treatments-xml/data/B7/11/B2/B711B23FFF0B86CCDA3FC681FDBD7C80.xml
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<document id="74A75A22C412F66414245FBF0A9B4204" ID-CLB-Dataset="56026" ID-DOI="10.11646/megataxa.3.1.1" ID-GBIF-Dataset="1ece557d-068b-465b-809e-0fc1110035b3" ID-ISSN="2703-3090" ID-Zenodo-Dep="6422776" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614076789781" checkinUser="plazi" docAuthor="Nakayama, Naohide" docDate="2020" docId="B711B23FFF0B86CCDA3FC681FDBD7C80" docLanguage="en" docName="megataxa.3.1.1.pdf" docOrigin="Megataxa 3 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Hymenogadus Gilbert &amp; Hubbs 1920" docType="treatment" docVersion="7" lastPageNumber="223" masterDocId="4B28CA47FFD68612D915C228FFE77D35" masterDocTitle="Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph" masterLastPageNumber="383" masterPageNumber="1" pageNumber="222" updateTime="1698930971026" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="46AE19297BEC63DF0F477A20B6AC6655">Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph</mods:title>
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<mods:namePart id="9255CF7A178695E0A8D77E39ACB4025F">Nakayama, Naohide</mods:namePart>
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<mods:affiliation id="5B431820A59A838EBE295BF5F77CFC98">Department of Marine Biology, School of Marine Science and Technology, Tokai University, 3 - 20 - 1 Orido, Shimizu, Shizuoka 424 - 8610, Japan ® naonakayama @ tsc. u-tokai. ac. jp or ®</mods:affiliation>
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<treatment id="B711B23FFF0B86CCDA3FC681FDBD7C80" ID-DOI="http://doi.org/10.5281/zenodo.6456244" ID-GBIF-Taxon="194588094" ID-Zenodo-Dep="6456244" LSID="urn:lsid:plazi:treatment:B711B23FFF0B86CCDA3FC681FDBD7C80" httpUri="http://treatment.plazi.org/id/B711B23FFF0B86CCDA3FC681FDBD7C80" lastPageId="222" lastPageNumber="223" pageId="221" pageNumber="222">
<subSubSection id="77A250A2FF0B86CFDA3FC681FAB179F6" box="[810,1366,1193,1220]" pageId="221" pageNumber="222" type="nomenclature">
<paragraph id="3F070329FF0B86CFDA3FC681FAB179F6" blockId="221.[810,1366,1193,1254]" box="[810,1366,1193,1220]" pageId="221" pageNumber="222">
<emphasis id="0DCCDF3BFF0B86CFDA3FC681FAB179F6" bold="true" box="[810,1366,1193,1220]" pageId="221" pageNumber="222">
Genus
<taxonomicName id="F8B878AAFF0B86CFDA94C681FAB179F6" ID-CoL="527N" ID-ENA="1508084" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[897,1366,1193,1220]" class="Actinopterygii" family="Macrouridae" genus="Hymenogadus" kingdom="Animalia" order="Gadiformes" pageId="221" pageNumber="222" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0B86CFDA94C681FBC879F1" bold="true" box="[897,1071,1193,1220]" italics="true" pageId="221" pageNumber="222">Hymenogadus</emphasis>
<bibRefCitation id="5B297ED8FF0B86CFDD22C681FAB179F6" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[1079,1366,1193,1219]" pageId="221" pageNumber="222" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF0B86CFDA3FC6E4FB5979D3" box="[810,1214,1227,1254]" pageId="221" pageNumber="222" type="vernacular_names">
<paragraph id="3F070329FF0B86CFDA3FC6E4FB5979D3" blockId="221.[810,1366,1193,1254]" box="[810,1214,1227,1254]" pageId="221" pageNumber="222">[Japanese name: Oni-sujidara-zoku]</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF0B86CFDA3FC73BFAD27858" pageId="221" pageNumber="222" type="reference_group">
<paragraph id="3F070329FF0B86CFDA3FC73BFAD27858" blockId="221.[810,1451,1299,1389]" pageId="221" pageNumber="222">
<taxonomicName id="F8B878AAFF0B86CFDA3FC73BFB09781C" ID-CoL="527N" authority="Gilbert &amp; Hubbs, 1920: 521" authorityName="Gilbert &amp; Hubbs" authorityPageNumber="521" authorityYear="1920" box="[810,1262,1299,1321]" class="Actinopterygii" family="Macrouridae" genus="Hymenogadus" kingdom="Animalia" order="Gadiformes" pageId="221" pageNumber="222" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0B86CFDA3FC73BFC54781D" box="[810,947,1299,1320]" italics="true" pageId="221" pageNumber="222">Hymenogadus</emphasis>
<bibRefCitation id="5B297ED8FF0B86CFDAD1C73BFB09781C" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[964,1262,1299,1321]" pageId="221" pageNumber="222" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920:521</bibRefCitation>
</taxonomicName>
(as subgenus of
<taxonomicName id="F8B878AAFF0B86CFDA4AC71DFBE3787F" ID-CoL="526Y" authorityName="Giglioli" authorityYear="1884" box="[863,1028,1333,1354]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="221" pageNumber="222" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0B86CFDA4AC71DFBE3787F" box="[863,1028,1333,1354]" italics="true" pageId="221" pageNumber="222">Hymenocephalus</emphasis>
</taxonomicName>
;
<typeStatus id="E003BD8BFF0B86CFDD0FC71DFBA4787F" box="[1050,1091,1333,1354]" pageId="221" pageNumber="222">type</typeStatus>
species:
<taxonomicName id="F8B878AAFF0B86CFDDBBC71DFBD97859" ID-CoL="3NJ8V" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="221" pageNumber="222" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="0DCCDF3BFF0B86CFDDBBC71DFA4C787F" box="[1198,1451,1333,1354]" italics="true" pageId="221" pageNumber="222">Hymenocephalus gracilis</emphasis>
<bibRefCitation id="5B297ED8FF0B86CFDA4AC77FFBD97859" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[863,1086,1367,1389]" pageId="221" pageNumber="222" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
</taxonomicName>
, by original designation).
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF0B86CFDA3FC7BFFBC77523" pageId="221" pageNumber="222" type="diagnosis">
<paragraph id="3F070329FF0B86CFDA3FC7BFFBC77523" blockId="221.[810,1452,1431,2070]" pageId="221" pageNumber="222">
<emphasis id="0DCCDF3BFF0B86CFDA3FC7BFFC437884" bold="true" box="[810,932,1431,1457]" pageId="221" pageNumber="222">Diagnosis.</emphasis>
Anus immediately anterior to anal-fin origin, surrounded by narrow periproct. Ventral striae, consisting of fine, parallel black lines over silvery layer on each side of isthmus, shoulder girdle, most of chest, and posterior to pelvic fins. Gular region covered with fine striations of black epithelium. Light organ developed as long tube, extending from anus to slightly anterior to pelvicfin bases; small lens at each end of light organ. Second spinous ray of first dorsal fin serrated along its leading edge. Pelvic-fin rays 8, outer ray slender, not expanded or ribbon-like distally. Teeth all small, in narrow villiform bands in both jaws. Trunk shallow, cylindrical in cross section. Head bones thin, fragile; head skin membranous, translucent. Nasal bones weakly ossified, not forming stout, dorsoventrally flattened processes. Chin barbel present. Branchiostegal rays 7. Species small, less than
<quantity id="F840AECCFF0B86CFDA3FC590FC8A7AE7" box="[810,877,1976,2002]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="221" pageNumber="222" unit="cm" value="13.0">13 cm</quantity>
TL. Body with silvery reflection when fresh. [From
<bibRefCitation id="5B297ED8FF0B86CFDA3FC5F2FBC77AC1" author="Iwamoto, T. &amp; Shao, K. - T. &amp; Ho, H. - C." box="[810,1056,2010,2036]" pageId="221" pageNumber="222" pagination="513 - 530" refId="ref289069" refString="Iwamoto, T., Shao, K. - T. &amp; Ho, H. - C. (2011) Elevation of Spicomacrurus (Gadiformes: Macrouridae) to generic status, with descriptions of two new species from the southwestern Pacific. Bulletin of Marine Science, 87, 513 - 530. http: // dx. doi. org / 10.5343 / bms. 2011.1004" type="journal article" year="2011">
Iwamoto
<emphasis id="0DCCDF3BFF0B86CFDA81C5F2FC2A7AC1" box="[916,973,2010,2036]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2011)
</bibRefCitation>
,
<bibRefCitation id="5B297ED8FF0B86CFDD3EC5F2FA907AC1" author="McMillan, P. &amp; Iwamoto, T." box="[1067,1399,2010,2036]" pageId="221" pageNumber="222" pagination="117 - 134" refId="ref294107" refString="McMillan, P. &amp; Iwamoto, T. (2014) Descriptions of four species of grenadier fishes of the genera Hymenocephalus and Hymenogadus (Teleostei, Gadiformes, Macrouridae) from the New Zealand region and Tasman Sea, including two new species of Hymenocephalus. Zootaxa, 3856, 117 - 134. http: // dx. doi. org / 10.11646 / zootaxa. 3856.1.5" type="journal article" year="2014">McMillan &amp; Iwamoto (2014)</bibRefCitation>
, and
<bibRefCitation id="5B297ED8FF0B86CFDA3FC5D4FBF37523" author="Schwarzhans, W." box="[810,1044,2043,2070]" pageId="221" pageNumber="222" pagination="1 - 73" refId="ref299271" refString="Schwarzhans, W. (2014) Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species. Zootaxa, 3888, 1 - 73. http: // dx. doi. org / 10.11646 / zootaxa. 3888.1.1" type="journal article" year="2014">Schwarzhans (2014)</bibRefCitation>
.]
</paragraph>
</subSubSection>
<subSubSection id="77A250A2FF0886CCD9A8C285FDBD7C80" pageId="222" pageNumber="223" type="discussion">
<paragraph id="3F070329FF0886CCD9A8C285FDBD7C80" blockId="222.[136,778,173,437]" pageId="222" pageNumber="223">
<emphasis id="0DCCDF3BFF0886CCD9A8C285FED67DF2" bold="true" box="[189,305,173,199]" pageId="222" pageNumber="223">Remarks.</emphasis>
<taxonomicName id="F8B878AAFF0886CCD82FC285FE3D7DF2" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[314,474,173,199]" class="Actinopterygii" family="Macrouridae" genus="Hymenogadus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0886CCD82FC285FE3D7DF2" box="[314,474,173,199]" italics="true" pageId="222" pageNumber="223">Hymenogadus</emphasis>
</taxonomicName>
was originally established as a subgenus of
<taxonomicName id="F8B878AAFF0886CCD84DC2E7FD2E7DDC" authority="Giglioli, 1884" authorityName="Giglioli" authorityYear="1884" box="[344,713,207,233]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0886CCD84DC2E7FDFE7DDC" box="[344,537,207,233]" italics="true" pageId="222" pageNumber="223">Hymenocephalus</emphasis>
<bibRefCitation id="5B297ED8FF0886CCDB31C2E7FD2E7DDC" author="Giglioli, E. H." box="[548,713,207,233]" pageId="222" pageNumber="223" pagination="198 - 270" refId="ref285386" refString="Giglioli, E. H. (1884) Esplorazione talassografica del Mediterraneo. In: Giglioli, E. H. &amp; Issel, A. (Eds.), Plagos, Saggio sulla Vita e sui Prodrotti del Mare. Instituto de' Sordo-muti, Genova, pp. 198 - 270. http: // dx. doi. org / 10.5962 / bhl. title. 42527" type="book chapter" year="1884">Giglioli, 1884</bibRefCitation>
</taxonomicName>
, with the type species
<taxonomicName id="F8B878AAFF0886CCD847C2DAFCE17C3E" authority="(Gilbert &amp; Hubbs, 1920)" baseAuthorityName="Gilbert &amp; Hubbs" baseAuthorityYear="1920" box="[338,774,241,267]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="0DCCDF3BFF0886CCD847C2DAFE377C3E" box="[338,464,241,267]" italics="true" pageId="222" pageNumber="223">H. gracilis</emphasis>
(
<bibRefCitation id="5B297ED8FF0886CCD8F1C2D9FD1A7C3E" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[484,765,241,267]" pageId="222" pageNumber="223" pagination="369 - 588" refId="ref286072" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (vol. 1, pt. 7), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert &amp; Hubbs, 1920</bibRefCitation>
)
</taxonomicName>
. Subsequently,
<bibRefCitation id="5B297ED8FF0886CCD824C33BFE1A7C18" author="Okamura, O." box="[305,509,275,301]" pageId="222" pageNumber="223" refId="ref297275" refString="Okamura, O. (1970 a) Fauna Japonica, Macrourina (Pisces). Academic Press of Japan, Tokyo, 216 pp., 64 pls." type="book" year="1970">Okamura (1970a)</bibRefCitation>
elevated this taxon to full generic rank, with description of another subgenus
<taxonomicName id="F8B878AAFF0886CCD99DC37FFED57C44" authorityName="Okamura" authorityYear="1970" box="[136,306,343,369]" class="Actinopterygii" family="Macrouridae" genus="Spicomacrurus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0886CCD99DC37FFED57C44" box="[136,306,343,369]" italics="true" pageId="222" pageNumber="223">Spicomacrurus</emphasis>
</taxonomicName>
. In addition to
<taxonomicName id="F8B878AAFF0886CCD8F7C370FDBC7C44" baseAuthorityName="Gilbert &amp; Hubbs" baseAuthorityYear="1920" box="[482,603,343,369]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="0DCCDF3BFF0886CCD8F7C370FDBC7C44" box="[482,603,343,369]" italics="true" pageId="222" pageNumber="223">H. gracilis</emphasis>
</taxonomicName>
,
<taxonomicName id="F8B878AAFF0886CCDB7FC37FFCED7C44" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[618,778,343,369]" class="Actinopterygii" family="Macrouridae" genus="Hymenogadus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="0DCCDF3BFF0886CCDB7FC37FFCED7C44" box="[618,778,343,369]" italics="true" pageId="222" pageNumber="223">Hymenogadus</emphasis>
</taxonomicName>
includes
<taxonomicName id="F8B878AAFF0886CCD9FBC352FD9D7CA6" authority="(Gilbert &amp; Hubbs, 1917)" baseAuthorityName="Gilbert &amp; Hubbs" baseAuthorityYear="1917" box="[238,634,377,403]" class="Actinopterygii" family="Macrouridae" genus="Hymenogadus" kingdom="Animalia" order="Gadiformes" pageId="222" pageNumber="223" phylum="Chordata" rank="species" species="tenuis">
<emphasis id="0DCCDF3BFF0886CCD9FBC352FEB17CA6" box="[238,342,378,403]" italics="true" pageId="222" pageNumber="223">H. tenuis</emphasis>
(
<bibRefCitation id="5B297ED8FF0886CCD873C351FD967CA6" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[358,625,377,403]" pageId="222" pageNumber="223" pagination="173 - 175" refId="ref286009" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1917) Description of Hymenocephalus tenuis, a new macruroid [sic] fish from the Hawaiian Islands. Proceedings of the United States National Museum, 54, 173 - 175. http: // dx. doi. org / 10.5479 / si. 00963801.54 - 2231.173" type="journal article" year="1917">Gilbert &amp; Hubbs, 1917</bibRefCitation>
)
</taxonomicName>
known only from the
<typeStatus id="E003BD8BFF0886CCD9FAC3B3FEB67C80" box="[239,337,411,437]" pageId="222" pageNumber="223" type="holotype">holotype</typeStatus>
collected from
<collectingRegion id="FD7CCDCBFF0886CCDB14C3B3FDB27C80" box="[513,597,411,437]" country="United States of America" name="Hawaii" pageId="222" pageNumber="223">Hawaii</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
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