287 lines
43 KiB
XML
287 lines
43 KiB
XML
<document id="829AFBF7BE0200DCC844D8926F2E8E65" ENCODING="UTF-8" ID-GBIF-Dataset="c0839ce9-2c24-405d-a4f7-bce70961357d" ModsDocID="z01343p055" checkinTime="1247131265830" checkinUser="thomas" docAuthor="Edward O. Murdy" docDate="2006" docId="92B743395B402090DE349E296CB71769" docLanguage="en" docName="2006_Murdy_gg1.xml" docOrigin="Zootaxa 1343" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:A2589DD6-BC0D-43DA-BFEA-A1E6BF042605" docTitle="Trypauchen Valenciennes in Cuvier & Valenciennes 1837" docType="treatment" docVersion="11" lastPageNumber="61" masterDocId="487E3E769909AC32A06D7ED6FD64B35B" masterDocTitle="A revision of the gobiid fish genus Trypauchen (Gobiidae: Amblyopinae)." masterLastPageNumber="68" masterPageNumber="55" pageNumber="57" updateTime="1701314493061" updateUser="plazi">
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<mods:title id="313EA256692722B4D28B97FECD83CE91">A revision of the gobiid fish genus Trypauchen (Gobiidae: Amblyopinae).</mods:title>
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<mods:namePart id="62657A5D6B1ACF4A2871CD18AE1945DB">Edward O. Murdy</mods:namePart>
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<mods:date id="35C0DC943DC0E880C35930B8BEA468DC">2006</mods:date>
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<treatment id="92B743395B402090DE349E296CB71769" ID-DOI="http://doi.org/10.5281/zenodo.6492148" ID-GBIF-Taxon="100122179" ID-Zenodo-Dep="6492148" LSID="urn:lsid:plazi:treatment:92B743395B402090DE349E296CB71769" httpUri="http://treatment.plazi.org/id/92B743395B402090DE349E296CB71769" lastPageNumber="61" pageNumber="57">
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<subSubSection id="D3C0E134234CEA7E93AC244D903FA425" type="nomenclature">
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<paragraph id="18E96BB43C83D42B70B57EE76D121864" pageNumber="57">
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<taxonomicName id="027729E622BD2A7C09EB3CAFB7ACD418" ID-CoL="647V6" ID-ENA="910305" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="57" phylum="Chordata" rank="genus">Trypauchen Valenciennes in Cuvier & Valenciennes, 1837</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C7FE693E43207AA259023465BA290C41" type="reference_group">
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<paragraph id="91108A431A67F7EAB9C62B42C9FF827D" pageNumber="57">
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<taxonomicName id="8EC412D7BE583B5760782D66E9608AE4" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="57" phylum="Chordata" rank="genus">Trypauchen Valenciennes in Cuvier & Valenciennes, 1837</taxonomicName>
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: 152 (type species:
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<taxonomicName id="D7332F0FE35155FA6A962149C5427923" LSID-ZBK="urn:lsid:zoobank.org:act:2B284409-AD4A-4D52-9285-9B4A368CED73" family="Gobiidae" genus="Gobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobius vagina Bloch & Schneider 1801:73" order="Perciformes" pageNumber="57" phylum="Chordata" rank="species" species="vagina">Gobius vagina Bloch and Schneider, 1801</taxonomicName>
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, by original designation and monotypy)
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</paragraph>
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</subSubSection>
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<subSubSection id="43C64983AF046F10EB025F368107CDDE" type="nomenclature">
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<paragraph id="D379A429E95466E8929255C400CEBD53" pageNumber="57">
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Included species.
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<taxonomicName id="51F41AB75C533786D5FE8459A361E843" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="57" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
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comprises two species:
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<taxonomicName id="448F777D74650C0FB23754A79CCF601F" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="57" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
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<taxonomicNameLabel id="5E3E03D1718E47F9701F9C3D62A24F79" rank="species">n. sp.</taxonomicNameLabel>
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and
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<taxonomicName id="FB553CF6BAD4F6F908A6CF08DAC2DFB6" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="57" phylum="Chordata" rank="species" species="vagina">T. vagina</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="663DD68824DB5EC00A4C2BED3DF3350B" type="etymology">
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<paragraph id="E60273703C8DAB7121066C55DEFC99B6" pageNumber="57">Etymology. From the Greek trypa (hole) and auchen (neck).</paragraph>
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</subSubSection>
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<subSubSection id="D7D0BE5AE6FDEFE0A186BFC54773317D" type="diagnosis">
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<paragraph id="5446CAACE75AF296B8EA1C805B27CE79" pageNumber="57">
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Diagnosis.
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<taxonomicName id="21BFE7041C17809721C82E1F7338C018" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="57" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
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is distinguishable from the other amblyopine genera with an opercular pouch in having: four pterygiophores anterior to the first hemal spine (rarely three); small funnel-like pelvic fin with I,4 or I,5 rays and well-developed interradial membrane; and abdomen with scales.
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</paragraph>
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</subSubSection>
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<subSubSection id="F46B4A3D84283EEC7FAC6B39A1C8B56F" type="description">
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<paragraph id="3D5F43F05D6F33F3C1E91B088F560BC4" pageNumber="57">
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Description. The following description of meristic counts (Table 1) is based on radiographs, and four cleared and stained specimens (one each of
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<taxonomicName id="0979D981ED4581702ED85419A2BE226A" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="57" phylum="Chordata" rank="species" species="vagina">T. vagina</taxonomicName>
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removed from AMS I.43479-001 and USNM 342640 and two of
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<taxonomicName id="B132F8A9C42D0863F87C3CF464DA8CD4" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="57" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
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from USNM 385741).
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</paragraph>
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<paragraph id="5A54A88D8DA5EFA67B8517D850C73BF4" lastPageNumber="58" pageNumber="57">
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Total dorsal-fin elements 46-58; first dorsal fin with six flexible spines (one specimen of
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<taxonomicName id="8C8BE24CBD20464789B8E26B81D1F7DA" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="57" phylum="Chordata" rank="species" species="vagina">T. vagina</taxonomicName>
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with five); all second dorsal-fin rays segmented and branched; dorsal-fin base long and broadly joined with caudal fin. Total anal-fin elements 39-50, all elements segmented and branched; anal-fin height less than second dorsal-fin height; anal-fin membrane broadly joined with caudal fin. Pectoral fin with 15-20 rays, crescent-shaped or falcate with upper rays longer than lower ones; all pectoral-fin rays segmented and branched distally. Pelvic-fin rays I,4 or I,5; broad frenum present; medial rays longest and completely united by interradial membrane; pelvic fins forming a small funnel that is slightly pointed posteriorly. Caudal fin with 17 segmented rays including 8+7 branched rays and an upper and lower simple ray; unsegmented procurrent rays 4, 2 dorsally and 2 ventrally.
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</paragraph>
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<paragraph id="59E3FA21A9223AB060BD927DEFFFC29C" pageNumber="58">Scales cycloid, present only on body, largest posteriorly, scales overlapping in caudal peduncle area. Longitudinal scale count 59-98, difficult to count with accuracy.</paragraph>
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<paragraph id="DC2BC1FAE57F9F862E5338C02C48FD7A" pageNumber="58">Typically, two rows of teeth in the upper and lower jaws; all teeth caninoid. Outer-row teeth much larger and stouter than those of inner row; lower-jaw teeth larger than upperjaw teeth. Outer row of upper jaw with 4-16 enlarged, canine teeth; 8-15 enlarged, canine teeth in outer row of lower jaw; numerous smaller canine teeth in inner row of upper and lower jaws. When mouth closed, upper jaw overlapped by lower jaw. No palatine or vomerine teeth present.</paragraph>
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<paragraph id="0830CE9CEDC0B75C42DE9292CD64694D" pageNumber="58">Tongue thick, tip rounded, free from floor of mouth. Gape wide, mouth oblique; maxilla extending posteriorly to the vertical with anterior half of orbit; posteriorly, near tip of maxilla, upper lip expanded into large fold that joins similar fold of lower lip at rictus, fold completely covers posterior part of jaws even when agape. No barbels on underside of head.</paragraph>
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<paragraph id="17EB884A70AB801B5368D3BFFE4E5B3A" pageNumber="58">Eye rudimentary and covered by skin in orbital depression, lens slightly larger than length of posterior naris. Posterior naris located on dorsoanterior rim of orbit; anterior naris at tip of small tube-like flap that slightly overhangs upper jaw. Chin, snout, and interorbital area with thickened flesh, which likely aids in burrow construction.</paragraph>
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<paragraph id="0C88EFEBCDEEF7386BF9B153DF58BE9F" pageNumber="58">Cephalic sensory canals and pores absent. Sensory papillae present on head and body, but difficult to observe without use of a stain. Prominent pouch of unknown function along the dorsal margin of the opercle (Fig. 1).</paragraph>
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<paragraph id="095741BC96F00FAA7359F663AAF7BD55" pageNumber="58">Gill rakers short, blunt, and not ossified; gill opening narrow, extending only the length of pectoral-fin base or slightly more ventrally.</paragraph>
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<paragraph id="889FDE7D23C652A361372BE3CF222725" pageNumber="58">Genital papilla large. In males, the papilla is pointed posteriorly, in females it is slightly rounded or truncate posteriorly.</paragraph>
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<paragraph id="AB5324298D48641287B01C826C5EF18E" pageNumber="58">Proportional measurements are given in Table 2.</paragraph>
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<paragraph id="24B11F329D52E3E2913C3D7DEB6F68C5" pageNumber="59">
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Osteology. Spinous dorsal-fin pterygiophore formula (PF) typically 3-1221 (one specimen of
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<taxonomicName id="82BE3D39861F6500140045879AA5EB00" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="vagina">T. vagina</taxonomicName>
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with 3-122 and one specimen of
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<taxonomicName id="D4A86A5471CE732BDB847AA583D74705" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="vagina">T. vagina</taxonomicName>
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with 3-1211). Precaudal vertebrae 10, caudal vertebrae 19-20 or 23-24. Pterygiophore of the second soft dorsal-fin ray (posteriormost pterygiophore inserting in 7th interneural space) with an autogenous middle radial. Typically four anal-fin pterygiophores anterior to first hemal spine, two specimens with three. Epurals 2. Basihyal spatulate. No gap between dorsal aspect of symplectic and hyomandibula as symplectic is broadly joined to hyomandibula. An anteriorly directed laminar extension of the preopercle articulates with the ventroposterior edge of the symplectic. Frontal crest low, without serrated edge. Frontals divided by a septum anteriorly into two open canals. Atlas with well-developed parapophyses, in contact with first epineural rib. Epineurals present from 1st precaudal vertebra to at least the 14th and sometimes the 18th caudal vertebra. Well-developed pleural ribs on 3rd to 9th precaudal vertebrae.
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</paragraph>
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<paragraph id="F01041D03B0FE07D49C9801DEBF916F3" lastPageNumber="60" pageNumber="59">
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Comparison of
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<taxonomicName id="D6363B7FDFE8D3C070CAAD5B2D5DB05F" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
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with other
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<taxonomicName id="FA02FE827FA95084C55936E5DD9382A9" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
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members and status of
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<taxonomicName id="67FE429CF50131F98C68B24C6FF1FC95" LSID-ZBK="urn:lsid:zoobank.org:act:61561745-3F6F-42C8-A119-9D66CCD6BC86" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen microcephalus Bleeker 1860:62" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="microcephalus">T. microcephalus</taxonomicName>
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,
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<taxonomicName id="1129E9075D34D19163AEBE26C5BB5234" LSID-ZBK="urn:lsid:zoobank.org:act:038A087D-3977-4460-AA08-52262AD6F331" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen raha Popta 1922:37" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="raha">T. raha</taxonomicName>
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,
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<taxonomicName id="B9B435CD1D75D25C86B8D7065CF3BC00" LSID-ZBK="urn:lsid:zoobank.org:act:C9A767E8-E9DB-4365-9A9E-3892269D64AD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen taenia Koumans 1953:279" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="taenia">T. taenia</taxonomicName>
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, and
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<taxonomicName id="4CA052A9376CCA36B158A2810ECA0849" LSID-ZBK="urn:lsid:zoobank.org:act:59A8240F-5DB2-44A3-97BB-1E5EC245BDDB" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen wakae Jordan & Snyder 1901:127" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="wakae">T. wakae</taxonomicName>
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. Based on their shared absence of an interneural gap, Birdsong et al (1988) created the monophyletic unit called the
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<taxonomicName id="953F8463FDD5C1ACC07A39298EB597E2" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
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that comprised
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<taxonomicName id="9FA1BBBA7A4AC3AC725C38A01B291F0E" LSID-ZBK="urn:lsid:zoobank.org:act:201333D9-EFB8-49D0-8B35-4AFBB11167B8" family="Gobiidae" genus="Amblyotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyotrypauchen Hora 1924:160" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Amblyotrypauchen</taxonomicName>
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,
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<taxonomicName id="40D86ACD152FF5FAE72FA69810FAC3BA" LSID-ZBK="urn:lsid:zoobank.org:act:3B55DB48-DA82-4437-9CD5-367CB961753C" family="Gobiidae" genus="Caragobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Caragobius Smith & Seale 1906:81" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Caragobius</taxonomicName>
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,
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<taxonomicName id="882EC80550C1F363C314EDEB525871EF" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
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, and
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<taxonomicName id="1A32E78CA84D60BB12837EAAB27C7F94" LSID-ZBK="urn:lsid:zoobank.org:act:143E3671-850E-4A13-92A6-83A88F89C67F" family="Gobiidae" genus="Trypauchenichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchenichthys Bleeker 1860:4, 63" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Trypauchenichthys</taxonomicName>
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. Murdy (2002) added
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<taxonomicName id="B6B2543DE484F5975213DA38D6ECF86E" LSID-ZBK="urn:lsid:zoobank.org:act:4A5D6F02-949A-4FA4-ABD3-2532A2FAF038" family="Gobiidae" genus="Ctenotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ctenotrypauchen Steindachner 1867:63" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Ctenotrypauchen</taxonomicName>
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and a new genus (
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<taxonomicName id="6A95112375F971D43AA914623F6D2B5F" LSID-ZBK="urn:lsid:zoobank.org:act:9A19C237-3130-4D14-9EF2-17A7E24F4A41" family="Gobiidae" genus="Karsten" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Karsten Murdy 2002:787" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Karsten</taxonomicName>
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) to the
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<taxonomicName id="2E9E2C72A6C338DE33061280659351F6" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
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and provided a key to the
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<taxonomicName id="F4084D0CB43C8BB487BF119924F86B37" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
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genera. Additionally, four species currently assigned to the genus
|
||
<taxonomicName id="6012E38421F19ABFCA963C8BDD15A2AE" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
|
||
are treated separately here as members of the
|
||
<taxonomicName id="B76AE7A8D6A4E85EFA325B35FDC2DCC1" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
:
|
||
<taxonomicName id="BA7B4B90D486B5405349E3A8EF56414B" LSID-ZBK="urn:lsid:zoobank.org:act:61561745-3F6F-42C8-A119-9D66CCD6BC86" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen microcephalus Bleeker 1860:62" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="microcephalus">T. microcephalus</taxonomicName>
|
||
,
|
||
<taxonomicName id="739D8D2F9D836F7FEC1CF65B876FDD76" LSID-ZBK="urn:lsid:zoobank.org:act:038A087D-3977-4460-AA08-52262AD6F331" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen raha Popta 1922:37" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="raha">T. raha</taxonomicName>
|
||
,
|
||
<taxonomicName id="38330F6994BF5D7C7B670E5E7B5AB5DE" LSID-ZBK="urn:lsid:zoobank.org:act:C9A767E8-E9DB-4365-9A9E-3892269D64AD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen taenia Koumans 1953:279" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="taenia">T. taenia</taxonomicName>
|
||
, and
|
||
<taxonomicName id="5DA123A8EE31FD82EBA96E9D8122E1F5" LSID-ZBK="urn:lsid:zoobank.org:act:59A8240F-5DB2-44A3-97BB-1E5EC245BDDB" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen wakae Jordan & Snyder 1901:127" order="Perciformes" pageNumber="59" phylum="Chordata" rank="species" species="wakae">T. wakae</taxonomicName>
|
||
. Monophyletic subunits of the
|
||
<taxonomicName id="632AED03FD62D32F9BD9951A104389AB" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="59" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
can be hypothesized based on the following characters:
|
||
</paragraph>
|
||
<paragraph id="91B885D5F4509B9C6E793D32F2FFDE5A" pageNumber="60">
|
||
Opercular pouch (Fig. 1).
|
||
<taxonomicName id="527F10EA637C214E080DB9E5195767F9" LSID-ZBK="urn:lsid:zoobank.org:act:3B55DB48-DA82-4437-9CD5-367CB961753C" family="Gobiidae" genus="Caragobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Caragobius Smith & Seale 1906:81" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">Caragobius</taxonomicName>
|
||
and
|
||
<taxonomicName id="BD8B9F270C0D28E5DDFFB1235EAD3F8D" LSID-ZBK="urn:lsid:zoobank.org:act:9A19C237-3130-4D14-9EF2-17A7E24F4A41" family="Gobiidae" genus="Karsten" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Karsten Murdy 2002:787" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">Karsten</taxonomicName>
|
||
lack an opercular pouch whereas the other members of the
|
||
<taxonomicName id="212B12B3D5282F95F6D7811633010FE3" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
possess an opercular pouch, a unique character not found in any other amblyopines nor in any other gobioids. Hence, the presence of an opercular pouch is considered the derived condition.
|
||
</paragraph>
|
||
<paragraph id="834868B0F561F4637CE8D87E58DAF591" pageNumber="60">
|
||
Number of anal-fin pterygiophores anterior to the first hemal spine (AP). All
|
||
<taxonomicName id="5D9B126BA150051D84524B0028D0972D" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
members with an opercular pouch typically have three or fewer AP except for
|
||
<taxonomicName id="15C64717D054C822A577EA8A0BBA0EDC" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="vagina">Trypauchen vagina</taxonomicName>
|
||
and
|
||
<taxonomicName id="F258770DE86CBBE74CC52EAA1C7E7E12" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
|
||
<taxonomicNameLabel id="8DA849886240E543E0EDB2F8228A64A5" rank="species">n.sp.</taxonomicNameLabel>
|
||
, which typically have four AP. As the sister group (
|
||
<taxonomicName id="07A5CDF1B646A5782080957B55EE1418" LSID-ZBK="urn:lsid:zoobank.org:act:3B55DB48-DA82-4437-9CD5-367CB961753C" family="Gobiidae" genus="Caragobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Caragobius Smith & Seale 1906:81" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">Caragobius</taxonomicName>
|
||
and
|
||
<taxonomicName id="777BC9A52411710D57E77E149E15D836" LSID-ZBK="urn:lsid:zoobank.org:act:9A19C237-3130-4D14-9EF2-17A7E24F4A41" family="Gobiidae" genus="Karsten" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Karsten Murdy 2002:787" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">Karsten</taxonomicName>
|
||
) to those
|
||
<taxonomicName id="5E254B4224BD206150C597F03C2FADE4" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
members that have an opercular pouch typically have four or more AP, the condition of typically having three or fewer AP is hypothesized as the derived condition.
|
||
</paragraph>
|
||
<paragraph id="1ADDEC7A9987D3041BC005F81230A142" pageNumber="60">
|
||
Structure of the pelvic fins (Fig. 2). United pelvic fins forming a cup- or funnel-shaped disc with the medial rays joined completely by an interradial membrane is the typical gobiid condition. This condition is considered plesiomorphic within the Gobiidae. All
|
||
<taxonomicName id="A2786B6907849ED815A49B4CF77F2C08" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
members with an opercular pouch have emarginate or separate pelvic fins except for
|
||
<taxonomicName id="FA9C846D717C83F4C8F6E0BECC422D30" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="vagina">Trypauchen vagina</taxonomicName>
|
||
and
|
||
<taxonomicName id="39E13586A0E95E478CE0B04D3C4E52D8" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
|
||
<taxonomicNameLabel id="0DD7D434EF1FCE080289EFB162C10F74" rank="species">n. sp.</taxonomicNameLabel>
|
||
Emarginate or separate pelvic fins are considered derived.
|
||
</paragraph>
|
||
<paragraph id="07E79EE0DF914FDFD1CE32DABF3B733B" pageNumber="60">
|
||
Presence of scales on the abdomen.
|
||
<taxonomicName id="3223B2C21C3DFDDA465E4F812DB982DD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="vagina">Trypauchen vagina</taxonomicName>
|
||
,
|
||
<taxonomicName id="ACF58F54A53DB227AE3C2AB5D21CE47C" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
|
||
<taxonomicNameLabel id="56AD903C447D84879AA34972284AFBE8" rank="species">n. sp.</taxonomicNameLabel>
|
||
, and all other
|
||
<taxonomicName id="BDFF06CA968EDB93644FED33A9FCCF1F" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
members with an opercular pouch have scales on the abdomen except
|
||
<taxonomicName id="E059DBEB39A88ABD942CC592BB2B4C12" LSID-ZBK="urn:lsid:zoobank.org:act:61561745-3F6F-42C8-A119-9D66CCD6BC86" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen microcephalus Bleeker 1860:62" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="microcephalus">T. microcephalus</taxonomicName>
|
||
,
|
||
<taxonomicName id="7523E3A8F3C4B61411667FE6CE20C8AE" LSID-ZBK="urn:lsid:zoobank.org:act:038A087D-3977-4460-AA08-52262AD6F331" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen raha Popta 1922:37" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="raha">T. raha</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C6E44B2BE0E574D235DBBDE7F02DD62A" LSID-ZBK="urn:lsid:zoobank.org:act:59A8240F-5DB2-44A3-97BB-1E5EC245BDDB" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen wakae Jordan & Snyder 1901:127" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="wakae">T. wakae</taxonomicName>
|
||
. The absence of scales on the abdomen is hypothesized as being the derived condition.
|
||
</paragraph>
|
||
<paragraph id="5B49CA40FA7F6BA829FB3C553006C092" lastPageNumber="61" pageNumber="60">
|
||
Frontal crest. All
|
||
<taxonomicName id="EB0C81628901D7D228FE32A48EBD53A3" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
members with an opercular pouch lack an exposed frontal crest except
|
||
<taxonomicName id="6AC9AD455086C3ED8E8934AB6C68BDDE" LSID-ZBK="urn:lsid:zoobank.org:act:4A5D6F02-949A-4FA4-ABD3-2532A2FAF038" family="Gobiidae" genus="Ctenotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ctenotrypauchen Steindachner 1867:63" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">Ctenotrypauchen</taxonomicName>
|
||
,
|
||
<taxonomicName id="25AB98ED89AB5F93CEB21F59A164AAE7" LSID-ZBK="urn:lsid:zoobank.org:act:143E3671-850E-4A13-92A6-83A88F89C67F" family="Gobiidae" genus="Trypauchenichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchenichthys Bleeker 1860:4, 63" order="Perciformes" pageNumber="60" phylum="Chordata" rank="genus">Trypauchenichthys</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2B08773DF784B2B6B969D96AB53FEE25" LSID-ZBK="urn:lsid:zoobank.org:act:C9A767E8-E9DB-4365-9A9E-3892269D64AD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen taenia Koumans 1953:279" order="Perciformes" pageNumber="60" phylum="Chordata" rank="species" species="taenia">T. taenia</taxonomicName>
|
||
, which all have an exposed, serrated frontal crest. An exposed, serrated frontal crest is considered derived.
|
||
</paragraph>
|
||
<paragraph id="8FDA0D16ABBA25C6AA45F125E44A56A6" pageNumber="61">
|
||
Presence of head scales. All
|
||
<taxonomicName id="451E54F149CC979785B6601BE4C6854A" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
members with an opercular pouch lack scales on the head except
|
||
<taxonomicName id="84178503DA6FFA0BA498FDEA9CBCDE2C" LSID-ZBK="urn:lsid:zoobank.org:act:201333D9-EFB8-49D0-8B35-4AFBB11167B8" family="Gobiidae" genus="Amblyotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyotrypauchen Hora 1924:160" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Amblyotrypauchen</taxonomicName>
|
||
(Murdy, 2003).
|
||
<taxonomicName id="117720E1801AAE7E0F306B053978EE55" LSID-ZBK="urn:lsid:zoobank.org:act:201333D9-EFB8-49D0-8B35-4AFBB11167B8" family="Gobiidae" genus="Amblyotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyotrypauchen Hora 1924:160" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Amblyotrypauchen</taxonomicName>
|
||
has scale patches on the head, cheek and opercle; this is considered a derived character.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D866FFAB9DB31AFFA8837C7E59A6DEB0" type="discussion">
|
||
<paragraph id="9E746FB6405931402B1D71D93B15CD42" pageNumber="61">
|
||
Based on the character polarities described above, I hypothesize that
|
||
<taxonomicName id="D16DE0C3DD5FE05318337DDC1D1C25A7" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="vagina">Trypauchen vagina</taxonomicName>
|
||
and
|
||
<taxonomicName id="B00EBDB2B55FAA2BFA2B6B6948A551D3" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="4EA05EB86D7AD5DB9B8E67576ED95C16" rank="species">n.sp.</taxonomicNameLabel>
|
||
form the sister group to
|
||
<taxonomicName id="38773D23AFB3F79D135338554F755B65" LSID-ZBK="urn:lsid:zoobank.org:act:201333D9-EFB8-49D0-8B35-4AFBB11167B8" family="Gobiidae" genus="Amblyotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Amblyotrypauchen Hora 1924:160" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Amblyotrypauchen</taxonomicName>
|
||
,
|
||
<taxonomicName id="AF59D197E37AC79FC7986DCCD7C0C2E5" LSID-ZBK="urn:lsid:zoobank.org:act:4A5D6F02-949A-4FA4-ABD3-2532A2FAF038" family="Gobiidae" genus="Ctenotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ctenotrypauchen Steindachner 1867:63" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Ctenotrypauchen</taxonomicName>
|
||
,
|
||
<taxonomicName id="C9BEC5D64F4E808FC7B84EC32B20EDBE" LSID-ZBK="urn:lsid:zoobank.org:act:143E3671-850E-4A13-92A6-83A88F89C67F" family="Gobiidae" genus="Trypauchenichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchenichthys Bleeker 1860:4, 63" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Trypauchenichthys</taxonomicName>
|
||
and the four remaining nominal species of
|
||
<taxonomicName id="F1026309348B3EEF1ECE91D24661802E" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
|
||
(
|
||
<taxonomicName id="806E658AB65C1A2DFF8E270B2E8C76E2" LSID-ZBK="urn:lsid:zoobank.org:act:61561745-3F6F-42C8-A119-9D66CCD6BC86" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen microcephalus Bleeker 1860:62" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="microcephalus">T. microcephalus</taxonomicName>
|
||
,
|
||
<taxonomicName id="86319F3C9CCBF077EA8F9CF1DC78FD93" LSID-ZBK="urn:lsid:zoobank.org:act:038A087D-3977-4460-AA08-52262AD6F331" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen raha Popta 1922:37" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="raha">T. raha</taxonomicName>
|
||
,
|
||
<taxonomicName id="E732115D44FF1E642FF6E96E55ED65E6" LSID-ZBK="urn:lsid:zoobank.org:act:C9A767E8-E9DB-4365-9A9E-3892269D64AD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen taenia Koumans 1953:279" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="taenia">T. taenia</taxonomicName>
|
||
and
|
||
<taxonomicName id="66CF66DD4F919B4C6D7C9FCFA573D78E" LSID-ZBK="urn:lsid:zoobank.org:act:59A8240F-5DB2-44A3-97BB-1E5EC245BDDB" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen wakae Jordan & Snyder 1901:127" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="wakae">T. wakae</taxonomicName>
|
||
). Three nominal species of
|
||
<taxonomicName id="5378F843DE123376E9E5DCC4A1C792BB" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
|
||
(i.e.,
|
||
<taxonomicName id="D936C20570ACD900EED57162B5EEDDC1" LSID-ZBK="urn:lsid:zoobank.org:act:61561745-3F6F-42C8-A119-9D66CCD6BC86" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen microcephalus Bleeker 1860:62" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="microcephalus">T. microcephalus</taxonomicName>
|
||
,
|
||
<taxonomicName id="3B10F676B4EA448C820516301AF1F70A" LSID-ZBK="urn:lsid:zoobank.org:act:038A087D-3977-4460-AA08-52262AD6F331" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen raha Popta 1922:37" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="raha">T. raha</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7456142AD6A65EB61EE3AF28343AA44C" LSID-ZBK="urn:lsid:zoobank.org:act:59A8240F-5DB2-44A3-97BB-1E5EC245BDDB" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen wakae Jordan & Snyder 1901:127" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="wakae">T. wakae</taxonomicName>
|
||
) are united by synapomorphies (i.e., three or fewer AP, emarginate pelvic fins, and absence of abdominal scales) not found in combination in any other
|
||
<taxonomicName id="02D1A9E824162B475194E9D2FF9737FF" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">‘Trypauchen’ group</taxonomicName>
|
||
genus and, thus, cannot be assigned at this time. As for
|
||
<taxonomicName id="DAD5311816052F335455BE51C9DF261C" LSID-ZBK="urn:lsid:zoobank.org:act:C9A767E8-E9DB-4365-9A9E-3892269D64AD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen taenia Koumans 1953:279" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="taenia">T. taenia</taxonomicName>
|
||
, this species is hypothesized to be a member of the monophyletic unit containing
|
||
<taxonomicName id="C638A7B70D258A53104A9C7A993C811C" LSID-ZBK="urn:lsid:zoobank.org:act:4A5D6F02-949A-4FA4-ABD3-2532A2FAF038" family="Gobiidae" genus="Ctenotrypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ctenotrypauchen Steindachner 1867:63" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Ctenotrypauchen</taxonomicName>
|
||
and
|
||
<taxonomicName id="AF30D2D8558F3DED22A34DF5FE19F83F" LSID-ZBK="urn:lsid:zoobank.org:act:143E3671-850E-4A13-92A6-83A88F89C67F" family="Gobiidae" genus="Trypauchenichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchenichthys Bleeker 1860:4, 63" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Trypauchenichthys</taxonomicName>
|
||
. These four nominal species of
|
||
<taxonomicName id="980D529B416358F8C737D6583BB8300F" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
|
||
,
|
||
<taxonomicName id="40E2F0AF6B8B0323C970DA49F9608E33" LSID-ZBK="urn:lsid:zoobank.org:act:61561745-3F6F-42C8-A119-9D66CCD6BC86" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen microcephalus Bleeker 1860:62" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="microcephalus">T. microcephalus</taxonomicName>
|
||
,
|
||
<taxonomicName id="BC04D25B73A39FACDCAA56ADED41D17D" LSID-ZBK="urn:lsid:zoobank.org:act:038A087D-3977-4460-AA08-52262AD6F331" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen raha Popta 1922:37" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="raha">T. raha</taxonomicName>
|
||
,
|
||
<taxonomicName id="727F7704C40380E9FCECB0CB31E70F3C" LSID-ZBK="urn:lsid:zoobank.org:act:C9A767E8-E9DB-4365-9A9E-3892269D64AD" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen taenia Koumans 1953:279" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="taenia">T. taenia</taxonomicName>
|
||
, and
|
||
<taxonomicName id="595DAC3E32F6E5A7B90D68813767EEDB" LSID-ZBK="urn:lsid:zoobank.org:act:59A8240F-5DB2-44A3-97BB-1E5EC245BDDB" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen wakae Jordan & Snyder 1901:127" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="wakae">T. wakae</taxonomicName>
|
||
, will be treated in greater detail in forthcoming papers as revisionary studies of amblyopine gobies continue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="58DCE05A9005393D8737F577811D3CB4" type="key">
|
||
<paragraph id="26887C7E6E76CF7284CE9852A11A2D3C" pageNumber="61">
|
||
Key to the species of
|
||
<taxonomicName id="13302F3EBDE6951842C128F82232E95C" LSID-ZBK="urn:lsid:zoobank.org:act:19ACDE46-06C7-4E91-B9A5-8F0ED3417A7B" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trypauchen Valenciennes 1837:152" order="Perciformes" pageNumber="61" phylum="Chordata" rank="genus">Trypauchen</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8DC850B902066E7B7F382AC851AA3D24" pageNumber="61">
|
||
1a. Total elements in dorsal fin 50-58; total elements in anal fin 43-50; caudal vertebrae 23 or 24, typically 24; SL typically 84% or more of total length; head length less than 18% of SL; predorsal length less than 22% of SL; prepelvic length less than 18% of SL; preanal length 36% or less of SL. (Kuwait, India, Thailand, Vietnam, Philippines, China) ...............................................................................................................
|
||
<taxonomicName id="5C30C93B3A843C83EDADAFDF199681B0" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="vagina">T. vagina</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="CAD32364CC10C1A86E5FD4625CC91B5B" pageNumber="61">
|
||
1b. Total elements in dorsal fin 46-52; total elements in anal fin 39-44; caudal vertebrae 19 or 20; SL 84% or less of total length; head length 18% or more of SL; predorsal length typically more than 22% of SL; prepelvic length 18% or more of SL; preanal length more than 36% of SL. (Myanmar, Thailand, Malaysia, Vietnam, China) ........... ..............................................................................................................
|
||
<taxonomicName id="5439B2CCFB89DFA35635DACA857D2536" family="Gobiidae" genus="Trypauchen" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="61" phylum="Chordata" rank="species" species="pelaeos">T. pelaeos</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="8174F839CC4CE6E5A7FB3231378E22BD" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |