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<mods:title id="1239412CF000982580E55B7084D0E487">Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)</mods:title>
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<mods:affiliation id="0044CF40905BD9FAC08AE315FE1451E5">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<mods:affiliation id="A354F32AEF3AA81753F8756F15461E59">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143 &amp; Department of Entomology, PO BOX 37012, MRC 169, Smithsonian Institution, Washington, DC 20013 - 7012, USA mathisw @ si. edu; https: // orcid. org / 0000 - 0002 - 5012 - 1762 &amp; Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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15. 
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<emphasis id="B90CEAB33A6F7B6FFF1AF9A9568BF992" bold="true" box="[195,481,1613,1640]" italics="true" pageId="60" pageNumber="61">Scatophila variofacialis</emphasis>
Sturtevant and Wheeler
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<figureCitation id="13432A243A6F7B6FFF4EF995566BF971" box="[151,257,1649,1675]" captionStart="FIGURE 110" captionStartId="54.[151,250,1228,1252]" captionTargetBox="[151,1435,584,1204]" captionTargetId="figure-199@54.[151,1435,584,1204]" captionTargetPageId="54" captionText="FIGURE 110. Distribution map of Scatophila bolwigi sp. nov. (squares), S. tuberculosa Cresson (triangles), and S. variofacialis Sturtevant &amp; Wheeler (circles)." figureDoi="http://doi.org/10.5281/zenodo.13210849" httpUri="https://zenodo.org/record/13210849/files/figure.png" pageId="60" pageNumber="61">Figs. 110</figureCitation>
, 
<figureCitation id="13432A243A6F7B6FFED5F9955605F971" box="[268,367,1649,1675]" captionStart-0="FIGURES 130–133" captionStart-1="FIGURES 134–136" captionStart-2="FIGURES 137–141" captionStartId-0="61.[151,264,1375,1399]" captionStartId-1="62.[151,264,1325,1349]" captionStartId-2="63.[151,264,1742,1766]" captionTargetBox-0="[163,1435,181,1349]" captionTargetBox-1="[159,1427,497,1293]" captionTargetBox-2="[151,1435,355,1719]" captionTargetId-0="figure-27@61.[151,1435,181,1351]" captionTargetId-1="figure-159@62.[151,1435,489,1301]" captionTargetId-2="figure-93@63.[151,1435,345,1719]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionTargetPageId-2="63" captionText-0="FIGURES 130–133. Head of Scatophila variofacialis Sturtevant &amp; Wheeler. 130. Male, latero-oblique view. 131. Male, lateral view. 132. Female, latero-oblique view. 133. Female, lateral view. Scale bar = 0.2 mm." captionText-1="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." captionText-2="FIGURES 137–141. Head, thorax and wing of Scatophila. 137. S. variofacialis Sturtevant &amp; Wheeler, head and thorax, dorsal view. 138–139. Wing, dorsal view 138. S. variofacialis Sturtevant &amp; Wheeler. 139. S. arenaria Cresson. 140. Head of S. arenaria Cresson, latero-oblique view. 141. Same, lateral view. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13210865" figureDoi-1="http://doi.org/10.5281/zenodo.13210867" figureDoi-2="http://doi.org/10.5281/zenodo.13210869" httpUri-0="https://zenodo.org/record/13210865/files/figure.png" httpUri-1="https://zenodo.org/record/13210867/files/figure.png" httpUri-2="https://zenodo.org/record/13210869/files/figure.png" pageId="60" pageNumber="61">130–138</figureCitation>
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<treatmentCitation id="0AD910B03A6F7B6FFF4EF94B5595F93D" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[151,767,1711,1735]" page="199" pageId="60" pageNumber="61" year="1954">
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<emphasis id="B90CEAB33A6F7B6FFF4EF94B56E7F93D" box="[151,397,1711,1735]" italics="true" pageId="60" pageNumber="61">Scatophila variofacialis</emphasis>
<bibRefCitation id="EFE94B503A6F7B6FFE4EF94B5595F93D" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[407,767,1711,1735]" pageId="60" pageNumber="61" pagination="151 - 257" refId="ref70834" refString="Sturtevant, A. H. &amp; Wheeler, M. R. (1954) Synopses of Nearctic Ephydridae (Diptera). Transactions of the American Entomological Society, 79, 151 - 257." type="journal article" year="1954">Sturtevant and Wheeler 1954: 199</bibRefCitation>
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.— 
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<bibRefCitation id="EFE94B503A6F7B6FFCC6F94B54A6F93D" author="Wirth, W. W." box="[799,972,1711,1735]" pageId="60" pageNumber="61" pagination="734 - 759" refId="ref70987" refString="Wirth, W. W. (1965) Ephydridae. In: Stone, A., Sabrosky, C. W., Wirth, W. W., Foote, R. W. &amp; Coulson, J. R. (Eds.), A Catalog of the Diptera of America North of Mexico. Handbook 276. U. S. Department of Agriculture, Washington, D. C., pp. 734 - 759." type="book chapter" year="1965">Wirth 1965: 759</bibRefCitation>
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[
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catalog].— 
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<bibRefCitation id="EFE94B503A6F7B6FFB72F94B52F6F93D" author="Zatwarnicki, T." box="[1195,1436,1711,1735]" pageId="60" pageNumber="61" pagination="277 - 298" refId="ref71260" refString="Zatwarnicki, T. (1987) New synonyms and records of Palearctic Scatophila (Diptera, Ephydridae). Polskie Pismo Entomologiczne, 57 (2), 277 - 298." type="journal article" year="1987">Zatwarnicki 1987: 295</bibRefCitation>
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[checklist].— 
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<bibRefCitation id="EFE94B503A6F7B6FFE94F92A55D9F91C" author="Zatwarnicki, T. &amp; Mathis, W. N." box="[333,691,1742,1766]" pageId="60" pageNumber="61" pagination="351 - 370" refId="ref71480" refString="Zatwarnicki, T. &amp; Mathis, W. N. (1994) Phylogeny and classification of the genus Scatophila Becker (Diptera: Ephydridae). Annales de la Societe Entomologique de France, 29 (4), 351 - 370. [1993] https: // doi. org / 10.1080 / 21686351.1993.12277698" type="journal article" year="1994">Zatwarnicki and Mathis 1994: 364</bibRefCitation>
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[phylogeny; 
<emphasis id="B90CEAB33A6F7B6FFCE2F92A54EDF91C" box="[827,903,1742,1766]" italics="true" pageId="60" pageNumber="61">noctula</emphasis>
group].— 
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<bibRefCitation id="EFE94B503A6F7B6FFC36F92A5238F91C" author="Mathis, W. N. &amp; Zatwarnicki, T." box="[1007,1362,1742,1766]" pageId="60" pageNumber="61" pagination="1 - 423" refId="ref70301" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). International Memoirs on Entomology, 4, i - vi + 1 - 423." type="journal article" year="1995">Mathis and Zatwarnicki 1995: 287</bibRefCitation>
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[world catalog].
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Diagnosis.—This species is distinguished from congeners by the following combination of characters: Small to moderately small shore flies, body length 
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.
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<emphasis id="B90CEAB33A6F7B6FFF1EF890566EF874" box="[199,260,1908,1934]" italics="true" pageId="60" pageNumber="61">Head</emphasis>
(
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, 
<figureCitation id="13432A243A6F7B6FFE1BF890569AF874" box="[450,496,1908,1934]" captionStart="FIGURES 137–141" captionStartId="63.[151,264,1742,1766]" captionTargetBox="[151,1435,355,1719]" captionTargetId="figure-93@63.[151,1435,345,1719]" captionTargetPageId="63" captionText="FIGURES 137–141. Head, thorax and wing of Scatophila. 137. S. variofacialis Sturtevant &amp; Wheeler, head and thorax, dorsal view. 138–139. Wing, dorsal view 138. S. variofacialis Sturtevant &amp; Wheeler. 139. S. arenaria Cresson. 140. Head of S. arenaria Cresson, latero-oblique view. 141. Same, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210869" httpUri="https://zenodo.org/record/13210869/files/figure.png" pageId="60" pageNumber="61">137</figureCitation>
): Mesofrons and fronto-orbits tan to brown. Face of 
<specimenCount id="9D7EFD283A6F7B6FFBE2F8905324F874" box="[1083,1102,1908,1934]" count="1" pageId="60" pageNumber="61" type="male">♂</specimenCount>
brown, slightly protuberant at ventral margin, with an elongate, medial depression bordered by dense curly setulae, especially laterally; face of 
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with some subshiny luster. Gena moderately short, slightly more than half height of basal flagellomere; gena-to-eye ratio 0.24–0.25.
</paragraph>
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<emphasis id="B90CEAB33A6E7B6EFF4EFABB560DFA8D" bold="true" box="[151,359,1375,1399]" pageId="61" pageNumber="62">FIGURES 130–133</emphasis>
. Head of 
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Sturtevant &amp; Wheeler.
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130. Male, latero-oblique view. 131. Male, lateral view. 132. Female, latero-oblique view. 133. Female, lateral view. Scale bar = 0.2 mm.
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<emphasis id="B90CEAB33A6E7B6EFF1EFA095672F9FD" box="[199,280,1517,1543]" italics="true" pageId="61" pageNumber="62">Thorax</emphasis>
(
<figureCitation id="13432A243A6E7B6EFEF0FA0956BAF9F2" box="[297,464,1517,1544]" captionStart="FIGURES 137–141" captionStartId="63.[151,264,1742,1766]" captionTargetBox="[151,1435,355,1719]" captionTargetId="figure-93@63.[151,1435,345,1719]" captionTargetPageId="63" captionText="FIGURES 137–141. Head, thorax and wing of Scatophila. 137. S. variofacialis Sturtevant &amp; Wheeler, head and thorax, dorsal view. 138–139. Wing, dorsal view 138. S. variofacialis Sturtevant &amp; Wheeler. 139. S. arenaria Cresson. 140. Head of S. arenaria Cresson, latero-oblique view. 141. Same, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210869" httpUri="https://zenodo.org/record/13210869/files/figure.png" pageId="61" pageNumber="62">Figs. 137–138</figureCitation>
): Mesonotum (
<figureCitation id="13432A243A6E7B6EFDA2FA0955B6F9F2" box="[635,732,1517,1544]" captionStart="FIGURES 137–141" captionStartId="63.[151,264,1742,1766]" captionTargetBox="[151,1435,355,1719]" captionTargetId="figure-93@63.[151,1435,345,1719]" captionTargetPageId="63" captionText="FIGURES 137–141. Head, thorax and wing of Scatophila. 137. S. variofacialis Sturtevant &amp; Wheeler, head and thorax, dorsal view. 138–139. Wing, dorsal view 138. S. variofacialis Sturtevant &amp; Wheeler. 139. S. arenaria Cresson. 140. Head of S. arenaria Cresson, latero-oblique view. 141. Same, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210869" httpUri="https://zenodo.org/record/13210869/files/figure.png" pageId="61" pageNumber="62">Fig. 137</figureCitation>
) mostly brown with grayish stripes along setal tracks; scutum mostly brown with variable pattern of linear gray areas medially and anterolaterad of acrostichal tracks; a posteromedial, prescutellar spot; a medial, gray stripe; presutural dorsocentral setae lacking. Wing (
<figureCitation id="13432A243A6E7B6EFB28F9D15231F9AA" box="[1265,1371,1589,1616]" captionStart="FIGURES 137–141" captionStartId="63.[151,264,1742,1766]" captionTargetBox="[151,1435,355,1719]" captionTargetId="figure-93@63.[151,1435,345,1719]" captionTargetPageId="63" captionText="FIGURES 137–141. Head, thorax and wing of Scatophila. 137. S. variofacialis Sturtevant &amp; Wheeler, head and thorax, dorsal view. 138–139. Wing, dorsal view 138. S. variofacialis Sturtevant &amp; Wheeler. 139. S. arenaria Cresson. 140. Head of S. arenaria Cresson, latero-oblique view. 141. Same, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210869" httpUri="https://zenodo.org/record/13210869/files/figure.png" pageId="61" pageNumber="62">Figs. 138</figureCitation>
) with moderate pattern of white spots; veins R 
<subScript id="17FC34E43A6E7B6EFD8CF9885504F980" attach="left" box="[597,622,1644,1658]" fontSize="6" pageId="61" pageNumber="62">2+3</subScript>
and R 
<subScript id="17FC34E43A6E7B6EFD6CF98855A5F980" attach="left" box="[693,719,1644,1658]" fontSize="6" pageId="61" pageNumber="62">4+5</subScript>
shallowly undulate near white spots; costal vein ratio 0.24–0.30; M 
<subScript id="17FC34E43A6E7B6EFF69F97457D2F964" attach="left" box="[176,184,1680,1694]" fontSize="6" pageId="61" pageNumber="62">1</subScript>
vein ratio 0.59–0.72. Legs dark; male midfemur lacking row of spine-like setulae along posteroventral margin Tarsi often yellowish to brownish red basally.
</paragraph>
<paragraph id="8BC736A13A6E7B6DFF1EF92157BFFE2B" blockId="61.[151,1436,1517,2012]" lastBlockId="62.[151,1437,151,465]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<emphasis id="B90CEAB33A6E7B6EFF1EF921565BF925" box="[199,305,1733,1759]" italics="true" pageId="61" pageNumber="62">Abdomen</emphasis>
(
<figureCitation id="13432A243A6E7B6EFE9DF9215685F925" box="[324,495,1733,1759]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="61" pageNumber="62">Figs. 134–136</figureCitation>
): Mostly black, subshiny, with some blackish gray areas basally on tergites 2–4; subshiny. Male terminalia (
<figureCitation id="13432A243A6E7B6EFE08F90D5511F8F9" box="[465,635,1769,1795]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="61" pageNumber="62">Figs. 134–136</figureCitation>
): Epandrium in posterior view (
<figureCitation id="13432A243A6E7B6EFC34F90D5325F8FE" box="[1005,1103,1769,1796]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="61" pageNumber="62">Fig. 134</figureCitation>
) generally irregularly round, about as wide as high without measurement of ventrolateral projections, dorsal margin slightly projected dorsad, thereafter sloped ventrolaterally to lateral margins, lateral margins widest at midheight, and at midheight with shallow, rounded bulge, ventrolateral corners with ventral, digitiform projections, ventral margin curved ventrally, tending to be flattened medially, epandrial surface bearing sparsely scattered setulae, more clustered around ventral angle at base of ventrolateral projections; cercal cavity higher than wide, oval, ventral margin pointed ventromedially; cerci in posterior view (
<figureCitation id="13432A243A6E7B6EFEB1F82556A3F826" box="[360,457,1985,2012]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="61" pageNumber="62">Fig. 134</figureCitation>
) hemispherical; aedeagus in lateral view (
<figureCitation id="13432A243A6E7B6EFC71F8255363F821" box="[936,1033,1985,2011]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="61" pageNumber="62">Fig. 136</figureCitation>
) squarish, only slightly higher than wide, ventral margin broadly rounded, dorsal excavation irregularly V-shaped with a short, thumb-like projection, anterior and posterior corners angulate; in ventral view (
<figureCitation id="13432A243A6D7B6DFCCAFF5F5419FF2F" box="[787,883,186,213]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="62" pageNumber="63">Fig. 135</figureCitation>
) squarish, only slightly longer than width, apical margin evenly concave; ventral aedeagal process shallowly C-shaped; phallapodeme in lateral view (
<figureCitation id="13432A243A6D7B6DFAE8FF3B52FEFF03" box="[1329,1428,223,249]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="62" pageNumber="63">Fig. 136</figureCitation>
) with distinct keel, long and rounded apically, extension toward base of aedeagus slightly longer and more robust, extension to postgonite tapered, shorter than opposite extension, in ventral view (
<figureCitation id="13432A243A6D7B6DFB90FEC353C1FEBB" box="[1097,1195,294,321]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="62" pageNumber="63">Fig. 135</figureCitation>
) shallowly Y-shaped with lateral extensions capitate apically, width of “Y” about nearly twice length; postgonite in lateral view (
<figureCitation id="13432A243A6D7B6DFAB6FEAF57A8FE73" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="62" pageNumber="63">Fig. 136</figureCitation>
) generally irregularly V-shaped, dorsal arm rectangular, posterior apex robustly digitiform, ventral arm parallel sided, narrow, shallowly curved; neohypandrium in lateral view (
<figureCitation id="13432A243A6D7B6DFC5AFE77548EFE57" box="[899,996,403,429]" captionStart="FIGURES 134–136" captionStartId="62.[151,264,1325,1349]" captionTargetBox="[159,1427,497,1293]" captionTargetId="figure-159@62.[151,1435,489,1301]" captionTargetPageId="62" captionText="FIGURES 134–136. Male terminalia of Scatophila variofacialis Sturtevant &amp; Wheeler. 134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="62" pageNumber="63">Fig. 136</figureCitation>
) narrow, parallel sided, deeply curved strap.
</paragraph>
<caption id="DF0766293A6D7B6DFF4EFAC9560EFA77" ID-DOI="http://doi.org/10.5281/zenodo.13210867" ID-Zenodo-Dep="13210867" httpUri="https://zenodo.org/record/13210867/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,264,1325,1349]" targetBox="[159,1427,497,1293]" targetPageId="62" targetType="figure">
<paragraph id="8BC736A13A6D7B6DFF4EFAC9560EFA77" blockId="62.[151,1437,1325,1421]" pageId="62" pageNumber="63">
<emphasis id="B90CEAB33A6D7B6DFF4EFAC95603FABF" bold="true" box="[151,361,1325,1349]" pageId="62" pageNumber="63">FIGURES 134–136</emphasis>
. Male terminalia of 
<taxonomicName id="4C784D223A6D7B6DFDE6FAC95348FABF" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[575,1058,1325,1349]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="variofacialis">
<emphasis id="B90CEAB33A6D7B6DFDE6FAC95459FABF" box="[575,819,1325,1349]" italics="true" pageId="62" pageNumber="63">Scatophila variofacialis</emphasis>
Sturtevant &amp; Wheeler.
</taxonomicName>
134. Epandrium and cerci, posterior view. 135. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 136. Same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A6D7B6DFF1EFA045684F99F" blockId="62.[151,1437,1503,2034]" pageId="62" pageNumber="63">
Type Material.—The 
<typeStatus id="54C388033A6D7B6DFE19FA3B5548FA03" box="[448,546,1503,1529]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="4C784D223A6D7B6DFD55FA3B54F6FA03" box="[652,924,1503,1529]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="variofacialis">
<emphasis id="B90CEAB33A6D7B6DFD55FA3B54F6FA03" box="[652,924,1503,1529]" italics="true" pageId="62" pageNumber="63">Scatophila variofacialis</emphasis>
</taxonomicName>
is labeled “Carson, Wash[ington]. 8-6-51 [
<date id="FFC610613A6D7B6DFA54FA3B566FF9E4" pageId="62" pageNumber="63" value="1951-08-06">6 Aug 1951</date>
][handwritten]/M.R. Wheeler/ 
<typeStatus id="54C388033A6D7B6DFD8CF9E05583F9E4" box="[597,745,1540,1566]" pageId="62" pageNumber="63" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C784D223A6D7B6DFD36F9E753EFF9E7" authority="Stvt &amp; Whlr" authorityName="Stvt &amp; Whlr" box="[751,1157,1539,1566]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="variofacialis">Scatophila variofacialis Stvt &amp; Whlr</taxonomicName>
[pink]/6687 TYPE [dark pink; number handwritten].” The 
<typeStatus id="54C388033A6D7B6DFDCBF9C3551EF9BB" box="[530,628,1575,1601]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
is double mounted (glued to a paper point), is in good condition, and is deposited in the ANSP (6687).
</paragraph>
<paragraph id="8BC736A13A6D7B6DFF1EF98B5509F954" blockId="62.[151,1437,1503,2034]" pageId="62" pageNumber="63">
Other Specimens Examined.— 
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<collectingCountry id="F36F76313A6D7B6DFDEFF994547EF970" box="[566,788,1648,1674]" name="United States of America" pageId="62" pageNumber="63">UNITED STATES</collectingCountry>
. 
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<collectingRegion id="49BCF8433A6D7B6DFCF1F99454A6F970" box="[808,972,1648,1674]" country="United States of America" name="California" pageId="62" pageNumber="63">CALIFORNIA</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A6D7B6DFC39F98B532DF973" box="[992,1095,1647,1673]" pageId="62" pageNumber="63">Alameda</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFB84F98B5263F973" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFB84F98B5263F973" box="[1117,1289,1647,1673]" country="United States of America" latitude="62.268333" longLatPrecision="102" longitude="-145.385" municipality="Alameda" name="Berkeley Hills" pageId="62" pageNumber="63" stateProvince="California">Berkeley Hills</location>
(
<geoCoordinate id="EE4C50663A6D7B6DFAFAF98B52F2F973" box="[1315,1432,1647,1673]" degrees="62" direction="north" minutes="16.1" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="62.268333">62°16.1'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFF4EF977564AF954" box="[151,288,1683,1710]" degrees="145" direction="west" minutes="23.1" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-145.385">145°23.1'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFEEDF97056D5F954" box="[308,447,1684,1710]" pageId="62" pageNumber="63" value="1908-04-11">
<collectingDate id="EF82E9893A6D7B6DFEEDF97056D5F954" box="[308,447,1684,1710]" pageId="62" pageNumber="63" value="1908-04-11">11 Apr 1908</collectingDate>
</date>
(
<specimenCount id="9D7EFD283A6D7B6DFE09F970569FF954" box="[464,501,1684,1710]" count="1" pageId="62" pageNumber="63" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A6D7B6DFDD8F970553DF954" box="[513,599,1684,1710]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A6D7B6DFF1EF95C54A5F898" blockId="62.[151,1437,1503,2034]" pageId="62" pageNumber="63">
<materialsCitation id="3B103CFC3A6D7B6DFF1EF95C56C0F90C" collectingDate="2006-06-07" collectionCode="USNM" collectorName="D. and W. N. Mathis" elevation="825" latitude="44.82" location="Goose Creek" longLatPrecision="92" longitude="-117.463165" municipality="Baker" pageId="62" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Oregon">
<emphasis id="B90CEAB33A6D7B6DFF1EF95C56F8F92B" box="[199,402,1719,1746]" italics="true" pageId="62" pageNumber="63">
<collectingRegion id="49BCF8433A6D7B6DFF1EF95C5656F928" box="[199,316,1720,1746]" country="United States of America" name="Oregon" pageId="62" pageNumber="63">OREGON</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A6D7B6DFE9FF95356E6F92B" box="[326,396,1719,1745]" pageId="62" pageNumber="63">Baker</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFE40F95C5541F92B" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFE40F95C5541F92B" box="[409,555,1719,1746]" latitude="44.82" longLatPrecision="92" longitude="-117.463165" municipality="Baker" name="Goose Creek" pageId="62" pageNumber="63" stateProvince="Oregon">Goose Creek</location>
(
<locationDeviation id="9A752F833A6D7B6DFDE0F9535477F92B" box="[569,797,1719,1746]" location="Baker City" pageId="62" pageNumber="63">
<quantity id="4C809B443A6D7B6DFDE0F95355EBF92B" box="[569,641,1719,1746]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="62" pageNumber="63" unit="km" value="35.0">35 km</quantity>
E Baker City
</locationDeviation>
; 
<geoCoordinate id="EE4C50663A6D7B6DFCF1F95C54F6F928" box="[808,924,1720,1746]" degrees="44" direction="north" minutes="49.2" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="44.82">44°49.2'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFC7EF95C5352F928" box="[935,1080,1720,1746]" degrees="117" direction="west" minutes="27.79" orientation="longitude" pageId="62" pageNumber="63" precision="9" value="-117.463165">117°27.79'W</geoCoordinate>
; 
<quantity id="4C809B443A6D7B6DFB9EF95353FBF928" box="[1095,1169,1719,1746]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="62" pageNumber="63" unit="m" value="825.0">
<elevation id="0055D1923A6D7B6DFB9EF95353FBF928" box="[1095,1169,1719,1746]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="62" pageNumber="63" unit="m" value="825.0">825 m</elevation>
</quantity>
), 
<date id="FFC610613A6D7B6DFB7CF95C5248F92B" box="[1189,1314,1719,1746]" pageId="62" pageNumber="63" value="2006-06-07">
<collectingDate id="EF82E9893A6D7B6DFB7CF95C5248F92B" box="[1189,1314,1719,1746]" pageId="62" pageNumber="63" value="2006-06-07">7 Jun 2006</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFAF4F95C5662F90F" pageId="62" pageNumber="63">D. and W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A6D7B6DFEC5F9385657F90C" box="[284,317,1756,1782]" count="1" pageId="62" pageNumber="63" type="female">1♀</specimenCount>
; 
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)
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. 
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<emphasis id="B90CEAB33A6D7B6DFE6CF9385564F90F" box="[437,526,1756,1781]" italics="true" pageId="62" pageNumber="63">
<collectingMunicipality id="6BA3ACDB3A6D7B6DFE6CF938556DF90F" box="[437,519,1756,1781]" pageId="62" pageNumber="63">Benton</collectingMunicipality>
:
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<location id="8EA7607A3A6D7B6DFDCFF93F55E3F90F" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFDCFF93F55E3F90F" box="[534,649,1755,1781]" latitude="44.54" longLatPrecision="665" longitude="-123.36667" municipality="Benton" name="Philomath" pageId="62" pageNumber="63" stateProvince="Oregon">Philomath</location>
(
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W; 
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), 
<date id="FFC610613A6D7B6DFBFAF93853D7F90C" box="[1059,1213,1756,1782]" pageId="62" pageNumber="63" value="1972-05-29">
<collectingDate id="EF82E9893A6D7B6DFBFAF93853D7F90C" box="[1059,1213,1756,1782]" pageId="62" pageNumber="63" value="1972-05-29">29 May 1972</collectingDate>
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, 
<collectorName id="268D53773A6D7B6DFB11F938520AF90F" box="[1224,1376,1755,1782]" pageId="62" pageNumber="63">W. N. Mathis</collectorName>
(
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; 
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)
</materialsCitation>
; 
<materialsCitation id="3B103CFC3A6D7B6DFEDBF91B52F3F8E0" box="[258,1433,1791,1818]" collectingDate="1972-05-29" collectionCode="USNM" collectorName="W. N. Mathis" latitude="44.5" location="Rock Creek" longLatPrecision="927" longitude="-123.44" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Oregon">
<location id="8EA7607A3A6D7B6DFEDBF91B56E3F8E3" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFEDBF91B56E3F8E3" box="[258,393,1791,1817]" latitude="44.5" longLatPrecision="927" longitude="-123.44" name="Rock Creek" pageId="62" pageNumber="63" stateProvince="Oregon">Rock Creek</location>
(
<locationDeviation id="9A752F833A6D7B6DFE40F91B55F6F8E3" box="[409,668,1791,1818]" location="Philomath" pageId="62" pageNumber="63">
<quantity id="4C809B443A6D7B6DFE40F91B5681F8E3" box="[409,491,1791,1818]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="62" pageNumber="63" unit="km" value="6.5">6.5 km</quantity>
SW Philomath
</locationDeviation>
; 
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, 
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), 
<date id="FFC610613A6D7B6DFC6DF8E4533AF8E0" box="[948,1104,1792,1818]" pageId="62" pageNumber="63" value="1972-05-29">
<collectingDate id="EF82E9893A6D7B6DFC6DF8E4533AF8E0" box="[948,1104,1792,1818]" pageId="62" pageNumber="63" value="1972-05-29">29 May 1972</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFB85F8E4539CF8E3" box="[1116,1270,1791,1818]" pageId="62" pageNumber="63">W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A6D7B6DFAD1F8E45247F8E0" box="[1288,1325,1792,1818]" count="1" pageId="62" pageNumber="63" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A6D7B6DFAE2F8E452FBF8E0" box="[1339,1425,1792,1818]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<emphasis id="B90CEAB33A6D7B6DFF4EF8C7567AF8C7" box="[151,272,1827,1853]" italics="true" pageId="62" pageNumber="63">
<materialsCitation id="3B103CFC3A6D7B6DFF4EF8C75660F8C7" box="[151,266,1827,1853]" location="Josephine" pageId="62" pageNumber="63" specimenCount="1" stateProvince="Oregon">
<location id="8EA7607A3A6D7B6DFF4EF8C75660F8C7" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFF4EF8C75660F8C7" box="[151,266,1827,1853]" name="Josephine" pageId="62" pageNumber="63" stateProvince="Oregon">Josephine</location>
</materialsCitation>
:
</emphasis>
<materialsCitation id="3B103CFC3A6D7B6DFECEF8C053C5F8C4" box="[279,1199,1827,1854]" collectingDate="1972-06-27" collectionCode="USNM" collectorName="W. N. Mathis" latitude="42.09833" location="Caves" longLatPrecision="114" longitude="-123.40667" pageId="62" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Oregon">
<collectingRegion id="49BCF8433A6D7B6DFECEF8C05601F8C4" box="[279,363,1828,1854]" country="United States of America" name="Oregon" pageId="62" pageNumber="63">Oregon</collectingRegion>
<location id="8EA7607A3A6D7B6DFEABF8C056DCF8C4" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFEABF8C056DCF8C4" box="[370,438,1828,1854]" latitude="42.09833" longLatPrecision="114" longitude="-123.40667" name="Caves" pageId="62" pageNumber="63" stateProvince="Oregon">Caves</location>
(
<geoCoordinate id="EE4C50663A6D7B6DFE1FF8C75551F8C4" box="[454,571,1827,1854]" degrees="42" direction="north" minutes="05.9" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="42.09833">42°05.9'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFD9FF8C055BAF8C4" box="[582,720,1828,1854]" degrees="123" direction="west" minutes="24.4" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-123.40667">123°24.4'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFD3AF8C05405F8C4" box="[739,879,1828,1854]" pageId="62" pageNumber="63" value="1972-06-27">
<collectingDate id="EF82E9893A6D7B6DFD3AF8C05405F8C4" box="[739,879,1828,1854]" pageId="62" pageNumber="63" value="1972-06-27">27 Jun 1972</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFCA3F8C05365F8C7" box="[890,1039,1827,1854]" pageId="62" pageNumber="63">W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A6D7B6DFBFBF8C05329F8C4" box="[1058,1091,1828,1854]" count="1" pageId="62" pageNumber="63" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A6D7B6DFB88F8C053CDF8C4" box="[1105,1191,1828,1854]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6D7B6DFB60F8C754A1F898" collectingDate="1972-03-20" collectionCode="USNM" collectorName="W. N. Mathis" latitude="44.78" location="Helmick State Park" longLatPrecision="113" longitude="-123.236664" municipality="Polk" pageId="62" pageNumber="63" specimenCount="2" specimenCount-male="2" stateProvince="Oregon">
<emphasis id="B90CEAB33A6D7B6DFB60F8C7539CF8C7" box="[1209,1270,1827,1853]" italics="true" pageId="62" pageNumber="63">
<collectingMunicipality id="6BA3ACDB3A6D7B6DFB60F8C75385F8C7" box="[1209,1263,1827,1853]" pageId="62" pageNumber="63">Polk</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFB24F8C757A0F89B" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFB24F8C757A0F89B" latitude="44.78" longLatPrecision="113" longitude="-123.236664" municipality="Polk" name="Helmick State Park" pageId="62" pageNumber="63" stateProvince="Oregon">Helmick State Park</location>
(
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, 
<geoCoordinate id="EE4C50663A6D7B6DFE83F8AC5689F898" box="[346,483,1864,1890]" degrees="123" direction="west" minutes="14.2" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-123.236664">123°14.2'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFE2EF8AC55E1F898" box="[503,651,1864,1890]" pageId="62" pageNumber="63" value="1972-03-20">
<collectingDate id="EF82E9893A6D7B6DFE2EF8AC55E1F898" box="[503,651,1864,1890]" pageId="62" pageNumber="63" value="1972-03-20">20 Mar 1972</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFD4CF8AC5446F89B" box="[661,812,1863,1890]" pageId="62" pageNumber="63">W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A6D7B6DFCE2F8AC540BF898" box="[827,865,1864,1890]" count="2" pageId="62" pageNumber="63" type="male">2♂</specimenCount>
; 
<collectionCode id="ED69AE643A6D7B6DFCB4F8AC54A9F898" box="[877,963,1864,1890]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C362652A3A6D7B13FF1EF8885357FD53" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63" type="description">
<paragraph id="8BC736A13A6D7B6CFF1EF888539BFF2F" blockId="62.[151,1437,1503,2034]" lastBlockId="63.[151,1436,150,321]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<materialsCitation id="3B103CFC3A6D7B6DFF1EF88852F3F87C" box="[199,1433,1899,1926]" collectingDate="1934-09-03" collectionCode="USNM" collectorName="A. L. Melander" latitude="48.08" location="Sequim Bay" longLatPrecision="110" longitude="-123.10167" municipality="Clallam" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Washington">
<emphasis id="B90CEAB33A6D7B6DFF1EF8885683F87F" box="[199,489,1899,1926]" italics="true" pageId="62" pageNumber="63">
<collectingRegion id="49BCF8433A6D7B6DFF1EF8885610F87C" box="[199,378,1900,1926]" country="United States of America" name="Washington" pageId="62" pageNumber="63">WASHINGTON</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A6D7B6DFE5FF88F5689F87F" box="[390,483,1899,1925]" pageId="62" pageNumber="63">Clallam</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFE2AF88F5515F87C" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFE2AF88F5515F87C" box="[499,639,1899,1926]" latitude="48.08" longLatPrecision="110" longitude="-123.10167" municipality="Clallam" name="Sequim Bay" pageId="62" pageNumber="63" stateProvince="Washington">Sequim Bay</location>
(
<geoCoordinate id="EE4C50663A6D7B6DFD49F888546FF87C" box="[656,773,1900,1926]" degrees="48" direction="north" minutes="04.8" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="48.08">48°04.8'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFCCAF88F54F6F87F" box="[787,924,1899,1925]" degrees="123" direction="west" minutes="06.1" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-123.10167">123°06.1'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFC6BF8885352F87C" box="[946,1080,1900,1926]" pageId="62" pageNumber="63" value="1934-09-03">
<collectingDate id="EF82E9893A6D7B6DFC6BF8885352F87C" box="[946,1080,1900,1926]" pageId="62" pageNumber="63" value="1934-09-03">3 Sep 1934</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFB9AF888539EF87F" box="[1091,1268,1899,1926]" pageId="62" pageNumber="63">A. L. Melander</collectorName>
(
<specimenCount id="9D7EFD283A6D7B6DFAD1F8885246F87C" box="[1288,1324,1900,1926]" count="1" pageId="62" pageNumber="63" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A6D7B6DFAE2F88852FBF87C" box="[1339,1425,1900,1926]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6D7B6DFF4EF86B530DF850" box="[151,1127,1935,1962]" collectingDate="1917-07-07" collectionCode="USNM" collectorName="A. L. Melander" latitude="44.628334" location="Vancouver" longLatPrecision="113" longitude="-122.668335" municipality="Clark" pageId="62" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="WASHINGTON">
<emphasis id="B90CEAB33A6D7B6DFF4EF86B578AF853" box="[151,224,1935,1961]" italics="true" pageId="62" pageNumber="63">
<collectingMunicipality id="6BA3ACDB3A6D7B6DFF4EF86B57B0F853" box="[151,218,1935,1961]" pageId="62" pageNumber="63">Clark</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFF31F874560AF850" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFF31F874560AF850" box="[232,352,1936,1962]" latitude="44.628334" longLatPrecision="113" longitude="-122.668335" municipality="Clark" name="Vancouver" pageId="62" pageNumber="63" stateProvince="WASHINGTON">Vancouver</location>
(
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, 
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), 
<date id="FFC610613A6D7B6DFD49F8745461F850" box="[656,779,1935,1962]" pageId="62" pageNumber="63" value="1917-07-07">
<collectingDate id="EF82E9893A6D7B6DFD49F8745461F850" box="[656,779,1935,1962]" pageId="62" pageNumber="63" value="1917-07-07">7 Jul 1917</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFCCCF87454AEF853" box="[789,964,1935,1962]" pageId="62" pageNumber="63">A. L. Melander</collectorName>
(
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; 
<collectionCode id="ED69AE643A6D7B6DFBD0F8745335F850" box="[1033,1119,1936,1962]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6D7B6DFBABF874544EF834" collectingDate="1935-06-27" collectionCode="USNM" collectorName="A. L. Melander" latitude="46.43667" location="Toledo" longLatPrecision="112" longitude="-122.84167" municipality="Lewis" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="WASHINGTON">
<emphasis id="B90CEAB33A6D7B6DFBABF87453D7F853" box="[1138,1213,1936,1961]" italics="true" pageId="62" pageNumber="63">
<collectingMunicipality id="6BA3ACDB3A6D7B6DFBABF87453DCF853" box="[1138,1206,1936,1961]" pageId="62" pageNumber="63">Lewis</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFB1CF86B5278F853" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFB1CF86B5278F853" box="[1221,1298,1935,1961]" latitude="46.43667" longLatPrecision="112" longitude="-122.84167" municipality="Lewis" name="Toledo" pageId="62" pageNumber="63" stateProvince="WASHINGTON">Toledo</location>
(
<geoCoordinate id="EE4C50663A6D7B6DFAFAF86B52F2F853" box="[1315,1432,1935,1961]" degrees="46" direction="north" minutes="26.2" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="46.43667">46°26.2'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFF4EF857564BF834" box="[151,289,1971,1998]" degrees="122" direction="west" minutes="50.5" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-122.84167">122°50.5'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFEEFF85056AFF834" box="[310,453,1971,1998]" pageId="62" pageNumber="63" value="1935-06-27">
<collectingDate id="EF82E9893A6D7B6DFEEFF85056AFF834" box="[310,453,1971,1998]" pageId="62" pageNumber="63" value="1935-06-27">27 Jun 1935</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFE09F85055EAF837" box="[464,640,1971,1998]" pageId="62" pageNumber="63">A. L. Melander</collectorName>
(
<specimenCount id="9D7EFD283A6D7B6DFD4AF85055DDF834" box="[659,695,1972,1998]" count="1" pageId="62" pageNumber="63" type="male">1♂</specimenCount>
; 
<collectionCode id="ED69AE643A6D7B6DFD1CF8505471F834" box="[709,795,1972,1998]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="62" pageNumber="63" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6D7B6DFCF6F857553AF808" collectingDate="1972-07-19" collectionCode="USNM" collectorName="W. J. Turner" latitude="48.491665" location="Twisp" longLatPrecision="110" longitude="-120.11833" municipality="Okanagan" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="WASHINGTON">
<emphasis id="B90CEAB33A6D7B6DFCF6F85754C7F837" box="[815,941,1971,1997]" italics="true" pageId="62" pageNumber="63">
<collectingMunicipality id="6BA3ACDB3A6D7B6DFCF6F85754CDF837" box="[815,935,1971,1997]" pageId="62" pageNumber="63">Okanagan</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6D7B6DFC6CF8575493F837" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFC6CF8575493F837" box="[949,1017,1971,1997]" latitude="48.491665" longLatPrecision="110" longitude="-120.11833" municipality="Okanagan" name="Twisp" pageId="62" pageNumber="63" stateProvince="WASHINGTON">Twisp</location>
(
<quantity id="4C809B443A6D7B6DFBD3F8575331F837" box="[1034,1115,1971,1998]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="62" pageNumber="63" unit="km" value="6.5">6.5 km</quantity>
E; 
<geoCoordinate id="EE4C50663A6D7B6DFB5CF8575393F834" box="[1157,1273,1971,1998]" degrees="48" direction="north" minutes="29.5" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="48.491665">48°29.5'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFADFF85052E1F834" box="[1286,1419,1972,1998]" degrees="120" direction="west" minutes="07.1" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-120.11833">120°07.1'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFF4EF83C5676F808" box="[151,284,2007,2034]" pageId="62" pageNumber="63" value="1972-07-19">
<collectingDate id="EF82E9893A6D7B6DFF4EF83C5676F808" box="[151,284,2007,2034]" pageId="62" pageNumber="63" value="1972-07-19">19 Jul 1972</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6DFEFEF83C56DAF808" box="[295,432,2008,2034]" pageId="62" pageNumber="63">W. J. Turner</collectorName>
(
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; 
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)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6D7B6CFD83F83C56EEFF4B" collectingDate="1971-08-04" collectionCode="USNM" collectorName="La Grande &amp; W. N. Mathis" lastPageId="63" lastPageNumber="64" latitude="46.835" location="Pierce" longLatPrecision="112" longitude="-122.31834" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="WASHINGTON">
<emphasis id="B90CEAB33A6D7B6DFD83F83C55C0F80B" box="[602,682,2008,2033]" italics="true" pageId="62" pageNumber="63">
<location id="8EA7607A3A6D7B6DFD83F83C55CFF80B" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6D7B6DFD83F83C55CFF80B" box="[602,677,2008,2033]" latitude="46.835" longLatPrecision="112" longitude="-122.31834" name="Pierce" pageId="62" pageNumber="63" stateProvince="WASHINGTON">Pierce</location>
:
</emphasis>
<collectorName id="268D53773A6D7B6DFD6BF83C5442F80B" box="[690,808,2007,2034]" pageId="62" pageNumber="63">La Grande</collectorName>
(
<quantity id="4C809B443A6D7B6DFCEEF83C54ECF80B" box="[823,902,2007,2034]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="62" pageNumber="63" unit="km" value="4.8">4.8 km</quantity>
N; 
<geoCoordinate id="EE4C50663A6D7B6DFC69F833534EF808" box="[944,1060,2007,2034]" degrees="46" direction="north" minutes="50.1" orientation="latitude" pageId="62" pageNumber="63" precision="92" value="46.835">46°50.1'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6D7B6DFBF6F83C53DEF808" box="[1071,1204,2008,2034]" degrees="122" direction="west" minutes="19.1" orientation="longitude" pageId="62" pageNumber="63" precision="92" value="-122.31834">122°19.1'W</geoCoordinate>
), 
<date id="FFC610613A6D7B6DFB15F83C523AF808" box="[1228,1360,2008,2034]" pageId="62" pageNumber="63" value="1971-08-04">
<collectingDate id="EF82E9893A6D7B6DFB15F83C523AF808" box="[1228,1360,2008,2034]" pageId="62" pageNumber="63" value="1971-08-04">4 Aug 1971</collectingDate>
</date>
, 
<collectorName id="268D53773A6D7B6CFA82F83C578FFF4B" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">W. N. Mathis</collectorName>
(
<specimenCount id="9D7EFD283A6C7B6CFF2FFF735670FF4B" box="[246,282,151,177]" count="1" pageId="63" pageNumber="64" type="male">1♂</specimenCount>
; 
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)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6C7B6CFE57FF735252FF4B" box="[398,1336,150,177]" collectingDate="1925-07-04" collectionCode="USNM" collectorName="A. L. Melander" latitude="47.146667" location="Puget" longLatPrecision="112" longitude="-122.793335" municipality="Thurston" pageId="63" pageNumber="64" specimenCount="2" specimenCount-male="2" stateProvince="WASHINGTON">
<emphasis id="B90CEAB33A6C7B6CFE57FF735696FF4B" box="[398,508,151,177]" italics="true" pageId="63" pageNumber="64">
<collectingMunicipality id="6BA3ACDB3A6C7B6CFE57FF73569CFF4B" box="[398,502,151,177]" pageId="63" pageNumber="64">Thurston</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A6C7B6CFDDAFF735529FF4B" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6C7B6CFDDAFF735529FF4B" box="[515,579,151,177]" latitude="47.146667" longLatPrecision="112" longitude="-122.793335" municipality="Thurston" name="Puget" pageId="63" pageNumber="64" stateProvince="WASHINGTON">Puget</location>
(
<geoCoordinate id="EE4C50663A6C7B6CFD88FF7355ACFF4B" box="[593,710,151,177]" degrees="47" direction="north" minutes="08.8" orientation="latitude" pageId="63" pageNumber="64" precision="92" value="47.146667">47°08.8'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6C7B6CFD08FF735430FF4B" box="[721,858,151,177]" degrees="122" direction="west" minutes="47.6" orientation="longitude" pageId="63" pageNumber="64" precision="92" value="-122.793335">122°47.6'W</geoCoordinate>
), 
<date id="FFC610613A6C7B6CFCB7FF73548EFF4B" box="[878,996,150,177]" pageId="63" pageNumber="64" value="1925-07-04">
<collectingDate id="EF82E9893A6C7B6CFCB7FF73548EFF4B" box="[878,996,150,177]" pageId="63" pageNumber="64" value="1925-07-04">4 Jul 1925</collectingDate>
</date>
, 
<collectorName id="268D53773A6C7B6CFC34FF7353F2FF4B" box="[1005,1176,151,177]" pageId="63" pageNumber="64">A. L. Melander</collectorName>
(
<specimenCount id="9D7EFD283A6C7B6CFB71FF7353A4FF4B" box="[1192,1230,151,177]" count="2" pageId="63" pageNumber="64" type="male">2♂</specimenCount>
; 
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)
</materialsCitation>
. 
<materialsCitation id="3B103CFC3A6C7B6CFA98FF735387FF2F" collectingDate="1972-07-16" collectionCode="USNM" collectorName="W. J. Turner" county="Yakima" latitude="46.701668" location="Bear Creek" longLatPrecision="112" longitude="-120.753334" municipality="Tieton" pageId="63" pageNumber="64" specimenCount="1" specimenCount-female="1" stateProvince="WASHINGTON">
<emphasis id="B90CEAB33A6C7B6CFA98FF7352F6FF4B" box="[1345,1436,151,177]" italics="true" pageId="63" pageNumber="64">
<collectingCounty id="62A64E2D3A6C7B6CFA98FF7352FCFF4B" box="[1345,1430,151,177]" pageId="63" pageNumber="64">Yakima</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BA3ACDB3A6C7B6CFF4EFF5F57B5FF2F" box="[151,223,187,213]" pageId="63" pageNumber="64">Tieton</collectingMunicipality>
(
<quantity id="4C809B443A6C7B6CFF37FF5F5626FF2F" box="[238,332,187,213]" metricMagnitude="4" metricUnit="m" metricValue="1.28" pageId="63" pageNumber="64" unit="km" value="12.8">12.8 km</quantity>
SW; 
<geoCoordinate id="EE4C50663A6C7B6CFE53FF5F5694FF2F" box="[394,510,187,213]" degrees="46" direction="north" minutes="42.1" orientation="latitude" pageId="63" pageNumber="64" precision="92" value="46.701668">46°42.1'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6C7B6CFDD3FF5E55F8FF2F" box="[522,658,186,213]" degrees="120" direction="west" minutes="45.2" orientation="longitude" pageId="63" pageNumber="64" precision="92" value="-120.753334">120°45.2'W</geoCoordinate>
; 
<location id="8EA7607A3A6C7B6CFD44FF5F544AFF2F" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6C7B6CFD44FF5F544AFF2F" box="[669,800,187,213]" county="Yakima" latitude="46.701668" longLatPrecision="112" longitude="-120.753334" municipality="Tieton" name="Bear Creek" pageId="63" pageNumber="64" stateProvince="WASHINGTON">Bear Creek</location>
), 
<date id="FFC610613A6C7B6CFCEAFF5F54D3FF2F" box="[819,953,187,213]" pageId="63" pageNumber="64" value="1972-07-16">
<collectingDate id="EF82E9893A6C7B6CFCEAFF5F54D3FF2F" box="[819,953,187,213]" pageId="63" pageNumber="64" value="1972-07-16">16 Jul 1972</collectingDate>
</date>
, 
<collectorName id="268D53773A6C7B6CFC1AFF5F5326FF2F" box="[963,1100,187,213]" pageId="63" pageNumber="64">W. J. Turner</collectorName>
(
<specimenCount id="9D7EFD283A6C7B6CFBB9FF5F53EBFF2F" box="[1120,1153,187,213]" count="1" pageId="63" pageNumber="64" type="female">1♀</specimenCount>
; 
<collectionCode id="ED69AE643A6C7B6CFB56FF5F538FFF2F" box="[1167,1253,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="63" pageNumber="64" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A6C7B6CFF1EFF3B53C7FF03" blockId="63.[151,1436,150,321]" box="[199,1197,222,249]" pageId="63" pageNumber="64">
<materialsCitation id="3B103CFC3A6C7B6CFF1EFF3B53C2FF03" box="[199,1192,222,249]" country="United States of America" latitude="45.726665" location="Carson" longLatPrecision="112" longitude="-121.81834" municipality="Skamania" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<typeStatus id="54C388033A6C7B6CFF1EFF3B5795FF03" box="[199,255,223,249]" pageId="63" pageNumber="64">Type</typeStatus>
Locality.— 
<collectingCountry id="F36F76313A6C7B6CFE5DFF3B5577FF03" box="[388,541,223,249]" name="United States of America" pageId="63" pageNumber="64">United States</collectingCountry>
. 
<collectingRegion id="49BCF8433A6C7B6CFDF1FF3B55DBFF03" box="[552,689,223,249]" country="United States of America" name="Washington" pageId="63" pageNumber="64">Washington</collectingRegion>
. 
<collectingMunicipality id="6BA3ACDB3A6C7B6CFD62FF3B5444FF03" box="[699,814,223,249]" pageId="63" pageNumber="64">Skamania</collectingMunicipality>
: 
<location id="8EA7607A3A6C7B6CFCE1FF3B54EDFF03" LSID="urn:lsid:plazi:treatment:03D187B73A6F7B13FF4EF9A95357FD53:8EA7607A3A6C7B6CFCE1FF3B54EDFF03" box="[824,903,223,249]" country="United States of America" latitude="45.726665" longLatPrecision="112" longitude="-121.81834" municipality="Skamania" name="Carson" pageId="63" pageNumber="64" stateProvince="Washington">Carson</location>
(
<geoCoordinate id="EE4C50663A6C7B6CFC4EFF3A5366FF03" box="[919,1036,222,249]" degrees="45" direction="north" minutes="43.6" orientation="latitude" pageId="63" pageNumber="64" precision="92" value="45.726665">45°43.6'N</geoCoordinate>
, 
<geoCoordinate id="EE4C50663A6C7B6CFBCEFF3B53F6FF03" box="[1047,1180,223,249]" degrees="121" direction="west" minutes="49.1" orientation="longitude" pageId="63" pageNumber="64" precision="92" value="-121.81834">121°49.1'W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A6C7B6CFF1EFEE7531FFEE7" blockId="63.[151,1436,150,321]" box="[199,1141,259,285]" pageId="63" pageNumber="64">
Distribution (
<figureCitation id="13432A243A6C7B6CFE87FEE756D7FEE7" box="[350,445,259,285]" captionStart="FIGURE 110" captionStartId="54.[151,250,1228,1252]" captionTargetBox="[151,1435,584,1204]" captionTargetId="figure-199@54.[151,1435,584,1204]" captionTargetPageId="54" captionText="FIGURE 110. Distribution map of Scatophila bolwigi sp. nov. (squares), S. tuberculosa Cresson (triangles), and S. variofacialis Sturtevant &amp; Wheeler (circles)." figureDoi="http://doi.org/10.5281/zenodo.13210849" httpUri="https://zenodo.org/record/13210849/files/figure.png" pageId="63" pageNumber="64">Fig. 110</figureCitation>
).— 
<emphasis id="B90CEAB33A6C7B6CFE3EFEE7553AFEE6" box="[487,592,259,284]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="4C784D223A6C7B6CFE3EFEE75523FEE6" box="[487,585,259,284]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
:
</emphasis>
<collectingCountry id="F36F76313A6C7B6CFD8EFEE75587FEE7" box="[599,749,259,285]" name="United States of America" pageId="63" pageNumber="64">United States</collectingCountry>
(
<collectingRegion id="49BCF8433A6C7B6CFD25FEE75418FEE7" box="[764,882,259,285]" country="United States of America" name="California" pageId="63" pageNumber="64">California</collectingRegion>
, 
<collectingRegion id="49BCF8433A6C7B6CFCA4FEE754BEFEE7" box="[893,980,259,285]" country="United States of America" name="Oregon" pageId="63" pageNumber="64">Oregon</collectingRegion>
, 
<collectingRegion id="49BCF8433A6C7B6CFC06FEE75300FEE7" box="[991,1130,259,285]" country="United States of America" name="Washington" pageId="63" pageNumber="64">Washington</collectingRegion>
).
</paragraph>
<paragraph id="8BC736A13A6C7B6CFF1EFEC35458FEBB" blockId="63.[151,1436,150,321]" box="[199,818,295,321]" pageId="63" pageNumber="64">
Remarks.—This species is placed in the 
<emphasis id="B90CEAB33A6C7B6CFD57FEC35588FEBB" box="[654,738,295,321]" italics="true" pageId="63" pageNumber="64">noctula</emphasis>
group.
</paragraph>
<caption id="DF0766293A6C7B6CFF4EF92A5481F8D4" ID-DOI="http://doi.org/10.5281/zenodo.13210869" ID-Zenodo-Dep="13210869" httpUri="https://zenodo.org/record/13210869/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,264,1742,1766]" targetBox="[151,1435,355,1719]" targetPageId="63" targetType="figure">
<paragraph id="8BC736A13A6C7B6CFF4EF92A5481F8D4" blockId="63.[151,1437,1742,1838]" pageId="63" pageNumber="64">
<emphasis id="B90CEAB33A6C7B6CFF4EF92A560DF91C" bold="true" box="[151,359,1742,1766]" pageId="63" pageNumber="64">FIGURES 137–141</emphasis>
. Head, thorax and wing of 
<taxonomicName id="4C784D223A6C7B6CFDA5F92A5582F91C" box="[636,744,1742,1766]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB33A6C7B6CFDA5F92A5582F91C" box="[636,744,1742,1766]" italics="true" pageId="63" pageNumber="64">Scatophila</emphasis>
</taxonomicName>
. 137. 
<taxonomicName id="4C784D223A6C7B6CFCFEF92B53CCF91C" authority="Sturtevant &amp; Wheeler" authorityName="Sturtevant &amp; Wheeler" box="[807,1190,1742,1766]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="variofacialis">
<emphasis id="B90CEAB33A6C7B6CFCFEF92B54AAF91C" box="[807,960,1742,1766]" italics="true" pageId="63" pageNumber="64">S. variofacialis</emphasis>
Sturtevant &amp; Wheeler
</taxonomicName>
, head and thorax, dorsal view. 138–139. Wing, dorsal view 138. 
<taxonomicName id="4C784D223A6C7B6CFD98F91754BEF8F0" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[577,980,1778,1802]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="variofacialis">
<emphasis id="B90CEAB33A6C7B6CFD98F91755B4F8F0" box="[577,734,1778,1802]" italics="true" pageId="63" pageNumber="64">S. variofacialis</emphasis>
Sturtevant &amp; Wheeler.
</taxonomicName>
139. 
<taxonomicName id="4C784D223A6C7B6CFBCAF9175381F8F0" authority="Cresson." authorityName="Cresson." box="[1043,1259,1778,1802]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B90CEAB33A6C7B6CFBCAF91753E2F8F0" box="[1043,1160,1779,1802]" italics="true" pageId="63" pageNumber="64">S. arenaria</emphasis>
Cresson.
</taxonomicName>
140. Head of 
<taxonomicName id="4C784D223A6C7B6CFA50F9175623F8D4" authority="Cresson" authorityName="Cresson" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B90CEAB33A6C7B6CFA50F9175785F8D4" italics="true" pageId="63" pageNumber="64">S. arenaria</emphasis>
Cresson
</taxonomicName>
, latero-oblique view. 141. Same, lateral view. Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A137B13FF4EFF7356A4FF4B" blockId="64.[151,462,151,178]" box="[151,462,151,178]" pageId="64" pageNumber="65">
<emphasis id="B90CEAB33A137B13FF4EFF7356A4FF4B" bold="true" box="[151,462,151,178]" pageId="64" pageNumber="65">
The 
<emphasis id="B90CEAB33A137B13FF16FF73561EFF48" bold="true" box="[207,372,151,178]" italics="true" pageId="64" pageNumber="65">quadriguttata</emphasis>
Group
</emphasis>
</paragraph>
<paragraph id="8BC736A13A137B13FF4EFF3B5389FE0F" blockId="64.[151,1437,223,682]" pageId="64" pageNumber="65">
Diagnosis.—Medial portion of male face often depressed. Ventral margin of epandrium generally with short, sharp, medial process, sometimes broadly convex (
<figureCitation id="13432A243A137B13FD4AFEE75468FEE7" box="[659,770,259,285]" captionStart="FIGURES 188–190" captionStartId="81.[151,264,1066,1090]" captionTargetBox="[159,1427,189,1033]" captionTargetId="figure-27@81.[151,1435,181,1041]" captionTargetPageId="81" captionText="FIGURES 188–190. Male terminalia of Scatophila disjuncta Cresson. 188. Epandrium and cerci, posterior view. 189. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 190. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210919" httpUri="https://zenodo.org/record/13210919/files/figure.png" pageId="64" pageNumber="65">Figs. 188</figureCitation>
); neohypandrium band-like, strongly deflected; postgonite lobate with convex ventral margin; proximal margin arcuately rounded or slightly incised; distal process of postgonite triangular, its apex narrowly pointed, ventral process long and band-like; aedeagus with ventral process, in dorsal view elongate, in lateral view ventral margin slightly rounded or in medial portion nearly straight; distal margin of the dorsal aedeagal opening bears variously developed lobes, which could be acuminate or with lateral processes; phallapodeme in lateral view (
<figureCitation id="13432A243A137B13FE3EFE53553BFE2B" box="[487,593,439,465]" captionStart="FIGURES 188–190" captionStartId="81.[151,264,1066,1090]" captionTargetBox="[159,1427,189,1033]" captionTargetId="figure-27@81.[151,1435,181,1041]" captionTargetPageId="81" captionText="FIGURES 188–190. Male terminalia of Scatophila disjuncta Cresson. 188. Epandrium and cerci, posterior view. 189. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 190. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210919" httpUri="https://zenodo.org/record/13210919/files/figure.png" pageId="64" pageNumber="65">Figs. 190</figureCitation>
, 
<figureCitation id="13432A243A137B13FD85FE5355EDFE2B" box="[604,647,439,465]" captionStart="FIGURES 199–201" captionStartId="84.[151,264,1581,1605]" captionTargetBox="[159,1427,713,1549]" captionTargetId="figure-264@84.[151,1435,705,1557]" captionTargetPageId="84" captionText="FIGURES 199–201. Male terminalia of Scatophila hirsuta Sturtevant &amp; Wheeler. 199. Epandrium and cerci, posterior view. 200. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 201. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210933" httpUri="https://zenodo.org/record/13210933/files/figure.png" pageId="64" pageNumber="65">201</figureCitation>
) deflected before its proximal top, in dorsal view with subapical lateral projections, lobate, surrounding the proximal portion of the aedeagus, remaining portion elongate.
</paragraph>
<paragraph id="8BC736A13A137B13FF1EFE1B541EFD7F" blockId="64.[151,1437,223,682]" pageId="64" pageNumber="65">
<taxonomicName id="4C784D223A137B13FF1EFE1B5614FDE3" box="[199,382,511,537]" pageId="64" pageNumber="65">Nearctic species</taxonomicName>
included in the 
<emphasis id="B90CEAB33A137B13FDEDFE1B55A4FDE3" box="[564,718,511,537]" italics="true" pageId="64" pageNumber="65">quadriguttata</emphasis>
group are: 
<taxonomicName id="4C784D223A137B13FC96FE1B53F9FDE3" authority="Cresson" authorityName="Cresson" box="[847,1171,511,537]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B90CEAB33A137B13FC96FE1B5345FDE3" box="[847,1071,511,537]" italics="true" pageId="64" pageNumber="65">Scatophila arenaria</emphasis>
Cresson
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FB46FE1B5641FDC7" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bipiliaris">
<emphasis id="B90CEAB33A137B13FB46FE1B5248FDE3" box="[1183,1314,511,537]" italics="true" pageId="64" pageNumber="65">S. bipiliaris</emphasis>
Sturtevant and Wheeler
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FEE0FDC75542FDC7" authority="Cresson" authorityName="Cresson" box="[313,552,547,573]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="B90CEAB33A137B13FEE0FDC756ABFDC7" box="[313,449,547,573]" italics="true" pageId="64" pageNumber="65">S. bisignata</emphasis>
Cresson
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FDEEFDC754BAFDC7" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" box="[567,976,547,573]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A137B13FDEEFDC755DEFDC7" box="[567,692,547,573]" italics="true" pageId="64" pageNumber="65">S. conifera</emphasis>
Sturtevant and Wheeler
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FC07FDC75262FDC7" authority="(Stenhammar)" baseAuthorityName="Stenhammar" baseAuthorityYear="1844" box="[990,1288,547,573]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="cribrata">
<emphasis id="B90CEAB33A137B13FC07FDC75330FDC7" box="[990,1114,547,573]" italics="true" pageId="64" pageNumber="65">S. cribrata</emphasis>
(Stenhammar)
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FACFFDC7579EFD9B" authority="Cresson" authorityName="Cresson" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="disjuncta">
<emphasis id="B90CEAB33A137B13FACFFDC752F7FDC7" box="[1302,1437,547,573]" italics="true" pageId="64" pageNumber="65">S. disjuncta</emphasis>
Cresson
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FEDAFDA355E7FD9B" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" box="[259,653,583,609]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B90CEAB33A137B13FEDAFDA3561BFD9B" box="[259,369,583,609]" italics="true" pageId="64" pageNumber="65">S. hirsuta</emphasis>
Sturtevant and Wheeler
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FD42FDA35447FD9B" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[667,813,583,609]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A137B13FD42FDA35447FD9B" box="[667,813,583,609]" italics="true" pageId="64" pageNumber="65">S. mystacina</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A137B13FCEFFDA354F9FD9B" bold="true" box="[822,915,583,609]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="A23F57C83A137B13FCEFFDA354F9FD9B" box="[822,915,583,609]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C784D223A137B13FC7CFDA35272FD9B" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" box="[933,1304,583,609]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A137B13FC7CFDA35496FD9B" box="[933,1020,583,609]" italics="true" pageId="64" pageNumber="65">S. picta</emphasis>
Sturtevant and Wheeler
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FAFFFDA356CBFD7F" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B90CEAB33A137B13FAFFFDA352F6FD9B" box="[1318,1436,583,609]" italics="true" pageId="64" pageNumber="65">S. pulchra</emphasis>
Sturtevant and Wheeler
</taxonomicName>
, 
<taxonomicName id="4C784D223A137B13FE75FD8F541EFD7F" authority="Sturtevant and Wheeler." authorityName="Sturtevant and Wheeler." box="[428,884,619,645]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="rubribrunnea">
<emphasis id="B90CEAB33A137B13FE75FD8F5535FD7F" box="[428,607,619,645]" italics="true" pageId="64" pageNumber="65">S. rubribrunnea</emphasis>
Sturtevant and Wheeler.
</taxonomicName>
</paragraph>
<paragraph id="8BC736A13A137B13FF1EFD6B5357FD53" blockId="64.[151,1437,223,682]" box="[199,1085,655,682]" pageId="64" pageNumber="65">
This is the most speciose species group in the 
<taxonomicName id="4C784D223A137B13FD15FD6B5441FD53" box="[716,811,655,681]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="97" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
Region with 11 species.
</paragraph>
</subSubSection>
</treatment>
</document>