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<document id="DC900A7F34FE85C451A38E933913D006" ID-DOI="10.11646/zootaxa.5322.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203657" ID-ZooBank="DD295EE6-AB3F-4377-900E-CC76F1C6BC2D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690885128246" checkinUser="plazi" docAuthor="Klompen, Hans &amp; Gerdeman, Beverly S." docDate="2023" docId="C55587C9FFEFB81D86C9CF46F53CFD1E" docLanguage="en" docName="zootaxa.5322.1.1.pdf" docOrigin="Zootaxa 5322 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5322.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heterozercon calakmulensis Klompen &amp; Gerdeman 2023, new species" docType="treatment" docVersion="6" lastPageNumber="36" masterDocId="396CFFB1FFCEB83E865EC861F07CFFD3" masterDocTitle="Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="34" updateTime="1726239911995" updateUser="tatiana">
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<mods:title id="1F7FAAE18A6BDCCB19B3701630FCB503">Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)</mods:title>
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<mods:namePart id="5ECCFAE60702F1BAAC78687BA38FA11A">Klompen, Hans</mods:namePart>
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<mods:namePart id="132AD27F7C05B0B2DFE73487086D641D">Gerdeman, Beverly S.</mods:namePart>
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<mods:date id="18F380653A55C2B525A0B906EA56BD5F">2023</mods:date>
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<treatment id="C55587C9FFEFB81D86C9CF46F53CFD1E" ID-DOI="http://doi.org/10.5281/zenodo.8203623" ID-Zenodo-Dep="8203623" LSID="urn:lsid:plazi:treatment:C55587C9FFEFB81D86C9CF46F53CFD1E" httpUri="http://treatment.plazi.org/id/C55587C9FFEFB81D86C9CF46F53CFD1E" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="05E66554FFEFB81F86C9CF46F2F9F893" box="[151,645,1830,1857]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="4D4336DFFFEFB81F86C9CF46F2F9F893" blockId="33.[151,645,1830,1893]" box="[151,645,1830,1857]" pageId="33" pageNumber="34">
<heading id="160B81B3FFEFB81F86C9CF46F2F9F893" bold="true" box="[151,645,1830,1857]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
<emphasis id="7F88EACDFFEFB81F86C9CF46F2F9F893" bold="true" box="[151,645,1830,1857]" pageId="33" pageNumber="34">
<taxonomicName id="8AFC4D5CFFEFB81F86C9CF46F192F892" authority="Klompen &amp; Gerdeman, 2023" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[151,494,1830,1857]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="calakmulensis" status="sp. nov.">
<emphasis id="7F88EACDFFEFB81F86C9CF46F192F892" bold="true" box="[151,494,1830,1857]" italics="true" pageId="33" pageNumber="34">Heterozercon calakmulensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64BB57B6FFEFB81F87A8CF46F2F9F893" box="[502,645,1830,1857]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="05E66554FFEFB81D86C9CF2AF53CFD1E" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="description">
<paragraph id="4D4336DFFFEFB81F86C9CF2AF25FF8B6" blockId="33.[151,645,1830,1893]" box="[151,547,1866,1893]" pageId="33" pageNumber="34">
(
<figureCitation id="D5C72A5AFFEFB81F86C1CF2AF15CF8B6" box="[159,288,1866,1893]" captionStart="FIGURES 1215" captionStartId="31.[151,264,1943,1967]" captionTargetBox="[215,1376,182,1917]" captionTargetId="figure-21@31.[212,1376,181,1920]" captionTargetPageId="31" captionText="FIGURES 1215. Heterozerconidae, chelicera males, photograph (top) and composite drawing (bottom): 1213, Afroheterozercon mahsbergi (OSAL 003045); 1415, Heterozercon calakmulensis (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203665" httpUri="https://zenodo.org/record/8203665/files/figure.png" pageId="33" pageNumber="34">Figs 1415</figureCitation>
,
<figureCitation id="D5C72A5AFFEFB81F8772CF2AF108F8B6" box="[300,372,1867,1893]" captionStart="FIGURES 3538" captionStartId="44.[151,264,1908,1932]" captionTargetBox="[217,1372,184,1882]" captionTargetId="figure-25@44.[212,1375,181,1885]" captionTargetPageId="44" captionText="FIGURES 3538. Heterozerconidae, gnathosoma, adults: 35, Maracazercon joliveti, female (paratype, #7471; red arrow: setal base-like structure on hypostome); 36, Maracazercon joliveti, male (paratype; #7475); 37, Heterozercon calakmulensis, female (CNAC 012444); 38, Heterozercon calakmulensis, male (CNAC 012443). Inset: gnathotectum (female and male) and labrum + salivary stylets." figureDoi="http://doi.org/10.5281/zenodo.8203683" httpUri="https://zenodo.org/record/8203683/files/figure.png" pageId="33" pageNumber="34">3738</figureCitation>
,
<figureCitation id="D5C72A5AFFEFB81F87DECF2AF1B4F8B6" box="[384,456,1867,1893]" captionStart="FIGURES 6768" captionStartId="52.[151,264,1957,1981]" captionTargetBox="[264,1325,1167,1929]" captionTargetId="figure-44@52.[258,1328,1161,1934]" captionTargetPageId="52" captionText="FIGURES 6768. Heterozerconidae, Heterozercon calakmulensis n. sp., idiosoma venter: 67, female (CNAC 012444); 68, male (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203717" httpUri="https://zenodo.org/record/8203717/files/figure.png" pageId="33" pageNumber="34">6768</figureCitation>
,
<figureCitation id="D5C72A5AFFEFB81F878ACF2AF18EF8B6" box="[468,498,1867,1893]" captionStart="FIGURES 8286" captionStartId="56.[151,264,1943,1967]" captionTargetBox="[209,1379,184,1916]" captionTargetId="figure-25@56.[206,1382,181,1919]" captionTargetPageId="56" captionText="FIGURES 8286. Heterozerconidae, leg I, male. 82, Afroheterozercon mahsbergi (OSAL 003042, published); 83, Maracazercon joliveti (paratype); 84, Heterozercon calakmulensis n. sp. (CNAC 012443); 85, Allozercon leytensis n. sp. (OSAL 053185); 86, Al. (Philippinozercon) makilingensis (OSAL 0053267, redrawn from Gerdeman et al. (2018))." figureDoi="http://doi.org/10.5281/zenodo.8203733" httpUri="https://zenodo.org/record/8203733/files/figure.png" pageId="33" pageNumber="34">84</figureCitation>
,
<figureCitation id="D5C72A5AFFEFB81F87A0CF2AF267F8B6" box="[510,539,1867,1893]" captionStart="FIGURES 9396" captionStartId="58.[151,264,1943,1967]" captionTargetBox="[234,1353,184,1917]" captionTargetId="figure-25@58.[231,1356,181,1920]" captionTargetPageId="58" captionText="FIGURES 9396. Heterozerconidae, leg II, male: 93, Maracazercon joliveti (paratype); 94, Heterozercon calakmulensis n. sp. (CNAC 012443); 95, Allozercon leytensis n. sp. (OSAL 053185); 96, Al. (Philippinozercon) makilingensis (OSAL 053267, redrawn from Gerdeman et al. (2018))." figureDoi="http://doi.org/10.5281/zenodo.8203737" httpUri="https://zenodo.org/record/8203737/files/figure.png" pageId="33" pageNumber="34">94</figureCitation>
)
</paragraph>
<paragraph id="4D4336DFFFEFB81F86C9CFF2F30CF826" blockId="33.[151,1436,1939,2037]" pageId="33" pageNumber="34">
Diagnosis. Spermatodactyl complex, with a rounded protrusion just below the pointed tip; distance between setae
<emphasis id="7F88EACDFFEFB81F86C9CFD6F0B1F802" box="[151,205,1975,2001]" italics="true" pageId="33" pageNumber="34">hyp3</emphasis>
and
<emphasis id="7F88EACDFFEFB81F875ACFD6F167F803" box="[260,283,1975,2000]" italics="true" pageId="33" pageNumber="34">sc</emphasis>
subequal to that between
<emphasis id="7F88EACDFFEFB81F8460CFD6F208F802" box="[574,628,1975,2001]" italics="true" pageId="33" pageNumber="34">hyp3</emphasis>
and
<emphasis id="7F88EACDFFEFB81F84F4CFD6F2AEF802" box="[682,722,1975,2001]" italics="true" pageId="33" pageNumber="34">hyp</emphasis>
2; sternitiventral shield without distinct ridges; setae
<emphasis id="7F88EACDFFEFB81F837DCFD6F53DF802" box="[1315,1345,1975,2001]" italics="true" pageId="33" pageNumber="34">Z5</emphasis>
minute, much smaller than
<emphasis id="7F88EACDFFEFB81F8733CFBAF1F5F826" box="[365,393,2011,2037]" italics="true" pageId="33" pageNumber="34">S5</emphasis>
; ventral and anal shields in male not fused.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699C8F6F1A0FF06" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Description. Female idiosoma length 834, width 698 (N=1); male idiosoma length 712, width 598 (N=1). All measurements in
<tableCitation id="007E0364FFECB81C8705C8DAF1ABFF06" box="[347,471,187,213]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="15.[152,245,151,175]" captionStartId-1="16.[158,251,151,175]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="TABLE 3. Comparative measurements (in µm) of newly described Heterozerconidae: females." captionText-1="TABLE 4. Comparative measurements (in µm) of newly described Heterozerconidae: males." pageId="34" pageNumber="35">Tables 34</tableCitation>
.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699C8BEF287FEB6" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Chelicera (
<figureCitation id="D5C72A5AFFECB81C871FC8BEF1BFFF2A" box="[321,451,222,249]" captionStart="FIGURES 1215" captionStartId="31.[151,264,1943,1967]" captionTargetBox="[215,1376,182,1917]" captionTargetId="figure-21@31.[212,1376,181,1920]" captionTargetPageId="31" captionText="FIGURES 1215. Heterozerconidae, chelicera males, photograph (top) and composite drawing (bottom): 1213, Afroheterozercon mahsbergi (OSAL 003045); 1415, Heterozercon calakmulensis (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203665" httpUri="https://zenodo.org/record/8203665/files/figure.png" pageId="34" pageNumber="35">Figs 1415</figureCitation>
). Basal part of fixed digit in female stout, length width ratio 12. Movable digit in adults with thin, straplike dorsal extensions. Movable digit in female elongate, digit length&gt;6 times basal width. Inside movable digit female with brush-like structure. Excrescences on male chelicera present, interdigital. Spermatodactyl on fixed digit of male chelicera, of the flaccid
<typeStatus id="9247887DFFECB81C849DC92AF28BFEB6" box="[707,759,331,357]" pageId="34" pageNumber="35">type</typeStatus>
.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699C90EF1B5FE02" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Palp. Axial outgrowth of palp trochanter in female and male distinct and membranous. Seta
<emphasis id="7F88EACDFFECB81C8296C90EF48EFE5A" box="[1224,1266,367,393]" italics="true" pageId="34" pageNumber="35">pd1</emphasis>
on femur present. Setae on femur long, thick spines. Seta
<emphasis id="7F88EACDFFECB81C84D2C9F2F2CCFE7E" box="[652,688,403,429]" italics="true" pageId="34" pageNumber="35">al2</emphasis>
and
<emphasis id="7F88EACDFFECB81C84B7C9F2F283FE7E" box="[745,767,403,429]" italics="true" pageId="34" pageNumber="35">pl</emphasis>
on genu present. Number of sensilla on palp tarsus 1011. Formula: 2651410/11.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699C9BAF523FD56" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Gnathosoma (
<figureCitation id="D5C72A5AFFECB81C873BC9BAF196FE26" box="[357,490,475,501]" captionStart="FIGURES 3538" captionStartId="44.[151,264,1908,1932]" captionTargetBox="[217,1372,184,1882]" captionTargetId="figure-25@44.[212,1375,181,1885]" captionTargetPageId="44" captionText="FIGURES 3538. Heterozerconidae, gnathosoma, adults: 35, Maracazercon joliveti, female (paratype, #7471; red arrow: setal base-like structure on hypostome); 36, Maracazercon joliveti, male (paratype; #7475); 37, Heterozercon calakmulensis, female (CNAC 012444); 38, Heterozercon calakmulensis, male (CNAC 012443). Inset: gnathotectum (female and male) and labrum + salivary stylets." figureDoi="http://doi.org/10.5281/zenodo.8203683" httpUri="https://zenodo.org/record/8203683/files/figure.png" pageId="34" pageNumber="35">Figs 3738</figureCitation>
). Gnathotectum of female without prominent points. Anterior margin gnathotectum not serrate. Overall shape of gnathotectum curved. Sexual differentiation in gnathotectum absent. Lateral lips enlarged to form a trough; posterior extension of the trough long, extending distinctly beyond insertions of setae
<emphasis id="7F88EACDFFECB81C8301CA42F5E9FDEE" box="[1375,1429,547,573]" italics="true" pageId="34" pageNumber="35">hyp2</emphasis>
. Distance between setae
<emphasis id="7F88EACDFFECB81C87F9CA26F1A1FDB2" box="[423,477,583,609]" italics="true" pageId="34" pageNumber="35">hyp3</emphasis>
and
<emphasis id="7F88EACDFFECB81C8448CA26F251FDB3" box="[534,557,583,608]" italics="true" pageId="34" pageNumber="35">sc</emphasis>
subequal to that between
<emphasis id="7F88EACDFFECB81C8509CA26F3F1FDB2" box="[855,909,583,609]" italics="true" pageId="34" pageNumber="35">hyp3</emphasis>
and
<emphasis id="7F88EACDFFECB81C8598CA26F380FDB2" box="[966,1020,583,609]" italics="true" pageId="34" pageNumber="35">hyp2</emphasis>
. Setae
<emphasis id="7F88EACDFFECB81C820ECA26F4FBFDB2" box="[1104,1159,583,609]" italics="true" pageId="34" pageNumber="35">hyp2</emphasis>
at least twice as long as
<emphasis id="7F88EACDFFECB81C86C9CA0AF0B1FD56" box="[151,205,619,645]" italics="true" pageId="34" pageNumber="35">hyp3</emphasis>
. Cornicula membranous without distal notch. Additional setal base-like structure on hypostome present.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699CAEEF53EFD22" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Dorsum. Dorsal shield in adults covering most of the dorsum; shield with distinct reticulation. Anterior dorsal margin in adults without elongate setae; setae
<emphasis id="7F88EACDFFECB81C84C6CAD2F2D1FD1E" box="[664,685,691,717]" italics="true" pageId="34" pageNumber="35">j1</emphasis>
absent or not distinct (e.g., minute). Antero-marginal area of dorsal shield in males without distinct spines. Median dorsal setae minute. Posterior dorsal setae (
<emphasis id="7F88EACDFFECB81C82CDCAB6F4A0FD22" box="[1171,1244,727,753]" italics="true" pageId="34" pageNumber="35">Z2Z4</emphasis>
) minute.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699CA9AF2EAFC1A" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Sternal area female (
<figureCitation id="D5C72A5AFFECB81C87F2CA9AF183FCC6" box="[428,511,763,789]" captionStart="FIGURES 6768" captionStartId="52.[151,264,1957,1981]" captionTargetBox="[264,1325,1167,1929]" captionTargetId="figure-44@52.[258,1328,1161,1934]" captionTargetPageId="52" captionText="FIGURES 6768. Heterozerconidae, Heterozercon calakmulensis n. sp., idiosoma venter: 67, female (CNAC 012444); 68, male (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203717" httpUri="https://zenodo.org/record/8203717/files/figure.png" pageId="34" pageNumber="35">Fig. 67</figureCitation>
). Areas near insertion sternal setae
<emphasis id="7F88EACDFFECB81C85DBCA9AF3DAFCC6" box="[901,934,763,789]" italics="true" pageId="34" pageNumber="35">st1</emphasis>
sclerotized, forming isolated platelets. Sclerites of sternal setae
<emphasis id="7F88EACDFFECB81C8731CB7EF1ECFCEA" box="[367,400,799,825]" italics="true" pageId="34" pageNumber="35">st1</emphasis>
and
<emphasis id="7F88EACDFFECB81C879ACB7EF198FCEA" box="[452,484,799,825]" italics="true" pageId="34" pageNumber="35">st2</emphasis>
not fused. Sternal setae
<emphasis id="7F88EACDFFECB81C84AECB7EF36DFCEA" box="[752,785,799,825]" italics="true" pageId="34" pageNumber="35">st1</emphasis>
and lyrifissures
<emphasis id="7F88EACDFFECB81C8599CB7EF395FCEA" box="[967,1001,799,825]" italics="true" pageId="34" pageNumber="35">iv1</emphasis>
inserted on the same shield. Areas near insertion sternal setae
<emphasis id="7F88EACDFFECB81C87CBCB22F1C9FC8E" box="[405,437,835,861]" italics="true" pageId="34" pageNumber="35">st2</emphasis>
sclerotized, forming isolated platelets. Areas near insertion sternal setae
<emphasis id="7F88EACDFFECB81C82ADCB22F568FC8E" box="[1267,1300,835,861]" italics="true" pageId="34" pageNumber="35">st3</emphasis>
sclerotized, fused to genitiventral shield. Metasternal setae
<emphasis id="7F88EACDFFECB81C84F1CB06F2ACFC52" box="[687,720,871,897]" italics="true" pageId="34" pageNumber="35">st4</emphasis>
absent. Areas near insertion sternal setae
<emphasis id="7F88EACDFFECB81C82F0CB06F4B2FC52" box="[1198,1230,871,897]" italics="true" pageId="34" pageNumber="35">st5</emphasis>
sclerotized, fused to genitiventral shield. Sternal lyrifissures
<emphasis id="7F88EACDFFECB81C8423CBEAF2E3FC76" box="[637,671,907,933]" italics="true" pageId="34" pageNumber="35">iv1</emphasis>
,
<emphasis id="7F88EACDFFECB81C84F1CBEAF2ADFC76" box="[687,721,907,933]" italics="true" pageId="34" pageNumber="35">iv2</emphasis>
, and
<emphasis id="7F88EACDFFECB81C854CCBEAF349FC76" box="[786,821,907,933]" italics="true" pageId="34" pageNumber="35">iv3</emphasis>
present. Distinct curved, sclerotized ridge on anterior margin of female genitiventral shield present.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699CBB2F31EFB72" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Sternal area male (
<figureCitation id="D5C72A5AFFECB81C87C6CBB2F190FC3E" box="[408,492,979,1005]" captionStart="FIGURES 6768" captionStartId="52.[151,264,1957,1981]" captionTargetBox="[264,1325,1167,1929]" captionTargetId="figure-44@52.[258,1328,1161,1934]" captionTargetPageId="52" captionText="FIGURES 6768. Heterozerconidae, Heterozercon calakmulensis n. sp., idiosoma venter: 67, female (CNAC 012444); 68, male (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203717" httpUri="https://zenodo.org/record/8203717/files/figure.png" pageId="34" pageNumber="35">Fig. 68</figureCitation>
). Areas near insertion sternal setae
<emphasis id="7F88EACDFFECB81C852BCBB2F3EAFC3E" box="[885,918,979,1005]" italics="true" pageId="34" pageNumber="35">st1</emphasis>
sclerotized, forming isolated platelets. Sternal setae
<emphasis id="7F88EACDFFECB81C8689CB96F084FBC2" box="[215,248,1015,1041]" italics="true" pageId="34" pageNumber="35">st1</emphasis>
and lyrifissures
<emphasis id="7F88EACDFFECB81C87EACB96F1AAFBC2" box="[436,470,1015,1041]" italics="true" pageId="34" pageNumber="35">iv1</emphasis>
inserted on the same shield. Sclerotized areas near insertion sternal setae
<emphasis id="7F88EACDFFECB81C8344CB96F546FBC2" box="[1306,1338,1015,1041]" italics="true" pageId="34" pageNumber="35">st2</emphasis>
forming isolated platelets. Sclerotized areas of sternal setae
<emphasis id="7F88EACDFFECB81C8481CC7AF37CFBE6" box="[735,768,1051,1077]" italics="true" pageId="34" pageNumber="35">st2</emphasis>
and
<emphasis id="7F88EACDFFECB81C8564CC7AF327FBE6" box="[826,859,1051,1077]" italics="true" pageId="34" pageNumber="35">st3</emphasis>
not fused. Sclerotized areas near insertions sternal setae
<emphasis id="7F88EACDFFECB81C8688CC5EF08BFB8A" box="[214,247,1087,1113]" italics="true" pageId="34" pageNumber="35">st3</emphasis>
and
<emphasis id="7F88EACDFFECB81C8772CC5EF130FB8A" box="[300,332,1087,1113]" italics="true" pageId="34" pageNumber="35">st5</emphasis>
fused with sternitiventral shield. Sternal lyrifissures
<emphasis id="7F88EACDFFECB81C85C4CC5EF3C0FB8A" box="[922,956,1087,1113]" italics="true" pageId="34" pageNumber="35">iv1</emphasis>
,
<emphasis id="7F88EACDFFECB81C8597CC5EF390FB8A" box="[969,1004,1087,1113]" italics="true" pageId="34" pageNumber="35">iv2</emphasis>
, and
<emphasis id="7F88EACDFFECB81C8276CC5EF437FB8A" box="[1064,1099,1087,1113]" italics="true" pageId="34" pageNumber="35">iv3</emphasis>
present. Male genital opening presternal. Genital shields not overlapping base of the tritosternum. Sternitiventral area in male relatively smooth, with a few cuticular bumps, otherwise surface faintly reticulate.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699CCCAF547F9DA" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Opisthogaster (
<figureCitation id="D5C72A5AFFECB81C872CCCCAF189FB16" box="[370,501,1195,1221]" captionStart="FIGURES 6768" captionStartId="52.[151,264,1957,1981]" captionTargetBox="[264,1325,1167,1929]" captionTargetId="figure-44@52.[258,1328,1161,1934]" captionTargetPageId="52" captionText="FIGURES 6768. Heterozerconidae, Heterozercon calakmulensis n. sp., idiosoma venter: 67, female (CNAC 012444); 68, male (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203717" httpUri="https://zenodo.org/record/8203717/files/figure.png" pageId="34" pageNumber="35">Figs 6768</figureCitation>
). Metapodal and sternitiventral shields in adults not fused. Posterolateral margin of metapodal shields rounded. Setae
<emphasis id="7F88EACDFFECB81C8448CCAEF23CFB3A" box="[534,576,1231,1257]" italics="true" pageId="34" pageNumber="35">Zv3</emphasis>
absent. Opisthogastral suckers present, posterior to ventral shield, of heterozerconid
<typeStatus id="9247887DFFECB81C86A1CC92F14FFADE" box="[255,307,1267,1293]" pageId="34" pageNumber="35">type</typeStatus>
. Secondary genital openings of female on axial side of opisthogastral suckers, associated with a few small sclerites (
<figureCitation id="D5C72A5AFFECB81C8716CD76F1E0FAE2" box="[328,412,1303,1329]" captionStart="FIGURES 6768" captionStartId="52.[151,264,1957,1981]" captionTargetBox="[264,1325,1167,1929]" captionTargetId="figure-44@52.[258,1328,1161,1934]" captionTargetPageId="52" captionText="FIGURES 6768. Heterozerconidae, Heterozercon calakmulensis n. sp., idiosoma venter: 67, female (CNAC 012444); 68, male (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203717" httpUri="https://zenodo.org/record/8203717/files/figure.png" pageId="34" pageNumber="35">Fig. 67</figureCitation>
, arrow). Apodemes extending from opisthogastral suckers absent. Ventral shield area with a single shield. Sternitiventral and anal shields in male partly fused, not fused in female. Setae
<emphasis id="7F88EACDFFECB81C829BCD5AF491FA86" box="[1221,1261,1339,1365]" italics="true" pageId="34" pageNumber="35">Sv2</emphasis>
not inserted on genitiventral (female) or sternitiventral (male) shield. Setae
<emphasis id="7F88EACDFFECB81C8568CD3EF322FAAA" box="[822,862,1375,1401]" italics="true" pageId="34" pageNumber="35">Sv3</emphasis>
inserted on genitiventral (female) or sternitiventral (male) shield. Setae
<emphasis id="7F88EACDFFECB81C87DECDE2F1D4FA4E" box="[384,424,1411,1437]" italics="true" pageId="34" pageNumber="35">Jv5</emphasis>
inserted anterior to ventrianal line. Insertion paranal setae (
<emphasis id="7F88EACDFFECB81C820FCDE2F411FA4F" box="[1105,1133,1411,1436]" italics="true" pageId="34" pageNumber="35">pa</emphasis>
) at level of anus. Postanal (
<emphasis id="7F88EACDFFECB81C86FFCDC6F0C1FA13" box="[161,189,1447,1472]" italics="true" pageId="34" pageNumber="35">po</emphasis>
) seta similar in length to paranal (
<emphasis id="7F88EACDFFECB81C8467CDC6F229FA13" box="[569,597,1447,1472]" italics="true" pageId="34" pageNumber="35">pa</emphasis>
) setae. Setae
<emphasis id="7F88EACDFFECB81C84B0CDC6F377FA12" box="[750,779,1447,1473]" italics="true" pageId="34" pageNumber="35">Z5</emphasis>
minute; setae
<emphasis id="7F88EACDFFECB81C85F2CDC6F3B4FA12" box="[940,968,1447,1473]" italics="true" pageId="34" pageNumber="35">S5</emphasis>
,
<emphasis id="7F88EACDFFECB81C858BCDC6F389FA12" box="[981,1013,1447,1473]" italics="true" pageId="34" pageNumber="35">R5</emphasis>
and
<emphasis id="7F88EACDFFECB81C8274CDC6F435FA12" box="[1066,1097,1447,1473]" italics="true" pageId="34" pageNumber="35">R4</emphasis>
of medium length. Number of marginal opisthosomal setae small, modified into hook-like spines. Lyrifissures
<emphasis id="7F88EACDFFECB81C824DCDAAF449FA36" box="[1043,1077,1483,1509]" italics="true" pageId="34" pageNumber="35">iv5</emphasis>
inserted near anterior margin of suckers. Postero-marginal shield in both sexes wide, extending lateral well beyond opisthogastral suckers.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699CE72F451F846" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
Legs (
<figureCitation id="D5C72A5AFFECB81C8750CE72F11BF9FE" box="[270,359,1555,1581]" captionStart="FIGURES 8286" captionStartId="56.[151,264,1943,1967]" captionTargetBox="[209,1379,184,1916]" captionTargetId="figure-25@56.[206,1382,181,1919]" captionTargetPageId="56" captionText="FIGURES 8286. Heterozerconidae, leg I, male. 82, Afroheterozercon mahsbergi (OSAL 003042, published); 83, Maracazercon joliveti (paratype); 84, Heterozercon calakmulensis n. sp. (CNAC 012443); 85, Allozercon leytensis n. sp. (OSAL 053185); 86, Al. (Philippinozercon) makilingensis (OSAL 0053267, redrawn from Gerdeman et al. (2018))." figureDoi="http://doi.org/10.5281/zenodo.8203733" httpUri="https://zenodo.org/record/8203733/files/figure.png" pageId="34" pageNumber="35">Figs 84</figureCitation>
,
<figureCitation id="D5C72A5AFFECB81C872ACE72F1E8F9FE" box="[372,404,1555,1581]" captionStart="FIGURES 9396" captionStartId="58.[151,264,1943,1967]" captionTargetBox="[234,1353,184,1917]" captionTargetId="figure-25@58.[231,1356,181,1920]" captionTargetPageId="58" captionText="FIGURES 9396. Heterozerconidae, leg II, male: 93, Maracazercon joliveti (paratype); 94, Heterozercon calakmulensis n. sp. (CNAC 012443); 95, Allozercon leytensis n. sp. (OSAL 053185); 96, Al. (Philippinozercon) makilingensis (OSAL 053267, redrawn from Gerdeman et al. (2018))." figureDoi="http://doi.org/10.5281/zenodo.8203737" httpUri="https://zenodo.org/record/8203737/files/figure.png" pageId="34" pageNumber="35">94</figureCitation>
). Tibiae and tarsi of legs I of similar width as rest of the leg. Acrotarsus on legs I present. Femora I setae
<emphasis id="7F88EACDFFECB81C8716CE56F10FF982" box="[328,371,1591,1617]" italics="true" pageId="34" pageNumber="35">ad3</emphasis>
and
<emphasis id="7F88EACDFFECB81C87F3CE56F1ADF982" box="[429,465,1591,1617]" italics="true" pageId="34" pageNumber="35">pl2</emphasis>
present, setae
<emphasis id="7F88EACDFFECB81C8425CE56F2E9F982" box="[635,661,1591,1617]" italics="true" pageId="34" pageNumber="35">v3</emphasis>
and
<emphasis id="7F88EACDFFECB81C8491CE56F295F982" box="[719,745,1591,1617]" italics="true" pageId="34" pageNumber="35">v4</emphasis>
absent. Genua I setae
<emphasis id="7F88EACDFFECB81C85B1CE56F465F982" box="[1007,1049,1591,1617]" italics="true" pageId="34" pageNumber="35">ad3</emphasis>
and
<emphasis id="7F88EACDFFECB81C820DCE56F400F982" box="[1107,1148,1591,1617]" italics="true" pageId="34" pageNumber="35">pd3</emphasis>
absent. Tibiae I seta
<emphasis id="7F88EACDFFECB81C832CCE56F5E0F982" box="[1394,1436,1591,1617]" italics="true" pageId="34" pageNumber="35">ad2</emphasis>
present, setae
<emphasis id="7F88EACDFFECB81C876BCE3AF11CF9A6" box="[309,352,1627,1653]" italics="true" pageId="34" pageNumber="35">ad3</emphasis>
,
<emphasis id="7F88EACDFFECB81C8730CE3AF1EAF9A6" box="[366,406,1627,1653]" italics="true" pageId="34" pageNumber="35">av2</emphasis>
and
<emphasis id="7F88EACDFFECB81C8793CE3AF189F9A6" box="[461,501,1627,1653]" italics="true" pageId="34" pageNumber="35">pv2</emphasis>
absent. Femora II setae
<emphasis id="7F88EACDFFECB81C8556CE3AF34DF9A6" box="[776,817,1627,1653]" italics="true" pageId="34" pageNumber="35">av2</emphasis>
and
<emphasis id="7F88EACDFFECB81C8536CE3AF3ECF9A6" box="[872,912,1627,1653]" italics="true" pageId="34" pageNumber="35">pv2</emphasis>
absent. Genua IIIV setae
<emphasis id="7F88EACDFFECB81C829CCE3AF490F9A6" box="[1218,1260,1627,1653]" italics="true" pageId="34" pageNumber="35">ad3</emphasis>
and
<emphasis id="7F88EACDFFECB81C837DCE3AF53BF9A6" box="[1315,1351,1627,1653]" italics="true" pageId="34" pageNumber="35">pl2</emphasis>
absent. Genua II seta
<emphasis id="7F88EACDFFECB81C876CCE1EF127F94A" box="[306,347,1663,1689]" italics="true" pageId="34" pageNumber="35">pd3</emphasis>
absent, genua IIIIV seta
<emphasis id="7F88EACDFFECB81C8421CE1EF2D5F94A" box="[639,681,1663,1689]" italics="true" pageId="34" pageNumber="35">pd3</emphasis>
present. Tibiae IIIV setae
<emphasis id="7F88EACDFFECB81C8583CE1EF47BF94A" box="[989,1031,1663,1689]" italics="true" pageId="34" pageNumber="35">ad2</emphasis>
,
<emphasis id="7F88EACDFFECB81C824ACE1EF442F94A" box="[1044,1086,1663,1689]" italics="true" pageId="34" pageNumber="35">pd3</emphasis>
and
<emphasis id="7F88EACDFFECB81C822DCE1EF4E4F94A" box="[1139,1176,1663,1689]" italics="true" pageId="34" pageNumber="35">pl2</emphasis>
absent. Femora III seta
<emphasis id="7F88EACDFFECB81C86C9CEC2F0CEF96E" box="[151,178,1699,1725]" italics="true" pageId="34" pageNumber="35">v3</emphasis>
absent. Tarsi IV setae
<emphasis id="7F88EACDFFECB81C87EBCEC2F1A2F96E" box="[437,478,1699,1725]" italics="true" pageId="34" pageNumber="35">av4</emphasis>
and
<emphasis id="7F88EACDFFECB81C8448CEC2F242F96E" box="[534,574,1699,1725]" italics="true" pageId="34" pageNumber="35">pv4</emphasis>
absent. Complete chaetotaxy in
<tableCitation id="007E0364FFECB81C85EFCEC2F474F96E" box="[945,1032,1699,1725]" captionStart="TABLE 2" captionStartId="14.[152,245,152,176]" captionTargetPageId="14" captionText="TABLE 2. Leg chaetotaxy in adult Heterozerconoidea." pageId="34" pageNumber="35">Table 2</tableCitation>
. Both coxae I setae in male distinct spines with rounded tips. Femora I setae
<emphasis id="7F88EACDFFECB81C843CCEA6F2FBF932" box="[610,647,1735,1761]" italics="true" pageId="34" pageNumber="35">al1</emphasis>
and
<emphasis id="7F88EACDFFECB81C84E3CEA6F29CF932" box="[701,736,1735,1761]" italics="true" pageId="34" pageNumber="35">
al
<quantity id="8A049B3AFFECB81C848DCEA6F29CF932" box="[723,736,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="34" pageNumber="35" unit="in" value="2.0">2</quantity>
</emphasis>
in male spine-like, in anteroventral position; setae
<emphasis id="7F88EACDFFECB81C8340CEA6F53EF932" box="[1310,1346,1735,1761]" italics="true" pageId="34" pageNumber="35">al1</emphasis>
and
<emphasis id="7F88EACDFFECB81C8326CEA6F5E0F932" box="[1400,1436,1735,1761]" italics="true" pageId="34" pageNumber="35">
al
<quantity id="8A049B3AFFECB81C83D1CEA6F5E0F932" box="[1423,1436,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="34" pageNumber="35" unit="in" value="2.0">2</quantity>
</emphasis>
in female sturdy setae, not spines. Femora I seta
<emphasis id="7F88EACDFFECB81C84E0CE8AF2A4F8D7" box="[702,728,1771,1796]" italics="true" pageId="34" pageNumber="35">av</emphasis>
and
<emphasis id="7F88EACDFFECB81C854FCE8AF357F8D7" box="[785,811,1771,1796]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
in male large straight spines; setae
<emphasis id="7F88EACDFFECB81C82E0CE8AF4A4F8D7" box="[1214,1240,1771,1796]" italics="true" pageId="34" pageNumber="35">av</emphasis>
and
<emphasis id="7F88EACDFFECB81C834ECE8AF556F8D7" box="[1296,1322,1771,1796]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
in female spines, but much shorter than in male. Femora II seta
<emphasis id="7F88EACDFFECB81C84A8CF6EF370F8FA" box="[758,780,1807,1833]" italics="true" pageId="34" pageNumber="35">al</emphasis>
in male spine-like and ventral in position, setae
<emphasis id="7F88EACDFFECB81C836ECF6EF536F8FB" box="[1328,1354,1807,1832]" italics="true" pageId="34" pageNumber="35">av</emphasis>
and
<emphasis id="7F88EACDFFECB81C83DCCF6EF5E0F8FB" box="[1410,1436,1807,1832]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
in male spines; setae
<emphasis id="7F88EACDFFECB81C87DBCF52F1E3F89F" box="[389,415,1843,1868]" italics="true" pageId="34" pageNumber="35">av</emphasis>
and
<emphasis id="7F88EACDFFECB81C8788CF52F18CF89F" box="[470,496,1843,1868]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
in female much thinner. Genua II seta
<emphasis id="7F88EACDFFECB81C85FACF52F3C2F89F" box="[932,958,1843,1868]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
and tibiae II seta
<emphasis id="7F88EACDFFECB81C82D9CF52F4DDF89F" box="[1159,1185,1843,1868]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
in male spines, barely half as long as seta
<emphasis id="7F88EACDFFECB81C872DCF36F1F1F8A3" box="[371,397,1879,1904]" italics="true" pageId="34" pageNumber="35">av</emphasis>
; setae
<emphasis id="7F88EACDFFECB81C8785CF36F189F8A3" box="[475,501,1879,1904]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
in female setiform and as long as
<emphasis id="7F88EACDFFECB81C8526CF36F3EEF8A3" box="[888,914,1879,1904]" italics="true" pageId="34" pageNumber="35">av</emphasis>
. Tarsi II seta
<emphasis id="7F88EACDFFECB81C8275CF36F433F8A2" box="[1067,1103,1879,1905]" italics="true" pageId="34" pageNumber="35">pl1</emphasis>
a spine in both sexes, thinner in female. Trochanters III in male with seta
<emphasis id="7F88EACDFFECB81C84DCCF1AF2E4F846" box="[642,664,1915,1941]" italics="true" pageId="34" pageNumber="35">al</emphasis>
spine-like, setae
<emphasis id="7F88EACDFFECB81C8504CF1AF308F847" box="[858,884,1915,1940]" italics="true" pageId="34" pageNumber="35">av</emphasis>
and
<emphasis id="7F88EACDFFECB81C85F5CF1AF3BDF846" box="[939,961,1915,1941]" italics="true" pageId="34" pageNumber="35">pl</emphasis>
setiform.
</paragraph>
<paragraph id="4D4336DFFFECB81C8699CFFEF091F80E" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="35">
<materialsCitation id="FD943C82FFECB81C8699CFFEF3B7F86A" box="[199,971,1951,1977]" collectionCode="CNAC" pageId="34" pageNumber="35" specimenCode="CNAC 012443, CNAC 012444" specimenCount="2" typeStatus="holotype">
Type depository.
<typeStatus id="9247887DFFECB81C87D3CFFEF189F86A" box="[397,501,1951,1977]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
male, accession number
<specimenCode id="1D5A9EA4FFECB81C8547CFFEF3BBF86A" box="[793,967,1951,1977]" collectionCode="CNAC" pageId="34" pageNumber="35">CNAC 012443</specimenCode>
,
</materialsCitation>
<materialsCitation id="FD943C82FFECB81C858ACFFEF091F80E" collectionCode="CNAC" pageId="34" pageNumber="35" specimenCode="CNAC 012443, CNAC 012444" specimenCount="2" typeStatus="paratype">
and female
<typeStatus id="9247887DFFECB81C8202CFFEF4C3F86A" box="[1116,1215,1951,1977]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
,
<specimenCode id="1D5A9EA4FFECB81C8292CFFEF507F86A" box="[1228,1403,1951,1977]" collectionCode="CNAC" pageId="34" pageNumber="35">CNAC 012444</specimenCode>
, at
<collectionCode id="2BEDAE1AFFECB81C86C9CFA2F094F80E" box="[151,232,1987,2013]" pageId="34" pageNumber="35">CNAC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="4D4336DFFFEDB81D8699C8F6F508FF06" blockId="35.[151,1437,150,717]" pageId="35" pageNumber="36">
<materialsCitation id="FD943C82FFEDB81D8699C8F6F322FF06" collectingDate="2005-07-05" collectionCode="CNAC" collectorName="Vazquez, M. M." country="Mexico" elevation="150" latitude="18.9473" location="Calakmul Biosphere Reserve" longLatPrecision="7" longitude="-89.3155" municipality="Material" pageId="35" pageNumber="36" specimenCode="CNAC 012443" specimenCount="1" specimenCount-male="1" stateProvince="Campeche" typeStatus="holotype">
<collectingMunicipality id="AD27ACA5FFEDB81D8699C8F6F15AFF62" box="[199,294,151,177]" pageId="35" pageNumber="36">Material</collectingMunicipality>
examined.
<collectingCountry id="35EB764FFFEDB81D87F6C8F6F27EFF62" box="[424,514,151,177]" name="Mexico" pageId="35" pageNumber="36">Mexico</collectingCountry>
:
<collectingRegion id="8F38F83DFFEDB81D8452C8F6F2F9FF62" box="[524,645,151,177]" country="Mexico" name="Campeche" pageId="35" pageNumber="36">Campeche</collectingRegion>
,
<location id="48236004FFEDB81D84CEC8F6F3A5FF62" LSID="urn:lsid:plazi:treatment:C55587C9FFEFB81D86C9CF46F53CFD1E:48236004FFEDB81D84CEC8F6F3A5FF62" box="[656,985,151,177]" country="Mexico" latitude="18.9473" longLatPrecision="7" longitude="-89.3155" municipality="Material" name="Calakmul Biosphere Reserve" pageId="35" pageNumber="36" stateProvince="Campeche">Calakmul Biosphere Reserve</location>
, Bel-Ha, ~
<quantity id="8A049B3AFFEDB81D820FC8F7F4E8FF62" box="[1105,1172,150,177]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="35" pageNumber="36" unit="m" value="150.0">
<elevation id="C6D1D1ECFFEDB81D820FC8F7F4E8FF62" box="[1105,1172,150,177]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="35" pageNumber="36" unit="m" value="150.0">150m</elevation>
</quantity>
,
<geoCoordinate id="28C85018FFEDB81D82C1C8F6F569FF62" box="[1183,1301,151,177]" degrees="18.9473" direction="north" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="18.9473">18.9473 N</geoCoordinate>
<geoCoordinate id="28C85018FFEDB81D8342C8F7F5E4FF62" box="[1308,1432,150,177]" degrees="89.3155" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="-89.3155">89.3155 W</geoCoordinate>
,
<date id="3942101FFFEDB81D86C9C8DBF164FF06" box="[151,280,186,213]" pageId="35" pageNumber="36" value="2005-07-05">
<collectingDate id="2906E9F7FFEDB81D86C9C8DBF164FF06" box="[151,280,186,213]" pageId="35" pageNumber="36" value="2005-07-05">5Jul-2005</collectingDate>
</date>
,
<collectorName id="E0095309FFEDB81D877DC8DAF1A0FF06" box="[291,476,187,213]" pageId="35" pageNumber="36">Vázquez, M. M.</collectorName>
,
<specimenCount id="5BFAFD56FFEDB81D87B9C8DAF249FF06" box="[487,565,187,213]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
,
<specimenCode id="1D5A9EA4FFEDB81D841FC8DAF291FF06" box="[577,749,187,213]" collectionCode="CNAC" pageId="35" pageNumber="36">CNAC 012443</specimenCode>
,
<typeStatus id="9247887DFFEDB81D84A6C8DAF322FF06" box="[760,862,187,213]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="FD943C82FFEDB81D8536C8DAF50CFF06" box="[872,1392,187,213]" collectingDate="2005-07-05" collectionCode="CNAC" collectorName="Vazquez, M. M." country="Mexico" elevation="150" latitude="18.9473" location="Calakmul Biosphere Reserve" longLatPrecision="7" longitude="-89.3155" municipality="Material" pageId="35" pageNumber="36" specimenCode="CNAC 012444" specimenCount="1" specimenCount-female="1" stateProvince="Campeche" typeStatus="paratype">
same data,
<specimenCount id="5BFAFD56FFEDB81D85B8C8DAF436FF06" box="[998,1098,187,213]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
,
<specimenCode id="1D5A9EA4FFEDB81D820BC8DAF57DFF06" box="[1109,1281,187,213]" collectionCode="CNAC" pageId="35" pageNumber="36">CNAC 012444</specimenCode>
,
<typeStatus id="9247887DFFEDB81D8352C8DAF50CFF06" box="[1292,1392,187,213]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="4D4336DFFFEDB81D8699C8BEF156FECE" blockId="35.[151,1437,150,717]" pageId="35" pageNumber="36">Etymology. The species name is derived from the collecting locality, near the ruins of the ancient Mayan city of Calakmul.</paragraph>
<paragraph id="4D4336DFFFEDB81D8699C946F1BDFDCA" blockId="35.[151,1437,150,717]" pageId="35" pageNumber="36">
Remarks.
<taxonomicName id="8AFC4D5CFFEDB81D8764C946F20BFE92" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[314,631,295,321]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="calakmulensis">
<emphasis id="7F88EACDFFEDB81D8764C946F20BFE92" box="[314,631,295,321]" italics="true" pageId="35" pageNumber="36">Heterozercon calakmulensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="8AFC4D5CFFEDB81D8550C946F3CBFE93" authorityName="Fain" authorityYear="1989" box="[782,951,295,321]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="microsuctus">
<emphasis id="7F88EACDFFEDB81D8550C946F35EFE93" box="[782,802,295,320]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D856FC946F3CBFE93" box="[817,951,295,320]" italics="true" pageId="35" pageNumber="36">microsuctus</emphasis>
</taxonomicName>
in the shape of the spermatodactyl (with a rounded protrusion near the tip which is absent in
<taxonomicName id="8AFC4D5CFFEDB81D8491C92AF30BFEB7" authorityName="Fain" authorityYear="1989" box="[719,887,331,357]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="microsuctus">
<emphasis id="7F88EACDFFEDB81D8491C92AF29FFEB7" box="[719,739,331,356]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D84AFC92AF30BFEB7" box="[753,887,331,356]" italics="true" pageId="35" pageNumber="36">microsuctus</emphasis>
</taxonomicName>
) and in the size of the opisthogastral suckers (of “standard” size vs. distinctly smaller than average in
<taxonomicName id="8AFC4D5CFFEDB81D84B2C90EF3E9FE5B" authorityName="Fain" authorityYear="1989" box="[748,917,367,393]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="microsuctus">
<emphasis id="7F88EACDFFEDB81D84B2C90EF37CFE5B" box="[748,768,367,392]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D8551C90EF3E9FE5B" box="[783,917,367,392]" italics="true" pageId="35" pageNumber="36">microsuctus</emphasis>
</taxonomicName>
). Distinguishing this species from the Berlese species,
<taxonomicName id="8AFC4D5CFFEDB81D86A9C9F2F1D8FE7E" authorityName="Berlese" authorityYear="1888" box="[247,420,403,429]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="degeneratus">
<emphasis id="7F88EACDFFEDB81D86A9C9F2F177FE7F" box="[247,267,403,428]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D8745C9F2F1D8FE7E" box="[283,420,403,429]" italics="true" pageId="35" pageNumber="36">degeneratus</emphasis>
</taxonomicName>
and
<taxonomicName id="8AFC4D5CFFEDB81D8783C9F2F24AFE7E" authorityName="Berlese" authorityYear="1902" box="[477,566,403,429]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="7F88EACDFFEDB81D8783C9F2F18DFE7F" box="[477,497,403,428]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D845EC9F2F24AFE7E" box="[512,566,403,429]" italics="true" pageId="35" pageNumber="36">latus</emphasis>
</taxonomicName>
, is more problematical given the lack of detail in these older descriptions. It appears that
<taxonomicName id="8AFC4D5CFFEDB81D877DC9D6F1B2FE02" authorityName="Berlese" authorityYear="1888" box="[291,462,439,465]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="degeneratus">
<emphasis id="7F88EACDFFEDB81D877DC9D6F14BFE03" box="[291,311,439,464]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D871BC9D6F1B2FE02" box="[325,462,439,465]" italics="true" pageId="35" pageNumber="36">degeneratus</emphasis>
</taxonomicName>
shares the relatively small opisthogastral suckers with
<taxonomicName id="8AFC4D5CFFEDB81D826FC9D6F4A4FE03" authorityName="Fain" authorityYear="1989" box="[1073,1240,439,465]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="microsuctus">
<emphasis id="7F88EACDFFEDB81D826FC9D6F439FE03" box="[1073,1093,439,464]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D820CC9D6F4A4FE03" box="[1106,1240,439,464]" italics="true" pageId="35" pageNumber="36">microsuctus</emphasis>
</taxonomicName>
(as already noted by
<bibRefCitation id="296D4B2EFFEDB81D86E5C9BAF134FE26" author="Fain, A." box="[187,328,475,501]" pageId="35" pageNumber="36" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain (1989))</bibRefCitation>
but neither
<bibRefCitation id="296D4B2EFFEDB81D878EC9BAF20BFE26" author="Berlese, A. &amp; Leonardi, G." box="[464,631,475,501]" pageId="35" pageNumber="36" pagination="12 - 18" refId="ref37203" refString="Berlese, A. &amp; Leonardi, G. (1902) Acari Sud Americani. Zoologischer Anzeiger, 25, 12 - 18." type="journal article" year="1902">Berlese (1902)</bibRefCitation>
or
<bibRefCitation id="296D4B2EFFEDB81D84C3C9BAF331FE26" author="Silvestri, F." box="[669,845,475,501]" pageId="35" pageNumber="36" pagination="171 - 234" refId="ref39685" refString="Silvestri, F. (1903) Contribuzione alla conoscenza dei Termitidi e Termitofili dell' America Meridionale. Parte II. Termitofili. Redia, Firenze, 1, 171 - 234, pls. I - VI. https: // doi. org / 10.5962 / bhl. title. 137413" type="journal article" year="1903">Silvestri (1903)</bibRefCitation>
provide sufficient information to distinguish
<taxonomicName id="8AFC4D5CFFEDB81D8310C9BAF16FFDCA" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="calakmulensis">
<emphasis id="7F88EACDFFEDB81D8310C9BAF51EFE27" box="[1358,1378,475,500]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D832FC9BAF16FFDCA" italics="true" pageId="35" pageNumber="36">calakmulensis</emphasis>
</taxonomicName>
from
<taxonomicName id="8AFC4D5CFFEDB81D8706C99EF1C6FDCA" box="[344,442,511,537]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="7F88EACDFFEDB81D8706C99EF110FDCB" box="[344,364,511,536]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D8724C99EF1C6FDCA" box="[378,442,511,537]" italics="true" pageId="35" pageNumber="36">latum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4D4336DFFFEDB81D8699CA42F53CFD1E" blockId="35.[151,1437,150,717]" pageId="35" pageNumber="36">
The designation of the medium pair of longer setae on the posterior margin sclerite as
<emphasis id="7F88EACDFFEDB81D82D2CA42F4D5FDEE" box="[1164,1193,547,573]" italics="true" pageId="35" pageNumber="36">S5</emphasis>
is based on comparative observations for other
<taxonomicName id="8AFC4D5CFFEDB81D879CCA26F2CBFDB2" box="[450,695,583,609]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7F88EACDFFEDB81D879CCA26F224FDB3" box="[450,600,583,608]" italics="true" pageId="35" pageNumber="36">Heterozercon</emphasis>
species.
</taxonomicName>
Some of these show the same arrangement as
<taxonomicName id="8AFC4D5CFFEDB81D82E7CA26F506FDB2" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[1209,1402,583,609]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="calakmulensis">
<emphasis id="7F88EACDFFEDB81D82E7CA26F4B1FDB3" box="[1209,1229,583,608]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D8285CA26F506FDB2" box="[1243,1402,583,609]" italics="true" pageId="35" pageNumber="36">calakmulensis</emphasis>
</taxonomicName>
, in others an added pair of distinctly longer setae is visible internal of, but very close to, the setae here designated as
<emphasis id="7F88EACDFFEDB81D8327CA0AF5E9FD56" box="[1401,1429,619,645]" italics="true" pageId="35" pageNumber="36">S5</emphasis>
. We consider that pair of setae
<emphasis id="7F88EACDFFEDB81D87B2CAEEF276FD7A" box="[492,522,655,681]" italics="true" pageId="35" pageNumber="36">Z5</emphasis>
. Because no such setae can be observed in
<taxonomicName id="8AFC4D5CFFEDB81D85AACAEEF4CAFD7A" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[1012,1206,655,681]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="calakmulensis">
<emphasis id="7F88EACDFFEDB81D85AACAEEF474FD7B" box="[1012,1032,655,680]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D8249CAEEF4CAFD7A" box="[1047,1206,655,681]" italics="true" pageId="35" pageNumber="36">calakmulensis</emphasis>
</taxonomicName>
we hypothesize that
<emphasis id="7F88EACDFFEDB81D86C9CAD2F0C9FD1E" box="[151,181,691,717]" italics="true" pageId="35" pageNumber="36">
Z
<quantity id="8A049B3AFFEDB81D86F6CAD2F0C9FD1E" box="[168,181,691,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="35" pageNumber="36" unit="in" value="5.0">5</quantity>
</emphasis>
in
<taxonomicName id="8AFC4D5CFFEDB81D8687CAD2F1E6FD1E" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[217,410,691,717]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="calakmulensis">
<emphasis id="7F88EACDFFEDB81D8687CAD2F091FD1F" box="[217,237,691,716]" italics="true" pageId="35" pageNumber="36">H</emphasis>
.
<emphasis id="7F88EACDFFEDB81D86A5CAD2F1E6FD1E" box="[251,410,691,717]" italics="true" pageId="35" pageNumber="36">calakmulensis</emphasis>
</taxonomicName>
are absent or, more likely, reduced to the size of the accessory setae on the sclerite.
</paragraph>
</subSubSection>
</treatment>
</document>