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<document id="44FB3AF5F127BC02514DB95CA72B1C26" ID-DOI="10.25226/bboc.v140i1.2020.a3" ID-ISSN="2513-9894" ID-Zenodo-Dep="13755707" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1718071637822" checkinUser="felipe" docAuthor="Silva, Héctor Gómez de, Pérez Villafaña, Mónica G., Cruz-Nieto, Javier &amp; Cruz-Nieto, Miguel Ángel" docDate="2020" docId="0381A348FFCECB3736CAA8B43932FC1C" docLanguage="en" docName="BullBritOrnitholClub.140.1.7-37.pdf" docOrigin="Bulletin of the British Ornithologists Club 140 (1)" docStyle="DocumentStyle:EFA7F293C51FD4806C5290561C16F223.2:BullBritOrnitholClub.2017-.journal_article" docStyleId="EFA7F293C51FD4806C5290561C16F223" docStyleName="BullBritOrnitholClub.2017-.journal_article" docStyleVersion="2" docTitle="Leptotila verreauxi subsp. capitalis Nelson 1898" docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="FFB8DB30FFDACB21365AAD593D0DFFC5" masterDocTitle="Are some of the birds endemic to the Tres Marías Islands (Mexico) species?" masterLastPageNumber="37" masterPageNumber="7" pageNumber="26" updateTime="1726182511137" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="B822ECEF76AD5092BF0B80EA2B24C296">Are some of the birds endemic to the Tres Marías Islands (Mexico) species?</mods:title>
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<mods:namePart id="E305310ED22F1FBF6095CBA2D70123C1">Silva, Héctor Gómez de</mods:namePart>
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<mods:namePart id="CBA2C1E6FDE761EC4645F7686E06BA55">Pérez Villafaña, Mónica G.</mods:namePart>
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<mods:namePart id="B319AA866CAE92066F50A8A4CA9854C0">Cruz-Nieto, Javier</mods:namePart>
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<mods:namePart id="E296BEAAC970C0A7125C254E83712CD4">Cruz-Nieto, Miguel Ángel</mods:namePart>
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<treatment id="0381A348FFCECB3736CAA8B43932FC1C" LSID="urn:lsid:plazi:treatment:0381A348FFCECB3736CAA8B43932FC1C" httpUri="http://treatment.plazi.org/id/0381A348FFCECB3736CAA8B43932FC1C" lastPageId="22" lastPageNumber="28" pageId="20" pageNumber="26">
<heading id="D0DFA532FFCECB3536CAA8B43E83F9C0" box="[144,910,1517,1541]" fontSize="10" level="2" pageId="20" pageNumber="26" reason="2">
<subSubSection id="C33241D5FFCECB3536CAA8B43FC5F9C0" box="[144,712,1517,1541]" pageId="20" pageNumber="26" type="nomenclature">
<paragraph id="8B97125EFFCECB3536CAA8B43FC5F9C0" blockId="20.[144,1164,1517,1701]" box="[144,712,1517,1541]" pageId="20" pageNumber="26">
<emphasis id="B95CCE4CFFCECB3536CAA8B43CAEF9C0" bold="true" box="[144,419,1517,1541]" pageId="20" pageNumber="26">WHITE-TIPPED DOVE</emphasis>
<taxonomicName id="4C2869DDFFCECB3537F0A8B43FC5F9C0" ID-CoL="5JBN3" authorityName="Nelson" authorityYear="1898" box="[426,712,1517,1541]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="20" pageNumber="26" phylum="Chordata" rank="subSpecies" species="verreauxi" subSpecies="capitalis">
<emphasis id="B95CCE4CFFCECB3537F0A8B43FC5F9C0" box="[426,712,1517,1541]" italics="true" pageId="20" pageNumber="26">Leptotila verreauxi capitalis</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33241D5FFCECB373494A8B43932FC1C" lastPageId="22" lastPageNumber="28" pageId="20" pageNumber="26" type="description">
<paragraph id="8B97125EFFCECB353494A8B43E83F9C0" blockId="20.[144,1164,1517,1701]" box="[718,910,1517,1541]" pageId="20" pageNumber="26">
(vs.
<taxonomicName id="4C2869DDFFCECB3534A7A8B73E8BF9C0" box="[765,902,1517,1541]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="20" pageNumber="26" phylum="Chordata" rank="variety" species="verreauxi" variety="angelica">
<emphasis id="B95CCE4CFFCECB3534A7A8B73E8BF9C0" box="[765,902,1517,1541]" italics="true" pageId="20" pageNumber="26">L. v. angelica</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="8B97125EFFCECB3736CAAB543ED3FEFC" blockId="20.[144,1164,1517,1701]" lastBlockId="22.[144,1165,129,985]" lastPageId="22" lastPageNumber="28" pageId="20" pageNumber="26">
<emphasis id="B95CCE4CFFCECB3536CAAB543DF1F9E0" box="[144,252,1549,1573]" italics="true" pageId="20" pageNumber="26">Coloration</emphasis>
.—Breast colour paler and less red in all specimens (
<bibRefCitation id="EFB96FAFFFCECB35350BAB543ED6F9E0" author="Grant, P. R." box="[849,987,1549,1573]" pageId="20" pageNumber="26" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grant 1965a</bibRefCitation>
,
<specimenCount id="9D2ED9D7FFCECB3535BFAB543931F9E0" box="[997,1084,1549,1573]" count="25" pageId="20" pageNumber="26" type="male">25 male</specimenCount>
and
<specimenCount id="9D2ED9D7FFCECB353229AB543DD6F980" count="21" pageId="20" pageNumber="26" type="female">21 female</specimenCount>
<emphasis id="B95CCE4CFFCECB3536BBAB743C3BF980" box="[225,310,1581,1605]" italics="true" pageId="20" pageNumber="26">capitalis</emphasis>
vs.
<specimenCount id="9D2ED9D7FFCECB353739AB743CB7F980" box="[355,442,1581,1605]" count="32" pageId="20" pageNumber="26" type="male">32 male</specimenCount>
and
<specimenCount id="9D2ED9D7FFCECB3537A8AB743F50F980" box="[498,605,1581,1605]" count="12" pageId="20" pageNumber="26" type="female">12 female</specimenCount>
<emphasis id="B95CCE4CFFCECB353439AB743FBBF980" box="[611,694,1581,1605]" italics="true" pageId="20" pageNumber="26">angelica</emphasis>
), although the single darkest
<emphasis id="B95CCE4CFFCECB35325FAB743957F980" box="[1029,1114,1581,1605]" italics="true" pageId="20" pageNumber="26">capitalis</emphasis>
was almost indistinguishable from the single palest
<emphasis id="B95CCE4CFFCECB3534FCAB143FF4F9A0" box="[678,761,1613,1637]" italics="true" pageId="20" pageNumber="26">angelica</emphasis>
. Also, 50% of island specimens had more extensive white throats than in all mainland birds, 75% of island birds had fewer and paler brown feathers on the thighs and flanks than 100% of mainland birds, and in
<emphasis id="B95CCE4CFFCECB353217ABD7395AF960" box="[1101,1111,1678,1701]" italics="true" pageId="20" pageNumber="26">c</emphasis>
.75% of island birds the vinous breast colour extended less far onto the belly than approximately 75% of mainland birds; hence island birds appear to have a larger, white abdomen (
<bibRefCitation id="EFB96FAFFFCFCB343210AB1F3DDEF9B8" author="Grant, P. R." pageId="21" pageNumber="27" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grant 1965a</bibRefCitation>
). The face is whiter due to the ear-coverts being white or whitish vs. usually pale pinkish grey, and because the white forehead reaches further posteriorly and contrasts sharply with the hindcrown / nape / postocular area (
<figureCitation id="13130EDBFFCCCB373553ADD83E54FF5C" box="[777,857,129,153]" captionStart="Figure 17" captionStartId="21.[144,205,1446,1465]" captionTargetBox="[144,1164,129,1430]" captionTargetId="figure-130@21.[144,1164,694,1433]" captionTargetPageId="21" captionText="Figure 17. Above: White-tipped Dove Leptotila verreauxi angelica, Cruz de Huanacaxtle, Nayarit, Mexico, April 2019 (Marie ONeill), below: L. v. capitalis, Isla María Cleofas, May 2016; note the contrasting white forecrown and cheeks, extensively whitish underparts, and darker brown back, wings and tail (Mónica G. Pérez Villafaña)" figureDoi="http://doi.org/10.5281/zenodo.13755735" httpUri="https://zenodo.org/record/13755735/files/figure.png" pageId="22" pageNumber="28">Fig. 17</figureCitation>
;
<bibRefCitation id="EFB96FAFFFCCCB373532ADD83EF4FF5C" author="Nelson, E. W." box="[872,1017,129,153]" pageId="22" pageNumber="28" pagination="5 - 11" refId="ref16130" refString="Nelson, E. W. 1898. Descriptions of new birds from the Tres Marias Islands, western Mexico. Proc. Biol. Soc. Wash. 72: 5 - 11." type="journal article" year="1898">Nelson 1898</bibRefCitation>
). The whiter face recalls
<collectingCountry id="F33F52CEFFCCCB373748ADF83C7EFF7C" box="[274,371,161,185]" name="Grenada" pageId="22" pageNumber="28">Grenada</collectingCountry>
Dove
<taxonomicName id="4C2869DDFFCCCB3737E7ADFB3F19FF7C" baseAuthorityName="Lawrence" baseAuthorityYear="1884" box="[445,532,161,185]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="species" species="wellsi">
<emphasis id="B95CCE4CFFCCCB3737E7ADFB3F19FF7C" box="[445,532,161,185]" italics="true" pageId="22" pageNumber="28">L. wellsi</emphasis>
</taxonomicName>
and both that and the more extensive white underparts resemble Caribbean Dove
<taxonomicName id="4C2869DDFFCCCB37379CAD9B3F56FF1C" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[454,603,194,217]" class="Aves" family="Accipitridae" genus="Buteo" kingdom="Animalia" order="Accipitriformes" pageId="22" pageNumber="28" phylum="Chordata" rank="species" species="jamaicensis">
<emphasis id="B95CCE4CFFCCCB37379CAD9B3F56FF1C" box="[454,603,194,217]" italics="true" pageId="22" pageNumber="28">L. jamaicensis</emphasis>
</taxonomicName>
. However, unlike those species, the ear-coverts and forehead / forecrown contrast with a pink rather than blue-grey hindcrown / nape / postocular area. Half of
<bibRefCitation id="EFB96FAFFFCCCB3737FEAC583F42FEDC" author="Grant, P. R." box="[420,591,257,281]" pageId="22" pageNumber="28" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grants (1965a)</bibRefCitation>
<emphasis id="B95CCE4CFFCCCB373402AC583FA0FEDC" box="[600,685,257,281]" italics="true" pageId="22" pageNumber="28">capitalis</emphasis>
sample was darker dorsally than 100% of his
<emphasis id="B95CCE4CFFCCCB3736EDAC783C07FEFC" box="[183,266,289,313]" italics="true" pageId="22" pageNumber="28">angelica</emphasis>
sample, consistent with
<bibRefCitation id="EFB96FAFFFCCCB37347BAC783FEFFEFC" author="Ridgway, R." box="[545,738,289,313]" pageId="22" pageNumber="28" pagination="1 - 543" refId="ref16832" refString="Ridgway, R. 1916. The birds of North and Middle America. Bull. US Natl. Mus. 50 (7): 1 - 543." type="journal article" year="1916">Ridgways (1916)</bibRefCitation>
diagnosis and
<figureCitation id="13130EDBFFCCCB3735D7AC783ED7FEFC" box="[909,986,289,313]" captionStart="Figure 17" captionStartId="21.[144,205,1446,1465]" captionTargetBox="[144,1164,129,1430]" captionTargetId="figure-130@21.[144,1164,694,1433]" captionTargetPageId="21" captionText="Figure 17. Above: White-tipped Dove Leptotila verreauxi angelica, Cruz de Huanacaxtle, Nayarit, Mexico, April 2019 (Marie ONeill), below: L. v. capitalis, Isla María Cleofas, May 2016; note the contrasting white forecrown and cheeks, extensively whitish underparts, and darker brown back, wings and tail (Mónica G. Pérez Villafaña)" figureDoi="http://doi.org/10.5281/zenodo.13755735" httpUri="https://zenodo.org/record/13755735/files/figure.png" pageId="22" pageNumber="28">Fig. 17</figureCitation>
.
</paragraph>
<caption id="DF5742D6FFCFCB3436CAA8FF3C2BF9C1" ID-DOI="http://doi.org/10.5281/zenodo.13755735" ID-Zenodo-Dep="13755735" httpUri="https://zenodo.org/record/13755735/files/figure.png" pageId="21" pageNumber="27" startId="21.[144,205,1446,1465]" targetBox="[144,1164,129,1430]" targetPageId="21" targetType="figure">
<paragraph id="8B97125EFFCFCB3436CAA8FF3C2BF9C1" blockId="21.[144,1165,1446,1540]" pageId="21" pageNumber="27">
Figure 17. Above: White-tipped Dove
<taxonomicName id="4C2869DDFFCFCB343453A8FF3ED2FA7C" authority=", Cruz de Huanacaxtle" authorityName="Cruz de Huanacaxtle" box="[521,991,1446,1466]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="21" pageNumber="27" phylum="Chordata" rank="subSpecies" species="verreauxi" subSpecies="angelica">
<emphasis id="B95CCE4CFFCFCB343453A8FF3FF3FA7C" box="[521,766,1446,1465]" italics="true" pageId="21" pageNumber="27">Leptotila verreauxi angelica</emphasis>
, Cruz de Huanacaxtle
</taxonomicName>
, Nayarit, Mexico, April 2019 (Marie ONeill), below:
<taxonomicName id="4C2869DDFFCFCB3437B9A8993F56FA17" authorityName="Nelson" authorityYear="1898" box="[483,603,1471,1490]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="21" pageNumber="27" phylum="Chordata" rank="variety" species="verreauxi" variety="capitalis">
<emphasis id="B95CCE4CFFCFCB3437B9A8993F56FA17" box="[483,603,1471,1490]" italics="true" pageId="21" pageNumber="27">L. v. capitalis</emphasis>
</taxonomicName>
, Isla María Cleofas, May 2016; note the contrasting white forecrown and cheeks, extensively whitish underparts, and darker brown back, wings and tail (Mónica G. Pérez Villafaña)
</paragraph>
</caption>
<paragraph id="8B97125EFFCCCB37369AAC183E43FE7D" blockId="22.[144,1165,129,985]" pageId="22" pageNumber="28">
<emphasis id="B95CCE4CFFCCCB37369AAC183C50FE9C" box="[192,349,321,345]" italics="true" pageId="22" pageNumber="28">Morphometrics</emphasis>
.—Wing, tarsus and bill longer (
<bibRefCitation id="EFB96FAFFFCCCB37349DAC183E5FFE9C" author="Grant, P. R." box="[711,850,321,345]" pageId="22" pageNumber="28" pagination="1 - 106" refId="ref14491" refString="Grant, P. R. 1965 a. A systematic study of the terrestrial birds of the Tres Marias Islands, Mexico. Postilla 90: 1 - 106." type="journal article" year="1965">Grant 1965a</bibRefCitation>
; largest effect size is for bill length of females, score 2) whereas the tail of females is shorter, and in both sexes the white on the tail tip is slightly shorter (score 1). The presence vs. absence of sexual dimorphism in tail length is a further difference between
<emphasis id="B95CCE4CFFCCCB37343DACF83FB1FE7C" box="[615,700,417,441]" italics="true" pageId="22" pageNumber="28">capitalis</emphasis>
and
<emphasis id="B95CCE4CFFCCCB3734AFACF83E45FE7C" box="[757,840,417,441]" italics="true" pageId="22" pageNumber="28">angelica</emphasis>
.
</paragraph>
<paragraph id="8B97125EFFCCCB37369AAC9B3932FC1C" blockId="22.[144,1165,129,985]" pageId="22" pageNumber="28">
<emphasis id="B95CCE4CFFCCCB37369AAC9B3CBCFE1C" box="[192,433,449,473]" italics="true" pageId="22" pageNumber="28">Reasons for uncertainty</emphasis>
.—The whiter underparts and face of
<emphasis id="B95CCE4CFFCCCB373514AC983EAEFE1C" box="[846,931,449,473]" italics="true" pageId="22" pageNumber="28">capitalis</emphasis>
resemble
<taxonomicName id="4C2869DDFFCCCB373248AC9B3DD5FE3C" authorityName="Salvin" authorityYear="1895" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="subSpecies" species="verreauxi" subSpecies="decolor">
<emphasis id="B95CCE4CFFCCCB373248AC9B3DD5FE3C" italics="true" pageId="22" pageNumber="28">L. verreauxi decolor</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2869DDFFCCCB3736B3ACBB3C8EFE3C" box="[233,387,481,505]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="variety" variety="decipiens">
<emphasis id="B95CCE4CFFCCCB3736B3ACBB3C8EFE3C" box="[233,387,481,505]" italics="true" pageId="22" pageNumber="28">L. v. decipiens</emphasis>
</taxonomicName>
and some
<taxonomicName id="4C2869DDFFCCCB373453ACBB3FAAFE3C" box="[521,679,482,505]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="variety" variety="verreauxi">
<emphasis id="B95CCE4CFFCCCB373453ACBB3FAAFE3C" box="[521,679,482,505]" italics="true" pageId="22" pageNumber="28">L. v. verreauxi</emphasis>
</taxonomicName>
(e.g. hbw.com/ibc/1016351 and hbw.com/ ibc/996812 from
<collectingCountry id="F33F52CEFFCCCB37371DAF583CB2FDDC" box="[327,447,513,537]" name="Costa Rica" pageId="22" pageNumber="28">Costa Rica</collectingCountry>
, hbw.com/ibc/980837 from
<collectingCountry id="F33F52CEFFCCCB3734AAAF583E50FDDC" box="[752,861,513,537]" name="Colombia" pageId="22" pageNumber="28">Colombia</collectingCountry>
and hbw.com/ibc/1086049, hbw.com/ibc/1002421 but not darker hbw.com/ibc/1002417 from the Lesser Antilles). Occasional White-tipped Doves from scatered localities in mainland
<collectingCountry id="F33F52CEFFCCCB373593AF183916FD9C" box="[969,1051,577,601]" name="Mexico" pageId="22" pageNumber="28">Mexico</collectingCountry>
resemble
<emphasis id="B95CCE4CFFCCCB3736CAAF383DE5FDBC" box="[144,232,609,633]" italics="true" pageId="22" pageNumber="28">capitalis</emphasis>
(e.g. htps://macaulaylibrary.org/asset/55312471, htps://macaulaylibrary.org/ asset/57930541, htps://macaulaylibrary.org/asset/57744481, and htps://macaulaylibrary. org/asset/39082171) at least in some features (e.g., htps://macaulaylibrary.org/asset/25545561 with whitish ear-coverts but forehead / forecrown showing litle contrast; or htps:// macaulaylibrary.org/asset/43367021 with forecrown less extensive and pinker breast, or htps://macaulaylibrary.org/asset/32166601 with very whitish flanks but strongly pinkish breast, malar region and ear-coverts). We hypothesise that the mutation(s) responsible for a whitish face and underparts contrasting with the mid-crown / nape / postocular area have arisen independently several times in
<taxonomicName id="4C2869DDFFCCCB373460AE383F99FCBC" authorityName="Swainson" authorityYear="1837" box="[570,660,865,889]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="genus">
<emphasis id="B95CCE4CFFCCCB373460AE383F99FCBC" box="[570,660,865,889]" italics="true" pageId="22" pageNumber="28">Leptotila</emphasis>
</taxonomicName>
, becoming fixed in
<emphasis id="B95CCE4CFFCCCB37352BAE383ECBFCBC" box="[881,966,865,889]" italics="true" pageId="22" pageNumber="28">capitalis</emphasis>
,
<emphasis id="B95CCE4CFFCCCB373589AE383916FCBC" box="[979,1051,865,889]" italics="true" pageId="22" pageNumber="28">decolor</emphasis>
,
<emphasis id="B95CCE4CFFCCCB373272AE383988FCBC" box="[1064,1157,865,889]" italics="true" pageId="22" pageNumber="28">decipiens</emphasis>
, Caribbean and
<collectingCountry id="F33F52CEFFCCCB373718AED83CAEFC5C" box="[322,419,897,921]" name="Grenada" pageId="22" pageNumber="28">Grenada</collectingCountry>
Doves, but not in
<taxonomicName id="4C2869DDFFCCCB3734D9AEDB3E12FC5D" box="[643,799,897,921]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="variety" variety="verreauxi">
<emphasis id="B95CCE4CFFCCCB3734D9AEDB3E12FC5D" box="[643,799,897,921]" italics="true" pageId="22" pageNumber="28">L. v. verreauxi</emphasis>
</taxonomicName>
or other races, similar to other known cases of parallel speciation (e.g.
<bibRefCitation id="EFB96FAFFFCCCB373402AEF83E2AFC7C" author="Cooper, E. A. &amp; Uy, J. A. C." box="[600,807,929,953]" pageId="22" pageNumber="28" pagination="3760 - 3774" refId="ref13906" refString="Cooper, E. A. &amp; Uy, J. A. C. 2017. Genomic evidence for convergent evolution of a key trait underlying divergence in island birds. Mol. Ecol. 26: 3760 - 3774." type="journal article" year="2017">Cooper &amp; Uy 2017</bibRefCitation>
and references therein). Clearly more research is needed to understand colour variation in
<taxonomicName id="4C2869DDFFCCCB37357EAE9B3EACFC1D" authorityName="Bonaparte" authorityYear="1855" box="[804,929,961,985]" class="Aves" family="Columbidae" genus="Leptotila" kingdom="Animalia" order="Columbiformes" pageId="22" pageNumber="28" phylum="Chordata" rank="species" species="verreauxi">
<emphasis id="B95CCE4CFFCCCB37357EAE9B3EACFC1D" box="[804,929,961,985]" italics="true" pageId="22" pageNumber="28">L. verreauxi</emphasis>
</taxonomicName>
and relatives.
</paragraph>
</subSubSection>
</heading>
</treatment>
</document>