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<document id="07F49899B54916D2B0420132806E741D" ID-CLB-Dataset="29628" ID-DOI="http://dx.doi.org/10.3897/zookeys.573.7878" ID-GBIF-Dataset="2774945f-2178-4677-98e2-643a3255134b" ID-PMC="PMC4829928" ID-Pensoft-Pub="1313-2970-573-217" ID-PubMed="27110169" ID-ZooBank="C88328D61FDE4E6FBB3B7085AFE98939" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-573-217" ModsDocOrigin="ZooKeys 573" ModsDocTitle="A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae)" checkinTime="1533733036323" checkinUser="pensoft" docAuthor="Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit &amp; Ernst, Crystal" docDate="2016" docId="923019B67466B066A46B990DFADF08EE" docLanguage="en" docName="ZooKeys 573: 217-256" docOrigin="ZooKeys 573" docSource="http://dx.doi.org/10.3897/zookeys.573.7878" docTitle="Liogluta Thomson 1858" docType="treatment" docVersion="6" lastPageNumber="219" masterDocId="FF889744AC561C4EFF9AA070FFD58558" masterDocTitle="A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae)" masterLastPageNumber="256" masterPageNumber="217" pageNumber="219" updateTime="1701381888426" updateUser="ExternalLinkService">
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<mods:title id="8A33842AD0BFFBA41BBC172B950FB719">A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae)</mods:title>
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<mods:namePart id="FC5C36CE06BCD261832C053F2B109CE4">Klimaszewski, Jan</mods:namePart>
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<mods:namePart id="92F9D601422FE7FF435C430B5863041D">Langor, David W.</mods:namePart>
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<mods:namePart id="156969EAE07D46E9D0B231C2E85190F5">Sikes, Derek</mods:namePart>
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<mods:namePart id="2CAEFD8031E53EFB411A01F3E89ABDC8">Bourdon, Caroline</mods:namePart>
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<subSubSection id="9D3134CB11B2C406020D71E0AE8C7104" pageId="2" pageNumber="219" type="nomenclature">
<paragraph id="26BC5CE48E381DB1DC1D7FC89BE18208" pageId="2" pageNumber="219">
<taxonomicName id="2A0FEE38413879F09C30ABC34941BDBA" ID-CoL="8VRSS" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">Liogluta Thomson, 1858</taxonomicName>
</paragraph>
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<paragraph id="39C3CE0D6EF42E94090F631654C67034" pageId="2" pageNumber="219">
<taxonomicName id="470E0521B97B9EFC0C55C058F254D106" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">Liogluta</taxonomicName>
Thomson, 1858: 35. Type species
<taxonomicName id="E11DEC8966B9AEB2DA8AE8725B8BBE38" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota umbonata" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="umbonata">Homalota umbonata</taxonomicName>
Erichson, 1839, by monotypy. As valid genus:
<bibRefCitation id="6266C1B328A25493A7FDC1C30BAA6A0A" author="Lohse, GA" editor="Freude, H" journalOrPublisher="Goecke &amp; Evers, Krefeld" pageId="25" pageNumber="242" pagination="7 - 72" title="Die Kaefer Mitteleuropas. Band 5. Staphylinidae II" year="1974">Lohse (1974)</bibRefCitation>
;
<bibRefCitation id="38B7A2559C33E5140D3F1A18E505A29A" author="Lohse, GA" journalOrPublisher="The Coleopterists Bulletin" pageId="25" pageNumber="242" pagination="121 - 202" title="Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae)." volume="44" year="1990">Lohse et al. (1990)</bibRefCitation>
.
</paragraph>
<paragraph id="BDAC1F8BB53258143FDDED6851B61BFC" pageId="2" pageNumber="219">
<taxonomicName id="1DC42171AEC0B582E5879C68D0B2B572" class="Insecta" family="Staphylinidae" genus="Anepsiota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anepsiota" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">Anepsiota</taxonomicName>
Casey, 1894: 321;
<bibRefCitation id="D746E56B1B1A5DA4E79624E9B70ED36D" author="Casey, TL" journalOrPublisher="Louis" pageId="24" pageNumber="241" pagination="125 - 434" title="Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St." volume="16" year="1906">Casey (1906</bibRefCitation>
: 335); as
<taxonomicName id="E05C8EEAEBE980B09B1082F6DE29D1E5" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Anepsiota)" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="subGenus" subGenus="Anepsiota">Atheta (Anepsiota)</taxonomicName>
:
<bibRefCitation id="E23D29C3787F0587EB2493533EA5182D" author="Casey, TL" journalOrPublisher="New Era Printing Co., Lancaster, Pennsylvania" pageId="24" pageNumber="241" pagination="1 - 183" title="New species of the staphylinid tribe Myrmedoniini." url="10.5962/bhl.title.1159" volumeTitle="Memoirs on the Coleoptera" year="1910">Casey (1910</bibRefCitation>
: 12),
<bibRefCitation id="B7822D46A6510F2BF344224D7E2109E8" author="Fenyes, A" editor="Wytsman, PA" journalOrPublisher="L. Desmet-Verteneuil, Bruxelles" pageId="24" pageNumber="241" pagination="111 - 414" title="Genera Insectorum" year="1920">Fenyes (1920</bibRefCitation>
: 203). Synonymized by
<bibRefCitation id="2F7809A275310E350DF265ADD62368E5" author="Bernhauer, M" journalOrPublisher="Junk, Berlin" pageId="24" pageNumber="241" title="Pars 82: Staphylinidae VI [pp. 499 - 988] In: Coleopterorum Catalogus. Volumen V. Staphylinidae. W." year="1926">Bernhauer and Scheerpeltz (1926</bibRefCitation>
: 656);
<bibRefCitation id="7D3CA5FD04744AE8EF47F0038CA88960" author="Moore, I" journalOrPublisher="University of California, Division of Agricultural Sciences" pageId="25" pageNumber="242" pagination="1 - 514" title="A catalogue of the Staphylinidae of America north of Mexico (Coleoptera)." volumeTitle="Special Publication" year="1975">Moore and Legner (1975</bibRefCitation>
: 350).
</paragraph>
<paragraph id="83B64A054A3F0355B4ABE43CE09C140F" pageId="2" pageNumber="219">
<taxonomicName id="3369240E8622A5334850C5CBEB4EE824" class="Insecta" family="Staphylinidae" genus="Athetota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Athetota" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">Athetota</taxonomicName>
Casey, 1906: 334. Synonymized with
<taxonomicName id="25CB7275477C70510A367A88EC7147C2" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Anepsiota)" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="subGenus" subGenus="Anepsiota">Atheta (Anepsiota)</taxonomicName>
by
<bibRefCitation id="DDB6F1B3186E6895D46C8A195DF527D9" author="Fenyes, A" editor="Wytsman, PA" journalOrPublisher="L. Desmet-Verteneuil, Bruxelles" pageId="24" pageNumber="241" pagination="111 - 414" title="Genera Insectorum" year="1920">Fenyes (1920</bibRefCitation>
: 203); as synonym of
<taxonomicName id="FBB2D4816806C7A24F637EDE7BEC13E9" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Liogluta)" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="subGenus" subGenus="Liogluta">Atheta (Liogluta)</taxonomicName>
:
<bibRefCitation id="E5C8B67FB21BE5A69768AB34232FBD6C" author="Bernhauer, M" journalOrPublisher="Junk, Berlin" pageId="24" pageNumber="241" title="Pars 82: Staphylinidae VI [pp. 499 - 988] In: Coleopterorum Catalogus. Volumen V. Staphylinidae. W." year="1926">Bernhauer and Scheerpeltz (1926</bibRefCitation>
: 656),
<bibRefCitation id="271B9C883A05CC6EF6DA08D05896098B" author="Moore, I" journalOrPublisher="University of California, Division of Agricultural Sciences" pageId="25" pageNumber="242" pagination="1 - 514" title="A catalogue of the Staphylinidae of America north of Mexico (Coleoptera)." volumeTitle="Special Publication" year="1975">Moore and Legner (1975</bibRefCitation>
: 350).
</paragraph>
<paragraph id="0F29B86AA9506B3D09A96D420B8345F5" pageId="2" pageNumber="219">
<taxonomicName id="1C992135264531FA90E30D9FC176C62A" class="Insecta" family="Staphylinidae" genus="Hypnota" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hypnota" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">Hypnota</taxonomicName>
Mulsant &amp; Rey, 1873: 591. Synonymized with
<taxonomicName id="03CCA211C0256B2B70C4571193B0D58C" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Liogluta)" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="subGenus" subGenus="Liogluta">Atheta (Liogluta)</taxonomicName>
by
<bibRefCitation id="9BC545C8FA4B8D4F536B38C5BDD3FC8F" author="Fenyes, A" editor="Wytsman, PA" journalOrPublisher="L. Desmet-Verteneuil, Bruxelles" pageId="24" pageNumber="241" pagination="111 - 414" title="Genera Insectorum" year="1920">Fenyes (1920</bibRefCitation>
: 203);
<bibRefCitation id="415B7E03100D1F724DAC9D7D5E7B5C3B" author="Bernhauer, M" journalOrPublisher="Junk, Berlin" pageId="24" pageNumber="241" title="Pars 82: Staphylinidae VI [pp. 499 - 988] In: Coleopterorum Catalogus. Volumen V. Staphylinidae. W." year="1926">Bernhauer and Scheerpeltz (1926</bibRefCitation>
: 656);
<bibRefCitation id="432052AB202EC1294AB06C27C9B7D407" author="Moore, I" journalOrPublisher="University of California, Division of Agricultural Sciences" pageId="25" pageNumber="242" pagination="1 - 514" title="A catalogue of the Staphylinidae of America north of Mexico (Coleoptera)." volumeTitle="Special Publication" year="1975">Moore and Legner (1975</bibRefCitation>
: 350).
</paragraph>
</subSubSection>
<subSubSection id="4E23BD51EFBC7E4F2F5AA03F9BBDE76E" pageId="2" pageNumber="219" type="diagnosis">
<paragraph id="2FF23E88E8FC0AB0DB5657CA19A71AF8" pageId="2" pageNumber="219">Diagnosis.</paragraph>
<paragraph id="437A84E5769E524266B60758767D9F93" pageId="2" pageNumber="219">Body length ranging from 2.8-5.4 mm, body narrowly subparallel (Figs 1, 8, 16, 23, 30, 35, 44, 50, 58, 65, 72, 79, 86, 90), moderately flattened; elytra and abdomen wider than head and pronotum; uniformly dark brown or reddish-brown with head and posterior abdomen dark brown to almost black; integument of forebody with strong meshed microsculpture; surface of elytra often granulose (Figs 50, 51); head with incomplete postocular carinae, postocular region of head long and in most species longer than diameter of eye; glossae split into two lobes; antennae long with articles VI-X subquadrate, slightly transverse or rarely slightly elongate; pronotum with hypomera fully visible in lateral view; pubescence of pronotum directed posteriad on midline of disc and posterolaterad to laterad on sides; mesocoxae narrowly separated, metasternal process short and acute; legs long, three basal articles of metatarsi highly elongate in most species; tarsal formula 5-5-5. Male. Apical margin of male tergite VIII often with broad, variably-shaped rectangular projection, with edge entire or bearing crenulation or variably shaped structures, with two lateral teeth and sometimes with additional median tooth; integument of disc often with broadly distributed asperate punctation near apex (Figs 3, 11, 18, 25, 38, 46, 53, 60, 67, 74, 81, 88, 92); median lobe of aedeagus simple with apical part variably shaped in lateral view (Figs 2, 9, 17, 24, 36, 45, 52, 59, 66, 73, 80, 87, 91). Female. Spermatheca with capsule narrowly club-shaped or tubular, with apical invagination moderate to deep, stem long, sinuate, looped or twisted posteriorly (Figs 7, 15, 22, 29, 34, 42, 43, 57, 64, 71, 78, 85, 96); sternite VIII with apical margin rounded or medially emarginate, antecostal suture arcuate, or slightly to distinctly sinuate (Figs 6, 14, 21, 28, 33, 41, 49, 56, 63, 70, 77, 84, 89, 95).</paragraph>
<caption id="23990C3DDAD6C1DD35A6EAC3ACA0C2ED" pageId="2" pageNumber="219">
<paragraph id="54444E2E43617179B3576BB8CC9A6DD4" pageId="2" pageNumber="219">
Figures 1-7.
<taxonomicName id="D35E5CE11B6398EFC4F876FFE258399A" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta terminalis" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="terminalis">Liogluta terminalis</taxonomicName>
(Casey): 1 habitus in dorsal view 2 median lobe of aedeagus in lateral view 3 male tergite VIII 4 male sternite VIII 5 female tergite VIII 6 female sternite VIII 7 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="922F117EE959A971C2E625D53C3D80AC" pageId="2" pageNumber="219">
<paragraph id="48FA905E8AC113C135E6708A53D780E9" pageId="2" pageNumber="219">
Figures 8-15.
<taxonomicName id="AA11AD3868DF6FEFCE6594F18B031E0A" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta trapezicollis" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="trapezicollis">Liogluta trapezicollis</taxonomicName>
Lohse: 8 habitus in dorsal view 9, 10 median lobe of aedeagus in lateral view 11 male tergite VIII 12 male sternite VIII 13 female tergite VIII 14 female sternite VIII 15 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="6A7CDE5845411174A50486251A097A33" pageId="2" pageNumber="219">
<paragraph id="3C966E1968B59FFC24AE78810E1B89BE" pageId="2" pageNumber="219">
Figures 16-22.
<taxonomicName id="D767593CA97F8550B8EF0BA6C59D6F65" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta quadricollis" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="quadricollis">Liogluta quadricollis</taxonomicName>
(Casey): 16 habitus in dorsal view 17 median lobe of aedeagus in lateral view 18 male tergite VIII 19 male sternite VIII [16-19 based on male from BC] 20 female tergite VIII 21 female sternite VIII 22 spermatheca (19-22 based on female holotype). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="70474D4B6FFD5578D6E4C3A284129FC4" pageId="2" pageNumber="219">
<paragraph id="6708EAEC55987598E288892397D13E00" pageId="2" pageNumber="219">
Figures 23-29.
<taxonomicName id="E1BECE46735F64F5DE9AC6F28D4A6EF0" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta wickhami" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="wickhami">Liogluta wickhami</taxonomicName>
(Casey): 23 habitus in dorsal view 24 median lobe of aedeagus in lateral view 25 male tergite VIII 26 male sternite VIII 27 female tergite VIII 28 female sternite VIII 29 spermatheca [23, 27-29 based on female holotype]. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="36A0EAACBA249FCCA7E50788CD1CC925" pageId="2" pageNumber="219">
<paragraph id="37ACE6070453CC3EE885B3F72E31A15E" pageId="2" pageNumber="219">
Figures 30-34.
<taxonomicName id="8269DB2CE83414BA8D38B1E98036EF37" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta vasta" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="vasta">Liogluta vasta</taxonomicName>
(
<normalizedToken id="1A130AE8F5C5F032FD143B039CB5C01C" originalValue="Mäklin">Maeklin</normalizedToken>
): 30 habitus in dorsal view 31 female pygidium (terminal segments) 32 female tergite VIII 33 female sternite VIII (30-33 based on female holotype) 34 spermatheca (based on YT specimen after Lohse and Smetana 1895). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="7480F7EE7ED5459765634114207FCFC3" pageId="2" pageNumber="219">
<paragraph id="54E3C3A483F7833D04D83FCC6BD4297C" pageId="2" pageNumber="219">
Figures 35-43.
<taxonomicName id="F53BACB7B213D4371FCF01B429E19ED4" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta nigropolita" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="nigropolita">Liogluta nigropolita</taxonomicName>
(Bernhauer): 35 habitus in dorsal view 36 median lobe of aedeagus in lateral view 37 median lobe of aedeagus in dorsal view (36, 37 after
<bibRefCitation id="A232EF4FF02CC58E2170231B738F7BF2" author="Lohse, GA" journalOrPublisher="The Coleopterists Bulletin" pageId="25" pageNumber="242" pagination="121 - 202" title="Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae)." volume="44" year="1990">Lohse et al. 1990</bibRefCitation>
) 38 male tergite VIII 39 male sternite VIII 40 female tergite VIII 41 female sternite VIII 42, 43 spermatheca (43 after
<bibRefCitation id="9D6777C13B568499900F7BED9BCCC613" author="Lohse, GA" journalOrPublisher="The Coleopterists Bulletin" pageId="25" pageNumber="242" pagination="121 - 202" title="Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae)." volume="44" year="1990">Lohse et al. 1990</bibRefCitation>
). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="42D54C75634DA12127426B558CE4885C" pageId="2" pageNumber="219">
<paragraph id="6A468EB41E15EB9391BE9848356C1A3A" pageId="2" pageNumber="219">
Figures 44-49.
<taxonomicName id="A1F2DDA8A1BEFCB17EFBAA65B439084F" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta nitens" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="nitens">Liogluta nitens</taxonomicName>
(
<normalizedToken id="C9CFFFBC46B11E461A4AE2B821735C1C" originalValue="Mäklin">Maeklin</normalizedToken>
): 44 habitus in dorsal view (morphotype with broad and long elytra) 45 median lobe of aedeagus in lateral view 46 male tergite VIII 47 male sternite VIII 48 female tergite VIII 49 female sternite VIII. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="22B706F6A63D3550DF39494BF94B1C70" pageId="2" pageNumber="219">
<paragraph id="9444CBBC703D6802747F6B8E00586CE9" pageId="2" pageNumber="219">
Figures 50-57.
<taxonomicName id="2CB5F8B863EC1016C053CDA264D03E1F" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta granulosa" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="granulosa">Liogluta granulosa</taxonomicName>
Lohse: 50 habitus in dorsal view 51 elytron 52 median lobe of aedeagus in lateral view 53 male tergite VIII 54 male sternite VIII [50-54 based on male holotype] 55 female tergite VIII 56 female sternite VIII 57 spermatheca [55-57 based on female from YT]. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="52135E66EA28736D6C53F6167C625AF1" pageId="2" pageNumber="219">
<paragraph id="9F488418C70942A8ADE0D77562A05645" pageId="2" pageNumber="219">
Figures 58-64.
<taxonomicName id="584779B1769DFF09A7452942E4907470" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta microgranulosa" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="microgranulosa">Liogluta microgranulosa</taxonomicName>
Klimaszewski &amp; Webster, sp. n.: 58 habitus in dorsal view 59 median lobe of aedeagus in lateral view 60 male tergite VIII 61 male sternite VIII 62 female tergite VIII 63 female sternite VIII 64 spermatheca (58-64 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="7C6A6E957A18773CA097E0A7770EF05A" pageId="2" pageNumber="219">
<paragraph id="2A7309AB21892ED0AC997CCCA1E1E71D" pageId="2" pageNumber="219">
Figures 65-71.
<taxonomicName id="64337FCE2E3A7B7370EDCF2FA8AA7513" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta castoris" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="castoris">Liogluta castoris</taxonomicName>
Klimaszewski &amp; Webster, sp. n.: 65 habitus in dorsal view 66 median lobe of aedeagus in lateral view 67 male tergite VIII 68 male sternite VIII 69 female tergite VIII 70 female sternite VIII 71 spermatheca (65-71 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="12E7E7542ABEBE93432E1FAF27AE5C3B" pageId="2" pageNumber="219">
<paragraph id="5F365CAF185FC3D502BDBF5921C77254" pageId="2" pageNumber="219">
Figures 72-78.
<taxonomicName id="92596EF88E49558CE7DA28031B19B7B3" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta pseudocastoris" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="pseudocastoris">Liogluta pseudocastoris</taxonomicName>
Klimaszewski &amp; Webster, sp. n.: 72 habitus in dorsal view 73 median lobe of aedeagus in lateral view 74 male tergite VIII 75 male sternite VIII 76 female tergite VIII 77 female sternite VIII 78 spermatheca (72-78 based on type specimens). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="AF8002480C5E1FEF185B88D1347775DD" pageId="2" pageNumber="219">
<paragraph id="9F68679915D28705A323519D8721BB55" pageId="2" pageNumber="219">
Figures 79-85.
<taxonomicName id="C4FD1EBBFCB516151BF14063EB6A571A" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta intermedia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="intermedia">Liogluta intermedia</taxonomicName>
Klimaszewski &amp; Langor: 79 habitus in dorsal view 80 median lobe of aedeagus in lateral view 81 male tergite VIII, 82 male sternite VIII 83 female tergite VIII 84 female sternite VIII 85 spermatheca. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="E0DD1910CDB083AF26BE90BA8358E403" pageId="2" pageNumber="219">
<paragraph id="551EBC5387F2A937351E8A81917A28B1" pageId="2" pageNumber="219">
Figures 86-89.
<taxonomicName id="3794AB9F78207D434992213CBF0D5BA0" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta atriventris" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="atriventris">Liogluta atriventris</taxonomicName>
(Casey): 86 habitus in dorsal view 87 median lobe of aedeagus in lateral view 88 male tergite VIII 89 male sternite VIII (86-89 based on male lectotype). Female unknown. Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="7C00C0DEBCD794F3F6D4791FEC675CD6" pageId="2" pageNumber="219">
<paragraph id="4148F7EA39B9ECFD7B7349E978340FD4" pageId="2" pageNumber="219">
Figures 90-96.
<taxonomicName id="EF514FDABE3F9B3B2D0837E94E1DAFB9" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta gigantea" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="gigantea">Liogluta gigantea</taxonomicName>
Klimaszewski &amp; Langor: 90 habitus in dorsal view 91 median lobe of aedeagus in lateral view 92 male tergite VIII 93 male sternite VIII 94 female tergite VIII 95 female sternite VIII 96 spermatheca (90, 94-96 based on holotype, 91-93 based on male from Ontario). Scale bar of habitus = 1 mm, remaining scale bars = 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D0B49067934DDF1EC9C2919A93C7C2F0" pageId="2" pageNumber="219" type="distribution">
<paragraph id="E6133190402F6A760B927120D82A6EEE" pageId="2" pageNumber="219">Distribution.</paragraph>
<paragraph id="C07EE8750F2F77483F34133E39BF53AD" pageId="2" pageNumber="219">
The genus is holarctic in distribution (
<bibRefCitation id="46D9875E22781D8795532B5F7992349B" author="Smetana, A" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="25" pageNumber="242" pagination="353 - 494" title="Subfamily Aleocharinae Fleming, 1921." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea" year="2004">Smetana 2004</bibRefCitation>
).
<bibRefCitation id="456B1A35C3BACDD822054236A761ECCD" author="Seevers, CH" journalOrPublisher="Fieldiana: Zoology" pageId="25" pageNumber="242" title="A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae)." volume="71" year="1978">Seevers (1978)</bibRefCitation>
mentioned a few species from Africa and Jamaica but these records need verification.
</paragraph>
</subSubSection>
<subSubSection id="5DBBF53F79DF0CFF045B9072F81E748D" lastPageId="5" lastPageNumber="222" pageId="2" pageNumber="219" type="key to canadian and alaskan species of liogluta">
<paragraph id="EA0E1A0F289725CFD218343B7ED5A9F0" pageId="2" pageNumber="219">
Key to Canadian and Alaskan species of
<taxonomicName id="05930BCB7AD57AA29886FA6B28F1A570" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">Liogluta</taxonomicName>
</paragraph>
<paragraph id="14D7F69B6CF3A4B27B4E54D4EA64CC34" lastPageId="5" lastPageNumber="222" pageId="2" pageNumber="219">
<table id="E1C6BD96F1A23E984B75483E028661DF" lastPageId="5" lastPageNumber="222" pageId="2" pageNumber="219">
<tr id="6517D66DE54113E5772B22D5F8EAD19C" pageId="2" pageNumber="219">
<td id="CB6FC95B15A72332B6531D0347096D90" colspan="1" pageId="2" pageNumber="219" rowspan="1">18162330</td>
<td id="C19652580D5B44712A7F0CD0373238A2" colspan="1" pageId="2" pageNumber="219" rowspan="1">2</td>
</tr>
<tr id="E6303B5D8EF3AA920E793F72A505BF8C" pageId="2" pageNumber="219">
<td id="A7F6B205B187346248833A7C42A8AC6E" colspan="1" pageId="2" pageNumber="219" rowspan="1">354450586572798690</td>
<td id="433DD3712F144824A59289C50DFCC297" colspan="1" pageId="2" pageNumber="219" rowspan="1">6</td>
</tr>
<tr id="7F2E3A7448BD11BAA4B51413B42B893E" pageId="2" pageNumber="219">
<td id="5B9BA8FD81F2CC4323992F8744A96E52" colspan="1" pageId="2" pageNumber="219" rowspan="1">16</td>
<td id="207625C9B37B004B7D3AEED1D48C6A41" colspan="1" pageId="2" pageNumber="219" rowspan="1">
<taxonomicName id="7FF42755376ADF73EBB16028F34E6DCB" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta quadricollis" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="quadricollis">Liogluta quadricollis</taxonomicName>
(Casey)
</td>
</tr>
<tr id="686E334780EF1C180DBE0B13CE48389C" pageId="2" pageNumber="219">
<td id="477D9E3AB4363F8E2612ABC1EB74507E" colspan="1" pageId="2" pageNumber="219" rowspan="1">182330</td>
<td id="D6EBB8C9DD6A3786A1B247AC06855EDA" colspan="1" pageId="2" pageNumber="219" rowspan="1">3</td>
</tr>
<tr id="44C352EC68C3544C8953ED47AD094735" pageId="2" pageNumber="219">
<td id="B6170081043C3F75DD6A04B853277280" colspan="1" pageId="2" pageNumber="219" rowspan="1">1</td>
<td id="E2363257B69F07A7751920EEF783D10A" colspan="1" pageId="2" pageNumber="219" rowspan="1">
<taxonomicName id="3995FD901879FAF1B942837EC1867771" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta terminalis" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="terminalis">Liogluta terminalis</taxonomicName>
(Casey)
</td>
</tr>
<tr id="1B3599229FA41DF64465BF1731746118" pageId="2" pageNumber="219">
<td id="EB9507509F36381B0611457988921D8B" colspan="1" pageId="2" pageNumber="219" rowspan="1">82330</td>
<td id="90424ED7DF0339CFAE928C3C06817005" colspan="1" pageId="2" pageNumber="219" rowspan="1">4</td>
</tr>
<tr id="2AEBA5D4F5F9B112EFD9669DC6C07DE0" lastPageId="3" lastPageNumber="220" pageId="2" pageNumber="219">
<td id="80831D912460EDF55315B2EA186243DD" colspan="1" lastPageId="3" lastPageNumber="220" pageId="2" pageNumber="219" rowspan="1">
8811910
<pageBreakToken id="613F5FFABFEE150C38E6787993754F3A" pageId="3" pageNumber="220" start="start">15</pageBreakToken>
</td>
<td id="553CFC5A0AB7F1618EB8B96991CA22CA" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="FECC9C5FF6106EF0CA3B7122AC25510E" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta trapezicollis" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="trapezicollis">Liogluta trapezicollis</taxonomicName>
Lohse
</td>
</tr>
<tr id="2F30432F5A4753686509D89AEEA51527" pageId="3" pageNumber="220">
<td id="524AE40577CD3017C38602164280BFAD" colspan="1" pageId="3" pageNumber="220" rowspan="1">2330242934</td>
<td id="188F7435536E38CABE123E326CC78B98" colspan="1" pageId="3" pageNumber="220" rowspan="1">5</td>
</tr>
<tr id="2B78319A0DCAB5FD530D99ABF87294AA" pageId="3" pageNumber="220">
<td id="70AB02E46BA9CCE2A976298AEA97FD86" colspan="1" pageId="3" pageNumber="220" rowspan="1">232529</td>
<td id="FD6072B91BFC01F9A17F73C706B3F5EB" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="F871F418B2B8D52B045385AD005F06F4" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta wickhami" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="wickhami">Liogluta wickhami</taxonomicName>
(Casey)
</td>
</tr>
<tr id="D5EE605AD1765ABA2657F83D22B4D2F6" pageId="3" pageNumber="220">
<td id="9F6F617AE02AAAD42E600761E5B44E52" colspan="1" pageId="3" pageNumber="220" rowspan="1">3034</td>
<td id="68D580B86534D40D9E94304E0C54CC00" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="9FCD944D76D5CC214E85939DAA316D56" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta vasta" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="vasta">Liogluta vasta</taxonomicName>
(
<normalizedToken id="55537DF93783C98D5E0B6AE0AB436DD6" originalValue="Mäklin">Maeklin</normalizedToken>
)
</td>
</tr>
<tr id="DB01AABBBA7DE6C04D37434785D13AFA" pageId="3" pageNumber="220">
<td id="BEB97CFF846B989649E3A70141F437BB" colspan="1" pageId="3" pageNumber="220" rowspan="1">354490384692</td>
<td id="3DF4DF1D2B585A77AD13E5B194CC828E" colspan="1" pageId="3" pageNumber="220" rowspan="1">7</td>
</tr>
<tr id="499FF89AB89DEB096436B795D64EDAE9" pageId="3" pageNumber="220">
<td id="CA4B4C9F2BFAE2C9BB3D13231E14DB42" colspan="1" pageId="3" pageNumber="220" rowspan="1">505865727986536067748188</td>
<td id="2835C11083EA8661F955A523B9870A88" colspan="1" pageId="3" pageNumber="220" rowspan="1">9</td>
</tr>
<tr id="20EDE5F1C9E11982DBC3CD97BE697AA2" pageId="3" pageNumber="220">
<td id="9BA91608E367DB41BFE5DA609AC53CE9" colspan="1" pageId="3" pageNumber="220" rowspan="1">909091-96</td>
<td id="6D3149C4291F14FB8D5178A5C34A35B0" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="3C7BB57004AAE4B5B1239609E0FD17C8" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta gigantea" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="gigantea">Liogluta gigantea</taxonomicName>
Klimaszewski &amp; Langor
</td>
</tr>
<tr id="E937CD9E52899AFF59ADEE6C37D0D237" pageId="3" pageNumber="220">
<td id="18DB1BAE663DB12A37374A1FCDAC902D" colspan="1" pageId="3" pageNumber="220" rowspan="1">
3544354436
<normalizedToken id="1783FFDB6E5DD5030EF29524ACA693C9" originalValue="–4345–">-4345-</normalizedToken>
49
</td>
<td id="A4EC2B6D15E60981C81D77080ABA91AC" colspan="1" pageId="3" pageNumber="220" rowspan="1">8</td>
</tr>
<tr id="239256B8A4FE8B42D55BDF482E27B183" pageId="3" pageNumber="220">
<td id="632A75EF4E40F192805AC32B2E7A4E28" colspan="1" pageId="3" pageNumber="220" rowspan="1">353538363739-43</td>
<td id="37FC5C3C3CAE35FD2B549C7A866BC95A" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="639840281481E77E8BE7328D2A2080D1" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta nigropolita" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="nigropolita">Liogluta nigropolita</taxonomicName>
(Bernhauer)
</td>
</tr>
<tr id="521B00ABF4C4E84BAF61F519EFD5A41A" pageId="3" pageNumber="220">
<td id="430FEE41DA67A817D6FE7434FA061190" colspan="1" pageId="3" pageNumber="220" rowspan="1">44444645-49</td>
<td id="FBD3F0E8070A21F80FE4424682F157F9" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="01DDA4102945F43D1F4ED662893709D0" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta nitens" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="nitens">Liogluta nitens</taxonomicName>
(Mklin)
</td>
</tr>
<tr id="87B7F9A64519CE22075199499DBA9BEA" pageId="3" pageNumber="220">
<td id="7ED39D3765F6F57462EE1AFE83683734" colspan="1" pageId="3" pageNumber="220" rowspan="1">505051535254-57</td>
<td id="8DB5DE8D921E75D42AEDE71D055C9EE7" colspan="1" pageId="3" pageNumber="220" rowspan="1">
<taxonomicName id="0CD3F64641040D2EB44C090764BED350" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta granulosa" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="granulosa">Liogluta granulosa</taxonomicName>
Lohse
</td>
</tr>
<tr id="B5D99D03D2D062045B0D25479DBA50AC" lastPageId="4" lastPageNumber="221" pageId="3" pageNumber="220">
<td id="93F845CA332DB457C01723EFF4D47C62" colspan="1" lastPageId="4" lastPageNumber="221" pageId="3" pageNumber="220" rowspan="1">
5865727986
<pageBreakToken id="D7C4F0CF9179F9D48B27331BEF4DEBDA" pageId="4" pageNumber="221" start="start">6067748188</pageBreakToken>
</td>
<td id="FA2C0182B0E86BA6CB7E786139329DA1" colspan="1" pageId="4" pageNumber="221" rowspan="1">10</td>
</tr>
<tr id="66CD7DE0B4881A1A9CDA77BC57C153F3" pageId="4" pageNumber="221">
<td id="1619EE58B803DFF376D338F1D87363B1" colspan="1" pageId="4" pageNumber="221" rowspan="1">86868887</td>
<td id="462836ED4D391F4D4B59CAB79D6C7965" colspan="1" pageId="4" pageNumber="221" rowspan="1">
<taxonomicName id="432A429EDFC8B0FB855C1C9E6858DD40" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta atriventris" order="Coleoptera" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="atriventris">Liogluta atriventris</taxonomicName>
(Casey)
</td>
</tr>
<tr id="20636235304EEE6FE3E9BD4AB063CB6E" pageId="4" pageNumber="221">
<td id="4C51D64A27C9C33EC154766F0ED86C8C" colspan="1" pageId="4" pageNumber="221" rowspan="1">5865727960677481</td>
<td id="61EA49356DE8D3B52A0A5D081CA5D95E" colspan="1" pageId="4" pageNumber="221" rowspan="1">11</td>
</tr>
<tr id="7D24AC587A4A1ED8A8410EE71FDBCB3E" pageId="4" pageNumber="221">
<td id="8D195B604FD2207CCD99B19030D1CBBA" colspan="1" pageId="4" pageNumber="221" rowspan="1">7979797981808485</td>
<td id="7D9CDDCA334518CE96E1864A5C17B4B8" colspan="1" pageId="4" pageNumber="221" rowspan="1">
<taxonomicName id="296F5BB290D2775188FED5E66BD4C09A" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta intermedia" order="Coleoptera" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="intermedia">Liogluta intermedia</taxonomicName>
Klimaszewski &amp; Langor
</td>
</tr>
<tr id="1090A8F3683EC75EC18A24FFAE2A3C92" pageId="4" pageNumber="221">
<td id="F4F9951FF3C5A7823143F39BFDCF500C" colspan="1" pageId="4" pageNumber="221" rowspan="1">586572586572586572586572606774596673637077647178</td>
<td id="729D792A1768E7914D28E561CAD35642" colspan="1" pageId="4" pageNumber="221" rowspan="1">12</td>
</tr>
<tr id="DE1E5A8E3FCCCD80878E094B31028EAC" pageId="4" pageNumber="221">
<td id="A07660693CA49235F927B5F42E1A3EC3" colspan="1" pageId="4" pageNumber="221" rowspan="1">6567667071</td>
<td id="524E666FC154ACAFAB0C378E4A45864E" colspan="1" pageId="4" pageNumber="221" rowspan="1">
<taxonomicName id="98F664149D7F71AAB2E9168C90F914AA" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta castoris" order="Coleoptera" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="castoris">Liogluta castoris</taxonomicName>
Klimaszewski &amp; Webster, sp. n.
</td>
</tr>
<tr id="8F49E835274B68A2532E868FA128ABA8" pageId="4" pageNumber="221">
<td id="419058B5FB8B3F18ED00AE5CB6585CD1" colspan="1" pageId="4" pageNumber="221" rowspan="1">58726074597363776478</td>
<td id="C81FAACEDE7C8449339DE3E3A03369D3" colspan="1" pageId="4" pageNumber="221" rowspan="1">13</td>
</tr>
<tr id="2E9D3065A47F033C2D9FDEBCBBE865A3" pageId="4" pageNumber="221">
<td id="5A1C14E5BAF2F4CBC3586B17D6EA784D" colspan="1" pageId="4" pageNumber="221" rowspan="1">727274737778</td>
<td id="D73BEA9443DC076BD01D7A8F4EEF6061" colspan="1" pageId="4" pageNumber="221" rowspan="1">
<taxonomicName id="A2DD6D22B2752C7E5EB54EBD6B51E661" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta pseudocastoris" order="Coleoptera" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="pseudocastoris">Liogluta pseudocastoris</taxonomicName>
Klimaszewski &amp; Webster, sp. n.
</td>
</tr>
<tr id="576E51E7F78AD9AA9A223B58948E0AC0" pageId="5" pageNumber="222">
<td id="5D6B8E7FC2DBAB0E980B1BD9E6BB518F" colspan="1" pageId="5" pageNumber="222" rowspan="1">
<pageBreakToken id="CF5A88AA9A14F330829C25411A2C195F" pageId="5" pageNumber="222" start="start">585860596364</pageBreakToken>
</td>
<td id="B53FDFB4DF506F2AC462A86490E1BD67" colspan="1" pageId="5" pageNumber="222" rowspan="1">
<taxonomicName id="028835D37ABB4A906A5E75A5A316B74D" class="Insecta" family="Staphylinidae" genus="Liogluta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogluta microgranulosa" order="Coleoptera" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="microgranulosa">Liogluta microgranulosa</taxonomicName>
Klimaszewski &amp; Webster, sp. n.
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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