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<mods:title id="4BA666040599544CA1DF584A0C3FAB3F">Phylogeny of the cleptoparasitic Megachilini genera Coelioxys and Radoszkowskiana, with the description of six new subgenera in Coelioxys (Hymenoptera: Megachilidae)</mods:title>
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<subSubSection id="C35F650FCC2771597C05171B0811AC7E" box="[874,1363,607,634]" pageId="51" pageNumber="405" type="nomenclature">
<paragraph id="8BFA3684CC2771597C05171B0811AC7E" blockId="51.[874,1363,607,634]" box="[874,1363,607,634]" pageId="51" pageNumber="405">
<heading id="D0B281E8CC2771597C05171B0811AC7E" box="[874,1363,607,634]" centered="true" fontSize="9" level="2" pageId="51" pageNumber="405" reason="2">
<taxonomicName id="4C454D07CC2771597C05171B0811AC7E" authority="PASTEELS, 1977" authorityName="Pasteels" authorityYear="1977" box="[874,1363,607,634]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="378" phylum="Arthropoda" rank="subGenus" subGenus="Torridapis">
<emphasis id="B931EA96CC2771597C05171B09C9AC7D" box="[874,1163,608,634]" italics="true" pageId="51" pageNumber="405">
<smallCapsWord id="8D1CA058CC2771597C05171B0EAAAC70" baselines="627,627" box="[874,1000,611,632]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Coelioxys" pageId="51" pageNumber="405">COELIOXYS</smallCapsWord>
(
<smallCapsWord id="8D1CA058CC2771597C94171A09C0AC70" baselines="627,627" box="[1019,1154,610,633]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Torridapis" pageId="51" pageNumber="405">TORRIDAPIS</smallCapsWord>
)
</emphasis>
<bibRefCitation id="EFD44B75CC2771597BFB17190811AC7E" author="Pasteels JJ" box="[1172,1363,607,634]" pageId="51" pageNumber="405" pagination="161 - 197" refId="ref43482" refString="Pasteels JJ. 1977. Les Megachilini parasites (Coelioxys s. l.) d'Afrique noire. Revue de Zoologie Africaine 91: 161-197." type="journal article" year="1977">
<smallCapsWord id="8D1CA058CC2771597BFB17190841AC7F" baselines="627,628" box="[1172,1283,609,633]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Pasteels" pageId="51" pageNumber="405">PASTEELS</smallCapsWord>
, 1977
</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C35F650FCC27715E7C4417C80F64AF30" lastPageId="52" lastPageNumber="406" pageId="51" pageNumber="405" type="materials_examined">
<paragraph id="8BFA3684CC2771597C4417C8086BACC2" blockId="51.[811,1321,687,709]" box="[811,1321,687,709]" pageId="51" pageNumber="405">
<emphasis id="B931EA96CC2771597C4417C80EFCACC2" box="[811,958,688,709]" italics="true" pageId="51" pageNumber="405">
<typeStatus id="54FE8826CC2771597C4417C80E23ACC2" box="[811,865,688,709]" pageId="51" pageNumber="405">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C454D07CC2771597CA217D70867ACC2" authority="Smith, 1854" authorityName="Smith" authorityYear="1854" box="[973,1317,687,709]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="torrida">
<emphasis id="B931EA96CC2771597CA217D709CCACC3" box="[973,1166,687,708]" italics="true" pageId="51" pageNumber="405">Coelioxys torrida</emphasis>
Smith, 1854
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA3684CC2771597C4417950909ABC6" blockId="51.[811,1426,748,1473]" pageId="51" pageNumber="405">
<emphasis id="B931EA96CC2771597C4417950EE0AD05" box="[811,930,749,770]" italics="true" pageId="51" pageNumber="405">Diagnosis:</emphasis>
With
<emphasis id="B931EA96CC2771597C9117950952AD05" box="[1022,1040,749,770]" italics="true" pageId="51" pageNumber="405">C</emphasis>
. (
<emphasis id="B931EA96CC2771597B47179409E9AD06" box="[1064,1195,748,769]" italics="true" pageId="51" pageNumber="405">Liothyrapis</emphasis>
) and the species
<emphasis id="B931EA96CC2771597C4416740E7FAD26" box="[811,829,780,801]" italics="true" pageId="51" pageNumber="405">C</emphasis>
.
<taxonomicName id="4C454D07CC2771597C2216730909AD27" authorityName="Benoist" authorityYear="1955" box="[845,1099,779,800]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="madagascariensis" subGenus="Allocoelioxys">
(
<emphasis id="B931EA96CC2771597C3A16730E24AD27" box="[853,870,779,800]" italics="true" pageId="51" pageNumber="405">A</emphasis>
.)
<emphasis id="B931EA96CC2771597C1016730909AD27" box="[895,1099,779,800]" italics="true" pageId="51" pageNumber="405">madagascariensis</emphasis>
</taxonomicName>
, this Old World subgenus is the only one within
<taxonomicName id="4C454D07CC2771597B7B1652093FAD38" box="[1044,1149,810,831]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="B931EA96CC2771597B7B1652093FAD38" box="[1044,1149,810,831]" italics="true" pageId="51" pageNumber="405">Coelioxys</emphasis>
</taxonomicName>
in which the compound eyes are bare (
<figureCitation id="137E2A01CC2771597CB416300962AD5A" box="[987,1056,840,862]" captionStart="Figures 4 and 5" captionStartId="22.[162,252,1021,1040]" captionTargetBox="[172,1435,430,992]" captionTargetId="figure-248@22.[169,1432,426,988]" captionTargetPageId="22" captionText="Figures 4 and 5. Female. Frontal view of head (for compound eye): 4, Coelioxys weinlandi; 5, Coelioxys zapoteca. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807005" httpUri="https://zenodo.org/record/14807005/files/figure.png" pageId="51" pageNumber="405">Fig. 4</figureCitation>
). Species of
<emphasis id="B931EA96CC2771597BDB16310984AD59" box="[1204,1222,841,862]" italics="true" pageId="51" pageNumber="405">C</emphasis>
. (
<emphasis id="B931EA96CC2771597BB11630081AAD5A" box="[1246,1368,840,861]" italics="true" pageId="51" pageNumber="405">Torridapis</emphasis>
) can be distinguished from
<taxonomicName id="4C454D07CC2771597B2A161F0870AD7B" authorityName="Benoist" authorityYear="1955" box="[1093,1330,871,892]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B931EA96CC2771597B2A161F0915AD7B" box="[1093,1111,871,892]" italics="true" pageId="51" pageNumber="405">C</emphasis>
.
<emphasis id="B931EA96CC2771597B09161F0870AD7B" box="[1126,1330,871,892]" italics="true" pageId="51" pageNumber="405">madagascariensis</emphasis>
</taxonomicName>
by the lateral margins of axilla neither elevated nor carinate (
<figureCitation id="137E2A01CC2771597CC016DC0942ADBE" box="[943,1024,932,954]" captionStart="Figures 813" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 813. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 11</figureCitation>
) (conspicuously elevated and carinate in the latter;
<figureCitation id="137E2A01CC2771597B5F16BA0934ADDF" box="[1072,1142,962,984]" captionStart="Figures 6 and 7" captionStartId="22.[162,252,1850,1869]" captionTargetBox="[169,1434,1127,1818]" captionTargetId="figure-275@22.[169,1433,1129,1818]" captionTargetPageId="22" captionText="Figures 6 and 7. Female. Dorsal view of mesoscutellum (for lateral margin of axilla): 6, Coelioxys madagascariensis; 7, Coelioxys verticalis. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807009" httpUri="https://zenodo.org/record/14807009/files/figure.png" pageId="51" pageNumber="405">Fig. 6</figureCitation>
). They can be separated from those of
<emphasis id="B931EA96CC2771597CA116990EA2ADF1" box="[974,992,993,1014]" italics="true" pageId="51" pageNumber="405">C</emphasis>
. (
<emphasis id="B931EA96CC2771597C9716990939ADF1" box="[1016,1147,993,1014]" italics="true" pageId="51" pageNumber="405">Liothyrapis</emphasis>
) by the following list of characteristics. Axilla long, free apical portion produced beyond the posterior margin of mesoscutellum, and curved apically (
<figureCitation id="137E2A01CC2771597A5D114508C6AA55" box="[1330,1412,1085,1107]" captionStart="Figures 813" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 813. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 11</figureCitation>
); mesoscutellum with a dorsal median carina at least basally (
<figureCitation id="137E2A01CC2771597CF911020EAAAA88" box="[918,1000,1146,1168]" captionStart="Figures 813" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 813. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 11</figureCitation>
). Female:
<collectionCode id="ED54AE41CC2771597B1A110209C4AA97" box="[1141,1158,1146,1168]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
6 lacking long, erect, thick hairs, with apex ending in an inconspicuous, short, spine-like process (
<figureCitation id="137E2A01CC2771597BED11CF0996AACB" box="[1154,1236,1207,1229]" captionStart="Figures 813" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 813. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 12</figureCitation>
);
<collectionCode id="ED54AE41CC2771597B9411CF084EAACA" box="[1275,1292,1207,1229]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
6 and S6 gradually tapering, greatly elongate, S6 folded, subacute apically, its lateral margins with an inconspicuous fringe composed of short brownish hairs (
<figureCitation id="137E2A01CC2771597CE810490E98AB41" box="[903,986,1329,1351]" captionStart="Figures 813" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 813. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 12</figureCitation>
). Male: hypostomal area lacking concavity;
<collectionCode id="ED54AE41CC2771597CCA10280EF4AB61" box="[933,950,1360,1382]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="405">T</collectionCode>
6 basally lacking, or with short, flattened lateral expanded areas, never with long, acute teeth (
<figureCitation id="137E2A01CC2771597C5C10F50EC4ABA5" box="[819,902,1421,1443]" captionStart="Figures 813" captionStartId="24.[162,252,1789,1808]" captionTargetBox="[201,1433,1019,1606]" captionTargetId="figure-546@24.[169,1433,1020,1607]" captionTargetPageId="24" captionText="Figures 813. 8, 9, female, Coelioxys apicata: 8, dorsal view of mesoscutellum; 9, dorsal view of T6 and S6. 10, male, Coelioxys decipiens: dorsal view of T6 (for lateral tooth of T6). 11, female, Coelioxys ducalis: dorsal view of mesoscutellum. 12, 13, Coelioxys weinlandi: 12, female, dorsal view of T6 and S6; 13, male, dorsal view of T6 (for lateral tooth of T6). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807011" httpUri="https://zenodo.org/record/14807011/files/figure.png" pageId="51" pageNumber="405">Fig. 13</figureCitation>
); S4 medially sulcate, hairy, and S6 long, ≥1.2
<emphasis id="B931EA96CC2771597C3410D50E2CABC6" bold="true" box="[859,878,1453,1473]" italics="true" pageId="51" pageNumber="405">9</emphasis>
longer than broad.
</paragraph>
<paragraph id="8BFA3684CC2771597C4410910E9EA8B1" blockId="51.[811,1426,1513,1718]" pageId="51" pageNumber="405">
<emphasis id="B931EA96CC2771597C4410910EEBABF9" box="[811,937,1513,1534]" italics="true" pageId="51" pageNumber="405">Comments:</emphasis>
Ten species are classified in this subgenus (
<bibRefCitation id="EFD44B75CC2771597CC3137F09ABA81A" author="Ascher JS &amp; Pickering J" box="[940,1257,1543,1565]" pageId="51" pageNumber="405" refId="ref42522" refString="Ascher JS, Pickering J. 2015. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available via http://www.discoverlife.org/mp/20q?guide=Apoidea_species [Accessed 10 March 2015]" type="url" year="2015">Ascher &amp; Pickering, 2015</bibRefCitation>
), with a new one described by
<bibRefCitation id="EFD44B75CC2771597B4E135E0849A83B" author="Rocha-Filho LC" box="[1057,1291,1574,1596]" pageId="51" pageNumber="405" pagination="9 - 28" refId="ref43631" refString="Rocha-Filho LC. 2016. A revision of the cleptoparasitic bee genus Coelioxys (Hymenoptera: Megachilidae) from Australia. European Journal of Entomology 113: 9-28." type="journal article" year="2016">Rocha-Filho (2016)</bibRefCitation>
; the four Afrotropical species were revised by
<bibRefCitation id="EFD44B75CC2771597B8E133C08D0A85D" author="Pasteels JJ" box="[1249,1426,1604,1626]" pageId="51" pageNumber="405" pagination="1 - 139" refId="ref43437" refString="Pasteels JJ. 1968. Revision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Annales Sciences Zoologiques, Mus <20> ee Royal de L'Afrique Centrale, Tervuren, Serie In- 8 167: 1-139." type="journal article" year="1968">Pasteels (1968)</bibRefCitation>
and classified as
<emphasis id="B931EA96CC2771597C9A131B093AA87F" box="[1013,1144,1635,1656]" italics="true" pageId="51" pageNumber="405">Liothyrapis</emphasis>
, the other species come from Indomalayan or Palaearctic regions,
<collectingCountry id="F3527614CC2771597A4F13F908D0A890" box="[1312,1426,1665,1687]" name="Mauritius" pageId="51" pageNumber="405">Mauritius</collectingCountry>
or
<collectingCountry id="F3527614CC2771597C2413D80E9BA8B1" box="[843,985,1696,1718]" name="Madagascar" pageId="51" pageNumber="405">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BFA3684CC27715E7C4413A60F64AF30" blockId="51.[811,1426,1757,1871]" lastBlockId="52.[162,776,199,312]" lastPageId="52" lastPageNumber="406" pageId="51" pageNumber="405">
<emphasis id="B931EA96CC2771597C4413A60EB5A8F4" box="[811,1015,1757,1779]" italics="true" pageId="51" pageNumber="405">List of species:</emphasis>
<taxonomicName id="4C454D07CC2771597B6A13A508CCA8F5" authority="Friese, 1911" authorityName="Friese" authorityYear="1911" box="[1029,1422,1757,1779]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B931EA96CC2771597B6A13A50993A8F5" box="[1029,1233,1757,1778]" italics="true" pageId="51" pageNumber="405">Coelioxys analis</emphasis>
Friese, 1911
</taxonomicName>
,
<taxonomicName id="4C454D07CC2771597C44138409EAA916" authority="Smith, 1875" authorityName="Smith" authorityYear="1875" box="[811,1192,1787,1810]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B931EA96CC2771597C4413840EBBA916" box="[811,1017,1788,1809]" italics="true" pageId="51" pageNumber="405">Coelioxys basalis</emphasis>
Smith, 1875
</taxonomicName>
,
<taxonomicName id="4C454D07CC2771597BAE13840E8BA937" authority="Smith, 1854" authorityName="Smith" authorityYear="1854" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B931EA96CC2771597BAE138408D0A916" box="[1217,1426,1788,1809]" italics="true" pageId="51" pageNumber="405">Coelioxys ducalis</emphasis>
Smith, 1854
</taxonomicName>
,
<taxonomicName id="4C454D07CC2771597C8A126208CCA928" authority="Smith, 1873" authorityName="Smith" authorityYear="1873" box="[997,1422,1818,1840]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="fenestrata">
<emphasis id="B931EA96CC2771597C8A1262099AA928" box="[997,1240,1818,1839]" italics="true" pageId="51" pageNumber="405">Coelioxys fenestrata</emphasis>
Smith, 1873
</taxonomicName>
,
<taxonomicName id="4C454D07CC2771597C44124109B6A948" authority="Rocha-Filho, 2016" authorityName="Rocha-Filho" authorityYear="2016" box="[811,1268,1849,1871]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="julia">
<emphasis id="B931EA96CC2771597C4412410EA9A949" box="[811,1003,1849,1870]" italics="true" pageId="51" pageNumber="405">Coelioxys julia</emphasis>
<bibRefCitation id="EFD44B75CC2771597B62124109B6A948" author="Rocha-Filho LC" box="[1037,1268,1849,1871]" pageId="51" pageNumber="405" pagination="9 - 28" refId="ref43631" refString="Rocha-Filho LC. 2016. A revision of the cleptoparasitic bee genus Coelioxys (Hymenoptera: Megachilidae) from Australia. European Journal of Entomology 113: 9-28." type="journal article" year="2016">Rocha-Filho, 2016</bibRefCitation>
</taxonomicName>
*,
<taxonomicName id="4C454D07CC27715E7A4612410C87AEDB" authority="Strand, 1913" authorityName="Strand" authorityYear="1913" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" lastPageId="52" lastPageNumber="406" order="Hymenoptera" pageId="51" pageNumber="405" phylum="Arthropoda" rank="species" species="kosemponis">
<emphasis id="B931EA96CC27715E7A4612410C66AEDB" italics="true" lastPageId="52" lastPageNumber="406" pageId="51" pageNumber="405">Coelioxys kosemponis</emphasis>
Strand, 1913
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7EBC15BF0D99AEFD" authority="Friese, 1913" authorityName="Friese" authorityYear="1913" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B931EA96CC20715E7EBC15BF0FF3AEDB" box="[467,689,199,220]" italics="true" pageId="52" pageNumber="406">Coelioxys maculata</emphasis>
Friese, 1913
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E69159D0E46AEFC" authority="Pasteels, 1968" authorityName="Pasteels" authorityYear="1968" box="[262,772,229,251]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="maculoides">
<emphasis id="B931EA96CC20715E7E69159D0F55AEFD" box="[262,535,229,250]" italics="true" pageId="52" pageNumber="406">Coelioxys maculoides</emphasis>
<bibRefCitation id="EFD44B75CC20715E7D51159D0E46AEFC" author="Pasteels JJ" box="[574,772,229,251]" pageId="52" pageNumber="406" pagination="1 - 139" refId="ref43437" refString="Pasteels JJ. 1968. Revision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Annales Sciences Zoologiques, Mus <20> ee Royal de L'Afrique Centrale, Tervuren, Serie In- 8 167: 1-139." type="journal article" year="1968">Pasteels, 1968</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7FCD147C0C51AF1E" authorityName="Smith" authorityYear="1854" box="[162,275,260,281]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="torrida">
<emphasis id="B931EA96CC20715E7FCD147C0DF6AF1E" box="[162,180,260,281]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7FAC147C0C51AF1E" box="[195,275,260,281]" italics="true" pageId="52" pageNumber="406">torrida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E45147C0F8BAF1D" authority="Pasteels, 1968" authorityName="Pasteels" authorityYear="1968" box="[298,713,260,282]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="torridula">
<emphasis id="B931EA96CC20715E7E45147C0F49AF1E" box="[298,523,260,281]" italics="true" pageId="52" pageNumber="406">Coelioxys torridula</emphasis>
<bibRefCitation id="EFD44B75CC20715E7D74147C0F8BAF1D" author="Pasteels JJ" box="[539,713,260,282]" pageId="52" pageNumber="406" pagination="1 - 139" refId="ref43437" refString="Pasteels JJ. 1968. Revision des Megachilidae (Hymenoptera Apoidea) de L'Afrique Noire. II. Le Genre Coelioxys. Annales Sciences Zoologiques, Mus <20> ee Royal de L'Afrique Centrale, Tervuren, Serie In- 8 167: 1-139." type="journal article" year="1968">Pasteels, 1968</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C454D07CC20715E7FCD145A0F60AF30" authority="Schulz, 1904" authorityName="Schulz" authorityYear="1904" box="[162,546,290,312]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="weinlandi">
<emphasis id="B931EA96CC20715E7FCD145A0CC7AF30" box="[162,389,290,311]" italics="true" pageId="52" pageNumber="406">Coelioxys weinlandi</emphasis>
Schulz, 1904
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>