197 lines
22 KiB
XML
197 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.407.7353" ID-GBIF-Dataset="941e1e73-0c38-4f51-942a-f8d61ff3e194" ID-PMC="PMC4042703" ID-Pensoft-Pub="1313-2970-407-55" ID-PubMed="24899831" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-407-55" ModsDocOrigin="ZooKeys 407" ModsDocTitle="Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Zabrini (Coleoptera, Carabidae)" checkinTime="1451245910087" checkinUser="pensoft" docAuthor="Kavanaugh, David H., Hieke, Fritz, Liang, Hongbin & Dong, Dazhi" docDate="2014" docId="022EAA58A87F9CB4B1D64D3303354ABE" docLanguage="en" docName="ZooKeys 407: 55-119" docOrigin="ZooKeys 407" docSource="http://dx.doi.org/10.3897/zookeys.407.7353" docTitle="Amara (Amara) congrua Morawitz 1863" docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="9B7CFF91FFAD355FFFEB8169FFEB117D" masterDocTitle="Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Zabrini (Coleoptera, Carabidae)" masterLastPageNumber="119" masterPageNumber="55" pageNumber="63" updateTime="1668158414670" updateUser="ExternalLinkService">
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<mods:title>Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Zabrini (Coleoptera, Carabidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hieke, Fritz</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dong, Dazhi</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>407</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>55</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.407.7353</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.407.7353</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-407-55</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052912" LSID="urn:lsid:plazi:treatment:022EAA58A87F9CB4B1D64D3303354ABE" httpUri="http://treatment.plazi.org/id/022EAA58A87F9CB4B1D64D3303354ABE" lastPageId="10" lastPageNumber="65" pageId="8" pageNumber="63">
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<subSubSection pageId="8" pageNumber="63" type="nomenclature">
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<paragraph pageId="8" pageNumber="63">
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<pageBreakToken pageId="8" pageNumber="63" start="start">2</pageBreakToken>
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.
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<taxonomicName LSID="http://species-id.net/wiki/Amara_congrua" authority="Morawitz, 1863" authorityName="Morawitz" authorityYear="1863" class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Amara) congrua" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="congrua" subGenus="Amara">Amara (Amara) congrua Morawitz, 1863</taxonomicName>
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Figs 5
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<normalizedToken originalValue="b–c">b-c</normalizedToken>
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, e, 6b, 7
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<normalizedToken originalValue="e–f">e-f</normalizedToken>
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, 10, 22b, 27-30
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="63" type="reference_group">
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara mongolica" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="mongolica">Amara mongolica</taxonomicName>
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Motschulsky, 1844: 185, nomen oblitum. Type material: Number of syntypes not specified and unknown, a fragmentary male specimen in ZMMU (
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<bibRefCitation author="Keleinikova, SI" journalOrPublisher="Sbornik Trudov Zoologicheskogo Muzeia MGU" pageId="42" pageNumber="97" pagination="183 - 224" title="V. I. Motschulsky's types of Coleoptera in the collection of the Zoological Museum MGU, 1. Carabidae." volume="15" year="1976">Keleinikova 1976</bibRefCitation>
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: 206) identified as holotype in that collection by Hieke. Type locality: "sur la
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<normalizedToken originalValue="frontière">frontiere</normalizedToken>
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de la Mongolie", probably northeastern China, Outer Manchuria (now Primorsky Krai of Russia). Synonymy suggested by
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<bibRefCitation author="Morawitz, A" journalOrPublisher="Memoires de l'Academie Imperiale des Sciences de St. Petersbourg" pageId="43" pageNumber="98" pagination="1 - 84" title="Beitrag zur Kaeferfauna der Insel Jesso, 1. Cicindelidae et Carabici." volume="6" year="1863 b">Morawitz 1863b</bibRefCitation>
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: 63, synonymized by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 116).
