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<document id="BDC58B93AFE6B9FCE629042DC5C2E1D8" ID-DOI="10.11646/phytotaxa.441.3.3" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727812835720" checkinUser="felipe" docAuthor="Mukherjee, Prasanta Kumar" docDate="2020" docId="039587D3281BFF80FF52E3B8FA63E63C" docLanguage="en" docName="phytotaxa.441.3.3.pdf" docOrigin="Phytotaxa 441 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.441.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Piper leptostachyum Miq. Syst. Piperac." docType="treatment" docVersion="1" lastPageNumber="267" masterDocId="FFACFFAB281FFF85FFDAE742FFF6E775" masterDocTitle="Nomenclatural notes on Piper (Piperaceae) from India III" masterLastPageNumber="273" masterPageNumber="263" pageNumber="267" updateTime="1727813926713" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4EE48018D2F5152FBEFC565F5F11DB82">Nomenclatural notes on Piper (Piperaceae) from India III</mods:title>
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<mods:namePart id="BE9E179227C23227F711DB718DCE52C6">Mukherjee, Prasanta Kumar</mods:namePart>
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<mods:title id="4287081BD0A316C0911F62521978E16D">Phytotaxa</mods:title>
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<mods:part id="103F44281C72A3691FBCB54CB13EEF49">
<mods:date id="7A22F059C222AB663A4EFD9B3E5396E9">2020</mods:date>
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<mods:identifier id="D00C770BD09C05AC84C77092603A13DA" type="DOI">10.11646/phytotaxa.441.3.3</mods:identifier>
<mods:identifier id="F27E9BBA28DF6F955F05EB4A3AAFC14F" type="ISSN">1179-3163</mods:identifier>
</mods:mods>
<treatment id="039587D3281BFF80FF52E3B8FA63E63C" LSID="urn:lsid:plazi:treatment:039587D3281BFF80FF52E3B8FA63E63C" httpUri="http://treatment.plazi.org/id/039587D3281BFF80FF52E3B8FA63E63C" lastPageId="5" lastPageNumber="267" pageId="4" pageNumber="267">
<subSubSection id="C326654E281BFF81FF52E3B8FD12E261" box="[136,740,1274,1301]" pageId="4" pageNumber="267" type="nomenclature">
<paragraph id="8B8336C5281BFF81FF52E3B8FD12E261" blockId="4.[136,740,1274,1301]" box="[136,740,1274,1301]" pageId="4" pageNumber="267">
<heading id="D0CB81A9281BFF81FF52E3B8FD12E261" box="[136,740,1274,1301]" fontSize="11" level="2" pageId="4" pageNumber="267" reason="4">
<emphasis id="B948EAD7281BFF81FF52E3B8FF6BE261" bold="true" box="[136,157,1274,1300]" pageId="4" pageNumber="267">6.</emphasis>
<taxonomicName id="4C3C4D46281BFF81FF7EE3B8FD9BE261" ID-CoL="4J4LN" authority="Miq. Syst. Piperac." box="[164,621,1274,1301]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="leptostachyum">
<emphasis id="B948EAD7281BFF81FF7EE3B8FE79E260" bold="true" box="[164,399,1274,1301]" italics="true" pageId="4" pageNumber="267">Piper leptostachyum</emphasis>
Miq. Syst. Piperac.
</taxonomicName>
315. 1844
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326654E281BFF80FF52E204FA63E63C" lastPageId="5" lastPageNumber="268" pageId="4" pageNumber="267" type="materials_examined">
<paragraph id="8B8336C5281BFF81FF52E204FD12E2F5" blockId="4.[136,1452,1350,1480]" pageId="4" pageNumber="267">
<emphasis id="B948EAD7281BFF81FF52E204FF2CE22E" bold="true" box="[136,218,1350,1371]" pageId="4" pageNumber="267">Type:—</emphasis>
<materialsCitation id="3B543C98281BFF81FF00E204FB10E22E" box="[218,1254,1350,1372]" collectionCode="K" country="Myanmar" county="Wallich" location="Nedaun" municipality="river Aatran" pageId="4" pageNumber="267" specimenCode="K001124408" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="F32B7655281BFF81FF00E204FEADE22E" box="[218,347,1350,1371]" name="Myanmar" pageId="4" pageNumber="267">MYANMAR</collectingCountry>
:
<location id="8EE3601E281BFF81FEBEE204FE47E229" LSID="urn:lsid:plazi:treatment:039587D3281BFF80FF52E3B8FA63E63C:8EE3601E281BFF81FEBEE204FE47E229" box="[356,433,1350,1372]" country="Myanmar" county="Wallich" municipality="river Aatran" name="Nedaun" pageId="4" pageNumber="267">Nedaun</location>
,
<collectingMunicipality id="6BE7ACBF281BFF81FE60E204FDD9E22E" box="[442,559,1350,1372]" pageId="4" pageNumber="267">river Aatran</collectingMunicipality>
,
<emphasis id="B948EAD7281BFF81FDE2E204FD76E22E" box="[568,640,1350,1371]" italics="true" pageId="4" pageNumber="267">
<collectingCounty id="62E24E49281BFF81FDE2E204FD76E22E" box="[568,640,1350,1371]" pageId="4" pageNumber="267">Wallich</collectingCounty>
</emphasis>
num. list no. 6649 (
<typeStatus id="54878867281BFF81FCE7E204FC61E229" box="[829,919,1350,1372]" pageId="4" pageNumber="267" type="lectotype">lectotype</typeStatus>
designated here)
<collectionCode id="ED2DAE00281BFF81FBE5E204FBA6E22E" box="[1087,1104,1350,1371]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
(
<specimenCode id="DB9A9EBE281BFF81FB86E204FB29E22E" box="[1116,1247,1350,1371]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K001124408</specimenCode>
)
</materialsCitation>
;
<typeStatus id="54878867281BFF81FB34E204FA9DE229" box="[1262,1387,1350,1372]" pageId="4" pageNumber="267" type="isolectotype">isolectotypes</typeStatus>
G-DC (G00206817 image!, U (U1476711 &amp;U1476712 image!),
</paragraph>
<paragraph id="8B8336C5281BFF81FF52E2CCFED4E2BD" blockId="4.[136,1452,1350,1480]" pageId="4" pageNumber="267">
Homotypic synonym:—
<taxonomicName id="4C3C4D46281BFF81FEABE2CCFD7AE2D1" authority="C. DC. Prodr." authorityName="C. DC. Prodr." box="[369,652,1422,1444]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="indicum">
<emphasis id="B948EAD7281BFF81FEABE2CCFE0CE2D6" box="[369,506,1422,1443]" italics="true" pageId="4" pageNumber="267">Piper indicum</emphasis>
C. DC. Prodr.
</taxonomicName>
16: 362. 1869.
<emphasis id="B948EAD7281BFF81FCC3E2CDFCCBE2D1" box="[793,829,1423,1444]" italics="true" pageId="4" pageNumber="267">non</emphasis>
Garsault Fig. Pl. Med. 3: t. 459. 1764.
<emphasis id="B948EAD7281BFF81FB66E2CDFED4E2BD" italics="true" pageId="4" pageNumber="267">nom. invalid opus utique oppresum.</emphasis>
</paragraph>
<paragraph id="8B8336C5281BFF81FF52E2B4FE62E0B5" blockId="4.[136,1452,1526,2093]" pageId="4" pageNumber="267">
<emphasis id="B948EAD7281BFF81FF52E2B4FF10E165" bold="true" box="[136,230,1526,1552]" pageId="4" pageNumber="267">Note:—</emphasis>
Miquel gave no locality but cited
<specimenCount id="9D3AFD4C281BFF81FDA8E2B5FCE8E165" box="[626,798,1526,1553]" count="2" pageId="4" pageNumber="267" type="generic">two specimens</specimenCount>
,
<emphasis id="B948EAD7281BFF81FCF1E2B4FC76E165" box="[811,896,1526,1552]" italics="true" pageId="4" pageNumber="267">Wallich</emphasis>
num. list nos. 1540 and 6649, while describing the species. Such specimens could be located at three places. The label on a sheet at U having two barcode numbers (U1476711 &amp; (U1476712) shows no. 1540 only. At
<collectionCode id="ED2DAE00281BFF81FD05E17DFD05E12C" box="[735,755,1599,1625]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
the specimen (
<collectionCode id="ED2DAE00281BFF81FC7BE17CFC43E12D" box="[929,949,1598,1624]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
001124408) bearing Wallichs annotation as
<taxonomicName id="4C3C4D46281BFF81FF52E121FEB0E109" box="[136,326,1634,1660]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="leptostachyum">
<emphasis id="B948EAD7281BFF81FF52E121FEB0E109" box="[136,326,1634,1660]" italics="true" pageId="4" pageNumber="267">P. leptostachyum</emphasis>
</taxonomicName>
has the label with both the
<emphasis id="B948EAD7281BFF81FDA7E120FD24E109" box="[637,722,1634,1660]" italics="true" pageId="4" pageNumber="267">Wallich</emphasis>
Num. list nos. 1540 and 6649. The G-DC specimen (G0020817) for
<emphasis id="B948EAD7281BFF81FF6AE1C4FEF3E1D5" box="[176,261,1670,1696]" italics="true" pageId="4" pageNumber="267">Wallich</emphasis>
Num. list no. 6649 was annotated as
<taxonomicName id="4C3C4D46281BFF81FD73E1C5FC90E1D5" box="[681,870,1670,1696]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="leptostachyum">
<emphasis id="B948EAD7281BFF81FD73E1C5FC90E1D5" box="[681,870,1670,1696]" italics="true" pageId="4" pageNumber="267">P. leptostachyum</emphasis>
</taxonomicName>
by Wallich and as
<taxonomicName id="4C3C4D46281BFF81FBE1E1C5FB2CE1D5" box="[1083,1242,1670,1696]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="indicum">
<emphasis id="B948EAD7281BFF81FBE1E1C5FB2CE1D5" box="[1083,1242,1670,1696]" italics="true" pageId="4" pageNumber="267">Piper indicum</emphasis>
</taxonomicName>
by C.de Candolle. The facts show that Hooker (1886: 95) was wrong to conclude that “Miquels citation of Wall. Cat. for this is an error.” Neither Miquel had designated a
<typeStatus id="54878867281BFF81FD8BE18CFD40E19D" box="[593,694,1742,1768]" pageId="4" pageNumber="267" type="holotype">holotype</typeStatus>
, nor any of the specimens cited above bears annotation by Miquel. Mention of
<collectionCode id="ED2DAE00281BFF81FED7E1B1FED7E078" box="[269,289,1779,1805]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
(
<collectionCode id="ED2DAE00281BFF81FEEAE1B0FEB2E079" box="[304,324,1778,1804]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
00124408) as the
<typeStatus id="54878867281BFF81FDD5E1B0FD87E079" box="[527,625,1778,1804]" pageId="4" pageNumber="267" type="holotype">holotype</typeStatus>
by Suwanphakdee
<emphasis id="B948EAD7281BFF81FC96E1B1FC73E079" box="[844,901,1778,1804]" italics="true" pageId="4" pageNumber="267">et al.</emphasis>
(2016) is an error (Art. 9.1, Turland
<emphasis id="B948EAD7281BFF81FAF9E1B1FAAAE079" box="[1315,1372,1778,1804]" italics="true" pageId="4" pageNumber="267">et al.</emphasis>
2018). So too by Mukherjee (2018) who had unnecessarily mentioned the specimen at G-DC (G00206817) as the
<typeStatus id="54878867281BFF81FA98E054FA5EE045" box="[1346,1448,1814,1840]" pageId="4" pageNumber="267" type="holotype">holotype</typeStatus>
. The best option should be to designate a
<typeStatus id="54878867281BFF81FDB9E078FD3CE021" box="[611,714,1850,1876]" pageId="4" pageNumber="267" type="lectotype">lectotype</typeStatus>
(Art. 9.2, 9.11 &amp; 9.12, Turland
<emphasis id="B948EAD7281BFF81FBE4E079FB8EE021" box="[1086,1144,1850,1876]" italics="true" pageId="4" pageNumber="267">et al.</emphasis>
2018). The specimen at
<collectionCode id="ED2DAE00281BFF81FA4DE079FA5DE020" box="[1431,1451,1851,1877]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
(
<collectionCode id="ED2DAE00281BFF81FF4AE01CFF52E00D" box="[144,164,1886,1912]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="267" type="Herbarium">K</collectionCode>
001124408) is designated here as the
<typeStatus id="54878867281BFF81FD9DE01CFD47E00D" box="[583,689,1886,1912]" pageId="4" pageNumber="267" type="lectotype">lectotype</typeStatus>
. This specimen is in best condition in preservation and labeling. The G-DC specimen is stored as
<emphasis id="B948EAD7281BFF81FE10E0C1FDBEE0E9" box="[458,584,1922,1948]" italics="true" pageId="4" pageNumber="267">
<taxonomicName id="4C3C4D46281BFF81FE10E0C1FDB2E0E9" box="[458,580,1922,1948]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="indicum">P. indicum</taxonomicName>
.
</emphasis>
The specimen at U lacks Wallichs 6649 number. Its locality is mentioned as “Nedaun, river Aatran”.
</paragraph>
<paragraph id="8B8336C5281BFF80FF67E088FA63E63C" blockId="4.[136,1452,1526,2093]" lastBlockId="5.[136,1452,158,329]" lastPageId="5" lastPageNumber="268" pageId="4" pageNumber="267">
Suwanphakdee
<emphasis id="B948EAD7281BFF81FEB4E089FE51E091" box="[366,423,1994,2020]" italics="true" pageId="4" pageNumber="267">et al.</emphasis>
(2016: 610) treated
<taxonomicName id="4C3C4D46281BFF81FD56E089FC53E091" authority="C. DC." authorityName="C. DC." box="[652,933,1994,2020]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="rhytidocarpum">
<emphasis id="B948EAD7281BFF81FD56E089FCB8E091" box="[652,846,1994,2020]" italics="true" pageId="4" pageNumber="267">P. rhytidocarpum</emphasis>
C. DC.
</taxonomicName>
and
<taxonomicName id="4C3C4D46281BFF81FC01E089FB07E091" authority="Miquel" authorityName="Miquel" box="[987,1265,1994,2020]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="leptostachyum">
<emphasis id="B948EAD7281BFF81FC01E089FB6FE091" box="[987,1177,1994,2020]" italics="true" pageId="4" pageNumber="267">P. leptostachyum</emphasis>
Miquel
</taxonomicName>
as synonymous.
<taxonomicName id="4C3C4D46281BFF81FF52E0ADFEB1EF7D" box="[136,327,2030,2056]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="leptostachyum">
<emphasis id="B948EAD7281BFF81FF52E0ADFEB1EF7D" box="[136,327,2030,2056]" italics="true" pageId="4" pageNumber="267">P. leptostachyum</emphasis>
</taxonomicName>
is characterized by oblique ovate elliptic leaves with unequal rounded base, veins 9, the upper two veins placed above 1/3 the length of the leaf are alternate; female spikes
<quantity id="4CC49B20281BFF81FC6BEF50FBC2EF58" box="[945,1076,2066,2093]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="267" unit="cm" value="17.5" valueMax="20.0" valueMin="15.0">15 to 20 cm</quantity>
long, with distantly placed fruits. Leaves in
<taxonomicName id="4C3C4D46281AFF80FEDBE7DDFE30E7CD" box="[257,454,158,184]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="rhytidocarpum">
<emphasis id="B948EAD7281AFF80FEDBE7DDFE30E7CD" box="[257,454,158,184]" italics="true" pageId="5" pageNumber="268">P. rhytidocarpum</emphasis>
</taxonomicName>
are ovate-orbicular, rounded equal base, female spikes to 30+ cm, fruits more or less closely spaced. Mukherjee (2018: 27) identified
<taxonomicName id="4C3C4D46281AFF80FD6CE781FC80E7A9" box="[694,886,194,220]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="leptostachyum">
<emphasis id="B948EAD7281AFF80FD6CE781FC80E7A9" box="[694,886,194,220]" italics="true" pageId="5" pageNumber="268">P. leptostachyum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C4D46281AFF80FC6AE781FB82E7A9" box="[944,1140,194,220]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="rhytidocarpum">
<emphasis id="B948EAD7281AFF80FC6AE781FB82E7A9" box="[944,1140,194,220]" italics="true" pageId="5" pageNumber="268">P. rhytidocarpum</emphasis>
</taxonomicName>
as distinct species and has made observations on their typification. The error in lectotypification of
<taxonomicName id="4C3C4D46281AFF80FC13E7A5FB7AE675" box="[969,1164,230,256]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="rhytidocarpum">
<emphasis id="B948EAD7281AFF80FC13E7A5FB7AE675" box="[969,1164,230,256]" italics="true" pageId="5" pageNumber="268">P. rhytidocarpum</emphasis>
</taxonomicName>
by Gilbert &amp; Xia (1994: 194) is repeated by Suwanphakdee
<emphasis id="B948EAD7281AFF80FDC1E649FDA3E651" box="[539,597,266,292]" italics="true" pageId="5" pageNumber="268">et al.</emphasis>
(2016: 610). There has been a minor error by Mukherjee (2018: 27) which needs correction; Barcode code number for the
<typeStatus id="54878867281AFF80FD47E66CFD09E63D" box="[669,767,302,328]" pageId="5" pageNumber="268" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3C4D46281AFF80FCFEE66DFC11E63D" box="[804,999,302,329]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="rhytidocarpum">
P.
<emphasis id="B948EAD7281AFF80FC9BE66CFC11E63D" box="[833,999,302,328]" italics="true" pageId="5" pageNumber="268">rhytidocarpum</emphasis>
</taxonomicName>
is (G00207056 and not (G00207656).
</paragraph>
</subSubSection>
</treatment>
</document>