247 lines
35 KiB
XML
247 lines
35 KiB
XML
<document id="241EA88F2B9832484559B257A9FD3502" ID-DOI="10.11646/zootaxa.1575.1.2" ID-ISSN="11755334" ID-Zenodo-Dep="5096879" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626202182340" checkinUser="felipe" docAuthor="Sasakawa, Kôji" docDate="2007" docId="921787CBFFBEFF942FE5FF56FB51FA97" docLanguage="en" docName="zt01575p045.pdf" docOrigin="Zootaxa 1575 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1575.1.2" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Bembidion (Desarmatocillenus) yakushimanum Sasakawa 2007, sp. nov." docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="6E2EFFB3FFB9FF932F72FFC1FFADFFAB" masterDocTitle="Taxonomic studies on the Bembidion (‘ Cillenus’) complex (Coleoptera: Carabidae): A revision of the subgeneric taxonomy and description of a new species from Japan" masterLastPageNumber="45" masterPageNumber="35" pageNumber="42" updateTime="1699003504360" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="45D8D927539309E6B392B847CB51706F">Taxonomic studies on the Bembidion (‘ Cillenus’) complex (Coleoptera: Carabidae): A revision of the subgeneric taxonomy and description of a new species from Japan</mods:title>
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<mods:namePart id="D51D47C455BE73D3EE44E3A9022FBFBF">Sasakawa, Kôji</mods:namePart>
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<mods:title id="46C51E8056AB310B9EB45E052D9C06BF">Zootaxa</mods:title>
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<mods:date id="23E6FB248A617BD965495749AFE301D9">2007</mods:date>
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<mods:number id="EBE6AD41A931EE5105421E0396A306F6">2007-09-05</mods:number>
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<mods:number id="855E4A05F746744CBF312F4331DD7616">1575</mods:number>
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<paragraph id="1A0136DDFFBEFF942FE5FF56FC92FF1A" blockId="7.[151,831,151,212]" box="[151,831,151,177]" pageId="7" pageNumber="42">
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<heading id="414981B1FFBEFF942FE5FF56FC92FF1A" bold="true" box="[151,831,151,177]" fontSize="11" level="1" pageId="7" pageNumber="42" reason="1">
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<emphasis id="28CAEACFFFBEFF942FE5FF56FC92FF1A" bold="true" box="[151,831,151,177]" pageId="7" pageNumber="42">
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<taxonomicName id="DDBE4D5EFFBEFF942FE5FF56FD77FF1A" ID-CoL="LDS6" authority="Sasakawa, 2007" authorityName="Sasakawa" authorityYear="2007" box="[151,730,151,177]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="yakushimanum" status="sp. nov." subGenus="Desarmatocillenus">
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<emphasis id="28CAEACFFFBEFF942FE5FF56FEB6FF1A" bold="true" box="[151,283,151,177]" italics="true" pageId="7" pageNumber="42">Bembidion</emphasis>
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(
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<emphasis id="28CAEACFFFBEFF942E5EFF56FDA3FF1A" bold="true" box="[300,526,151,177]" italics="true" pageId="7" pageNumber="42">Desarmatocillenus</emphasis>
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)
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<emphasis id="28CAEACFFFBEFF942D6DFF56FD77FF1A" bold="true" box="[543,730,151,177]" italics="true" pageId="7" pageNumber="42">yakushimanum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="33F957B4FFBEFF942D90FF56FC92FF1A" box="[738,831,151,177]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="52A46556FFBEFF942FE5FF7BFE15FF7F" box="[151,440,186,212]" pageId="7" pageNumber="42" type="description">
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<paragraph id="1A0136DDFFBEFF942FE5FF7BFE15FF7F" blockId="7.[151,831,151,212]" box="[151,440,186,212]" pageId="7" pageNumber="42">
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(
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<figureCitation id="82852A58FFBEFF942FEDFF7BFE81FF7F" box="[159,300,186,212]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Figs. 4, 6, 8</figureCitation>
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,
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<figureCitation id="82852A58FFBEFF942E4BFF7BFEF5FF7F" box="[313,344,186,212]" captionStart="FIGURES 10–11" captionStartId="5.[151,269,1831,1855]" captionTargetBox="[313,1277,1574,1793]" captionTargetId="figure-325@5.[292,1300,1569,1794]" captionTargetPageId="5" captionText="FIGURES 10–11. Aedeagus of Bembidion yakushimanum (10, left lateral view) and B. tsutsuii (11, left lateral view of apical part). Scale: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096883" httpUri="https://zenodo.org/record/5096883/files/figure.png" pageId="7" pageNumber="42">10</figureCitation>
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,
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<figureCitation id="82852A58FFBEFF942E17FF7BFE1DFF7F" box="[357,432,186,212]" captionStart="FIGURES 12–24" captionStartId="6.[151,269,1049,1073]" captionTargetBox="[151,1436,194,1041]" captionTargetId="figure-352@6.[151,1436,194,1041]" captionTargetPageId="6" captionText="FIGURES 12–24. Endophalli (left anterolateral view, right lateral view) of Bembidion spp. (12–21) and an outgroup species, Blemus discus (22), and spermathecae of Bembidion spp. (23–24). 12, 23. B. sinicum; 13. B. kasaharai; 14. B. seticorne; 15. B. yokohamae; 16. B. aestuarii; 17. B. sumaoi; 18. B. tokunoshimanum; 19, 24. B. yakushimanum; 20. B. tsutsuii; 21. B. umi. Note that the endophallus of B. yokohamae (15) is not fully inflated because of damage during dissection. Numbers indicate character-number and character-state in Table 1. Scales: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096885" httpUri="https://zenodo.org/record/5096885/files/figure.png" pageId="7" pageNumber="42">19, 24</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="52A46556FFBEFF942FE5FECBFCB4FE2A" pageId="7" pageNumber="42" type="reference_group">
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<paragraph id="1A0136DDFFBEFF942FE5FECBFB89FE8A" blockId="7.[151,1060,266,385]" box="[151,1060,266,289]" pageId="7" pageNumber="42">
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<treatmentCitationGroup id="3AAE11F3FFBEFF942FE5FECBFB89FE8A" box="[151,1060,266,289]" pageId="7" pageNumber="42">
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<taxonomicName id="DDBE4D5EFFBEFF942FE5FECBFDEBFE8A" ID-CoL="8L3BK" authorityName="Ueno" authorityYear="1954" box="[151,582,266,289]" class="Insecta" family="Carabidae" genus="Armatocillenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="tsutsuii" subGenus="Corallicillenus">
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<emphasis id="28CAEACFFFBEFF942FE5FECBFE97FE8A" box="[151,314,266,289]" italics="true" pageId="7" pageNumber="42">Armatocillenus</emphasis>
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(
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<emphasis id="28CAEACFFFBEFF942E3BFECBFE44FE8A" box="[329,489,266,289]" italics="true" pageId="7" pageNumber="42">Corallicillenus</emphasis>
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)
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<emphasis id="28CAEACFFFBEFF942E8BFECBFDEBFE8A" box="[505,582,266,289]" italics="true" pageId="7" pageNumber="42">tsutsuii</emphasis>
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</taxonomicName>
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(part):
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<treatmentCitation id="9B1F10CCFFBEFF942DE4FECBFCF1FE8A" author="Nakane, T." box="[662,860,266,289]" page="103" pageId="7" pageNumber="42" year="1956">
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<bibRefCitation id="7E2F4B2CFFBEFF942DE4FECBFCF1FE8A" author="Nakane, T." box="[662,860,266,289]" pageId="7" pageNumber="42" pagination="101 - 104" refId="ref6886" refString="Nakane, T. (1956) On a new species of Armatocillenus and its allies in Japan. Insecta Matcumurana, 19, 101 - 104." type="journal article" year="1956">Nakane 1956: 103</bibRefCitation>
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||
</treatmentCitation>
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||
.
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<treatmentCitation id="9B1F10CCFFBEFF942C14FECBFB8DFE8A" author="Nakane, T." box="[870,1056,266,289]" page="26" pageId="7" pageNumber="42" year="1963">
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<bibRefCitation id="7E2F4B2CFFBEFF942C14FECBFB8DFE8A" author="Nakane, T." box="[870,1056,266,289]" pageId="7" pageNumber="42" pagination="22 - 54" refId="ref6914" refString="Nakane, T. (1963) Harpalidae. In: Nakane, T., Ohbayashi, K., Nomura, S. & Kurosawa, Y. (Eds), Iconographia Insectorum Japonicorum Colore Naturali Edita II (Coleoptera). Hokuryukan, Tokyo, pp. 22 - 54 (incl. pls. 11 - 27). (in Japanese)" type="book chapter" year="1963">Nakane 1963: 26</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="1A0136DDFFBEFF942FE5FEEBFC91FEEA" blockId="7.[151,1060,266,385]" box="[151,828,298,321]" pageId="7" pageNumber="42">
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<treatmentCitationGroup id="3AAE11F3FFBEFF942FE5FEEBFC91FEEA" box="[151,828,298,321]" pageId="7" pageNumber="42">
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<taxonomicName id="DDBE4D5EFFBEFF942FE5FEEBFDBBFEEA" ID-CoL="8LXBS" baseAuthorityName="Ueno" baseAuthorityYear="1954" box="[151,534,298,321]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="tsutsuii" subGenus="Corallicillenus">
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||
<emphasis id="28CAEACFFFBEFF942FE5FEEBFEA7FEEA" box="[151,266,298,321]" italics="true" pageId="7" pageNumber="42">Bembidion</emphasis>
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||
(
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<emphasis id="28CAEACFFFBEFF942E68FEEBFE17FEEA" box="[282,442,298,321]" italics="true" pageId="7" pageNumber="42">Corallicillenus</emphasis>
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||
)
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<emphasis id="28CAEACFFFBEFF942EBBFEEBFDBBFEEA" box="[457,534,298,321]" italics="true" pageId="7" pageNumber="42">tsutsuii</emphasis>
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</taxonomicName>
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(part):
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||
<treatmentCitation id="9B1F10CCFFBEFF942D14FEEBFC95FEEA" author="Lindroth, C. H." box="[614,824,298,321]" page="188" pageId="7" pageNumber="42" year="1980">
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||
<bibRefCitation id="7E2F4B2CFFBEFF942D14FEEBFC95FEEA" author="Lindroth, C. H." box="[614,824,298,321]" pageId="7" pageNumber="42" pagination="179 - 205" refId="ref6634" refString="Lindroth, C. H. (1980) A revisionary study of the taxon Cillenus Samouelle, 1829 and related forms (Coeloptera [sic]: Carabidae, Bembidiini). Entomologica Scandinavica, 11, 179 - 205." type="journal article" year="1980">Lindroth 1980: 188</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="1A0136DDFFBEFF942FE5FE8BFCE1FECA" blockId="7.[151,1060,266,385]" box="[151,844,330,353]" pageId="7" pageNumber="42">
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<treatmentCitationGroup id="3AAE11F3FFBEFF942FE5FE8BFCE1FECA" box="[151,844,330,353]" pageId="7" pageNumber="42">
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<taxonomicName id="DDBE4D5EFFBEFF942FE5FE8BFE20FECA" ID-CoL="8L3BK" authorityName="Ueno" authorityYear="1954" box="[151,397,330,353]" class="Insecta" family="Carabidae" genus="Armatocillenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="tsutsuii">
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<emphasis id="28CAEACFFFBEFF942FE5FE8BFE20FECA" box="[151,397,330,353]" italics="true" pageId="7" pageNumber="42">Armatocillenus tsutsuii</emphasis>
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</taxonomicName>
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(part):
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<treatmentCitation id="9B1F10CCFFBEFF942EACFE8BFD23FECA" author="Morita, S." box="[478,654,330,353]" page="92" pageId="7" pageNumber="42" year="1985">
|
||
<bibRefCitation id="7E2F4B2CFFBEFF942EACFE8BFD23FECA" author="Morita, S." box="[478,654,330,353]" pageId="7" pageNumber="42" pagination="89 - 102" refId="ref6747" refString="Morita, S. (1985) Carabidae (Bembidiinae, Patrobinae). In: Ueno, S. - I., Kurosawa, Y. & Sato, M. (Eds), The Coleoptera of Japan in color. Vol. 2. Hoikusha, Osaka, pp. 89 - 102 (incl. pls. 17 - 19). (in Japanese)" type="book chapter" year="1985">Morita 1985: 92</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
<treatmentCitation id="9B1F10CCFFBEFF942DEAFE8BFCE5FECA" author="Morita, S." box="[664,840,330,353]" page="17" pageId="7" pageNumber="42" year="1993">
|
||
<bibRefCitation id="7E2F4B2CFFBEFF942DEAFE8BFCE5FECA" author="Morita, S." box="[664,840,330,353]" pageId="7" pageNumber="42" pagination="17 - 22" refId="ref6818" refString="Morita, S. (1993) Marine carabid beetles from Japan. The Natrure and Insects, 28 (11), 17 - 22. (in Japanese)" type="journal article" year="1993">Morita 1993: 17</bibRefCitation>
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="1A0136DDFFBEFF942FE5FEABFCB4FE2A" blockId="7.[151,1060,266,385]" box="[151,793,362,385]" pageId="7" pageNumber="42">
|
||
<taxonomicName id="DDBE4D5EFFBEFF942FE5FEABFE51FE2A" ID-CoL="8LBSQ" baseAuthorityName="Ueno" baseAuthorityYear="1954" box="[151,508,362,385]" class="Insecta" family="Carabidae" genus="Cillenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="tsutsuii" subGenus="Corallicillenus">
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<emphasis id="28CAEACFFFBEFF942FE5FEABFF5DFE2A" box="[151,240,362,385]" italics="true" pageId="7" pageNumber="42">Cillenus</emphasis>
|
||
(
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<emphasis id="28CAEACFFFBEFF942F8DFEABFE32FE2A" box="[255,415,362,385]" italics="true" pageId="7" pageNumber="42">Corallicillenus</emphasis>
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||
)
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||
<emphasis id="28CAEACFFFBEFF942EDDFEABFE51FE2A" box="[431,508,362,385]" italics="true" pageId="7" pageNumber="42">tsutsuii</emphasis>
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||
</taxonomicName>
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||
(part):
|
||
<bibRefCitation id="7E2F4B2CFFBEFF942D39FEABFCB8FE2A" author="Marggi, W. A. & Huber, C. & Muller-Motzfeld, G. & Hartmann, M." box="[587,789,362,385]" pageId="7" pageNumber="42" pagination="241 - 273" refId="ref6676" refString="Marggi, W. A., Huber, C., Muller-Motzfeld, G. & Hartmann, M. (2003) Subtribe Bembidiina Stephens, 1827. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 241 - 273." type="book chapter" year="2003">
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||
Marggi
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||
<emphasis id="28CAEACFFFBEFF942DD2FEABFD7BFE2A" box="[672,726,362,385]" italics="true" pageId="7" pageNumber="42">et al.</emphasis>
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||
2003
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||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52A46556FFBEFF942FE5FE73FE1CFE5F" pageId="7" pageNumber="42" type="diagnosis">
|
||
<paragraph id="1A0136DDFFBEFF942FE5FE73FE1CFE5F" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
<emphasis id="28CAEACFFFBEFF942FE5FE73FEBAFE67" bold="true" box="[151,279,434,460]" pageId="7" pageNumber="42">Diagnosis.</emphasis>
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||
Similar to
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<taxonomicName id="DDBE4D5EFFBEFF942EE9FE73FDBCFE60" baseAuthorityName="Ueno" baseAuthorityYear="1954" box="[411,529,434,459]" class="Insecta" family="Carabidae" genus="Cillenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="tsutsuii">
|
||
<emphasis id="28CAEACFFFBEFF942EE9FE73FDBCFE60" box="[411,529,434,459]" italics="true" pageId="7" pageNumber="42">B. tsutsuii</emphasis>
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||
</taxonomicName>
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||
(
|
||
<figureCitation id="82852A58FFBEFF942D52FE73FDDEFE67" box="[544,627,434,460]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="82852A58FFBEFF942DF2FE73FD33FE67" box="[640,670,434,460]" captionStart="FIGURES 10–11" captionStartId="5.[151,269,1831,1855]" captionTargetBox="[313,1277,1574,1793]" captionTargetId="figure-325@5.[292,1300,1569,1794]" captionTargetPageId="5" captionText="FIGURES 10–11. Aedeagus of Bembidion yakushimanum (10, left lateral view) and B. tsutsuii (11, left lateral view of apical part). Scale: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096883" httpUri="https://zenodo.org/record/5096883/files/figure.png" pageId="7" pageNumber="42">11</figureCitation>
|
||
,
|
||
<figureCitation id="82852A58FFBEFF942DDEFE73FD60FE67" box="[684,717,434,460]" captionStart="FIGURES 12–24" captionStartId="6.[151,269,1049,1073]" captionTargetBox="[151,1436,194,1041]" captionTargetId="figure-352@6.[151,1436,194,1041]" captionTargetPageId="6" captionText="FIGURES 12–24. Endophalli (left anterolateral view, right lateral view) of Bembidion spp. (12–21) and an outgroup species, Blemus discus (22), and spermathecae of Bembidion spp. (23–24). 12, 23. B. sinicum; 13. B. kasaharai; 14. B. seticorne; 15. B. yokohamae; 16. B. aestuarii; 17. B. sumaoi; 18. B. tokunoshimanum; 19, 24. B. yakushimanum; 20. B. tsutsuii; 21. B. umi. Note that the endophallus of B. yokohamae (15) is not fully inflated because of damage during dissection. Numbers indicate character-number and character-state in Table 1. Scales: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096885" httpUri="https://zenodo.org/record/5096885/files/figure.png" pageId="7" pageNumber="42">20</figureCitation>
|
||
), but distinguished by blackish elytra (
|
||
<figureCitation id="82852A58FFBEFF942BE6FE73FB76FE67" box="[1172,1243,434,460]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Fig. 4</figureCitation>
|
||
) and a narrower aedeagal apex (
|
||
<figureCitation id="82852A58FFBEFF942E3EFE1BFE0EFE5F" box="[332,419,474,500]" captionStart="FIGURES 10–11" captionStartId="5.[151,269,1831,1855]" captionTargetBox="[313,1277,1574,1793]" captionTargetId="figure-325@5.[292,1300,1569,1794]" captionTargetPageId="5" captionText="FIGURES 10–11. Aedeagus of Bembidion yakushimanum (10, left lateral view) and B. tsutsuii (11, left lateral view of apical part). Scale: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096883" httpUri="https://zenodo.org/record/5096883/files/figure.png" pageId="7" pageNumber="42">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52A46556FFBEFF942FB5FDC3FB96FB6F" pageId="7" pageNumber="42" type="description">
|
||
<paragraph id="1A0136DDFFBEFF942FB5FDC3FDE3FD3F" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
<emphasis id="28CAEACFFFBEFF942FB5FDC3FEF2FDB7" bold="true" box="[199,351,514,540]" pageId="7" pageNumber="42">Description.</emphasis>
|
||
Body length (measured from mandible apices to elytral end):
|
||
<specimenCount id="0CB8FD54FFBEFF942B46FDC3FBE5FDBD" box="[1076,1096,514,534]" pageId="7" pageNumber="42" type="male">♂</specimenCount>
|
||
, 4.12–4.27 mm (mean ± SD: 4.19 ±
|
||
<quantity id="DD469B38FFBEFF942F98FDEBFECFFDEF" box="[234,354,554,580]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="42" unit="mm" value="74.0">0.074 mm</quantity>
|
||
) n=4;
|
||
<specimenCount id="0CB8FD54FFBEFF942EC2FDEBFE69FD95" box="[432,452,554,574]" pageId="7" pageNumber="42" type="female">♀</specimenCount>
|
||
, 4.14 mm, n=1. Dorsal surface of head, pronotum, elytral lateral margins and interval 1 reddish brown, remaining elytra blackish brown. Appendages light brown except for blackish brown to black inner margin of mandible apex.
|
||
</paragraph>
|
||
<paragraph id="1A0136DDFFBEFF942FB5FD63FC08FCA7" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
Head (
|
||
<figureCitation id="82852A58FFBEFF942E66FD63FEF6FD17" box="[276,347,674,700]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Fig. 4</figureCitation>
|
||
) longer than wide, with length from mandible apex to neck base 1.28–1.35 times longer than width between eyes; left mandible slightly convex at subapical inner margin; eyes moderate and almost flat, with the posterior edges in front of the posterior supraorbital setae.
|
||
</paragraph>
|
||
<paragraph id="1A0136DDFFBEFF942FB5FCDBFD19FC2F" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
Pronotum (
|
||
<figureCitation id="82852A58FFBEFF942E39FCDBFE3FFC9F" box="[331,402,794,820]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Fig. 4</figureCitation>
|
||
) 1.33–1.39 times wider than long; maximum width 1.19 times wider than anterior margin; 1.40–1.43 times wider than posterior margin; anterior angles notably produced, with apices narrowly rounded; hind angles right to somewhat acute.
|
||
</paragraph>
|
||
<paragraph id="1A0136DDFFBEFF942FB5FC53FCF4FB8F" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
Elytra (
|
||
<figureCitation id="82852A58FFBEFF942E6DFC53FECBFC07" box="[287,358,914,940]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Fig. 4</figureCitation>
|
||
) almost parallel-sided, 1.56–1.63 times longer than wide, with lateral margins weakly arcuate throughout; shoulders distinct; anterior discal setigerous puncture on basal 2/5, posterior one on apical 1/4; outermost intervals truncate and denticulate at subapical angle (
|
||
<figureCitation id="82852A58FFBEFF942C09FC23FC6FFC57" box="[891,962,994,1020]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Fig. 6</figureCitation>
|
||
). Hind trochanter (
|
||
<figureCitation id="82852A58FFBEFF942BEDFC23FB48FC57" box="[1183,1253,994,1020]" captionStart="FIGURES 1–9" captionStartId="5.[151,269,1353,1377]" captionTargetBox="[151,1436,797,1326]" captionTargetId="figure-322@5.[151,1436,797,1326]" captionTargetPageId="5" captionText="FIGURES 1–9. Habitus (1–5), elytral outermost intervals (6–7), and hind trochanter (8-9) of representative Bembidion species. 1. B. sinicum, male; 2. B. yokohamae, male (Heguri-gawa Riv); 3, 9. B. aestuarii, male (Rv. Obitsugawa). 4, 8. B. yakushimanum, holotype male; 5. B. tsutsuii, male; 6. B. yakushimanum, paratype female. 7. B. aestuarii, female (Riv. Tama). Arrows indicate posterior end of elytral outermost-intervals. Scales: 2.0 mm for 1, 1.0 mm for 2–5, 0.5 mm for 6– 9." figureDoi="http://doi.org/10.5281/zenodo.5096881" httpUri="https://zenodo.org/record/5096881/files/figure.png" pageId="7" pageNumber="42">Fig. 8</figureCitation>
|
||
) short, about 1/ 3 as long as hind femur; apex rounded. Hind wings reduced.
|
||
</paragraph>
|
||
<paragraph id="1A0136DDFFBEFF942FB5FBF3FEB5FB37" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
Aedeagus (
|
||
<figureCitation id="82852A58FFBEFF942E3EFBF3FE0FFBE7" box="[332,418,1074,1100]" captionStart="FIGURES 10–11" captionStartId="5.[151,269,1831,1855]" captionTargetBox="[313,1277,1574,1793]" captionTargetId="figure-325@5.[292,1300,1569,1794]" captionTargetPageId="5" captionText="FIGURES 10–11. Aedeagus of Bembidion yakushimanum (10, left lateral view) and B. tsutsuii (11, left lateral view of apical part). Scale: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096883" httpUri="https://zenodo.org/record/5096883/files/figure.png" pageId="7" pageNumber="42">Fig. 10</figureCitation>
|
||
) with apex relatively narrow. Parameres each with 2 setae at apex. Endophallus (
|
||
<figureCitation id="82852A58FFBEFF942A1EFBF3FF18FBDF" captionStart="FIGURES 12–24" captionStartId="6.[151,269,1049,1073]" captionTargetBox="[151,1436,194,1041]" captionTargetId="figure-352@6.[151,1436,194,1041]" captionTargetPageId="6" captionText="FIGURES 12–24. Endophalli (left anterolateral view, right lateral view) of Bembidion spp. (12–21) and an outgroup species, Blemus discus (22), and spermathecae of Bembidion spp. (23–24). 12, 23. B. sinicum; 13. B. kasaharai; 14. B. seticorne; 15. B. yokohamae; 16. B. aestuarii; 17. B. sumaoi; 18. B. tokunoshimanum; 19, 24. B. yakushimanum; 20. B. tsutsuii; 21. B. umi. Note that the endophallus of B. yokohamae (15) is not fully inflated because of damage during dissection. Numbers indicate character-number and character-state in Table 1. Scales: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096885" httpUri="https://zenodo.org/record/5096885/files/figure.png" pageId="7" pageNumber="42">Fig. 19</figureCitation>
|
||
) with 3 lobes; ventromedian lobe small; dorsomedian lobe large, with apex narrowly rounded; gonoporal lobe small.
|
||
</paragraph>
|
||
<paragraph id="1A0136DDFFBEFF942FB5FB6BFB96FB6F" blockId="7.[151,1436,434,1340]" box="[199,1083,1194,1220]" pageId="7" pageNumber="42">
|
||
Sclerotized part of spermathecal apex (
|
||
<figureCitation id="82852A58FFBEFF942DE2FB6BFD4BFB6F" box="[656,742,1194,1220]" captionStart="FIGURES 12–24" captionStartId="6.[151,269,1049,1073]" captionTargetBox="[151,1436,194,1041]" captionTargetId="figure-352@6.[151,1436,194,1041]" captionTargetPageId="6" captionText="FIGURES 12–24. Endophalli (left anterolateral view, right lateral view) of Bembidion spp. (12–21) and an outgroup species, Blemus discus (22), and spermathecae of Bembidion spp. (23–24). 12, 23. B. sinicum; 13. B. kasaharai; 14. B. seticorne; 15. B. yokohamae; 16. B. aestuarii; 17. B. sumaoi; 18. B. tokunoshimanum; 19, 24. B. yakushimanum; 20. B. tsutsuii; 21. B. umi. Note that the endophallus of B. yokohamae (15) is not fully inflated because of damage during dissection. Numbers indicate character-number and character-state in Table 1. Scales: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5096885" httpUri="https://zenodo.org/record/5096885/files/figure.png" pageId="7" pageNumber="42">Fig. 24</figureCitation>
|
||
) two times longer than wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52A46556FFBEFF942FB5FB13FC6AFABF" pageId="7" pageNumber="42" type="materials_examined">
|
||
<paragraph id="1A0136DDFFBEFF942FB5FB13FC6AFABF" blockId="7.[151,1436,434,1340]" pageId="7" pageNumber="42">
|
||
<emphasis id="28CAEACFFFBEFF942FB5FB13FE27FB47" bold="true" box="[199,394,1234,1260]" pageId="7" pageNumber="42">Type materials.</emphasis>
|
||
<materialsCitation id="AAD63C80FFBEFF942EE6FB13FAA3FB47" box="[404,1294,1234,1260]" collectingDate="1984-07-22" collectorName="Yaku Is. & M. Tao" location="Anbo" pageId="7" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="holotype">
|
||
<typeStatus id="C505887FFFBEFF942EE6FB13FDA9FB47" box="[404,516,1234,1260]" pageId="7" pageNumber="42" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<specimenCount id="0CB8FD54FFBEFF942D66FB13FD85FB4D" box="[532,552,1234,1254]" pageId="7" pageNumber="42" type="male">♂</specimenCount>
|
||
, ‘
|
||
<location id="1F616006FFBEFF942D49FB13FD29FB47" LSID="urn:lsid:plazi:treatment:921787CBFFBEFF942FE5FF56FB51FA97:1F616006FFBEFF942D49FB13FD29FB47" box="[571,644,1234,1260]" name="Anbo" pageId="7" pageNumber="42" stateProvince="Kagoshima">Anbo</location>
|
||
,
|
||
<collectorName id="B74B530BFFBEFF942DE0FB13FD58FB47" box="[658,757,1234,1260]" pageId="7" pageNumber="42">Yaku Is.</collectorName>
|
||
<collectingRegion id="D87AF83FFFBEFF942D8DFB13FC6BFB47" box="[767,966,1234,1260]" country="Japan" name="Kagoshima" pageId="7" pageNumber="42">Kagoshima Pref.</collectingRegion>
|
||
<date id="6E00101DFFBEFF942CA2FB13FBDFFB47" box="[976,1138,1234,1260]" pageId="7" pageNumber="42" value="1984-07-22">
|
||
<collectingDate id="7E44E9F5FFBEFF942CA2FB13FBDFFB47" box="[976,1138,1234,1260]" pageId="7" pageNumber="42" value="1984-07-22">22. VII. 1984</collectingDate>
|
||
</date>
|
||
<collectorName id="B74B530BFFBEFF942B0EFB13FB7FFB47" box="[1148,1234,1234,1260]" pageId="7" pageNumber="42">M. Tao</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
’;
|
||
<materialsCitation id="AAD63C80FFBEFF942A51FB13FC6EFABF" collectingDate="1984-07-22" collectionCode="NSMT" collectorName="Yaku Is. & M. Tao" location="Anbo" pageId="7" pageNumber="42" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Kagoshima" typeStatus="paratype">
|
||
<typeStatus id="C505887FFFBEFF942A51FB13FA35FB47" box="[1315,1432,1234,1260]" pageId="7" pageNumber="42" type="paratype">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="0CB8FD54FFBEFF942FE5FB3BFF61FAA5" box="[151,204,1274,1300]" pageId="7" pageNumber="42" type="male">3♂♂</specimenCount>
|
||
<specimenCount id="0CB8FD54FFBEFF942FBEFB3BFF42FAA5" box="[204,239,1274,1300]" pageId="7" pageNumber="42" type="female">1♀</specimenCount>
|
||
, same data as holotype. Deposited in the collection of
|
||
<collectionCode id="7CAFAE18FFBEFF942C02FB3BFC6EFABF" box="[880,963,1274,1300]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="7" pageNumber="42">NSMT</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52A46556FFBEFF942FB5FAE3FB51FA97" box="[199,1276,1314,1340]" pageId="7" pageNumber="42" type="etymology">
|
||
<paragraph id="1A0136DDFFBEFF942FB5FAE3FB51FA97" blockId="7.[151,1436,434,1340]" box="[199,1276,1314,1340]" pageId="7" pageNumber="42">
|
||
<emphasis id="28CAEACFFFBEFF942FB5FAE3FEFEFA97" bold="true" box="[199,339,1314,1340]" pageId="7" pageNumber="42">Etymology.</emphasis>
|
||
Named after the
|
||
<typeStatus id="C505887FFFBEFF942D6DFAE3FDFCFA97" box="[543,593,1314,1340]" pageId="7" pageNumber="42">type</typeStatus>
|
||
locality, Yakushima Island,
|
||
<collectingRegion id="D87AF83FFFBEFF942CEFFAE3FB0BFA97" box="[925,1190,1314,1340]" country="Japan" name="Kagoshima" pageId="7" pageNumber="42">Kagoshima Prefecture</collectingRegion>
|
||
,
|
||
<collectingCountry id="62A9764DFFBEFF942BC3FAE3FB55FA97" box="[1201,1272,1314,1340]" name="Japan" pageId="7" pageNumber="42">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |