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<document id="EC62C374B5698C30FF0C58556D50152B" ID-CLB-Dataset="32777" ID-DOI="10.11646/zootaxa.4282.3.2" ID-GBIF-Dataset="13966ea2-f556-4d72-917e-a1b7dd85d0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="827888" ID-ZooBank="797FDD34-9129-4EC4-8A8F-1B7DEBA77E06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500107109175" checkinUser="plazi" docAuthor="Kluge, Nikita J. &amp; Novikova, Eugenia A." docDate="2017" docId="8E2587C2FFAF68299ED4FD7C47F4F872" docLanguage="en" docName="zootaxa.4282.3.2.pdf" docOrigin="Zootaxa 4282 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anafroptilum odontostylus Kluge &amp; Novikova, 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="468" masterDocId="721CFFBAFFA568269E43FFC84520FF98" masterDocTitle="Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region" masterLastPageNumber="472" masterPageNumber="453" pageNumber="463" updateTime="1698464953366" updateUser="ExternalLinkService">
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<mods:title id="70A6A2E619671A1F097A2D752399BCA1">Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region</mods:title>
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<mods:name id="3BFBF9492DA6148F4E843AD7E472FF5E" type="personal">
<mods:role id="4A9952D24F4BE180B4160FDBA246B823">
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<mods:namePart id="3F71C6DE5655F7DFC06DDA41770B7952">Kluge, Nikita J.</mods:namePart>
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<mods:namePart id="A87EA1020A8755A77ADD83D9152B0115">Novikova, Eugenia A.</mods:namePart>
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<mods:typeOfResource id="356CA9AB8437C48FDF190513CFE0052A">text</mods:typeOfResource>
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<mods:title id="917C67C49DB0B9571459257BA0BF694C">Zootaxa</mods:title>
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<mods:date id="C23734E5FF671FDD9F7234B4E1E73375">2017</mods:date>
<mods:detail id="0CD0B8B66E45535A6D9E8D40036D219C" type="volume">
<mods:number id="FD0C1767431F7F748CB0B6DFAA93BABB">4282</mods:number>
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<mods:number id="4EF0006472A59D325D380B86B37ACB3A">3</mods:number>
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<mods:classification id="01781A77E0AE21E613A9CDDE529781FB">journal article</mods:classification>
<mods:identifier id="57D6DDB057CDF300EC2D1108523EF95B" type="CLB-Dataset">32777</mods:identifier>
<mods:identifier id="7BC504B6AEB153DB71DF1345619A3898" type="DOI">10.11646/zootaxa.4282.3.2</mods:identifier>
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<treatment id="8E2587C2FFAF68299ED4FD7C47F4F872" ID-DOI="http://doi.org/10.5281/zenodo.6028653" ID-GBIF-Taxon="132548381" ID-Zenodo-Dep="6028653" LSID="urn:lsid:plazi:treatment:8E2587C2FFAF68299ED4FD7C47F4F872" httpUri="http://treatment.plazi.org/id/8E2587C2FFAF68299ED4FD7C47F4F872" lastPageId="15" lastPageNumber="468" pageId="10" pageNumber="463">
<subSubSection id="4E96655FFFAF682C9ED4FD7C4407FD77" pageId="10" pageNumber="463" type="nomenclature">
<paragraph id="063336D4FFAF682C9ED4FD7C4703FD56" blockId="10.[151,547,692,751]" box="[151,547,692,718]" pageId="10" pageNumber="463">
<heading id="5D7B81B8FFAF682C9ED4FD7C4703FD56" bold="true" box="[151,547,692,718]" fontSize="11" level="1" pageId="10" pageNumber="463" reason="1">
<emphasis id="34F8EAC6FFAF682C9ED4FD7C4703FD56" bold="true" box="[151,547,692,718]" pageId="10" pageNumber="463">
<taxonomicName id="C18C4D57FFAF682C9ED4FD7C44FAFD56" box="[151,474,692,718]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFAF682C9ED4FD7C44FAFD56" bold="true" box="[151,474,692,718]" italics="true" pageId="10" pageNumber="463">Anafroptilum odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFAF682C9FA2FD7C4703FD56" box="[481,547,692,718]" pageId="10" pageNumber="463" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="063336D4FFAF682C9ED4FD1E4407FD77" blockId="10.[151,547,692,751]" box="[151,295,726,751]" pageId="10" pageNumber="463">
(
<figureCitation id="9EB72A51FFAF682C9EDCFD1E443FFD77" box="[159,287,726,751]" captionStart-0="FIGURES 54 – 59" captionStart-1="FIGURES 60 – 64" captionStart-2="FIGURES 65 – 68" captionStart-3="FIGURES 69 – 77" captionStart-4="FIGURE 78" captionStart-5="FIGURES 79 – 81" captionStart-6="FIGURES 82 – 85" captionStartId-0="11.[151,264,1943,1966]" captionStartId-1="12.[151,264,1198,1221]" captionStartId-2="13.[151,264,1895,1918]" captionStartId-3="14.[151,264,1940,1963]" captionStartId-4="15.[151,250,1478,1500]" captionStartId-5="16.[151,264,1498,1521]" captionStartId-6="17.[151,264,1964,1987]" captionTargetBox-0="[170,1414,193,1927]" captionTargetBox-1="[151,1436,193,1177]" captionTargetBox-2="[151,1436,193,1874]" captionTargetBox-3="[200,1394,209,1894]" captionTargetBox-4="[190,1386,217,1416]" captionTargetBox-5="[176,1371,195,1448]" captionTargetBox-6="[152,1435,769,1943]" captionTargetId-0="figure@11.[170,1414,193,1927]" captionTargetId-1="figure@12.[151,1436,193,1177]" captionTargetId-2="figure@13.[151,1436,193,1874]" captionTargetId-3="figure@14.[170,1417,193,1919]" captionTargetId-4="figure@15.[170,1417,193,1457]" captionTargetId-5="figure@16.[173,1410,185,1455]" captionTargetId-6="figure@17.[151,1436,769,1943]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="17" captionText-0="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." captionText-1="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." captionText-2="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." captionText-3="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." captionText-4="FIGURE 78. Anafroptilum odontostylus sp. n., labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." captionText-5="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." captionText-6="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri-0="https://zenodo.org/record/827904/files/figure.png" httpUri-1="https://zenodo.org/record/827906/files/figure.png" httpUri-2="https://zenodo.org/record/827908/files/figure.png" httpUri-3="https://zenodo.org/record/827910/files/figure.png" httpUri-4="https://zenodo.org/record/827912/files/figure.png" httpUri-5="https://zenodo.org/record/827914/files/figure.png" httpUri-6="https://zenodo.org/record/827916/files/figure.png" pageId="10" pageNumber="463">Figs 54–85</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAF682C9ED4FCD64012FCAF" box="[151,1330,798,823]" pageId="10" pageNumber="463" type="etymology">
<paragraph id="063336D4FFAF682C9ED4FCD64012FCAF" blockId="10.[151,1437,798,1975]" box="[151,1330,798,823]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9ED4FCD6443EFCAF" bold="true" box="[151,286,798,823]" pageId="10" pageNumber="463">Etymology.</emphasis>
Referring to denticle on inner side of the first segment of male imaginal gonostylus (
<figureCitation id="9EB72A51FFAF682C9A9BFCD6400AFCAF" box="[1240,1322,798,823]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="10" pageNumber="463">Fig. 79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAF682C9E84FC8A464CFB7F" pageId="10" pageNumber="463" type="materials_examined">
<paragraph id="063336D4FFAF682C9E84FC8A44FAFC3B" blockId="10.[151,1437,798,1975]" pageId="10" pageNumber="463">
<materialsCitation id="B6E43C89FFAF682C9E84FC8A46A1FCE7" ID-GBIF-Occurrence="1572324440" collectingDate="2015-02-09" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" county="Pai" location="Province" municipality="river Pai" pageId="10" pageNumber="463" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="34F8EAC6FFAF682C9E84FC8A4492FCC3" bold="true" box="[199,434,834,859]" pageId="10" pageNumber="463">Material examined.</emphasis>
<typeStatus id="D9378876FFAF682C9FF8FC8A4708FCC3" box="[443,552,834,859]" pageId="10" pageNumber="463" type="holotype">Holotype</typeStatus>
: L-S-I
<specimenCount id="108AFD5DFFAF682C9C30FC8B47A6FCC3" box="[627,646,835,859]" pageId="10" pageNumber="463" type="male">♂</specimenCount>
{specimen [XIII] (1) 2015},
<collectingCountry id="7E9B7644FFAF682C9D99FC8B414EFCC3" box="[986,1134,835,859]" name="Thailand" pageId="10" pageNumber="463">THAILAND</collectingCountry>
, Mae-Hong-Son
<location id="0353600FFFAF682C9B72FC8A40B7FCC3" LSID="urn:lsid:plazi:treatment:8E2587C2FFAF68299ED4FD7C47F4F872:0353600FFFAF682C9B72FC8A40B7FCC3" box="[1329,1431,834,859]" country="Thailand" county="Pai" municipality="river Pai" name="Province" pageId="10" pageNumber="463">Province</location>
,
<collectingMunicipality id="E657ACAEFFAF682C9ED4FCAE45D9FCE7" box="[151,249,870,895]" pageId="10" pageNumber="463">river Pai</collectingMunicipality>
upstream of
<collectingCounty id="EF524E58FFAF682C9FD1FCAE4498FCE7" box="[402,440,870,895]" pageId="10" pageNumber="463">Pai</collectingCounty>
,
<collectingDate id="6276E9FCFFAF682C9F85FCAE470FFCE7" box="[454,559,870,895]" pageId="10" pageNumber="463" value="2015-02-09">
<date id="72321014FFAF682C9F85FCAE470FFCE7" box="[454,559,870,895]" pageId="10" pageNumber="463" value="2015-02-09">9.II.2015</date>
</collectingDate>
, coll.
<collectorName id="AB795302FFAF682C9C34FCAF47C0FCE7" box="[631,736,870,895]" pageId="10" pageNumber="463">N. Kluge</collectorName>
&amp;
<collectorName id="AB795302FFAF682C9D44FCAF46A1FCE7" box="[775,897,870,895]" pageId="10" pageNumber="463">L. Sheyko</collectorName>
</materialsCitation>
.
<materialsCitation id="B6E43C89FFAF682C9DCEFCAF44F7FC3B" ID-GBIF-Occurrence="1572324442" collectingDate="2015-02-08" collectorName="N. Kluge &amp; L. Sheyko" country="Thailand" county="Pai" location="Province" municipality="river Pai" pageId="10" pageNumber="463" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="D9378876FFAF682C9DCEFCAF46D3FCE7" box="[909,1011,871,895]" pageId="10" pageNumber="463" type="paratype">Paratype</typeStatus>
: the same locality,
<collectingDate id="6276E9FCFFAF682C9A93FCAE4019FCE7" box="[1232,1337,870,895]" pageId="10" pageNumber="463" value="2015-02-08">
<date id="72321014FFAF682C9A93FCAE4019FCE7" box="[1232,1337,870,895]" pageId="10" pageNumber="463" value="2015-02-08">8.II.2015</date>
</collectingDate>
, coll.
<collectorName id="AB795302FFAF682C9BC2FCAF45FBFC3B" pageId="10" pageNumber="463">N. Kluge</collectorName>
&amp;
<collectorName id="AB795302FFAF682C9EBCFC434456FC3B" box="[255,374,906,931]" pageId="10" pageNumber="463">L. Sheyko</collectorName>
: L-S-I
<specimenCount id="108AFD5DFFAF682C9F80FC4344F7FC3B" box="[451,471,907,931]" pageId="10" pageNumber="463" type="male">♂</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="063336D4FFAF682C9E84FC66464CFB7F" blockId="10.[151,1437,798,1975]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9E84FC6647EEFC5F" bold="true" box="[199,718,942,967]" pageId="10" pageNumber="463">
Descriptions.
<emphasis id="34F8EAC6FFAF682C9F31FC67449DFC5E" bold="true" box="[370,445,943,966]" italics="true" pageId="10" pageNumber="463">Larva.</emphasis>
Cuticular coloration.
</emphasis>
Head with light brown and colorless areas. Pronotum and mesonotum with light brown and colorless areas; protoptera light brownish with colorless stripes corresponding to longitudinal veins (
<figureCitation id="9EB72A51FFAF682C9F33FC3E44E5FB97" box="[368,453,1014,1039]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="10" pageNumber="463">Fig. 56</figureCitation>
). Metanotum and thoracic pleura light brown, thoracic sterna and membranes colorless. Legs nearly colorless; apex of femur, base of tibia and outer side of tarsus diffusively tinged with light brownish. Abdomen with light brown and colorless areas identical in two examined individuals (
<figureCitation id="9EB72A51FFAF682C9A39FBF641F0FBCF" box="[1146,1232,1086,1111]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="10" pageNumber="463">Fig. 54</figureCitation>
): terga II and VII with most extensive brown coloration, terga III and V with light median area, other terga lighter; abdominal sterna at most part colorless, with anterior margins brown. Cerci at most part light brownish, colorless at base, with posterior margin of each 4th segment slightly darkened; paracercus lighter than cerci (
<figureCitation id="9EB72A51FFAF682C9A3BFB6241EFFB5B" box="[1144,1231,1194,1219]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="10" pageNumber="463">Fig. 54</figureCitation>
). Each swimming seta of cerci and paracercus brown proximally, colorless distally.
</paragraph>
</subSubSection>
<subSubSection id="4E96655FFFAF68299E84FB3A47F4F872" lastPageId="15" lastPageNumber="468" pageId="10" pageNumber="463" type="description">
<paragraph id="063336D4FFAF682C9E84FB3A465FFA93" blockId="10.[151,1437,798,1975]" box="[199,895,1266,1291]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9E84FB3A44C5FA93" bold="true" box="[199,485,1266,1291]" pageId="10" pageNumber="463">Hypodermal coloration.</emphasis>
Unknown (probably not expressed).
</paragraph>
<paragraph id="063336D4FFAF682C9E84FADE4741F923" blockId="10.[151,1437,798,1975]" pageId="10" pageNumber="463">
<emphasis id="34F8EAC6FFAF682C9E84FADE441FFAB7" bold="true" box="[199,319,1302,1327]" pageId="10" pageNumber="463">Structure.</emphasis>
Frons between antennae with flat elevation; frontal suture relative long and obtuse-angled (as in
<figureCitation id="9EB72A51FFAF682C9ED4FAF245CAFACB" box="[151,234,1338,1363]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Fig. 37</figureCitation>
). Labrum of usual shape; dorsal surface with irregularly situated setae [without constant pair of submedian setae and/or pair of latero-distal setal rows – see
<taxonomicName id="C18C4D57FFAF682C9C83FA964674FAEE" box="[704,852,1374,1398]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9C83FA964674FAEE" box="[704,852,1374,1398]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(9)] (
<figureCitation id="9EB72A51FFAF682C9DD6FA9646CDFAEF" box="[917,1005,1374,1399]" captionStart="FIGURES 60 – 64" captionStartId="12.[151,264,1198,1221]" captionTargetBox="[151,1436,193,1177]" captionTargetId="figure@12.[151,1436,193,1177]" captionTargetPageId="12" captionText="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="10" pageNumber="463">Figs 60</figureCitation>
,
<figureCitation id="9EB72A51FFAF682C9DBAFA964139FAEF" box="[1017,1049,1374,1399]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="10" pageNumber="463">76</figureCitation>
). Both mandibles with incisor and kinetodontium deeply separated [see
<taxonomicName id="C18C4D57FFAF682C9C20FA4B47D7FA03" box="[611,759,1411,1435]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9C20FA4B47D7FA03" box="[611,759,1411,1435]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(10)]; planes of incisor, kinetodontium and prostheca perpendicular to plane of mandible (
<figureCitation id="9EB72A51FFAF682C9C71FA6E4795FA27" box="[562,693,1446,1471]" captionStart="FIGURES 60 – 64" captionStartId="12.[151,264,1198,1221]" captionTargetBox="[151,1436,193,1177]" captionTargetId="figure@12.[151,1436,193,1177]" captionTargetPageId="12" captionText="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="10" pageNumber="463">Figs 61–62</figureCitation>
); denticles of incisors, kinetodontia and prosthecae similar to that of
<taxonomicName id="C18C4D57FFAF682C9EFAFA034478FA7B" box="[185,344,1483,1507]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="species" species="orthostylus">
<emphasis id="34F8EAC6FFAF682C9EFAFA034478FA7B" box="[185,344,1483,1507]" italics="true" pageId="10" pageNumber="463">A. orthostylus</emphasis>
</taxonomicName>
(as in
<figureCitation id="9EB72A51FFAF682C9FEEFA024712FA7B" box="[429,562,1482,1507]" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Figs 38–39</figureCitation>
). Molar processes of left mandible as long as distal molar projection; each process terminated by short apical lamella (
<figureCitation id="9EB72A51FFAF682C9C3FFA2647F0F99F" box="[636,720,1518,1543]" captionStart="FIGURES 60 – 64" captionStartId="12.[151,264,1198,1221]" captionTargetBox="[151,1436,193,1177]" captionTargetId="figure@12.[151,1436,193,1177]" captionTargetPageId="12" captionText="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="10" pageNumber="463">Fig. 63</figureCitation>
: al). Maxilla with 3 canines, 3 dentisetae, 4–5 setae proximad of dentisetae and 9–10 setae of inner ventral row; all canines, dentisetae and setae slender, pointed and bent at the same direction (
<figureCitation id="9EB72A51FFAF682C9F10F9FE4489F9D7" box="[339,425,1590,1615]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="10" pageNumber="463">Fig. 77</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAF682C9FB6F9FE47A9F9D6" box="[501,649,1590,1614]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9FB6F9FE47A9F9D6" box="[501,649,1590,1614]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(12)]. Maxillary palp 3-segmented, all three segments of subequal length (
<figureCitation id="9EB72A51FFAF682C9EAFF992441FF9EB" box="[236,319,1626,1651]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="10" pageNumber="463">Fig. 77</figureCitation>
). Labium with glossae and paraglossae not narrowed near bases; setae of glossa not shortened; setae of dorso-median row of paraglossa located near median margin; labial palp with 3rd segment widened apically and truncate (
<figureCitation id="9EB72A51FFAF682C9F43F96A4472F923" box="[256,338,1698,1723]" captionStart="FIGURE 78" captionStartId="15.[151,250,1478,1500]" captionTargetBox="[190,1386,217,1416]" captionTargetId="figure@15.[170,1417,193,1457]" captionTargetPageId="15" captionText="FIGURE 78. Anafroptilum odontostylus sp. n., labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa." httpUri="https://zenodo.org/record/827912/files/figure.png" pageId="10" pageNumber="463">Fig. 78</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAF682C9FD6F96B4709F923" box="[405,553,1699,1723]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9FD6F96B4709F923" box="[405,553,1699,1723]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(2)].
</paragraph>
<paragraph id="063336D4FFAF682C9E84F90E416DF82F" blockId="10.[151,1437,798,1975]" pageId="10" pageNumber="463">
Vestiges of hind protoptera represented by blunt projections. Legs as in A.
<taxonomicName id="C18C4D57FFAF682C9A7FF90E419DF946" box="[1084,1213,1734,1758]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="species" species="kazlauskasi">
<emphasis id="34F8EAC6FFAF682C9A7FF90E419DF946" box="[1084,1213,1734,1758]" italics="true" pageId="10" pageNumber="463">kazlauskasi</emphasis>
</taxonomicName>
(
<bibRefCitation id="621D4B25FFAF682C9A93F90E45C2F89B" author="Kluge" pageId="10" pageNumber="463" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2012">
Kluge 2012:
<figureCitation id="9EB72A51FFAF682C9B28F90E45C2F89B" captionStart="FIGURES 37 – 43" captionStartId="6.[151,264,1794,1817]" captionTargetBox="[201,1383,200,1772]" captionTargetId="figure@6.[201,1395,193,1781]" captionTargetPageId="6" captionText="FIGURES 37 – 43. Anafroptilum orthostylus sp. n. 37, part of eXuviae of larval head with labrum, bases of antennae and frontal suture; 38 – 39, incisor, kinetodontium and prostheca of left and right mandibles, view from base of mandible; 40, paraproct; 41, genitals of male imago (on the left half, gonovectal muscles and hidden part of penis shown by interrupted lines, gonostylar muscle not shown; at right half gonostylar muscle shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 42, eXuviae of gonostyli of subimago; 43, larval protogonostyli and subimaginal gonostyli developing under larval cuticle (40, 42, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; in, incisor; kd, kinetodontoum; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); prs, prostheca; usg, unistyliger." httpUri="https://zenodo.org/record/827898/files/figure.png" pageId="10" pageNumber="463">Figs 38–40</figureCitation>
</bibRefCitation>
): femora nearly parallel-sided, fore femur widest at middle, middle and hind femora widest at apex; apex of each femur finely serrate (
<figureCitation id="9EB72A51FFAF682C9F86F8C6473BF8BF" box="[453,539,1806,1831]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="10" pageNumber="463">Fig. 59</figureCitation>
); each femur without stout setae on outer margin and apex; only inner margin with few small stout setae [see
<taxonomicName id="C18C4D57FFAF682C9C40F8FB47B7F8D3" box="[515,663,1843,1867]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9C40F8FB47B7F8D3" box="[515,663,1843,1867]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(14)]; fore tibia without patella-tibial suture, middle and hind tibia with patella-tibial suture [see
<taxonomicName id="C18C4D57FFAF682C9FA0F89E4757F8F6" box="[483,631,1878,1902]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAF682C9FA0F89E4757F8F6" box="[483,631,1878,1902]" italics="true" pageId="10" pageNumber="463">Anafroptilum</emphasis>
</taxonomicName>
(1)]; each tibia with short ark of fine setae at outer side near base; outer side of tibia and tarsus without stout setae, inner side of tibia and tarsus with sparse spine-like setae. Claw with two equal regular rows of small denticles, reaching less than 1/2 of claw length (
<figureCitation id="9EB72A51FFAF682C9DAFF8564160F82F" box="[1004,1088,1950,1975]" captionStart="FIGURES 60 – 64" captionStartId="12.[151,264,1198,1221]" captionTargetBox="[151,1436,193,1177]" captionTargetId="figure@12.[151,1436,193,1177]" captionTargetPageId="12" captionText="FIGURES 60 – 64. Anafroptilum odontostylus sp. n., larva. 60, labrum; 61 – 62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection." httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="10" pageNumber="463">Fig. 64</figureCitation>
).
</paragraph>
<caption id="52F3665CFFAE682D9ED4F85F4618F873" httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="11" pageNumber="464" targetBox="[170,1414,193,1927]" targetPageId="11">
<paragraph id="063336D4FFAE682D9ED4F85F4618F873" blockId="11.[151,1436,1943,2027]" pageId="11" pageNumber="464">
<emphasis id="34F8EAC6FFAE682D9ED4F85F4473F835" bold="true" box="[151,339,1943,1966]" pageId="11" pageNumber="464">FIGURES 54–59.</emphasis>
<taxonomicName id="C18C4D57FFAE682D9F1AF8504745F836" box="[345,613,1944,1966]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="464" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFAE682D9F1AF8504745F836" box="[345,613,1944,1966]" italics="true" pageId="11" pageNumber="464">Anafroptilum odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFAE682D9C28F8504783F835" box="[619,675,1944,1965]" pageId="11" pageNumber="464" rank="species">
<emphasis id="34F8EAC6FFAE682D9C28F8504783F835" bold="true" box="[619,675,1944,1965]" pageId="11" pageNumber="464">sp. n.</emphasis>
</taxonomicNameLabel>
, larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54–55, 57–59, holotype).
</paragraph>
</caption>
<caption id="52F3665CFFA9682A9ED4FB664052FB7B" httpUri="https://zenodo.org/record/827906/files/figure.png" pageId="12" pageNumber="465" targetBox="[151,1436,193,1177]" targetPageId="12">
<paragraph id="063336D4FFA9682A9ED4FB664052FB7B" blockId="12.[151,1436,1198,1251]" pageId="12" pageNumber="465">
<emphasis id="34F8EAC6FFA9682A9ED4FB664478FB5C" bold="true" box="[151,344,1198,1221]" pageId="12" pageNumber="465">FIGURES 60–64.</emphasis>
<taxonomicName id="C18C4D57FFA9682A9F21FB674753FB5D" box="[354,627,1199,1221]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="465" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA9682A9F21FB674753FB5D" box="[354,627,1199,1221]" italics="true" pageId="12" pageNumber="465">Anafroptilum odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA9682A9C3EFB67479BFB5C" box="[637,699,1199,1220]" pageId="12" pageNumber="465" rank="species">
<emphasis id="34F8EAC6FFA9682A9C3EFB67479BFB5C" bold="true" box="[637,699,1199,1220]" pageId="12" pageNumber="465">sp. n.</emphasis>
</taxonomicNameLabel>
, larva. 60, labrum; 61–62, left and right mandibles; 63, mola of left mandible; 64, claw (60, 61, 64, holotype). Abbreviations: al, apical lamella of molar process; dmp, distal molar projection.
</paragraph>
</caption>
<paragraph id="063336D4FFA9682A9E84FAC345F6F883" blockId="12.[151,1437,1291,2036]" pageId="12" pageNumber="465">
Abdominal terga and sterna with small denticles and sparse narrow scales (
<figureCitation id="9EB72A51FFA9682A9A4EFAC341B0FABC" box="[1037,1168,1291,1316]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="12" pageNumber="465">Figs 65–68</figureCitation>
); sterna with irregularly situated simple long setae (
<figureCitation id="9EB72A51FFA9682A9F86FAE64739FADF" box="[453,537,1326,1351]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="12" pageNumber="465">Fig. 67</figureCitation>
). Posterior margin of abdominal tergum I without denticles; posterior margins of terga II–VIII with blunt brown denticles (
<figureCitation id="9EB72A51FFA9682A9C2FFA9B47E0FAF4" box="[620,704,1363,1388]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="12" pageNumber="465">Fig. 65</figureCitation>
); posterior margin of tergum XI with blunt, rounded and pointed denticles, without denticles behind pair of submedian pores (
<figureCitation id="9EB72A51FFA9682A9D0DFABE4684FA17" box="[846,932,1398,1423]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="12" pageNumber="465">Fig. 66</figureCitation>
); tergum X with pointed denticles (
<figureCitation id="9EB72A51FFA9682A9B79FABE40AFFA17" box="[1338,1423,1398,1423]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="12" pageNumber="465">Fig. 58</figureCitation>
). Posterior margin of abdominal sterna I–IV without denticles; posterior margin of sterna V–VIII with blunt and split denticles (
<figureCitation id="9EB72A51FFA9682A9F4DFA764445FA4F" box="[270,357,1470,1495]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="12" pageNumber="465">Fig. 67</figureCitation>
); posterior margin of sternum IX of mature male larva with prominent semicircular smooth protogonostyli and dense stick-like denticles between them (
<figureCitation id="9EB72A51FFA9682A9D0CFA2B4685FA64" box="[847,933,1507,1532]" captionStart="FIGURES 65 – 68" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[151,1436,193,1874]" captionTargetId="figure@13.[151,1436,193,1874]" captionTargetPageId="13" captionText="FIGURES 65 – 68. Anafroptilum odontostylus sp. n., eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX." httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="12" pageNumber="465">Fig. 68</figureCitation>
). Paraprocts with pointed denticles on apex and median side (
<figureCitation id="9EB72A51FFA9682A9F23F9CE4494F987" box="[352,436,1542,1567]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="12" pageNumber="465">Fig. 81</figureCitation>
). Lateral denticles either absent, or present only on distal part of abdominal segment IX (
<figureCitation id="9EB72A51FFA9682A9EDCF9E345D2F9DC" box="[159,242,1579,1604]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="12" pageNumber="465">Fig. 57</figureCitation>
) [see
<taxonomicName id="C18C4D57FFA9682A9F79F9E344EEF9DB" box="[314,462,1579,1603]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA9682A9F79F9E344EEF9DB" box="[314,462,1579,1603]" italics="true" pageId="12" pageNumber="465">Anafroptilum</emphasis>
</taxonomicName>
(3)]. Each tergalius I–VII with long costal and anal marginal ribs, without denticles, distal margin without rib; tergalius I asymmetrically widened in distal part; tergalius II asymmetrically widened in midlength; tergalii II–VI asymmetrically widened in proximal part; tergalius VII nearly symmetrical (
<figureCitation id="9EB72A51FFA9682A9B4EF9BB40B0F914" box="[1293,1424,1651,1676]" captionStart="FIGURES 69 – 77" captionStartId="14.[151,264,1940,1963]" captionTargetBox="[200,1394,209,1894]" captionTargetId="figure@14.[170,1417,193,1919]" captionTargetPageId="14" captionText="FIGURES 69 – 77. Anafroptilum odontostylus sp. n., larvae. 69 – 75, tergalii I – VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c 1, c 2, c 3, maXillary canines; ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="12" pageNumber="465">Figs 69–75</figureCitation>
). Cerci with small denticles on all segments; one simple or composed larger spine can be present on lateral side of cercus near tip; among two examined individuals, the
<typeStatus id="D9378876FFA9682A9D51F9734654F94C" box="[786,884,1723,1748]" pageId="12" pageNumber="465" type="holotype">holotype</typeStatus>
has double spine on each cercus (
<figureCitation id="9EB72A51FFA9682A9B49F9734041F94C" box="[1290,1377,1723,1748]" captionStart="FIGURES 54 – 59" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[170,1414,193,1927]" captionTargetId="figure@11.[170,1414,193,1927]" captionTargetPageId="11" captionText="FIGURES 54 – 59. Anafroptilum odontostylus sp. n., larva. 54, eXuviae of abdominal sterna (left) and terga (right); 55, enlarged portion of cercus; 56, eXuviae of left half of pronotum and mesonotum; 57, lateral margin of IX abdominal segment; 58, abdominal tergum X; 59, apeX of femur (54 – 55, 57 – 59, holotype)." httpUri="https://zenodo.org/record/827904/files/figure.png" pageId="12" pageNumber="465">Fig. 55</figureCitation>
); the
<typeStatus id="D9378876FFA9682A9ED4F92845D7F960" box="[151,247,1760,1784]" pageId="12" pageNumber="465" type="paratype">paratype</typeStatus>
has single spine on left cercus and vestigial spine on right cercus. Outer side of cercus without additional setae.
</paragraph>
<paragraph id="063336D4FFA9682A9E84F8EE465CF857" blockId="12.[151,1437,1291,2036]" pageId="12" pageNumber="465">
<emphasis id="34F8EAC6FFA9682A9E84F8EE4767F8A7" bold="true" box="[199,583,1830,1855]" pageId="12" pageNumber="465">
<emphasis id="34F8EAC6FFA9682A9E84F8EE441CF8A7" bold="true" box="[199,316,1830,1855]" italics="true" pageId="12" pageNumber="465">Subimago</emphasis>
. Cuticular coloration.
</emphasis>
Head colorless. Pronotum and prosternum light brown. Mesonotum nearly colorless with brown microtrichia, certain sutures light brown (
<figureCitation id="9EB72A51FFA9682A9D2EF88346E3F8FB" box="[877,963,1867,1892]" captionStart="FIGURES 82 – 85" captionStartId="17.[151,264,1964,1987]" captionTargetBox="[152,1435,769,1943]" captionTargetId="figure@17.[151,1436,769,1943]" captionTargetPageId="17" captionText="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri="https://zenodo.org/record/827916/files/figure.png" pageId="12" pageNumber="465">Fig. 83</figureCitation>
). Thoracic pleura and sterna at most part nearly colorless with brown microtrichia, certain sclerites light brow (
<figureCitation id="9EB72A51FFA9682A9DF6F8A6412BF81F" box="[949,1035,1902,1927]" captionStart="FIGURES 82 – 85" captionStartId="17.[151,264,1964,1987]" captionTargetBox="[152,1435,769,1943]" captionTargetId="figure@17.[151,1436,769,1943]" captionTargetPageId="17" captionText="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri="https://zenodo.org/record/827916/files/figure.png" pageId="12" pageNumber="465">Fig. 84</figureCitation>
). Legs nearly colorless with brown microtrichia. Abdominal terga and sterna nearly colorless with brown microtrichia; terga with very light brownish sublateral markings. Cerci very light brownish with brown spines.
</paragraph>
<paragraph id="063336D4FFA9682A9E84F81341E9F86B" blockId="12.[151,1437,1291,2036]" box="[199,1225,2011,2036]" pageId="12" pageNumber="465">
<emphasis id="34F8EAC6FFA9682A9E84F8134408F86C" bold="true" box="[199,296,2011,2036]" pageId="12" pageNumber="465">Texture.</emphasis>
On all legs all tarsomeres covered by pointed microlepides [see
<taxonomicName id="C18C4D57FFA9682A9DBEF81341B1F86B" box="[1021,1169,2011,2035]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFA9682A9DBEF81341B1F86B" box="[1021,1169,2011,2035]" italics="true" pageId="12" pageNumber="465">Anafroptilum</emphasis>
</taxonomicName>
(7)].
</paragraph>
<caption id="52F3665CFFA8682B9ED4F8AF4152F823" httpUri="https://zenodo.org/record/827908/files/figure.png" pageId="13" pageNumber="466" targetBox="[151,1436,193,1874]" targetPageId="13">
<paragraph id="063336D4FFA8682B9ED4F8AF4152F823" blockId="13.[151,1436,1895,1979]" pageId="13" pageNumber="466">
<emphasis id="34F8EAC6FFA8682B9ED4F8AF4475F8E5" bold="true" box="[151,341,1895,1918]" pageId="13" pageNumber="466">FIGURES 65–68.</emphasis>
<taxonomicName id="C18C4D57FFA8682B9F1EF8A0474BF8E6" box="[349,619,1896,1918]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="466" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFA8682B9F1EF8A0474BF8E6" box="[349,619,1896,1918]" italics="true" pageId="13" pageNumber="466">Anafroptilum odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFA8682B9C30F8A0478DF8E5" box="[627,685,1896,1917]" pageId="13" pageNumber="466" rank="species">
<emphasis id="34F8EAC6FFA8682B9C30F8A0478DF8E5" bold="true" box="[627,685,1896,1917]" pageId="13" pageNumber="466">sp. n.</emphasis>
</taxonomicNameLabel>
, eXuviae of larval abdomen. 65, median part of tergum III; 66, posterior part of tergum IX; 67, median part of sternum VIII; 68, posterior part of sternum IX of male with protogonostyli. Arrows show pairs of submedian pores; abbreviations: sVIII, sIX, sterna VIII and IX; tIII, tIX, terga III and IX.
</paragraph>
</caption>
<caption id="52F3665CFFAB68289ED4F85C4156F851" httpUri="https://zenodo.org/record/827910/files/figure.png" pageId="14" pageNumber="467" targetBox="[200,1394,209,1894]" targetPageId="14">
<paragraph id="063336D4FFAB68289ED4F85C4156F851" blockId="14.[151,1436,1940,1993]" pageId="14" pageNumber="467">
<emphasis id="34F8EAC6FFAB68289ED4F85C4474F832" bold="true" box="[151,340,1940,1963]" pageId="14" pageNumber="467">FIGURES 69–77.</emphasis>
<taxonomicName id="C18C4D57FFAB68289F19F85C4747F832" box="[346,615,1940,1962]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="467" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFAB68289F19F85C4747F832" box="[346,615,1940,1962]" italics="true" pageId="14" pageNumber="467">Anafroptilum odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFAB68289C2EF85D4787F832" box="[621,679,1941,1962]" pageId="14" pageNumber="467" rank="species">
<emphasis id="34F8EAC6FFAB68289C2EF85D4787F832" bold="true" box="[621,679,1941,1962]" pageId="14" pageNumber="467">sp. n.</emphasis>
</taxonomicNameLabel>
, larvae. 69–75, tergalii I–VII; 76, labrum and part of clypeus; 77, maXilla, dorsal view (76, holotype). Abbreviations: c1, c2, c3, maXillary canines; ds1, ds2, ds3, dentisetae.
</paragraph>
</caption>
<caption id="52F3665CFFAA68299ED4FA0E445BFA62" httpUri="https://zenodo.org/record/827912/files/figure.png" pageId="15" pageNumber="468" targetBox="[190,1386,217,1416]" targetPageId="15">
<paragraph id="063336D4FFAA68299ED4FA0E445BFA62" blockId="15.[151,1436,1478,1530]" pageId="15" pageNumber="468">
<emphasis id="34F8EAC6FFAA68299ED4FA0E4401FA43" bold="true" box="[151,289,1478,1500]" pageId="15" pageNumber="468">FIGURE 78.</emphasis>
<taxonomicName id="C18C4D57FFAA68299F6BFA0E4716FA44" box="[296,566,1478,1500]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="468" phylum="Arthropoda" rank="species" species="odontostylus" status="sp. nov.">
<emphasis id="34F8EAC6FFAA68299F6BFA0E4716FA44" box="[296,566,1478,1500]" italics="true" pageId="15" pageNumber="468">Anafroptilum odontostylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FCB57BDFFAA68299C7EFA0F4758FA44" box="[573,632,1479,1500]" pageId="15" pageNumber="468" rank="species">
<emphasis id="34F8EAC6FFAA68299C7EFA0F4758FA44" bold="true" box="[573,632,1479,1500]" pageId="15" pageNumber="468">sp. n.</emphasis>
</taxonomicNameLabel>
, labium, ventral view (left) and dorsal view (right). Abbreviation: mr, median row of setae of glossa.
</paragraph>
</caption>
<paragraph id="063336D4FFAA68299E84F9EB464FF977" blockId="15.[151,1437,1570,2027]" pageId="15" pageNumber="468">
<emphasis id="34F8EAC6FFAA68299E84F9EB4479F9A3" bold="true" box="[199,345,1570,1595]" italics="true" pageId="15" pageNumber="468">Imago, male</emphasis>
(
<figureCitation id="9EB72A51FFAA68299F2AF9EA449FF9A3" box="[361,447,1570,1595]" captionStart="FIGURES 82 – 85" captionStartId="17.[151,264,1964,1987]" captionTargetBox="[152,1435,769,1943]" captionTargetId="figure@17.[151,1436,769,1943]" captionTargetPageId="17" captionText="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri="https://zenodo.org/record/827916/files/figure.png" pageId="15" pageNumber="468">Fig. 82</figureCitation>
). Turbinate eyes with stem light yellow basally, grayish-white distally; facetted surface light brownish. Thorax light brownish with ocher. Legs pale ocher. In
<typeStatus id="D9378876FFAA68299DECF98E4134F9C7" box="[943,1044,1606,1631]" pageId="15" pageNumber="468" type="holotype">holotype</typeStatus>
, proportion of femur / tibia / tarsal segments of fore leg 70: 115: 3: 56: 42: 29: 14; middle leg 56: 56: 20: 7: 3: 12; hind leg 56: 50: 17: 6: 3: 12. Tarsus of middle and hind leg with one apical spine on 2nd tarsomere (primary 3rd tarsomere). Fore wing with membrane and veins colorless; pterostigma with 2–3 simple oblique veins. Hind wings absent. Abdomen nearly colorless, VII–X segments pale ocher. Cerci unicolor light ocher.
</paragraph>
<paragraph id="063336D4FFAA68299E84F9324763F83B" blockId="15.[151,1437,1570,2027]" pageId="15" pageNumber="468">
Unistyligers and gonostyli colorless (
<figureCitation id="9EB72A51FFAA68299C39F93247F6F88B" box="[634,726,1786,1811]" captionStart="FIGURES 82 – 85" captionStartId="17.[151,264,1964,1987]" captionTargetBox="[152,1435,769,1943]" captionTargetId="figure@17.[151,1436,769,1943]" captionTargetPageId="17" captionText="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri="https://zenodo.org/record/827916/files/figure.png" pageId="15" pageNumber="468">Figs 85</figureCitation>
). Unistyligers nearly meet together by their bases; styligeral muscle wide, attached to bases of unistyligers (
<figureCitation id="9EB72A51FFAA68299CFDF8D64636F8AF" box="[702,790,1822,1847]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="15" pageNumber="468">Fig. 79</figureCitation>
). Finger-like sclerotized projection between unistyligers (
<figureCitation id="9EB72A51FFAA68299EDCF88A4403F8C3" box="[159,291,1858,1883]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="15" pageNumber="468">Fig. 79, 80</figureCitation>
,
<figureCitation id="9EB72A51FFAA68299F72F88A4470F8C3" box="[305,336,1858,1883]" captionStart="FIGURES 82 – 85" captionStartId="17.[151,264,1964,1987]" captionTargetBox="[152,1435,769,1943]" captionTargetId="figure@17.[151,1436,769,1943]" captionTargetPageId="17" captionText="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri="https://zenodo.org/record/827916/files/figure.png" pageId="15" pageNumber="468">85</figureCitation>
). 1st segment of gonostylus with prominent projection on inner side; 2nd segment arched; 3rd segment relatively large (
<figureCitation id="9EB72A51FFAA68299FF6F8AE472DF8E7" box="[437,525,1894,1919]" captionStart="FIGURES 79 – 81" captionStartId="16.[151,264,1498,1521]" captionTargetBox="[176,1371,195,1448]" captionTargetId="figure@16.[173,1410,185,1455]" captionTargetPageId="16" captionText="FIGURES 79 – 81. Anafroptilum odontostylus sp. n. 79, genitals of male imago; 80, gonostyli of male subimago; 81, larval paraproct (79 – 80, holotype). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/827914/files/figure.png" pageId="15" pageNumber="468">Figs 79</figureCitation>
,
<figureCitation id="9EB72A51FFAA68299C5AF8AE4716F8E7" box="[537,566,1894,1919]" captionStart="FIGURES 82 – 85" captionStartId="17.[151,264,1964,1987]" captionTargetBox="[152,1435,769,1943]" captionTargetId="figure@17.[151,1436,769,1943]" captionTargetPageId="17" captionText="FIGURES 82 – 85. Anafroptilum odontostylus sp. n. 82, male imago; 83, right half of eXuviae of subimaginal mesonotum; 84, eXuviae of subimaginal pleuron; 85, genitals of male imago (85, holotype)." httpUri="https://zenodo.org/record/827916/files/figure.png" pageId="15" pageNumber="468">85</figureCitation>
) [see
<taxonomicName id="C18C4D57FFAA68299C3FF8AE4631F8E6" box="[636,785,1894,1918]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="34F8EAC6FFAA68299C3FF8AE4631F8E6" box="[636,785,1894,1918]" italics="true" pageId="15" pageNumber="468">Anafroptilum</emphasis>
</taxonomicName>
(15)]. Penis brown; external part of penis wide roundishtriangular, with apical margin smooth.
</paragraph>
<paragraph id="063336D4FFAA68299E84F865472FF85F" blockId="15.[151,1437,1570,2027]" box="[199,527,1965,1991]" pageId="15" pageNumber="468">
<emphasis id="34F8EAC6FFAA68299E84F86544B2F85E" bold="true" box="[199,402,1965,1990]" italics="true" pageId="15" pageNumber="468">Female and eggs.</emphasis>
Unknown.
</paragraph>
<paragraph id="063336D4FFAA68299E84F81A47F4F872" blockId="15.[151,1437,1570,2027]" box="[199,724,2002,2027]" pageId="15" pageNumber="468">
<emphasis id="34F8EAC6FFAA68299E84F81A446EF873" bold="true" box="[199,334,2002,2027]" pageId="15" pageNumber="468">Dimension.</emphasis>
Fore wing length of male 3.5 mm.
</paragraph>
</subSubSection>
</treatment>
</document>
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