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</paragraph>
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Amara) congrua" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="congrua" subGenus="Amara">Amara (Amara) congrua</taxonomicName>
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Morawitz, 1863b: 62. Type material: Number of syntypes not specified, but at least several, with 5 syntypes in ZIN; lectotype male and 1 paralectotype designated by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 25), the 3 other syntypes (paralectotypes) are adults of
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara chalcites" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chalcites">Amara chalcites</taxonomicName>
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Dejean. Type locality: Japan: Hakodate.
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</paragraph>
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara zimmermanni" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="zimmermanni">Amara zimmermanni</taxonomicName>
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Putzeys, 1875: LI (
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<normalizedToken originalValue="röm">roem</normalizedToken>
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. 51) [nec Heer, 1837: 38]. Type material: Number of syntypes not specified, but at least several, with 3 syntypes in IRSNB and 2 in DEI (
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<bibRefCitation author="Doebler, H" journalOrPublisher="Beitraege zur Entomologie, Deutsches Entomologisches Institut" pageId="40" pageNumber="95" pagination="99 - 150" title="Katalog der in den Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - XII (Coleoptera: Carabidae)." volume="28" year="1975">
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<normalizedToken originalValue="Döbler">Doebler</normalizedToken>
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1975
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</bibRefCitation>
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: 148); two of those syntypes are adults of
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara chalcites" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chalcites">Amara chalcites</taxonomicName>
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Dejean, and the specimen from Kyoto in IRSNB, which has been considered as
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<normalizedToken originalValue="“Type”">"Type"</normalizedToken>
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, is an adult of
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara congrua" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="congrua">Amara congrua</taxonomicName>
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Morawitz. Type locality: Japan, Kyoto. Synonymized by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 23), previously synonymized incorrectly with
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara chalcites" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chalcites">Amara chalcites</taxonomicName>
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Dejean by
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<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="40" pageNumber="95" pagination="1 - 5" title="Additions to the list of geadephagous Coleoptera of Japan, with synonimic and other remarks." volume="1876" year="1876">Bates (1876</bibRefCitation>
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: 4).
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</paragraph>
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara striatella" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="striatella">Amara striatella</taxonomicName>
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Putzeys, 1875: LII (
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<normalizedToken originalValue="röm">roem</normalizedToken>
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. 52). Type material: Number of syntypes not specified, but 4 of them in IRSNB; lectotype male designated by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 27); one of those syntypes is an adult of
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara chalcites" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chalcites">Amara chalcites</taxonomicName>
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Dejean. Type locality: Japan, Kyoto. Synonymized by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 23), previously synonymized incorrectly with
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara chalcites" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="chalcites">Amara chalcites</taxonomicName>
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Dejean (as "var. striatella") by
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<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="40" pageNumber="95" pagination="205 - 290" title="11. Supplement to the Geodephagous Coleoptera of Japan, chiefly from the collection of Mr. George Lewis, made during his second visit, from February, 1880, to September, 1881." volume="1883" year="1883">Bates (1883</bibRefCitation>
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: 242).
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</paragraph>
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Amara) mandzhurica" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="mandzhurica" subGenus="Amara">Amara (Amara) mandzhurica</taxonomicName>
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Lutshnik, 1935: 257. Type material: Holotype female (not a male as stated by Lutshnik) in ZIN. Type locality: China: Manchuria:
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<normalizedToken originalValue="“Ertzendjantzy”">"Ertzendjantzy"</normalizedToken>
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. Synonymized by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 58).
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</paragraph>
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Amara) ovatoides" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="ovatoides" subGenus="Amara">Amara (Amara) ovatoides</taxonomicName>
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Baliani, 1943: 38. Type material: Holotype female (not a male as stated by Baliani, but as can be seen in his Fig. 1), in MCSNG. Type locality: China, Shanghai. Synonymized by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="215 - 326" title="Revision der Amara-Untergattung Percosia Zimm. und Bemerkungen zu anderen Amara-Arten (Col., Carabidae)." volume="25" year="1978">Hieke (1978</bibRefCitation>
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: 314).
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</paragraph>
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<paragraph pageId="8" pageNumber="63">
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara (Amara) abnormalis" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="abnormalis" subGenus="Amara">Amara (Amara) abnormalis</taxonomicName>
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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, 1956: 213. Type material: Holotype and 1 paratype in ZFMK, 2 paratypes in NMPC. Type locality: China, Fujian, Kuatun. Synonymized by
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<bibRefCitation author="Hieke, F" journalOrPublisher="Deutsche Entomologische Zeitschrift (N. F.)" pageId="41" pageNumber="96" pagination="1 - 125" title="Beitrag zur Synonymie der palaearktischen Amara-Arten (Coleoptera, Carabidae)." volume="20" year="1973">Hieke (1973</bibRefCitation>
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: 2).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63" type="diagnosis">
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<paragraph pageId="8" pageNumber="63">Diagnosis.</paragraph>
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<paragraph lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63">
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Adults of this species (Fig. 10) can be distinguished from those of all other species in the region by the following combination of character states: body length 9-10 mm; base of the pronotum evenly convex from one side to the other, outer basal impressions absent or only very faintly suggested; elytra with parascutellar pore puncture present; medial protibial spurs simple; tarsomere 5 of hind tarsi with two or (in a few specimens) three pairs of setae ventrally (Fig. 6b); last abdominal sternite of male with one pair (Fig. 5e)
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<pageBreakToken pageId="9" pageNumber="64" start="start">and</pageBreakToken>
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female with two pairs (Fig. 5c) of setiferous punctures near hind margin; sclerites of internal sac of median lobe of male aedeagus with form as in Fig. 7
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<normalizedToken originalValue="e–f">e-f</normalizedToken>
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.
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</paragraph>
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<caption pageId="9" pageNumber="64">
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<paragraph pageId="9" pageNumber="64">
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Figure 7. Apical and subapical region of median lobe (dorsal aspect) and apex of right paramere (medial aspect) of aedeagus of males.
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<normalizedToken originalValue="a–b">a-b</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara silvestrii" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="silvestrii">Amara silvestrii</taxonomicName>
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Baliani
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<normalizedToken originalValue="c–d">c-d</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara shaanxiensis" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="shaanxiensis">Amara shaanxiensis</taxonomicName>
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Hieke
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<normalizedToken originalValue="e–f">e-f</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara congrua" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="congrua">Amara congrua</taxonomicName>
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Morawitz.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="64" type="habitat distribution">
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<paragraph pageId="9" pageNumber="64">Habitat distribution.</paragraph>
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<paragraph pageId="9" pageNumber="64">
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Specimens of this species were collected in daytime from under stones and other cover in open roadside areas with scattered grasses and shrubs (Fig. 22b), at the edges of agricultural fields, including wet and dry rice paddies, and on open banks of streams, and in these same habitats at night, when beetles were found active on bare substrate. Specimens were found at elevations ranging from 892 to 2000 m. The only other
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="genus">Amara</taxonomicName>
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species members of which were found syntopic with those of
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara congrua" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="congrua">Amara congrua</taxonomicName>
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was
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<taxonomicName class="Insecta" family="Carabidae" genus="Amara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amara latithorax" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="latithorax">Amara latithorax</taxonomicName>
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, with adults of both species found together at only one site at an elevation of 1223 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="64" type="geographical distribution within the gaoligong shan">
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<paragraph pageId="9" pageNumber="64">Geographical distribution within the Gaoligong Shan.</paragraph>
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<paragraph pageId="9" pageNumber="64">Fig. 10d. We examined a total of 33 specimens (17 males and 16 females) from the following localities: Fugong County: Lumadeng Township (Chihengdi village, 26.98749°, 98.86310°, 1230 m, 29 April 2004, D.H. Kavanaugh & H.B. Liang collectors [3 males and 2 females; CAS, IOZ]); Shangpa Township (road on west side of Nu Jiang S of Shangpa, 26.88952°, 98.86539°, 1223 m, 22 April 2004, D.H. Kavanaugh & C.E. Griswold collectors [1 male and 1 female; CAS]). Gongshan County: Cikai Township (27.27064°, 98.66557°, 1500 m, 28 June 2000, D.H. Kavanaugh & H.B. Liang collectors [1 male; CAS]); Heiwadi village (27.78594°, 98.60117°, 1880-2000 m, 28 June 2000, D.H. Kavanaugh & H.B. Liang collectors [1 female; IOZ]). Lushui County: Liuku Township (Laimo village, 25.82767°, 98.85120°, 892 m, 15 May 2004, H.B. Liang & X.Y. Li collectors [1 male and 1 female; CAS, IOZ]). Tengchong County: Hehua Township (3.8 km S of Hehua along Daying Jiang at Nangyan village, 24.93873°, 98.38444°, 1140 m, 2 June 2006, D.H. Kavanaugh, R.L. Brett & D.Z. Dong collectors [1 female; CAS]), (5.4 km S of Hehua along Daying Jiang at Dengma village, 24.92346°, 98.38612°, 1105 m, 2 June 2006, D.H. Kavanaugh, R.L. Brett & D.Z. Dong collectors [1 female; IOZ]; Jietou Township (stream 0.7 km N of Jietou, 25.43128°, 98.64773°, 1564 m, 22 May 2006, D.H. Kavanaugh, R.L. Brett & H.B. Liang collectors [3 males and 2 females; CAS, IOZ]); Qingshui Township (Liangyang village in Rehai area, 24.94919°, 98.44921°, 1450 m, 1 June 2006, D.H. Kavanaugh, R.L. Brett, H.D. Liang, D.Z. Dong & P. Hu collectors [1 female; IOZ]); Qushi Township (Longchuan Jiang at Qinqiao in Qinqiao village, 25.27250°, 98.60083°, 1464 m, 6 June 2006, D.H. Kavanaugh, R.L. Brett & D.Z. Dong collectors [7 males and 5 females; CAS, IOZ, ZMHB]); Shangying Township (Bawan-Tengchong Road KM 63, 25.02917°, 98.66917°, 1360 m, 19 October 2003, H.B. Liang & X.C. Shi collectors [1 female; IOZ]); Wuhe Township (west bank of Longchuan Jiang at Tongjiazhuang village, 24.89284°, 98.67439°, 1210 m, H.B. Liang collector 24 May 2005 [1 male; IOZ]).</paragraph>
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<paragraph pageId="9" pageNumber="64">Members of this species were collected from the northern to the southern parts of the study area (Core Areas 2, 3, 5 and 6), but they were found only on the eastern side of the mountain range in northern and central areas (Core Areas 2,3 and 5) and only on the western versant in the southern part (Core Area 6). This distribution pattern is most likely an artifact of inadequate sampling on the western slope of the mountain range, much of which is in Myanmar.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="65" type="overall geographical distribution">
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<paragraph pageId="10" pageNumber="65">
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<pageBreakToken pageId="10" pageNumber="65" start="start">Overall</pageBreakToken>
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geographical distribution.
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</paragraph>
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<paragraph pageId="10" pageNumber="65">Fig. 27. This species has been recorded from China (Beijing, Fujian, Gansu, Guizhou, Hebei, Heilongjiang, Hongkong, Hubei, Jiangsu, Jiangxi, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanghai, Sichuan, Yunnan and Zhejian Provinces), Japan (southern half), Laos, North and South Korea, Myanmar (extreme north), Russia (Primorskij Krai), Taiwan and Vietnam (extreme north).</paragraph>
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</subSubSection>
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</treatment>
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</document> |