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<mods:title id="DA2DA9341843DF00CB0648A26E587E2A">Phylogeny of the cleptoparasitic Megachilini genera Coelioxys and Radoszkowskiana, with the description of six new subgenera in Coelioxys (Hymenoptera: Megachilidae)</mods:title>
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<heading id="D0B281E8CC20715E7FAA14170FA7AF82" box="[197,741,363,390]" centered="true" fontSize="9" level="2" pageId="52" pageNumber="406" reason="2">
<taxonomicName id="4C454D07CC20715E7FAA14170FA7AF82" authority="MITCHELL, 1973" authorityName="Mitchell" authorityYear="1973" box="[197,741,363,390]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="383" phylum="Arthropoda" rank="subGenus" subGenus="Xerocoelioxys">
<emphasis id="B931EA96CC20715E7FAA14170F55AF82" box="[197,535,363,389]" italics="true" pageId="52" pageNumber="406">
<smallCapsWord id="8D1CA058CC20715E7FAA14170C01AF85" baselines="383,382" box="[197,323,367,388]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Coelioxys" pageId="52" pageNumber="406">COELIOXYS</smallCapsWord>
(
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)
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<bibRefCitation id="EFD44B75CC20715E7D4F14150FA7AF82" author="Mitchell TB" box="[544,741,363,390]" pageId="52" pageNumber="406" refId="ref43263" refString="Mitchell TB. 1973. A subgeneric revision of the genus Coelioxys of the Western Hemisphere (Hymenoptera: Megachilidae). Raleigh: Department of Entomology, North Carolina State University." type="book" year="1973">
<smallCapsWord id="8D1CA058CC20715E7D4F14150FD4AF84" baselines="383,383" box="[544,662,365,389]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mitchell" pageId="52" pageNumber="406">MITCHELL</smallCapsWord>
, 1973
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<emphasis id="B931EA96CC20715E7FCD14C40C76AFD6" box="[162,308,444,465]" italics="true" pageId="52" pageNumber="406">
<typeStatus id="54FE8826CC20715E7FCD14C40D9AAFD6" box="[162,216,444,465]" pageId="52" pageNumber="406">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C454D07CC20715E7E2C14C30FDBAFD7" authority="Cresson, 1872" authorityName="Cresson" authorityYear="1872" box="[323,665,443,464]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="edita">
<emphasis id="B931EA96CC20715E7E2C14C30CAFAFD7" box="[323,493,443,464]" italics="true" pageId="52" pageNumber="406">Coelioxys edita</emphasis>
Cresson, 1872
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.
</paragraph>
<paragraph id="8BFA3684CC20715E7FCD14810C33AA76" blockId="52.[162,777,504,1137]" pageId="52" pageNumber="406">
<emphasis id="B931EA96CC20715E7FCD14810C5BAC09" box="[162,281,505,526]" italics="true" pageId="52" pageNumber="406">Diagnosis:</emphasis>
Species in this subgenus are very similar to
<emphasis id="B931EA96CC20715E7FD0176F0D93AC2B" box="[191,209,535,556]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7F86176F0C3CAC2B" box="[233,382,535,556]" italics="true" pageId="52" pageNumber="406">
<taxonomicName id="4C454D07CC20715E7F86176F0C10AC2B" box="[233,338,535,556]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
<taxonomicNameLabel id="A20257EDCC20715E7E34176F0C3CAC2B" box="[347,382,535,556]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
) spp., but can be differentiated by the pronotal lobe distinctly lamellate (Fig. 31) or carinate throughout, whereas in species of
<emphasis id="B931EA96CC20715E7FCD170B0DF6AC8F" box="[162,180,627,648]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7FA4170A0C23AC8F" box="[203,353,626,648]" italics="true" pageId="52" pageNumber="406">
<taxonomicName id="4C454D07CC20715E7FA4170A0C76AC80" box="[203,308,626,647]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
<taxonomicNameLabel id="A20257EDCC20715E7E51170B0C23AC8F" box="[318,353,627,648]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
) the pronotal lobe is conspicuously rounded on its outer margin and weakly carinate on its inner margin (
<figureCitation id="137E2A01CC20715E7E0517D70CFFACC2" box="[362,445,687,709]" captionStart="Figures 30 and 31" captionStartId="28.[162,252,820,839]" captionTargetBox="[170,1433,198,787]" captionTargetId="figure-156@28.[170,1433,198,787]" captionTargetPageId="28" captionText="Figures 30 and 31. Female, frontal view of pronotal lobe (outer margin indicated by red arrow): 30, Coelioxys sodalis; 31, Coelioxys grindeliae. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807021" httpUri="https://zenodo.org/record/14807021/files/figure.png" pageId="52" pageNumber="406">Fig. 30</figureCitation>
). Another difference between species of these two subgenera is in the mesosomal pilosity, which is short, composed of subappressed or appressed hairs, forming marginal lines or spots on the mesoscutum in
<emphasis id="B931EA96CC20715E7EFE16520CE1AD38" box="[401,419,810,831]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7ED516520F12AD38" box="[442,592,810,831]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
) spp. (Fig. 31), contrasting with the long and dense pubescence in species of
<emphasis id="B931EA96CC20715E7E70161F0C73AD7B" box="[287,305,871,892]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7E26161F0C9DAD7A" box="[329,479,871,893]" italics="true" pageId="52" pageNumber="406">
<taxonomicName id="4C454D07CC20715E7E26161F0CF0AD7B" box="[329,434,871,892]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
<taxonomicNameLabel id="A20257EDCC20715E7ED416100C9DAD7A" box="[443,479,872,893]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="137E2A01CC20715E7E93161F0F0DAD7B" box="[508,591,871,893]" captionStart="Figures 30 and 31" captionStartId="28.[162,252,820,839]" captionTargetBox="[170,1433,198,787]" captionTargetId="figure-156@28.[170,1433,198,787]" captionTargetPageId="28" captionText="Figures 30 and 31. Female, frontal view of pronotal lobe (outer margin indicated by red arrow): 30, Coelioxys sodalis; 31, Coelioxys grindeliae. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807021" httpUri="https://zenodo.org/record/14807021/files/figure.png" pageId="52" pageNumber="406">Fig. 30</figureCitation>
). The following combination of characteristics together with the two preceding ones separate species of
<emphasis id="B931EA96CC20715E7D2F16DC0F10ADBE" box="[576,594,932,953]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7D0516DC0E42ADBE" box="[618,768,932,953]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
) from all other subgenera. Mesoscutum coarsely and densely punctate, no interspaces linear to shorter than puncture diameter (as in
<figureCitation id="137E2A01CC20715E7D0811780FB9AA12" box="[615,763,1023,1046]" captionStart="Figures 83–86" captionStartId="45.[148,238,1329,1348]" captionTargetBox="[155,1417,198,1296]" captionTargetId="figure-283@45.[155,1417,198,1296]" captionTargetPageId="45" captionText="Figures 83–86. Female. 83, 84, Coelioxys rufescens: 83, dorsal view of mesoscutellum (for pilosity on anterior margin of mesoscutellum); 84, dorsal view of terga (for apical tergal fasciae). 85, 86, Coelioxys conoidea: 85, dorsal view of mesoscutellum (for pilosity on anterior margin of mesoscutellum); 86, dorsal view of terga (for apical tergal fasciae). Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807051" httpUri="https://zenodo.org/record/14807051/files/figure.png" pageId="52" pageNumber="406">Figs 83, 85</figureCitation>
); postgradular grooves of
<collectionCode id="ED54AE41CC20715E7D4811660F7AAA34" box="[551,568,1054,1075]" name="Tavera, Department of Geology and Geophysics" pageId="52" pageNumber="406">T</collectionCode>
2
<emphasis id="B931EA96CC20715E7D2A11590F13AA34" box="[581,593,1057,1075]" italics="true" pageId="52" pageNumber="406">–</emphasis>
<collectionCode id="ED54AE41CC20715E7D3E11660F21AA34" box="[593,611,1054,1075]" name="Tavera, Department of Geology and Geophysics" pageId="52" pageNumber="406">T</collectionCode>
3 complete, uninterrupted medially (
<figureCitation id="137E2A01CC20715E7EAF11450F56AA55" box="[448,532,1085,1107]" captionStart="Figures 24 and 25" captionStartId="26.[162,252,1593,1612]" captionText="Figures 24 and 25. Female, dorsal view of T2–T3 (for postgradular grooves): 24, Coelioxys rufitarsis; 25, Coelioxys apacheorum. Scale bars: 1.5 mm." pageId="52" pageNumber="406">Fig. 24</figureCitation>
), and gradulus of
<collectionCode id="ED54AE41CC20715E7D8511450FB9AA55" box="[746,763,1085,1106]" name="Tavera, Department of Geology and Geophysics" pageId="52" pageNumber="406">T</collectionCode>
2 bowed posteriorly.
</paragraph>
<paragraph id="8BFA3684CC20715E7FCD11E10FCBA81A" blockId="52.[162,776,1176,1565]" pageId="52" pageNumber="406">
<emphasis id="B931EA96CC20715E7FCD11E10C5DAAA9" box="[162,287,1177,1198]" italics="true" pageId="52" pageNumber="406">Comments:</emphasis>
<bibRefCitation id="EFD44B75CC20715E7E4111E00C8DAAA9" author="Baker JR" box="[302,463,1176,1198]" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker (1975)</bibRefCitation>
revised and keyed ten species in this subgenus. As a result of our phylogenies,
<taxonomicName id="4C454D07CC20715E7E2D11AE0CB1AAEC" authorityName="Baker" authorityYear="1975" box="[322,499,1238,1259]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="serricaudata">
<emphasis id="B931EA96CC20715E7E2D11AE0C16AAEC" box="[322,340,1238,1259]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7E0C11AE0CB1AAEC" box="[355,499,1238,1259]" italics="true" pageId="52" pageNumber="406">serricaudata</emphasis>
</taxonomicName>
, previously placed in
<emphasis id="B931EA96CC20715E7FCD118C0DF6AB0E" box="[162,180,1268,1289]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7FA4118C0C23AB0D" box="[203,353,1268,1290]" italics="true" pageId="52" pageNumber="406">
<taxonomicName id="4C454D07CC20715E7FA4118C0C76AB0E" box="[203,308,1268,1289]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
<taxonomicNameLabel id="A20257EDCC20715E7E51118D0C23AB0D" box="[318,353,1269,1290]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
), is now regarded as belonging to
<emphasis id="B931EA96CC20715E7FCD106B0DF6AB2F" box="[162,180,1299,1320]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7FA4106B0C23AB2F" box="[203,353,1299,1320]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
). Also, considering the diagnostic characteristics of this subgenus, the other three North American species assigned by
<bibRefCitation id="EFD44B75CC20715E7D3C10280FABAB61" author="Baker JR" box="[595,745,1360,1382]" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker (1975)</bibRefCitation>
to
<emphasis id="B931EA96CC20715E7FCD10170DF6AB83" box="[162,180,1391,1412]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7FA410160C23AB83" box="[203,353,1390,1412]" italics="true" pageId="52" pageNumber="406">
<taxonomicName id="4C454D07CC20715E7FA410160C76AB84" box="[203,308,1390,1411]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="genus" sensu="stricto">Coelioxys</taxonomicName>
<taxonomicNameLabel id="A20257EDCC20715E7E5110170C23AB83" box="[318,353,1391,1412]" pageId="52" pageNumber="406" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
),
<taxonomicName id="4C454D07CC20715E7EEF10160D99ABA5" authority="Cockerell, 1912" authorityName="Cockerell" authorityYear="1912" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="hirsutissima">
<emphasis id="B931EA96CC20715E7EEF10160FC5AB84" box="[384,647,1390,1411]" italics="true" pageId="52" pageNumber="406">Coelioxys hirsutissima</emphasis>
Cockerell, 1912
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7F9D10F50F84ABA5" authority="Cockerell, 1912" authorityName="Cockerell" authorityYear="1912" box="[242,710,1421,1443]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B931EA96CC20715E7F9D10F50CB5ABA5" box="[242,503,1421,1442]" italics="true" pageId="52" pageNumber="406">Coelioxys immaculata</emphasis>
Cockerell, 1912
</taxonomicName>
, and
<taxonomicName id="4C454D07CC20715E7FCD10D30F57ABC6" authority="Baker, 1975" authorityName="Baker" authorityYear="1975" box="[162,533,1451,1473]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="mitchelli">
<emphasis id="B931EA96CC20715E7FCD10D30C38ABC7" box="[162,378,1451,1472]" italics="true" pageId="52" pageNumber="406">Coelioxys mitchelli</emphasis>
<bibRefCitation id="EFD44B75CC20715E7EEA10D30F57ABC6" author="Baker JR" box="[389,533,1451,1473]" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
</taxonomicName>
, should be classified as
<emphasis id="B931EA96CC20715E7F8110B20C42ABD8" box="[238,256,1482,1503]" italics="true" pageId="52" pageNumber="406">C</emphasis>
. (
<emphasis id="B931EA96CC20715E7E7710B20CECABD8" box="[280,430,1482,1503]" italics="true" pageId="52" pageNumber="406">Xerocoelioxys</emphasis>
). This subgenus is predominantly Nearctic but with some species reaching the Neotropical region of
<collectingCountry id="F3527614CC20715E7D5D137F0FC4A81A" box="[562,646,1543,1565]" name="Mexico" pageId="52" pageNumber="406">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="8BFA3684CC20715E7FCD133D0C1BA949" blockId="52.[162,776,1604,1901]" pageId="52" pageNumber="406">
<emphasis id="B931EA96CC20715E7FCD133D0C26A85D" box="[162,356,1604,1626]" italics="true" pageId="52" pageNumber="406">List of species:</emphasis>
<taxonomicName id="4C454D07CC20715E7E1C133C0E46A85E" authority="Cresson, 1878" authorityName="Cresson" authorityYear="1878" box="[371,772,1604,1626]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="B931EA96CC20715E7E1C133C0F7EA85D" box="[371,572,1604,1626]" italics="true" pageId="52" pageNumber="406">Coelioxys aperta</emphasis>
Cresson, 1878
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7FCD131B0F69A87E" authority="Hill, 1936" authorityName="Hill" authorityYear="1936" box="[162,555,1635,1657]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="bisoncornua">
<emphasis id="B931EA96CC20715E7FCD131B0CE6A87F" box="[162,420,1635,1656]" italics="true" pageId="52" pageNumber="406">Coelioxys bisoncornua</emphasis>
Hill, 1936
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7D51131B0C0BA890" authority="Mitchell, 1962" authorityName="Mitchell" authorityYear="1962" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="boharti">
<emphasis id="B931EA96CC20715E7D51131B0E4AA87F" box="[574,776,1635,1656]" italics="true" pageId="52" pageNumber="406">Coelioxys boharti</emphasis>
Mitchell, 1962
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E3913FA0CF3A891" authorityName="Cresson" authorityYear="1872" box="[342,433,1665,1687]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="edita">
<emphasis id="B931EA96CC20715E7E3913FA0C2AA890" box="[342,360,1666,1687]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7E1713F90CF3A891" box="[376,433,1665,1686]" italics="true" pageId="52" pageNumber="406">edita</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7EAE13F90D99A8B1" authority="Mitchell, 1962" authorityName="Mitchell" authorityYear="1962" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="galactiae">
<emphasis id="B931EA96CC20715E7EAE13F90FDBA891" box="[449,665,1665,1686]" italics="true" pageId="52" pageNumber="406">Coelioxys galactiae</emphasis>
Mitchell, 1962
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E6913D80E46A8B2" authority="Cockerell, 1900" authorityName="Cockerell" authorityYear="1900" box="[262,772,1696,1718]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="grindeliae">
<emphasis id="B931EA96CC20715E7E6913D80F48A8B2" box="[262,522,1696,1717]" italics="true" pageId="52" pageNumber="406">Coelioxys grindeliae</emphasis>
Cockerell, 1900
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7FCD13C70C10A8D4" authorityName="Cockerell" authorityYear="1912" box="[162,338,1726,1748]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="hirsutissima">
<emphasis id="B931EA96CC20715E7FCD13C70C10A8D4" box="[162,338,1726,1748]" italics="true" pageId="52" pageNumber="406">C. hirsutissima</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E1B13C70F5CA8D4" authorityName="Cockerell" authorityYear="1912" box="[372,542,1726,1748]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B931EA96CC20715E7E1B13C70CC4A8D3" box="[372,390,1727,1748]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7EFA13C60F5CA8D4" box="[405,542,1726,1747]" italics="true" pageId="52" pageNumber="406">immaculata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7D2F13C60C15A8F5" authority="Cockerell, 1921" authorityName="Cockerell" authorityYear="1921" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="mesae">
<emphasis id="B931EA96CC20715E7D2F13C60E4AA8D3" box="[576,776,1726,1748]" italics="true" pageId="52" pageNumber="406">Coelioxys mesae</emphasis>
Cockerell, 1921
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E0813A50CAEA8F5" authorityName="Baker" authorityYear="1975" box="[359,492,1757,1778]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="mitchelli">
<emphasis id="B931EA96CC20715E7E0813A50C3BA8F5" box="[359,377,1757,1778]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7EE713A50CAEA8F5" box="[392,492,1757,1778]" italics="true" pageId="52" pageNumber="406">mitchelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E9013A50D99A916" authority="Baker, 1975" authorityName="Baker" authorityYear="1975" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="nodis">
<emphasis id="B931EA96CC20715E7E9013A50FF3A8F5" box="[511,689,1757,1778]" italics="true" pageId="52" pageNumber="406">Coelioxys nodis</emphasis>
<bibRefCitation id="EFD44B75CC20715E7DD313A50D99A916" author="Baker JR" pageId="52" pageNumber="406" pagination="649 - 730" refId="ref42640" refString="Baker JR. 1975. Taxonomy of five Neartic subgenera of Coelioxys (Hymenoptera: Megachilidae). The University of Kansas Science Bulletin 50: 649-730." type="journal article" year="1975">Baker, 1975</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E7F13840E46A916" authority="Crawford, 1914" authorityName="Crawford" authorityYear="1914" box="[272,772,1788,1810]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B931EA96CC20715E7E7F13840CB7A916" box="[272,501,1788,1809]" italics="true" pageId="52" pageNumber="406">Coelioxys piercei</emphasis>
Crawford, 1914
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7FCD12630C11A928" authorityName="Baker" authorityYear="1975" box="[162,339,1818,1840]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="serricaudata">
<emphasis id="B931EA96CC20715E7FCD12630DF6A937" box="[162,180,1819,1840]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7FAC12620C11A928" box="[195,339,1818,1839]" italics="true" pageId="52" pageNumber="406">serricaudata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D07CC20715E7E0A12630C82A937" box="[357,448,1819,1840]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="B931EA96CC20715E7E0A12630C35A937" box="[357,375,1819,1840]" italics="true" pageId="52" pageNumber="406">C</emphasis>
.
<emphasis id="B931EA96CC20715E7EE912630C82A937" box="[390,448,1819,1840]" italics="true" pageId="52" pageNumber="406">soror</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C454D07CC20715E7D6912620C17A949" authority="Cockerell, 1909" authorityName="Cockerell" authorityYear="1909" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="406" phylum="Arthropoda" rank="species" species="soledadensis">
<emphasis id="B931EA96CC20715E7D6912620E4AA928" box="[518,776,1818,1839]" italics="true" pageId="52" pageNumber="406">Coelioxys soledadensis</emphasis>
Cockerell, 1909
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA3684CC20715E7FD5122F0FF1A96A" blockId="52.[162,776,1604,1901]" box="[186,691,1879,1901]" pageId="52" pageNumber="406">Six new subgenera are described as follows.</paragraph>
<paragraph id="8BFA3684CC20715E7C3115AE083BAEEC" blockId="52.[833,1430,210,328]" box="[862,1401,210,237]" pageId="52" pageNumber="406">
<taxonomicName id="4C454D07CC20715E7C3115AE0990AEEA" box="[862,1234,210,237]" class="Insecta" family="Megachilidae" genus="Coelioxys" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="390" phylum="Arthropoda" rank="subGenus" subGenus="Austrocleptria">
<smallCapsWord id="8D1CA058CC20715E7C3115AE0EA6AEED" baselines="230" box="[862,996,214,235]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="oelioxys" pageId="52" pageNumber="406">
<emphasis id="B931EA96CC20715E7C3115AE0EA6AEED" bold="true" box="[862,996,214,235]" italics="true" pageId="52" pageNumber="406">COELIOXYS</emphasis>
</smallCapsWord>
<emphasis id="B931EA96CC20715E7C8215AA0EBAAEEA" bold="true" box="[1005,1016,210,237]" pageId="52" pageNumber="406">(</emphasis>
<smallCapsWord id="8D1CA058CC20715E7C9715AC0984AEED" baselines="230,230" box="[1016,1222,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Austrocleptria" pageId="52" pageNumber="406">
<emphasis id="B931EA96CC20715E7C9715AC0984AEED" bold="true" box="[1016,1222,212,236]" italics="true" pageId="52" pageNumber="406">AUSTROCLEPTRIA</emphasis>
</smallCapsWord>
<emphasis id="B931EA96CC20715E7BA815AA0990AEEA" bold="true" box="[1223,1234,210,237]" pageId="52" pageNumber="406">)</emphasis>
</taxonomicName>
<smallCapsWord id="8D1CA058CC20715E7BB415AC0871AEEA" baselines="230,231" box="[1243,1331,211,237]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Rocha" pageId="52" pageNumber="406">
<collectionCode id="ED54AE41CC20715E7BB415AC0871AEEA" box="[1243,1331,211,237]" pageId="52" pageNumber="406">ROCHA-</collectionCode>
</smallCapsWord>
<smallCapsWord id="8D1CA058CC20715E7A5C15AC083BAEEC" baselines="230,231" box="[1331,1401,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Filho" pageId="52" pageNumber="406">
<collectionCode id="ED54AE41CC20715E7A5C15AC083BAEEC" box="[1331,1401,212,236]" pageId="52" pageNumber="406">FILHO</collectionCode>
</smallCapsWord>
</paragraph>
</subSubSection>
</treatment>
</document>
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