287 lines
37 KiB
XML
287 lines
37 KiB
XML
<document id="0099E7E5D48EF4E605BAC631E84A4B59" ID-ISSN="0024-4082" ID-ZooBank="7882D0C-5833-4DB4-98D9-77600EEC22C3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738702329404" checkinUser="plazi" docAuthor="Puillandre, Nicolas, Fedosov, Alexander E., Zaharias, Paul, Aznar-Cormano, Laetitia & Kantor, Yuri I." docDate="2017" docId="03AD4855FFC8C03F900EF916FA90FCB0" docLanguage="en" docName="zlx012.pdf" docOrigin="Zoological Journal of the Linnean Society 181" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lophiotoma semfala Puillandre, Fedosov, Zaharias, Aznar-Cormano & Kantor, 2017, SP. NOV." docType="treatment" docVersion="1" lastPageNumber="258" masterDocId="FF94302DFFC5C0309114FF8EFFF3FFFB" masterDocTitle="A quest for the lost types of Lophiotoma (Gastropoda: Conoidea: Turridae): integrative taxonomy in a nomenclatural mess" masterLastPageNumber="271" masterPageNumber="243" pageNumber="256" updateTime="1738703087694" updateUser="GgImagineBatch">
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<mods:mods id="E5C3036743B0DE919A2410F81737F442" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2D93BEC861997BC69CC450C23F5CEE90">
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<mods:title id="6F2DD261132EBBFB2B5104E50DBEAEEC">A quest for the lost types of Lophiotoma (Gastropoda: Conoidea: Turridae): integrative taxonomy in a nomenclatural mess</mods:title>
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<mods:name id="B2F0CB59B47C4AB07D191E3A0B12281D" type="personal">
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<mods:namePart id="7D53AAB0EB50BF6BCB9EAB1CC250EE8A">Puillandre, Nicolas</mods:namePart>
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<mods:roleTerm id="15A87EC1FDA174FE30D441EA9494052E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ABA279AD9BFAB52297F3AA12E076EDEB">Fedosov, Alexander E.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="8485530089E4182AE43EC7F97E3F8545">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EDDF9575C936D1DC546D0668E01AD4CA">Zaharias, Paul</mods:namePart>
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</mods:name>
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<mods:roleTerm id="BB27671AB9AF64EC4B446E38D9ECBA8D">Author</mods:roleTerm>
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<mods:namePart id="0B9BE8CE6FD969878D05F76744B2A5DD">Aznar-Cormano, Laetitia</mods:namePart>
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</mods:name>
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<mods:name id="D62F6CF85020FA2A1B8F9EE1443AD2E0" type="personal">
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<mods:roleTerm id="909ECB3477F5CDE77333256015B04763">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E5FE70BC8BCCAF50C59F7053A17E3A4">Kantor, Yuri I.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8EEE532B5EC8E47B510DFA5968DA2A4D">text</mods:typeOfResource>
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<mods:title id="6CD418EE50F4DC8C7F77FAE2A661063A">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="6BAA6A9D4D1E2D5648A10E7CB7C55EE3">
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<mods:date id="44D57F1678FED5838237B012CC8A0AFA">2017</mods:date>
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<mods:detail id="264CE7EFBE539D0D2FE23A04B27F5B89" type="volume">
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<mods:number id="A2EBC288770EF9F8C6A8564C30C1422D">181</mods:number>
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<mods:start id="F0FEF2FE924D7E5E8B987E0BFF893B5E">243</mods:start>
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<treatment id="03AD4855FFC8C03F900EF916FA90FCB0" LSID="urn:lsid:plazi:treatment:03AD4855FFC8C03F900EF916FA90FCB0" httpUri="http://treatment.plazi.org/id/03AD4855FFC8C03F900EF916FA90FCB0" lastPageId="15" lastPageNumber="258" pageId="13" pageNumber="256">
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<subSubSection id="C31EAAC8FFC8C03D900EF916FD60F954" box="[282,659,1687,1711]" pageId="13" pageNumber="256" type="nomenclature">
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<paragraph id="8BBBF943FFC8C03D900EF916FD60F954" blockId="13.[282,659,1687,1711]" box="[282,659,1687,1711]" pageId="13" pageNumber="256">
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<heading id="D0F34E2FFFC8C03D900EF916FD60F954" box="[282,659,1687,1711]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="256" reason="2">
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<taxonomicName id="4C0482C0FFC8C03D900EF916FDDAF955" ID-CoL="3W2GJ" authorityName="Puillandre & Fedosov & Zaharias & Aznar-Cormano & Kantor" authorityYear="2017" box="[282,553,1688,1711]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="256" phylum="Mollusca" rank="species" species="semfala" status="sp. nov.">
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<smallCapsWord id="8D5D6F9FFFC8C03D900EF916FE4AF955" baselines="1705,1705" box="[282,441,1688,1711]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lophiotoma" pageId="13" pageNumber="256">LOPHIOTOMA</smallCapsWord>
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<smallCapsWord id="8D5D6F9FFFC8C03D90D4F915FDDAF955" baselines="1705" box="[448,553,1691,1710]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="semfala" pageId="13" pageNumber="256">SEMFALA</smallCapsWord>
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</taxonomicName>
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<emphasis id="B9702551FFC8C03D9325F915FD60F954" bold="true" box="[561,659,1687,1711]" pageId="13" pageNumber="256">
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<taxonomicNameLabel id="A243982AFFC8C03D9325F915FD60F954" box="[561,659,1687,1711]" pageId="13" pageNumber="256" rank="species">
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<smallCapsWord id="8D5D6F9FFFC8C03D9325F915FDBDF955" baselines="1705" box="[561,590,1691,1710]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="13" pageNumber="256">SP</smallCapsWord>
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.
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<smallCapsWord id="8D5D6F9FFFC8C03D934FF915FD7FF955" baselines="1705" box="[603,652,1691,1710]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="13" pageNumber="256">NOV</smallCapsWord>
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.
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</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EAAC8FFC8C03F90B8F930FA90FCB0" lastPageId="15" lastPageNumber="258" pageId="13" pageNumber="256" type="description">
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<paragraph id="8BBBF943FFC8C03D90B8F930FDF2F92C" blockId="13.[428,513,1726,1751]" box="[428,513,1726,1751]" pageId="13" pageNumber="256">
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(
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<figureCitation id="133FE5C6FFC8C03D90A1F930FE0AF92C" box="[437,505,1726,1751]" captionStart="Figure 7" captionStartId="14.[144,223,1384,1406]" captionTargetBox="[144,1429,195,1344]" captionTargetId="figure-217@14.[144,1429,195,1344]" captionTargetPageId="14" captionText="Figure 7. Lophiotoma semfala sp. nov. (A–C) Holotype, MNHN IM-2007-41337, SANTO 2006, st. LD35, SL 41.8 mm. (D) MNHN IM-2013-14504, PAPUA NIUGINI, st. PD41, SL 29.5 mm. (E) MNHN IM-2007-40830, Philippines, PANGLAO 2004, st. R62, SL 35.7 mm. (F–G) Dark form, MNHN IM-2013-4019, PAPUA NIUGINI, st. PD39, SL 12.4 mm (F – at the same scale as other shells, F′ enlarged). (G) Lateral view of the protoconch. (H) MNHN IM-2013-14965, PAPUA NIUGINI, st. PD45, SL 26.8 mm. All shells (except F′) at the same scale." pageId="13" pageNumber="256">
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<smallCapsWord id="8D5D6F9FFFC8C03D90A1F930FE2EF92D" baselines="1745,1745" box="[437,477,1726,1751]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Fig" pageId="13" pageNumber="256">FIG</smallCapsWord>
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. 7
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</figureCitation>
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)
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</paragraph>
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<paragraph id="8BBBF943FFC8C03D91B7F970FEA9F8C8" blockId="13.[163,779,1790,1843]" pageId="13" pageNumber="256">
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<uri id="FF95F541FFC8C03D91B7F970FEA9F8C8" pageId="13" pageNumber="256">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA2C396FFC8C03D90ADF970FEA9F8C8" pageId="13" pageNumber="256">9E586542-B03B-4133-93F7- 55B79FFD5A33</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BBBF943FFC8C03D91B7F8DBFAF5FEE2" blockId="13.[163,714,1876,1898]" lastBlockId="13.[827,1444,197,281]" pageId="13" pageNumber="256">
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<emphasis id="B9702551FFC8C03D91B7F8DBFEB4F892" box="[163,327,1876,1898]" italics="true" pageId="13" pageNumber="256">Type material:</emphasis>
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<materialsCitation id="3B6CF31EFFC8C03D905AF8DAFAF5FEE2" collectionCode="MNHN" collectorName="Aore I. Aimbuei Bay" country="Vanuatu" elevation="6" latitude="-15.546667" location="Aimbuei Bay" longLatPrecision="128" longitude="167.19333" pageId="13" pageNumber="256" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54BF47E1FFC8C03D905AF8DAFE47F891" box="[334,436,1876,1898]" pageId="13" pageNumber="256" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED156186FFC8C03D90AFF8DAFDFCF892" box="[443,527,1876,1897]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="256">MNHN</collectionCode>
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IM-2007-41337.
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<emphasis id="B9702551FFC8C03D922FFF49FC39FF20" box="[827,970,198,220]" italics="true" pageId="13" pageNumber="256">Type locality</emphasis>
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:
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<collectingCountry id="F313B9D3FFC8C03D92C1FF49FBCAFF27" box="[981,1081,199,220]" name="Vanuatu" pageId="13" pageNumber="256">Vanuatu</collectingCountry>
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,
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<collectorName id="26F19C95FFC8C03D9557FF48FAD4FF27" box="[1091,1319,198,220]" pageId="13" pageNumber="256">
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Aoré I.
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<location id="8EDBAF98FFC8C03D9581FF48FAD4FF27" LSID="urn:lsid:plazi:treatment:03AD4855FFC8C03F900EF916FA90FCB0:8EDBAF98FFC8C03D9581FF48FAD4FF27" box="[1173,1319,198,220]" country="Vanuatu" latitude="-15.546667" longLatPrecision="128" longitude="167.19333" name="Aimbuei Bay" pageId="13" pageNumber="256">Aimbuei Bay</location>
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</collectorName>
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,
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<geoCoordinate id="EE309F84FFC8C03D9426FF48FA6DFF20" box="[1330,1438,197,220]" degrees="15" direction="south" minutes="32.8" orientation="latitude" pageId="13" pageNumber="256" precision="92" value="-15.546667">15°32.8′S</geoCoordinate>
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,
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<geoCoordinate id="EE309F84FFC8C03D922FFF6BFC30FF01" box="[827,963,228,251]" degrees="167" direction="east" minutes="11.6" orientation="longitude" pageId="13" pageNumber="256" precision="92" value="167.19333">167°11.6′E</geoCoordinate>
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, white coral sand,
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<quantity id="4CFC54A6FFC8C03D95ADFF6BFAF5FF01" box="[1209,1286,229,250]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="13" pageNumber="256" unit="m" value="5.5" valueMax="8.0" valueMin="3.0">
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||
<elevation id="00291E70FFC8C03D95ADFF6BFAF5FF01" box="[1209,1286,229,250]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="13" pageNumber="256" unit="m" value="5.5" valueMax="8.0" valueMin="3.0">3–8 m</elevation>
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</quantity>
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(Expedition SANTO 2006, st. LD35, R/V
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<taxonomicName id="4C0482C0FFC8C03D956AFE8AFB58FEE2" box="[1150,1195,260,281]" class="Gastropoda" family="Muricidae" genus="Alis" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="256" phylum="Mollusca" rank="genus">
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||
<emphasis id="B9702551FFC8C03D956AFE8AFB58FEE2" box="[1150,1195,260,281]" italics="true" pageId="13" pageNumber="256">Alis</emphasis>
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</taxonomicName>
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annex).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BBBF943FFC8C03D922FFEC4FBFCFE47" blockId="13.[827,1443,330,444]" pageId="13" pageNumber="256">
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<emphasis id="B9702551FFC8C03D922FFEC4FC4FFEA4" box="[827,956,330,351]" italics="true" pageId="13" pageNumber="256">Etymology:</emphasis>
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semfala – the ‘same’ in Bislama, the creole language, one of the official languages of
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<collectingCountry id="F313B9D3FFC8C03D942EFEE7FA6CFE85" box="[1338,1439,361,382]" name="Vanuatu" pageId="13" pageNumber="256">Vanuatu</collectingCountry>
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. Used as noun in apposition to reflect the similarity to
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<taxonomicName id="4C0482C0FFC8C03D922FFE28FBFBFE47" box="[827,1032,422,444]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="256" phylum="Mollusca" rank="species" species="acuta">
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<emphasis id="B9702551FFC8C03D922FFE28FBFBFE47" box="[827,1032,422,444]" italics="true" pageId="13" pageNumber="256">Lophiotoma acuta</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BBBF943FFC8C03E922FFE60FD39F8F4" blockId="13.[827,1443,493,1895]" lastBlockId="14.[145,761,1570,1807]" lastPageId="14" lastPageNumber="257" pageId="13" pageNumber="256">
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||
<emphasis id="B9702551FFC8C03D922FFE60FB18FDF9" box="[827,1259,493,515]" italics="true" pageId="13" pageNumber="256">
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||
Description (
|
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<typeStatus id="54BF47E1FFC8C03D92CFFE63FBB1FDF9" box="[987,1090,493,514]" pageId="13" pageNumber="256" type="holotype">holotype</typeStatus>
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) (
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<figureCitation id="133FE5C6FFC8C03D9574FE60FB2AFDF9" box="[1120,1241,493,515]" captionStart="Figure 7" captionStartId="14.[144,223,1384,1406]" captionTargetBox="[144,1429,195,1344]" captionTargetId="figure-217@14.[144,1429,195,1344]" captionTargetPageId="14" captionText="Figure 7. Lophiotoma semfala sp. nov. (A–C) Holotype, MNHN IM-2007-41337, SANTO 2006, st. LD35, SL 41.8 mm. (D) MNHN IM-2013-14504, PAPUA NIUGINI, st. PD41, SL 29.5 mm. (E) MNHN IM-2007-40830, Philippines, PANGLAO 2004, st. R62, SL 35.7 mm. (F–G) Dark form, MNHN IM-2013-4019, PAPUA NIUGINI, st. PD39, SL 12.4 mm (F – at the same scale as other shells, F′ enlarged). (G) Lateral view of the protoconch. (H) MNHN IM-2013-14965, PAPUA NIUGINI, st. PD45, SL 26.8 mm. All shells (except F′) at the same scale." pageId="13" pageNumber="256">Fig. 7A–C</figureCitation>
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):
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</emphasis>
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Shell medium thick, narrow fusiform, spire high, siphonal canal long narrow, slightly inclined to left. Protoconch conical, eroded, rendering exact whorl count and sculpture examination doubtful, of about three evenly convex whorls. Protoconch diameter
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<quantity id="4CFC54A6FFC8C03D95F1FD08FABFFD67" box="[1253,1356,646,668]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="256" unit="mm" value="0.73">0.73 mm</quantity>
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, height
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<quantity id="4CFC54A6FFC8C03D922FFD2BFC55FD41" box="[827,934,677,699]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="256" unit="mm" value="0.85">0.85 mm</quantity>
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||
. Teleoconch whorls angulated at shoulder, ten in total. Suture very shallow, indistinct, subsutural region wide, distinctly concave, subsutural cord low, triangular in profile, with three angular ridges on last whorl, central one strongest. On upper teleoconch whorls, only central ridge persists. Subsutural region smooth on upper teleoconch whorls, with one spiral ridge appearing on fourth, two on sixth, three on seventh and five on last whorl. Paired sinus cords strongest forming angulated shoulder. On upper whorls both cords nearly equal in size, obtusely triangular, on penultimate and last whorls cords more angulate, although still rounded on top, only on last whorl upper cord distinctly stronger than lower. Base of spire whorls smooth on first four whorl, with one spiral cord on fifth to sixth whorls, starting from seventh whorl number of cords gradually increases, and penultimate whorl with seven narrow cords of slightly different size, median much stronger; interspaces three to four times broader than cords. Base of last whorl with three major spiral cords and several riblets between them, canal with 20 cords, becoming gradually broader, lower and more closely spaced anteriorly. Shell base gradually narrowing towards narrow and long nearly straight siphonal canal. Aperture pear shaped, outer lip concave in upper part and weakly convex below shoulder, gradually passing into canal. Anal sinus deep, with nearly parallel sides, with straight posterior margin, parallel to shell axis; outer lip in side-view rounded and opisthocline, stromboid notch well defined. Growth lines indistinct, closely spaced. Shell creamy, protoconch and three first teleoconch whorls very light brown. Subsutural cord with regularly spaced brown spots, not extending beyond cord, broader on last three whorls. Sinus cords with distinct dark brown regularly spaced spots occupying whole width of cord and separate on each cord, minor spiral cords with dense brown flecks. Aperture creamy inside. Measurements: SL
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<quantity id="4CFC54A6FFC8C03D9277F8BCFC39F8BC" box="[867,970,1842,1863]" metricMagnitude="-2" metricUnit="m" metricValue="4.18" pageId="13" pageNumber="256" unit="mm" value="41.8">41.8 mm</quantity>
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, AL (with canal)
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<quantity id="4CFC54A6FFC8C03D9580F8BCFB08F8BC" box="[1172,1275,1842,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="13" pageNumber="256" unit="mm" value="20.9">20.9 mm</quantity>
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, SW
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<quantity id="4CFC54A6FFC8C03D9423F8BCFA6CF8BC" box="[1335,1439,1842,1864]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="13" pageNumber="256" unit="mm" value="10.6">10.6 mm</quantity>
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. Radula examined in
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<specimenCount id="9D0232CAFFC8C03D953CF8DFFB11F89C" box="[1064,1250,1873,1895]" count="3" pageId="13" pageNumber="256" type="generic">three specimens</specimenCount>
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||
, two from
|
||
<collectingCountry id="F313B9D3FFC8C03E944EF8DFFED3F9C3" lastPageId="14" lastPageNumber="257" name="Papua New Guinea" pageId="13" pageNumber="256">Papua New Guinea</collectingCountry>
|
||
and one from the
|
||
<collectingCountry id="F313B9D3FFCBC03E90E1F9ACFD89F9C3" box="[501,634,1570,1592]" name="Philippines" pageId="14" pageNumber="257">Philippines</collectingCountry>
|
||
, very similar in all examined specimens (
|
||
<figureCitation id="133FE5C6FFCBC03E90E1F9CFFDBAF9AD" box="[501,585,1600,1623]" captionStart="Figure 5" captionStartId="11.[163,243,1392,1414]" captionTargetBox="[259,1347,195,1352]" captionTargetId="figure-202@11.[259,1347,195,1352]" captionTargetPageId="11" captionText="Figure 5. Radulae of studied Lophiotoma. (A–B) Lophiotoma acuta (Perry, 1811). (A) MNHN IM-2013-14235, PAPUA NIUGINI st. PD33, SL 32.1 mm. (B) MNHN IM-2013-14505, PAPUA NIUGINI st. PD41, SL 21.6. (C) Lophiotoma polytropa (Helbling, 1779), MNHN uncatalogued, PANGLAO 2004, st. M50. (D) Lophiotoma brevicaudata (Reeve, 1843), MNHN IM-2007-40994, SANTO 2006, st. DB12, SL 16.8 mm. (E) Lophiotoma picturata (Weinkauff, 1876), MNHN IM-2013-53422. (F) Lophiotoma bratasusa sp. nov., holotype. (G) Lophiotoma jickelii (Weinkauff, 1875), neotype. (H) Lophiotoma semfala sp. nov., MNHN IM-2013-14504. (I) Lophiotoma kina sp. nov., holotype. Scale bars 50 µm." pageId="14" pageNumber="257">Fig. 5H</figureCitation>
|
||
). Radula membrane medium long, of 33–50 rows of teeth of which 9–16 not fully formed. Marginal teeth duplex. Anterior (inner) half solid, narrowly lanceolate, dorso-ventrally compressed with sharp lateral cutting edges. In posterior half major and accessory limbs rather thin, bifurcate at about 45° angle. Central formation absent.
|
||
</paragraph>
|
||
<caption id="DF7BA9CBFFCBC03E9184FAE6FCE6FA08" pageId="14" pageNumber="257" startId="14.[144,223,1384,1406]" targetBox="[144,1429,195,1344]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BBBF943FFCBC03E9184FAE6FCE6FA08" blockId="14.[144,1425,1383,1523]" pageId="14" pageNumber="257">
|
||
<emphasis id="B9702551FFCBC03E9184FAE6FDA7FA86" bold="true" box="[144,596,1383,1406]" pageId="14" pageNumber="257">
|
||
Figure 7.
|
||
<taxonomicName id="4C0482C0FFCBC03E9011FAE9FE05FA86" authorityName="Puillandre & Fedosov & Zaharias & Aznar-Cormano & Kantor" authorityYear="2017" box="[261,502,1383,1405]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="257" phylum="Mollusca" rank="species" species="semfala" status="sp. nov.">
|
||
<emphasis id="B9702551FFCBC03E9011FAE9FE05FA86" bold="true" box="[261,502,1383,1405]" italics="true" pageId="14" pageNumber="257">Lophiotoma semfala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A243982AFFCBC03E90E9FAE6FDA7FA86" box="[509,596,1384,1405]" pageId="14" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A–C) Holotype, MNHN IM-2007-41337, SANTO 2006, st. LD35, SL 41.8 mm. (D) MNHN IM-2013-14504, PAPUA NIUGINI, st. PD41, SL 29.5 mm. (E) MNHN IM-2007-40830, Philippines, PANGLAO 2004, st. R62, SL 35.7 mm. (F–G) Dark form, MNHN IM-2013-4019, PAPUA NIUGINI, st. PD39, SL 12.4 mm (F – at the same scale as other shells, F′ enlarged). (G) Lateral view of the protoconch. (H) MNHN IM-2013-14965, PAPUA NIUGINI, st. PD45, SL 26.8 mm. All shells (except F′) at the same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBBF943FFCBC03F9185F8B9FDA9F993" blockId="14.[145,761,1847,1900]" lastBlockId="15.[163,779,791,1640]" lastPageId="15" lastPageNumber="258" pageId="14" pageNumber="257">
|
||
<emphasis id="B9702551FFCBC03E9185F8B9FF0BF8B7" box="[145,248,1847,1868]" italics="true" pageId="14" pageNumber="257">Remarks</emphasis>
|
||
: The new species is represented only by
|
||
<specimenCount id="9D0232CAFFCBC03E93CCF8B9FEE3F897" count="6" pageId="14" pageNumber="257" type="generic">six specimens</specimenCount>
|
||
, including the
|
||
<typeStatus id="54BF47E1FFCBC03E90DEF8D8FDC2F897" box="[458,561,1878,1900]" pageId="14" pageNumber="257" type="holotype">holotype</typeStatus>
|
||
and despite the limited material, two rather distinct forms can be recognized. The ‘light’ form that includes the
|
||
<typeStatus id="54BF47E1FFCBC03E9442F9CFFCAEF98E" pageId="14" pageNumber="257" type="holotype">holotype</typeStatus>
|
||
has fewer brown spots and the base colour is uniformly creamy. The brown spots on the subsutural cord are in most specimens confined to the cord itself and do not extend beyond, but in the
|
||
<typeStatus id="54BF47E1FFCBC03E9438F935FA63F92A" box="[1324,1424,1723,1745]" pageId="14" pageNumber="257" type="holotype">holotype</typeStatus>
|
||
on some whorls there are brownish blurred extensions of the spots to the subsutural region. Available specimens other than the
|
||
<typeStatus id="54BF47E1FFCBC03E9570F899FB39F8D6" box="[1124,1226,1815,1837]" pageId="14" pageNumber="257" type="holotype">holotype</typeStatus>
|
||
are smaller and less speckled. The ‘dark’ form is represented by
|
||
<specimenCount id="9D0232CAFFCBC03E9473F8B8FC52F890" count="2" pageId="14" pageNumber="257" type="generic">two specimens</specimenCount>
|
||
only, one being juvenile (
|
||
<figureCitation id="133FE5C6FFCBC03E95D1F8DBFAC4F891" box="[1221,1335,1877,1899]" captionStart="Figure 7" captionStartId="14.[144,223,1384,1406]" captionTargetBox="[144,1429,195,1344]" captionTargetId="figure-217@14.[144,1429,195,1344]" captionTargetPageId="14" captionText="Figure 7. Lophiotoma semfala sp. nov. (A–C) Holotype, MNHN IM-2007-41337, SANTO 2006, st. LD35, SL 41.8 mm. (D) MNHN IM-2013-14504, PAPUA NIUGINI, st. PD41, SL 29.5 mm. (E) MNHN IM-2007-40830, Philippines, PANGLAO 2004, st. R62, SL 35.7 mm. (F–G) Dark form, MNHN IM-2013-4019, PAPUA NIUGINI, st. PD39, SL 12.4 mm (F – at the same scale as other shells, F′ enlarged). (G) Lateral view of the protoconch. (H) MNHN IM-2013-14965, PAPUA NIUGINI, st. PD45, SL 26.8 mm. All shells (except F′) at the same scale." pageId="14" pageNumber="257">Fig. 7F–G</figureCitation>
|
||
). It has slightly darker base colour, with a light brown shell base and canal and with the subsutural region below the subsutural cord uniformly brown. There was no correlation between geographic distributions, since
|
||
<specimenCount id="9D0232CAFFCAC03F91B7FC1CFEB7FC5C" box="[163,324,913,935]" count="1" pageId="15" pageNumber="258" type="generic">one specimen</specimenCount>
|
||
of the dark form was collected in the
|
||
<collectingCountry id="F313B9D3FFCAC03F91B7FC3EFEDFFC3D" box="[163,300,944,966]" name="Philippines" pageId="15" pageNumber="258">Philippines</collectingCountry>
|
||
, while another in
|
||
<collectingCountry id="F313B9D3FFCAC03F9311FC3EFD18FC3D" box="[517,747,944,966]" name="Papua New Guinea" pageId="15" pageNumber="258">Papua New Guinea</collectingCountry>
|
||
at similar depths. The sinus cords of the adult specimen of the dark form are also sharper on top on the last whorl. On most parts of the teleoconch whorls, the sinus cords are either similar in size, or the lower even slightly more pronounced, than the upper, but on the last whorl the situation is reversed. An intact protoconch persists only in the juvenile of the dark form (
|
||
<figureCitation id="133FE5C6FFCAC03F91F9FB2BFEB6FB41" box="[237,325,1189,1211]" captionStart="Figure 7" captionStartId="14.[144,223,1384,1406]" captionTargetBox="[144,1429,195,1344]" captionTargetId="figure-217@14.[144,1429,195,1344]" captionTargetPageId="14" captionText="Figure 7. Lophiotoma semfala sp. nov. (A–C) Holotype, MNHN IM-2007-41337, SANTO 2006, st. LD35, SL 41.8 mm. (D) MNHN IM-2013-14504, PAPUA NIUGINI, st. PD41, SL 29.5 mm. (E) MNHN IM-2007-40830, Philippines, PANGLAO 2004, st. R62, SL 35.7 mm. (F–G) Dark form, MNHN IM-2013-4019, PAPUA NIUGINI, st. PD39, SL 12.4 mm (F – at the same scale as other shells, F′ enlarged). (G) Lateral view of the protoconch. (H) MNHN IM-2013-14965, PAPUA NIUGINI, st. PD45, SL 26.8 mm. All shells (except F′) at the same scale." pageId="15" pageNumber="258">Fig. 7G</figureCitation>
|
||
), it consists of 2.75 whorls, diameter
|
||
<quantity id="4CFC54A6FFCAC03F91B7FB4AFEF8FB22" box="[163,267,1220,1242]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="15" pageNumber="258" unit="mm" value="0.68">0.68 mm</quantity>
|
||
, height 0.73, which is significantly smaller than in
|
||
<typeStatus id="54BF47E1FFCAC03F9010FB6CFE9FFB03" box="[260,364,1250,1272]" pageId="15" pageNumber="258" type="holotype">holotype</typeStatus>
|
||
, although the existing material is insufficient for estimates of variation. The species is extremely similar to
|
||
<taxonomicName id="4C0482C0FFCAC03F90B6FAAEFD8FFACD" box="[418,636,1312,1334]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="acuta">
|
||
<emphasis id="B9702551FFCAC03F90B6FAAEFD8FFACD" box="[418,636,1312,1334]" italics="true" pageId="15" pageNumber="258">Lophiotoma acuta</emphasis>
|
||
</taxonomicName>
|
||
, which also has dark and light forms. It can be distinguished in most cases by being less pronounced and more rounded on the top sinus cords, providing a less angulated appearance to the shell shoulder, as well as the cords being more similar in size (
|
||
<figureCitation id="133FE5C6FFCAC03F9330FA37FD97FA34" box="[548,612,1465,1487]" captionStart="Figure 6" captionStartId="12.[145,224,1297,1319]" captionTargetBox="[353,1217,195,1257]" captionTargetId="figure-265@12.[353,1217,195,1257]" captionTargetPageId="12" captionText="Figure 6. Anal sinus and spiral sculpture of different species of Lophiotoma. (A) Lophiotoma acuta (Perry, 1811), MNHN IM-2007-41179. (B) Lophiotoma acuta, MNHN IM-2009-29711, SL 50.9 mm. (C) Lophiotoma semfala sp. nov., holotype, MNHN IM-2007-41337, SL 41.8 mm. (D) Lophiotoma semfala sp. nov., dark form, MNHN IM-2007-40830, SL 35.7 mm. (E) Lophiotoma kina sp. nov., holotype, MNHN IM-2013-16307, SL 31.0 mm. (F) Lophiotoma jickelii (Weinkauff, 1875), neotype, MNHN IM-2013-13275, SL 39.4 mm. Arrows indicate diagnostic details of the sculpture." pageId="15" pageNumber="258">Fig. 6</figureCitation>
|
||
, compare A, B with C, D), and domination of the lower cord over the higher one on the teleoconch whorls. The protoconch of
|
||
<taxonomicName id="4C0482C0FFCAC03F91D4F99BFE66F9D0" box="[192,405,1557,1579]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="acuta">
|
||
<emphasis id="B9702551FFCAC03F91D4F99BFE66F9D0" box="[192,405,1557,1579]" italics="true" pageId="15" pageNumber="258">Lophiotoma acuta</emphasis>
|
||
</taxonomicName>
|
||
is slightly larger (
|
||
<figureCitation id="133FE5C6FFCAC03F9366F99BFD47F9D1" box="[626,692,1557,1579]" captionStart="Figure 8" captionStartId="15.[163,244,639,661]" captionTargetBox="[165,1439,198,597]" captionTargetId="figure-585@15.[163,1442,196,600]" captionTargetPageId="15" captionText="Figure 8. Scatterplot of protoconch measurements in studied species of Lophiotoma. D: protoconch diameter, mm; H: exposed height, mm. (A) Diamonds: L. semfala sp. nov.; squares: L. acuta; triangles: L. kina sp. nov.; circles: L. jickelii. (B) Diamonds: L. bratasusa sp. nov.; squares: L. picturata." pageId="15" pageNumber="258">Fig. 8</figureCitation>
|
||
), while the radula is longer (consists of 55–80 rows of teeth vs. 33–50 rows in
|
||
<taxonomicName id="4C0482C0FFCAC03F9060F9DDFE1EF99C" authorityName="Puillandre & Fedosov & Zaharias & Aznar-Cormano & Kantor" authorityYear="2017" box="[372,493,1618,1640]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="semfala" status="sp. nov.">
|
||
<emphasis id="B9702551FFCAC03F9060F9DDFE1EF99C" box="[372,493,1618,1640]" italics="true" pageId="15" pageNumber="258">L. semfala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A243982AFFCAC03F90E1F9DDFDBDF993" box="[501,590,1619,1640]" pageId="15" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7BA9CBFFCAC03F91B7FDF1FCE4FD2B" pageId="15" pageNumber="258" startId="15.[163,244,639,661]" targetBox="[165,1439,198,597]" targetPageId="15" targetType="figure">
|
||
<paragraph id="8BBBF943FFCAC03F91B7FDF1FCE4FD2B" blockId="15.[163,1443,639,720]" pageId="15" pageNumber="258">
|
||
<emphasis id="B9702551FFCAC03F91B7FDF1FEFCFD6F" bold="true" box="[163,271,639,661]" pageId="15" pageNumber="258">Figure 8.</emphasis>
|
||
Scatterplot of protoconch measurements in studied species of
|
||
<emphasis id="B9702551FFCAC03F92A8FDF1FBB7FD6E" box="[956,1092,639,661]" italics="true" pageId="15" pageNumber="258">
|
||
<taxonomicName id="4C0482C0FFCAC03F92A8FDF1FBB3FD6E" authorityName="CASEY" authorityYear="1904" box="[956,1088,639,661]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="genus">Lophiotoma</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
D: protoconch diameter, mm; H: exposed height, mm. (A) Diamonds:
|
||
<emphasis id="B9702551FFCAC03F930BFD13FD04FD49" bold="true" box="[543,759,668,690]" pageId="15" pageNumber="258">
|
||
<taxonomicName id="4C0482C0FFCAC03F930BFD13FD6AFD49" authorityName="Puillandre & Fedosov & Zaharias & Aznar-Cormano & Kantor" authorityYear="2017" box="[543,665,668,690]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="semfala" status="sp. nov.">
|
||
<emphasis id="B9702551FFCAC03F930BFD13FD6AFD49" bold="true" box="[543,665,668,690]" italics="true" pageId="15" pageNumber="258">L. semfala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A243982AFFCAC03F93B4FD13FD04FD49" box="[672,759,669,690]" pageId="15" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; squares:
|
||
<taxonomicName id="4C0482C0FFCAC03F9276FD13FC44FD49" box="[866,951,669,690]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="acuta">
|
||
<emphasis id="B9702551FFCAC03F9276FD13FC44FD49" box="[866,951,669,690]" italics="true" pageId="15" pageNumber="258">L. acuta</emphasis>
|
||
</taxonomicName>
|
||
; triangles:
|
||
<emphasis id="B9702551FFCAC03F953AFD13FB13FD49" bold="true" box="[1070,1248,668,690]" pageId="15" pageNumber="258">
|
||
<taxonomicName id="4C0482C0FFCAC03F953AFD13FB72FD49" authorityName="Puillandre & Fedosov & Zaharias & Aznar-Cormano & Kantor" authorityYear="2017" box="[1070,1153,668,690]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="kina" status="sp. nov.">
|
||
<emphasis id="B9702551FFCAC03F953AFD13FB72FD49" bold="true" box="[1070,1153,668,690]" italics="true" pageId="15" pageNumber="258">L. kina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A243982AFFCAC03F959DFD13FB13FD49" box="[1161,1248,669,690]" pageId="15" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; circles:
|
||
<taxonomicName id="4C0482C0FFCAC03F942FFD13FA6EFD49" authorityName="Weinkauff" authorityYear="1875" box="[1339,1437,668,690]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="jickelii">
|
||
<emphasis id="B9702551FFCAC03F942FFD13FA6EFD49" box="[1339,1437,668,690]" italics="true" pageId="15" pageNumber="258">L. jickelii</emphasis>
|
||
</taxonomicName>
|
||
. (B) Diamonds:
|
||
<emphasis id="B9702551FFCAC03F902AFD34FDDEFD34" bold="true" box="[318,557,697,719]" pageId="15" pageNumber="258">
|
||
<taxonomicName id="4C0482C0FFCAC03F902AFD34FE23FD34" authorityName="Puillandre & Fedosov & Zaharias & Aznar-Cormano & Kantor" authorityYear="2017" box="[318,464,697,719]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="bratasusa" status="sp. nov.">
|
||
<emphasis id="B9702551FFCAC03F902AFD34FE23FD34" bold="true" box="[318,464,697,719]" italics="true" pageId="15" pageNumber="258">L. bratasusa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A243982AFFCAC03F90C3FD34FDDEFD34" box="[471,557,698,719]" pageId="15" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; squares:
|
||
<taxonomicName id="4C0482C0FFCAC03F9382FD34FCE1FD2B" baseAuthorityName="Weinkauff" baseAuthorityYear="1876" box="[662,786,698,720]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="picturata">
|
||
<emphasis id="B9702551FFCAC03F9382FD34FCE1FD2B" box="[662,786,698,720]" italics="true" pageId="15" pageNumber="258">L. picturata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBBF943FFCAC03F91B7F917FA90FCB0" blockId="15.[163,779,1689,1895]" lastBlockId="15.[827,1442,790,843]" pageId="15" pageNumber="258">
|
||
<emphasis id="B9702551FFCAC03F91B7F917FEDDF955" box="[163,302,1689,1710]" italics="true" pageId="15" pageNumber="258">Distribution</emphasis>
|
||
: The species was found in the
|
||
<collectingCountry id="F313B9D3FFCAC03F9390F917FCF4F954" box="[644,775,1689,1711]" name="Philippines" pageId="15" pageNumber="258">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="F313B9D3FFCAC03F91B7F936FE75F935" box="[163,390,1720,1742]" name="Papua New Guinea" pageId="15" pageNumber="258">Papua New Guinea</collectingCountry>
|
||
and
|
||
<collectingCountry id="F313B9D3FFCAC03F90D6F936FDDAF936" box="[450,553,1720,1741]" name="Vanuatu" pageId="15" pageNumber="258">Vanuatu</collectingCountry>
|
||
. In all these localities, it is sympatric with
|
||
<emphasis id="B9702551FFCAC03F90ACF959FDE8F917" box="[440,539,1751,1772]" italics="true" pageId="15" pageNumber="258">
|
||
<taxonomicName id="4C0482C0FFCAC03F90ACF959FDE5F917" box="[440,534,1751,1772]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="acuta">L. acuta</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Judging from available material (only six sequenced specimens), it is much more rare than
|
||
<taxonomicName id="4C0482C0FFCAC03F904EF89AFE4BF8D2" box="[346,440,1812,1833]" class="Gastropoda" family="Turridae" genus="Lophiotoma" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="258" phylum="Mollusca" rank="species" species="acuta">
|
||
<emphasis id="B9702551FFCAC03F904EF89AFE4BF8D2" box="[346,440,1812,1833]" italics="true" pageId="15" pageNumber="258">L. acuta</emphasis>
|
||
</taxonomicName>
|
||
, for which we had sequenced more than
|
||
<specimenCount id="9D0232CAFFCAC03F9020F8BCFE02F8B3" box="[308,497,1842,1864]" count="160" pageId="15" pageNumber="258" type="generic">160 specimens</specimenCount>
|
||
. Although we did not sequence any specimens from
|
||
<collectingCountry id="F313B9D3FFCAC03F90EFF8DCFD59F89C" box="[507,682,1873,1895]" name="New Caledonia" pageId="15" pageNumber="258">New Caledonia</collectingCountry>
|
||
, judging from the shell characters the species is also found in
|
||
<collectingCountry id="F313B9D3FFCAC03F922FFCB8FC1BFCB0" box="[827,1000,821,843]" name="New Caledonia" pageId="15" pageNumber="258">New Caledonia</collectingCountry>
|
||
(uncatalogued MNHN material).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |