature flowers, is geographically closest and morphologically most similar to
<taxonomicName id="19BCF5E1FFC45946FD8FF93304F78786" box="[603,702,1669,1693]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="45" pageNumber="86" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="ECC85270FFC45946FD8FF93304F78786" box="[603,702,1669,1693]" italics="true" pageId="45" pageNumber="86">L. koelzii</emphasis>
</taxonomicName>
. Like that species, it occurs in an apparently dry habitat and has seasonally deciduous foliage. Both of these characteristics suggest that the undetermined plant has four stamens. Other characteristics in common with
<taxonomicName id="19BCF5E1FFC65944FC71FF6A024E81E8" box="[933,1031,219,243]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="47" pageNumber="88" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="ECC85270FFC65944FC71FF6A024E81E8" box="[933,1031,219,243]" italics="true" pageId="47" pageNumber="88">L. koelzii</emphasis>
</taxonomicName>
include erect to flexuose eglandular trichomes on the rachis, peduncles, and pedicels; ovate-elliptic calyx lobes, and externally glabrous corollas. It differs from specimens of
<taxonomicName id="19BCF5E1FFC65944FCC9FEAA05C88028" box="[797,897,283,307]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="47" pageNumber="88" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="ECC85270FFC65944FCC9FEAA05C88028" box="[797,897,283,307]" italics="true" pageId="47" pageNumber="88">L. koelzii</emphasis>
</taxonomicName>
studied herein by having pubescent young stems, shorter calyces (
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long, although the flowers are not fully mature), and longer (
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) capsules that are pubescent only at the apex. Disposition of this plant awaits further studies and/or collections.
</paragraph>
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14.
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<emphasis id="ECC85270FFC75945FF2BF9E207EE877C" box="[255,423,1620,1639]" italics="true" pageId="46" pageNumber="87">Louteridium koelzii</emphasis>
</taxonomicName>
. A. Apex of shoot with flush of new leaves (
<emphasis id="ECC85270FFC75945FCFBF9E205B6877C" box="[815,1023,1620,1639]" italics="true" pageId="46" pageNumber="87">McVaugh &amp; Koelz 1797</emphasis>
). B. Branch with inflorescence in flower (
<emphasis id="ECC85270FFC75945FE9AF9DB0455879B" box="[334,540,1645,1665]" italics="true" pageId="46" pageNumber="87">McVaugh &amp; Koelz 1507</emphasis>
). C. Leaf (
<emphasis id="ECC85270FFC75945FDA8F9DB0502879B" box="[636,843,1645,1665]" italics="true" pageId="46" pageNumber="87">McVaugh &amp; Koelz 1797</emphasis>
). D. Corolla bud opened (
<emphasis id="ECC85270FFC75945FBFAF9DB06A98781" italics="true" pageId="46" pageNumber="87">McVaugh &amp; Koelz 1797</emphasis>
). E. Dehisced corolla with epipetalous stamens (
<emphasis id="ECC85270FFC75945FD76F93105788781" box="[674,817,1671,1690]" italics="true" pageId="46" pageNumber="87">McVaugh 26180</emphasis>
). F. Distal portion of style and stigma (
<emphasis id="ECC85270FFC75945FF56F916074687A8" box="[130,271,1696,1715]" italics="true" pageId="46" pageNumber="87">McVaugh 26180</emphasis>
). G. Partially dehisced capsule with calyx and pedicel (
<emphasis id="ECC85270FFC75945FCD7F91605D887A8" box="[771,913,1696,1715]" italics="true" pageId="46" pageNumber="87">McVaugh 26180</emphasis>
). H. Seed (
<emphasis id="ECC85270FFC75945FC2FF91602C187A8" box="[1019,1160,1696,1715]" italics="true" pageId="46" pageNumber="87">McVaugh 26180</emphasis>
). Drawn by Karin Douthit, copyright reserved to University of Michigan Herbarium, used with permission.
</paragraph>
</caption>
<paragraph id="DE038E62FFC65944FF7FFE2D04328308" blockId="47.[123,1174,219,532]" pageId="47" pageNumber="88">
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</emphasis>
.—
<materialsCitation id="6ED4843FFFC65944FD8DFE2D06C280C8" collectionCode="R" country="Mexico" location="12 - 13 km SW of Pihuamo" pageId="47" pageNumber="88" specimenCount="1" stateProvince="Jalisco">
<collectingCountry id="A6ABCEF2FFC65944FD8DFE2D048B80A8" box="[601,706,411,435]" name="Mexico" pageId="47" pageNumber="88">MEXICO</collectingCountry>
.
<emphasis id="ECC85270FFC65944FD00FE2D056880A8" bold="true" box="[724,801,411,435]" pageId="47" pageNumber="88">
<collectingRegion id="1C784080FFC65944FD00FE2D056880A8" box="[724,801,411,435]" country="Mexico" name="Jalisco" pageId="47" pageNumber="88">Jalisco</collectingRegion>
</emphasis>
: ca.
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SW of Pihuamo
</location>
,
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<emphasis id="ECC85270FFC65944FF5FFE0A06DB80C8" box="[139,146,444,467]" italics="true" pageId="47" pageNumber="88">.</emphasis>
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26180
</emphasis>
(
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</document>

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T.F. Daniel,
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Distr. Pochutla,
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!
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!
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!
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!
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!
<collectingCountry id="A6ABCEF2FFCD594FFED2FB0B076185CE" box="[262,296,1213,1237]" name="United States of America" pageId="36" pageNumber="77">US</collectingCountry>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFCD594DFF7FFB53051B80A8" lastPageId="38" lastPageNumber="79" pageId="36" pageNumber="77" type="description">
<paragraph id="DE038E62FFCD594EFF7FFB530515827B" blockId="36.[123,1174,1253,1726]" lastBlockId="37.[123,1174,219,864]" lastPageId="37" lastPageNumber="78" pageId="36" pageNumber="77">
Shrubs to trees to
<quantity id="19442387FFCD594FFEA4FB5307EE85E7" box="[368,423,1253,1277]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="77" unit="m" value="12.0">12 m</quantity>
tall, sometimes epipetric. Older (woody) stems quadrate, lenticellate, irregularly striate-sulcate, lacking trichomes; younger (herbaceous) stems subquadrate-sulcate, sparsely lenticellate, irregularly fissured, evenly pubescent with erect to flexuose simple and dendritic (sparse) eglandular trichomes &lt;0.1
<quantity id="19442387FFCD594FFDEBFAF304DB8447" box="[575,658,1349,1373]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="77" unit="mm" value="0.5">0.5 mm</quantity>
long. Leaves seasonally deciduous, ± clustered at apex of old growth or at apex of an otherwise leafless shoot of new growth, petiolate, petioles to
<quantity id="19442387FFCD594FFFAFFA33068E8487" box="[123,199,1413,1437]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="36" pageNumber="77" unit="mm" value="65.0">65 mm</quantity>
long, blades subsucculent, ovate to elliptic to broadly elliptic,
<quantity id="19442387FFCD594FFCAFFA3305B48487" box="[891,1021,1413,1437]" metricMagnitude="-1" metricUnit="m" metricValue="1.33" metricValueMax="1.9" metricValueMin="0.76" pageId="36" pageNumber="77" unit="mm" value="133.0" valueMax="190.0" valueMin="76.0">76190 mm</quantity>
long,
<quantity id="19442387FFCD594FFB91FA3306ED84A7" metricMagnitude="-2" metricUnit="m" metricValue="8.1" metricValueMax="12.2" metricValueMin="4.0" pageId="36" pageNumber="77" unit="mm" value="81.0" valueMax="122.0" valueMin="40.0">40122 mm</quantity>
wide, 1.42.5 × longer than wide, apiculate to acuminate at apex, rounded to acute to attenuate at base, adaxial surface pubescent with simple and dendritic eglandular trichomes, trichomes soon becoming ± restricted to proximal portion or to midvein, abaxial surface pubescent (especially along veins) with dendritic trichomes to
<quantity id="19442387FFCD594FFDEDF9B304C48706" box="[569,653,1541,1565]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="77" unit="mm" value="0.5">0.5 mm</quantity>
long, midvein often pinkish or reddish, margin entire (sometimes undulate and appearing subcrenate). Inflorescence a terminal subsessile to pedunculate raceme to
<quantity id="19442387FFCD594FFEA6F9F307838746" box="[370,458,1605,1629]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="36" pageNumber="77" unit="mm" value="220.0">220 mm</quantity>
long, peduncle to
<quantity id="19442387FFCD594FFD46F9F304958746" box="[658,732,1605,1629]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="77" unit="mm" value="50.0">50 mm</quantity>
long, pubescent like young stems, rachis pubescent like young stems; dichasia opposite or alternate, sessile, 1-flowered, to
<quantity id="19442387FFCD594FFBD9F9D302128766" box="[1037,1115,1637,1661]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="36" pageNumber="77" unit="mm" value="47.0">47 mm</quantity>
long (excluding corollas). Bracts caducous, lanceolate to lance-ovate,
<quantity id="19442387FFCD594FFCE5F93305EA8786" box="[817,931,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="36" pageNumber="77" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">1116 mm</quantity>
long, 2.
<quantity id="19442387FFCD594FFC2FF933021A8786" box="[1019,1107,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="36" pageNumber="77" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
wide, abaxially pubescent with simple and dendritic trichomes 0.05
<quantity id="19442387FFCD594FFCC8F910052787A6" box="[796,878,1702,1726]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="77" unit="mm" value="0.2">0.2 mm</quantity>
long. Bracteoles caducous, lanceolate to lance-elliptic,
<quantity id="19442387FFCC594EFE70FF6D07B481E9" box="[420,509,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="37" pageNumber="78" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
long,
<quantity id="19442387FFCC594EFD97FF6D04D581E9" box="[579,668,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="37" pageNumber="78" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide, abaxially pubescent like bracts. Flowers pedicellate, pedicels
<quantity id="19442387FFCC594EFE8AFF4A079B8008" box="[350,466,252,276]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="4.6" metricValueMin="2.1" pageId="37" pageNumber="78" unit="mm" value="33.5" valueMax="46.0" valueMin="21.0">2146 mm</quantity>
long, pubescent like rachis or with the trichomes to
<quantity id="19442387FFCC594EFBC3FF4A021F8008" box="[1047,1110,252,276]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="78" unit="mm" value="1.0">1 mm</quantity>
long. Calyx
<quantity id="19442387FFCC594EFF16FEAA077C802F" box="[194,309,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="37" pageNumber="78" unit="mm" value="24.5" valueMax="32.0" valueMin="17.0">1732 mm</quantity>
long, lobes fused at base for 1
<quantity id="19442387FFCC594EFD5BFEAA04A8802F" box="[655,737,284,308]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="78" unit="mm" value="1.5">1.5 mm</quantity>
, subhomomorphic to subheteromorphic, membranaceous, subelliptic to ovate-elliptic to subrhombic-obovate, rounded to acute at apex, abaxially pubescent with mostly dendritic trichomes 0.1
<quantity id="19442387FFCC594EFCD5FEE80511806E" box="[769,856,350,374]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="78" unit="mm" value="0.5">0.5 mm</quantity>
long, posterior lobe planar,
<quantity id="19442387FFCC594EFFAFFEC906BB808D" box="[123,242,383,407]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="37" pageNumber="78" unit="mm" value="24.5" valueMax="32.0" valueMin="17.0">1732 mm</quantity>
long,
<quantity id="19442387FFCC594EFE95FEC907F1808D" box="[321,440,383,407]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.9" metricValueMin="1.0" pageId="37" pageNumber="78" unit="mm" value="14.5" valueMax="19.0" valueMin="10.0">1019 mm</quantity>
wide, usually slightly larger and sometimes more conspicuously venose than lateral lobes, major veins often maroon, lateral lobes
<quantity id="19442387FFCC594EFCECFE2905E380AC" box="[824,938,415,439]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="3.1" metricValueMin="2.0" pageId="37" pageNumber="78" unit="mm" value="25.5" valueMax="31.0" valueMin="20.0">2031 mm</quantity>
long,
<quantity id="19442387FFCC594EFC3BFE29021D80AC" box="[1007,1108,415,439]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="37" pageNumber="78" unit="mm" value="13.0" valueMax="18.0" valueMin="8.0">818 mm</quantity>
wide. Corolla light green or greenish yellow, sometimes with maroon on limb (especially at base of lobes) and distal portion of throat, externally glabrous (inconspicuously glandular punctate but lacking elongate trichomes),
<quantity id="19442387FFCC594EFE14FDB7047E8302" box="[448,567,513,537]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="6.2" metricValueMin="5.0" pageId="37" pageNumber="78" unit="mm" value="56.0" valueMax="62.0" valueMin="50.0">5062 mm</quantity>
long, tube
<quantity id="19442387FFCC594EFD6BFDB7057E8302" box="[703,823,513,537]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="37" pageNumber="78" unit="mm" value="36.0" valueMax="37.0" valueMin="35.0">3537 mm</quantity>
long, narrow proximal portion
<quantity id="19442387FFCC594EFFAFFD9406A48322" box="[123,237,546,570]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="37" pageNumber="78" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">1115 mm</quantity>
long, 610.
<quantity id="19442387FFCC594EFEBBFD9407E48322" box="[367,429,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="78" unit="mm" value="5.0">5 mm</quantity>
diameter near midpoint, throat
<quantity id="19442387FFCC594EFCD0FD94053E8322" box="[772,887,546,570]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="37" pageNumber="78" unit="mm" value="22.0" valueMax="24.0" valueMin="20.0">2024 mm</quantity>
long,
<quantity id="19442387FFCC594EFC69FD9402798322" box="[957,1072,546,570]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="37" pageNumber="78" unit="mm" value="30.0" valueMax="35.0" valueMin="25.0">2535 mm</quantity>
diameter at mouth, lobes recurved to recoiled, broadly ovate to subtriangular,
<quantity id="19442387FFCC594EFCB1FDF505938341" box="[869,986,579,603]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="37" pageNumber="78" unit="mm" value="16.5" valueMax="20.0" valueMin="13.0">1320 mm</quantity>
long,
<quantity id="19442387FFCC594EFBF5FDF502DC8341" box="[1057,1173,579,603]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="2.1" metricValueMin="1.0" pageId="37" pageNumber="78" unit="mm" value="15.5" valueMax="21.0" valueMin="10.0">1021 mm</quantity>
wide, entire at apex. Stamens 4,
<quantity id="19442387FFCC594EFE06FDD5040D8360" box="[466,580,611,635]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="37" pageNumber="78" unit="mm" value="70.0" valueMax="80.0" valueMin="60.0">6080 mm</quantity>
long, filaments glabrous distally, glabrous or pubescent with eglandular trichomes near base, thecae 810.
<quantity id="19442387FFCC594EFD74FD3204968380" box="[672,735,644,668]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="78" unit="mm" value="5.0">5 mm</quantity>
long; staminode 1, rodlike, 0.
<quantity id="19442387FFCC594EFBF9FD3202DC8380" box="[1069,1173,644,668]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="4.0" metricValueMin="0.6" pageId="37" pageNumber="78" unit="mm" value="23.0" valueMax="40.0" valueMin="6.0">640 mm</quantity>
long. Style
<quantity id="19442387FFCC594EFF21FD13073D83A7" box="[245,372,677,701]" metricMagnitude="-2" metricUnit="m" metricValue="8.55" metricValueMax="10.1" metricValueMin="7.0" pageId="37" pageNumber="78" unit="mm" value="85.5" valueMax="101.0" valueMin="70.0">70101 mm</quantity>
long, distally glabrous, pubescent with eglandular and glandular trichomes near base, stigma equally 2-lobed, lobes broadly elliptic to broadly ovate-triangular,
<quantity id="19442387FFCC594EFBD4FD73021183C6" box="[1024,1112,709,733]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="37" pageNumber="78" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, 1
<quantity id="19442387FFCC594EFF41FD5006A183E5" box="[149,232,742,766]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="78" unit="mm" value="1.4">1.4 mm</quantity>
wide. Capsule
<quantity id="19442387FFCC594EFE45FD50044C83E5" box="[401,517,742,766]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.8" metricValueMin="2.1" pageId="37" pageNumber="78" unit="mm" value="24.5" valueMax="28.0" valueMin="21.0">2128 mm</quantity>
long, 6.59.
<quantity id="19442387FFCC594EFD5BFD50048483E5" box="[655,717,742,766]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="78" unit="mm" value="5.0">5 mm</quantity>
in diameter, densely pubescent with erect glandular trichomes 0.05
<quantity id="19442387FFCC594EFE40FCB107AF8205" box="[404,486,775,799]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="78" unit="mm" value="0.5">0.5 mm</quantity>
long and with an overstory (sometimes sparse) of erect to flexuose (sometimes dendritic) eglandular trichomes to
<quantity id="19442387FFCC594EFD70FC9E04BC8224" box="[676,757,808,832]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="78" unit="mm" value="1.4">1.4 mm</quantity>
long, stipe 2.5
<quantity id="19442387FFCC594EFC75FC9E05BB8224" box="[929,1010,807,832]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="37" pageNumber="78" unit="mm" value="3.5">3.5 mm</quantity>
long. Seeds up to 16 per capsule, 5.
<quantity id="19442387FFCC594EFE80FCFE07E5827B" box="[340,428,840,864]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="37" pageNumber="78" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">27 mm</quantity>
long, 56.
<quantity id="19442387FFCC594EFDF4FCFE0417827B" box="[544,606,840,864]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="78" unit="mm" value="4.0">4 mm</quantity>
wide, surfaces smooth.
</paragraph>
<paragraph id="DE038E62FFCC594EFF7FFCC204C3856B" blockId="37.[123,1174,884,1726]" pageId="37" pageNumber="78">
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<emphasis id="ECC85270FFCC594EFF7FFCC207788290" bold="true" box="[171,305,884,908]" pageId="37" pageNumber="78">PHENOLOGY</emphasis>
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.— Flowering: FebruaryMarch; fruiting: MarchApril. Based on field observations and cultivated plants (
<emphasis id="ECC85270FFCC594EFE75FC2304CC82B6" box="[417,645,917,941]" italics="true" pageId="37" pageNumber="78">Daniel et al. 11894cv</emphasis>
), flowering occurs on leafless (or nearly leafless) plants during the dry season. Near the end of the dry season (e.g., late MarchApril) when flowering is waning and fruits are mature, a new flush of vegetative growth appears from axils of clustered leaf scars at the base of the inflorescence, which eventually falls away. New internodal stem elongation (e.g., the young stems of the description above) takes place from the axil of a leaf scar on the old growth and terminates in a cluster of new leaves and/or between at least one of the pairs of leaves in the cluster and the remaining cluster.
</paragraph>
<paragraph id="DE038E62FFCC594EFF7FFBCC05C287A6" blockId="37.[123,1174,884,1726]" pageId="37" pageNumber="78">
Field observations of
<emphasis id="ECC85270FFCC594EFE45FBCC04158589" box="[401,604,1146,1170]" italics="true" pageId="37" pageNumber="78">Daniel et al. 11894</emphasis>
over three days (
<date id="AA02A8A2FFCC594EFCC7FBCC05BE8589" box="[787,1015,1146,1170]" pageId="37" pageNumber="78" value="2012-02-24" valueMax="2012-02-26" valueMin="2012-02-24">2426 February 2012</date>
) revealed: day 1 between 08:0009:00 (light) — corollas mostly fallen, only a few from previous night still attached to tree; day 2 between 19:0020:00 (dark) — all corollas open, ca. 100 seen, bats active around plants but none seen visiting flowers, no floral odor detected and no nectar visible in saccate tube of undissected corollas, stigma extended ca.
<quantity id="19442387FFCC594EFD16FB4B04B1840F" box="[706,760,1277,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="78" unit="cm" value="1.0">1 cm</quantity>
beyond anthers on fresh flowers, stigma of 7 flowers examined for pollen (all pollinated); day 3 between 17:3018:30 (light) — many corollas open and many others still in bud, corollas open fully (including recurving of corolla lobes) in 15 to 20 seconds, open flowers actively visited by Cinnamon Hummingbird (
<taxonomicName id="19BCF5E1FFCC594EFBE3FAE806FF848C" baseAuthorityName="Delattre" baseAuthorityYear="1843" class="Aves" family="Trochilidae" genus="Amazilia" kingdom="Animalia" order="Apodiformes" pageId="37" pageNumber="78" phylum="Chordata" rank="species" species="rutila">
<emphasis id="ECC85270FFCC594EFBE3FAE806FF848C" italics="true" pageId="37" pageNumber="78">Amazilia rutila</emphasis>
</taxonomicName>
, species det. by Jeff Chemnick), birds probe for nectar once or twice at same flower before moving on, nectar is located behind a barrier (seen in dissected flowers) at the base of the tube, nectar not visible in the saccate throat, birds appear to contact anthers with head or back and presumably contact the stigma on some visits as well, birds visit between 510 flowers on a single plant or on multiple plants before moving away from an area or resting on a branch, 2 flowers were observed to open and when subsequently visited by a hummingbird (one flower visited once, other flower visited twice) they were checked for pollination (no pollen observed on stigma of either), small bees or flies also observed visiting flowers but they only contact anthers, buds continue opening until full darkness (at 18:30); day 3 between 18:3020:30 (dark) — hummingbirds no longer active, bats very active around plants but none observed visiting flowers.
</paragraph>
<paragraph id="DE038E62FFCF594DFF7FFF6D051B80A8" blockId="38.[123,1174,219,1205]" pageId="38" pageNumber="79">
On a single flower of
<emphasis id="ECC85270FFCF594DFE4BFF6D04C081E8" box="[415,649,219,243]" italics="true" pageId="38" pageNumber="79">Daniel et al. 11894cv</emphasis>
grown in San Francisco, corolla lobes began to separate at 15:50 and the corolla was fully open (lobes spreading 90° with respect to the throat or reflexed) with the stamens and style fully exserted by 16:15. By 18:00, all corolla lobes were at least partially recoiled. At 23:00, 6.5 μl of nectar was encountered in the nectar chamber, and the stigma was dusted with pollen from the anthers. At 07:15 the next day, 72.1 μl of nectar was recovered from the nectar chamber. The corolla abscised and fell from the persisting flower at 07:45. Thus, the corolla of the pollinated flower persisted for ca. 16 hours.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFCF594DFF7FFE0A053F8368" pageId="38" pageNumber="79" type="distribution">
<paragraph id="DE038E62FFCF594DFF7FFE0A02718308" blockId="38.[123,1174,219,1205]" pageId="38" pageNumber="79">
<emphasis id="ECC85270FFCF594DFF7FFE0A079D80C9" bold="true" box="[171,468,444,468]" pageId="38" pageNumber="79">
<smallCapsWord id="D8E518BEFFCF594DFF7FFE0A070A80C9" baselines="462,461" box="[171,323,444,468]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="38" pageNumber="79">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="D8E518BEFFCF594DFE9DFE09073D80C9" baselines="461" box="[329,372,447,466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="38" pageNumber="79">AND</smallCapsWord>
<smallCapsWord id="D8E518BEFFCF594DFEAFFE09079D80C9" baselines="461" box="[379,468,447,466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="habitat" pageId="38" pageNumber="79">HABITAT</smallCapsWord>
</emphasis>
.—
<materialsCitation id="6ED4843FFFCF594DFE29FE0D02718308" county="Southern" elevation="675" location="Isthmus of Tehuantepec" municipality="Sierra Madre del Sur" pageId="38" pageNumber="79" specimenCount="1" stateProvince="Mexico">
<collectingCounty id="3762F6EEFFCF594DFE29FE0D041480C8" box="[509,605,443,467]" pageId="38" pageNumber="79">Southern</collectingCounty>
<collectingRegion id="1C784080FFCF594DFDB0FE0D04FC80C8" box="[612,693,443,467]" country="Mexico" name="Mexico" pageId="38" pageNumber="79">Mexico</collectingRegion>
(
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;
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) in the
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and the
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; plants occur on limestone (often karstic) slopes of streams in tropical deciduous forests and tropical subdeciduous forests at elevations from
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.
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</paragraph>
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.—
<figureCitation id="468792E7FFCF594DFEADFDAD07B68328" box="[377,511,539,564]" captionStart="FIGURE" captionStartId="5.[155,167,1561,1580]" captionText="FIGURE 2. Louteridium brevicalyx (A-E) and L. dendropilosum (F-L). A. Habit and habitat. B. Trunk with adventitious roots on rocks. C. Six-year old cultivated plant with clusters of new leaves. D. Flower. E. Shoot apex showing clustered leaf scars. F. Base of trunk with adventitious roots on rocks. G. Six-year old cultivated plant with clusters of new leaves. H. Shoot apex showing clustered leaf scars. I Young inflorescence with bracts, bracteoles, and flower buds. J. Flower. K. Corolla showing invagination along base of throat. L. Internal view of corolla showing invaginations along base of throat. (A, B, Daniel &amp; Steinmann 11913; C-E, Daniel &amp; Steinmann 11913cv; F, Daniel et al. 11784; G-L, Daniel et al. 11894cv). Photos by T. Daniel." pageId="38" pageNumber="79">Figure 2FL</figureCitation>
;
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, fig. 8).
</paragraph>
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.— A discussion and preliminary conservation assessment of Endangered (En) was proposed for this species (B1, a, b;
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) by
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.
</paragraph>
</subSubSection>
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<emphasis id="ECC85270FFCF594DFF7FFDCA07638389" bold="true" box="[171,298,636,660]" pageId="38" pageNumber="79">DISCUSSION</emphasis>
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.—
<taxonomicName id="19BCF5E1FFCF594DFE81FDCA04CB838F" box="[341,642,636,660]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="38" pageNumber="79" phylum="Tracheophyta" rank="species" species="dendropilosum">
<emphasis id="ECC85270FFCF594DFE81FDCA04CB838F" box="[341,642,636,660]" italics="true" pageId="38" pageNumber="79">Louteridium dendropilosum</emphasis>
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is unique among congeners by its dendritic trichomes, which are present on both vegetative and reproductive organs. Trichomes of other species may consist of one or more cells and be either glandular or eglandular, but they are not branched. Bracts and bracteoles are caducous prior to maturation of the flowers associated with them; thus, they are rarely encountered and are not present on any of the wild-collected specimens. The data on their shape, size, and pubescence noted above were taken just prior to their dehiscence from the young inflorescence of a cultivated plant (
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).
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.—
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:
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,
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,
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, ca.
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downstream from jct.
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SE of Rancho Dioon
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toward
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, et al. 11894
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(
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),
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(
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,
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), cultivated plants of
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grown from seeds in
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,
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,
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(
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);
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,
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[sic]
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,
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, antena microondas [ca.
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.
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1396
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(
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,
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);
<collectorName id="7349EBB4FFCF594DFECCFB8A07FE854F" box="[280,439,1084,1108]" pageId="38" pageNumber="79">Distr. Pochutla</collectorName>
,
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,
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</quantity>
de la terracería sobre la vereda rumbo
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,
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,
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<emphasis id="ECC85270FFCF594DFD1DFBEB0490856F" box="[713,729,1117,1140]" italics="true" pageId="38" pageNumber="79">A</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFCF594DFD0DFBEB0519856F" box="[729,848,1117,1140]" italics="true" pageId="38" pageNumber="79">
.
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<location id="DB63D8B9FFCF594DFD3DFBEB057A856F" LSID="urn:lsid:plazi:treatment:56153F74FFCD594DFF7FFBAA053785AF:DB63D8B9FFCF594DFD3DFBEB057A856F" box="[745,819,1117,1140]" country="Mexico" name="Saynes" pageId="38" pageNumber="79" stateProvince="Oaxaca">Saynes</location>
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</materialsCitation>
.
</emphasis>
<materialsCitation id="6ED4843FFFCF594DFC8DFBEB046085AF" collectionCode="MEXU, SERO, S" collectorName="Distr. Juchitan &amp; Mpio. El Barrio &amp; Cerro Palmasola" country="Mexico" latitude="16.742367" longLatPrecision="1" longitude="-95.093346" pageId="38" pageNumber="79" specimenCount="1" stateProvince="Oaxaca">
<emphasis id="ECC85270FFCF594DFC8DFBEB0599856E" box="[857,976,1116,1141]" italics="true" pageId="38" pageNumber="79">et al. 3831</emphasis>
(
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,
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);
<collectorName id="7349EBB4FFCF594DFFAFFBCA075E858F" box="[123,279,1148,1172]" pageId="38" pageNumber="79">Distr. Juchitán</collectorName>
,
<collectorName id="7349EBB4FFCF594DFEF2FBCA079A858F" box="[294,467,1148,1172]" pageId="38" pageNumber="79">Mpio. El Barrio</collectorName>
, parte alta del
<collectorName id="7349EBB4FFCF594DFDACFBCB0562858F" box="[632,811,1148,1173]" pageId="38" pageNumber="79">Cerro Palmasola</collectorName>
, junto a la antena de microondas [ca.
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,
<geoCoordinate id="BB88E8A5FFCF594DFE8FFB2A044C85AF" box="[347,517,1180,1205]" degrees="095" direction="west" minutes="5" orientation="longitude" pageId="38" pageNumber="79" precision="1" seconds="36. 04" value="-95.093346">095°536.04”W</geoCoordinate>
],
<collectionCode id="B8AD16A7FFCF594DFDC8FB2B046085AF" box="[540,553,1181,1204]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="38" pageNumber="79" type="Herbarium">
<emphasis id="ECC85270FFCF594DFDC8FB2B046085AF" box="[540,553,1181,1204]" italics="true" pageId="38" pageNumber="79">S</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFCF594DFDFDFB2B04B885AF" box="[553,753,1180,1205]" italics="true" pageId="38" pageNumber="79">
.
<materialsCitation id="6ED4843FFFCF594DFDE3FB2B04E485AF" box="[567,685,1181,1205]" collectionCode="R" country="Mexico" location="Zamudio" pageId="38" pageNumber="79" specimenCount="1" stateProvince="Oaxaca">
<location id="DB63D8B9FFCF594DFDE3FB2B04DF85AE" LSID="urn:lsid:plazi:treatment:56153F74FFCD594DFF7FFBAA053785AF:DB63D8B9FFCF594DFDE3FB2B04DF85AE" box="[567,662,1181,1205]" country="Mexico" name="Zamudio" pageId="38" pageNumber="79" stateProvince="Oaxaca">Zamudio</location>
<collectionCode id="B8AD16A7FFCF594DFD49FB2B04E485AF" box="[669,685,1181,1204]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="79">R</collectionCode>
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. 6352
</emphasis>
(CAS, IEB).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="25C22A403C37FA66DE57F0D84A69641D">Louteridium (Acanthaceae: Acanthoideae: Ruellieae: Trichantherinae): Taxonomy, Phylogeny, Reproductive Biology, and Conservation</mods:title>
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<subSubSection id="96A6DDE9FFD15953FF7FFEEB021980AE" pageId="56" pageNumber="97" type="nomenclature">
<paragraph id="DE038E62FFD15953FF7FFEEB021980AE" blockId="56.[123,1174,348,438]" pageId="56" pageNumber="97">
<emphasis id="ECC85270FFD15953FF7FFEEB06F6806E" bold="true" box="[171,191,349,373]" pageId="56" pageNumber="97">9.</emphasis>
<treatmentCitation id="5F1DA873FFD15953FF1CFEEA05F3806E" author="Brandegee" box="[200,954,348,373]" journal="Univ. Calif. Publ. Bot." page="68" pageId="56" pageNumber="97" score="66.63618723694691" volume="6" year="1914">
<taxonomicName id="19BCF5E1FFD15953FF1CFEEA0572806E" ID-CoL="6QNQQ" authority="Brandegee, Univ. Calif. Publ. Bot." box="[200,827,348,373]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="56" pageNumber="97" phylum="Tracheophyta" rank="species" species="purpusii">
<emphasis id="ECC85270FFD15953FF1CFEEA07F1806E" bold="true" box="[200,440,348,373]" italics="true" pageId="56" pageNumber="97">Louteridium purpusii</emphasis>
<materialsCitation id="6ED4843FFFD15953FE16FEEB0572806E" box="[450,827,349,373]" collectingDate="1913-08" collectionCode="TYPE, C, UC, BM, F, GH, MO, NY" country="Mexico" latitude="15.173594" location="Finca Irlanda" longLatPrecision="1" longitude="-92.336174" pageId="56" pageNumber="97" specimenCount="2" stateProvince="Chiapas" typeStatus="holotype">
Brandegee,
<emphasis id="ECC85270FFD15953FD90FEE80572806E" box="[580,827,349,373]" italics="true" pageId="56" pageNumber="97">Univ. Calif. Publ. Bot.</emphasis>
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</taxonomicName>
6:68. 1914
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.
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<emphasis id="ECC85270FFD15953FC1EFEEB024D806F" bold="true" box="[970,1028,349,373]" pageId="56" pageNumber="97">TYPE</emphasis>
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.—
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.
<emphasis id="ECC85270FFD15953FFAFFECB0691808E" bold="true" box="[123,216,381,405]" pageId="56" pageNumber="97">
<collectingRegion id="1C784080FFD15953FFAFFECB0691808E" box="[123,216,381,405]" country="Mexico" name="Chiapas" pageId="56" pageNumber="97">Chiapas</collectingRegion>
</emphasis>
:
<location id="DB63D8B9FFD15953FF31FECB073B808E" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD15953FF31FECB073B808E" box="[229,370,381,405]" country="Mexico" latitude="15.173594" longLatPrecision="1" longitude="-92.336174" name="Finca Irlanda" pageId="56" pageNumber="97" stateProvince="Chiapas">Finca Irlanda</location>
[ca.
<geoCoordinate id="BB88E8A5FFD15953FE71FECB0401808E" box="[421,584,381,405]" degrees="15" direction="north" minutes="10" orientation="latitude" pageId="56" pageNumber="97" precision="1" seconds="24. 94" value="15.173594">15°1024.94”N</geoCoordinate>
,
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<collectingDate id="BA46514AFFD15953FCF5FEC805C4808E" box="[801,909,381,405]" pageId="56" pageNumber="97" value="1913-08">VIII-1913</collectingDate>
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(flr, frt),
<emphasis id="ECC85270FFD15953FC3BFEC802DC808E" box="[1007,1173,381,405]" italics="true" pageId="56" pageNumber="97">C. Purpus 6969</emphasis>
(
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: UC!;
<typeStatus id="010730C0FFD15953FEF8FE2807CA80AD" box="[300,387,414,438]" pageId="56" pageNumber="97" type="isotype">isotypes</typeStatus>
: BM!, EAP-image seen, F!, GH!, MO!, NY!, US!, Z-image seen).
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFD15953FF7FFE7F05498767" pageId="56" pageNumber="97" type="description">
<paragraph id="DE038E62FFD15953FF7FFE7F04E3874B" blockId="56.[123,1174,457,1617]" pageId="56" pageNumber="97">
Shrubs or trees to
<quantity id="19442387FFD15953FEBBFE7F07D080FA" box="[367,409,457,481]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="56" pageNumber="97" unit="m" value="9.0">9 m</quantity>
tall, terrestrial. Older (woody) stems subquadrate-sulcate, ± scurfy and lenticellate, often puberulent like young stems; younger (herbaceous) stems quadrate to quadrate-sulcate and ± irregularly striate, glabrous or ± evenly puberulent with erect to retrorse to antrorse eglandular trichomes to
<quantity id="19442387FFD15953FE56FD9D07AA8359" box="[386,483,554,579]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="56" pageNumber="97" unit="mm" value="0.05">0.05 mm</quantity>
long. Leaves apparently not all seasonally deciduous, ± evenly distributed along young stems, petiolate, petioles to
<quantity id="19442387FFD15953FD61FDFD05488378" box="[693,769,587,611]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="56" pageNumber="97" unit="mm" value="40.0">40 mm</quantity>
long, blades membranaceous, ovate-elliptic to obovate-elliptic to obovate,
<quantity id="19442387FFD15953FDC0FDDA04EB8398" box="[532,674,620,644]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.8" metricValueMin="1.3" pageId="56" pageNumber="97" unit="mm" value="255.0" valueMax="380.0" valueMin="130.0">130380 mm</quantity>
long,
<quantity id="19442387FFD15953FD33FDDA052F8398" box="[743,870,619,644]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="11.5" metricValueMin="4.0" pageId="56" pageNumber="97" unit="mm" value="77.5" valueMax="115.0" valueMin="40.0">40115 mm</quantity>
wide, 2.14.3 × longer than wide, acuminate to acute-apiculate at apex, acute to attenuate at base, adaxial surface glabrous or puberulent like young stems, abaxial surface paler than abaxial surface, glabrous or pubescent with mostly antrorse eglandular trichomes 0.05
<quantity id="19442387FFD15953FD9DFD7B04D383FF" box="[585,666,717,741]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="56" pageNumber="97" unit="mm" value="0.1">0.1 mm</quantity>
long along major veins, margin often undulate, entire to sinuate to crenate to crenate-dentate. Inflorescence a terminal pedunculate raceme to racemose thyrse to
<quantity id="19442387FFD15953FECBFCB8072B823E" box="[287,354,782,806]" metricMagnitude="-1" metricUnit="m" metricValue="6.3" pageId="56" pageNumber="97" unit="cm" value="63.0">63 cm</quantity>
long (including peduncles and excluding corollas), peduncle to
<quantity id="19442387FFD15953FBC1FCB80211823E" box="[1045,1112,782,806]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="97" unit="cm" value="20.0">20 cm</quantity>
long, glabrous or pubescent like young stems, rachis puberulent with (retrorse to) erect to flexuose to antrorse eglandular trichomes to
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long; dichasia modified by very short expansion between pairs of succeeding flowers with the congested dichasial axis becoming a ± fan-shaped to multi- and tortuously-branched racemelike lateral short-shoot to
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long, opposite (and alternate?), sessile to subsessile, 3many-flowered, to
<quantity id="19442387FFD15953FDEAFC0604C182DC" box="[574,648,944,968]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="56" pageNumber="97" unit="mm" value="60.0">60 mm</quantity>
long (excluding corollas), dichasial peduncles (if present) to
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long, pubescent like rachis. Bracts caducous, not seen. Bracteoles and secondary bracteoles caducous, lance-linear to linear-elliptic,
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long, 12.
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wide, abaxially pubescent like rachis. Flowers pedicellate, pedicels to
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long, puberulent with erect to retrorse (to antrorse) eglandular trichomes to
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long. Calyx
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long, lobes distinct or fused at base up to
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, heteromorphic, membranaceous, abaxially puberulent like pedicels and sometimes with a few sparse stipitate glands up to
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long as well, also inconspicuously glandular-punctate, posterior lobe conduplicate, ovate to elliptic to obovate,
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long, slightly shorter than to slightly longer than lateral lobes, 8.
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wide, acuminate to falcate at apex, lateral lobes sublunate,
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long,
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wide, acute to subfalcate at apex. Corolla cream to yellow-greenish and tinged with maroon,
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long, externally pubescent with stipitate glandular trichomes 0.05
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long (sometimes with flexuose eglandular trichomes to
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long on or near margins of lobes as well), tube
<quantity id="19442387FFD15953FDA2FA8304A08456" box="[630,745,1333,1358]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="56" pageNumber="97" unit="mm" value="29.0" valueMax="33.0" valueMin="25.0">2533 mm</quantity>
long, narrow proximal portion
<quantity id="19442387FFD15953FBE9FA8002DC8456" box="[1085,1173,1333,1358]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="56" pageNumber="97" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long,
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in diameter near midpoint, throat
<quantity id="19442387FFD15953FDB4FAE0049B8476" box="[608,722,1366,1390]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="56" pageNumber="97" unit="mm" value="26.0" valueMax="30.0" valueMin="22.0">2230 mm</quantity>
long,
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diameter at mouth, lobes spreading to recurved, triangular to ovate,
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long,
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wide, rounded at apex. Stamens 2,
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long, filaments glabrous distally, pubescent proximally with eglandular trichomes, thecae
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long; staminodes (if present) not seen. Style
<quantity id="19442387FFD15953FC55FA0105BD84D4" box="[897,1012,1463,1487]" metricMagnitude="-2" metricUnit="m" metricValue="8.85" metricValueMax="9.2" metricValueMin="8.5" pageId="56" pageNumber="97" unit="mm" value="88.5" valueMax="92.0" valueMin="85.0">8592 mm</quantity>
long, glabrous distally (not seen proximally), stigma equally 2-lobed, lobes elliptic, 1.4
<quantity id="19442387FFD15953FC77FA6E05AA84F4" box="[931,995,1496,1520]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="97" unit="mm" value="2.0">2 mm</quantity>
long, width not determined. Capsule
<quantity id="19442387FFD15953FE88FA4E0784870B" box="[348,461,1528,1552]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="56" pageNumber="97" unit="mm" value="26.0" valueMax="30.0" valueMin="22.0">2230 mm</quantity>
long, 57.
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in diameter, densely pubescent with stipitate glandular trichomes (0.05) 0.1
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long, stipe 3
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long. Seeds up to 12 per capsule, 4.5
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long, 4.5
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wide, surfaces smooth.
</paragraph>
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<emphasis id="ECC85270FFD15953FF7FF9D307788760" bold="true" box="[171,305,1637,1661]" pageId="56" pageNumber="97">PHENOLOGY</emphasis>
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.— Flowering and fruiting: AugustMarch.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFD15952FF7FF9330511838E" lastPageId="57" lastPageNumber="98" pageId="56" pageNumber="97" type="distribution">
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<smallCapsWord id="D8E518BEFFD15953FEAEF93E079A8780" baselines="1686" box="[378,467,1672,1691]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="habitat" pageId="56" pageNumber="97">HABITAT</smallCapsWord>
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.— Southern Mexico (Chiapas),
<collectingCountry id="A6ABCEF2FFD15953FCFBF93305EA8786" box="[815,931,1669,1693]" name="Guatemala" pageId="56" pageNumber="97">Guatemala</collectingCountry>
(
<figureCitation id="468792E7FFD15953FC66F93305B78786" box="[946,1022,1669,1693]" captionStart="FIGURE" captionStartId="20.[161,173,1071,1090]" captionTargetBox="[124,1173,223,1050]" captionTargetId="figure-1373@20.[120,1176,216,1055]" captionTargetPageId="20" captionText="FIGURE 10. Map of southern Mexico (showing states) and northern Central America with distributions of Louteridium chartaceum, L. donnell-smithii, and L. purpusii. Symbols may pertain to multiple collections." figureDoi="http://doi.org/10.5281/zenodo.13155729" httpUri="https://zenodo.org/record/13155729/files/figure.png" pageId="56" pageNumber="97">Fig. 10</figureCitation>
); plants occur in wooded ravines in montane rain forests on the Pacific versant at elevations from
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.
</paragraph>
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<emphasis id="ECC85270FFD05952FF7FFF6D070C81EA" bold="true" box="[171,325,219,243]" pageId="57" pageNumber="98">ILLUSTRATION</emphasis>
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.—
<figureCitation id="468792E7FFD05952FEBAFF6D079581E9" box="[366,476,219,243]" captionStart="FIGURE" captionStartId="10.[155,167,1596,1615]" captionText="FIGURE 4 (left). Flowers of Louteridium spp. A, B. L. costaricense (González 1977), lateral and frontal views. C. L. donnell-smithii (Daniel &amp; Véliz 11337), cream to green-yellow colored form. D. L. donnell-smithii (Daniel et al. 11356), chestnut to maroon colored form. E. L. mexicanum (Breedlove &amp; Daniel 70879cv). F. L. purpusii (Velásquez et al. 658). G. L. parayi (Daniel &amp; Wendt 5805) showing basally saccate dorsal calyx lobe on two buds at left. H. L. parayi (Breedlove 24816). Photos by D. Breedlove (H), T. Daniel (C-E, G), J. González (A-B), and L. Velásquez (F), used with permission." pageId="57" pageNumber="98">
Figure
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</paragraph>
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.— “palo de agua,” (
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Quarles
<taxonomicName id="19BCF5E1FFD05952FDA2FF4A04A38008" authority="Ufford" authorityName="Ufford" box="[630,746,251,275]" class="Magnoliopsida" family="Acanthaceae" genus="Harpochilus" kingdom="Plantae" order="Lamiales" pageId="57" pageNumber="98" phylum="Tracheophyta" rank="species" species="van">van Ufford</taxonomicName>
137
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).
</paragraph>
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.—
<taxonomicName id="19BCF5E1FFD05952FEACFEAD04148028" box="[376,605,283,307]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="57" pageNumber="98" phylum="Tracheophyta" rank="species" species="purpusii">
<emphasis id="ECC85270FFD05952FEACFEAD04148028" box="[376,605,283,307]" italics="true" pageId="57" pageNumber="98">Louteridium purpusii</emphasis>
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has an EOO of
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. Two prominent threats are evident throughout most or all of its known distribution, coffee agriculture and volcanic eruptions. Numerous active volcanos lie within and adjacent to the entire EOO of this species. Most of the occurrences of the species are within the prime growing elevations (i.e.,
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) for
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<emphasis id="ECC85270FFD05952FBB2FECA06BF80AF" italics="true" pageId="57" pageNumber="98">Coffea arabica</emphasis>
</taxonomicName>
L., and within two important coffee-growing regions (the Soconusco region of
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,
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and the departments of
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and
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in
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).
<collectingCountry id="A6ABCEF2FFD05952FBC1FE0B022F80CE" box="[1045,1126,445,469]" name="Mexico" pageId="57" pageNumber="98">Mexico</collectingCountry>
and
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are two of the top 10 coffee-producing countries in the world (
<bibRefCitation id="BA2DF393FFD05952FC52FE6B020D80EE" author="ANONYMOUS" box="[902,1092,477,501]" pageId="57" pageNumber="98" refId="ref46952" refString="ANONYMOUS. 2003. The coffee economy. Frontline World, Facts &amp; Stats. &lt;www. pbs. org / frontlineworld / stories / guatemala. mexico / facts. html&gt; [accessed 24 May 2018]" type="book" year="2003">Anonymous 2003</bibRefCitation>
, 2018). Increasing demand for coffee worldwide (e.g.,
<bibRefCitation id="BA2DF393FFD05952FDABFE4B0562830E" author="LORENZETTI, L." box="[639,811,509,533]" pageId="57" pageNumber="98" refId="ref48776" refString="LORENZETTI, L. 2016. Americans' coffee guzzling is pushing bean prices higher. Fortune. &lt;www. f ortune. com / 2016 / 07 / 01 / americans-coffee-prices /&gt; [accessed 24 May 2018]" type="journal volume" year="2016">Lorenzetti 2016</bibRefCitation>
) has or could result in increased conversion of natural vegetation in the EOO of this species to agriculture. Thus, coffee production would appear to be the more immediate threat to
<taxonomicName id="19BCF5E1FFD05952FD56FD8804BF834E" box="[642,758,574,597]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="57" pageNumber="98" phylum="Tracheophyta" rank="species" species="purpusii">
<emphasis id="ECC85270FFD05952FD56FD8804BF834E" box="[642,758,574,597]" italics="true" pageId="57" pageNumber="98">L. purpusii</emphasis>
</taxonomicName>
. However, based on either threat, there would appear to be a single location for the species. A preliminary conservation assessment of Endangered (EN; B1, a, b;
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) is proposed for this species.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFD05952FF7FFD28059A8707" pageId="57" pageNumber="98" type="discussion">
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<emphasis id="ECC85270FFD05952FF7FFD28076383AE" bold="true" box="[171,298,670,694]" pageId="57" pageNumber="98">DISCUSSION</emphasis>
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.— This species resembles some plants of
<taxonomicName id="19BCF5E1FFD05952FD3EFD29053783AD" box="[746,894,671,694]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="57" pageNumber="98" phylum="Tracheophyta" rank="species" species="mexicanum">
<emphasis id="ECC85270FFD05952FD3EFD29053783AD" box="[746,894,671,694]" italics="true" pageId="57" pageNumber="98">L. mexicanum</emphasis>
</taxonomicName>
(see discussion under that species), but differs by its more obovate leaves and pubescence. Additional morphological tendencies that distinguish these species include the more racemose (vs. thyrsoid) inflorescences of
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<emphasis id="ECC85270FFD05952FFAFFCB606A6820C" box="[123,239,768,791]" italics="true" pageId="57" pageNumber="98">L. purpusii</emphasis>
</taxonomicName>
(with dichasial peduncle lengths of 0 to 2 (vs. 1 to 22) mm; the caducous bracteoles (vs. usually persistent in
<taxonomicName id="19BCF5E1FFD05952FE8BFC9607B1822C" box="[351,504,800,823]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="57" pageNumber="98" phylum="Tracheophyta" rank="species" species="mexicanum">
<emphasis id="ECC85270FFD05952FE8BFC9607B1822C" box="[351,504,800,823]" italics="true" pageId="57" pageNumber="98">L. mexicanum</emphasis>
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); and the generally longer styles (85 to 92 vs.
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long), capsules (22 to 30 vs.
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long), and capsular stipes (3 to 3.5 vs.
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long). The distributional ranges of these two species are not known to overlap and are restricted to different versants of
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and
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.
</paragraph>
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.—
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<collectingCountry id="A6ABCEF2FFD05952FD9AFC1604FE82A3" box="[590,695,928,952]" name="Mexico" pageId="57" pageNumber="98">MEXICO</collectingCountry>
.
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<collectingRegion id="1C784080FFD05952FD12FC16056A82A3" box="[710,803,928,952]" country="Mexico" name="Chiapas" pageId="57" pageNumber="98">Chiapas</collectingRegion>
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:
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,
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of
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[ca.
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],
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D.
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&amp;
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<emphasis id="ECC85270FFD05952FCB6FC77052082C3" box="[866,873,961,984]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FCA6FC7605B582C3" box="[882,1020,960,984]" italics="true" pageId="57" pageNumber="98">
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31613
</emphasis>
(
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,
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,
<collectionCode id="B8AD16A7FFD05952FFAFFC5706EF82E2" box="[123,166,993,1017]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="57" pageNumber="98" type="Herbarium">MO</collectionCode>
);
<collectorName id="7349EBB4FFD05952FF68FC56070882E3" box="[188,321,992,1016]" pageId="57" pageNumber="98">Finca Nubes</collectorName>
[ca.
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,
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],
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<emphasis id="ECC85270FFD05952FD25FC5704B782E3" box="[753,766,993,1016]" italics="true" pageId="57" pageNumber="98">F</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FD2AFC57054C82E3" box="[766,773,993,1016]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FCDFFC5605EA82E2" box="[779,931,992,1017]" italics="true" pageId="57" pageNumber="98">
<collectingMunicipality id="3E671418FFD05952FCDFFC56052182E3" box="[779,872,992,1016]" pageId="57" pageNumber="98">Miranda</collectingMunicipality>
1723
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC66FC57059A82E3" box="[946,979,993,1016]" country="USA" lsid="urn:lsid:biocol.org:col:14627" name="University of Texas at Austin" pageId="57" pageNumber="98" type="Herbarium">LL</collectionCode>
,
<collectingCountry id="A6ABCEF2FFD05952FC0BFC57024882E2" box="[991,1025,993,1017]" name="United States of America" pageId="57" pageNumber="98">US</collectingCountry>
); from [Finca San Antonio]
<location id="DB63D8B9FFD05952FEDBFBB707CA8502" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FEDBFBB707CA8502" box="[271,387,1025,1049]" country="United States of America" municipality="Miranda" name="Chicharras" pageId="57" pageNumber="98">Chicharras</location>
,
<collectionCode id="B8AD16A7FFD05952FE44FBB707E98503" box="[400,416,1025,1048]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="57" pageNumber="98" type="Herbarium">
<emphasis id="ECC85270FFD05952FE44FBB707E98503" box="[400,416,1025,1048]" italics="true" pageId="57" pageNumber="98">E</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FE74FBB707EE8503" box="[416,423,1025,1048]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FE79FBB7047A8502" box="[429,563,1025,1049]" italics="true" pageId="57" pageNumber="98">
<collectingMunicipality id="3E671418FFD05952FE79FBB707BE8502" box="[429,503,1025,1049]" pageId="57" pageNumber="98">Nelson</collectingMunicipality>
3749
</emphasis>
(
<collectingCountry id="A6ABCEF2FFD05952FD97FBB7042C8502" box="[579,613,1025,1049]" name="United States of America" pageId="57" pageNumber="98">US</collectingCountry>
);
<location id="DB63D8B9FFD05952FDA8FBB7050B8502" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FDA8FBB7050B8502" box="[636,834,1025,1049]" country="United States of America" latitude="15.123922" longLatPrecision="1" longitude="-92.24591" municipality="Nelson" name="Finca San Antonio" pageId="57" pageNumber="98">Finca San Antonio</location>
[Chicharras; ca.
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,
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],
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<emphasis id="ECC85270FFD05952FE9FFB9407138522" box="[331,346,1058,1081]" italics="true" pageId="57" pageNumber="98">L</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FE8EFB9407288522" box="[346,353,1058,1081]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FEBEFB97043B8522" box="[362,626,1057,1081]" italics="true" pageId="57" pageNumber="98">
<location id="DB63D8B9FFD05952FEBEFB9707898522" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FEBEFB9707898522" box="[362,448,1057,1081]" country="United States of America" name="Quarles" pageId="57" pageNumber="98">Quarles</location>
<taxonomicName id="19BCF5E1FFD05952FE1EFB9404098522" authority="Ufford" authorityName="Ufford" box="[458,576,1057,1081]" class="Magnoliopsida" family="Acanthaceae" genus="Harpochilus" kingdom="Plantae" order="Lamiales" pageId="57" pageNumber="98" phylum="Tracheophyta" rank="species" species="van">van Ufford</taxonomicName>
137
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FD51FB9704D18523" box="[645,664,1057,1080]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="57" pageNumber="98" type="Herbarium">U</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6ED4843FFFD05952FD65FB97049D8542" collectionCode="C" country="Guatemala" location="Sta. Maria Ikibal" pageId="57" pageNumber="98" specimenCount="1" stateProvince="Quezaltenango">
<collectingCountry id="A6ABCEF2FFD05952FD65FB97051E8522" box="[689,855,1057,1081]" name="Guatemala" pageId="57" pageNumber="98">GUATEMALA</collectingCountry>
.
<emphasis id="ECC85270FFD05952FCBCFB97025B8522" bold="true" box="[872,1042,1057,1081]" pageId="57" pageNumber="98">
<collectingRegion id="1C784080FFD05952FCBCFB97025B8522" box="[872,1042,1057,1081]" country="Guatemala" name="Quezaltenango" pageId="57" pageNumber="98">Quezaltenango</collectingRegion>
</emphasis>
:
<location id="DB63D8B9FFD05952FBF7FB9706F08542" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FBF7FB9706F08542" country="Guatemala" name="Sta. Maria Ikibal" pageId="57" pageNumber="98" stateProvince="Quezaltenango">Sta. María Ikibál</location>
[not located, possibly Santa María de Jesús],
<collectionCode id="B8AD16A7FFD05952FD16FBF4049D8542" box="[706,724,1090,1113]" country="Denmark" name="University of Copenhagen" pageId="57" pageNumber="98" type="Herbarium">
<emphasis id="ECC85270FFD05952FD16FBF4049D8542" box="[706,724,1090,1113]" italics="true" pageId="57" pageNumber="98">C</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFD05952FD00FBF405C78542" box="[724,910,1089,1113]" italics="true" pageId="57" pageNumber="98">
.
<materialsCitation id="6ED4843FFFD05952FD31FBF705CE8542" box="[741,903,1089,1113]" collectionCode="O" country="Guatemala" location="Bernoulli" pageId="57" pageNumber="98" specimenCount="1" stateProvince="Quezaltenango">
<location id="DB63D8B9FFD05952FD31FBF705038542" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FD31FBF705038542" box="[741,842,1089,1113]" country="Guatemala" name="Bernoulli" pageId="57" pageNumber="98" stateProvince="Quezaltenango">Bernoulli</location>
&amp;
<collectionCode id="B8AD16A7FFD05952FCA0FBF405CE8542" box="[884,903,1090,1113]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="57" pageNumber="98" type="Herbarium">O</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="6ED4843FFFD05952FC4DFBF406C38562" collectionCode="K, S" country="Guatemala" location="Cario" pageId="57" pageNumber="98" specimenCount="1" stateProvince="Quezaltenango">
<emphasis id="ECC85270FFD05952FC4DFBF4025F8542" box="[921,1046,1089,1113]" italics="true" pageId="57" pageNumber="98">
<location id="DB63D8B9FFD05952FC4DFBF4059E8542" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FC4DFBF4059E8542" box="[921,983,1090,1113]" country="Guatemala" name="Cario" pageId="57" pageNumber="98" stateProvince="Quezaltenango">Cario</location>
2254
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FBFEFBF402748542" box="[1066,1085,1090,1113]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="57" pageNumber="98" type="Herbarium">K</collectionCode>
); lower
<collectionCode id="B8AD16A7FFD05952FFAFFBD706C38562" box="[123,138,1121,1145]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="57" pageNumber="98" type="Herbarium">S</collectionCode>
</materialsCitation>
<materialsCitation id="6ED4843FFFD05952FF46FBD704728562" box="[146,571,1121,1145]" collectionCode="W" country="Guatemala" location="Volcan Santa Maria" pageId="57" pageNumber="98" specimenCount="1" stateProvince="Quezaltenango">
slope of
<location id="DB63D8B9FFD05952FF24FBD7078F8562" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FF24FBD7078F8562" box="[240,454,1121,1145]" country="Guatemala" name="Volcan Santa Maria" pageId="57" pageNumber="98" stateProvince="Quezaltenango">Volcán Santa María</location>
,
<quantity id="19442387FFD05952FE01FBD704538562" box="[469,538,1121,1145]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="57" pageNumber="98" unit="mi" value="0.3">0.3 mi</quantity>
<collectionCode id="B8AD16A7FFD05952FDF6FBD404728562" box="[546,571,1122,1145]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="57" pageNumber="98" type="Herbarium">W</collectionCode>
</materialsCitation>
<materialsCitation id="6ED4843FFFD05952FD97FBD7022F8562" box="[579,1126,1121,1145]" collectionCode="KM" country="Guatemala" location="Finca Pirineos" pageId="57" pageNumber="98" specimenCode="KM 97" specimenCount="1" stateProvince="Quezaltenango">
of jct.
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road with hwy. 95 at
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(
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); de la represa
<collectorName id="7349EBB4FFD05952FBA8FB3406B485A1" pageId="57" pageNumber="98">de Santa María</collectorName>
al
<collectorName id="7349EBB4FFD05952FECAFB1407E385A1" box="[286,426,1186,1210]" pageId="57" pageNumber="98">Volcán Pecul</collectorName>
[ca.
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<emphasis id="ECC85270FFD05952FCB4FB15020385A1" box="[864,1098,1186,1210]" italics="true" pageId="57" pageNumber="98">
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(
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);
<collectorName id="7349EBB4FFD05952FFAFFB74072A85C1" box="[123,355,1218,1242]" pageId="57" pageNumber="98">San José Buena Vista</collectorName>
,
<collectorName id="7349EBB4FFD05952FEA5FB7407A585C1" box="[369,492,1218,1242]" pageId="57" pageNumber="98">Costa Cuca</collectorName>
, [ca.
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,
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],
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<emphasis id="ECC85270FFD05952FC7FFB7505F385C1" box="[939,954,1219,1242]" italics="true" pageId="57" pageNumber="98">L</emphasis>
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<emphasis id="ECC85270FFD05952FC6EFB75058885C1" box="[954,961,1219,1242]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FC1CFB74022F85C1" box="[968,1126,1218,1242]" italics="true" pageId="57" pageNumber="98">
<location id="DB63D8B9FFD05952FC1CFB74027F85C1" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FC1CFB74027F85C1" box="[968,1078,1218,1242]" country="United States of America" latitude="14.653394" longLatPrecision="1" longitude="-91.590385" name="Rodriguez" pageId="57" pageNumber="98">Rodríguez</location>
444
</emphasis>
(
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);
<collectorName id="7349EBB4FFD05952FFAFFB54068C85E1" box="[123,197,1250,1274]" pageId="57" pageNumber="98">Palmar</collectorName>
, [ca.
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,
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],
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<emphasis id="ECC85270FFD05952FD59FB5504D485E1" box="[653,669,1251,1274]" italics="true" pageId="57" pageNumber="98">A</emphasis>
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<emphasis id="ECC85270FFD05952FD49FB5504ED85E1" box="[669,676,1251,1274]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FD7AFB54057C85E1" box="[686,821,1250,1274]" italics="true" pageId="57" pageNumber="98">
<collectingMunicipality id="3E671418FFD05952FD7AFB5404BC85E1" box="[686,757,1250,1274]" pageId="57" pageNumber="98">Skutch</collectingMunicipality>
1451
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(
<collectionCode id="B8AD16A7FFD05952FC9CFB55051C85E1" box="[840,853,1251,1274]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
,
<collectingCountry id="A6ABCEF2FFD05952FCB1FB5405CE85E1" box="[869,903,1250,1274]" name="United States of America" pageId="57" pageNumber="98">US</collectingCountry>
);
<collectorName id="7349EBB4FFD05952FC76FB54020A85E1" box="[930,1091,1250,1274]" pageId="57" pageNumber="98">Finca Pireneos</collectorName>
, below
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, [ca.
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,
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<emphasis id="ECC85270FFD05952FD02FAB504AA8401" box="[726,739,1283,1306]" italics="true" pageId="57" pageNumber="98">P</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FD37FAB504A38401" box="[739,746,1283,1306]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FD25FAB405DE8401" box="[753,919,1282,1306]" italics="true" pageId="57" pageNumber="98">
<location id="DB63D8B9FFD05952FD25FAB405048401" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FD25FAB405048401" box="[753,845,1282,1306]" country="United States of America" name="Standley" pageId="57" pageNumber="98">Standley</location>
68259
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC72FAB505FA8401" box="[934,947,1283,1306]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
,
<collectingCountry id="A6ABCEF2FFD05952FC15FAB505AA8400" box="[961,995,1283,1307]" name="United States of America" pageId="57" pageNumber="98">US</collectingCountry>
); along old road
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,
<collectionCode id="B8AD16A7FFD05952FDDBFA9504558421" box="[527,540,1315,1338]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="57" pageNumber="98" type="Herbarium">
<emphasis id="ECC85270FFD05952FDDBFA9504558421" box="[527,540,1315,1338]" italics="true" pageId="57" pageNumber="98">P</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FDC8FA95046A8421" box="[540,547,1315,1338]" italics="true" pageId="57" pageNumber="98">.</emphasis>
<materialsCitation id="6ED4843FFFD05952FDFEFA9504828460" collectionCode="F, J" collectorName="Quebrada San Geronimo &amp; Volcan Santa Maria" country="United States of America" location="Finca Pirineos" municipality="Standley" pageId="57" pageNumber="98" specimenCount="1">
<emphasis id="ECC85270FFD05952FDFEFA9504998420" box="[554,720,1315,1339]" italics="true" pageId="57" pageNumber="98">
<collectingMunicipality id="3E671418FFD05952FDFEFA9504CF8420" box="[554,646,1315,1339]" pageId="57" pageNumber="98">Standley</collectingMunicipality>
86714
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FD0BFA9504A78421" box="[735,750,1315,1338]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
);
<emphasis id="ECC85270FFD05952FCD2FA9505008420" box="[774,841,1315,1339]" italics="true" pageId="57" pageNumber="98">86726</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC8CFA95052E8421" box="[856,871,1315,1338]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
),
<emphasis id="ECC85270FFD05952FCAAFA9505888420" box="[894,961,1315,1339]" italics="true" pageId="57" pageNumber="98">86749</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC05FA9505A98421" box="[977,992,1315,1338]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
),
<emphasis id="ECC85270FFD05952FC22FA9502708420" box="[1014,1081,1315,1339]" italics="true" pageId="57" pageNumber="98">86750</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FB9DFA95021F8421" box="[1097,1110,1315,1338]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
,
<collectingCountry id="A6ABCEF2FFD05952FBB0FA9502CF8420" box="[1124,1158,1315,1339]" name="United States of America" pageId="57" pageNumber="98">US</collectingCountry>
),
<emphasis id="ECC85270FFD05952FFAFFAF506F78440" box="[123,190,1347,1371]" italics="true" pageId="57" pageNumber="98">86763</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FF1AFAF506948441" box="[206,221,1347,1370]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
); along
<collectorName id="7349EBB4FFD05952FEECFAF5040A8440" box="[312,579,1347,1371]" pageId="57" pageNumber="98">Quebrada San Gerónimo</collectorName>
,
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, lower S-facing slopes of
<collectorName id="7349EBB4FFD05952FBDDFAF506F28460" pageId="57" pageNumber="98">Volcán Santa María</collectorName>
,
<locationDeviation id="CFB19740FFD05952FF13FAD504FB8460" box="[199,690,1379,1403]" firstLocation="Santa Maria de Jesus" pageId="57" pageNumber="98" secondLocation="Calahuache">between Santa María de Jesús and Calahuaché</locationDeviation>
,
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<emphasis id="ECC85270FFD05952FD6BFAD204828460" box="[703,715,1380,1403]" italics="true" pageId="57" pageNumber="98">J</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FD1FFAD2049B8460" box="[715,722,1380,1403]" italics="true" pageId="57" pageNumber="98">.</emphasis>
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<emphasis id="ECC85270FFD05952FD0CFAD505D38460" box="[728,922,1379,1403]" italics="true" pageId="57" pageNumber="98">
<location id="DB63D8B9FFD05952FD0CFAD505188460" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FD0CFAD505188460" box="[728,849,1379,1403]" country="United States of America" name="Steyermark" pageId="57" pageNumber="98">Steyermark</location>
33435
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC7CFAD205FE8460" box="[936,951,1380,1403]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
); off
<collectorName id="7349EBB4FFD05952FC26FAD202628460" box="[1010,1067,1380,1404]" pageId="57" pageNumber="98">Hwy.</collectorName>
95 at
<specimenCode id="8E1A2619FFD05952FBBEFAD206EA8487" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="57" pageNumber="98">KM 197</specimenCode>
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,
<materialsCitation id="6ED4843FFFD05952FF67FA3505D384A0" collectionCode="DUKE, L" collectorName="D. Stone &amp; Colomba &amp; Finca San Francisco Pie de la Cuesta" country="United States of America" latitude="14.728069" location="Finca Pirineos" longLatPrecision="1" longitude="-91.7179" pageId="57" pageNumber="98" specimenCount="1">
old road to
<location id="DB63D8B9FFD05952FEE1FA3507988480" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FEE1FA3507988480" box="[309,465,1411,1435]" country="United States of America" latitude="14.728069" longLatPrecision="1" longitude="-91.7179" name="Finca Pirineos" pageId="57" pageNumber="98">Finca Pirineos</location>
, ca.
<quantity id="19442387FFD05952FDDEFA3204198480" box="[522,592,1411,1436]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="57" pageNumber="98" unit="mi" value="0.5">0.5 mi</quantity>
from main hwy.,
<emphasis id="ECC85270FFD05952FCCDFA3205FF8480" box="[793,950,1411,1435]" italics="true" pageId="57" pageNumber="98">
<collectorName id="7349EBB4FFD05952FCCDFA3205318480" box="[793,888,1412,1435]" pageId="57" pageNumber="98">D. Stone</collectorName>
3086
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC1DFA32025A8480" box="[969,1043,1412,1435]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15531" lsid="urn:lsid:biocol.org:col:15531" name="Duke University" pageId="57" pageNumber="98" type="Herbarium">DUKE</collectionCode>
);
<collectorName id="7349EBB4FFD05952FBF9FA3502C68480" box="[1069,1167,1411,1435]" pageId="57" pageNumber="98">Colomba</collectorName>
,
<collectorName id="7349EBB4FFD05952FFAFFA12044084A7" box="[123,521,1444,1468]" pageId="57" pageNumber="98">Finca San Francisco Pie de la Cuesta</collectorName>
,
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,
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,
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<emphasis id="ECC85270FFD05952FC5FFA1205D384A0" box="[907,922,1444,1467]" italics="true" pageId="57" pageNumber="98">L</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFD05952FC4EFA12020684A0" box="[922,1103,1444,1468]" italics="true" pageId="57" pageNumber="98">
.
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&amp; W.
</emphasis>
<materialsCitation id="6ED4843FFFD05952FB82FA12043084E7" collectionCode="CAS, J" collectorName="Finca El Porvenir" country="United States of America" county="Lopez" latitude="14.960161" location="Volcan Tajumulco" longLatPrecision="1" longitude="-91.94465" municipality="Rio Chopal" pageId="57" pageNumber="98" specimenCount="1" stateProvince="San Marcos">
<emphasis id="ECC85270FFD05952FB82FA1206F984C7" italics="true" pageId="57" pageNumber="98">
<collectingCounty id="3762F6EEFFD05952FB82FA1202DF84A0" box="[1110,1174,1444,1467]" pageId="57" pageNumber="98">López</collectingCounty>
1792
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FF16FA7206BF84C7" box="[194,246,1476,1500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="57" pageNumber="98" type="Museum">CAS</collectionCode>
).
<emphasis id="ECC85270FFD05952FEDAFA7207DF84C7" bold="true" box="[270,406,1476,1500]" pageId="57" pageNumber="98">
<collectingRegion id="1C784080FFD05952FEDAFA7207DF84C7" box="[270,406,1476,1500]" country="United States of America" name="San Marcos" pageId="57" pageNumber="98">San Marcos</collectingRegion>
</emphasis>
:
<collectorName id="7349EBB4FFD05952FE73FA72042E84C7" box="[423,615,1476,1500]" pageId="57" pageNumber="98">Finca El Porvenir</collectorName>
along
<collectingMunicipality id="3E671418FFD05952FD67FA72057984C7" box="[691,816,1476,1500]" pageId="57" pageNumber="98">Río Chopal</collectingMunicipality>
, S-facing slopes of
<location id="DB63D8B9FFD05952FBD8FA7206F784E7" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FBD8FA7206F784E7" country="United States of America" county="Lopez" latitude="14.960161" longLatPrecision="1" longitude="-91.94465" municipality="Rio Chopal" name="Volcan Tajumulco" pageId="57" pageNumber="98" stateProvince="San Marcos">Volcán Tajumulco</location>
[ca.
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,
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],
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</materialsCitation>
.
<materialsCitation id="6ED4843FFFD05952FD52FA52020084E7" box="[646,1097,1508,1532]" collectionCode="F" collectorName="El Tumbador" country="United States of America" location="Steyermark" pageId="57" pageNumber="98" specimenCount="1" stateProvince="San Marcos">
<emphasis id="ECC85270FFD05952FD52FA52050084E7" box="[646,841,1508,1532]" italics="true" pageId="57" pageNumber="98">
<location id="DB63D8B9FFD05952FD52FA5204B684E7" LSID="urn:lsid:plazi:treatment:56153F74FFD15952FF7FFEEB059A8707:DB63D8B9FFD05952FD52FA5204B684E7" box="[646,767,1508,1532]" country="United States of America" name="Steyermark" pageId="57" pageNumber="98" stateProvince="San Marcos">Steyermark</location>
37501
</emphasis>
(
<collectionCode id="B8AD16A7FFD05952FC83FA53052F84E7" box="[855,870,1509,1532]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="57" pageNumber="98" type="Herbarium">F</collectionCode>
);
<collectorName id="7349EBB4FFD05952FCA9FA52024E84E7" box="[893,1031,1508,1532]" pageId="57" pageNumber="98">El Tumbador</collectorName>
, finca
</materialsCitation>
<materialsCitation id="6ED4843FFFD05952FB84FA5304058707" collectionCode="L" country="Colombia" latitude="14.855528" location="Nueva Granada" longLatPrecision="1" longitude="-91.88497" pageId="57" pageNumber="98" specimenCount="1">
<collectingCountry id="A6ABCEF2FFD05952FB84FA53069C8707" name="Colombia" pageId="57" pageNumber="98">Nueva Granada</collectingCountry>
,
<geoCoordinate id="BB88E8A5FFD05952FF37F9B207308707" box="[227,377,1540,1564]" degrees="14" direction="north" minutes="51" orientation="latitude" pageId="57" pageNumber="98" precision="1" seconds="19. 9" value="14.855528">14°5119.9”N</geoCoordinate>
,
<geoCoordinate id="BB88E8A5FFD05952FE5CF9B204668707" box="[392,559,1540,1564]" degrees="091" direction="west" minutes="53" orientation="longitude" pageId="57" pageNumber="98" precision="1" seconds="05. 9" value="-91.88497">091°5305.9”W</geoCoordinate>
,
<collectionCode id="B8AD16A7FFD05952FDE9F9B304058707" box="[573,588,1541,1564]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="57" pageNumber="98" type="Herbarium">
<emphasis id="ECC85270FFD05952FDE9F9B304058707" box="[573,588,1541,1564]" italics="true" pageId="57" pageNumber="98">L</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFD05952FD98F9B305798707" box="[588,816,1540,1564]" italics="true" pageId="57" pageNumber="98">
.
<taxonomicName id="19BCF5E1FFD05952FD8EF9B2048A8707" box="[602,707,1540,1564]" class="Insecta" family="Crambidae" genus="Velasquez" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="98" phylum="Arthropoda" rank="genus">Velásquez</taxonomicName>
et al. 658
</emphasis>
(BIGU, CAS).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,542 @@
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<mods:title id="8EE4C91A415667F381211C97DB4F19D5">Louteridium (Acanthaceae: Acanthoideae: Ruellieae: Trichantherinae): Taxonomy, Phylogeny, Reproductive Biology, and Conservation</mods:title>
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<treatment id="56153F74FFF65949FF7FFAB604C982F9" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9" httpUri="http://treatment.plazi.org/id/56153F74FFF65949FF7FFAB604C982F9" lastPageId="34" lastPageNumber="75" pageId="31" pageNumber="72">
<subSubSection id="96A6DDE9FFF65974FF7FFAB604138462" pageId="31" pageNumber="72" type="nomenclature">
<paragraph id="DE038E62FFF65974FF7FFAB604138462" blockId="31.[123,1174,1280,1401]" pageId="31" pageNumber="72">
<emphasis id="ECC85270FFF65974FF7FFAB606F68402" bold="true" box="[171,191,1280,1305]" pageId="31" pageNumber="72">2.</emphasis>
<treatmentCitation id="5F1DA873FFF65974FF1DFAB602618402" author="Leonard" box="[201,1064,1280,1305]" journal="Publ. Carnegie Inst. Wash." page="197" pageId="31" pageNumber="72" score="430.3814197489919" volume="461" year="1936">
<taxonomicName id="19BCF5E1FFF65974FF1DFAB605C98403" ID-CoL="72NQR" authority="Leonard, Publ. Carnegie Inst. Wash." box="[201,896,1280,1305]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="31" pageNumber="72" phylum="Tracheophyta" rank="species" species="chartaceum">
<emphasis id="ECC85270FFF65974FF1DFAB607A98403" bold="true" box="[201,480,1280,1304]" italics="true" pageId="31" pageNumber="72">Louteridium chartaceum</emphasis>
<materialsCitation id="6ED4843FFFF65974FE3FFAB705C98403" box="[491,896,1281,1305]" collectingDate="1935-03-24" collectionCode="TYPE, P, ARIZ, K, LL, MICH, MO, NY" country="Belize" latitude="17.38713" location="Sibun River" longLatPrecision="1" longitude="-88.45053" municipality="Gracie Rock" pageId="31" pageNumber="72" specimenCount="2" stateProvince="Belize" typeStatus="holotype">
Leonard,
<emphasis id="ECC85270FFF65974FD81FAB705C98403" box="[597,896,1281,1305]" italics="true" pageId="31" pageNumber="72">Publ. Carnegie Inst. Wash.</emphasis>
</materialsCitation>
</taxonomicName>
461:197. 1936
</treatmentCitation>
.
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<emphasis id="ECC85270FFF65974FBEEFAB7023D840C" bold="true" box="[1082,1140,1281,1305]" pageId="31" pageNumber="72">TYPE</emphasis>
</smallCapsWord>
.—
<collectingCountry id="A6ABCEF2FFF65974FFAFFA97069E8423" box="[123,215,1313,1336]" name="Belize" pageId="31" pageNumber="72">BELIZE</collectingCountry>
.
<emphasis id="ECC85270FFF65974FF37FA97076E8422" bold="true" box="[227,295,1313,1337]" pageId="31" pageNumber="72">
<collectingRegion id="1C784080FFF65974FF37FA97076E8422" box="[227,295,1313,1337]" country="Belize" name="Belize" pageId="31" pageNumber="72">Belize</collectingRegion>
</emphasis>
:
<collectingMunicipality id="3E671418FFF65974FEE1FA9707F38422" box="[309,442,1313,1337]" pageId="31" pageNumber="72">Gracie Rock</collectingMunicipality>
,
<location id="DB63D8B9FFF65974FE13FA97040E8422" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFF65974FE13FA97040E8422" box="[455,583,1313,1337]" country="Belize" latitude="17.38713" longLatPrecision="1" longitude="-88.45053" municipality="Gracie Rock" name="Sibun River" pageId="31" pageNumber="72" stateProvince="Belize">Sibun River</location>
[ca.
<geoCoordinate id="BB88E8A5FFF65974FDAFFA9705578423" box="[635,798,1313,1337]" degrees="17" direction="north" minutes="23" orientation="latitude" pageId="31" pageNumber="72" precision="1" seconds="13. 67" value="17.38713">17°2313.67”N</geoCoordinate>
,
<geoCoordinate id="BB88E8A5FFF65974FCFFFA97059C8423" box="[811,981,1313,1337]" degrees="088" direction="west" minutes="27" orientation="longitude" pageId="31" pageNumber="72" precision="1" seconds="1. 91" value="-88.45053">088°271.91”W</geoCoordinate>
],
<date id="AA02A8A2FFF65974FC3FFA97022E8422" box="[1003,1127,1313,1337]" pageId="31" pageNumber="72" value="1935-03-24">
<collectingDate id="BA46514AFFF65974FC3FFA97022E8422" box="[1003,1127,1313,1337]" pageId="31" pageNumber="72" value="1935-03-24">24-III-1935</collectingDate>
</date>
(flr, frt),
<emphasis id="ECC85270FFF65974FF79FAF407188442" box="[173,337,1345,1369]" italics="true" pageId="31" pageNumber="72">P. Gentle 1526</emphasis>
(
<typeStatus id="010730C0FFF65974FEB0FAF707888442" box="[356,449,1345,1369]" pageId="31" pageNumber="72" type="holotype">holotype</typeStatus>
: US-1589669!;
<typeStatus id="010730C0FFF65974FDA1FAF704858442" box="[629,716,1345,1369]" pageId="31" pageNumber="72" type="isotype">isotypes</typeStatus>
: A-image seen, ARIZ!, K!, LL!, MICH!, MO!, NY!, US-1589668!, WIS-image seen).
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFF6594AFF7FFA3A054585C8" lastPageId="33" lastPageNumber="74" pageId="31" pageNumber="72" type="description">
<paragraph id="DE038E62FFF6594AFF7FFA3A056C853F" blockId="31.[123,1174,1420,1573]" lastBlockId="33.[123,1174,219,1060]" lastPageId="33" lastPageNumber="74" pageId="31" pageNumber="72">
Shrubs to trees to
<quantity id="19442387FFF65974FEA5FA3A07E684BF" box="[369,431,1420,1444]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="31" pageNumber="72" unit="m" value="3.5">3.5 m</quantity>
tall, often epipetric. Older (woody) stems irregularly striate, ± lenticellate, lacking trichomes; younger (herbaceous) stems subterete to subquadrate, glabrous. Leaves seasonally deciduous, ± clustered just proximal to inflorescence, petiolate (or distal leaves subsessile), petioles to
<quantity id="19442387FFF65974FEE5FA5B0737871F" box="[305,382,1517,1541]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="72" unit="mm" value="70.0">70 mm</quantity>
long, blades subsucculent, ovate to elliptic (to obovate-elliptic),
<quantity id="19442387FFF65974FB91FA5B06ED873F" metricMagnitude="-1" metricUnit="m" metricValue="1.775" metricValueMax="2.85" metricValueMin="0.7" pageId="31" pageNumber="72" unit="mm" value="177.5" valueMax="285.0" valueMin="70.0">70285 mm</quantity>
long,
<quantity id="19442387FFF65974FF3EF9BB0717873F" box="[234,350,1549,1573]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="9.5" metricValueMin="3.0" pageId="31" pageNumber="72" unit="mm" value="62.5" valueMax="95.0" valueMin="30.0">3095 mm</quantity>
wide, 1.73.7 × longer than wide, acuminate to subfalcate at apex, rounded to acute (distal leaves) to attenuate (proximal leaves) at base, surfaces glabrous, margin entire and sometimes with inconspicuous marginal swellings/encrustations. Inflorescence a terminal (raceme to) racemose thyrse to thyrse to
<quantity id="19442387FFC8594AFE06FEAA0463802F" box="[466,554,284,308]" metricMagnitude="-1" metricUnit="m" metricValue="5.8" pageId="33" pageNumber="74" unit="mm" value="580.0">580 mm</quantity>
long (including peduncle and excluding corollas), peduncle to
<quantity id="19442387FFC8594AFF6AFE8B075F804E" box="[190,278,317,341]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="33" pageNumber="74" unit="mm" value="240.0">240 mm</quantity>
long, glabrous, rachis glabrous; dichasia expanded or sometimes modified by very short expansion between pairs of succeeding flowers with the congested dichasial axis becoming a ± linear racemelike lateral short-shoot to
<quantity id="19442387FFC8594AFDE6FEC90434808D" box="[562,637,383,407]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="74" unit="mm" value="10.0">10 mm</quantity>
long, opposite, sessile to pedunculate, (1) many-flowered, to
<quantity id="19442387FFC8594AFEDDFE29071E80AC" box="[265,343,415,439]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="33" pageNumber="74" unit="mm" value="70.0">70 mm</quantity>
long (excluding corollas), dichasial peduncles (if present) to
<quantity id="19442387FFC8594AFBD3FE29021C80AC" box="[1031,1109,415,439]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="33" pageNumber="74" unit="mm" value="24.0">24 mm</quantity>
long, glabrous. Bracts caducous, subfoliose, ovate to lanceolate,
<quantity id="19442387FFC8594AFCDAFE7605CC80CC" box="[782,901,448,472]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.2" metricValueMin="1.8" pageId="33" pageNumber="74" unit="mm" value="35.0" valueMax="52.0" valueMin="18.0">1852 mm</quantity>
long, 3.
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wide, reduced in size toward apex, abaxially glabrous. Bracteoles caducous, triangular,
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long, 1.5
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wide, abaxially glabrous. Flowers pedicellate, pedicels
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long, glabrous. Calyx 1826 (
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fruit) mm long, lobes fused at base for 1
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, subheteromorphic, subsucculent, linear to lance-linear,
<quantity id="19442387FFC8594AFE35FDF4041F8341" box="[481,598,578,602]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="33" pageNumber="74" unit="mm" value="20.0" valueMax="25.0" valueMin="15.0">1525 mm</quantity>
long,
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wide, (subrounded) to acute at apex, abaxially glandular-punctate (punctations to
<quantity id="19442387FFC8594AFDB6FDD504FF8360" box="[610,694,611,635]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="74" unit="mm" value="0.1">0.1 mm</quantity>
in diameter, sometimes sparse) but lacking elongate trichomes, posterior lobe subplanar to subconduplicate,
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long, equal to or short- er than lateral lobes,
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wide, lateral lobes
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long, 3.
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wide, all lobes (subrounded to) acute at apex. Corolla light green or greenish yellow with lobes usually becoming slightly maroon-tinged and darker maroon at distal tips with age,
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long, externally glabrous, tube
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long, narrow proximal portion
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long,
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in diameter near midpoint, throat
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long,
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in diameter at mouth, lobes recurved to recoiled, oblong to ovate-elliptic, 1214.
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long, 8.510.
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wide, rounded and bifid at apex. Stamens 4,
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long, distally glabrous, pubescent near base with eglandular trichomes, thecae yellowish green, 6.59.
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long; staminode 1, ± triangular, 0.1
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long. Style
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long, glabrous, stigma unequally 2-lobed and ± obliquely funnelform, 1.5
<quantity id="19442387FFC8594AFC6CFC1C05B082D9" box="[952,1017,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="74" unit="mm" value="2.0">2 mm</quantity>
long, lobes ± elliptic, 1
<quantity id="19442387FFC8594AFF24FC7D070D82F9" box="[240,324,971,995]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="74" unit="mm" value="1.5">1.5 mm</quantity>
long, 0.6
<quantity id="19442387FFC8594AFE69FC7D07B482F9" box="[445,509,971,995]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="74" unit="mm" value="1.0">1 mm</quantity>
wide. Capsule (20)
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long,
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in diameter, glabrous, stipe 1.5
<quantity id="19442387FFC8594AFE9FFC5A07CE8518" box="[331,391,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="74" unit="mm" value="2.0">2 mm</quantity>
long. Seeds up to 16 per capsule, 4
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long,
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wide, surfaces with subconic to low rounded protrusions or becoming smooth.
</paragraph>
<caption id="8AC3DEEAFFF65974FF4FF9F5045987A7" pageId="31" pageNumber="72" startId="31.[155,167,1603,1622]" targetType="figure">
<paragraph id="DE038E62FFF65974FF4FF9F5045987A7" blockId="31.[123,1174,1602,1724]" pageId="31" pageNumber="72">
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12 (right).
<taxonomicName id="19BCF5E1FFF65974FE95F9F4045E874E" box="[321,535,1602,1621]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="31" pageNumber="72" phylum="Tracheophyta" rank="species" species="chartaceum">
<emphasis id="ECC85270FFF65974FE95F9F4045E874E" box="[321,535,1602,1621]" italics="true" pageId="31" pageNumber="72">Louteridium chartaceum</emphasis>
</taxonomicName>
(
<emphasis id="ECC85270FFF65974FDF2F9F504DE874E" box="[550,663,1602,1621]" italics="true" pageId="31" pageNumber="72">Brewer 7176</emphasis>
). A. Flower. B. Habit of young plant showing clusters of leaves. C. Trunk of mature plant growing on rocky slope. D. Portion of mature leaf showing red to pink veins and marginal swellings/encrustations. E. Older stem showing swelling above nodes, raised lenticels, and leaf scars. F. Inflorescence with lateral short shoots, persistent calyces on old flowers, and a nearly mature floral bud. Photos by S. Brewer (to whom copyrights are reserved), used with permission.
</paragraph>
</caption>
<paragraph id="DE038E62FFC8594AFF7FFB8E054585C8" blockId="33.[123,1174,1080,1726]" pageId="33" pageNumber="74">
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<emphasis id="ECC85270FFC8594AFF7FFB8E07788554" bold="true" box="[171,305,1080,1104]" pageId="33" pageNumber="74">PHENOLOGY</emphasis>
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.— Flowering: JanuaryMarch; fruiting: FebruaryMarch, June. In the two populations of this species observed for ca. one hour each, floral buds were present on the plants at both sites during daylight hours (morning for
<emphasis id="ECC85270FFC8594AFDB3FBCC04A58589" box="[615,748,1146,1170]" italics="true" pageId="33" pageNumber="74">Daniel 8294</emphasis>
and afternoon for
<emphasis id="ECC85270FFC8594AFC62FBCC06F985A9" italics="true" pageId="33" pageNumber="74">Daniel &amp; Butterwick 5905</emphasis>
), and mature fallen flowers were present on the ground at one site (
<emphasis id="ECC85270FFC8594AFC44FB2C025C85A9" box="[912,1045,1178,1202]" italics="true" pageId="33" pageNumber="74">Daniel 8294</emphasis>
). Likewise,
<emphasis id="ECC85270FFC8594AFFAFFB0A070B85C8" box="[123,322,1211,1235]" italics="true" pageId="33" pageNumber="74">Brewer et al. 7176</emphasis>
notes that the corollas fall in the morning.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFC85949FF7FFB6A020380E3" lastPageId="34" lastPageNumber="75" pageId="33" pageNumber="74" type="distribution">
<paragraph id="DE038E62FFC8594AFF7FFB6A053D842E" blockId="33.[123,1174,1080,1726]" pageId="33" pageNumber="74">
<emphasis id="ECC85270FFC8594AFF7FFB6A079385E9" bold="true" box="[171,474,1244,1268]" pageId="33" pageNumber="74">
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<smallCapsWord id="D8E518BEFFC8594AFE55FB69079385E9" baselines="1261" box="[385,474,1247,1266]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="habitat" pageId="33" pageNumber="74">HABITAT</smallCapsWord>
</emphasis>
.—
<materialsCitation id="6ED4843FFFC8594AFDD1FB6A053D842E" country="Belize" county="East-central" elevation="65" location="Sabido" municipality="Meerman" pageId="33" pageNumber="74" specimenCount="1" stateProvince="Cayo">
<collectingCounty id="3762F6EEFFC8594AFDD1FB6A04CD85EF" box="[517,644,1244,1268]" pageId="33" pageNumber="74">East-central</collectingCounty>
<collectingCountry id="A6ABCEF2FFC8594AFD5BFB6A049A85EF" box="[655,723,1244,1268]" name="Belize" pageId="33" pageNumber="74">Belize</collectingCountry>
(
<collectingCountry id="A6ABCEF2FFC8594AFD31FB6A056085EF" box="[741,809,1244,1268]" name="Belize" pageId="33" pageNumber="74">Belize</collectingCountry>
,
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;
<figureCitation id="468792E7FFC8594AFC57FB6A059B85EF" box="[899,978,1244,1268]" captionStart="FIGURE" captionStartId="20.[161,173,1071,1090]" captionTargetBox="[124,1173,223,1050]" captionTargetId="figure-1373@20.[120,1176,216,1055]" captionTargetPageId="20" captionText="FIGURE 10. Map of southern Mexico (showing states) and northern Central America with distributions of Louteridium chartaceum, L. donnell-smithii, and L. purpusii. Symbols may pertain to multiple collections." figureDoi="http://doi.org/10.5281/zenodo.13155729" httpUri="https://zenodo.org/record/13155729/files/figure.png" pageId="33" pageNumber="74">Fig. 10</figureCitation>
); plants occur on steep limestone slopes in evergreen seasonal forests (tropical evergreen seasonal broadleaf lowland forest fide
<collectingMunicipality id="3E671418FFC8594AFF24FAA8071F842E" box="[240,342,1310,1333]" pageId="33" pageNumber="74">Meerman</collectingMunicipality>
and
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2001) at elevations from
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<elevation id="55916951FFC8594AFD3FFAAB0524842E" box="[747,877,1309,1333]" metricMagnitude="1" metricUnit="m" metricValue="6.5" metricValueMax="12.0" metricValueMin="1.0" pageId="33" pageNumber="74" unit="m" value="65.0" valueMax="120.0" valueMin="10.0">10 to 120 m</elevation>
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.
</materialsCitation>
</paragraph>
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<emphasis id="ECC85270FFC8594AFF7FFA880719844F" bold="true" box="[171,336,1342,1366]" pageId="33" pageNumber="74">ILLUSTRATIONS</emphasis>
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.—
<figureCitation id="468792E7FFC8594AFEADFA8807A8844D" box="[377,481,1342,1366]" captionStart="FIGURE" captionStartId="31.[155,167,1603,1622]" captionText="FIGURE 12 (right). Louteridium chartaceum (Brewer 7176). A. Flower. B. Habit of young plant showing clusters of leaves. C. Trunk of mature plant growing on rocky slope. D. Portion of mature leaf showing red to pink veins and marginal swellings/encrustations. E. Older stem showing swelling above nodes, raised lenticels, and leaf scars. F. Inflorescence with lateral short shoots, persistent calyces on old flowers, and a nearly mature floral bud. Photos by S. Brewer (to whom copyrights are reserved), used with permission." pageId="33" pageNumber="74">Figure 12</figureCitation>
;
<bibRefCitation id="BA2DF393FFC8594AFE3BFA8804F4844D" author="LEONARD, E. C." box="[495,701,1342,1366]" pageId="33" pageNumber="74" pagination="191 - 238" refId="ref48702" refString="LEONARD, E. C. 1936. The Acanthaceae of the Yucatan Peninsula. Carnegie Institute of Washington Publication 461: 191 - 238." type="journal article" year="1936">Leonard (1936:197</bibRefCitation>
, fig. 1).
</paragraph>
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<emphasis id="ECC85270FFC8594AFF7FFAE80707846E" bold="true" box="[171,334,1374,1398]" pageId="33" pageNumber="74">CONSERVATION</emphasis>
</smallCapsWord>
.—
<bibRefCitation id="BA2DF393FFC8594AFEA8FAE8045E846C" author="DANIEL, T. F." box="[380,535,1374,1399]" pageId="33" pageNumber="74" pagination="161 - 174" refId="ref47733" refString="DANIEL, T. F. 1997. Catalog of the Acanthaceae of Belize with taxonomic and phytogeographic notes. Contributions from the University of Michigan Herbarium 21: 161 - 174." type="journal article" year="1997">Daniel (1997)</bibRefCitation>
noted that the population at the
<typeStatus id="010730C0FFC8594AFC4DFAE9058E846C" box="[921,967,1375,1399]" pageId="33" pageNumber="74">type</typeStatus>
locality had been destroyed by quarrying activities, but that other populations had been located on several of the small, isolated, and steep limestone hills in the southern portion of the coastal plain in
<collectingRegion id="1C784080FFC8594AFB85FA16068284C2" country="Belize" name="Belize" pageId="33" pageNumber="74">Belize District</collectingRegion>
.
<taxonomicName id="19BCF5E1FFC8594AFF0EFA7707AD84C2" box="[218,484,1473,1497]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="33" pageNumber="74" phylum="Tracheophyta" rank="species" species="chartaceum">
<emphasis id="ECC85270FFC8594AFF0EFA7707AD84C2" box="[218,484,1473,1497]" italics="true" pageId="33" pageNumber="74">Louteridium chartaceum</emphasis>
</taxonomicName>
has since (i.e., in 2005) been located in
<collectingRegion id="1C784080FFC8594AFC76FA77027A84C2" box="[930,1075,1473,1497]" country="Belize" name="Cayo" pageId="33" pageNumber="74">Cayo District</collectingRegion>
, as well. Plants occur in at least three protected areas, and the species EOO (
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<superScript id="29C9232AFFC8594AFC75FA5705E384E9" attach="left" box="[929,938,1505,1522]" fontSize="7" pageId="33" pageNumber="74">2</superScript>
) includes a portion of another one. Ecologist and botanist Steven Brewer (in litt.,
<date id="AA02A8A2FFC8594AFD25F9B405378701" box="[753,894,1538,1562]" pageId="33" pageNumber="74" value="2018-05-14">14 May 2018</date>
) indicates that the species “is common though not abundant on ridge-tops and exposed limestone along the north side of the Maya Mountains in
<collectingRegion id="1C784080FFC8594AFE84F9F507C18740" box="[336,392,1603,1627]" country="Belize" name="Cayo" pageId="33" pageNumber="74">Cayo</collectingRegion>
and
<collectingCountry id="A6ABCEF2FFC8594AFE6FF9F507B68740" box="[443,511,1603,1627]" name="Belize" pageId="33" pageNumber="74">Belize</collectingCountry>
districts.” Brewer (in litt.,
<date id="AA02A8A2FFC8594AFCCFF9F505E18740" box="[795,936,1603,1628]" pageId="33" pageNumber="74" value="2018-05-22">22 May 2018</date>
) also notes that in this region, potential threats to all of the limestone forests include land conversion via agriculture (both in and around protected areas), encroachment or settlement in protected areas, and fire (either natural or “escaping” flames from regular burning used to clear nearby agricultural fields or bush). In the same communication, Brewer also observed that, “
<taxonomicName id="19BCF5E1FFCB5949FD31FF6A05C981E8" box="[741,896,219,243]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="34" pageNumber="75" phylum="Tracheophyta" rank="species" species="chartaceum">
<emphasis id="ECC85270FFCB5949FD31FF6A05C981E8" box="[741,896,219,243]" italics="true" pageId="34" pageNumber="75">L. chartaceum</emphasis>
</taxonomicName>
(though I doubt the same is true for
<taxonomicName id="19BCF5E1FFCB5949FF2CFF4A07F3800F" box="[248,442,252,276]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="34" pageNumber="75" phylum="Tracheophyta" rank="species" species="donnell-smithii">
<emphasis id="ECC85270FFCB5949FF2CFF4A07F3800F" box="[248,442,252,276]" italics="true" pageId="34" pageNumber="75">L. donnell-smithii</emphasis>
</taxonomicName>
) is surprisingly resilient from the effects of even very hot fires. Natural light fires are occasional on those limestone hills and so it seems that most species there are tolerant or resilient in the face of light fires.” Scant change in vegetation cover within the EOO is evident from the historical imagery (1969 to 2016) seen via
<bibRefCitation id="BA2DF393FFCB5949FCDAFEE8025C806D" author="GOOGLE EARTH PRO" box="[782,1045,350,374]" pageId="34" pageNumber="75" refId="ref48440" refString="GOOGLE EARTH PRO. 2018. Version 7. 3. 1. 4507 (64 - bit). &lt;https: // earth. google. com / download-earth. html&gt; [accessed 24 May 2018]" type="url" year="2018">Google Earth Pro (2018)</bibRefCitation>
. If the three protected areas in which the species occurs are treated either as a single or multiple locations, and the unprotected areas are treated as another location under threat (i.e., conversion of nearby, and partly within the EOO, land to agriculture as an inferred or projected threat), a preliminary conservation assessment of Endangered (EN; B1, a, b) can be proposed for
<taxonomicName id="19BCF5E1FFCB5949FC7CFE57020A80E2" box="[936,1091,481,505]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="34" pageNumber="75" phylum="Tracheophyta" rank="species" species="chartaceum">
<emphasis id="ECC85270FFCB5949FC7CFE57020A80E2" box="[936,1091,481,505]" italics="true" pageId="34" pageNumber="75">L. chartaceum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFCB5949FF7FFDB704C982F9" pageId="34" pageNumber="75" type="discussion">
<paragraph id="DE038E62FFCB5949FF7FFDB7027D8360" blockId="34.[123,1174,219,995]" pageId="34" pageNumber="75">
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<emphasis id="ECC85270FFCB5949FF7FFDB707638303" bold="true" box="[171,298,513,537]" pageId="34" pageNumber="75">DISCUSSION</emphasis>
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.— Plants grown in San Francisco from cuttings of
<taxonomicName id="19BCF5E1FFCB5949FC86FDB702128302" box="[850,1115,513,537]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="34" pageNumber="75" phylum="Tracheophyta" rank="species" species="chartaceum">
<emphasis id="ECC85270FFCB5949FC86FDB702128302" box="[850,1115,513,537]" italics="true" pageId="34" pageNumber="75">Louteridium chartaceum</emphasis>
</taxonomicName>
from the
<typeStatus id="010730C0FFCB5949FF76FD9406998321" box="[162,208,546,570]" pageId="34" pageNumber="75">type</typeStatus>
locality (
<emphasis id="ECC85270FFCB5949FEE2FD9404208321" box="[310,617,546,570]" italics="true" pageId="34" pageNumber="75">Daniel &amp; Butterwick 5905cv</emphasis>
) produced floral buds that attained
<quantity id="19442387FFCB5949FC32FD9402798322" box="[998,1072,546,570]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="34" pageNumber="75" unit="mm" value="35.0">35 mm</quantity>
in length but abscised prior to opening. Both thyrses and racemose thyrses are present on
<emphasis id="ECC85270FFCB5949FC1FFDF502C78341" box="[971,1166,578,603]" italics="true" pageId="34" pageNumber="75">Brewer et al. 7176</emphasis>
. Bracteoles of a pair are sometimes fused basally for about
<quantity id="19442387FFCB5949FD26FDD505798360" box="[754,816,611,635]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="75" unit="mm" value="1.0">1 mm</quantity>
, at least along one side.
</paragraph>
<paragraph id="DE038E62FFCB5949FF7FFD3204C982F9" blockId="34.[123,1174,219,995]" pageId="34" pageNumber="75">
<emphasis id="ECC85270FFCB5949FF7FFD3204688381" bold="true" box="[171,545,644,668]" pageId="34" pageNumber="75">
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</emphasis>
.—
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<collectingCountry id="A6ABCEF2FFCB5949FD9DFD3204EC8380" box="[585,677,644,667]" name="Belize" pageId="34" pageNumber="75">BELIZE</collectingCountry>
.
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<collectingRegion id="1C784080FFCB5949FD66FD3204BF8387" box="[690,758,644,668]" country="Belize" name="Belize" pageId="34" pageNumber="75">Belize</collectingRegion>
</emphasis>
:
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,
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<collectionCode id="B8AD16A7FFCB5949FBBCFD3202318380" box="[1128,1144,644,667]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="34" pageNumber="75" type="Herbarium">E</collectionCode>
of Coastal Road
</locationDeviation>
,
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,
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,
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<emphasis id="ECC85270FFCB5949FD9FFD13041183A7" box="[587,600,677,700]" italics="true" pageId="34" pageNumber="75">S</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFCB5949FD8CFD13041683A7" box="[600,607,677,700]" italics="true" pageId="34" pageNumber="75">.</emphasis>
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<collectingCounty id="3762F6EEFFCB5949FDB3FD1304FC83A7" box="[615,693,677,700]" pageId="34" pageNumber="75">Brewer</collectingCounty>
et al. 7176
</emphasis>
(MO-image seen); along
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near milepost 18, ca.
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<collectionCode id="B8AD16A7FFCB5949FE0CFD7007B883C6" box="[472,497,710,733]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="34" pageNumber="75" type="Herbarium">W</collectionCode>
of
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toward
<collectorName id="7349EBB4FFCB5949FD31FD7005C183C6" box="[741,904,709,733]" pageId="34" pageNumber="75">La Democracia</collectorName>
, ca.
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,
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,
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<emphasis id="ECC85270FFCB5949FFAFFD5106C183E5" box="[123,136,743,766]" italics="true" pageId="34" pageNumber="75">T</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFCB5949FF5CFD5106C683E5" box="[136,143,743,766]" italics="true" pageId="34" pageNumber="75">.</emphasis>
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<emphasis id="ECC85270FFCB5949FF4CFD50076983E5" box="[152,288,742,766]" italics="true" pageId="34" pageNumber="75">
<location id="DB63D8B9FFCB5949FF4CFD5006A883E5" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FF4CFD5006A883E5" box="[152,225,742,766]" country="United States of America" name="Daniel" pageId="34" pageNumber="75">Daniel</location>
8294
</emphasis>
(
<collectionCode id="B8AD16A7FFCB5949FEE7FD51071E83E5" box="[307,343,743,766]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="34" pageNumber="75">BR</collectionCode>
,
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,
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,
<collectionCode id="B8AD16A7FFCB5949FDDFFD50041883E5" box="[523,593,742,766]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="34" pageNumber="75" type="Herbarium">MICH</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FDB5FD5004C583E5" box="[609,652,742,766]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="34" pageNumber="75" type="Herbarium">MO</collectionCode>
,
<collectingCountry id="A6ABCEF2FFCB5949FD49FD5004F683E5" box="[669,703,742,766]" name="United States of America" pageId="34" pageNumber="75">US</collectingCountry>
)
</materialsCitation>
;
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<collectingCounty id="3762F6EEFFCB5949FD0DFD5005DF83E5" box="[729,918,742,766]" pageId="34" pageNumber="75">Gracie Rock Hill</collectingCounty>
near
<collectingMunicipality id="3E671418FFCB5949FC0CFD5002DF83E5" box="[984,1174,742,766]" pageId="34" pageNumber="75">Rockville Quarry</collectingMunicipality>
between
<location id="DB63D8B9FFCB5949FF34FCB107358205" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FF34FCB107358205" box="[224,380,775,799]" country="United States of America" county="Gracie Rock Hill" latitude="17.38713" longLatPrecision="1" longitude="-88.45053" municipality="Rockville Quarry" name="Western Hwy." pageId="34" pageNumber="75">Western Hwy.</location>
and
<location id="DB63D8B9FFCB5949FE6FFCB104098204" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FE6FFCB104098204" box="[443,576,775,799]" country="United States of America" county="Gracie Rock Hill" latitude="17.38713" longLatPrecision="1" longitude="-88.45053" municipality="Rockville Quarry" name="Sibun River" pageId="34" pageNumber="75">Sibun River</location>
, ca.
<locationDeviation id="CFB19740FFCB5949FDABFCB105E38204" box="[639,938,775,799]" location="Belize City" pageId="34" pageNumber="75">
<quantity id="19442387FFCB5949FDABFCB104818204" box="[639,712,775,799]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="75" unit="km" value="30.0">30 km</quantity>
SW of Belize City
</locationDeviation>
[ca.
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,
<geoCoordinate id="BB88E8A5FFCB5949FFAFFC9E076D8224" box="[123,292,808,832]" degrees="088" direction="west" minutes="27" orientation="longitude" pageId="34" pageNumber="75" precision="1" seconds="1. 91" value="-88.45053">088°271.91”W</geoCoordinate>
],
<collectionCode id="B8AD16A7FFCB5949FEEEFC9E070E8224" box="[314,327,808,831]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="75">
<emphasis id="ECC85270FFCB5949FEEEFC9E070E8224" box="[314,327,808,831]" italics="true" pageId="34" pageNumber="75">T</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFCB5949FE93FC9E07928224" box="[327,475,807,831]" italics="true" pageId="34" pageNumber="75">
.
<materialsCitation id="6ED4843FFFCB5949FE80FC91079D8224" box="[340,468,807,831]" collectionCode="M" country="United States of America" location="Daniel" pageId="34" pageNumber="75" specimenCount="1">
<location id="DB63D8B9FFCB5949FE80FC9107D48224" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FE80FC9107D48224" box="[340,413,807,831]" country="United States of America" name="Daniel" pageId="34" pageNumber="75">Daniel</location>
&amp;
<collectionCode id="B8AD16A7FFCB5949FE6AFC9E079D8224" box="[446,468,808,831]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="34" pageNumber="75" type="Herbarium">M</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="6ED4843FFFCB5949FE35FC91047F829A" collectionCode="C, CAS, K, MICH, MO, NY, S, J" collectorName="San Francisco &amp; Gracie Rock" country="United States of America" location="Western Hwy." municipality="Butterwick" pageId="34" pageNumber="75" specimenCount="1" stateProvince="California">
<emphasis id="ECC85270FFCB5949FE35FC9104D9825B" box="[481,656,807,832]" italics="true" pageId="34" pageNumber="75">
<collectingMunicipality id="3E671418FFCB5949FE35FC91041C8224" box="[481,597,807,831]" pageId="34" pageNumber="75">Butterwick</collectingMunicipality>
5905
</emphasis>
(
<collectionCode id="B8AD16A7FFCB5949FD74FC9E04FB825B" box="[672,690,808,832]" country="Denmark" name="University of Copenhagen" pageId="34" pageNumber="75" type="Herbarium">C</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FD6AFC9E04BB825B" box="[702,754,808,832]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="75" type="Museum">CAS</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FD2BFC9E055B8224" box="[767,786,808,831]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="34" pageNumber="75" type="Herbarium">K</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FCCBFC9E052C825B" box="[799,869,808,832]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="34" pageNumber="75" type="Herbarium">MICH</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FCA6FC9E05D4825B" box="[882,925,808,832]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="34" pageNumber="75" type="Herbarium">MO</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FC7EFC9E05988224" box="[938,977,808,831]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="34" pageNumber="75" type="Herbarium">NY</collectionCode>
), cultivated plants of this collection grown from cuttings in
<collectorName id="7349EBB4FFCB5949FDEFFCFE049A827B" box="[571,723,840,864]" pageId="34" pageNumber="75">San Francisco</collectorName>
,
<collectingRegion id="1C784080FFCB5949FD36FCFE0507827B" box="[738,846,840,864]" country="United States of America" name="California" pageId="34" pageNumber="75">California</collectingRegion>
,
<emphasis id="ECC85270FFCB5949FC89FCFE05E3827B" box="[861,938,840,864]" italics="true" pageId="34" pageNumber="75">5905cv</emphasis>
(
<collectionCode id="B8AD16A7FFCB5949FC6FFCFE05A6827B" box="[955,1007,840,864]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="75" type="Museum">CAS</collectionCode>
);
<collectorName id="7349EBB4FFCB5949FBD3FCFE02C7827B" box="[1031,1166,840,864]" pageId="34" pageNumber="75">Gracie Rock</collectorName>
, 1.5
<locationDeviation id="CFB19740FFCB5949FF7DFCDF0708829A" box="[169,321,873,897]" location="Mile" pageId="34" pageNumber="75">
<quantity id="19442387FFCB5949FF7DFCDF0691829A" box="[169,216,873,897]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="34" pageNumber="75" unit="mi" value="4.0">4 mi</quantity>
<collectionCode id="B8AD16A7FFCB5949FF0AFCDF06A4829A" box="[222,237,873,897]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="34" pageNumber="75" type="Herbarium">S</collectionCode>
of Mile
</locationDeviation>
22 on
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,
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10959
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(
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,
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,
<collectionCode id="B8AD16A7FFCB5949FCBCFCDF05DA829A" box="[872,915,873,897]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="34" pageNumber="75" type="Herbarium">MO</collectionCode>
),
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<emphasis id="ECC85270FFCB5949FC7DFCDC05F0829A" box="[937,953,874,897]" italics="true" pageId="34" pageNumber="75">R</emphasis>
</collectionCode>
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<emphasis id="ECC85270FFCB5949FC6DFCDC0200829A" box="[953,1097,873,897]" italics="true" pageId="34" pageNumber="75">
.
<materialsCitation id="6ED4843FFFCB5949FC12FCDC020B829A" box="[966,1090,873,897]" collectionCode="J" country="United States of America" location="Liesner" pageId="34" pageNumber="75" specimenCount="1">
<location id="DB63D8B9FFCB5949FC12FCDC025F829A" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FC12FCDC025F829A" box="[966,1046,874,897]" country="United States of America" name="Liesner" pageId="34" pageNumber="75">Liesner</location>
&amp;
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</materialsCitation>
.
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<materialsCitation id="6ED4843FFFCB5949FB9AFCDC049A82D9" collectionCode="BM, DUKE, MO, NY, TEX, W" collectorName="Cayo &amp; Yaha" country="United States of America" county="Dwyer" location="Banks of Indian Creek" municipality="Manatee Forest Reserve" pageId="34" pageNumber="75" specimenCount="1">
<emphasis id="ECC85270FFCB5949FB9AFCDC06F982B9" italics="true" pageId="34" pageNumber="75">
<collectingCounty id="3762F6EEFFCB5949FB9AFCDC02DC829A" box="[1102,1173,874,897]" pageId="34" pageNumber="75">Dwyer</collectingCounty>
1485
</emphasis>
(
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,
<collectionCode id="B8AD16A7FFCB5949FF29FC3C070E82BA" box="[253,327,906,929]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15531" lsid="urn:lsid:biocol.org:col:15531" name="Duke University" pageId="34" pageNumber="75" type="Herbarium">DUKE</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FE83FC3C07CB82B9" box="[343,386,906,930]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="34" pageNumber="75" type="Herbarium">MO</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FE47FC3C07FF82BA" box="[403,438,906,929]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="34" pageNumber="75" type="Herbarium">NY</collectionCode>
,
<collectionCode id="B8AD16A7FFCB5949FE12FC3C07B382BA" box="[454,506,906,929]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15684" lsid="urn:lsid:biocol.org:col:15684" name="University of Texas at Austin" pageId="34" pageNumber="75" type="Herbarium">TEX</collectionCode>
).
<emphasis id="ECC85270FFCB5949FDC7FC3F040782BA" bold="true" box="[531,590,905,929]" pageId="34" pageNumber="75">
<collectorName id="7349EBB4FFCB5949FDC7FC3F040782BA" box="[531,590,905,929]" pageId="34" pageNumber="75">Cayo</collectorName>
</emphasis>
:
<collectingMunicipality id="3E671418FFCB5949FD8BFC3C052E82BA" box="[607,871,906,929]" pageId="34" pageNumber="75">Manatee Forest Reserve</collectingMunicipality>
,
<location id="DB63D8B9FFCB5949FCACFC3F023B82BA" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FCACFC3F023B82BA" box="[888,1138,905,929]" country="United States of America" county="Dwyer" municipality="Manatee Forest Reserve" name="Banks of Indian Creek" pageId="34" pageNumber="75">Banks of Indian Creek</location>
, at confluence of
<collectorName id="7349EBB4FFCB5949FEFBFC1C072F82D9" box="[303,358,938,962]" pageId="34" pageNumber="75">Yaha</collectorName>
and
<location id="DB63D8B9FFCB5949FE66FC1C040482D9" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FE66FC1C040482D9" box="[434,589,938,962]" country="United States of America" county="Dwyer" municipality="Manatee Forest Reserve" name="Indian Creek" pageId="34" pageNumber="75">Indian Creek</location>
,
<quantity id="19442387FFCB5949FDB2FC1C04E082D9" box="[614,681,938,962]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="75" unit="km" value="1.0">1 km</quantity>
<collectionCode id="B8AD16A7FFCB5949FD6EFC1D049A82D9" box="[698,723,939,962]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="34" pageNumber="75" type="Herbarium">W</collectionCode>
</materialsCitation>
<materialsCitation id="6ED4843FFFCB5949FD31FC1C070A82F8" collectionCode="H" country="United States of America" latitude="17.215055" location="Daylight Cave" longLatPrecision="1" longitude="-88.55539" pageId="34" pageNumber="75" specimenCount="1">
from
<location id="DB63D8B9FFCB5949FCF9FC1C059182D9" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FCF9FC1C059182D9" box="[813,984,938,962]" country="United States of America" latitude="17.215055" longLatPrecision="1" longitude="-88.55539" name="Daylight Cave" pageId="34" pageNumber="75">Daylight Cave</location>
,
<geoCoordinate id="BB88E8A5FFCB5949FC26FC1C02C782D9" box="[1010,1166,938,962]" degrees="17" direction="north" minutes="12" orientation="latitude" pageId="34" pageNumber="75" precision="1" seconds="54. 2" value="17.215055">17°1254.2”N</geoCoordinate>
,
<geoCoordinate id="BB88E8A5FFCB5949FFAFFC7D076B82F9" box="[123,290,971,995]" degrees="088" direction="west" minutes="33" orientation="longitude" pageId="34" pageNumber="75" precision="1" seconds="19. 4" value="-88.55539">088°3319.4”W</geoCoordinate>
,
<collectionCode id="B8AD16A7FFCB5949FEE4FC7A070A82F8" box="[304,323,972,995]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="34" pageNumber="75" type="Herbarium">
<emphasis id="ECC85270FFCB5949FEE4FC7A070A82F8" box="[304,323,972,995]" italics="true" pageId="34" pageNumber="75">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="ECC85270FFCB5949FE97FC7A070382F8" box="[323,330,972,995]" italics="true" pageId="34" pageNumber="75">.</emphasis>
<materialsCitation id="6ED4843FFFCB5949FE85FC7D043082F8" box="[337,633,971,995]" collectionCode="BM" country="United States of America" location="Harris" municipality="Baden" pageId="34" pageNumber="75" specimenCount="1">
<emphasis id="ECC85270FFCB5949FE85FC7D047F82F8" box="[337,566,971,995]" italics="true" pageId="34" pageNumber="75">
<collectingMunicipality id="3E671418FFCB5949FE85FC7D07DC82F8" box="[337,405,971,995]" pageId="34" pageNumber="75">Baden</collectingMunicipality>
&amp; D.
<location id="DB63D8B9FFCB5949FE0EFC7A046882F8" LSID="urn:lsid:plazi:treatment:56153F74FFF65949FF7FFAB604C982F9:DB63D8B9FFCB5949FE0EFC7A046882F8" box="[474,545,972,995]" country="United States of America" municipality="Baden" name="Harris" pageId="34" pageNumber="75">Harris</location>
3
</emphasis>
(
<collectionCode id="B8AD16A7FFCB5949FD92FC7D042682F9" box="[582,623,971,994]" country="United Kingdom" name="Bristol Museum" pageId="34" pageNumber="75">BM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,383 @@
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<mods:title id="9F2D466F1EF84AEEA6837CADEB67183B">Louteridium (Acanthaceae: Acanthoideae: Ruellieae: Trichantherinae): Taxonomy, Phylogeny, Reproductive Biology, and Conservation</mods:title>
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A. Richardson,
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17:66. 1972
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[ca.
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(flr, frt),
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et al. 15825
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(
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:
<collectingCountry id="A6ABCEF2FFF75975FD51FDAD04EE8328" box="[645,679,539,563]" name="United States of America" pageId="30" pageNumber="71">US</collectingCountry>
!;
<typeStatus id="010730C0FFF75975FD6BFDAD05458328" box="[703,780,539,563]" pageId="30" pageNumber="71" type="isotype">isotype</typeStatus>
:
<collectionCode id="B8AD16A7FFF75975FCCFFDAA050B8328" box="[795,834,540,563]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="30" pageNumber="71" type="Herbarium">NY</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFF75974FF7FFDFE021F8028" lastPageId="31" lastPageNumber="72" pageId="30" pageNumber="71" type="description">
<paragraph id="DE038E62FFF75975FF7FFDFE0766878A" blockId="30.[123,1174,583,1682]" pageId="30" pageNumber="71">
Trees to
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tall, mostly epipetric. Older (woody) stems subquadrate to terete, irregularly fissured, sometimes ± scurfy, often lenticellate, lacking trichomes; younger (herbaceous) stems terete (when fresh), becoming flattened on drying, sparsely lenticellate, irregularly fissured, lacking trichomes. Leaves seasonally deciduous, clustered at shoot apex, petiolate, petioles to
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long, sometimes tinged with pink or red, blades subsucculent, narrowly ovate to ovate to elliptic,
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long,
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wide, 1.82.7 × longer than wide, acute to acuminate at apex, acute to attenuate at base, surfaces glabrous (or rarely with very few scattered flexuose eglandular trichomes mostly on veins on abaxial surface, the trichomes especially evident on immature leaves), midvein sometimes pinkish to reddish purple (especially adaxially), margin entire. Inflorescence a terminal sessile or pedunculate thyrse to
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long (including peduncle, if present, and excluding corollas), peduncle (if present) pubescent with erect to flexuose glandular trichomes 0.05
<quantity id="19442387FFF75975FBD9FC3B021782BF" box="[1037,1118,909,933]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="71" unit="mm" value="0.7">0.7 mm</quantity>
long (glandular pubescent), rachis glandular pubescent and often with scattered erect to flexuose eglandular trichomes to
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long as well; dichasia opposite or alternate, pedunculate, 1many-flowered, to
<quantity id="19442387FFF75975FF03FC59077B851D" box="[215,306,1007,1031]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="71" unit="mm" value="120.0">120 mm</quantity>
long (excluding corollas), dichasial peduncles
<quantity id="19442387FFF75975FCEEFC5805E6851D" box="[826,943,1006,1030]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="6.0" metricValueMin="1.5" pageId="30" pageNumber="71" unit="mm" value="37.5" valueMax="60.0" valueMin="15.0">1560 mm</quantity>
long, pubescent like rachises. Bracts caducous (rarely present), lanceolate to lance-ovate, 6.57.
<quantity id="19442387FFF75975FC43FBB9059A853D" box="[919,979,1039,1063]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="71" unit="mm" value="5.0">5 mm</quantity>
long, 2.5
<quantity id="19442387FFF75975FB91FBB902DC853D" box="[1093,1173,1039,1063]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="71" unit="mm" value="2.8">2.8 mm</quantity>
wide, abaxially pubescent like rachis (except sometimes with more eglandular trichomes). Bracteoles and secondary bracteoles caducous, lance-subulate to linear to ovate-elliptic,
<quantity id="19442387FFF75975FC28FBE6021F8573" box="[1020,1110,1104,1128]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="30" pageNumber="71" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">27 mm</quantity>
long, 0.9
<quantity id="19442387FFF75975FF7DFBC706AE8593" box="[169,231,1137,1161]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="71" unit="mm" value="2.0">2 mm</quantity>
wide, abaxially pubescent like rachis or with more eglandular trichomes. Flowers pedicellate, pedicels
<quantity id="19442387FFF75975FEF9FB2707E985B2" box="[301,416,1169,1193]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="30" pageNumber="71" unit="mm" value="37.5" valueMax="50.0" valueMin="25.0">2550 mm</quantity>
long, glandular pubescent. Calyx
<quantity id="19442387FFF75975FCC6FB2705CC85B2" box="[786,901,1169,1193]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="30" pageNumber="71" unit="mm" value="13.0" valueMax="16.0" valueMin="10.0">1016 mm</quantity>
long, lobes fused at base for 1.5
<quantity id="19442387FFF75975FF03FB04075585D2" box="[215,284,1202,1226]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="71" unit="mm" value="3.0">3 mm</quantity>
, subhomomorphic to subheteromorphic, membranaceous, lanceolate to ovate, 7.
<quantity id="19442387FFF75975FF5BFB6406BF85F1" box="[143,246,1234,1258]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="30" pageNumber="71" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">512 mm</quantity>
long,
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wide, subequal or with posterior lobe either slightly larger or smaller than lateral lobes, acute at apex, abaxially pubescent with erect to flexuose glandular and eglandular trichomes 0.05
<quantity id="19442387FFF75975FE99FAA207E98430" box="[333,416,1300,1324]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="71" unit="mm" value="0.3">0.3 mm</quantity>
long, posterior lobe subplanar to subconduplicate. Corollas greenish,
<quantity id="19442387FFF75975FFAFFA8206A68457" box="[123,239,1332,1356]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.5" metricValueMin="3.5" pageId="30" pageNumber="71" unit="mm" value="50.0" valueMax="65.0" valueMin="35.0">3565 mm</quantity>
long, externally pubescent with erect to flexuose glandular and eglandular trichomes 0.05
<quantity id="19442387FFF75975FF63FAE307438477" box="[183,266,1365,1389]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="71" unit="mm" value="0.3">0.3 mm</quantity>
long, tube
<quantity id="19442387FFF75975FE5FFAE204498477" box="[395,512,1364,1389]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="30" pageNumber="71" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">2530 mm</quantity>
long, narrow proximal portion
<quantity id="19442387FFF75975FC8BFAE3058E8477" box="[863,967,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="30" pageNumber="71" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">712 mm</quantity>
long,
<quantity id="19442387FFF75975FBC4FAE3023E8477" box="[1040,1143,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="30" pageNumber="71" unit="mm" value="9.0" valueMax="11.0" valueMin="7.0">711 mm</quantity>
in diameter near midpoint, throat
<quantity id="19442387FFF75975FE1EFAC304748496" box="[458,573,1397,1421]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="30" pageNumber="71" unit="mm" value="20.0" valueMax="25.0" valueMin="15.0">1525 mm</quantity>
long,
<quantity id="19442387FFF75975FD57FAC304BE8496" box="[643,759,1397,1421]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="30" pageNumber="71" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">2630 mm</quantity>
in diameter at mouth, lobes spreading to recoiled, ovate to subcircular,
<quantity id="19442387FFF75975FE07FA20040D84B6" box="[467,580,1430,1454]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.4" metricValueMin="1.5" pageId="30" pageNumber="71" unit="mm" value="19.5" valueMax="24.0" valueMin="15.0">1524 mm</quantity>
long,
<quantity id="19442387FFF75975FD52FA2004B184B6" box="[646,760,1430,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.4" metricValueMin="1.6" pageId="30" pageNumber="71" unit="mm" value="20.0" valueMax="24.0" valueMin="16.0">1624 mm</quantity>
wide, rounded and emarginate at apex. Stamens 4,
<quantity id="19442387FFF75975FF23FA00072384D5" box="[247,362,1462,1486]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="30" pageNumber="71" unit="mm" value="72.5" valueMax="80.0" valueMin="65.0">6580 mm</quantity>
long, filaments distally glabrous, pubescent with eglandular trichomes near base, thecae
<quantity id="19442387FFF75975FED0FA61072084F5" box="[260,361,1495,1519]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="30" pageNumber="71" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
long; staminode 0. Style
<quantity id="19442387FFF75975FDAAFA6104B884F5" box="[638,753,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" metricValueMax="7.7" metricValueMin="7.5" pageId="30" pageNumber="71" unit="mm" value="76.0" valueMax="77.0" valueMin="75.0">7577 mm</quantity>
long, glabrous distally, pubescent near base with eglandular trichomes, stigma subequally 2-lobed and ± funnelform, lobes elliptic to subspheric,
<quantity id="19442387FFF75975FF2AF9AE07138734" box="[254,346,1560,1584]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="30" pageNumber="71" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, 1.5
<quantity id="19442387FFF75975FE02F9AE045E8734" box="[470,535,1560,1584]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="71" unit="mm" value="3.0">3 mm</quantity>
wide. Capsules
<quantity id="19442387FFF75975FD00F9AE05038734" box="[724,842,1560,1584]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="30" pageNumber="71" unit="mm" value="37.0" valueMax="42.0" valueMin="32.0">3242 mm</quantity>
long, 69.
<quantity id="19442387FFF75975FC12F9AE024E8734" box="[966,1031,1560,1584]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="71" unit="mm" value="5.0">5 mm</quantity>
in diameter, pubescent with erect to flexuose eglandular and subsessile to stipitate glandular trichomes less than 0.05
<quantity id="19442387FFF75975FF63F9EF074E876A" box="[183,263,1625,1649]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="30" pageNumber="71" unit="mm" value="0.4">0.4 mm</quantity>
long, stipe 25.
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long. Seeds 1624 per capsule, 5.
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long, 4.
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wide, surfaces smooth.
</paragraph>
<paragraph id="DE038E62FFF75974FF7FF910021F8028" blockId="30.[171,1173,1701,1726]" lastBlockId="31.[123,1174,219,1240]" lastPageId="31" lastPageNumber="72" pageId="30" pageNumber="71">
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<emphasis id="ECC85270FFF75975FF7FF910077887A7" bold="true" box="[171,305,1702,1726]" pageId="30" pageNumber="71">PHENOLOGY</emphasis>
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.— Flowering: March; fruiting: March. Field observations on
<emphasis id="ECC85270FFF75974FC31F913074981E8" italics="true" lastPageId="31" lastPageNumber="72" pageId="30" pageNumber="71">Daniel &amp; Steinmann 11913</emphasis>
reveal that no flowers were open between 16:0017:30 on the first day of observations, but several very large floral buds were present. On the following day, all of the large buds were found to be fully open at 08:30, and no visitors were noted over the course of an hour.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFF65974FF7FFE8A026783CE" pageId="31" pageNumber="72" type="distribution">
<paragraph id="DE038E62FFF65974FF7FFE8A07368088" blockId="31.[123,1174,219,1240]" pageId="31" pageNumber="72">
<emphasis id="ECC85270FFF65974FF7FFE8A07948049" bold="true" box="[171,477,316,340]" pageId="31" pageNumber="72">
<smallCapsWord id="D8E518BEFFF65974FF7FFE8A070A8049" baselines="334,333" box="[171,323,316,340]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="31" pageNumber="72">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="D8E518BEFFF65974FE9AFE8907308049" baselines="333" box="[334,377,319,338]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="31" pageNumber="72">AND</smallCapsWord>
<smallCapsWord id="D8E518BEFFF65974FE50FE8907948049" baselines="333" box="[388,477,319,338]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="habitat" pageId="31" pageNumber="72">HABITAT</smallCapsWord>
</emphasis>
.— West-central
<collectingRegion id="1C784080FFF65974FD4FFE8D04A58048" box="[667,748,315,339]" country="Mexico" name="Mexico" pageId="31" pageNumber="72">Mexico</collectingRegion>
(
<collectingRegion id="1C784080FFF65974FCD5FE8D053E8048" box="[769,887,315,339]" country="Mexico" name="Michoacan de Ocampo" pageId="31" pageNumber="72">Michoacán</collectingRegion>
;
<figureCitation id="468792E7FFF65974FC5EFE8D0584804F" box="[906,973,315,340]" captionStart="FIGURE" captionStartId="19.[155,167,950,969]" captionTargetBox="[127,1170,219,938]" captionTargetId="figure-78@19.[120,1176,216,945]" captionTargetPageId="19" captionText="FIGURE 8. Map of Mexico (showing states) and northern Central America with distributions of species of Louteridium endemic to Mexico: L. brevicalyx, L. dendropilosum, L. koelzii, L. parayi, L. rzedowskianum, and L. tamaulipense. Sym- bols may pertain to multiple collections." figureDoi="http://doi.org/10.5281/zenodo.13155723" httpUri="https://zenodo.org/record/13155723/files/figure.png" pageId="31" pageNumber="72">Fig. 8</figureCitation>
); plants occur on exposed limestone, often on steep slopes of arroyos, in tropical subdeciduous forests at elevations from 200 to 590 meters.
</paragraph>
<paragraph id="DE038E62FFF65974FF7FFE2A050280AF" blockId="31.[123,1174,219,1240]" box="[171,843,412,436]" pageId="31" pageNumber="72">
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<emphasis id="ECC85270FFF65974FF7FFE2A071980A9" bold="true" box="[171,336,412,436]" pageId="31" pageNumber="72">ILLUSTRATIONS</emphasis>
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.—
<figureCitation id="468792E7FFF65974FEADFE2A044D80AF" box="[377,516,412,436]" captionStart="FIGURE" captionStartId="5.[155,167,1561,1580]" captionText="FIGURE 2. Louteridium brevicalyx (A-E) and L. dendropilosum (F-L). A. Habit and habitat. B. Trunk with adventitious roots on rocks. C. Six-year old cultivated plant with clusters of new leaves. D. Flower. E. Shoot apex showing clustered leaf scars. F. Base of trunk with adventitious roots on rocks. G. Six-year old cultivated plant with clusters of new leaves. H. Shoot apex showing clustered leaf scars. I Young inflorescence with bracts, bracteoles, and flower buds. J. Flower. K. Corolla showing invagination along base of throat. L. Internal view of corolla showing invaginations along base of throat. (A, B, Daniel &amp; Steinmann 11913; C-E, Daniel &amp; Steinmann 11913cv; F, Daniel et al. 11784; G-L, Daniel et al. 11894cv). Photos by T. Daniel." pageId="31" pageNumber="72">Figure 2AE</figureCitation>
;
<bibRefCitation id="BA2DF393FFF65974FDC7FE2A04BD80AF" author="RICHARDSON, A." box="[531,756,412,436]" pageId="31" pageNumber="72" pagination="63 - 76" refId="ref49361" refString="RICHARDSON, A. 1972. Revision of Louteridium (Acanthaceae). Tulane Studies in Zoology and Botany 17: 63 - 76." type="journal article" year="1972">Richardson (1972:66</bibRefCitation>
, fig. 2).
</paragraph>
<paragraph id="DE038E62FFF65974FF7FFE0A026783CE" blockId="31.[123,1174,219,1240]" pageId="31" pageNumber="72">
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<emphasis id="ECC85270FFF65974FF7FFE0A070780C8" bold="true" box="[171,334,444,468]" pageId="31" pageNumber="72">CONSERVATION</emphasis>
</smallCapsWord>
.— Except for
<taxonomicName id="19BCF5E1FFF65974FE3BFE0B04FB80CF" baseAuthorityName="Kruse" baseAuthorityYear="1380" box="[495,690,444,468]" class="Magnoliopsida" family="Acanthaceae" genus="Louteridium" kingdom="Plantae" order="Lamiales" pageId="31" pageNumber="72" phylum="Tracheophyta" rank="species" species="rzedowskianum">
<emphasis id="ECC85270FFF65974FE3BFE0B04FB80CF" box="[495,690,444,468]" italics="true" pageId="31" pageNumber="72">L. rzedowskianum</emphasis>
</taxonomicName>
, which is known from a single site,
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<emphasis id="ECC85270FFF65974FBEEFE0B06FA80EF" italics="true" pageId="31" pageNumber="72">L. brevicalyx</emphasis>
</taxonomicName>
has the smallest known EOO (
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) among its congeners. At the site of
<emphasis id="ECC85270FFF65974FC38FE6A06B6830E" italics="true" pageId="31" pageNumber="72">Daniel &amp; Steinmann 11913</emphasis>
, about 100 plants were seen growing, mostly epipetrically, in an area of ca.
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<superScript id="29C9232AFFF65974FB58FE4A02DC8316" attach="left" box="[1164,1173,508,525]" fontSize="7" pageId="31" pageNumber="72">2</superScript>
on exposed limestone slopes in forest. No protected land appears to be within the species EOO. Evident threats to this species appear to be large-scale mining in the vicinity of
<emphasis id="ECC85270FFF65974FC3CFD8B06B7836E" italics="true" pageId="31" pageNumber="72">Daniel &amp; Steinmann 11913</emphasis>
(ca.
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distant) and deforestation for agriculture (including cattle) that is evident within the small EOO and in adjacent regions. Given the size of the EOO and apparent major threat (i.e., deforestation for agriculture), a single location is evident. A preliminary conservation assessment of Critically Endangered (CR; B1, a, b;
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) is proposed for
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<emphasis id="ECC85270FFF65974FC49FD08026E83CD" box="[925,1063,702,726]" italics="true" pageId="31" pageNumber="72">L. brevicalyx</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="96A6DDE9FFF65974FF7FFD6805E985C3" pageId="31" pageNumber="72" type="discussion">
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.— Leaves are absent on the field specimens examined. Information on young stems and leaves provided above is from cultivated plants (
<emphasis id="ECC85270FFF65974FD23FD480275820E" box="[759,1084,766,790]" italics="true" pageId="31" pageNumber="72">Daniel &amp; Steinmann 11913cv)</emphasis>
. Leaves of these plants may not have attained the maximum size encountered in natural settings. The inflorescence, which appears before the leaves, arises from the apex of the previous seasons woody growth and bears a stalked (pedunculate) thyrse or a sessile rachis subtended by axillary dichasia (sessile thyrse). A stem forms with the flush of new clustered leaves later in the season.
</paragraph>
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Most calyces have the three lobes that are characteristic of the genus. Five lobes with three prominent lobes and two reduced lobes are evident on
<emphasis id="ECC85270FFF65974FD19FC7605E882CC" box="[717,929,959,983]" italics="true" pageId="31" pageNumber="72">Hinton et al. 15843</emphasis>
; the reduced lobes are
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long and 0.8 to
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wide. Five-lobed calyces in inflorescences are likely teratological and suggest suppression rather than fusion of lobes for typical plants with three-lobed calyces.
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.
</emphasis>
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.
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<collectingRegion id="1C784080FFF65974FD07FBA90518852C" box="[723,849,1055,1079]" country="Mexico" name="Michoacan de Ocampo" pageId="31" pageNumber="72">Michoacán</collectingRegion>
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:
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,
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.7 carretera
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,
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,
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,
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.
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&amp;
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.
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5484
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(
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,
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); between
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and
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,
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of Aquila
</locationDeviation>
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,
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,
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,
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.
<collectorName id="7349EBB4FFF65974FE8CFB3607E88583" box="[344,417,1152,1176]" pageId="31" pageNumber="72">Daniel</collectorName>
&amp;
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.
<collectorName id="7349EBB4FFF65974FE0AFB360406858C" box="[478,591,1152,1175]" pageId="31" pageNumber="72">Steinmann</collectorName>
11913
</emphasis>
(
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,
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,
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,
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,
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,
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)
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, cultivat- ed plants of this collection grown from seed in San Francisco,
<collectingRegion id="1C784080FFF65974FCCBFB1605C285A3" box="[799,907,1184,1208]" country="United States of America" name="California" pageId="31" pageNumber="72">California</collectingRegion>
,
<emphasis id="ECC85270FFF65974FC4EFB1605BB85A3" box="[922,1010,1184,1208]" italics="true" pageId="31" pageNumber="72">11913cv</emphasis>
(CAS); Aquila [ca.
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,
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],
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<collectionCode id="B8AD16A7FFF65974FDFDFB77047585C3" box="[553,572,1217,1240]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="31" pageNumber="72" type="Herbarium">G</collectionCode>
.
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et al. 15843
</emphasis>
(
<collectingCountry id="A6ABCEF2FFF65974FCF8FB7605D985C3" box="[812,912,1216,1240]" isEnumeration="true" pageId="31" pageNumber="72">RSA, US</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,247 @@
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<taxonomicName id="2CE981E59776FFBDFFDFFEB5D22BC078" authority="Meehean, 1940" authorityName="Meehean" authorityYear="1940" box="[123,512,351,375]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="floridensis">
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<emphasis id="D99D26749776FFBDFFDFFEB5D164C078" bold="true" box="[123,335,351,375]" italics="true" pageId="1" pageNumber="134">Argulus floridensis</emphasis>
<bibRefCitation id="8F7887979776FFBDFEF2FEB5D22BC078" author="MEEHEAN, O. L." box="[342,512,351,375]" pageId="1" pageNumber="134" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean, 1940</bibRefCitation>
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<collectingRegion id="292D34849776FFBDFFDFFE95D0EBC09A" box="[123,192,383,405]" country="United States of America" name="Florida" pageId="1" pageNumber="134">Florida</collectingRegion>
fishlouse Figures 16.
</paragraph>
</subSubSection>
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</emphasis>
.—
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(unique), adult male,
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total length [
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Meehean 1940:
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.85 mm],
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77810 [see note], Key West,
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, 1884, collector: A.E. Verrill [= Addison Emery Verrill]. Note:
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, p. 467) listed the catalog number as “77810 (old number 60589);” mentioned here in case of relevant information in historical records under the old number.
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.— Two respiratory areas with smaller oblong “area” anterior to slightly larger oblong posterior “area;” single postantennal spines very broad; male with 77 support rods in suction cup; 1419 sclerites per rod in male; mouth tube with at least 19 scales on basal half; basal plate of second maxilla with 3 stout spines, scales covering nearly entire raised pad, one stout, naked seta on posterior margin of pad; accessory spines on thorax large, very broad base on raised pad with many scales, tapering to rounded tips; first 2 pairs of legs lacking flagella; posteroventral surface of coxae of second legs of male expanded into a broadly-rounded lobe that is covered with many small and closely-arranged scales; third legs of male with two pegs issuing from cleft on dorsal side of legs and two fingerlike papillae; small fleshy lobe at posterior of thorax; male abdomen bilobate with testes extending to the level of the anal sinus base; caudal rami long, slen- der, basal in anal sinus.
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.— Total length
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[
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Meehean 1940:
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.85 mm]. Carapace shape is “ovate, sinuses shallow, cephalic areas broad...; alae reaching abdomen [
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].” Carapace length [
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Meehean 1940:
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.88 mm]. Maximum carapace width [
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Meehean 1940:
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.28 mm]. Pair of compound eyes anteriorly with diameters 250 (left) and 220 (right) µm. Transverse distance between eyes
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(830 µm). Nauplius eye with one anterior and two posterior ocelli. Sclerotized dorsal ridges not forked anterior of eyes. Ventrally, carapace with small, posteriorly-projecting spines along outer margin [due to the deteriorated condition of the carapace alae, additional details cannot be included here]. Respiratory areas consist of two oblong-shaped areas, anterior one slightly smaller than posterior [from
<bibRefCitation id="8F7887979776FFBDFE8CFAC2D1E3C44F" author="MEEHEAN, O. L." box="[296,456,1320,1344]" pageId="1" pageNumber="134" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean 1940</bibRefCitation>
] (Fig. 1). No pigmentation present (but probably lost in preservative and/or due to clearing). Thorax compressed dorsoventrally, 4-segmented, with 2 pairs of posteriorly-projecting spines ventrally. Spines digitate with broad base, anterior pair (accessory spines) larger than posterior pair (postmaxillary spines), both pairs angled mesially. Accessory spines between basal segments of second maxillae; very broad base on a raised pad covered with many scales; spines tapering to round- ed tips (
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). Male with ovoid fleshy lobe at posterior of fourth thoracic segment between natatory lobes (Figs. 3A, 4). Thorax with
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1.
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:
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</paragraph>
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shape of anterior and posterior scales scattered on ventral surface; thoracic segment at base of third respiratory areas (redrawn from leg produced laterally into expanded anterolateral lobe with scales
<bibRefCitation id="8F7887979776FFBDFCC3F967D3CDC7AF" author="MEEHEAN, O. L." box="[871,998,1677,1696]" pageId="1" pageNumber="134" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean 1940</bibRefCitation>
). with secondary sexual structures (Figs.
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3.
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: A) ventral view of legs 2, 3, and 4 3A, B). (precoxa, coxa, and basis only); exopods, endopods, setae, and sensil-
</paragraph>
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2.
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: Second maxilla, raised pad with scales (adjacent on thorax),
</paragraph>
<paragraph id="EB56FA669775FFBEFFDFFD7BD0DEC3B2" blockId="2.[123,549,605,701]" pageId="2" pageNumber="135">accessory spine, and postmaxillary spine. Scale bar = 200 µm.</paragraph>
</caption>
<paragraph id="EB56FA669775FFBEFF0EFB59D2DBC5D1" blockId="2.[122,1174,723,1708]" pageId="2" pageNumber="135">First legs lack secondary modifica- lae omitted. B) dorsal view of leg 3 (precoxa, coxa, and basis only). tions. Second legs, the posterior ventral Scale bar = 500 µm.</paragraph>
<paragraph id="EB56FA669775FFBEFFDEFB19D0CEC484" blockId="2.[122,1174,723,1708]" pageId="2" pageNumber="135">
margin of coxae expanded into broadly-rounded lobe with many small, closely-arranged scales. Third legs, dorsal anterior coxae with broadly-rounded hump armed with many scales directed proximally; distal to this hump are two smaller, unarmed protuberances; at anterior distal margin of coxae an anteriorly-directed, fleshy projection (“upright fingerlike papilla” of
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) (Fig. 3A).
</paragraph>
<paragraph id="EB56FA669775FFBEFF0EFA79D3B9C784" blockId="2.[122,1174,723,1708]" pageId="2" pageNumber="135">
Third legs with two sclerotized pegs in cleft on dorsal side at distal margins of coxae (or fused coxae+bases); pegs with a single, large horn and orifice distally; sclerotized ridge at anterior dorsal of coxa/basis junction; one fleshy papilla extends proximally (mesially) from sclerotized ridge anterior of two pegs and overhanging cleft (“saddle-shaped depression” of
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). Posterior to pegs and cleft a triangular-shaped lobe covered with scales (Fig. 3B). Posterior margins of both precoxae+coxae and bases of fourth legs with natatory lobes bearing plumose setae. Anterior of bases a sclerotized ridge lies on basal half, opposed by recurved, finger-like appendage covered with fine scales distally; tiny spines or scales on its tip (Fig. 3A).
</paragraph>
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Abdomen bilobate. Abdomen length
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[
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Meehean 1940:
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.58 mm]; maximum width
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[
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.35 mm]. Anal sinus length
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(720 µm) [
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:
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]. Caudal rami paired, long, slender, near base of anal sinus (could not see the setae clearly, usually there are 5 simple setae at the tip of each). Tips of abdominal lobes narrowly rounded, not pointy. Testes extending to level of anal sinus base (
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). Scales absent on ventral surface of abdomen; spines or scales absent along margins of abdominal lobes.
</paragraph>
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4.
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: abdomen; fleshy lobe at
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5.
<taxonomicName id="2CE981E59774FFBFFCACFD29D381C3D9" authorityName="Meehean" authorityYear="1940" box="[776,938,707,726]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749774FFBFFCACFD29D381C3D9" box="[776,938,707,726]" italics="true" pageId="3" pageNumber="136">Argulus floridensis</emphasis>
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: first and second antennae. posterior of thorax is shown as a dashed outline at anterior of The fleshy segments of the first antennae were obscured or abdomen. Setae not shown on caudal rami. Scale bar = 500 missing on this structure but are included as dashed outlines. µm. Scale bar = 200 µm.
</paragraph>
</caption>
<paragraph id="EB56FA669774FFBFFF0EFC0BD363C2F6" blockId="3.[121,1174,833,1722]" box="[170,840,993,1018]" pageId="3" pageNumber="136">Mouth tube long, slender, with at least 19 scales on basal half.</paragraph>
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First antennae 4-segmented. First segment (basal segment) sclerotized, large with stout posteriorly-projecting posterior spine, narrowly rounded at tip; second segment sclerotized with small recurved spine anteriorly with narrow, blunt tip, posteriorly-projecting medial spine with rounded tip, and large recurved terminal spine; third and fourth fleshy segments missing or obscured on the
<typeStatus id="345244C49774FFBFFFDEFB6BD0FCC596" box="[122,215,1153,1177]" pageId="3" pageNumber="136" type="holotype">holotype</typeStatus>
(Fig. 5). Width of first antennae (mean of both antennae, µm) 530540 (535) in male. Second antennae 5-segmented, fleshy. First two segments larger; remaining three thin, cylindrical; basal segment bears posteriorly-projecting posterior spine with narrowly-rounded tip. Postantennal spines single (as opposed to double in some taxa), very wide at base, tapering to narrowly-rounded tip (Fig. 5).
</paragraph>
<paragraph id="EB56FA669774FFBFFF0EFAABD14BC716" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">First maxillae modified into suction cups in adult. In male, first maxillae inner diameter 680 µm and outer diameter 1,050 µm (n = 1). Number of sclerites per support rod in male 1419 (n = 25 support rods) [excludes abnormal rods with missing sclerites]. Basal (proximal) sclerite usually subquadrangular, shape variable; distal sclerites imbricate bowls, plates (Fig. 6). Suction cup with 29 sensilla on inside circumference.</paragraph>
<paragraph id="EB56FA669774FFBFFF0EF9CBD4BEC762" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">
Second maxillae 5-segmented with broad basal plate bearing three stout spines with the mesial spine broader than the other two (
<figureCitation id="73D2E6E39774FFBFFF26F98BD094C776" box="[130,191,1633,1657]" captionStart="FIGURE" captionStartId="2.[155,167,605,624]" captionTargetBox="[123,1159,219,1131]" captionTargetId="figure-1945@2.[575,1176,216,1134]" captionTargetPageId="2" captionText="FIGURE 2. Argulus floridensis: Second maxil- la, raised pad with scales (adjacent on thorax), accessory spine, and postmaxillary spine. Scale bar = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13155433" httpUri="https://zenodo.org/record/13155433/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
). Basal plate with elevated pad bearing 43 scales and one stout
<superScript id="1C9C572E9774FFBFFC33F9B0D4BEC762" attach="right" box="[919,1173,1626,1645]" fontSize="8" pageId="3" pageNumber="136">
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6.
<taxonomicName id="2CE981E59774FFBFFC4AF9B0D4A4C762" authorityName="Meehean" authorityYear="1940" box="[1006,1167,1626,1645]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749774FFBFFC4AF9B0D4A4C762" box="[1006,1167,1626,1645]" italics="true" pageId="3" pageNumber="136">Argulus floridensis</emphasis>
</taxonomicName>
:
</superScript>
</paragraph>
<paragraph id="EB56FA669774FFBFFCD3F99ED3FBC7B5" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">Two support rods from suction seta posteriorly. Bi-dentate to multidentate scales on ventral surfaces cup (right first maxilla), Scale bar of segments 2 and 3; segment 4 lacking armature. Distal segment with = 100 µm,</paragraph>
<paragraph id="EB56FA669773FFB8FFDFFF31D1F9C01C" blockId="4.[123,1174,219,374]" pageId="4" pageNumber="137">2 sharp claws (1 broken/missing) and one blunt, elongate lobe positioned above claws (did not see a small sensillum at tip of lobe).</paragraph>
</subSubSection>
<subSubSection id="A3F3A9ED9773FFB8FF0FFEF6D1A8C03B" box="[171,387,284,308]" pageId="4" pageNumber="137" type="biology_ecology">
<paragraph id="EB56FA669773FFB8FF0FFEF6D1A8C03B" blockId="4.[123,1174,219,374]" box="[171,387,284,308]" pageId="4" pageNumber="137">
<smallCapsWord id="EDB06CBA9773FFB8FF0FFEF6D0C2C03C" baselines="302,302" box="[171,233,284,308]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Host" pageId="4" pageNumber="137">
<emphasis id="D99D26749773FFB8FF0FFEF6D0C2C03C" bold="true" box="[171,233,284,308]" pageId="4" pageNumber="137">HOST</emphasis>
</smallCapsWord>
.— Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A3F3A9ED9773FFB8FF0FFED7D41AC079" pageId="4" pageNumber="137" type="etymology">
<paragraph id="EB56FA669773FFB8FF0FFED7D41AC079" blockId="4.[123,1174,219,374]" pageId="4" pageNumber="137">
<smallCapsWord id="EDB06CBA9773FFB8FF0FFED7D11EC05C" baselines="335,334" box="[171,309,317,341]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Etymology" pageId="4" pageNumber="137">
<emphasis id="D99D26749773FFB8FF0FFED7D11EC05C" bold="true" box="[171,309,317,341]" pageId="4" pageNumber="137">ETYMOLOGY</emphasis>
</smallCapsWord>
.— The specific name,
<taxonomicName id="2CE981E59773FFB8FD89FED7D2B5C05A" authorityName="Meehean" authorityYear="1940" box="[557,670,317,341]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749773FFB8FD89FED7D2B5C05A" box="[557,670,317,341]" italics="true" pageId="4" pageNumber="137">floridensis</emphasis>
</taxonomicName>
, is derived from the state,
<collectingRegion id="292D34849773FFB8FC1CFED7D42EC05A" box="[952,1029,317,341]" country="United States of America" name="Florida" pageId="4" pageNumber="137">Florida</collectingRegion>
,
<materialsCitation id="5B81F03B9773FFB8FBB6FED7D41AC079" collectionCode="Host" location="Florida" pageId="4" pageNumber="137" specimenCount="1" stateProvince="Florida">
in which the
<typeStatus id="345244C49773FFB8FFDFFEB4D082C079" box="[123,169,350,374]" pageId="4" pageNumber="137">type</typeStatus>
locality is located and has an adjectival suffix, ensis, meaning “of or from
<collectingRegion id="292D34849773FFB8FC75FEB7D435C07A" box="[977,1054,349,373]" country="United States of America" name="Florida" pageId="4" pageNumber="137">Florida</collectingRegion>
.”
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,123 @@
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<mods:title id="FC1EB8026FF63854BE99D54018656FE3">Atlas of the Reptiles of Libya</mods:title>
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<heading id="B64481ECFFC5FFDBFFADFB3AFD9F0ACF" box="[123,544,1195,1220]" fontSize="10" level="2" pageId="110" pageNumber="265" reason="4">
<treatmentCitationGroup id="CDA311AEFFC5FFDBFFADFB3AFD9F0ACF" box="[123,544,1195,1220]" pageId="110" pageNumber="265">
<treatmentCitation id="6C121091FFC5FFDBFFADFB3AFD9F0ACF" author="GMELIN, J. F." box="[123,544,1195,1220]" page="1041" pageId="110" pageNumber="265" year="1789">
<taxonomicName id="2AB34D03FFC5FFDBFFADFB3AFD9F0ACF" ID-CoL="7C3D6" authority="Gmelin, 1789: 1041" authorityName="Gmelin, 1789:" authorityPageNumber="1041" authorityYear="1041" box="[123,544,1195,1220]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="hermanni">
<emphasis id="DFC7EA92FFC5FFDBFFADFB3AFD9F0ACF" bold="true" box="[123,544,1195,1220]" pageId="110" pageNumber="265">
<emphasis id="DFC7EA92FFC5FFDBFFADFB3AFEFC0AC8" bold="true" box="[123,323,1195,1219]" italics="true" pageId="110" pageNumber="265">Testudo hermanni</emphasis>
<bibRefCitation id="89224B71FFC5FFDBFE9CFB3DFD9F0ACF" author="GMELIN, J. F." box="[330,544,1196,1220]" pageId="110" pageNumber="265" pagination="1038 - 1516" refId="ref86212" refString="GMELIN, J. F. 1789. Caroli a Linne ... Systema Naturae per Regna Tria Natural, Secundum Classes, Ordines, Genera, Species, cum Characteribus Differentilis, Synonymis, Locis. Tomus I, Editio Decima Tertia, Aucta, Reformata. Pars III. Amphibia et Pisces. Georg. Emanuel Beer, Lipsiae [Leipzig], Germany. Pp. 1038 - 1516." type="book chapter" year="1789">Gmelin, 1789:1041</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="ED0C3680FFC5FFDBFF7DFB47FD500B25" blockId="110.[123,1173,1238,1327]" pageId="110" pageNumber="265">
Reported by
<bibRefCitation id="89224B71FFC5FFDBFEE0FB47FDAD0AE4" author="LUISELLI, L. &amp; F. CIVETTA &amp; S. MASCIOLA &amp; G. MILANA &amp; G. AMORI" box="[310,530,1238,1263]" pageId="110" pageNumber="265" pagination="365 - 380" refId="ref91827" refString="LUISELLI, L., F. CIVETTA, S. MASCIOLA, G. MILANA, AND G. AMORI. 2012. Spatial turnover and knowledge gaps for African chelonians mirror those of African small mammals: conservation implications. Tropical Conservation Science, 5 (3): 365 - 380." type="journal article" year="2012">Luiselli et al. (2012)</bibRefCitation>
from
<collectingCountry id="95A47610FFC5FFDBFD83FB47FD2C0AE5" box="[597,659,1238,1262]" name="Libya" pageId="110" pageNumber="265">Libya</collectingCountry>
, but without specific data. Although introduced individuals may exist, there are no documented records of this species outside of southern Europe and its associated islands (Sindaco and Jeremčenko 2008).
</paragraph>
</subSubSection>
<subSubSection id="A5A9650BFFC5FFDBFFADFADFFCF1081B" pageId="110" pageNumber="265" type="reference_group">
<paragraph id="ED0C3680FFC5FFDBFFADFADFFE530B8D" blockId="110.[123,1173,1358,1414]" pageId="110" pageNumber="265">
<emphasis id="DFC7EA92FFC5FFDBFFADFADFFE530B8D" bold="true" pageId="110" pageNumber="265">
<taxonomicName id="2AB34D03FFC5FFDBFFADFADFFD920B6D" ID-CoL="55JTW" authority="Schoepff, 1793: 58" authorityName="Schoepff" authorityPageNumber="58" authorityYear="1793" box="[123,557,1358,1383]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="marginata">
<emphasis id="DFC7EA92FFC5FFDBFFADFADFFEF00B6D" bold="true" box="[123,335,1358,1382]" italics="true" pageId="110" pageNumber="265">Testudo marginata</emphasis>
Schoepff, 1793:58
</taxonomicName>
, pl. XI, pl. XII, fig. 1. [German edition]/Schoepff, 1793:52, pl. XI, pl. XII, fig. 1.
</emphasis>
</paragraph>
<paragraph id="ED0C3680FFC5FFDBFF7DFA08FCF1081B" blockId="110.[122,1173,1433,1553]" pageId="110" pageNumber="265">
Reported by
<bibRefCitation id="89224B71FFC5FFDBFEE1FA08FDAA0BBA" author="LUISELLI, L. &amp; F. CIVETTA &amp; S. MASCIOLA &amp; G. MILANA &amp; G. AMORI" box="[311,533,1433,1457]" pageId="110" pageNumber="265" pagination="365 - 380" refId="ref91827" refString="LUISELLI, L., F. CIVETTA, S. MASCIOLA, G. MILANA, AND G. AMORI. 2012. Spatial turnover and knowledge gaps for African chelonians mirror those of African small mammals: conservation implications. Tropical Conservation Science, 5 (3): 365 - 380." type="journal article" year="2012">Luiselli et al. (2012)</bibRefCitation>
from
<collectingCountry id="95A47610FFC5FFDBFD8FFA08FD280BBA" box="[601,663,1433,1457]" name="Libya" pageId="110" pageNumber="265">Libya</collectingCountry>
, but without specific data.
<bibRefCitation id="89224B71FFC5FFDBFC17FA08FBF70BBA" author="PETERS, W. C. H." box="[961,1096,1433,1457]" pageId="110" pageNumber="265" pagination="305 - 309" refId="ref94694" refString="PETERS, W. C. H. 1880. Uber die von Hrn. Gerhard Rohlfs und Dr. A. Stecker auf der Reise nach der Oase Kufra gesammelten Amphibien. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, 1880: 305 - 309, 1 pl. (part)." type="journal article" year="1880">Peters (1880</bibRefCitation>
,
<bibRefCitation id="89224B71FFC5FFDBFB81FA08FB330BBA" author="PETERS, W. C. H." box="[1111,1164,1433,1457]" pageId="110" pageNumber="265" refId="ref94750" refString="PETERS, W. C. H. 1881. Amphibien der Expedition nach Kufra. Pages 365 - 369 in Rohlfs, G., Kufra. Reise von Tripolis nach der Oase Kufra. F. A. Brockhaus, Leipzig, Germany. viii + 559 pp., 7 plates, 22 foldout tables, 3 foldout maps." type="book" year="1881">1881</bibRefCitation>
) reported
<taxonomicName id="2AB34D03FFC5FFDBFF0BFA28FE390BDA" ID-CoL="55JQ4" authorityName="Strauch" authorityYear="1862" box="[221,390,1465,1489]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="campanulata">
<emphasis id="DFC7EA92FFC5FFDBFF0BFA28FE390BDA" box="[221,390,1465,1489]" italics="true" pageId="110" pageNumber="265">T. campanulata</emphasis>
</taxonomicName>
[=
<taxonomicName id="2AB34D03FFC5FFDBFE65FA28FD810BDB" ID-CoL="55JTW" authorityName="Schoepff" authorityYear="1793" box="[435,574,1465,1488]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="marginata">
<emphasis id="DFC7EA92FFC5FFDBFE65FA28FD810BDB" box="[435,574,1465,1488]" italics="true" pageId="110" pageNumber="265">T. marginata</emphasis>
</taxonomicName>
] from Bir-Milrha. However, Schneider and
<bibRefCitation id="89224B71FFC5FFDBFBFCFA28FF7D0BFA" author="SCHNEIDER, C. &amp; W. SCHNEIDER" pageId="110" pageNumber="265" pagination="141 - 152" refId="ref96910" refString="SCHNEIDER, C., AND W. SCHNEIDER. 2008. The Egyptian tortoise, Testudo kleinmanni Lortet, 1883 in Libya. Salamandra, 44 (3): 141 - 152." type="journal article" year="2008">Schneider (2008)</bibRefCitation>
argued that the likely species intended was
<taxonomicName id="2AB34D03FFC5FFDBFD75FA48FC8B0BFA" ID-CoL="55JTJ" authorityName=", Sindaco and Jeremcenko" authorityYear="2008" box="[675,820,1497,1521]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="kleinmanni">
<emphasis id="DFC7EA92FFC5FFDBFD75FA48FC8B0BFA" box="[675,820,1497,1521]" italics="true" pageId="110" pageNumber="265">T. kleinmanni</emphasis>
</taxonomicName>
, based on the similarity of plastron pattern between adult
<taxonomicName id="2AB34D03FFC5FFDBFE4DFA68FD93081A" ID-CoL="55JTJ" authorityName=", Sindaco and Jeremcenko" authorityYear="2008" box="[411,556,1529,1553]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="kleinmanni">
<emphasis id="DFC7EA92FFC5FFDBFE4DFA68FD93081A" box="[411,556,1529,1553]" italics="true" pageId="110" pageNumber="265">T. kleinmanni</emphasis>
</taxonomicName>
and juvenile
<emphasis id="DFC7EA92FFC5FFDBFD68FA68FCF1081B" box="[702,846,1529,1552]" italics="true" pageId="110" pageNumber="265">
<taxonomicName id="2AB34D03FFC5FFDBFD68FA68FCF8081B" ID-CoL="55JTW" authorityName="Schoepff" authorityYear="1793" box="[702,839,1529,1552]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="110" pageNumber="265" phylum="Chordata" rank="species" species="marginata">T. marginata</taxonomicName>
.
</emphasis>
</paragraph>
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</treatment>
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