292 lines
31 KiB
XML
292 lines
31 KiB
XML
<document id="039FCBED0522772EDB9E63E3FD94EE13" ID-DOI="10.11646/phytotaxa.441.3.3" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727812835720" checkinUser="felipe" docAuthor="Mukherjee, Prasanta Kumar" docDate="2020" docId="039587D3281CFF81FF52E3F0FB7BE3B9" docLanguage="en" docName="phytotaxa.441.3.3.pdf" docOrigin="Phytotaxa 441 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.441.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Piper lamarckianum P. K. Mukh. 2020, nom. nov." docType="treatment" docVersion="1" lastPageNumber="267" masterDocId="FFACFFAB281FFF85FFDAE742FFF6E775" masterDocTitle="Nomenclatural notes on Piper (Piperaceae) from India III" masterLastPageNumber="273" masterPageNumber="263" pageNumber="266" updateTime="1727813926713" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="AB68AAC184760D807D974C3F2A62C9F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="975284BF090ABE00387FF68FDC8FDC10">
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<mods:title id="430D4A6375547A1424CFB0113F7FAE62">Nomenclatural notes on Piper (Piperaceae) from India III</mods:title>
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</mods:titleInfo>
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<mods:name id="7E725C7BF11EBE33ED4903F1FF27496E" type="personal">
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<mods:role id="8C81C043167FF100F1DA76CC36E5872F">
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<mods:roleTerm id="D98687C40C8BD85D69B2A4F40088C89E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F78D251D1227B426D2BFFAA3A9FC9F3E">Mukherjee, Prasanta Kumar</mods:namePart>
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<mods:typeOfResource id="A063CAAE25A30B93D0FC1CB826630D40">text</mods:typeOfResource>
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<mods:titleInfo id="D070B87D14C740C47FCEDA6EDB984AA2">
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<mods:title id="F378A9CD2A95005819D87775A68A0D2F">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F0AB8FCB28B949920017EAEA72886B11">
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<mods:date id="8A22A12C1B08FF57E77D15566DBF367F">2020</mods:date>
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<mods:detail id="E477B828854F9FD2F3EE2EE7600FFCFA" type="pubDate">
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<mods:number id="01D2C31222FAC8A51FE58BE53E502775">2020-05-05</mods:number>
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<mods:detail id="4624262ED9913294C07DEAFE601BF2EB" type="volume">
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<mods:number id="7E0AFDC76EE8E0E16690C52FD5154739">441</mods:number>
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</mods:detail>
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<mods:detail id="C2482B0A0E30CDC501E9865534EC9356" type="issue">
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<mods:number id="39D53945292F52DD17A2ACB5ABA7DF40">3</mods:number>
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<mods:start id="131AFD4C16FEA37B2634DF8757705D38">263</mods:start>
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<mods:end id="48EE42220555ED3625814B010359A691">273</mods:end>
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<mods:location id="DD3F0769982EB8A2722CD79315E1254A">
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<mods:url id="7C8B15A5C579364FD86B953F632E59A0">http://dx.doi.org/10.11646/phytotaxa.441.3.3</mods:url>
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</mods:location>
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<mods:classification id="D404B5F88AFFA03F7E93C72916D5B1A6">journal article</mods:classification>
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<mods:identifier id="E8DA61C918ED6AD2189D94D4AD30B60C" type="DOI">10.11646/phytotaxa.441.3.3</mods:identifier>
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<mods:identifier id="C7A96D12D4B26462B17499AB5607FC71" type="ISSN">1179-3163</mods:identifier>
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</mods:mods>
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<treatment id="039587D3281CFF81FF52E3F0FB7BE3B9" LSID="urn:lsid:plazi:treatment:039587D3281CFF81FF52E3F0FB7BE3B9" httpUri="http://treatment.plazi.org/id/039587D3281CFF81FF52E3F0FB7BE3B9" lastPageId="4" lastPageNumber="267" pageId="3" pageNumber="266">
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<subSubSection id="C326654E281CFF86FF52E3F0FD6EE3B8" box="[136,664,1202,1229]" pageId="3" pageNumber="266" type="nomenclature">
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<paragraph id="8B8336C5281CFF86FF52E3F0FD6EE3B8" blockId="3.[136,664,1202,1229]" box="[136,664,1202,1229]" pageId="3" pageNumber="266">
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<heading id="D0CB81A9281CFF86FF52E3F0FD6EE3B8" box="[136,664,1202,1229]" fontSize="11" level="2" pageId="3" pageNumber="266" reason="4">
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<emphasis id="B948EAD7281CFF86FF52E3F0FF6BE3B9" bold="true" box="[136,157,1202,1228]" pageId="3" pageNumber="266">5.</emphasis>
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<taxonomicName id="4C3C4D46281CFF86FF7EE3F0FDD7E3B9" ID-CoL="7Y86K" authority="P. K. Mukh." authorityName="P. K. Mukh." authorityYear="2020" box="[164,545,1202,1229]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="266" phylum="Tracheophyta" rank="species" species="lamarckianum" status="nom. nov.">
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<emphasis id="B948EAD7281CFF86FF7EE3F0FE65E3B8" bold="true" box="[164,403,1202,1229]" italics="true" pageId="3" pageNumber="266">Piper lamarckianum</emphasis>
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P. K. Mukh.
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</taxonomicName>
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<emphasis id="B948EAD7281CFF86FDFDE3F1FD6EE3B8" bold="true" box="[551,664,1203,1229]" italics="true" pageId="3" pageNumber="266">
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<taxonomicNameLabel id="A27B57AC281CFF86FDFDE3F1FD6EE3B8" box="[551,664,1203,1229]" pageId="3" pageNumber="266">nom. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C326654E281CFF86FF52E3BCFB36E266" box="[136,1216,1278,1300]" pageId="3" pageNumber="266" type="description">
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<paragraph id="8B8336C5281CFF86FF52E3BCFB36E266" blockId="3.[136,1452,1278,1516]" box="[136,1216,1278,1300]" pageId="3" pageNumber="266">
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<taxonomicName id="4C3C4D46281CFF86FF52E3BCFE22E261" authority="Lam. Tabl. Encycl." box="[136,468,1278,1300]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="266" phylum="Tracheophyta" rank="species" species="sylvestre">
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<emphasis id="B948EAD7281CFF86FF52E3BCFEE0E266" box="[136,278,1278,1299]" italics="true" pageId="3" pageNumber="266">Piper sylvestre</emphasis>
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Lam. Tabl. Encycl.
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</taxonomicName>
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1: 79. 1791
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<emphasis id="B948EAD7281CFF86FD96E3BDFD05E261" box="[588,755,1278,1300]" italics="true" pageId="3" pageNumber="266">nom. illeg. et non</emphasis>
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Loureiro. Fl. Cochinch. 1: 30. 1790.
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<emphasis id="B948EAD7281CFF86FB80E3BDFB4CE266" box="[1114,1210,1278,1300]" italics="true" pageId="3" pageNumber="266">nom. illeg</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326654E281CFF86FF52E260FD08E2B2" pageId="3" pageNumber="266" type="materials_examined">
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<paragraph id="8B8336C5281CFF86FF52E260FD08E2B2" blockId="3.[136,1452,1278,1516]" pageId="3" pageNumber="266">
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<emphasis id="B948EAD7281CFF86FF52E260FF2CE242" bold="true" box="[136,218,1314,1335]" pageId="3" pageNumber="266">Type:—</emphasis>
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<materialsCitation id="3B543C98281CFF86FF00E260FD90E229" collectorName="Huber & France & Lamarck Herbarium P" country="India" location="INDIA" pageId="3" pageNumber="266" specimenCount="1" typeStatus="lectotype">
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<collectingCountry id="F32B7655281CFF86FF00E260FEEBE242" box="[218,285,1314,1335]" name="India" pageId="3" pageNumber="266">INDIA</collectingCountry>
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&
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<collectingCountry id="F32B7655281CFF86FEE2E260FE37E242" box="[312,449,1314,1335]" name="Mauritius" pageId="3" pageNumber="266">MAURITIUS</collectingCountry>
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: “Insula Franciae, Malabaria, Philippinis (?)” (
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<typeStatus id="54878867281CFF86FCADE260FC27E24D" box="[887,977,1314,1336]" pageId="3" pageNumber="266" type="lectotype">lectotype</typeStatus>
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designated by
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<collectorName id="26C95313281CFF86FB86E260FB6EE24D" box="[1116,1176,1314,1336]" pageId="3" pageNumber="266">Huber</collectorName>
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(1987: 280): Isle-de-
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<collectorName id="26C95313281CFF86FABEE260FA5EE242" box="[1380,1448,1314,1335]" pageId="3" pageNumber="266">France</collectorName>
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,
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<collectorName id="26C95313281CFF86FF67E204FE60E22E" box="[189,406,1350,1371]" pageId="3" pageNumber="266">
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<emphasis id="B948EAD7281CFF86FF67E204FE75E22E" box="[189,387,1350,1371]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P
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</collectorName>
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(P00381410 image!)
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</materialsCitation>
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;
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<typeStatus id="54878867281CFF86FDABE204FD3EE22E" box="[625,712,1350,1371]" pageId="3" pageNumber="266" type="syntype">syntypes</typeStatus>
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; Isle-de-France,
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<emphasis id="B948EAD7281CFF86FCB3E204FBD9E22E" box="[873,1071,1350,1371]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P (P0038409 image!); Leponias, Male′ malabaro ou Leponias, Male′,
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<emphasis id="B948EAD7281CFF86FE33E228FD46E20A" box="[489,688,1386,1407]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P (P00381405 image!), Leponias, Male′,
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<emphasis id="B948EAD7281CFF86FB96E228FAE5E20A" box="[1100,1299,1386,1407]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P (P 00381406 image!), without locality,
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<emphasis id="B948EAD7281CFF86FE61E2CCFD75E2D6" box="[443,643,1422,1443]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P (P 00381407), without locality,
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<emphasis id="B948EAD7281CFF86FC0CE2CCFB6BE2D6" box="[982,1181,1422,1443]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P(P00381408) and without locality,
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<emphasis id="B948EAD7281CFF86FECAE2F0FE23E2B2" box="[272,469,1458,1479]" italics="true" pageId="3" pageNumber="266">Lamarck Herbarium</emphasis>
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P (P00381403 & P00381404).
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</paragraph>
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</subSubSection>
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<subSubSection id="C326654E281CFF81FF52E294FB7BE3B9" lastPageId="4" lastPageNumber="267" pageId="3" pageNumber="266" type="distribution">
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<paragraph id="8B8336C5281CFF86FF52E294FB95E299" blockId="3.[136,1452,1278,1516]" box="[136,1123,1494,1516]" pageId="3" pageNumber="266">
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<emphasis id="B948EAD7281CFF86FF52E294FED3E299" bold="true" box="[136,293,1494,1516]" pageId="3" pageNumber="266">Distribution:—</emphasis>
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<collectingCountry id="F32B7655281CFF86FEFFE294FE9BE29E" box="[293,365,1494,1515]" name="India" pageId="3" pageNumber="266">INDIA</collectingCountry>
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:
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<collectingRegion id="49F8F827281CFF86FEACE294FE4FE299" box="[374,441,1494,1516]" country="India" name="Kerala" pageId="3" pageNumber="266">Kerala</collectingRegion>
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;
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<collectingCountry id="F32B7655281CFF86FE19E294FDB3E29E" box="[451,581,1494,1515]" name="Sri Lanka" pageId="3" pageNumber="266">SRI LANKA</collectingCountry>
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,
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<collectingCountry id="F32B7655281CFF86FD94E294FD3AE29E" box="[590,716,1494,1515]" name="Maldives" pageId="3" pageNumber="266">MALDIVES</collectingCountry>
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, &
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<collectingCountry id="F32B7655281CFF86FD35E294FC82E29E" box="[751,884,1494,1515]" name="Mauritius" pageId="3" pageNumber="266">MAURITIUS</collectingCountry>
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ISLANDS (introduced).
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</paragraph>
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<paragraph id="8B8336C5281CFF81FF52E158FA68E7A9" blockId="3.[136,1452,1562,2093]" lastBlockId="4.[136,1452,158,1229]" lastPageId="4" lastPageNumber="267" pageId="3" pageNumber="266">
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<emphasis id="B948EAD7281CFF86FF52E158FF10E141" bold="true" box="[136,230,1562,1588]" pageId="3" pageNumber="266">Note:—</emphasis>
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Lamarck (1791: 79) has given a wider distribution for the species:
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<collectingCountry id="F32B7655281CFF86FC1CE158FBC0E141" box="[966,1078,1562,1588]" name="Mauritius" pageId="3" pageNumber="266">Mauritius</collectingCountry>
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, Malabar in
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<collectingCountry id="F32B7655281CFF86FB65E158FB0FE141" box="[1215,1273,1562,1588]" name="India" pageId="3" pageNumber="266">India</collectingCountry>
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and
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<collectingCountry id="F32B7655281CFF86FAF7E158FA5DE141" box="[1325,1451,1562,1588]" name="Philippines" pageId="3" pageNumber="266">Philippines</collectingCountry>
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which appear unnaturally a wider area. H. Huber (1987: 280) designated ‘Ile-de-France’ (
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<collectingCountry id="F32B7655281CFF86FB78E17CFAEEE12D" box="[1186,1304,1598,1624]" name="Mauritius" pageId="3" pageNumber="266">Mauritius</collectingCountry>
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!) at P as the nomenclatural type (
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<typeStatus id="54878867281CFF86FEAFE120FE11E109" box="[373,487,1634,1660]" pageId="3" pageNumber="266" type="lectotype">lectotype</typeStatus>
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!). There is one such specimen having a male spike at P (P00381410), but it carries no such annotation by Huber. However, the specimen is in poor state of representation for proper recognition. Notwithstanding these limitations, it is accepted as the
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<typeStatus id="54878867281CFF86FD22E1E8FCA9E1B1" box="[760,863,1706,1732]" pageId="3" pageNumber="266" type="lectotype">lectotype</typeStatus>
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(Art. 9.19, Turland
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<emphasis id="B948EAD7281CFF86FB99E1E9FB8BE1B1" box="[1091,1149,1706,1732]" italics="true" pageId="3" pageNumber="266">et al.</emphasis>
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2018). Friedmann in 1995 annotated a specimen at Lamarck’s herbarium (P00381405) as
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<taxonomicName id="4C3C4D46281CFF86FCBAE18DFBFFE19D" box="[864,1033,1742,1768]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="266" phylum="Tracheophyta" rank="species" species="sylvestre">
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<emphasis id="B948EAD7281CFF86FCBAE18DFBFFE19D" box="[864,1033,1742,1768]" italics="true" pageId="3" pageNumber="266">Piper sylvestre</emphasis>
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</taxonomicName>
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as ‘type’ without the mention of its locality. P00381406 also bears the annotation by Friedmann as
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<emphasis id="B948EAD7281CFF86FC91E1B1FC0EE079" box="[843,1016,1778,1804]" italics="true" pageId="3" pageNumber="266">
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<taxonomicName id="4C3C4D46281CFF86FC91E1B1FC03E079" box="[843,1013,1778,1804]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="266" phylum="Tracheophyta" rank="species" species="sylvestre">Piper sylvestre</taxonomicName>
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.
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</emphasis>
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Two more specimens at P (P00381407 and P00381408 respectively) were also annotated by Friedmann as
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<taxonomicName id="4C3C4D46281CFF86FC4BE055FBF9E045" box="[913,1039,1814,1840]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="266" phylum="Tracheophyta" rank="species" species="sylvestre">
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<emphasis id="B948EAD7281CFF86FC4BE055FBF9E045" box="[913,1039,1814,1840]" italics="true" pageId="3" pageNumber="266">P. sylvestre</emphasis>
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</taxonomicName>
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, but these appear to be the juvenile state of the species. Of all the specimens at Lamarck’s herbarium, the best one is (P00381409) which was annotated as
|
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<taxonomicName id="4C3C4D46281CFF86FF52E01DFED8E00D" box="[136,302,1886,1912]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="266" phylum="Tracheophyta" rank="species" species="sylvestre">
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<emphasis id="B948EAD7281CFF86FF52E01DFED8E00D" box="[136,302,1886,1912]" italics="true" pageId="3" pageNumber="266">Piper sylvestre</emphasis>
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</taxonomicName>
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by P. Sonnerat. It carries detached leaves and male and female spikes which look similar to those in the illustrations at Lamarck’s herbarium (P00381403 & P00381404, which are drawn side by side and labeled as “ou les Poivriers de la cote Malabarae”). The illustration as mentioned definitely indicates that the species is from the Malabar region of
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<collectingCountry id="F32B7655281CFF86FF28E088FED8E091" box="[242,302,1994,2020]" name="India" pageId="3" pageNumber="266">India</collectingCountry>
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, confirming the view of E. D. Merrill (1923: 18) that “As currently interpreted this is an Indian species”. Its distribution according to Huber (1987: 281) is southern
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<collectingCountry id="F32B7655281CFF86FCCAE0ACFCBCEF7D" box="[784,842,2030,2056]" name="India" pageId="3" pageNumber="266">India</collectingCountry>
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and
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<collectingCountry id="F32B7655281CFF86FCA7E0ACFC1AEF7D" box="[893,1004,2030,2056]" name="Sri Lanka" pageId="3" pageNumber="266">Sri Lanka</collectingCountry>
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, as well as the Mascarene Islands, where it is in cultivation or naturalized. Earlier, Hooker (l886: 93) refrained from citing any habitat or description outside of
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<collectingCountry id="F32B7655281BFF81FF52E7DCFF33E7CD" box="[136,197,158,184]" name="India" pageId="4" pageNumber="267">India</collectingCountry>
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. He stated that he had seen in Hooker’s herbarium a male specimen from
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<collectingCountry id="F32B7655281BFF81FBDEE7DCFB87E7CD" box="[1028,1137,158,184]" name="Mauritius" pageId="4" pageNumber="267">Mauritius</collectingCountry>
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annotated as
|
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<taxonomicName id="4C3C4D46281BFF81FAD1E7DDFA7EE7CD" box="[1291,1416,158,184]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
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<emphasis id="B948EAD7281BFF81FAD1E7DDFA7EE7CD" box="[1291,1416,158,184]" italics="true" pageId="4" pageNumber="267">P. sylvestre</emphasis>
|
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</taxonomicName>
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by Miquel and believed it to be a garden specimen. His statement of its distribution from Assam and Sylhet is doubtful.
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</paragraph>
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<paragraph id="8B8336C5281BFF81FF52E7A4FE6EE411" blockId="4.[136,1452,158,1229]" pageId="4" pageNumber="267">
|
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Opinions differ on the identity of
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<taxonomicName id="4C3C4D46281BFF81FDCDE7A5FB90E675" authority="Lour. C. L. Willdenow (1831: 673)" authorityName="Lour. C. L. Willdenow" authorityYear="1831" box="[535,1126,230,257]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
|
||
<emphasis id="B948EAD7281BFF81FDCDE7A5FD37E675" box="[535,705,230,256]" italics="true" pageId="4" pageNumber="267">Piper sylvestre</emphasis>
|
||
Lour. C. L. Willdenow (1831: 673)
|
||
</taxonomicName>
|
||
treated both Loureiro’s and Lamarck’s species as synonymous. Miquel (1843: 293) identified it with
|
||
<taxonomicName id="4C3C4D46281BFF81FC61E648FA51E651" authority="(Blume) Miquel (1843: 293)" authorityName="Miquel" authorityPageNumber="293" authorityYear="1843" baseAuthorityName="Blume" box="[955,1447,266,292]" class="Magnoliopsida" family="Piperaceae" genus="Cubeba" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="canina">
|
||
<emphasis id="B948EAD7281BFF81FC61E648FB93E651" box="[955,1125,266,292]" italics="true" pageId="4" pageNumber="267">Cubeba canina</emphasis>
|
||
(Blume) Miquel(1843: 293)
|
||
</taxonomicName>
|
||
. Merrill (1935:127) failed to identify this with any of the species of
|
||
<taxonomicName id="4C3C4D46281BFF81FC57E66CFBF0E63D" box="[909,1030,302,328]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
|
||
described from Indo-China by C. de Candolle (1810). C. Kunth (1840: 179) identified
|
||
<taxonomicName id="4C3C4D46281BFF81FD1AE611FC5DE619" authority="Lamarck" authorityName="Lamarck" box="[704,939,338,364]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
|
||
<emphasis id="B948EAD7281BFF81FD1AE611FCC8E619" box="[704,830,338,364]" italics="true" pageId="4" pageNumber="267">P. sylvestre</emphasis>
|
||
Lamarck
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C3C4D46281BFF81FC36E611FAD7E619" authority="Link (1821: 37)" authorityName="Link" authorityPageNumber="37" authorityYear="1821" box="[1004,1313,338,364]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="spurium">
|
||
<emphasis id="B948EAD7281BFF81FC36E611FB93E619" box="[1004,1125,339,364]" italics="true" pageId="4" pageNumber="267">P. spurium</emphasis>
|
||
Link (1821: 37)
|
||
</taxonomicName>
|
||
<emphasis id="B948EAD7281BFF81FAF0E611FAA2E619" box="[1322,1364,339,364]" italics="true" pageId="4" pageNumber="267">non</emphasis>
|
||
Forster ex Miquel (1839: 36). H. F.Link (1821: 37) did not cite a specimen (
|
||
<typeStatus id="54878867281BFF81FC49E634FC30E6E5" box="[915,966,374,400]" pageId="4" pageNumber="267">type</typeStatus>
|
||
) for his name. Link’s collections at B were destroyed during war (Stafleu & Cowan 1981: 65). A search for any surviving specimen of Link’s material elsewhere (as indicated by Stafleu & Cowan) was unsuccessful. As it stands at present, the exact identity of Link’s species cannot be established. Link described leaves as unequally cordate, measuring
|
||
<quantity id="4CC49B20281BFF81FC54E6A0FBF0E689" box="[910,1030,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="4" pageNumber="267" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 inches</quantity>
|
||
× 5–
|
||
<quantity id="4CC49B20281BFF81FBE1E6A0FB6EE689" box="[1083,1176,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="267" unit="in" value="4.0">4 inches</quantity>
|
||
, meaning they are ovoid. The leaf shape and measures are unlike the ones in
|
||
<taxonomicName id="4C3C4D46281BFF81FD60E545FCC3E555" box="[698,821,518,544]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
|
||
<emphasis id="B948EAD7281BFF81FD60E545FCC3E555" box="[698,821,518,544]" italics="true" pageId="4" pageNumber="267">P. sylvestre</emphasis>
|
||
</taxonomicName>
|
||
. Plate
|
||
<quantity id="4CC49B20281BFF81FCA5E544FC24E555" box="[895,978,518,544]" metricMagnitude="1" metricUnit="m" metricValue="4.91998" pageId="4" pageNumber="267" unit="in" value="1937.0">1937 in</quantity>
|
||
Wight’s Icon (1853), named as
|
||
<taxonomicName id="4C3C4D46281BFF81FAEAE545FA5DE555" box="[1328,1451,518,544]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
|
||
<emphasis id="B948EAD7281BFF81FAEAE545FA5DE555" box="[1328,1451,518,544]" italics="true" pageId="4" pageNumber="267">P. sylvestre</emphasis>
|
||
</taxonomicName>
|
||
shows a plant that is completely different from those present in Lamarck’s herbarium at P. It matches the description given for
|
||
<taxonomicName id="4C3C4D46281BFF81FF28E50DFE47E51D" box="[242,433,590,616]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="argyrophyllum">
|
||
<emphasis id="B948EAD7281BFF81FF28E50DFE47E51D" box="[242,433,590,616]" italics="true" pageId="4" pageNumber="267">P. argyrophyllum</emphasis>
|
||
</taxonomicName>
|
||
var. 2 by Hooker (1886: 94) and is named here as a distinct species
|
||
<taxonomicName id="4C3C4D46281BFF81FB40E50DFF02E5F9" authority="P. K. Mukh." authorityName="P. K. Mukh." authorityYear="2020" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="travancorianum" status="nom. nov.">
|
||
<emphasis id="B948EAD7281BFF81FB40E50DFA64E51D" box="[1178,1426,591,616]" italics="true" pageId="4" pageNumber="267">Piper travancorianum</emphasis>
|
||
P. K. Mukh.
|
||
</taxonomicName>
|
||
<emphasis id="B948EAD7281BFF81FF20E531FE90E5F9" box="[250,358,627,652]" italics="true" pageId="4" pageNumber="267">
|
||
<taxonomicNameLabel id="A27B57AC281BFF81FF20E531FE90E5F9" box="[250,358,627,652]" pageId="4" pageNumber="267">nom. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
G. H K. Thwaites (1861: 292) believed it to be same as
|
||
<taxonomicName id="4C3C4D46281BFF81FC0DE531FF5BE5C5" authority="Miquel. Hooker (1886: 93)" authorityName="Miquel. Hooker" authorityPageNumber="93" authorityYear="1886" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="hymenophyllum">
|
||
<emphasis id="B948EAD7281BFF81FC0DE531FB55E5F9" box="[983,1187,626,652]" italics="true" pageId="4" pageNumber="267">P. hymenophyllum</emphasis>
|
||
Miquel. Hooker (1886: 93)
|
||
</taxonomicName>
|
||
admitted that he found difficulties to define the species from the narrow-leaved forms of
|
||
<taxonomicName id="4C3C4D46281BFF81FB79E5D5FACBE5C5" box="[1187,1341,663,688]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="attenuatum">
|
||
<emphasis id="B948EAD7281BFF81FB79E5D5FACBE5C5" box="[1187,1341,663,688]" italics="true" pageId="4" pageNumber="267">P. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
and from some states of
|
||
<taxonomicName id="4C3C4D46281BFF81FEF5E5F9FE19E5A1" box="[303,495,698,724]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="argyrophyllum">
|
||
<emphasis id="B948EAD7281BFF81FEF5E5F9FE19E5A1" box="[303,495,698,724]" italics="true" pageId="4" pageNumber="267">P. argyrophyllum</emphasis>
|
||
</taxonomicName>
|
||
by description, “but feel sure being it is quite distinct from the former”. As observed in the present study, narrow leaved forms of
|
||
<taxonomicName id="4C3C4D46281BFF81FDACE59DFCF8E58D" box="[630,782,735,760]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="attenuatum">
|
||
<emphasis id="B948EAD7281BFF81FDACE59DFCF8E58D" box="[630,782,735,760]" italics="true" pageId="4" pageNumber="267">P. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
mentioned by Hooker is hard to be found. H. Trimen (1895: 429) expressed doubts about distinguishing it from
|
||
<emphasis id="B948EAD7281BFF81FD0CE440FC60E469" box="[726,918,770,796]" italics="true" pageId="4" pageNumber="267">
|
||
<taxonomicName id="4C3C4D46281BFF81FD0CE440FC65E469" box="[726,915,770,796]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="argyrophyllum">P.argyrophyllum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Characters of
|
||
<taxonomicName id="4C3C4D46281BFF81FBE5E441FAF9E469" authority="Lamk." authorityName="Lamk." box="[1087,1295,770,796]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
|
||
<emphasis id="B948EAD7281BFF81FBE5E441FB4BE469" box="[1087,1213,770,796]" italics="true" pageId="4" pageNumber="267">P. sylvestre</emphasis>
|
||
Lamk.
|
||
</taxonomicName>
|
||
as well as its differences with other species, like
|
||
<taxonomicName id="4C3C4D46281BFF81FDC5E465FD01E435" authority="Linn." authorityName="Linn." box="[543,759,806,832]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="malamiris">
|
||
<emphasis id="B948EAD7281BFF81FDC5E465FD59E435" box="[543,687,806,832]" italics="true" pageId="4" pageNumber="267">P. malamiris</emphasis>
|
||
Linn.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C4D46281BFF81FCDEE465FC48E435" authority="Vahl" authorityName="Vahl" authorityYear="1804" box="[772,958,806,832]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="diffusum">
|
||
<emphasis id="B948EAD7281BFF81FCDEE465FC74E435" box="[772,898,806,832]" italics="true" pageId="4" pageNumber="267">P. diffusum</emphasis>
|
||
Vahl
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D46281BFF81FC23E465FB4DE435" box="[1017,1211,806,832]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="argyrophyllum">
|
||
<emphasis id="B948EAD7281BFF81FC23E465FB4DE435" box="[1017,1211,806,832]" italics="true" pageId="4" pageNumber="267">P. argyrophyllum</emphasis>
|
||
</taxonomicName>
|
||
, have been noted by Huber (1987: 282–283).
|
||
</paragraph>
|
||
<paragraph id="8B8336C5281BFF81FF67E42CFD88E3DD" blockId="4.[136,1452,158,1229]" pageId="4" pageNumber="267">
|
||
Hooker (1886: 93) had expressed some reservation on the identity of the species and its distinctiveness from
|
||
<taxonomicName id="4C3C4D46281BFF81FA4DE42DFEDAE4D9" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="argyrophyllum">
|
||
<emphasis id="B948EAD7281BFF81FA4DE42DFEDAE4D9" italics="true" pageId="4" pageNumber="267">P. argyrophyllum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D46281BFF81FEB3E4D1FDFAE4D9" box="[361,524,915,941]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="attenuatum">
|
||
P.
|
||
<emphasis id="B948EAD7281BFF81FE50E4D1FDFAE4D9" box="[394,524,915,940]" italics="true" pageId="4" pageNumber="267">attenuatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAD7281BFF81FDD1E4D1FDF9E4D9" box="[523,527,915,940]" italics="true" pageId="4" pageNumber="267">.</emphasis>
|
||
These species can be distinguished from each other principally by their leaves. Leaves in
|
||
<taxonomicName id="4C3C4D46281BFF81FEDBE4F5FE76E4A5" box="[257,384,950,976]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="sylvestre">
|
||
<emphasis id="B948EAD7281BFF81FEDBE4F5FE76E4A5" box="[257,384,950,976]" italics="true" pageId="4" pageNumber="267">P. sylvestre</emphasis>
|
||
</taxonomicName>
|
||
are ovate, acuminate at the apex, broadly rounded or slightly unequally cordate at the base. Spikes of both sexes are long and slender. Leaves of
|
||
<taxonomicName id="4C3C4D46281BFF81FD3FE499FC76E481" box="[741,896,987,1012]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="attenuatum">
|
||
<emphasis id="B948EAD7281BFF81FD3FE499FC76E481" box="[741,896,987,1012]" italics="true" pageId="4" pageNumber="267">P. attenuatum</emphasis>
|
||
</taxonomicName>
|
||
are orbicular-ovate, mostly as broad as long and are deeply cordate or broadly truncate at the base and abruptly acuminate at the apex. It has long slender spikes of both sexes, bracts are decurrent with raised margins and fruits are globose. Leaves in
|
||
<taxonomicName id="4C3C4D46281BFF81FB93E361FAFFE349" box="[1097,1289,1058,1084]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="argyrophyllum">
|
||
<emphasis id="B948EAD7281BFF81FB93E361FAFFE349" box="[1097,1289,1058,1084]" italics="true" pageId="4" pageNumber="267">P. argyrophyllum</emphasis>
|
||
</taxonomicName>
|
||
are lanceolate to elliptic, sometimes whitish/silvery beneath with a rounded or acute sub equal base and a caudate/ acuminate apex. Its female spikes are shorter or equal the leaves in length and the peduncle equals the petiole, whereas fruits are ovate-elliptic, and subdensely placed on the rachis.
|
||
</paragraph>
|
||
<paragraph id="8B8336C5281BFF81FF67E3F0FB7BE3B9" blockId="4.[136,1452,158,1229]" box="[189,1165,1202,1229]" pageId="4" pageNumber="267">The specific epithet is proposed to commemorate Jean-Baptiste Lamarck (1744–1829).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |